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<document id="A177627355CA15A2B3E99F70D38DA909" ID-DOI="10.11646/zootaxa.3915.4.1" ID-GBIF-Dataset="01f159e7-e94a-4062-bf53-ea130633c06e" ID-ISSN="1175-5326" ID-Zenodo-Dep="241155" ID-ZooBank="7BC2D981-BCBC-448B-8F85-4D1DD1097BDD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460973632132" checkinUser="plazi" docAuthor="Monjaraz-Ruedas, Rodrigo &amp; Francke, Oscar F." docDate="2015" docId="03931F53FFE3FFFAFF68FDACFA6A0A49" docLanguage="en" docName="zt03915p490.pdf" docOrigin="Zootaxa 3915 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mayazomus kaamuul Monjaraz-Ruedas &amp; Francke, 2015, new species" docType="treatment" docVersion="8" lastPageNumber="478" masterDocId="FFAA672BFFFBFFE1FFFFFFE8FFFB0B76" masterDocTitle="Taxonomic revision of the genus Mayazomus Reddell &amp; Cokendolpher, 1995 (Schizomida: Hubbardiidae), with description of five new species from Chiapas, Mexico" masterLastPageNumber="490" masterPageNumber="451" pageNumber="475" updateTime="1698559400589" updateUser="plazi">
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<mods:title id="CD9CBF13B8E3B102DC47DBAC3150B6E0">Taxonomic revision of the genus Mayazomus Reddell &amp; Cokendolpher, 1995 (Schizomida: Hubbardiidae), with description of five new species from Chiapas, Mexico</mods:title>
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<mods:namePart id="63325BD473BEA4604487CE6BB9D3F46C">Monjaraz-Ruedas, Rodrigo</mods:namePart>
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<mods:date id="678C2487AE13866DFCAE70886ABF15E9">2015</mods:date>
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<treatment id="03931F53FFE3FFFAFF68FDACFA6A0A49" ID-DOI="http://doi.org/10.5281/zenodo.6113433" ID-GBIF-Taxon="119594053" ID-Zenodo-Dep="6113433" LSID="urn:lsid:plazi:treatment:03931F53FFE3FFFAFF68FDACFA6A0A49" httpUri="http://treatment.plazi.org/id/03931F53FFE3FFFAFF68FDACFA6A0A49" lastPageId="27" lastPageNumber="478" pageId="24" pageNumber="475">
<subSubSection id="C320FDCEFFE3FFF9FF68FDACFE4F09F6" pageId="24" pageNumber="475" type="nomenclature">
<paragraph id="8B85AE45FFE3FFF9FF68FDACFDC20929" blockId="24.[151,569,580,640]" box="[151,569,580,607]" pageId="24" pageNumber="475">
<heading id="D0CD1929FFE3FFF9FF68FDACFDC20929" bold="true" box="[151,569,580,607]" fontSize="11" level="1" pageId="24" pageNumber="475" reason="1">
<emphasis id="B94E7257FFE3FFF9FF68FDACFDC20929" bold="true" box="[151,569,580,607]" pageId="24" pageNumber="475">
<taxonomicName id="4C3AD5C6FFE3FFF9FF68FDACFE610928" ID-CoL="3YH7P" box="[151,410,580,606]" class="Arachnida" family="Hubbardiidae" genus="Mayazomus" kingdom="Animalia" order="Schizomida" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="kaamuul" status="sp. nov.">
<emphasis id="B94E7257FFE3FFF9FF68FDACFE610928" bold="true" box="[151,410,580,606]" italics="true" pageId="24" pageNumber="475">Mayazomus kaamuul</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27DCF2CFFE3FFF9FE56FDADFDC20929" box="[425,569,581,607]" pageId="24" pageNumber="475" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B85AE45FFE3FFF9FF68FD8FFE4F09F6" blockId="24.[151,569,580,640]" box="[151,436,615,640]" pageId="24" pageNumber="475">
<figureCitation id="1301B2C0FFE3FFF9FF68FD8FFEE409F6" box="[151,287,615,640]" captionStart-0="FIGURES 65, 66" captionStart-1="FIGURES 67 70" captionStart-2="FIGURES 71 78" captionStartId-0="25.[151,264,1967,1989]" captionStartId-1="26.[151,264,1217,1239]" captionStartId-2="27.[151,264,1607,1629]" captionTargetBox-0="[166,1408,663,1946]" captionTargetBox-1="[150,1435,193,1195]" captionTargetBox-2="[269,1310,380,1585]" captionTargetId-0="figure@25.[155,1432,661,1946]" captionTargetId-1="figure@26.[150,1436,193,1195]" captionTargetId-2="figure@27.[269,1318,375,1585]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="27" captionText-0="FIGURES 65, 66. Mayazomus kaamuul new species. Male holotype. Habitus: 65. Dorsal view. Female paratype. Habitus: 66. Dorsal view. Scale bar = 2 mm." captionText-1="FIGURES 67 70. Mayazomus kaamuul new species. Male. Chelicera (67 69): 67, mesal view. 68, fixed finger. 69, movable finger. Female. Spermathecae: 70, dorsal view. Scale bar (Fig. 70) = 0.05 mm." captionText-2="FIGURES 71 78. Mayazomus kaamuul new species. Male holotype. Flagellum (71 73): 71, dorsal view. 72, ventral view. 73, lateral view. Female. Flagellum (74 76): 74, dorsal view. 75, ventral view. 76, lateral view. Heteromorphic male. Pedipalp (77, 78): 77, ectal view. 78. mesal view. Scale bars = 0.2 mm." httpUri-0="https://zenodo.org/record/241171/files/figure.png" httpUri-1="https://zenodo.org/record/241172/files/figure.png" httpUri-2="https://zenodo.org/record/241173/files/figure.png" pageId="24" pageNumber="475">Figs. 6578</figureCitation>
,
<figureCitation id="1301B2C0FFE3FFF9FED4FD80FEAC09F6" box="[299,343,616,640]" captionStart="FIGURE 112" captionStartId="38.[151,250,1147,1169]" captionTargetBox="[151,1436,193,1125]" captionTargetId="figure@38.[151,1436,193,1126]" captionTargetPageId="38" captionText="FIGURE 112. Distribution map of the known species of the genus Mayazomus, showing the type localities of the five new species: M. tzotzil, M. aluxe, M. kaamuul, M. yaax and M. loobil" httpUri="https://zenodo.org/record/241182/files/figure.png" pageId="24" pageNumber="475">112</figureCitation>
,
<tableCitation id="C6B89BFEFFE3FFF9FE9DFD8FFE4F09F6" box="[354,436,615,640]" captionStart="TABLE 1" captionStartId="17.[151,239,548,571]" captionTargetBox="[159,1373,638,2029]" captionTargetPageId="17" captionText="TABLE 1. Measurements (mm) of Mayazomus tzotzil new species and Mayazomus kaamuul new species." httpUri="http://table.plazi.org/id/DF45FECDFFEAFFF0FF68FDCCFAFA094D" pageId="24" pageNumber="475" tableUuid="DF45FECDFFEAFFF0FF68FDCCFAFA094D">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C320FDCEFFE3FFF9FF68FD47FD3B08D6" pageId="24" pageNumber="475" type="materials_examined">
<paragraph id="8B85AE45FFE3FFF9FF68FD47FD3B08D6" blockId="24.[151,1437,687,2009]" pageId="24" pageNumber="475">
<emphasis id="B94E7257FFE3FFF9FF68FD47FEBE09BE" bold="true" box="[151,325,687,712]" pageId="24" pageNumber="475">
<typeStatus id="548110E7FFE3FFF9FF68FD47FF2909BE" box="[151,210,687,712]" pageId="24" pageNumber="475">Type</typeStatus>
material.
