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<document id="D4A04DB30270C6DB967231DF9BECA6C8" ID-CLB-Dataset="9488" ID-DOI="10.11646/zootaxa.4881.3.9" ID-GBIF-Dataset="bba05b9e-f544-488d-abf4-222aec8c64e7" ID-ISSN="1175-5326" ID-Zenodo-Dep="4283982" ID-ZooBank="DCA548C5-E514-4C1F-86F6-8D99360738E9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1605982853407" checkinUser="plazi" docAuthor="Nikoshe, Akash P., Meshram, Naresh M., Stuti &amp; Dey, Debjani" docDate="2020" docId="0393062EFFE2622EFF12F9A2D30EFEB2" docLanguage="en" docName="zootaxa.4881.3.9.pdf" docOrigin="Zootaxa 4881 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hecalus shanayai Nikoshe &amp; Meshram &amp; Dey 2020, sp. nov." docType="treatment" docUuid="9931A78C-EE3F-4BF2-8FC1-EE213C98D838" docUuidSource="ZooBank" docVersion="13" lastPageNumber="578" masterDocId="FFAA7E56FFE16228FF85FFBAD143FFF3" masterDocTitle="Indian Hecalina (Hemiptera: Cicadellidae: Deltocephalinae: Hecalini) with description of three new species" masterLastPageNumber="585" masterPageNumber="573" pageNumber="576" updateTime="1698847506462" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5417ED340475675B0A7C4F809B4981D1">Indian Hecalina (Hemiptera: Cicadellidae: Deltocephalinae: Hecalini) with description of three new species</mods:title>
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<mods:namePart id="79279A5D6B282CFFF7981E39EA936A19">Nikoshe, Akash P.</mods:namePart>
<mods:affiliation id="3E8154D0B5936ACF4F05A32D0AB7EBDF">Division of Entomology, Indian Council of Agricultural Research-Indian Agricultural Research Institute, New Delhi 110012, INDIA http: // orcid. org / 0000 - 0002 - 2783 - 2240</mods:affiliation>
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<mods:namePart id="85707E7892DAAC9ADD1AA52003299F53">Meshram, Naresh M.</mods:namePart>
<mods:affiliation id="921CE91633DEC7684788D160DF1ED19E">Division of Entomology, Indian Council of Agricultural Research-Indian Agricultural Research Institute, New Delhi 110012, INDIA http: // orcid. org / 0000 - 0002 - 2783 - 2240 &amp; https: // orcid. org / 0000 - 0002 - 7020 - 2084</mods:affiliation>
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<mods:namePart id="A8CD79B00437BB7D74C1F74AA6F6A55D">Stuti</mods:namePart>
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<mods:namePart id="ED81F73A497924768536FDD144924FD7">Dey, Debjani</mods:namePart>
<mods:affiliation id="63207CA1A79FDDC9BA77C09AF4AE7A74">Division of Entomology, Indian Council of Agricultural Research-Indian Agricultural Research Institute, New Delhi 110012, INDIA http: // orcid. org / 0000 - 0002 - 2783 - 2240 &amp; https: // orcid. org / 0000 - 0002 - 6956 - 239 X</mods:affiliation>
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<mods:title id="17B3E2D37BAF5B6163F8C6CED5308152">Zootaxa</mods:title>
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<mods:date id="1634DA788D5CEF4B62A05B667EB66BDD">2020</mods:date>
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<mods:number id="DD2799EA7595ADE1B9C7CEE896475D72">2020-11-20</mods:number>
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<mods:number id="659C1E0FD5563A879DCC46FF1C572920">4881</mods:number>
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<mods:identifier id="74560CA73A93F453CF46C41E6E34DBB2" type="DOI">10.11646/zootaxa.4881.3.9</mods:identifier>
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<treatment id="0393062EFFE2622EFF12F9A2D30EFEB2" ID-DOI="http://doi.org/10.5281/zenodo.4338242" ID-GBIF-Taxon="170532532" ID-Zenodo-Dep="4338242" LSID="urn:lsid:zoobank.org:act:9931A78C-EE3F-4BF2-8FC1-EE213C98D838" httpUri="http://treatment.plazi.org/id/0393062EFFE2622EFF12F9A2D30EFEB2" lastPageId="6" lastPageNumber="578" pageId="3" pageNumber="576">
<subSubSection id="C320E4B3FFE2622BFF12F9A2D3A4F9C0" box="[151,743,1560,1587]" pageId="3" pageNumber="576" type="nomenclature">
<paragraph id="8B85B738FFE2622BFF12F9A2D3A4F9C0" blockId="3.[151,743,1560,1623]" box="[151,743,1560,1587]" pageId="3" pageNumber="576">
<heading id="D0CD0054FFE2622BFF12F9A2D3A4F9C0" bold="true" box="[151,743,1560,1587]" fontSize="11" level="1" pageId="3" pageNumber="576" reason="1">
<emphasis id="B94E6B2AFFE2622BFF12F9A2D3A4F9C0" bold="true" box="[151,743,1560,1587]" pageId="3" pageNumber="576">
<taxonomicName id="4C3ACCBBFFE2622BFF12F9A2D33EF9C0" ID-CoL="8RRQY" authority="Nikoshe &amp; Meshram" authorityName="Nikoshe &amp; Meshram &amp; Dey" authorityYear="2020" box="[151,637,1560,1587]" class="Insecta" family="Cicadellidae" genus="Hecalus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="shanayai" status="sp. nov.">
