treatments-xml/data/03/8F/F8/038FF87B8C50FFD2811DA130CF0997A5.xml

<document id="74096B4EB111AED19F251C67998154A4" ID-CLB-Dataset="24179" ID-DOI="10.5852/ejt.2020.600" ID-GBIF-Dataset="89ad98bf-c070-4f7d-8b85-fa67a52f0996" ID-Zenodo-Dep="3637773" ID-ZooBank="A0FAB2BB-5CEF-42CB-9014-41655BC12D79" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1580885958890" checkinUser="plazi" docAuthor="Stöhr, Sabine, Weber, Alexandra Anh-Thu, Boissin, Emilie &amp; Chenuil, Anne" docDate="2020" docId="038FF87B8C50FFD2811DA130CF0997A5" docLanguage="en" docName="ejt-600_stoehr_weber_boissin.pdf.imf" docOrigin="European Journal of Taxonomy 600" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ophioderma zibrowii Stöhr &amp; Weber &amp; Boissin &amp; Chenuil 2020, sp. nov." docType="treatment" docUuid="7FB245FD-C8DF-402A-A7B1-FA7D048AFB04" docUuidSource="ZooBank" docVersion="9" lastPageNumber="32" masterDocId="FFB680038C4BFFCD8374A277CE10947B" masterDocTitle="Resolving the Ophioderma longicauda (Echinodermata: Ophiuroidea) cryptic species complex: five sisters, three of them new" masterLastPageNumber="37" masterPageNumber="1" pageNumber="28" updateTime="1698780426651" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="3403D139A31029E101EB25BCC69D1C5B">Resolving the Ophioderma longicauda (Echinodermata: Ophiuroidea) cryptic species complex: five sisters, three of them new</mods:title>
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<mods:namePart id="D7787AEB86D151BEB0CFD23F028CDFB5">Stöhr, Sabine</mods:namePart>
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<mods:namePart id="969D02AF686553BFB0AD5E2ED9235ABD">Weber, Alexandra Anh-Thu</mods:namePart>
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<mods:namePart id="8C6809AF909AEAB9F8940E6D48E9324D">Boissin, Emilie</mods:namePart>
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<mods:namePart id="1A28A9E94C68FEF755B30F2700F6642F">Chenuil, Anne</mods:namePart>
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<treatment id="038FF87B8C50FFD2811DA130CF0997A5" ID-DOI="http://doi.org/10.5281/zenodo.3638047" ID-GBIF-Taxon="161944092" ID-Zenodo-Dep="3638047" LSID="urn:lsid:zoobank.org:act:7FB245FD-C8DF-402A-A7B1-FA7D048AFB04" httpUri="http://treatment.plazi.org/id/038FF87B8C50FFD2811DA130CF0997A5" lastPageId="31" lastPageNumber="32" pageId="27" pageNumber="28">
<subSubSection id="C33C1AE68C50FFD6811DA130CADE97FC" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="8B99496D8C50FFD6811DA130CDDA9719" blockId="27.[357,1230,839,938]" box="[617,970,839,866]" pageId="27" pageNumber="28">
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<taxonomicName id="4C2632EE8C50FFD6811DA130CD789719" authority="Stöhr &amp; Weber &amp; Boissin &amp; Chenuil, 2020" authorityName="Stöhr &amp; Weber &amp; Boissin &amp; Chenuil" authorityYear="2020" box="[617,872,839,866]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="27" pageNumber="28" phylum="Echinodermata" rank="species" species="zibrowii" status="sp. nov.">
<emphasis id="B952957F8C50FFD6811DA130CD789719" bold="true" box="[617,872,839,866]" italics="true" pageId="27" pageNumber="28">Ophioderma zibrowii</emphasis>
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urn:lsid:zoobank.org:act: 
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</paragraph>
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<subSubSection id="C33C1AE68C50FFD681D2A1E7CD9E97D1" box="[678,910,912,938]" pageId="27" pageNumber="28" type="description">
<paragraph id="8B99496D8C50FFD681D2A1E7CD9E97D1" blockId="27.[357,1230,839,938]" box="[678,910,912,938]" pageId="27" pageNumber="28">
<figureCitation id="131D55E88C50FFD681D2A1E7CD3B97D1" box="[678,811,912,938]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="28.[189,232,1895,1921]" captionStartId-1="30.[189,232,1718,1744]" captionTargetBox-0="[196,1391,717,1854]" captionTargetBox-1="[189,1398,630,1678]" captionTargetId-0="figure@28.[196,1391,717,1854]" captionTargetId-1="figure@30.[189,1398,630,1678]" captionTargetPageId-0="28" captionTargetPageId-1="30" captionText-0="Fig. 11. Ophioderma zibrowii sp. nov., holotype (SMNH-Type-9188), digital images. A. Dorsal aspect, note the naked radial shields. B. Ventral aspect. C. Ventral arm. D. Dorsal arm, note the entire plates. E. Jaw. F. Brooded juveniles in interradius (arrow). Abbreviations: DAP = dorsal arm plate; M = madreporite; OS = oral shield; RS = radial shield; VAP = ventral arm plate." captionText-1="Fig. 12. SEM images of skeletal elements of Ophioderma zibrowii sp. nov., from non-type specimen of 15.4 mm disc diameter (SMNH-178455), Crete, Greece. A. Radial shield, external aspect, white arrows point to larger peripheral pores. B. Radial shield, internal aspect. C. Scale-free part magnified from Fig. 12A, note the absence of pores on the denser area (naked in intact animal, see Fig. 11A). D. Dental plate fragments. E. Adradial genital plate, lateral aspect. F. Distal end of Fig. 12E. G. Abradial genital plate. H. Oral plate, abradial face. I. Oral plate, adradial face, black arrows point to oral papillae sockets and pores. J. Arm vertebra, proximal face. K. Arm vertebra, distal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; Sp = spurs; vent = ventral." figureDoi-0="http://doi.org/10.5281/zenodo.3637795" figureDoi-1="http://doi.org/10.5281/zenodo.3637797" httpUri-0="https://zenodo.org/record/3637795/files/figure.png" httpUri-1="https://zenodo.org/record/3637797/files/figure.png" pageId="27" pageNumber="28">Figs 11–12</figureCitation>
; 
<tableCitation id="C6A47CD68C50FFD68043A1E7CD9E97D1" box="[823,910,912,938]" captionStart="Table 1" captionStartId="6.[182,251,686,714]" captionTargetBox="[192,1907,832,1596]" captionText="Table 1. Key characters that distinguish the species of Ophioderma Müller &amp; Troschel, 1840 in the Mediterranean Sea, the Northeast and Tropical Atlantic Ocean, formerly included in a cryptic complex under the name O. longicauda (Bruzelius, 1805). For O. guineense Greeff, 1882, the highest numbers of arm spines are from the syntypes. DAP = dorsal arm plate." httpUri="http://table.plazi.org/id/DF5919E58C4DFFCB83C2A0D9CDE1976A" pageId="27" pageNumber="28" tableUuid="DF5919E58C4DFFCB83C2A0D9CDE1976A">Table 1</tableCitation>
</paragraph>
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<heading id="D0D1FE018C50FFD683C9A1A5CF269797" bold="true" box="[189,310,978,1004]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="B952957F8C50FFD683C9A1A5CF269797" bold="true" box="[189,310,978,1004]" pageId="27" pageNumber="28">Diagnosis</emphasis>
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Usually naked radial shields. Seven arm spines. Eight oral papillae. Colour pattern greenish brown to dark green variegated on dorsal and ventral disc, arms banded, with light and dark spots along distal dorsal plate edges, oral frame and ventral arms lighter. Largest known size 
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dd. Brooder.