</emphasis>
<collectingCountry id="F32DEED5FFE3FFF9FEB3FD58FE4409BE" box="[332,447,688,712]" name="Mexico" pageId="24" pageNumber="475">MEXICO</collectingCountry>
:
<emphasis id="B94E7257FFE3FFF9FE35FD47FDDC09BE" box="[458,551,687,712]" italics="true" pageId="24" pageNumber="475">Chiapas</emphasis>
: Heteromorphic male
<typeStatus id="548110E7FFE3FFF9FCD9FD47FC7309BE" box="[806,904,687,712]" pageId="24" pageNumber="475" type="holotype">holotype</typeStatus>
(CNAN-T0860) [
<date id="FF848885FFE3FFF9FBACFD58FB1109BE" box="[1107,1258,687,712]" pageId="24" pageNumber="475" value="2013-07-23">23 July 2013</date>
, O. Francke, A. Valdez, C. Santibáñez, J. Mendoza, K. Zárate, R. Monjaraz] from outside of Cueva de las Abejas (
<geoCoordinate id="EE0EC882FFE3FFF9FAF1FD3CFA63099B" box="[1294,1432,724,749]" direction="north" orientation="latitude" pageId="24" pageNumber="475" precision="1" value="16.84874">16.84874ºN</geoCoordinate>
,
<geoCoordinate id="EE0EC882FFE3FFF9FF68FD10FEED0866" box="[151,278,760,784]" direction="west" orientation="longitude" pageId="24" pageNumber="475" precision="5" value="-93.2433">93.2433ºW</geoCoordinate>
, 1190 masl), Municipio San Fernando.
<typeStatus id="548110E7FFE3FFF9FD2CFD10FCBE0866" box="[723,837,760,784]" pageId="24" pageNumber="475" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D3C65CCFFE3FFF9FCB0FD10FC470866" box="[847,956,759,784]" pageId="24" pageNumber="475" type="female">2 females</specimenCount>
(CNAN-T0861), same data as
<typeStatus id="548110E7FFE3FFF9FAE2FD1FFA780866" box="[1309,1411,759,784]" pageId="24" pageNumber="475" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D3C65CCFFE3FFF9FA71FD10FF180843" pageId="24" pageNumber="475" type="female">1 female</specimenCount>
(CNAN-T0862) [
<date id="FF848885FFE3FFF9FE50FCF4FDB20842" box="[431,585,796,820]" pageId="24" pageNumber="475" value="2011-06-19">19 June 2011</date>
, O. Francke, A. Valdez, C. Santibáñez, J. Cruz, G. Contreras, K. Zarate, R. Monjaraz], same locality data as
<typeStatus id="548110E7FFE3FFF9FDE6FCD7FD84082E" box="[537,639,831,856]" pageId="24" pageNumber="475" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D3C65CCFFE3FFF9FD72FCA8FD0A082E" box="[653,753,831,856]" pageId="24" pageNumber="475" type="female">1 female</specimenCount>
(CNAN-T0863) [
<date id="FF848885FFE3FFF9FC3EFCD7FB9B082E" box="[961,1120,831,856]" pageId="24" pageNumber="475" value="2011-06-15">15 June 2011</date>
, O. Francke, A. Valdez, C. Santibáñez, J. Cruz, G. Contreras, K. Zarate, R. Monjaraz] from outside Cueva de Paso Burro, (
<geoCoordinate id="EE0EC882FFE3FFF9FAF1FC8CFA63080B" box="[1294,1432,868,893]" direction="north" orientation="latitude" pageId="24" pageNumber="475" precision="1" value="16.82883">16.82883ºN</geoCoordinate>
,
<geoCoordinate id="EE0EC882FFE3FFF9FF68FC60FEED08D6" box="[151,278,904,928]" direction="west" orientation="longitude" pageId="24" pageNumber="475" precision="5" value="-93.2433">93.2433ºW</geoCoordinate>
,
<specimenCount id="9D3C65CCFFE3FFF9FEDFFC6FFE8908D6" box="[288,370,903,928]" pageId="24" pageNumber="475" type="generic">
<quantity id="4CC203A0FFE3FFF9FEDFFC6FFE9508D6" box="[288,366,903,928]" metricMagnitude="2" metricUnit="m" metricValue="7.67" pageId="24" pageNumber="475" unit="m" value="767.0">767 m</quantity>
.