<emphasis id="B94E6B2AFFE2622BFF12F9A2D02CF9C0" bold="true" box="[151,367,1560,1587]" italics="true" pageId="3" pageNumber="576">Hecalus shanayai</emphasis>
Nikoshe &amp; Meshram
</taxonomicName>
,
<taxonomicNameLabel id="A27DD651FFE2622BFD0FF9A3D3A4F9C0" box="[650,743,1561,1587]" pageId="3" pageNumber="576" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C320E4B3FFE2622DFF12F987D41EF8B3" lastPageId="5" lastPageNumber="578" pageId="3" pageNumber="576" type="description">
<paragraph id="8B85B738FFE2622BFF12F987D0D3F9A4" blockId="3.[151,743,1560,1623]" box="[151,400,1597,1623]" pageId="3" pageNumber="576">
(
<figureCitation id="1301ABBDFFE2622BFF1AF987D076F9A4" box="[159,309,1597,1623]" captionStart="FIGURES 1–15" captionStartId="4.[152,270,1794,1818]" captionTargetBox="[175,1410,191,1759]" captionTargetId="figure-24@4.[171,1417,179,1768]" captionTargetPageId="4" captionText="FIGURES 1–15. Males of Hecalina species 1–5. Dorsal habitus 1. Hecalus ghaurii Rao &amp; Ramakrishnan; 2. H. lutescens (Distant); 3. H. shanayai sp. nov.; 4. H. tumidus sp. nov.; 5. Thomsonia assymmetricus sp. nov.; 6–10. Lateral habitus. 6. Hecalus ghaurii Rao &amp; Ramakrishnan; 7. H. lutescens (Distant); 8. H. shanayai sp. nov.; 9. H. tumidus sp. nov.; 10. T. assymmetricus sp. nov.; 11–15. Face 11. Hecalus ghaurii Rao &amp; Ramakrishnan; 12. H. lutescens (Distant); 13. H. shanayai sp. nov.; 14. H. tumidus sp. nov.; 15. T. assymmetricus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4283984" httpUri="https://zenodo.org/record/4283984/files/figure.png" pageId="3" pageNumber="576">Figs. 3, 8, 13</figureCitation>
,
<figureCitation id="1301ABBDFFE2622BFEC4F987D0CAF9A4" box="[321,393,1597,1623]" captionStart="FIGURES 34–40" captionStartId="7.[152,270,1977,2001]" captionTargetBox="[173,1404,643,1940]" captionTargetId="figure-210@7.[169,1418,633,1953]" captionTargetPageId="7" captionText="FIGURES 34–40. Genitalia male Hecalus shanayai sp. nov. 34. Pygofer; 35. Subgenital plate; 36. Style; 37. Aedeagus dorsal view; 38. Aedeagus, lateral view; 39. Aedeagus, ventral view; 40. Connective." figureDoi="http://doi.org/10.5281/zenodo.4283990" httpUri="https://zenodo.org/record/4283990/files/figure.png" pageId="3" pageNumber="576">34–40</figureCitation>
)
</paragraph>
<paragraph id="8B85B738FFE2622BFF12F93FD0A9F8F8" blockId="3.[151,1437,1669,2020]" pageId="3" pageNumber="576">
Male (
<figureCitation id="1301ABBDFFE2622BFF5AF93FD004F96C" box="[223,327,1669,1695]" captionStart="FIGURES 1–15" captionStartId="4.[152,270,1794,1818]" captionTargetBox="[175,1410,191,1759]" captionTargetId="figure-24@4.[171,1417,179,1768]" captionTargetPageId="4" captionText="FIGURES 1–15. Males of Hecalina species 1–5. Dorsal habitus 1. Hecalus ghaurii Rao &amp; Ramakrishnan; 2. H. lutescens (Distant); 3. H. shanayai sp. nov.; 4. H. tumidus sp. nov.; 5. Thomsonia assymmetricus sp. nov.; 6–10. Lateral habitus. 6. Hecalus ghaurii Rao &amp; Ramakrishnan; 7. H. lutescens (Distant); 8. H. shanayai sp. nov.; 9. H. tumidus sp. nov.; 10. T. assymmetricus sp. nov.; 11–15. Face 11. Hecalus ghaurii Rao &amp; Ramakrishnan; 12. H. lutescens (Distant); 13. H. shanayai sp. nov.; 14. H. tumidus sp. nov.; 15. T. assymmetricus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4283984" httpUri="https://zenodo.org/record/4283984/files/figure.png" pageId="3" pageNumber="576">Figs. 3, 8</figureCitation>
) Color yellowish green to green. Anterior margin of head with transverse submarginal fuscous line, face (
<figureCitation id="1301ABBDFFE2622BFF53F913D064F930" box="[214,295,1705,1731]" captionStart="FIGURES 1–15" captionStartId="4.[152,270,1794,1818]" captionTargetBox="[175,1410,191,1759]" captionTargetId="figure-24@4.[171,1417,179,1768]" captionTargetPageId="4" captionText="FIGURES 1–15. Males of Hecalina species 1–5. Dorsal habitus 1. Hecalus ghaurii Rao &amp; Ramakrishnan; 2. H. lutescens (Distant); 3. H. shanayai sp. nov.; 4. H. tumidus sp. nov.; 5. Thomsonia assymmetricus sp. nov.; 6–10. Lateral habitus. 6. Hecalus ghaurii Rao &amp; Ramakrishnan; 7. H. lutescens (Distant); 8. H. shanayai sp. nov.; 9. H. tumidus sp. nov.; 10. T. assymmetricus sp. nov.; 11–15. Face 11. Hecalus ghaurii Rao &amp; Ramakrishnan; 12. H. lutescens (Distant); 13. H. shanayai sp. nov.; 14. H. tumidus sp. nov.; 15. T. assymmetricus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4283984" httpUri="https://zenodo.org/record/4283984/files/figure.png" pageId="3" pageNumber="576">Fig. 13</figureCitation>
) brown, gena black, frontoclypeus transverse black striae invisible in middle. Compound eyes black, ocelli yellow. Forewings subhyaline, with diffuse fuscous transverse band across costal margin. Bases of tibial macrosetae and tarsi light brown.
