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<document id="6D5E8CD6935C558C83425DD911B825DD" ID-DOI="10.5281/zenodo.188392" ID-GBIF-Dataset="4683115e-4971-4b8f-bb75-1a3f9b104521" ID-ISSN="1175-5326" ID-Zenodo-Dep="188392" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461090263440" checkinUser="plazi" docAuthor="Pitts, James P., Wilson, Joseph S., Williams, Kevin A. &amp; Boehme, Nicole F." docDate="2009" docId="03933237FFDF980C039EFEFE5756FB16" docLanguage="en" docName="zt02131p053.pdf" docOrigin="Zootaxa 2131" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Dasymutilla imperialis Manley and Pitts" docType="treatment" docVersion="9" lastPageNumber="18" masterDocId="FFAA4A4FFFCE981D0309FFCC5617FFA2" masterDocTitle="Velvet ants (Hymenoptera: Mutillidae) of the Algodones sand dunes of California, USA" masterLastPageNumber="53" masterPageNumber="1" pageNumber="18" updateTime="1698588327638" updateUser="plazi">
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<mods:title id="6A2C0E993699E85C384B52229BF39907">Velvet ants (Hymenoptera: Mutillidae) of the Algodones sand dunes of California, USA</mods:title>
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<mods:namePart id="17549FB3DC60E6F410749C133AF7A267">Pitts, James P.</mods:namePart>
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<mods:namePart id="6DFB43F2C815361E78E63347F3DC175B">Boehme, Nicole F.</mods:namePart>
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<subSubSection id="C320D0AAFFDF980C039EFEFE5491FEEE" box="[151,646,306,332]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8B858321FFDF980C039EFEFE5491FEEE" blockId="17.[151,646,306,332]" box="[151,646,306,332]" pageId="17" pageNumber="18">
<heading id="D0CD344DFFDF980C039EFEFE5491FEEE" bold="true" box="[151,646,306,332]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<taxonomicName id="4C3AF8A2FFDF980C039EFEFE5491FEEE" authority="Manley and Pitts" authorityName="Manley and Pitts" box="[151,646,306,332]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="imperialis">
<emphasis id="B94E5F33FFDF980C039EFEFE5491FEEE" bold="true" box="[151,646,306,332]" pageId="17" pageNumber="18">
<emphasis id="B94E5F33FFDF980C039EFEFE57BEFEEE" bold="true" box="[151,425,306,332]" italics="true" pageId="17" pageNumber="18">Dasymutilla imperialis</emphasis>
Manley and Pitts
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C320D0AAFFDF980C039EFEB055FDFE11" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="8B858321FFDF980C039EFEB055FDFE11" blockId="17.[151,1436,380,435]" pageId="17" pageNumber="18">
<taxonomicName id="4C3AF8A2FFDF980C039EFEB0548AFE31" authority="Manley and Pitts, 2004" authorityName="Manley and Pitts" authorityYear="2004" box="[151,669,380,403]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="imperialis">
<emphasis id="B94E5F33FFDF980C039EFEB05798FE31" box="[151,399,380,403]" italics="true" pageId="17" pageNumber="18">Dasymutilla imperialis</emphasis>
<bibRefCitation id="EFABFED0FFDF980C0290FEB0548AFE31" author="Manley" box="[409,669,380,403]" pageId="17" pageNumber="52" refString="Manley, D. G. &amp; Pitts, J. P. (2004) Two new black species of Dasymutilla (Hymenoptera: Mutillidae) from Texas and California. Journal of the Kansas Entomological Societ y, 77, 644 - 649." type="journal article" year="2004">Manley and Pitts, 2004</bibRefCitation>
</taxonomicName>
. J. Kan. Ent. Soc. 77: 646. Male.
<typeStatus id="54813D83FFDF980C0713FEB0526AFE31" box="[1050,1149,380,403]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
data: California,
<taxonomicName id="4C3AF8A2FFDF980C064BFEB057ACFE11" authority="County, Glamis Dunes" authorityName="County, Glamis Dunes" class="Anthozoa" family="Aurelianidae" genus="Imperial" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actiniaria" pageId="17" pageNumber="18" phylum="Cnidaria" rank="genus">Imperial County, Glamis Dunes</taxonomicName>
,
<quantity id="4CC22EC4FFDF980C02CFFE5057EDFE11" box="[454,506,412,435]" metricMagnitude="3" metricUnit="m" metricValue="1.609344" pageId="17" pageNumber="18" unit="mi" value="1.0">1 mi</quantity>
. W Glamis,
<date id="FF84A5E1FFDF980C0177FE5054E0FE11" box="[638,759,412,435]" pageId="17" pageNumber="18" value="1988-10-07">7 Oct 1988</date>
, T. Griswold (EMUS).