</specimenCount>
a.s.l.), Municipio Berriozabal.
</paragraph>
</subSubSection>
<subSubSection id="C320FDCEFFE3FFF9FF38FC44FBB90F7B" pageId="24" pageNumber="475" type="etymology">
<paragraph id="8B85AE45FFE3FFF9FF38FC44FBB90F7B" blockId="24.[151,1437,687,2009]" pageId="24" pageNumber="475">
<emphasis id="B94E7257FFE3FFF9FF38FC44FEB508B3" bold="true" box="[199,334,940,965]" pageId="24" pageNumber="475">Etymology.</emphasis>
The specific name is a noun formed by the combination of two Mayan words: “Kaa”, which means two or double, and “Muul” which means mound or swelling; referring to the two swellings present in the dorsal surface of the male flagellum, which is a diagnostic character for the species.
</paragraph>
</subSubSection>
<subSubSection id="C320FDCEFFE3FFF9FF38FBFFFC9C0E96" pageId="24" pageNumber="475" type="diagnosis">
<paragraph id="8B85AE45FFE3FFF9FF38FBFFFC9C0E96" blockId="24.[151,1437,687,2009]" pageId="24" pageNumber="475">
<emphasis id="B94E7257FFE3FFF9FF38FBFFFEBA0F46" bold="true" box="[199,321,1047,1072]" pageId="24" pageNumber="475">Diagnosis.</emphasis>
Males can be distinguished by the pedipalp trochanter having a small trapezoidal apical process; pedipalp femur with spiniform setiferous tubercles
<emphasis id="B94E7257FFE3FFF9FD2BFBD4FCFB0F22" box="[724,768,1084,1108]" italics="true" pageId="24" pageNumber="475">Fv1</emphasis>
and
<emphasis id="B94E7257FFE3FFF9FCC9FBD4FC990F22" box="[822,866,1084,1108]" italics="true" pageId="24" pageNumber="475">Fv2</emphasis>
reduced and rounded (
<figureCitation id="1301B2C0FFE3FFF9FB9CFBD4FB4D0F22" box="[1123,1206,1084,1109]" captionStart="FIGURES 71 78" captionStartId="27.[151,264,1607,1629]" captionTargetBox="[269,1310,380,1585]" captionTargetId="figure@27.[269,1318,375,1585]" captionTargetPageId="27" captionText="FIGURES 71 78. Mayazomus kaamuul new species. Male holotype. Flagellum (71 73): 71, dorsal view. 72, ventral view. 73, lateral view. Female. Flagellum (74 76): 74, dorsal view. 75, ventral view. 76, lateral view. Heteromorphic male. Pedipalp (77, 78): 77, ectal view. 78. mesal view. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/241173/files/figure.png" pageId="24" pageNumber="475">Fig. 77</figureCitation>
), mesal surface with a small blunt apophysis (FAP); pedipalp tibia with a weak, triangular mesal apophysis, placed medially (TMA); and by the presence of one pair of circular dorso-submedian swellings and a posterior depression in the dorsal surface of the flagellum (
<figureCitation id="1301B2C0FFE3FFF9FE3EFB4FFDB00FB6" box="[449,587,1191,1216]" captionStart="FIGURES 71 78" captionStartId="27.[151,264,1607,1629]" captionTargetBox="[269,1310,380,1585]" captionTargetId="figure@27.[269,1318,375,1585]" captionTargetPageId="27" captionText="FIGURES 71 78. Mayazomus kaamuul new species. Male holotype. Flagellum (71 73): 71, dorsal view. 72, ventral view. 73, lateral view. Female. Flagellum (74 76): 74, dorsal view. 75, ventral view. 76, lateral view. Heteromorphic male. Pedipalp (77, 78): 77, ectal view. 78. mesal view. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/241173/files/figure.png" pageId="24" pageNumber="475">Figs. 7173</figureCitation>
) Females can by distinguished by the spermathecae having lateral lobes acuminate; median lobes long and slender, inverse J-shaped, almost three times longer than lateral lobes (
<figureCitation id="1301B2C0FFE3FFF9FAC4FB24FA6B0F92" box="[1339,1424,1228,1253]" captionStart="FIGURES 67 70" captionStartId="26.[151,264,1217,1239]" captionTargetBox="[150,1435,193,1195]" captionTargetId="figure@26.[150,1436,193,1195]" captionTargetPageId="26" captionText="FIGURES 67 70. Mayazomus kaamuul new species. Male. Chelicera (67 69): 67, mesal view. 68, fixed finger. 69, movable finger. Female. Spermathecae: 70, dorsal view. Scale bar (Fig. 70) = 0.05 mm." httpUri="https://zenodo.org/record/241172/files/figure.png" pageId="24" pageNumber="475">Fig. 70</figureCitation>
).
<taxonomicName id="4C3AD5C6FFE3FFF9FF68FB19FE7F0E7E" box="[151,388,1263,1288]" class="Arachnida" family="Hubbardiidae" genus="Mayazomus" kingdom="Animalia" order="Schizomida" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="kaamuul">
<emphasis id="B94E7257FFE3FFF9FF68FB19FE7F0E7E" box="[151,388,1263,1288]" italics="true" pageId="24" pageNumber="475">Mayazomus kaamuul</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C3AD5C6FFE3FFF9FDFFFB19FCEA0E7E" box="[512,785,1263,1288]" class="Arachnida" family="Hubbardiidae" genus="Mayazomus" kingdom="Animalia" order="Schizomida" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="hoffmannae">
<emphasis id="B94E7257FFE3FFF9FDFFFB19FCEA0E7E" box="[512,785,1263,1288]" italics="true" pageId="24" pageNumber="475">Mayazomus hoffmannae</emphasis>
</taxonomicName>
having a trapezoidal apical process, a blunt apophysis and median lobes about three times longer than lateral lobes, but they differ in pedipalp size,
<taxonomicName id="4C3AD5C6FFE3FFF9FB32FAFDFA860E5B" box="[1229,1405,1300,1325]" class="Arachnida" family="Hubbardiidae" genus="Mayazomus" kingdom="Animalia" order="Schizomida" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="hoffmannae">
<emphasis id="B94E7257FFE3FFF9FB32FAFDFB1F0E5A" box="[1229,1252,1301,1324]" italics="true" pageId="24" pageNumber="475">M</emphasis>
.