</paragraph>
<paragraph id="8B85B738FFE2622BFF42F8AFD408F868" blockId="3.[151,1437,1669,2020]" pageId="3" pageNumber="576">Head including eyes as long as width of pronotum. Head produced in front, crown length 0.7x shorter than width between eyes. Face as long as wide; anterior margin of pronotum not extending beyond eyes, more or less parallel with eyes. Frontoclypeus longer than wide; frontal suture, terminating laterad of ocelli. Ocelli near anterior margin, very closely oppressed to eyes. Pronotum length 0.2x as long as wide and 0.9x length of scutellum.</paragraph>
<paragraph id="8B85B738FFE2622DFF42F81FD1ADF8EF" blockId="3.[151,1437,1669,2020]" lastBlockId="5.[151,1437,1686,2037]" lastPageId="5" lastPageNumber="578" pageId="3" pageNumber="576">
<emphasis id="B94E6B2AFFE2622BFF42F81FD034F84C" bold="true" box="[199,375,1957,1983]" pageId="3" pageNumber="576">Male genitalia.</emphasis>
Pygofer (
<figureCitation id="1301ABBDFFE2622BFE66F81FD370F84C" box="[483,563,1957,1983]" captionStart="FIGURES 34–40" captionStartId="7.[152,270,1977,2001]" captionTargetBox="[173,1404,643,1940]" captionTargetId="figure-210@7.[169,1418,633,1953]" captionTargetPageId="7" captionText="FIGURES 34–40. Genitalia male Hecalus shanayai sp. nov. 34. Pygofer; 35. Subgenital plate; 36. Style; 37. Aedeagus dorsal view; 38. Aedeagus, lateral view; 39. Aedeagus, ventral view; 40. Connective." figureDoi="http://doi.org/10.5281/zenodo.4283990" httpUri="https://zenodo.org/record/4283990/files/figure.png" pageId="3" pageNumber="576">Fig. 34</figureCitation>
) longer than wide, posterior margin triangular, with posterior ½ densely setose. Style (
<figureCitation id="1301ABBDFFE2622BFF65F873D073F817" box="[224,304,1993,2020]" captionStart="FIGURES 34–40" captionStartId="7.[152,270,1977,2001]" captionTargetBox="[173,1404,643,1940]" captionTargetId="figure-210@7.[169,1418,633,1953]" captionTargetPageId="7" captionText="FIGURES 34–40. Genitalia male Hecalus shanayai sp. nov. 34. Pygofer; 35. Subgenital plate; 36. Style; 37. Aedeagus dorsal view; 38. Aedeagus, lateral view; 39. Aedeagus, ventral view; 40. Connective." figureDoi="http://doi.org/10.5281/zenodo.4283990" httpUri="https://zenodo.org/record/4283990/files/figure.png" pageId="3" pageNumber="576">Fig. 36</figureCitation>
) with preapical lobe obtusely angulate, with few microsetae, apophysis short, 0.25 of the total length. Connective (
<figureCitation id="1301ABBDFFE4622DFEADF92CD038F943" box="[296,379,1686,1712]" captionStart="FIGURES 34–40" captionStartId="7.[152,270,1977,2001]" captionTargetBox="[173,1404,643,1940]" captionTargetId="figure-210@7.[169,1418,633,1953]" captionTargetPageId="7" captionText="FIGURES 34–40. Genitalia male Hecalus shanayai sp. nov. 34. Pygofer; 35. Subgenital plate; 36. Style; 37. Aedeagus dorsal view; 38. Aedeagus, lateral view; 39. Aedeagus, ventral view; 40. Connective." figureDoi="http://doi.org/10.5281/zenodo.4283990" httpUri="https://zenodo.org/record/4283990/files/figure.png" pageId="5" pageNumber="578">Fig. 40</figureCitation>
) with stem 2x shorter than arms. Aedeagal shaft narrowed apically, broad medially in lateral view, foliate in apical 0.2 with sparsely serrated margin, dorsal margin with short subapical tooth, with pair of slender apical processes half as long as shaft extended anterolaterad, gonopore subapical on ventral margin (
<figureCitation id="1301ABBDFFE4622DFAE1F964D1A2F8EF" captionStart="FIGURES 34–40" captionStartId="7.[152,270,1977,2001]" captionTargetBox="[173,1404,643,1940]" captionTargetId="figure-210@7.[169,1418,633,1953]" captionTargetPageId="7" captionText="FIGURES 34–40. Genitalia male Hecalus shanayai sp. nov. 34. Pygofer; 35. Subgenital plate; 36. Style; 37. Aedeagus dorsal view; 38. Aedeagus, lateral view; 39. Aedeagus, ventral view; 40. Connective." figureDoi="http://doi.org/10.5281/zenodo.4283990" httpUri="https://zenodo.org/record/4283990/files/figure.png" pageId="5" pageNumber="578">Figs. 37–39</figureCitation>
).