</paragraph>
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<subSubSection id="C33C1AE68C50FFD683C9A6FACF809165" pageId="27" pageNumber="28" type="etymology">
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<heading id="D0D1FE018C50FFD683C9A6FACF5490DC" bold="true" box="[189,324,1165,1191]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="B952957F8C50FFD683C9A6FACF5490DC" bold="true" box="[189,324,1165,1191]" pageId="27" pageNumber="28">Etymology</emphasis>
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The species is named after Helmut Zibrowius for his important contributions in collecting material for this study. The existence of brooding 
<taxonomicName id="4C2632EE8C50FFD681F3A697CD0B9081" box="[647,795,1248,1274]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="27" pageNumber="28" phylum="Echinodermata" rank="genus">
<emphasis id="B952957F8C50FFD681F3A697CD0B9081" box="[647,795,1248,1274]" italics="true" pageId="27" pageNumber="28">Ophioderma</emphasis>
</taxonomicName>
in the Mediterranean Sea might still be unknown without his effort.
</paragraph>
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<heading id="D0D1FE018C50FFD683C9A731CFBA911B" bold="true" box="[189,426,1350,1376]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="B952957F8C50FFD683C9A731CFBA911B" bold="true" box="[189,426,1350,1376]" pageId="27" pageNumber="28">Material examined</emphasis>
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<paragraph id="8B99496D8C50FFD6839EA701CF4C91EB" blockId="27.[189,1398,1398,1497]" box="[234,348,1398,1424]" pageId="27" pageNumber="28">
<emphasis id="B952957F8C50FFD6839EA701CF4C91EB" bold="true" box="[234,348,1398,1424]" pageId="27" pageNumber="28">
<typeStatus id="549DF7CF8C50FFD6839EA701CF4C91EB" box="[234,348,1398,1424]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B99496D8C50FFD683C9A7EDCCCD91A2" blockId="27.[189,1398,1398,1497]" pageId="27" pageNumber="28">
<materialsCitation id="3B4E43308C50FFD683C9A7EDCCCD91A2" ID-GBIF-Occurrence="2563048059" collectingDate="2012-05" collectorName="A. Weber" location="Agios Pavlos" pageId="27" pageNumber="28" specimenCount="1" specimenCount-female="1" stateProvince="Kriti" typeStatus="holotype">
MEDITERRANEAN SEA • 
<specimenCount id="9D2082E48C50FFD6816DA7ECCC3F91CE" box="[537,559,1435,1461]" pageId="27" pageNumber="28" type="female">♀</specimenCount>
(brooding); southern 
<collectingRegion id="49E2878F8C50FFD6804DA7EDCD6C91CF" box="[825,892,1434,1460]" country="Greece" name="Kriti" pageId="27" pageNumber="28">Crete</collectingRegion>
, 
<location id="8EF91FB68C50FFD680FDA7EDCA3C91CF" LSID="urn:lsid:plazi:treatment:038FF87B8C50FFD2811DA130CF0997A5:8EF91FB68C50FFD680FDA7EDCA3C91CF" box="[905,1068,1434,1460]" name="Agios Pavlos" pageId="27" pageNumber="28" stateProvince="Kriti">Agios Pavlos</location>
; 
<date id="FF986FAD8C50FFD6874EA7ECCAA991CF" box="[1082,1209,1434,1461]" pageId="27" pageNumber="28" value="2012-05">
<collectingDate id="EFDC96458C50FFD6874EA7ECCAA991CF" box="[1082,1209,1434,1461]" pageId="27" pageNumber="28" value="2012-05">May 2012</collectingDate>
</date>
; 
<collectorName id="26D32CBB8C50FFD687B2A7EDCB2991CF" box="[1222,1337,1434,1460]" pageId="27" pageNumber="28">A. Weber</collectorName>
leg.; preserved in 95% ethanol; SMNH-Type-9188.
</materialsCitation>
</paragraph>
<paragraph id="8B99496D8C50FFD6839EA476CF779260" blockId="27.[189,1399,1537,1672]" box="[234,359,1537,1563]" pageId="27" pageNumber="28">
<emphasis id="B952957F8C50FFD6839EA476CF779260" bold="true" box="[234,359,1537,1563]" pageId="27" pageNumber="28">
<typeStatus id="549DF7CF8C50FFD6839EA476CF779260" box="[234,359,1537,1563]" pageId="27" pageNumber="28" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B99496D8C50FFD683B5A452CC7892FC" blockId="27.[189,1399,1537,1672]" pageId="27" pageNumber="28">
MEDITERRANEAN SEA • 
<materialsCitation id="3B4E43308C50FFD68178A452CADE9244" ID-GBIF-Occurrence="2563048063" box="[524,1230,1573,1600]" collectingDate="2012-05" collectorName="A. Weber" location="Agios Pavlos" pageId="27" pageNumber="28" specimenCount="1" specimenCount-female="1" stateProvince="Kriti" typeStatus="paratype">
<specimenCount id="9D2082E48C50FFD68178A452CC259244" box="[524,565,1573,1599]" pageId="27" pageNumber="28" type="female">1 ♀</specimenCount>
(brooding); same data as for holotype; SMNH-Type-9189
</materialsCitation>
• 
<materialsCitation id="3B4E43308C50FFD68795A452CC7392FC" ID-GBIF-Occurrence="2563048062" collectingDate="2005-08-10" collectorName="R. Graille" country="Greece" county="Agios Georgios" location="Symi Island" municipality="Rhodes" pageId="27" pageNumber="28" specimenCount="7" typeStatus="paratype">
<specimenCount id="9D2082E48C50FFD68795A452CB629244" box="[1249,1394,1573,1599]" pageId="27" pageNumber="28" type="generic">7 specimens</specimenCount>
; 
<collectingCountry id="F33109FD8C50FFD683C9A43ECF039218" box="[189,275,1609,1635]" name="Greece" pageId="27" pageNumber="28">Greece</collectingCountry>
, 
<location id="8EF91FB68C50FFD68255A43ECFA49218" LSID="urn:lsid:plazi:treatment:038FF87B8C50FFD2811DA130CF0997A5:8EF91FB68C50FFD68255A43ECFA49218" box="[289,436,1609,1635]" country="Greece" county="Agios Georgios" municipality="Rhodes" name="Symi Island" pageId="27" pageNumber="28">Symi Island</location>
, near 
<collectingMunicipality id="6BFDD3178C50FFD6828AA43ECC499218" box="[510,601,1609,1635]" pageId="27" pageNumber="28">Rhodes</collectingMunicipality>
, 
<collectingCounty id="62F831E18C50FFD68111A43ECD349218" box="[613,804,1609,1635]" pageId="27" pageNumber="28">Agios Georgios</collectingCounty>
; 
<date id="FF986FAD8C50FFD68046A43ECDC9921F" box="[818,985,1609,1636]" pageId="27" pageNumber="28" value="2005-08-10">
<collectingDate id="EFDC96458C50FFD68046A43ECDC9921F" box="[818,985,1609,1636]" pageId="27" pageNumber="28" value="2005-08-10">10 Aug. 2005</collectingDate>
</date>
; 
<collectorName id="26D32CBB8C50FFD68093A43DCA4D9218" box="[999,1117,1609,1636]" pageId="27" pageNumber="28">R. Graille</collectorName>
leg.; preserved in 95% ethanol; SMNH-Type-9190 to 9196
</materialsCitation>
.