<emphasis id="B94E7257FFE3FFF9FB07FAFCFA860E5B" box="[1272,1405,1300,1325]" italics="true" pageId="24" pageNumber="475">hoffmannae</emphasis>
</taxonomicName>
is proportionately longer (3.61 times longer than propeltidium length) than
<taxonomicName id="4C3AD5C6FFE3FFF9FC0BFAD1FB850E26" box="[1012,1150,1335,1360]" class="Arachnida" family="Hubbardiidae" genus="Mayazomus" kingdom="Animalia" order="Schizomida" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="kaamuul">
<emphasis id="B94E7257FFE3FFF9FC0BFAD1FB850E26" box="[1012,1150,1335,1360]" italics="true" pageId="24" pageNumber="475">M. kaamuul</emphasis>
</taxonomicName>
(2.82 times longer than propeltidium length). The male flagellum of
<taxonomicName id="4C3AD5C6FFE3FFF9FD77FAB5FCF50E03" box="[648,782,1372,1397]" class="Arachnida" family="Hubbardiidae" genus="Mayazomus" kingdom="Animalia" order="Schizomida" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="kaamuul">
<emphasis id="B94E7257FFE3FFF9FD77FAB5FCF50E03" box="[648,782,1372,1397]" italics="true" pageId="24" pageNumber="475">M. kaamuul</emphasis>
</taxonomicName>
presents a pair of rounded dorso-submedian swellings that are diagnostic and unique in the genus. The tibial apophysis is shorter and less pronounced in
<taxonomicName id="4C3AD5C6FFE3FFF9FB3FFA69FABD0EEE" box="[1216,1350,1407,1432]" class="Arachnida" family="Hubbardiidae" genus="Mayazomus" kingdom="Animalia" order="Schizomida" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="kaamuul">
<emphasis id="B94E7257FFE3FFF9FB3FFA69FABD0EEE" box="[1216,1350,1407,1432]" italics="true" pageId="24" pageNumber="475">M. kaamuul</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C3AD5C6FFE3FFF9FF68FA4DFEB80ECB" box="[151,323,1444,1469]" class="Arachnida" family="Hubbardiidae" genus="Mayazomus" kingdom="Animalia" order="Schizomida" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="hoffmannae">
<emphasis id="B94E7257FFE3FFF9FF68FA4DFF550ECA" box="[151,174,1445,1468]" italics="true" pageId="24" pageNumber="475">M</emphasis>
.
<emphasis id="B94E7257FFE3FFF9FF40FA4CFEB80ECB" box="[191,323,1444,1469]" italics="true" pageId="24" pageNumber="475">hoffmannae</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFABD3B4FFE3FFF9FEAAFA4CFCB20ECB" author="Reddell" box="[341,841,1444,1469]" pageId="24" pageNumber="475" refString="Reddell, J. R. &amp; Cokendolpher, J. C. (1986) New species and records of Schizomus (Arachnida: Schizomida) from Mexico. Texas Memorial Museum, Speleological Monographs, 1, 31 - 38." type="journal article" year="1986">Reddell &amp; Cokendolpher, 1986: figs. 610</bibRefCitation>
). Spermathecae of
<taxonomicName id="4C3AD5C6FFE3FFF9FBD7FA4DFB3C0ECB" box="[1064,1223,1444,1469]" class="Arachnida" family="Hubbardiidae" genus="Mayazomus" kingdom="Animalia" order="Schizomida" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="hoffmanae">
<emphasis id="B94E7257FFE3FFF9FBD7FA4DFB3C0ECB" box="[1064,1223,1444,1469]" italics="true" pageId="24" pageNumber="475">M. hoffmanae</emphasis>
</taxonomicName>
present small and slender lateral lobes whereas
<taxonomicName id="4C3AD5C6FFE3FFF9FE1FFA21FD9D0E96" box="[480,614,1479,1504]" class="Arachnida" family="Hubbardiidae" genus="Mayazomus" kingdom="Animalia" order="Schizomida" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="kaamuul">
<emphasis id="B94E7257FFE3FFF9FE1FFA21FD9D0E96" box="[480,614,1479,1504]" italics="true" pageId="24" pageNumber="475">M. kaamuul</emphasis>
</taxonomicName>
has wide lateral lobes.
</paragraph>
</subSubSection>
<subSubSection id="C320FDCEFFE3FFFAFF38FA04FC260BA2" lastPageId="27" lastPageNumber="478" pageId="24" pageNumber="475" type="description">
<paragraph id="8B85AE45FFE3FFF9FF38FA04FC930D06" blockId="24.[151,1437,687,2009]" pageId="24" pageNumber="475">
<emphasis id="B94E7257FFE3FFF9FF38FA04FEA20D73" bold="true" box="[199,345,1516,1541]" pageId="24" pageNumber="475">Description.</emphasis>
<emphasis id="B94E7257FFE3FFF9FE9AFA04FD260D73" box="[357,733,1516,1541]" italics="true" pageId="24" pageNumber="475">
Heteromorphic male (
<typeStatus id="548110E7FFE3FFF9FD96FA04FD350D73" box="[617,718,1516,1541]" pageId="24" pageNumber="475" type="holotype">holotype</typeStatus>
):
</emphasis>
Dark brownish. Prosoma: Propeltidium with two setae on anterior process and two pairs of dorsal setae; ocular spots asymmetrical and irregular. Mesopeltidial plates 0.26 wide, 0.05 long; gap between the plates 0.26. Metapeltidium 0.40 long, 0.67 wide. Anterior sternum with 10 setae, plus two sternopophysial setae; posterior sternum with six setae.