</paragraph>
<caption id="DF45E7B0FFE5622CFF1DF8B8D334F858" ID-DOI="http://doi.org/10.5281/zenodo.4283984" ID-Zenodo-Dep="4283984" httpUri="https://zenodo.org/record/4283984/files/figure.png" pageId="4" pageNumber="577" startId="4.[152,270,1794,1818]" targetBox="[175,1410,191,1759]" targetPageId="4">
<paragraph id="8B85B738FFE5622CFF1DF8B8D334F858" blockId="4.[151,1437,1794,1963]" pageId="4" pageNumber="577">
<emphasis id="B94E6B2AFFE5622CFF1DF8B8D00FF8E8" bold="true" box="[152,332,1794,1819]" pageId="4" pageNumber="577">FIGURES 1–15.</emphasis>
Males of
<taxonomicName id="4C3ACCBBFFE5622CFE2AF8B9D34FF8E8" box="[431,524,1795,1819]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="576" phylum="Arthropoda" rank="subTribe" subTribe="Hecalina">
<emphasis id="B94E6B2AFFE5622CFE2AF8B9D34FF8E8" box="[431,524,1795,1819]" italics="true" pageId="4" pageNumber="577">Hecalina</emphasis>
</taxonomicName>
species 1–5. Dorsal habitus 1.
<taxonomicName id="4C3ACCBBFFE5622CFCC4F8B9D584F8E8" authority="Rao &amp; Ramakrishnan" authorityName="Rama Subba Rao &amp; Ramakrishnan" authorityYear="1990" box="[833,1223,1795,1819]" class="Insecta" family="Cicadellidae" genus="Hecalus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="ghaurii">
<emphasis id="B94E6B2AFFE5622CFCC4F8B9D2A6F8E8" box="[833,997,1795,1819]" italics="true" pageId="4" pageNumber="577">Hecalus ghaurii</emphasis>
Rao &amp; Ramakrishnan
</taxonomicName>
; 2.
<taxonomicName id="4C3ACCBBFFE5622CFB6DF8B9D18BF8CC" authority="(Distant)" baseAuthorityName="Distant" class="Insecta" family="Cicadellidae" genus="Parabolocratus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="lutescens">
<emphasis id="B94E6B2AFFE5622CFB6DF8B9D420F8E8" box="[1256,1379,1795,1819]" italics="true" pageId="4" pageNumber="577">H. lutescens</emphasis>
(Distant)
</taxonomicName>
; 3.
<taxonomicName id="4C3ACCBBFFE5622CFF68F89DD02BF8CC" authority="Nikoshe &amp; Meshram &amp; Dey, 2020" authorityName="Nikoshe &amp; Meshram &amp; Dey" authorityYear="2020" box="[237,360,1831,1855]" class="Insecta" family="Cicadellidae" genus="Hecalus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="shanayai" status="sp. nov.">
<emphasis id="B94E6B2AFFE5622CFF68F89DD02BF8CC" box="[237,360,1831,1855]" italics="true" pageId="4" pageNumber="577">H. shanayai</emphasis>
</taxonomicName>
<emphasis id="B94E6B2AFFE5622CFEEAF89DD083F8CC" bold="true" box="[367,448,1831,1855]" pageId="4" pageNumber="577">
<taxonomicNameLabel id="A27DD651FFE5622CFEEAF89DD083F8CC" box="[367,448,1831,1855]" pageId="4" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 4.
<taxonomicName id="4C3ACCBBFFE5622CFE6DF89DD31BF8CC" authority="Nikoshe &amp; Meshram &amp; Dey, 2020" authorityName="Nikoshe &amp; Meshram &amp; Dey" authorityYear="2020" box="[488,600,1831,1855]" class="Insecta" family="Cicadellidae" genus="Hecalus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="tumidus" status="sp. nov.">
<emphasis id="B94E6B2AFFE5622CFE6DF89DD31BF8CC" box="[488,600,1831,1855]" italics="true" pageId="4" pageNumber="577">H. tumidus</emphasis>
</taxonomicName>
<emphasis id="B94E6B2AFFE5622CFDDAF89DD3F3F8CC" bold="true" box="[607,688,1831,1855]" pageId="4" pageNumber="577">
<taxonomicNameLabel id="A27DD651FFE5622CFDDAF89DD3F3F8CC" box="[607,688,1831,1855]" pageId="4" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 5.