</paragraph>
<paragraph id="8B99496D8C50FFD6839EA4D8CFB692B2" blockId="27.[189,1399,1711,2027]" box="[234,422,1711,1737]" pageId="27" pageNumber="28">
<emphasis id="B952957F8C50FFD6839EA4D8CFB692B2" bold="true" box="[234,422,1711,1737]" pageId="27" pageNumber="28">Other material</emphasis>
</paragraph>
<paragraph id="8B99496D8C50FFD683C9A4A3CB7F9390" blockId="27.[189,1399,1711,2027]" pageId="27" pageNumber="28">
MEDITERRANEAN SEA • 
<materialsCitation id="3B4E43308C50FFD6817BA4A3CA15934D" ID-GBIF-Occurrence="2563048035" collectingDate="2008-07-02" collectionCode="SMNH" collectorName="Emilie Aegea" country="Greece" location="Plaka Beach" municipality="Elounda" pageId="27" pageNumber="28" specimenCode="SMNH-102712, 113690, 13697, 11370, 113704, 113709, 113712" specimenCount="18" stateProvince="Kriti">
<specimenCount id="9D2082E48C50FFD6817BA4A3CCBD9295" box="[527,685,1748,1774]" pageId="27" pageNumber="28" type="generic">18 specimens</specimenCount>
(mitochondrial COI lineage L3, nuclear cluster C5); 
<collectingCountry id="F33109FD8C50FFD68668A4A3CB629295" box="[1308,1394,1748,1774]" name="Greece" pageId="27" pageNumber="28">Greece</collectingCountry>
, northwestern 
<collectingRegion id="49E2878F8C50FFD6822BA48FCFB29369" box="[351,418,1784,1810]" country="Greece" name="Kriti" pageId="27" pageNumber="28">Crete</collectingRegion>
, 
<collectingMunicipality id="6BFDD3178C50FFD682DBA48FCC049369" box="[431,532,1784,1810]" pageId="27" pageNumber="28">Elounda</collectingMunicipality>
, 
<location id="8EF91FB68C50FFD68154A48FCCA79369" LSID="urn:lsid:plazi:treatment:038FF87B8C50FFD2811DA130CF0997A5:8EF91FB68C50FFD68154A48FCCA79369" box="[544,695,1784,1810]" country="Greece" municipality="Elounda" name="Plaka Beach" pageId="27" pageNumber="28" stateProvince="Kriti">Plaka Beach</location>
; 
<date id="FF986FAD8C50FFD681B1A48FCD5C9369" box="[709,844,1784,1810]" pageId="27" pageNumber="28" value="2008-07-02">
<collectingDate id="EFDC96458C50FFD681B1A48FCD5C9369" box="[709,844,1784,1810]" pageId="27" pageNumber="28" value="2008-07-02">2 Jul. 2008</collectingDate>
</date>
; 
<collectorName id="26D32CBB8C50FFD6802EA48FCDEA9369" box="[858,1018,1784,1810]" pageId="27" pageNumber="28">Emilie Aegea</collectorName>
leg.; preserved in 95% ethanol; 
<specimenCode id="DB80E1168C50FFD683C9A56BCF66934D" box="[189,374,1820,1846]" collectionCode="SMNH-" pageId="27" pageNumber="28">SMNH-102712</specimenCode>
, 
<specimenCode id="DB80E1168C50FFD682F7A56BCFCA934D" box="[387,474,1820,1846]" collectionCode="SMNH-" pageId="27" pageNumber="28">113690</specimenCode>
to 
<specimenCode id="DB80E1168C50FFD68175A56BCC5D934D" box="[513,589,1820,1846]" collectionCode="SMNH-" pageId="27" pageNumber="28">13697</specimenCode>
, 
<specimenCode id="DB80E1168C50FFD6812EA56BCCB2934D" box="[602,674,1820,1846]" collectionCode="SMNH-" pageId="27" pageNumber="28">11370</specimenCode>
to 
<specimenCode id="DB80E1168C50FFD681BEA56BCD34934D" box="[714,804,1820,1846]" collectionCode="SMNH-" pageId="27" pageNumber="28">113704</specimenCode>
, 
<specimenCode id="DB80E1168C50FFD68044A56BCD97934D" box="[816,903,1820,1846]" collectionCode="SMNH-" pageId="27" pageNumber="28">113709</specimenCode>
to 
<specimenCode id="DB80E1168C50FFD680DAA56BCA15934D" box="[942,1029,1820,1846]" collectionCode="SMNH-" pageId="27" pageNumber="28">113712</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B4E43308C50FFD68754A56BCF6D93D9" ID-GBIF-Occurrence="2563048061" collectingDate="2005-09-09" collectionCode="SMNH" collectorName="H. Zibrowius" country="Greece" location="Kallithea Bay" municipality="Rhodes" pageId="27" pageNumber="28" specimenCode="SMNH-99657, 113982, 113985, 113987, 113990, 113992, 114003, 114005, 114006" specimenCount="21">
<specimenCount id="9D2082E48C50FFD68754A56BCAD0934D" box="[1056,1216,1820,1846]" pageId="27" pageNumber="28" type="generic">21 specimens</specimenCount>
(mitochondrial COI lineage L3, nuclear cluster C5); 
<collectingCountry id="F33109FD8C50FFD6810FA537CCC19321" box="[635,721,1856,1882]" name="Greece" pageId="27" pageNumber="28">Greece</collectingCountry>
, 
<collectingMunicipality id="6BFDD3178C50FFD681ABA537CD2A9321" box="[735,826,1856,1882]" pageId="27" pageNumber="28">Rhodes</collectingMunicipality>
, 
<location id="8EF91FB68C50FFD68033A537CDFE9320" LSID="urn:lsid:plazi:treatment:038FF87B8C50FFD2811DA130CF0997A5:8EF91FB68C50FFD68033A537CDFE9320" box="[839,1006,1856,1883]" country="Greece" municipality="Rhodes" name="Kallithea Bay" pageId="27" pageNumber="28">Kallithea Bay</location>
; 
<date id="FF986FAD8C50FFD68088A537CA9E9320" box="[1020,1166,1856,1883]" pageId="27" pageNumber="28" value="2005-09-09">
<collectingDate id="EFDC96458C50FFD68088A537CA9E9320" box="[1020,1166,1856,1883]" pageId="27" pageNumber="28" value="2005-09-09">9 Sep. 2005</collectingDate>
</date>
; 
<collectorName id="26D32CBB8C50FFD687E8A536CB2B9321" box="[1180,1339,1856,1883]" pageId="27" pageNumber="28">H. Zibrowius</collectorName>
leg.; preserved in 95% ethanol; 
<specimenCode id="DB80E1168C50FFD68179A513CCA89304" box="[525,696,1892,1919]" collectionCode="SMNH-" pageId="27" pageNumber="28">SMNH-99657</specimenCode>
, 
<specimenCode id="DB80E1168C50FFD681BDA513CD339305" box="[713,803,1892,1918]" collectionCode="SMNH-" pageId="27" pageNumber="28">113982</specimenCode>
, 
<specimenCode id="DB80E1168C50FFD68040A513CD9E9304" box="[820,910,1892,1919]" collectionCode="SMNH-" pageId="27" pageNumber="28">113985</specimenCode>
, 
<specimenCode id="DB80E1168C50FFD680EBA513CDE69305" box="[927,1014,1892,1918]" collectionCode="SMNH-" pageId="27" pageNumber="28">113987</specimenCode>
to 
<specimenCode id="DB80E1168C50FFD68752A513CA909305" box="[1062,1152,1892,1918]" collectionCode="SMNH-" pageId="27" pageNumber="28">113990</specimenCode>
, 
<specimenCode id="DB80E1168C50FFD687E5A513CAF89305" box="[1169,1256,1892,1918]" collectionCode="SMNH-" pageId="27" pageNumber="28">113992</specimenCode>
to 
<specimenCode id="DB80E1168C50FFD6866CA513CB629305" box="[1304,1394,1892,1918]" collectionCode="SMNH-" pageId="27" pageNumber="28">114003</specimenCode>
, 
<specimenCode id="DB80E1168C50FFD683C9A5FFCF0793D8" box="[189,279,1928,1955]" collectionCode="SMNH-" pageId="27" pageNumber="28">114005</specimenCode>
, 
<specimenCode id="DB80E1168C50FFD68252A5FFCF6D93D9" box="[294,381,1928,1954]" collectionCode="SMNH-" pageId="27" pageNumber="28">114006</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B4E43308C50FFD682E8A5FECDEE93BC" ID-GBIF-Occurrence="2563048043" collectingDate="2012-05" collectionCode="SMNH" collectorName="A. Weber" country="Greece" location="Crete" pageId="27" pageNumber="28" specimenCode="SMNH-147070" specimenCount="41" stateProvince="Kriti">