</paragraph>
<paragraph id="8B85AE45FFE3FFF9FF38F994FE6D0C1B" blockId="24.[151,1437,687,2009]" pageId="24" pageNumber="475">
<emphasis id="B94E7257FFE3FFF9FF38F994FEB90DE3" box="[199,322,1660,1685]" italics="true" pageId="24" pageNumber="475">Chelicerae</emphasis>
(
<figureCitation id="1301B2C0FFE3FFF9FEADF994FE250DE3" box="[338,478,1660,1685]" captionStart="FIGURES 67 70" captionStartId="26.[151,264,1217,1239]" captionTargetBox="[150,1435,193,1195]" captionTargetId="figure@26.[150,1436,193,1195]" captionTargetPageId="26" captionText="FIGURES 67 70. Mayazomus kaamuul new species. Male. Chelicera (67 69): 67, mesal view. 68, fixed finger. 69, movable finger. Female. Spermathecae: 70, dorsal view. Scale bar (Fig. 70) = 0.05 mm." httpUri="https://zenodo.org/record/241172/files/figure.png" pageId="24" pageNumber="475">Figs. 6769</figureCitation>
): Movable finger: Serrula with 20 teeth. Fixed finger with 5 smaller teeth between 2 primary teeth. Setation: Setal group formula: 1:3, 2:6, 3:4, 4:2, 5:9, 6:1. G1 (setae group 1) with three spatulate setae, covered with a few spiniform spicules (
<figureCitation id="1301B2C0FFE3FFF9FD69F92CFD1C0DAB" box="[662,743,1732,1757]" captionStart="FIGURES 67 70" captionStartId="26.[151,264,1217,1239]" captionTargetBox="[150,1435,193,1195]" captionTargetId="figure@26.[150,1436,193,1195]" captionTargetPageId="26" captionText="FIGURES 67 70. Mayazomus kaamuul new species. Male. Chelicera (67 69): 67, mesal view. 68, fixed finger. 69, movable finger. Female. Spermathecae: 70, dorsal view. Scale bar (Fig. 70) = 0.05 mm." httpUri="https://zenodo.org/record/241172/files/figure.png" pageId="24" pageNumber="475">Fig. 69</figureCitation>
) starting at middle of the shaft; G2 composed of six feathered setae, subequal in length, and shorter than movable finger length; G3 with four setae, feathered apically and smooth basally, two shorter, about 2/3 of long setae; G4 consisting of two setae, smooth, short and thick, spine-like; G5 with nine similar sized setae, feathered apically and longer than fixed finger; G6 with one smooth seta, 1/2 of movable finger length.
</paragraph>
<paragraph id="8B85AE45FFE3FFF8FF38F89FFB620AFE" blockId="24.[151,1437,687,2009]" lastBlockId="25.[151,1437,151,608]" lastPageId="25" lastPageNumber="476" pageId="24" pageNumber="475">
<emphasis id="B94E7257FFE3FFF9FF38F89FFECC0CE6" box="[199,311,1911,1936]" italics="true" pageId="24" pageNumber="475">Pedipalps</emphasis>
(
<figureCitation id="1301B2C0FFE3FFF9FEBAF89FFE350CE6" box="[325,462,1911,1936]" captionStart="FIGURES 71 78" captionStartId="27.[151,264,1607,1629]" captionTargetBox="[269,1310,380,1585]" captionTargetId="figure@27.[269,1318,375,1585]" captionTargetPageId="27" captionText="FIGURES 71 78. Mayazomus kaamuul new species. Male holotype. Flagellum (71 73): 71, dorsal view. 72, ventral view. 73, lateral view. Female. Flagellum (74 76): 74, dorsal view. 75, ventral view. 76, lateral view. Heteromorphic male. Pedipalp (77, 78): 77, ectal view. 78. mesal view. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/241173/files/figure.png" pageId="24" pageNumber="475">Figs. 77, 78</figureCitation>
): Robust; 2.82 times longer than propeltidium length. Trochanter long, 2.6 times longer than high; apical process triangular, greater than 45º; with a row of 10 setae on ventral margin; mesal surface with a row of three setae near ventral margin, plus two setae near dorsal margin; with a small median mesal spur. Femur wide, 1.66 times longer than high, elongated distally; distal margin straight, ventral margin flattened; distal margin of ectal surface with spiniform setiferous tubercles
<emphasis id="B94E7257FFE2FFF8FD1CFF54FCF50BA2" box="[739,782,188,212]" italics="true" pageId="25" pageNumber="476">Fv1</emphasis>
and
<emphasis id="B94E7257FFE2FFF8FCB6FF54FC8F0BA2" box="[841,884,188,212]" italics="true" pageId="25" pageNumber="476">Fv2</emphasis>
close to each other, the tubercle is globular and small; mesal surface with two ventral plus two dorsal spiniform setae (2+2); distal margin of the mesal surface with a small, blunt apophysis like a spine (
<figureCitation id="1301B2C0FFE2FFF8FDC4FEECFD770A6A" box="[571,652,260,285]" captionStart="FIGURES 71 78" captionStartId="27.[151,264,1607,1629]" captionTargetBox="[269,1310,380,1585]" captionTargetId="figure@27.[269,1318,375,1585]" captionTargetPageId="27" captionText="FIGURES 71 78. Mayazomus kaamuul new species. Male holotype. Flagellum (71 73): 71, dorsal view. 72, ventral view. 73, lateral view. Female. Flagellum (74 76): 74, dorsal view. 75, ventral view. 76, lateral view. Heteromorphic male. Pedipalp (77, 78): 77, ectal view. 78. mesal view. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/241173/files/figure.png" pageId="25" pageNumber="476">Fig. 78</figureCitation>
) (FAP). Patella strongly curved, inverse U-shaped; flattened ventrally and apically widened; with scattered setae and without armature. Tibia elongated, with scattered setae; with one apophysis on ventromesal surface opposable to tarsus, situated medially (TMA). Tarsus short, 2.86 times longer than wide. Claw about 1/2 as long as dorsal length of tarsus; spurs asymmetrical, 0.06 long.