<taxonomicName id="4C3ACCBBFFE5622CFD5DF89DD2A7F8CD" authority="Nikoshe &amp; Meshram &amp; Dey, 2020" authorityName="Nikoshe &amp; Meshram &amp; Dey" authorityYear="2020" box="[728,996,1831,1855]" class="Insecta" family="Cicadellidae" genus="Thomsonia" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="assymmetricus" status="sp. nov.">
<emphasis id="B94E6B2AFFE5622CFD5DF89DD2A7F8CD" box="[728,996,1831,1855]" italics="true" pageId="4" pageNumber="577">Thomsonia assymmetricus</emphasis>
</taxonomicName>
<emphasis id="B94E6B2AFFE5622CFC6EF89DD57FF8CC" bold="true" box="[1003,1084,1831,1855]" pageId="4" pageNumber="577">
<taxonomicNameLabel id="A27DD651FFE5622CFC6EF89DD57FF8CC" box="[1003,1084,1831,1855]" pageId="4" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 6–10. Lateral habitus. 6.
<taxonomicName id="4C3ACCBBFFE5622CFACCF89DD089F890" authority="Rao &amp; Ramakrishnan" authorityName="Rama Subba Rao &amp; Ramakrishnan" authorityYear="1990" class="Insecta" family="Cicadellidae" genus="Hecalus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="ghaurii">
<emphasis id="B94E6B2AFFE5622CFACCF89DD1A1F890" italics="true" pageId="4" pageNumber="577">Hecalus ghaurii</emphasis>
Rao &amp; Ramakrishnan
</taxonomicName>
; 7.
<taxonomicName id="4C3ACCBBFFE5622CFE6BF8F1D38CF890" authority="(Distant)" baseAuthorityName="Distant" box="[494,719,1867,1891]" class="Insecta" family="Cicadellidae" genus="Parabolocratus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="lutescens">
<emphasis id="B94E6B2AFFE5622CFE6BF8F1D328F890" box="[494,619,1867,1891]" italics="true" pageId="4" pageNumber="577">H. lutescens</emphasis>
(Distant)
</taxonomicName>
; 8.
<taxonomicName id="4C3ACCBBFFE5622CFD71F8F1D22CF890" authority="Nikoshe &amp; Meshram &amp; Dey, 2020" authorityName="Nikoshe &amp; Meshram &amp; Dey" authorityYear="2020" box="[756,879,1867,1891]" class="Insecta" family="Cicadellidae" genus="Hecalus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="shanayai" status="sp. nov.">
<emphasis id="B94E6B2AFFE5622CFD71F8F1D22CF890" box="[756,879,1867,1891]" italics="true" pageId="4" pageNumber="577">H. shanayai</emphasis>
</taxonomicName>
<emphasis id="B94E6B2AFFE5622CFCF3F8F1D284F890" bold="true" box="[886,967,1867,1891]" pageId="4" pageNumber="577">
<taxonomicNameLabel id="A27DD651FFE5622CFCF3F8F1D284F890" box="[886,967,1867,1891]" pageId="4" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 9.
<taxonomicName id="4C3ACCBBFFE5622CFC75F8F1D523F890" authority="Nikoshe &amp; Meshram &amp; Dey, 2020" authorityName="Nikoshe &amp; Meshram &amp; Dey" authorityYear="2020" box="[1008,1120,1867,1891]" class="Insecta" family="Cicadellidae" genus="Hecalus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="tumidus" status="sp. nov.">
<emphasis id="B94E6B2AFFE5622CFC75F8F1D523F890" box="[1008,1120,1867,1891]" italics="true" pageId="4" pageNumber="577">H. tumidus</emphasis>
</taxonomicName>
<emphasis id="B94E6B2AFFE5622CFBE2F8F1D5FBF890" bold="true" box="[1127,1208,1867,1891]" pageId="4" pageNumber="577">
<taxonomicNameLabel id="A27DD651FFE5622CFBE2F8F1D5FBF890" box="[1127,1208,1867,1891]" pageId="4" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 10.
<taxonomicName id="4C3ACCBBFFE5622CFB68F8F1D4DFF891" authority="Nikoshe &amp; Meshram &amp; Dey, 2020" authorityName="Nikoshe &amp; Meshram &amp; Dey" authorityYear="2020" box="[1261,1436,1867,1890]" class="Insecta" family="Cicadellidae" genus="Thomsonia" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="assymmetricus" status="sp. nov.">
<emphasis id="B94E6B2AFFE5622CFB68F8F1D4DFF891" box="[1261,1436,1867,1890]" italics="true" pageId="4" pageNumber="577">T. assymmetricus</emphasis>
</taxonomicName>
<emphasis id="B94E6B2AFFE5622CFF12F8D5D1AAF874" bold="true" box="[151,233,1903,1927]" pageId="4" pageNumber="577">
<taxonomicNameLabel id="A27DD651FFE5622CFF12F8D5D1AAF874" box="[151,233,1903,1927]" pageId="4" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 11–15. Face 11.
<taxonomicName id="4C3ACCBBFFE5622CFE27F8D5D272F874" authority="Rao &amp; Ramakrishnan" authorityName="Rama Subba Rao &amp; Ramakrishnan" authorityYear="1990" box="[418,817,1903,1927]" class="Insecta" family="Cicadellidae" genus="Hecalus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="ghaurii">
<emphasis id="B94E6B2AFFE5622CFE27F8D5D30BF874" box="[418,584,1903,1927]" italics="true" pageId="4" pageNumber="577">Hecalus ghaurii</emphasis>
Rao &amp; Ramakrishnan
</taxonomicName>
; 12.