<specimenCount id="9D2082E48C50FFD682E8A5FECC2E93D9" box="[412,574,1928,1955]" pageId="27" pageNumber="28" type="generic">41 specimens</specimenCount>
(mitochondrial COI lineage L3, nuclear cluster C5); 
<collectingCountry id="F33109FD8C50FFD687BEA5FECB3093D8" box="[1226,1312,1929,1955]" name="Greece" pageId="27" pageNumber="28">Greece</collectingCountry>
, 
<collectingRegion id="49E2878F8C50FFD6865BA5FECB6293D8" box="[1327,1394,1929,1955]" country="Greece" name="Kriti" pageId="27" pageNumber="28">Crete</collectingRegion>
; 
<date id="FF986FAD8C50FFD683C9A5DACF2B93BC" box="[189,315,1965,1991]" pageId="27" pageNumber="28" value="2012-05">
<collectingDate id="EFDC96458C50FFD683C9A5DACF2B93BC" box="[189,315,1965,1991]" pageId="27" pageNumber="28" value="2012-05">May 2012</collectingDate>
</date>
; 
<collectorName id="26D32CBB8C50FFD6823CA5DACFAA93BD" box="[328,442,1964,1991]" pageId="27" pageNumber="28">A. Weber</collectorName>
leg.; preserved in 95% ethanol; 
<specimenCode id="DB80E1168C50FFD68033A5DACDEE93BC" box="[839,1022,1965,1991]" collectionCode="SMNH-" pageId="27" pageNumber="28">SMNH-147070</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B4E43308C50FFD68769A5DACB7B9390" ID-GBIF-Occurrence="2563048041" collectingDate="2012-05" collectionCode="SMNH" collectorName="A. Weber" country="Greece" location="Crete" pageId="27" pageNumber="28" specimenCode="SMNH-178455" specimenCount="1" stateProvince="Kriti">
<specimenCount id="9D2082E48C50FFD68769A5DACAB493BD" box="[1053,1188,1964,1991]" pageId="27" pageNumber="28" type="generic">1 specimen</specimenCount>
(dd = 
<quantity id="4CDEE4888C50FFD6878EA5DBCB7A93BC" box="[1274,1386,1964,1991]" metricMagnitude="-2" metricUnit="m" metricValue="1.54" pageId="27" pageNumber="28" unit="mm" value="15.4">15.4 mm</quantity>
), dried dissociated ossicles and ossicles on two SEM stubs; same data as for preceding; 
<specimenCode id="DB80E1168C50FFD687C5A5A7CB7B9390" box="[1201,1387,2000,2027]" collectionCode="SMNH-" pageId="27" pageNumber="28">SMNH-178455</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33C1AE68C57FFD383C9A37CCCA995C9" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29" type="description">
<paragraph id="8B99496D8C57FFD183C9A37CCF5E955E" blockId="28.[189,334,267,293]" box="[189,334,267,293]" pageId="28" pageNumber="29">
<heading id="D0D1FE018C57FFD183C9A37CCF5E955E" bold="true" box="[189,334,267,293]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B952957F8C57FFD183C9A37CCF5E955E" bold="true" box="[189,334,267,293]" pageId="28" pageNumber="29">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B99496D8C57FFD1839EA34DCFD9952F" blockId="28.[189,1399,313,657]" box="[234,457,313,340]" pageId="28" pageNumber="29">
<emphasis id="B952957F8C57FFD1839EA34DCF4C952F" bold="true" box="[234,348,314,340]" pageId="28" pageNumber="29">
<typeStatus id="549DF7CF8C57FFD1839EA34DCF4C952F" box="[234,348,314,340]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="131D55E88C57FFD18218A34ECFD0952F" box="[364,448,313,340]" captionStart="Fig" captionStartId="28.[189,232,1895,1921]" captionTargetBox="[196,1391,717,1854]" captionTargetId="figure@28.[196,1391,717,1854]" captionTargetPageId="28" captionText="Fig. 11. Ophioderma zibrowii sp. nov., holotype (SMNH-Type-9188), digital images. A. Dorsal aspect, note the naked radial shields. B. Ventral aspect. C. Ventral arm. D. Dorsal arm, note the entire plates. E. Jaw. F. Brooded juveniles in interradius (arrow). Abbreviations: DAP = dorsal arm plate; M = madreporite; OS = oral shield; RS = radial shield; VAP = ventral arm plate." figureDoi="http://doi.org/10.5281/zenodo.3637795" httpUri="https://zenodo.org/record/3637795/files/figure.png" pageId="28" pageNumber="29">Fig. 11</figureCitation>
)
</paragraph>
<paragraph id="8B99496D8C57FFD083C9A32BCA3C95CE" blockId="28.[189,1399,313,657]" lastBlockId="29.[189,1399,267,437]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
Disc diameter 
<quantity id="4CDEE4888C57FFD1821AA32ACFCC950C" box="[366,476,349,375]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="28" pageNumber="29" unit="mm" value="14.3">14.3 mm</quantity>
, one intact arm, all others removed for molecular analysis. Dorsal disc with dense cover of granules, radial shields naked, widely separated, visible part oval (
<figureCitation id="131D55E88C57FFD187FDA3F7CAE795E1" box="[1161,1271,384,410]" captionStart="Fig" captionStartId="28.[189,232,1895,1921]" captionTargetBox="[196,1391,717,1854]" captionTargetId="figure@28.[196,1391,717,1854]" captionTargetPageId="28" captionText="Fig. 11. Ophioderma zibrowii sp. nov., holotype (SMNH-Type-9188), digital images. A. Dorsal aspect, note the naked radial shields. B. Ventral aspect. C. Ventral arm. D. Dorsal arm, note the entire plates. E. Jaw. F. Brooded juveniles in interradius (arrow). Abbreviations: DAP = dorsal arm plate; M = madreporite; OS = oral shield; RS = radial shield; VAP = ventral arm plate." figureDoi="http://doi.org/10.5281/zenodo.3637795" httpUri="https://zenodo.org/record/3637795/files/figure.png" pageId="28" pageNumber="29">Fig. 11A</figureCitation>
). Granules and scales crowd onto basal arms. Dorsal arm plates single on all joints, flat, overlapping, broadly fanshaped, twice as wide as long, with straight edges, wider distally than proximally (
<figureCitation id="131D55E88C57FFD1870FA3B1CAC7959B" box="[1147,1239,454,480]" captionStart="Fig" captionStartId="2.[189,232,1515,1541]" captionTargetBox="[194,1381,269,1466]" captionTargetId="figure@2.[193,1395,265,1467]" captionTargetPageId="2" captionText="Fig. 1. Phylogenetic relationships and distribution of species of Ophioderma Müller &amp; Troschel, 1840 in the Northeast and Tropical Atlantic Ocean and the Mediterranean Sea. A. Simplified phylogenetic relationships among species of Ophioderma described in this study, modified from Weber et al. (2019). Summary of reproductive strategy: S = broadcast spawner; B = brooder. Correspondence between species names, nuclear genetic clusters and COI mitochondrial lineages is summarized from Boissin et al. (2011) and Weber et al. (2019). C6 is tentatively assigned to O. zibrowii sp. nov. because it occurs far from the type locality and brooding species are expected to have low dispersal rates, leading to low gene flow between populations. B. Distribution of species of Ophioderma in the Northeast and Tropical Atlantic Ocean. C. Distribution of species of Ophioderma in the Mediterranean Sea." figureDoi="http://doi.org/10.5281/zenodo.3637775" httpUri="https://zenodo.org/record/3637775/files/figure.png" pageId="28" pageNumber="29">Fig. 1D</figureCitation>