</paragraph>
<paragraph id="8B85AE45FFE2FFF8FF38FE7DFAA10ADB" blockId="25.[151,1437,151,608]" box="[199,1370,404,429]" pageId="25" pageNumber="476">
<emphasis id="B94E7257FFE2FFF8FF38FE7DFEFD0ADA" box="[199,262,405,428]" italics="true" pageId="25" pageNumber="476">Legs:</emphasis>
Leg I, basitarsal-telotarsal proportions: 20: 3: 3: 4: 4: 4: 10. Femur IV 2.46 times longer than high.
</paragraph>
<paragraph id="8B85AE45FFE2FFF8FF38FE5FFBFB0916" blockId="25.[151,1437,151,608]" pageId="25" pageNumber="476">
<emphasis id="B94E7257FFE2FFF8FF38FE5FFE9F0AA6" box="[199,356,439,464]" italics="true" pageId="25" pageNumber="476">Opisthosoma:</emphasis>
Tergite I with two pairs microsetae anteriorly plus one pair macrosetae 2+1; tergite II with three pair microsetae anteriorly and two pairs macrosetae 3+2; tergites IIIVII with one pair dorsal setae each; tergites VIIIIX with one pair dorsal and one pair lateral setae; segments XXI telescoped with one pair lateral setae and five ventral setae; segment XII with one dorsal, two lateral and four ventral pairs of setae, without posterodorsal process. Sternites with scattered setae; genital aperture with many microsetae.
</paragraph>
<caption id="DF45FECDFFE2FFF8FF68F847FE2C0C92" httpUri="https://zenodo.org/record/241171/files/figure.png" pageId="25" pageNumber="476" targetBox="[166,1408,663,1946]" targetPageId="25">
<paragraph id="8B85AE45FFE2FFF8FF68F847FE2C0C92" blockId="25.[151,1436,1967,2020]" pageId="25" pageNumber="476">
<emphasis id="B94E7257FFE2FFF8FF68F847FEAF0CB3" bold="true" box="[151,340,1967,1989]" pageId="25" pageNumber="476">FIGURES 65, 66.</emphasis>
<taxonomicName id="4C3AD5C6FFE2FFF8FEA5F859FDCA0CB3" box="[346,561,1967,1990]" class="Arachnida" family="Hubbardiidae" genus="Mayazomus" kingdom="Animalia" order="Schizomida" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="kaamuul" status="sp. nov.">
<emphasis id="B94E7257FFE2FFF8FEA5F859FDCA0CB3" box="[346,561,1967,1990]" italics="true" pageId="25" pageNumber="476">Mayazomus kaamuul</emphasis>
</taxonomicName>
<emphasis id="B94E7257FFE2FFF8FDC7F847FD410CB3" bold="true" box="[568,698,1967,1989]" pageId="25" pageNumber="476">
<taxonomicNameLabel id="A27DCF2CFFE2FFF8FDC7F847FD4D0CB3" box="[568,694,1967,1989]" pageId="25" pageNumber="476" rank="species">new species</taxonomicNameLabel>
.
</emphasis>
Male holotype. Habitus: 65. Dorsal view. Female paratype. Habitus: 66. Dorsal view. Scale bar = 2 mm.
</paragraph>
</caption>
<caption id="DF45FECDFFE1FFFBFF68FB29FC4B0F83" httpUri="https://zenodo.org/record/241172/files/figure.png" pageId="26" pageNumber="477" targetBox="[150,1435,193,1195]" targetPageId="26">
<paragraph id="8B85AE45FFE1FFFBFF68FB29FC4B0F83" blockId="26.[151,1436,1217,1269]" pageId="26" pageNumber="477">
<emphasis id="B94E7257FFE1FFFBFF68FB29FEAE0FA1" bold="true" box="[151,341,1217,1239]" pageId="26" pageNumber="477">FIGURES 6770.</emphasis>
<taxonomicName id="4C3AD5C6FFE1FFFBFEA4FB2AFDC80FA1" box="[347,563,1217,1239]" class="Arachnida" family="Hubbardiidae" genus="Mayazomus" kingdom="Animalia" order="Schizomida" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="kaamuul" status="sp. nov.">
<emphasis id="B94E7257FFE1FFFBFEA4FB2AFDC80FA1" box="[347,563,1217,1239]" italics="true" pageId="26" pageNumber="477">Mayazomus kaamuul</emphasis>
</taxonomicName>
<emphasis id="B94E7257FFE1FFFBFDC5FB29FD460FA1" bold="true" box="[570,701,1217,1239]" pageId="26" pageNumber="477">
<taxonomicNameLabel id="A27DCF2CFFE1FFFBFDC5FB29FD420FA1" box="[570,697,1217,1239]" pageId="26" pageNumber="477" rank="species">new species</taxonomicNameLabel>
.
</emphasis>
Male. Chelicera (6769): 67, mesal view. 68, fixed finger. 69, movable finger. Female. Spermathecae: 70, dorsal view. Scale bar (Fig. 70) = 0.05 mm.