<taxonomicName id="4C3ACCBBFFE5622CFCE6F8D5D506F874" authority="(Distant)" baseAuthorityName="Distant" box="[867,1093,1903,1927]" class="Insecta" family="Cicadellidae" genus="Parabolocratus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="lutescens">
<emphasis id="B94E6B2AFFE5622CFCE6F8D5D2A3F874" box="[867,992,1903,1927]" italics="true" pageId="4" pageNumber="577">H. lutescens</emphasis>
(Distant)
</taxonomicName>
; 13.
<taxonomicName id="4C3ACCBBFFE5622CFBFDF8D5D5B0F874" authority="Nikoshe &amp; Meshram &amp; Dey, 2020" authorityName="Nikoshe &amp; Meshram &amp; Dey" authorityYear="2020" box="[1144,1267,1903,1927]" class="Insecta" family="Cicadellidae" genus="Hecalus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="shanayai" status="sp. nov.">
<emphasis id="B94E6B2AFFE5622CFBFDF8D5D5B0F874" box="[1144,1267,1903,1927]" italics="true" pageId="4" pageNumber="577">H. shanayai</emphasis>
</taxonomicName>
<emphasis id="B94E6B2AFFE5622CFB7EF8D5D40EF874" bold="true" box="[1275,1357,1903,1927]" pageId="4" pageNumber="577">
<taxonomicNameLabel id="A27DD651FFE5622CFB7EF8D5D40EF874" box="[1275,1357,1903,1927]" pageId="4" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 14.
<taxonomicName id="4C3ACCBBFFE5622CFA01F8D5D1A4F858" authority="Nikoshe &amp; Meshram &amp; Dey, 2020" authorityName="Nikoshe &amp; Meshram &amp; Dey" authorityYear="2020" class="Insecta" family="Cicadellidae" genus="Hecalus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="tumidus" status="sp. nov.">
<emphasis id="B94E6B2AFFE5622CFA01F8D5D1A4F858" italics="true" pageId="4" pageNumber="577">H. tumidus</emphasis>
</taxonomicName>
<emphasis id="B94E6B2AFFE5622CFF6BF829D07DF858" bold="true" box="[238,318,1939,1963]" pageId="4" pageNumber="577">
<taxonomicNameLabel id="A27DD651FFE5622CFF6BF829D07DF858" box="[238,318,1939,1963]" pageId="4" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 15.
<taxonomicName id="4C3ACCBBFFE5622CFEF7F829D363F859" authority="Nikoshe &amp; Meshram &amp; Dey, 2020" authorityName="Nikoshe &amp; Meshram &amp; Dey" authorityYear="2020" box="[370,544,1939,1962]" class="Insecta" family="Cicadellidae" genus="Thomsonia" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="assymmetricus" status="sp. nov.">
<emphasis id="B94E6B2AFFE5622CFEF7F829D363F859" box="[370,544,1939,1962]" italics="true" pageId="4" pageNumber="577">T. assymmetricus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DD651FFE5622CFDA3F829D334F858" box="[550,631,1939,1963]" pageId="4" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="DF45E7B0FFE4622DFF1DF991D4CAF99B" ID-DOI="http://doi.org/10.5281/zenodo.4283986" ID-Zenodo-Dep="4283986" httpUri="https://zenodo.org/record/4283986/files/figure.png" pageId="5" pageNumber="578" startId="5.[152,270,1579,1603]" targetBox="[185,1404,184,1549]" targetPageId="5">
<paragraph id="8B85B738FFE4622DFF1DF991D4CAF99B" blockId="5.[151,1436,1579,1640]" pageId="5" pageNumber="578">
<emphasis id="B94E6B2AFFE4622DFF1DF991D018F9B7" bold="true" box="[152,347,1579,1604]" pageId="5" pageNumber="578">FIGURES 16–26.</emphasis>
Genitalia male
<taxonomicName id="4C3ACCBBFFE4622DFE78F996D2C7F9B7" authority="Rao &amp; Ramakrishnan" authorityName="Rama Subba Rao &amp; Ramakrishnan" authorityYear="1990" box="[509,900,1580,1604]" class="Insecta" family="Cicadellidae" genus="Hecalus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="ghaurii">
<emphasis id="B94E6B2AFFE4622DFE78F996D3E1F9B7" box="[509,674,1580,1604]" italics="true" pageId="5" pageNumber="578">Hecalus ghaurii</emphasis>
Rao &amp; Ramakrishnan
</taxonomicName>
16. Pygofer; 17. Subgenital plate; 18. Style; 19. Aedeagus dorsal view; 20. Aedeagus, lateral view; 21. Aedeagus, ventral view; 22. Connective; 23–26. Label data on Holotype.
</paragraph>
</caption>
<paragraph id="8B85B738FFE4622DFF42F89CD41EF8B3" blockId="5.[151,1437,1686,2037]" box="[199,1373,1830,1856]" pageId="5" pageNumber="578">
<emphasis id="B94E6B2AFFE4622DFF42F89CD086F8B3" bold="true" box="[199,453,1830,1856]" pageId="5" pageNumber="578">Measurements (mm).</emphasis>
Male 4.63 long, 1.3 wide across eyes, 1.01 wide across hind margin of pronotum.