). Up to seven conical, slightly pointed arm spines, about equal in length, half as long as arm joint, ventralmost spine thicker than others and blunt (
<figureCitation id="131D55E88C57FFD18155A07BCC6F965C" box="[545,639,524,551]" captionStart="Fig" captionStartId="19.[189,232,1930,1956]" captionTargetBox="[189,1398,797,1888]" captionTargetId="figure@19.[189,1398,797,1888]" captionTargetPageId="19" captionText="Fig. 7. Ophioderma africana sp. nov., digital images. A–G. Holotype (SMNH-Type-7484). A. Dorsal aspect. B. Dorsal disc. C. Ventral aspect. D–E. Two dorsal arms, note multiple dorsal arm plates on both. F. Ventral arm. G. Mouth. H. Some paratypes (SMNH-Type-7487)." figureDoi="http://doi.org/10.5281/zenodo.3637787" httpUri="https://zenodo.org/record/3637787/files/figure.png" pageId="28" pageNumber="29">Fig. 7C</figureCitation>
). Ventral disc covered by granules similar to dorsal disc. Adoral shields and oral plates covered by granules, oral shields naked (
<figureCitation id="131D55E88C57FFD180D8A047CA079631" box="[940,1047,560,586]" captionStart="Fig" captionStartId="28.[189,232,1895,1921]" captionTargetBox="[196,1391,717,1854]" captionTargetId="figure@28.[196,1391,717,1854]" captionTargetPageId="28" captionText="Fig. 11. Ophioderma zibrowii sp. nov., holotype (SMNH-Type-9188), digital images. A. Dorsal aspect, note the naked radial shields. B. Ventral aspect. C. Ventral arm. D. Dorsal arm, note the entire plates. E. Jaw. F. Brooded juveniles in interradius (arrow). Abbreviations: DAP = dorsal arm plate; M = madreporite; OS = oral shield; RS = radial shield; VAP = ventral arm plate." figureDoi="http://doi.org/10.5281/zenodo.3637795" httpUri="https://zenodo.org/record/3637795/files/figure.png" pageId="28" pageNumber="29">Fig. 11B</figureCitation>
). Seven to eight oral papillae, Lyman’s ossicle elongated, pointing into mouth slit, adoral shield spines two, block shaped, other papillae smaller, conical (
<figureCitation id="131D55E88C57FFD182FEA001CFE596EB" box="[394,501,630,656]" captionStart="Fig" captionStartId="28.[189,232,1895,1921]" captionTargetBox="[196,1391,717,1854]" captionTargetId="figure@28.[196,1391,717,1854]" captionTargetPageId="28" captionText="Fig. 11. Ophioderma zibrowii sp. nov., holotype (SMNH-Type-9188), digital images. A. Dorsal aspect, note the naked radial shields. B. Ventral aspect. C. Ventral arm. D. Dorsal arm, note the entire plates. E. Jaw. F. Brooded juveniles in interradius (arrow). Abbreviations: DAP = dorsal arm plate; M = madreporite; OS = oral shield; RS = radial shield; VAP = ventral arm plate." figureDoi="http://doi.org/10.5281/zenodo.3637795" httpUri="https://zenodo.org/record/3637795/files/figure.png" pageId="28" pageNumber="29">Fig. 11E</figureCitation>
). Tooth papillae larger, conical, two in line with oral papillae. Oral shields triangular, twice as wide as long, straight distal edge, obtuse rounded proximal angle. Madreporite as wide as long. Adoral shield long, narrow, extending around the lateral edge of the oral shield, separating it from the arm. Adoral shields covered with granules, similar to disc granules. Proximal genital slits parallel to and as long as first and second joints. Distal genital slit parallel to and as long as fourth joint, at distance from disc edge. Bursae filled with brooded juveniles (
<figureCitation id="131D55E88C56FFD080C3A3EDCA0F95CE" box="[951,1055,410,437]" captionStart="Fig" captionStartId="28.[189,232,1895,1921]" captionTargetBox="[196,1391,717,1854]" captionTargetId="figure@28.[196,1391,717,1854]" captionTargetPageId="28" captionText="Fig. 11. Ophioderma zibrowii sp. nov., holotype (SMNH-Type-9188), digital images. A. Dorsal aspect, note the naked radial shields. B. Ventral aspect. C. Ventral arm. D. Dorsal arm, note the entire plates. E. Jaw. F. Brooded juveniles in interradius (arrow). Abbreviations: DAP = dorsal arm plate; M = madreporite; OS = oral shield; RS = radial shield; VAP = ventral arm plate." figureDoi="http://doi.org/10.5281/zenodo.3637795" httpUri="https://zenodo.org/record/3637795/files/figure.png" pageId="29" pageNumber="30">Fig. 11F</figureCitation>
).
</paragraph>
<caption id="DF5919E58C57FFD183C9A510CA3F9390" ID-DOI="http://doi.org/10.5281/zenodo.3637795" ID-Zenodo-Dep="3637795" httpUri="https://zenodo.org/record/3637795/files/figure.png" pageId="28" pageNumber="29" startId="28.[189,232,1895,1921]" targetBox="[196,1391,717,1854]" targetPageId="28">
<paragraph id="8B99496D8C57FFD183C9A510CA3F9390" blockId="28.[189,1399,1895,2027]" pageId="28" pageNumber="29">
<emphasis id="B952957F8C57FFD183C9A510CF0893FA" bold="true" box="[189,280,1895,1921]" pageId="28" pageNumber="29">Fig. 11.</emphasis>
<taxonomicName id="4C2632EE8C57FFD1826BA510CC0A93FA" authority="Stöhr &amp; Weber &amp; Boissin &amp; Chenuil, 2020" authorityName="Stöhr &amp; Weber &amp; Boissin &amp; Chenuil" authorityYear="2020" box="[287,538,1895,1921]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="28" pageNumber="29" phylum="Echinodermata" rank="species" species="zibrowii" status="sp. nov.">
<emphasis id="B952957F8C57FFD1826BA510CC0A93FA" box="[287,538,1895,1921]" italics="true" pageId="28" pageNumber="29">Ophioderma zibrowii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26128048C57FFD18155A51FCC6E93F9" box="[545,638,1896,1922]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (SMNH-Type-9188), digital images. 
<emphasis id="B952957F8C57FFD187D8A510CAD193FA" bold="true" box="[1196,1217,1895,1921]" pageId="28" pageNumber="29">A</emphasis>
. Dorsal aspect, note the naked radial shields. 
<emphasis id="B952957F8C57FFD18156A5FDCC2693DF" bold="true" box="[546,566,1930,1956]" pageId="28" pageNumber="29">B</emphasis>
. Ventral aspect. 
<emphasis id="B952957F8C57FFD18188A5FDCD0193DF" bold="true" box="[764,785,1930,1956]" pageId="28" pageNumber="29">C</emphasis>
. Ventral arm. 
<emphasis id="B952957F8C57FFD180C8A5FDCDC193DF" bold="true" box="[956,977,1930,1956]" pageId="28" pageNumber="29">D</emphasis>
. Dorsal arm, note the entire plates. 
<emphasis id="B952957F8C57FFD183C9A5D9CEC193B3" bold="true" box="[189,209,1966,1992]" pageId="28" pageNumber="29">E</emphasis>
. Jaw. 
<emphasis id="B952957F8C57FFD18257A5D9CF2593B3" bold="true" box="[291,309,1966,1992]" pageId="28" pageNumber="29">F</emphasis>
. Brooded juveniles in interradius (arrow). Abbreviations: DAP = dorsal arm plate; M = madreporite; OS = oral shield; RS = radial shield; VAP = ventral arm plate.