</paragraph>
</subSubSection>
<subSubSection id="C320D0AAFFDF980C039EFE2E57C8FDD2" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="8B858321FFDF980C039EFE2E57C8FDD2" blockId="17.[151,1437,482,1204]" pageId="17" pageNumber="18">
<emphasis id="B94E5F33FFDF980C039EFE2E5764FE59" box="[151,371,482,507]" italics="true" pageId="17" pageNumber="18">Diagnosis of male.</emphasis>
This species (
<figureCitation id="13019FA4FFDF980C0113FE2E5478FE5E" box="[538,623,482,508]" captionStart="FIGURES 47 55" captionStartId="11.[151,269,1657,1681]" captionTargetBox="[153,1424,450,1656]" captionTargetId="figure@11.[151,1436,411,1668]" captionTargetPageId="11" captionText="FIGURES 47 55. Habitus, lateral view. Dasymutilla atricuada, 47. female and 48. male; D. gloriosa, 49. female and 50. male; D. imperialis, 51. male; D. magnifica, 52. female and 53. male; and D. nocturna, 54. female and 55. male." httpUri="https://zenodo.org/record/188397/files/figure.png" pageId="17" pageNumber="18">Fig. 51</figureCitation>
) has the following combination of characters: the integument and the setae are entirely black, the anterior margin of the pronotum is emarginate and glabrous medially, and S2 has an oval pit filled with setae positioned just posterior to the center of the sternite. The genitalia were illustrated by
<bibRefCitation id="EFABFED0FFDF980C03B5FD9A57CBFDD2" author="Manley" box="[188,476,598,624]" pageId="17" pageNumber="52" refString="Manley, D. G. &amp; Pitts, J. P. (2004) Two new black species of Dasymutilla (Hymenoptera: Mutillidae) from Texas and California. Journal of the Kansas Entomological Societ y, 77, 644 - 649." type="journal article" year="2004">Manley and Pitts (2004)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C320D0AAFFDF980C03CCFDB157B0FD35" box="[197,423,637,663]" pageId="17" pageNumber="18" type="description">
<paragraph id="8B858321FFDF980C03CCFDB157B0FD35" blockId="17.[151,1437,482,1204]" box="[197,423,637,663]" pageId="17" pageNumber="18">
<emphasis id="B94E5F33FFDF980C03CCFDB15733FD34" box="[197,292,637,662]" italics="true" pageId="17" pageNumber="18">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C320D0AAFFDF980C03CCFD6854B7FCA9" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="8B858321FFDF980C03CCFD6854B7FCA9" blockId="17.[151,1437,482,1204]" pageId="17" pageNumber="18">
<emphasis id="B94E5F33FFDF980C03CCFD6857AEFD1F" box="[197,441,676,701]" italics="true" pageId="17" pageNumber="18">Material examined.</emphasis>
<emphasis id="B94E5F33FFDF980C02CBFD6F545DFD1F" bold="true" box="[450,586,675,701]" pageId="17" pageNumber="18">California</emphasis>
,
<taxonomicName id="4C3AF8A2FFDF980C0152FD6852A4FD1C" authority="Co., Algodones Dunes, Niland-Glamis" authorityName="Co., Algodones Dunes, Niland-Glamis" box="[603,1203,676,702]" class="Anthozoa" family="Aurelianidae" genus="Imperial" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actiniaria" pageId="17" pageNumber="18" phylum="Cnidaria" rank="genus">
<emphasis id="B94E5F33FFDF980C0152FD6854ECFD1F" box="[603,763,676,701]" italics="true" pageId="17" pageNumber="18">Imperial Co.</emphasis>
, Algodones Dunes, Niland-Glamis
</taxonomicName>
Road,
<quantity id="4CC22EC4FFDF980C0605FD685374FD1C" box="[1292,1379,676,702]" metricMagnitude="3" metricUnit="m" metricValue="7.4" pageId="17" pageNumber="18" unit="km" value="7.4">7.4 km</quantity>
NW Glamis,
<specimenCount id="9D3C48A8FFDF980C03F2FD06575AFD46" box="[251,333,714,740]" pageId="17" pageNumber="18" type="male">1 male</specimenCount>
,
<date id="FF84A5E1FFDF980C0250FD065445FD46" box="[345,594,714,740]" pageId="17" pageNumber="18" value="2008-09-22" valueMax="2008-11-15" valueMin="2008-09-22">22.Sep15.Nov.2008</date>
, E. Dreyfus (
<collectionCode id="ED2B1BE4FFDF980C01E4FD065552FD46" box="[749,837,714,740]" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="17" pageNumber="18">UCDC</collectionCode>
); Glamis Dunes, Gecko Campground, SW Glamis,
<specimenCount id="9D3C48A8FFDF980C039EFD3D56E3FCA9" box="[151,244,753,779]" pageId="17" pageNumber="18" type="male">4 males</specimenCount>
,
<date id="FF84A5E1FFDF980C0209FD3D5783FCA9" box="[256,404,753,779]" pageId="17" pageNumber="18" value="1987-09-28">28.Sep.1987</date>
, T. Griswold (
<collectionCode id="ED2B1BE4FFDF980C0135FD3D5483FCA9" box="[572,660,753,779]" pageId="17" pageNumber="18">EMUS</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C320D0AAFFDF980C03CCFCD45469FC90" box="[197,638,792,818]" pageId="17" pageNumber="18" type="distribution">
<paragraph id="8B858321FFDF980C03CCFCD45469FC90" blockId="17.[151,1437,482,1204]" box="[197,638,792,818]" pageId="17" pageNumber="18">
<emphasis id="B94E5F33FFDF980C03CCFCD4574BFC93" box="[197,348,792,817]" italics="true" pageId="17" pageNumber="18">Distribution.</emphasis>
Algodones Sand Dunes.