</paragraph>
</subSubSection>
<subSubSection id="C320E4B3FFE4622DFF42F8F0D2A9F87B" pageId="5" pageNumber="578" type="materials_examined">
<paragraph id="8B85B738FFE4622DFF42F8F0D2A9F87B" blockId="5.[151,1437,1686,2037]" pageId="5" pageNumber="578">
<emphasis id="B94E6B2AFFE4622DFF42F8F0D036F897" bold="true" box="[199,373,1866,1892]" pageId="5" pageNumber="578">Type material.</emphasis>
<materialsCitation id="3B52BD65FFE4622DFEF8F8F0D0BEF87B" ID-GBIF-Occurrence="2985584305" collectingDate="2015-09-22" collectionCode="NPC" collectorName="Coll. Akash Nikoshe" country="India" latitude="21.403" location="Chikhaldara" longLatPrecision="7" longitude="77.3268" pageId="5" pageNumber="578" specimenCount="1" stateProvince="Maharashtra" typeStatus="holotype">
<typeStatus id="5481099AFFE4622DFEF8F8F0D0A6F897" box="[381,485,1866,1892]" pageId="5" pageNumber="578" type="holotype">Holotype</typeStatus>
³,
<collectingCountry id="F32DF7A8FFE4622DFD95F8F0D327F897" box="[528,612,1866,1892]" name="India" pageId="5" pageNumber="578">INDIA</collectingCountry>
:
<collectingRegion id="49FE79DAFFE4622DFDEAF8F0D3BCF897" box="[623,767,1866,1892]" country="India" name="Maharashtra" pageId="5" pageNumber="578">Maharashtra</collectingRegion>
:
<location id="8EE5E1E3FFE4622DFC8FF8F0D2D0F897" LSID="urn:lsid:plazi:treatment:0393062EFFE2622EFF12F9A2D30EFEB2:8EE5E1E3FFE4622DFC8FF8F0D2D0F897" box="[778,915,1866,1892]" country="India" latitude="21.403" longLatPrecision="7" longitude="77.3268" name="Chikhaldara" pageId="5" pageNumber="578" stateProvince="Maharashtra">Chikhaldara</location>
(
<geoCoordinate id="EE0ED1FFFFE4622DFC27F8F0D56AF897" box="[930,1065,1866,1892]" degrees="21.4030" direction="north" orientation="latitude" pageId="5" pageNumber="578" precision="5" value="21.403">21.4030° N</geoCoordinate>
,
<geoCoordinate id="EE0ED1FFFFE4622DFBB1F8F0D5F5F897" box="[1076,1206,1866,1892]" degrees="77.3268" direction="east" orientation="longitude" pageId="5" pageNumber="578" precision="5" value="77.3268">77.3268° E</geoCoordinate>
),
<date id="FF8491F8FFE4622DFB4EF8F0D406F896" box="[1227,1349,1866,1893]" pageId="5" pageNumber="578" value="2015-09-22">
<collectingDate id="EFC06810FFE4622DFB4EF8F0D406F896" box="[1227,1349,1866,1893]" pageId="5" pageNumber="578" value="2015-09-22">22.ix.2015</collectingDate>
</date>
,
<collectingMethod id="527BCF2FFFE4622DFAD7F8F0D1FAF87B" pageId="5" pageNumber="578">Sweep net</collectingMethod>
<collectorName id="26CFD2EEFFE4622DFF45F8D4D0EAF87B" box="[192,425,1902,1928]" pageId="5" pageNumber="578">Coll. Akash Nikoshe</collectorName>
(
<collectionCode id="ED2B2FFDFFE4622DFE3DF8D4D0B6F87B" box="[440,501,1902,1928]" country="India" httpUri="http://grbio.org/cool/nwky-2b63" name="National Pusa Collection" pageId="5" pageNumber="578">NPC</collectionCode>
)
</materialsCitation>
.
<typeStatus id="5481099AFFE4622DFD82F8D4D334F87B" box="[519,631,1902,1928]" pageId="5" pageNumber="578" type="paratype">Paratypes</typeStatus>
, 6 ³ with same data as holotype.
</paragraph>
</subSubSection>
<subSubSection id="C320E4B3FFE4622DFF42F828D2CAF823" pageId="5" pageNumber="578" type="etymology">
<paragraph id="8B85B738FFE4622DFF42F828D2CAF823" blockId="5.[151,1437,1686,2037]" pageId="5" pageNumber="578">
<emphasis id="B94E6B2AFFE4622DFF42F828D00CF85F" bold="true" box="[199,335,1938,1964]" pageId="5" pageNumber="578">Etymology.</emphasis>
This species is named in honour of “Miss Shanaya” (daughter of the corresponding author) for her support to the author to pursue his passion in leafhopper taxonomy.