</paragraph>
</caption>
<paragraph id="8B99496D8C56FFD083C9A3AACAC296FD" blockId="29.[189,1399,477,646]" pageId="29" pageNumber="30">
Dorsal disc slightly variegated dark brown-green with grey, dorsal arms greyish green with brown bands, each light and dark band about three joints wide, a series of larger white and smaller black spots along the distal edge of each dorsal arm plate (
<figureCitation id="131D55E88C56FFD081A7A053CD709644" box="[723,864,548,575]" captionStart="Fig" captionStartId="28.[189,232,1895,1921]" captionTargetBox="[196,1391,717,1854]" captionTargetId="figure@28.[196,1391,717,1854]" captionTargetPageId="28" captionText="Fig. 11. Ophioderma zibrowii sp. nov., holotype (SMNH-Type-9188), digital images. A. Dorsal aspect, note the naked radial shields. B. Ventral aspect. C. Ventral arm. D. Dorsal arm, note the entire plates. E. Jaw. F. Brooded juveniles in interradius (arrow). Abbreviations: DAP = dorsal arm plate; M = madreporite; OS = oral shield; RS = radial shield; VAP = ventral arm plate." figureDoi="http://doi.org/10.5281/zenodo.3637795" httpUri="https://zenodo.org/record/3637795/files/figure.png" pageId="29" pageNumber="30">Fig. 11A, D</figureCitation>
). Ventral disc as dark as dorsal, oral frame and ventral arms lighter grey-green (
<figureCitation id="131D55E88C56FFD0814AA03FCCBA9619" box="[574,682,584,610]" captionStart="Fig" captionStartId="28.[189,232,1895,1921]" captionTargetBox="[196,1391,717,1854]" captionTargetId="figure@28.[196,1391,717,1854]" captionTargetPageId="28" captionText="Fig. 11. Ophioderma zibrowii sp. nov., holotype (SMNH-Type-9188), digital images. A. Dorsal aspect, note the naked radial shields. B. Ventral aspect. C. Ventral arm. D. Dorsal arm, note the entire plates. E. Jaw. F. Brooded juveniles in interradius (arrow). Abbreviations: DAP = dorsal arm plate; M = madreporite; OS = oral shield; RS = radial shield; VAP = ventral arm plate." figureDoi="http://doi.org/10.5281/zenodo.3637795" httpUri="https://zenodo.org/record/3637795/files/figure.png" pageId="29" pageNumber="30">Fig. 11B</figureCitation>
). Arm spines grey-green with white tips, except ventralmost spine, which is white (
<figureCitation id="131D55E88C56FFD082B1A01BCC0796FD" box="[453,535,620,646]" captionStart="Fig" captionStartId="28.[189,232,1895,1921]" captionTargetBox="[196,1391,717,1854]" captionTargetId="figure@28.[196,1391,717,1854]" captionTargetPageId="28" captionText="Fig. 11. Ophioderma zibrowii sp. nov., holotype (SMNH-Type-9188), digital images. A. Dorsal aspect, note the naked radial shields. B. Ventral aspect. C. Ventral arm. D. Dorsal arm, note the entire plates. E. Jaw. F. Brooded juveniles in interradius (arrow). Abbreviations: DAP = dorsal arm plate; M = madreporite; OS = oral shield; RS = radial shield; VAP = ventral arm plate." figureDoi="http://doi.org/10.5281/zenodo.3637795" httpUri="https://zenodo.org/record/3637795/files/figure.png" pageId="29" pageNumber="30">Fig. 11</figureCitation>
C–D). Oral papillae and tentacle scales white (
<figureCitation id="131D55E88C56FFD0874EA01BCAD596FD" box="[1082,1221,620,646]" captionStart="Fig" captionStartId="28.[189,232,1895,1921]" captionTargetBox="[196,1391,717,1854]" captionTargetId="figure@28.[196,1391,717,1854]" captionTargetPageId="28" captionText="Fig. 11. Ophioderma zibrowii sp. nov., holotype (SMNH-Type-9188), digital images. A. Dorsal aspect, note the naked radial shields. B. Ventral aspect. C. Ventral arm. D. Dorsal arm, note the entire plates. E. Jaw. F. Brooded juveniles in interradius (arrow). Abbreviations: DAP = dorsal arm plate; M = madreporite; OS = oral shield; RS = radial shield; VAP = ventral arm plate." figureDoi="http://doi.org/10.5281/zenodo.3637795" httpUri="https://zenodo.org/record/3637795/files/figure.png" pageId="29" pageNumber="30">Fig. 11C, E</figureCitation>
).
</paragraph>
<paragraph id="8B99496D8C56FFD0839EA0D9CF7796B3" blockId="29.[189,1399,686,963]" box="[234,359,686,712]" pageId="29" pageNumber="30">
<emphasis id="B952957F8C56FFD0839EA0D9CF7796B3" bold="true" box="[234,359,686,712]" pageId="29" pageNumber="30">
<typeStatus id="549DF7CF8C56FFD0839EA0D9CF7796B3" box="[234,359,686,712]" pageId="29" pageNumber="30" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B99496D8C56FFD083C9A0A5CC2B97B8" blockId="29.[189,1399,686,963]" pageId="29" pageNumber="30">
Disc diameter 
<quantity id="4CDEE4888C56FFD08211A0A5CC129697" box="[357,514,722,749]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" metricValueMax="1.35" metricValueMin="0.97" pageId="29" pageNumber="30" unit="mm" value="11.6" valueMax="13.5" valueMin="9.7">9.7–13.5 mm</quantity>
.All have up to seven arm spines. Most 
<typeStatus id="549DF7CF8C56FFD080B3A0A5CA279697" box="[967,1079,722,748]" pageId="29" pageNumber="30" type="paratype">paratypes</typeStatus>
have a green colour pattern similar to the 
<typeStatus id="549DF7CF8C56FFD08229A082CFD79774" box="[349,455,757,783]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
, one specimen is olive-brown. One 
<typeStatus id="549DF7CF8C56FFD0801CA081CDDD976B" box="[872,973,758,784]" pageId="29" pageNumber="30" type="paratype">paratype</typeStatus>
with granule-covered radial shields, all others naked. Oral papillae eight, in two specimens nine, but variable between jaws in an individual. Two specimens with two dorsal arm plates on 2–3 proximal joints on 1–3 arms. One specimen with 2–3 dorsal arm plates on proximal joints on three arms, on seven joints on one of those arms. One specimen has a parasitic snail (
<taxonomicName id="4C2632EE8C56FFD082DAA1F2CCF197E4" authority="Monterosato, 1872" authorityName="Monterosato" authorityYear="1872" box="[430,737,901,927]" class="Gastropoda" family="Eulimidae" genus="Ersilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="genus">
<emphasis id="B952957F8C56FFD082DAA1F2CFEE97E4" box="[430,510,901,927]" italics="true" pageId="29" pageNumber="30">Ersilia</emphasis>
Monterosato, 1872
</taxonomicName>
sp.) on one arm. All 
<typeStatus id="549DF7CF8C56FFD080A6A1F2CA5297E4" box="[978,1090,901,927]" pageId="29" pageNumber="30" type="paratype">paratypes</typeStatus>
lack part of an arm, which was used for molecular analysis.
</paragraph>
<paragraph id="8B99496D8C56FFD0839EA19CCC66907E" blockId="29.[189,1399,1003,1459]" box="[234,630,1003,1029]" pageId="29" pageNumber="30">
<emphasis id="B952957F8C56FFD0839EA19CCFB4907E" bold="true" box="[234,420,1003,1029]" pageId="29" pageNumber="30">Non-paratypes</emphasis>
(SMNH-147070)
</paragraph>
<paragraph id="8B99496D8C56FFD083C9A679CB7491C8" blockId="29.[189,1399,1003,1459]" pageId="29" pageNumber="30">
Disc diameter 
<quantity id="4CDEE4888C56FFD08210A679CC119052" box="[356,513,1038,1065]" metricMagnitude="-2" metricUnit="m" metricValue="1.4449999999999998" metricValueMax="2.05" metricValueMin="0.84" pageId="29" pageNumber="30" unit="mm" value="14.45" valueMax="20.5" valueMin="8.4">8.4–20.5 mm</quantity>
. Specimens smaller than 
<quantity id="4CDEE4888C56FFD08056A678CD639052" box="[802,883,1039,1065]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="29" pageNumber="30" unit="mm" value="10.0">10 mm</quantity>
dd had six arm spines, all others seven. Most specimens have a green colour pattern similar to the 
<typeStatus id="549DF7CF8C56FFD08046A645CD8C9037" box="[818,924,1074,1100]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
, several were dark olive with numerous white granules scattered across the disc. Out of the 
<specimenCount id="9D2082E48C56FFD08047A621CDC8900B" box="[819,984,1110,1136]" pageId="29" pageNumber="30" type="generic">42 specimens</specimenCount>
, a single one had granule-covered radial shields, all others with naked radial shields. Most of the specimens above 
<quantity id="4CDEE4888C56FFD08706A60DCAD590EE" box="[1138,1221,1146,1173]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="29" pageNumber="30" unit="mm" value="10.0">10 mm</quantity>
dd had at least two proximal arm joints on at least one arm with two dorsal arm plates, up to seven joints and several arms with double plates in one animal. The plates were usually of equal size or there was a hairline crack along the midline of the plate. Rarely, a third dorsal arm plate was present on one or two joints. Oral papillae eight, rarely nine, but variable between jaws in an individual and not correlated to size. These specimens were used for thermotolerance and regeneration experiments. These specimens were kept in an aquarium for a year where experiments of thermotolerance and regeneration were conducted. Therefore, most arms show signs of regeneration. One specimen had bright white distal arms but no sign of healed breaks. Maximum size recorded for 
<taxonomicName id="4C2632EE8C56FFD081AEA7EECD4D91C9" authority="Stöhr &amp; Weber &amp; Boissin &amp; Chenuil, 2020" authorityName="Stöhr &amp; Weber &amp; Boissin &amp; Chenuil" authorityYear="2020" box="[730,861,1432,1459]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="29" pageNumber="30" phylum="Echinodermata" rank="species" species="zibrowii" status="sp. nov.">
<emphasis id="B952957F8C56FFD081AEA7EECCFF91C8" box="[730,751,1433,1459]" italics="true" pageId="29" pageNumber="30">O</emphasis>
. 