</paragraph>
</caption>
<paragraph id="8B85AE45FFE1FFFBFF38FAF4FEE30D78" blockId="26.[151,1437,1308,2017]" pageId="26" pageNumber="477">
<emphasis id="B94E7257FFE1FFFBFF38FAF4FEC00E43" box="[199,315,1308,1333]" italics="true" pageId="26" pageNumber="477">Flagellum</emphasis>
(
<figureCitation id="1301B2C0FFE1FFFBFEB3FAF4FE210E43" box="[332,474,1308,1333]" captionStart="FIGURES 71 78" captionStartId="27.[151,264,1607,1629]" captionTargetBox="[269,1310,380,1585]" captionTargetId="figure@27.[269,1318,375,1585]" captionTargetPageId="27" captionText="FIGURES 71 78. Mayazomus kaamuul new species. Male holotype. Flagellum (71 73): 71, dorsal view. 72, ventral view. 73, lateral view. Female. Flagellum (74 76): 74, dorsal view. 75, ventral view. 76, lateral view. Heteromorphic male. Pedipalp (77, 78): 77, ectal view. 78. mesal view. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/241173/files/figure.png" pageId="26" pageNumber="477">Figs. 7173</figureCitation>
): dorsoventrally flattened, cordate in shape; 1.35 times longer than wide, 2.4 times longer than pedicel length; with one dorso-posterior depression, and a pair of dorso-submedian swellings. Setation:
<emphasis id="B94E7257FFE1FFFBFF68FA8DFF360E0B" box="[151,205,1381,1405]" italics="true" pageId="26" pageNumber="477">Dm1</emphasis>
situated over the bulb base;
<emphasis id="B94E7257FFE1FFFBFDE3FA8DFDA80E0B" box="[540,595,1381,1405]" italics="true" pageId="26" pageNumber="477">Dm4</emphasis>
situated distally;
<emphasis id="B94E7257FFE1FFFBFCDEFA8DFCB10E08" box="[801,842,1381,1406]" italics="true" pageId="26" pageNumber="477">Dl1</emphasis>
reduced, 1/3 length of
<emphasis id="B94E7257FFE1FFFBFBA4FA8DFB790E08" box="[1115,1154,1381,1406]" italics="true" pageId="26" pageNumber="477">Vl1</emphasis>
;
<emphasis id="B94E7257FFE1FFFBFB6CFA8DFB460E08" box="[1171,1213,1381,1406]" italics="true" pageId="26" pageNumber="477">Dl1</emphasis>
positioned at same level as
<emphasis id="B94E7257FFE1FFFBFF0CFA60FEE20ED7" box="[243,281,1416,1441]" italics="true" pageId="26" pageNumber="477">Vl1</emphasis>
;
<emphasis id="B94E7257FFE1FFFBFED7FA60FEA90ED7" box="[296,338,1416,1441]" italics="true" pageId="26" pageNumber="477">Dl3</emphasis>
at same level as
<emphasis id="B94E7257FFE1FFFBFDEEFA60FDC30ED7" box="[529,568,1416,1441]" italics="true" pageId="26" pageNumber="477">Vl2</emphasis>
; pair
<emphasis id="B94E7257FFE1FFFBFD86FA61FD570ED7" box="[633,684,1417,1441]" italics="true" pageId="26" pageNumber="477">Vm2</emphasis>
present; seta
<emphasis id="B94E7257FFE1FFFBFCB9FA61FC820ED7" box="[838,889,1417,1441]" italics="true" pageId="26" pageNumber="477">Vm1</emphasis>
at same level as
<emphasis id="B94E7257FFE1FFFBFBC7FA61FB900ED7" box="[1080,1131,1417,1441]" italics="true" pageId="26" pageNumber="477">Vm2</emphasis>
pair;
<emphasis id="B94E7257FFE1FFFBFB53FA61FB1B0ED6" box="[1196,1248,1417,1440]" italics="true" pageId="26" pageNumber="477">Vm5</emphasis>
posterior to
<emphasis id="B94E7257FFE1FFFBFA91FA60FA6E0ED7" box="[1390,1429,1416,1441]" italics="true" pageId="26" pageNumber="477">Vl1</emphasis>
. With one pair of antero-dorsal microsetae between
<emphasis id="B94E7257FFE1FFFBFCFDFA45FCC30EB3" box="[770,824,1453,1477]" italics="true" pageId="26" pageNumber="477">Dm1</emphasis>
and
<emphasis id="B94E7257FFE1FFFBFC83FA45FC5E0EB0" box="[892,933,1453,1478]" italics="true" pageId="26" pageNumber="477">Dl1</emphasis>
nearest to
<emphasis id="B94E7257FFE1FFFBFBCBFA45FBA60EB0" box="[1076,1117,1453,1478]" italics="true" pageId="26" pageNumber="477">Dl1</emphasis>
; one pair of antero-lateral microsetae on flagellar pedicel, and one pair of irregular clumps between
<emphasis id="B94E7257FFE1FFFBFBF4FA38FBCA0E9F" box="[1035,1073,1488,1513]" italics="true" pageId="26" pageNumber="477">Vl1</emphasis>
and
<emphasis id="B94E7257FFE1FFFBFB8CFA38FB610E9F" box="[1139,1178,1488,1513]" italics="true" pageId="26" pageNumber="477">Vl2</emphasis>
, each composed of 6 microsetae.
</paragraph>
<paragraph id="8B85AE45FFE1FFFBFF38F9F0FF080CC8" blockId="26.[151,1437,1308,2017]" pageId="26" pageNumber="477">
<emphasis id="B94E7257FFE1FFFBFF38F9F0FEE00D47" box="[199,283,1560,1585]" italics="true" pageId="26" pageNumber="477">Female</emphasis>
(
<typeStatus id="548110E7FFE1FFFBFED4F9F2FE740D47" box="[299,399,1562,1585]" pageId="26" pageNumber="477" type="paratype">
<emphasis id="B94E7257FFE1FFFBFED4F9F2FE740D47" box="[299,399,1562,1585]" italics="true" pageId="26" pageNumber="477">paratype</emphasis>
</typeStatus>
): Similar to the heteromorphic male, except in pedipalp development, 1.93 times longer than propeltidium length, the female pedipalp does not have armature or any modification on the segments, with a slight curvature in the patella; females present the same femoral apical process shape than males and the same position of the spinose setae on mesodistal margin. Other differences: body longer and more robust than male (
<tableCitation id="C6B89BFEFFE1FFFBFACDF96DFA710DEB" box="[1330,1418,1669,1694]" captionStart="TABLE 1" captionStartId="17.[151,239,548,571]" captionTargetBox="[159,1373,638,2029]" captionTargetPageId="17" captionText="TABLE 1. Measurements (mm) of Mayazomus tzotzil new species and Mayazomus kaamuul new species." httpUri="http://table.plazi.org/id/DF45FECDFFEAFFF0FF68FDCCFAFA094D" pageId="26" pageNumber="477" tableUuid="DF45FECDFFEAFFF0FF68FDCCFAFA094D">Table 1</tableCitation>