</paragraph>
</subSubSection>
<subSubSection id="C320E4B3FFE4622EFF42F860D30EFEB2" lastPageId="6" lastPageNumber="579" pageId="5" pageNumber="578" type="discussion">
<paragraph id="8B85B738FFE4622EFF42F860D30EFEB2" blockId="5.[151,1437,1686,2037]" lastBlockId="6.[151,1437,151,321]" lastPageId="6" lastPageNumber="579" pageId="5" pageNumber="578">
<emphasis id="B94E6B2AFFE4622DFF42F860D078F807" bold="true" box="[199,315,2010,2036]" pageId="5" pageNumber="578">Remarks.</emphasis>
<taxonomicName id="4C3ACCBBFFE4622DFEC1F860D34DF807" authority="Nikoshe &amp; Meshram &amp; Dey, 2020" authorityName="Nikoshe &amp; Meshram &amp; Dey" authorityYear="2020" box="[324,526,2010,2036]" class="Insecta" family="Cicadellidae" genus="Hecalus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="shanayai" status="sp. nov.">
<emphasis id="B94E6B2AFFE4622DFEC1F860D34DF807" box="[324,526,2010,2036]" italics="true" pageId="5" pageNumber="578">Hecalus shanayai</emphasis>
</taxonomicName>
<emphasis id="B94E6B2AFFE4622DFD92F860D330F807" bold="true" box="[535,627,2010,2036]" pageId="5" pageNumber="578">
<taxonomicNameLabel id="A27DD651FFE4622DFD92F860D330F807" box="[535,627,2010,2036]" pageId="5" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
externally resembles
<taxonomicName id="4C3ACCBBFFE4622DFCEBF861D525F806" authority="(Distant)" baseAuthorityName="Distant" box="[878,1126,2010,2037]" class="Insecta" family="Cicadellidae" genus="Parabolocratus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="lutescens">
<emphasis id="B94E6B2AFFE4622DFCEBF861D2BAF807" box="[878,1017,2010,2036]" italics="true" pageId="5" pageNumber="578">H. lutescens</emphasis>
(Distant)
</taxonomicName>
(
<figureCitation id="1301ABBDFFE4622DFBF2F860D451F807" box="[1143,1298,2010,2036]" captionStart="FIGURES 1–15" captionStartId="4.[152,270,1794,1818]" captionTargetBox="[175,1410,191,1759]" captionTargetId="figure-24@4.[171,1417,179,1768]" captionTargetPageId="4" captionText="FIGURES 1–15. Males of Hecalina species 1–5. Dorsal habitus 1. Hecalus ghaurii Rao &amp; Ramakrishnan; 2. H. lutescens (Distant); 3. H. shanayai sp. nov.; 4. H. tumidus sp. nov.; 5. Thomsonia assymmetricus sp. nov.; 6–10. Lateral habitus. 6. Hecalus ghaurii Rao &amp; Ramakrishnan; 7. H. lutescens (Distant); 8. H. shanayai sp. nov.; 9. H. tumidus sp. nov.; 10. T. assymmetricus sp. nov.; 11–15. Face 11. Hecalus ghaurii Rao &amp; Ramakrishnan; 12. H. lutescens (Distant); 13. H. shanayai sp. nov.; 14. H. tumidus sp. nov.; 15. T. assymmetricus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4283984" httpUri="https://zenodo.org/record/4283984/files/figure.png" pageId="5" pageNumber="578">Figs. 2, 7, 12</figureCitation>
,
<figureCitation id="1301ABBDFFE4622DFAA5F860D42BF807" box="[1312,1384,2010,2036]" captionStart="FIGURES 27–33" captionStartId="6.[152,270,1585,1609]" captionTargetBox="[202,1386,355,1550]" captionTargetId="figure-107@6.[192,1395,345,1562]" captionTargetPageId="6" captionText="FIGURES 27–33. Genitalia male Hecalus lutescens (Distant) 27. Pygofer; 28. Subgenital plate; 29. Style; 30. Aedeagus dorsal view; 31. Aedeagus, lateral view; 32. Aedeagus, ventral view; 33. Connective." figureDoi="http://doi.org/10.5281/zenodo.4283988" httpUri="https://zenodo.org/record/4283988/files/figure.png" pageId="5" pageNumber="578">27–33</figureCitation>
) but differs in having the aedeagal shaft foliate in the apical 0.2 with a sparsely serrated margin, the dorsal margin with a short subapical tooth, and lacking mid-dorsal lateral expansions which are more distinct in the latter. From
<taxonomicName id="4C3ACCBBFFE7622EFA04FF01D0B3FF0A" authority="Rao and Ramakrishnan" authorityName="Rao and Ramakrishnan" class="Insecta" family="Cicadellidae" genus="Hecalus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="579" phylum="Arthropoda" rank="species" species="ghauri">
<emphasis id="B94E6B2AFFE7622EFA04FF01D1A1FF0A" italics="true" pageId="6" pageNumber="579">H. ghauri</emphasis>
Rao and Ramakrishnan
</taxonomicName>
, (
<figureCitation id="1301ABBDFFE7622EFD87FF65D384FF0A" box="[514,711,223,249]" captionStart="FIGURES 16–26" captionStartId="5.[152,270,1579,1603]" captionTargetBox="[185,1404,184,1549]" captionTargetId="figure-19@5.[167,1419,181,1556]" captionTargetPageId="5" captionText="FIGURES 16–26. Genitalia male Hecalus ghaurii Rao &amp; Ramakrishnan 16. Pygofer; 17. Subgenital plate; 18. Style; 19. Aedeagus dorsal view; 20. Aedeagus, lateral view; 21. Aedeagus, ventral view; 22. Connective; 23–26. Label data on Holotype." figureDoi="http://doi.org/10.5281/zenodo.4283986" httpUri="https://zenodo.org/record/4283986/files/figure.png" pageId="6" pageNumber="579">Figs. 19, 20 &amp; 21</figureCitation>
) it can be distinguished by the aedeagal shaft foliate in the apical 0.2 with a sparsely serrated margin, the dorsal margin with a short subapical tooth, and lacking mid-dorsal lateral expansions more distinct in the former.
</paragraph>
</subSubSection>
</treatment>
</document>
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