<emphasis id="B952957F8C56FFD0818AA7EFCD4D91C9" box="[766,861,1432,1458]" italics="true" pageId="29" pageNumber="30">zibrowii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26128048C56FFD08010A7EECDAF91C8" box="[868,959,1433,1459]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
was 
<quantity id="4CDEE4888C56FFD0808FA7EECA5D91C8" box="[1019,1101,1433,1459]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="29" pageNumber="30" unit="mm" value="21.0">21 mm</quantity>
dd (
<bibRefCitation id="EFB7349C8C56FFD087F4A7EFCB4891C8" author="Weber" box="[1152,1368,1432,1459]" etAl="et al." firstAuthor="Weber" journalOrPublisher="Comptes Rendus Biologies" pageId="29" pageNumber="30" pagination="553 - 560" part="337" refId="ref19581" refString="Weber A. A. - T., Stohr S. &amp; Chenuil A. 2014. Genetic data, reproduction season and reproductive strategy data support the existence of biological species in Ophioderma longicauda. Comptes Rendus Biologies 337: 553 - 560. https: // doi. org / 10.1016 / j. crvi. 2014.07.007" title="Genetic data, reproduction season and reproductive strategy data support the existence of biological species in Ophioderma longicauda" type="journal article" year="2014">
Weber 
<emphasis id="B952957F8C56FFD087A2A7EECB0191C9" box="[1238,1297,1432,1458]" italics="true" pageId="29" pageNumber="30">et al.</emphasis>
2014
</bibRefCitation>
).
</paragraph>
<paragraph id="8B99496D8C56FFD0839EA7ADCC20918E" blockId="29.[189,1399,1498,2027]" box="[234,560,1498,1525]" pageId="29" pageNumber="30">
<emphasis id="B952957F8C56FFD0839EA7ADCFD2918E" bold="true" box="[234,450,1498,1525]" pageId="29" pageNumber="30">Skeletal elements</emphasis>
(
<figureCitation id="131D55E88C56FFD082A6A7ADCC37918E" box="[466,551,1498,1525]" captionStart="Fig" captionStartId="30.[189,232,1718,1744]" captionTargetBox="[189,1398,630,1678]" captionTargetId="figure@30.[189,1398,630,1678]" captionTargetPageId="30" captionText="Fig. 12. SEM images of skeletal elements of Ophioderma zibrowii sp. nov., from non-type specimen of 15.4 mm disc diameter (SMNH-178455), Crete, Greece. A. Radial shield, external aspect, white arrows point to larger peripheral pores. B. Radial shield, internal aspect. C. Scale-free part magnified from Fig. 12A, note the absence of pores on the denser area (naked in intact animal, see Fig. 11A). D. Dental plate fragments. E. Adradial genital plate, lateral aspect. F. Distal end of Fig. 12E. G. Abradial genital plate. H. Oral plate, abradial face. I. Oral plate, adradial face, black arrows point to oral papillae sockets and pores. J. Arm vertebra, proximal face. K. Arm vertebra, distal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; Sp = spurs; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.3637797" httpUri="https://zenodo.org/record/3637797/files/figure.png" pageId="29" pageNumber="30">Fig. 12</figureCitation>
)
</paragraph>
<paragraph id="8B99496D8C56FFD383C9A788CCA995C9" blockId="29.[189,1399,1498,2027]" lastBlockId="30.[189,1398,267,435]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
Non-type specimen of 
<quantity id="4CDEE4888C56FFD082BFA789CC249262" box="[459,564,1534,1561]" metricMagnitude="-2" metricUnit="m" metricValue="1.54" pageId="29" pageNumber="30" unit="mm" value="15.4">15.4 mm</quantity>
dd (SMNH-178455), completely dissociated. Radial shields elongated isosceles triangular, external surface with embossed oval of more finely meshed stereom in center of domed and thickened proximal half (only visible part in intact skeleton), row of eight pores curves around proximal dome (
<figureCitation id="131D55E88C56FFD082A7A41DCC4E92FF" box="[467,606,1642,1668]" captionStart="Fig" captionStartId="30.[189,232,1718,1744]" captionTargetBox="[189,1398,630,1678]" captionTargetId="figure@30.[189,1398,630,1678]" captionTargetPageId="30" captionText="Fig. 12. SEM images of skeletal elements of Ophioderma zibrowii sp. nov., from non-type specimen of 15.4 mm disc diameter (SMNH-178455), Crete, Greece. A. Radial shield, external aspect, white arrows point to larger peripheral pores. B. Radial shield, internal aspect. C. Scale-free part magnified from Fig. 12A, note the absence of pores on the denser area (naked in intact animal, see Fig. 11A). D. Dental plate fragments. E. Adradial genital plate, lateral aspect. F. Distal end of Fig. 12E. G. Abradial genital plate. H. Oral plate, abradial face. I. Oral plate, adradial face, black arrows point to oral papillae sockets and pores. J. Arm vertebra, proximal face. K. Arm vertebra, distal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; Sp = spurs; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.3637797" httpUri="https://zenodo.org/record/3637797/files/figure.png" pageId="29" pageNumber="30">Fig. 12A, C</figureCitation>
). On inner side of radial shield, a large pore on distal half and a round patch with denser stereom, two low distal bulbs separated by a furrow (
<figureCitation id="131D55E88C56FFD08777A4F9CA7F92D3" box="[1027,1135,1678,1704]" captionStart="Fig" captionStartId="30.[189,232,1718,1744]" captionTargetBox="[189,1398,630,1678]" captionTargetId="figure@30.[189,1398,630,1678]" captionTargetPageId="30" captionText="Fig. 12. SEM images of skeletal elements of Ophioderma zibrowii sp. nov., from non-type specimen of 15.4 mm disc diameter (SMNH-178455), Crete, Greece. A. Radial shield, external aspect, white arrows point to larger peripheral pores. B. Radial shield, internal aspect. C. Scale-free part magnified from Fig. 12A, note the absence of pores on the denser area (naked in intact animal, see Fig. 11A). D. Dental plate fragments. E. Adradial genital plate, lateral aspect. F. Distal end of Fig. 12E. G. Abradial genital plate. H. Oral plate, abradial face. I. Oral plate, adradial face, black arrows point to oral papillae sockets and pores. J. Arm vertebra, proximal face. K. Arm vertebra, distal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; Sp = spurs; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.3637797" httpUri="https://zenodo.org/record/3637797/files/figure.png" pageId="29" pageNumber="30">Fig. 12B</figureCitation>
). Dental plate consists of several pieces, sockets not penetrating the plate, bordered by low rim, tooth papillae sockets smaller, round, tooth sockets wide, oval (
<figureCitation id="131D55E88C56FFD0814EA4A2CCB79294" box="[570,679,1749,1775]" captionStart="Fig" captionStartId="30.[189,232,1718,1744]" captionTargetBox="[189,1398,630,1678]" captionTargetId="figure@30.[189,1398,630,1678]" captionTargetPageId="30" captionText="Fig. 12. SEM images of skeletal elements of Ophioderma zibrowii sp. nov., from non-type specimen of 15.4 mm disc diameter (SMNH-178455), Crete, Greece. A. Radial shield, external aspect, white arrows point to larger peripheral pores. B. Radial shield, internal aspect. C. Scale-free part magnified from Fig. 12A, note the absence of pores on the denser area (naked in intact animal, see Fig. 11A). D. Dental plate fragments. E. Adradial genital plate, lateral aspect. F. Distal end of Fig. 12E. G. Abradial genital plate. H. Oral plate, abradial face. I. Oral plate, adradial face, black arrows point to oral papillae sockets and pores. J. Arm vertebra, proximal face. K. Arm vertebra, distal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; Sp = spurs; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.3637797" httpUri="https://zenodo.org/record/3637797/files/figure.png" pageId="29" pageNumber="30">Fig. 12D</figureCitation>