</paragraph>
</subSubSection>
<subSubSection id="C320D0AAFFDF980C03CCFCF35756FB16" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8B858321FFDF980C03CCFCF35351FBE2" blockId="17.[151,1437,482,1204]" pageId="17" pageNumber="18">
<emphasis id="B94E5F33FFDF980C03CCFCF35723FCFA" box="[197,308,831,856]" italics="true" pageId="17" pageNumber="18">Remarks.</emphasis>
Only seven specimens have been recorded previously: the female is unknown. We reexamined four of them and also found a new specimen. The female is unknown, and possibly has never been collected. This species seems to be endemic to the Algodones Sand Dunes. This species may be synonymous with
<emphasis id="B94E5F33FFDF980C0676FC405716FC69" italics="true" pageId="17" pageNumber="18">
<taxonomicName id="4C3AF8A2FFDF980C0676FC4056EBFC69" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="gloriosa">D. gloriosa</taxonomicName>
,
</emphasis>
and can only be separated from
<taxonomicName id="4C3AF8A2FFDF980C018EFC7E5519FC69" box="[647,782,946,971]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="gloriosa">
<emphasis id="B94E5F33FFDF980C018EFC7E5519FC69" box="[647,782,946,971]" italics="true" pageId="17" pageNumber="18">D. gloriosa</emphasis>
</taxonomicName>
by setal coloration and slightly larger size. Because of their large size and the potential for endemism, we hesitate to synonymize
<taxonomicName id="4C3AF8A2FFDF980C0723FC1552DCFC50" box="[1066,1227,985,1010]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="imperialis">
<emphasis id="B94E5F33FFDF980C0723FC1552DCFC50" box="[1066,1227,985,1010]" italics="true" pageId="17" pageNumber="18">D. imperialis</emphasis>
</taxonomicName>
with
<taxonomicName id="4C3AF8A2FFDF980C061BFC15538AFC50" box="[1298,1437,985,1010]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="gloriosa">
<emphasis id="B94E5F33FFDF980C061BFC15538AFC50" box="[1298,1437,985,1010]" italics="true" pageId="17" pageNumber="18">D. gloriosa</emphasis>
</taxonomicName>
without further evidence. Molecular data would be helpful for clarify this difficulty, but we were unable to extract viable DNA from the newest specimen possibly due to the length of time it was left in the trap.
</paragraph>
<paragraph id="8B858321FFDF980C03CCFB815756FB16" blockId="17.[151,1437,482,1204]" pageId="17" pageNumber="18">
<bibRefCitation id="EFABFED0FFDF980C03CCFB8157E5FBC5" author="Manley" box="[197,498,1101,1127]" pageId="17" pageNumber="52" refString="Manley, D. G. &amp; Pitts, J. P. (2004) Two new black species of Dasymutilla (Hymenoptera: Mutillidae) from Texas and California. Journal of the Kansas Entomological Societ y, 77, 644 - 649." type="journal article" year="2004">Manley and Pitts (2004)</bibRefCitation>
described this species as lacking an apical fringe of setae on the pygidium. However, examination of the
<typeStatus id="54813D83FFDF980C02FBFBB85472FB2C" box="[498,613,1140,1166]" pageId="17" pageNumber="18" type="paratype">paratypes</typeStatus>
, however, shows variation in the presence or absence of this fringe, as in
<taxonomicName id="4C3AF8A2FFDF980C03BCFB57572DFB16" box="[181,314,1179,1204]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="gloriosa">
<emphasis id="B94E5F33FFDF980C03BCFB57572DFB16" box="[181,314,1179,1204]" italics="true" pageId="17" pageNumber="18">D. gloriosa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>