); setation as on male. Flagellum (
<figureCitation id="1301B2C0FFE1FFFBFE02F940FD7F0DB7" box="[509,644,1704,1729]" captionStart="FIGURES 71 78" captionStartId="27.[151,264,1607,1629]" captionTargetBox="[269,1310,380,1585]" captionTargetId="figure@27.[269,1318,375,1585]" captionTargetPageId="27" captionText="FIGURES 71 78. Mayazomus kaamuul new species. Male holotype. Flagellum (71 73): 71, dorsal view. 72, ventral view. 73, lateral view. Female. Flagellum (74 76): 74, dorsal view. 75, ventral view. 76, lateral view. Heteromorphic male. Pedipalp (77, 78): 77, ectal view. 78. mesal view. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/241173/files/figure.png" pageId="26" pageNumber="477">Figs. 7476</figureCitation>
) with three annuli. Setation: seta
<emphasis id="B94E7257FFE1FFFBFC01F940FBD30DB7" box="[1022,1064,1704,1729]" italics="true" pageId="26" pageNumber="477">Dl1</emphasis>
not reduced and at same level as
<emphasis id="B94E7257FFE1FFFBFF68F925FF450D90" box="[151,190,1741,1766]" italics="true" pageId="26" pageNumber="477">Vl1</emphasis>
;
<emphasis id="B94E7257FFE1FFFBFF2FF925FF010D90" box="[208,250,1741,1766]" italics="true" pageId="26" pageNumber="477">Dl3</emphasis>
posterior to
<emphasis id="B94E7257FFE1FFFBFE6BF925FE400D90" box="[404,443,1741,1766]" italics="true" pageId="26" pageNumber="477">Vl2</emphasis>
; seta
<emphasis id="B94E7257FFE1FFFBFDFBF925FDC30D93" box="[516,568,1741,1765]" italics="true" pageId="26" pageNumber="477">Vm1</emphasis>
at same level as
<emphasis id="B94E7257FFE1FFFBFCF6F925FC7F0D93" box="[777,900,1741,1765]" italics="true" pageId="26" pageNumber="477">Vm2; Vm2</emphasis>
reduced. Segment III with one lateral pair of microsetae near to
<emphasis id="B94E7257FFE1FFFBFE90F919FE590C7F" box="[367,418,1777,1801]" italics="true" pageId="26" pageNumber="477">Vm4</emphasis>
. Segment IV with one pair of distolateral microsetae near to
<emphasis id="B94E7257FFE1FFFBFBA2F918FB700C7F" box="[1117,1163,1776,1801]" italics="true" pageId="26" pageNumber="477">Vl2.</emphasis>
Spermathecae (
<figureCitation id="1301B2C0FFE1FFFBFABEF918FA6F0C7F" box="[1345,1428,1776,1801]" captionStart="FIGURES 67 70" captionStartId="26.[151,264,1217,1239]" captionTargetBox="[150,1435,193,1195]" captionTargetId="figure@26.[150,1436,193,1195]" captionTargetPageId="26" captionText="FIGURES 67 70. Mayazomus kaamuul new species. Male. Chelicera (67 69): 67, mesal view. 68, fixed finger. 69, movable finger. Female. Spermathecae: 70, dorsal view. Scale bar (Fig. 70) = 0.05 mm." httpUri="https://zenodo.org/record/241172/files/figure.png" pageId="26" pageNumber="477">Fig. 70</figureCitation>
) without bulbs, with 4 lobes; median pair curved, inverse J-shaped and longer than lateral pair; lateral pair curved on inner face and flat on external face ending in a pointed tip and wider than median pair; both pairs with duct openings. Gonopod long and thin; chitinized arc U- shaped, widened at the base in the posterior branch, with irregular margins, lateral tip not sclerotized. Chelicera: serrula with 17 teeth. Setal group formula 1:3, 2:6, 3:6, 4:2, 5:8, 6:1.
</paragraph>
<paragraph id="8B85AE45FFE1FFFAFF38F820FC260BA2" blockId="26.[151,1437,1308,2017]" lastBlockId="27.[151,1436,151,320]" lastPageId="27" lastPageNumber="478" pageId="26" pageNumber="477">
<emphasis id="B94E7257FFE1FFFBFF38F820FEC00C97" bold="true" box="[199,315,1992,2017]" pageId="26" pageNumber="477">Variation.</emphasis>
Homeomorphic male unknown. Female total length 3.884.88 (n=4). Cheliceral movable finger with 17 to 20 small teeth on serrula (mode= 20). Fixed finger of chelicerae with 4 to 5 (mode= 5) smaller teeth between 2 primary teeth. Variation in setal groups: G3 (46) and G5 (89).
</paragraph>
</subSubSection>
<subSubSection id="C320FDCEFFE0FFFAFF38FF37FA6A0A49" pageId="27" pageNumber="478" type="distribution">
<paragraph id="8B85AE45FFE0FFFAFF38FF37FA6A0A49" blockId="27.[151,1436,151,320]" pageId="27" pageNumber="478">
<emphasis id="B94E7257FFE0FFFAFF38FF37FE9B0B8E" bold="true" box="[199,352,223,248]" pageId="27" pageNumber="478">Distribution.</emphasis>
This species is distributed in the medium-height rainforest located NW of Tuxtla Gutierrez, Chiapas, in the municipalities of San Fernando and Berriozabal (
<figureCitation id="1301B2C0FFE0FFFAFC64FEECFBFA0A6A" box="[923,1025,260,285]" captionStart="FIGURE 112" captionStartId="38.[151,250,1147,1169]" captionTargetBox="[151,1436,193,1125]" captionTargetId="figure@38.[151,1436,193,1126]" captionTargetPageId="38" captionText="FIGURE 112. Distribution map of the known species of the genus Mayazomus, showing the type localities of the five new species: M. tzotzil, M. aluxe, M. kaamuul, M. yaax and M. loobil" httpUri="https://zenodo.org/record/241182/files/figure.png" pageId="27" pageNumber="478">Fig. 112</figureCitation>
). The specimens were collected in perturbed areas, under rocks and rotten logs; the only male known was found near a cornfield in a fairly dry zone.
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