). Adradial genital plate blade-like flat with bulbous distal end and large pore, distal pit with knob on one side, three ridges on other side (
<figureCitation id="131D55E88C56FFD0876BA48ECA629368" box="[1055,1138,1785,1811]" captionStart="Fig" captionStartId="30.[189,232,1718,1744]" captionTargetBox="[189,1398,630,1678]" captionTargetId="figure@30.[189,1398,630,1678]" captionTargetPageId="30" captionText="Fig. 12. SEM images of skeletal elements of Ophioderma zibrowii sp. nov., from non-type specimen of 15.4 mm disc diameter (SMNH-178455), Crete, Greece. A. Radial shield, external aspect, white arrows point to larger peripheral pores. B. Radial shield, internal aspect. C. Scale-free part magnified from Fig. 12A, note the absence of pores on the denser area (naked in intact animal, see Fig. 11A). D. Dental plate fragments. E. Adradial genital plate, lateral aspect. F. Distal end of Fig. 12E. G. Abradial genital plate. H. Oral plate, abradial face. I. Oral plate, adradial face, black arrows point to oral papillae sockets and pores. J. Arm vertebra, proximal face. K. Arm vertebra, distal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; Sp = spurs; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.3637797" httpUri="https://zenodo.org/record/3637797/files/figure.png" pageId="29" pageNumber="30">Fig. 12</figureCitation>
E–F). Abradial genital plate much smaller and scale-like flat, distally wider than proximally (
<figureCitation id="131D55E88C56FFD08098A56ACA49934C" box="[1004,1113,1821,1847]" captionStart="Fig" captionStartId="30.[189,232,1718,1744]" captionTargetBox="[189,1398,630,1678]" captionTargetId="figure@30.[189,1398,630,1678]" captionTargetPageId="30" captionText="Fig. 12. SEM images of skeletal elements of Ophioderma zibrowii sp. nov., from non-type specimen of 15.4 mm disc diameter (SMNH-178455), Crete, Greece. A. Radial shield, external aspect, white arrows point to larger peripheral pores. B. Radial shield, internal aspect. C. Scale-free part magnified from Fig. 12A, note the absence of pores on the denser area (naked in intact animal, see Fig. 11A). D. Dental plate fragments. E. Adradial genital plate, lateral aspect. F. Distal end of Fig. 12E. G. Abradial genital plate. H. Oral plate, abradial face. I. Oral plate, adradial face, black arrows point to oral papillae sockets and pores. J. Arm vertebra, proximal face. K. Arm vertebra, distal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; Sp = spurs; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.3637797" httpUri="https://zenodo.org/record/3637797/files/figure.png" pageId="29" pageNumber="30">Fig. 12G</figureCitation>
). Oral plates longer than high, middle part markedly lower than ends, abradial muscle fossa rounded triangular, not reaching upper edge (
<figureCitation id="131D55E88C56FFD08239A513CFAA9304" box="[333,442,1892,1919]" captionStart="Fig" captionStartId="30.[189,232,1718,1744]" captionTargetBox="[189,1398,630,1678]" captionTargetId="figure@30.[189,1398,630,1678]" captionTargetPageId="30" captionText="Fig. 12. SEM images of skeletal elements of Ophioderma zibrowii sp. nov., from non-type specimen of 15.4 mm disc diameter (SMNH-178455), Crete, Greece. A. Radial shield, external aspect, white arrows point to larger peripheral pores. B. Radial shield, internal aspect. C. Scale-free part magnified from Fig. 12A, note the absence of pores on the denser area (naked in intact animal, see Fig. 11A). D. Dental plate fragments. E. Adradial genital plate, lateral aspect. F. Distal end of Fig. 12E. G. Abradial genital plate. H. Oral plate, abradial face. I. Oral plate, adradial face, black arrows point to oral papillae sockets and pores. J. Arm vertebra, proximal face. K. Arm vertebra, distal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; Sp = spurs; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.3637797" httpUri="https://zenodo.org/record/3637797/files/figure.png" pageId="29" pageNumber="30">Fig. 12H</figureCitation>
). On adradial oral plate proximoventral part four oral papilla sockets and pores, at proximoventral edge four granule sockets and pores (
<figureCitation id="131D55E88C56FFD0805FA5FFCD9C93D9" box="[811,908,1928,1954]" captionStart="Fig" captionStartId="30.[189,232,1718,1744]" captionTargetBox="[189,1398,630,1678]" captionTargetId="figure@30.[189,1398,630,1678]" captionTargetPageId="30" captionText="Fig. 12. SEM images of skeletal elements of Ophioderma zibrowii sp. nov., from non-type specimen of 15.4 mm disc diameter (SMNH-178455), Crete, Greece. A. Radial shield, external aspect, white arrows point to larger peripheral pores. B. Radial shield, internal aspect. C. Scale-free part magnified from Fig. 12A, note the absence of pores on the denser area (naked in intact animal, see Fig. 11A). D. Dental plate fragments. E. Adradial genital plate, lateral aspect. F. Distal end of Fig. 12E. G. Abradial genital plate. H. Oral plate, abradial face. I. Oral plate, adradial face, black arrows point to oral papillae sockets and pores. J. Arm vertebra, proximal face. K. Arm vertebra, distal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; Sp = spurs; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.3637797" httpUri="https://zenodo.org/record/3637797/files/figure.png" pageId="29" pageNumber="30">Fig. 12I</figureCitation>
). Vertebrae typically zygospondylus, with large dorsal and small ventral muscle flanges, no growth rings obvious (
<figureCitation id="131D55E88C56FFD08773A5DBCA4B93BD" box="[1031,1115,1964,1990]" captionStart="Fig" captionStartId="30.[189,232,1718,1744]" captionTargetBox="[189,1398,630,1678]" captionTargetId="figure@30.[189,1398,630,1678]" captionTargetPageId="30" captionText="Fig. 12. SEM images of skeletal elements of Ophioderma zibrowii sp. nov., from non-type specimen of 15.4 mm disc diameter (SMNH-178455), Crete, Greece. A. Radial shield, external aspect, white arrows point to larger peripheral pores. B. Radial shield, internal aspect. C. Scale-free part magnified from Fig. 12A, note the absence of pores on the denser area (naked in intact animal, see Fig. 11A). D. Dental plate fragments. E. Adradial genital plate, lateral aspect. F. Distal end of Fig. 12E. G. Abradial genital plate. H. Oral plate, abradial face. I. Oral plate, adradial face, black arrows point to oral papillae sockets and pores. J. Arm vertebra, proximal face. K. Arm vertebra, distal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; Sp = spurs; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.3637797" httpUri="https://zenodo.org/record/3637797/files/figure.png" pageId="29" pageNumber="30">Fig. 12</figureCitation>
J–K). Lateral arm plates compact, weakly curved around the arm, strongly convex distally, concave proximally, with ventral excavation for tube foot opening (
<figureCitation id="131D55E88C55FFD38125A37CCCB5955E" box="[593,677,267,293]" captionStart="Fig" captionStartId="30.[189,232,1718,1744]" captionTargetBox="[189,1398,630,1678]" captionTargetId="figure@30.[189,1398,630,1678]" captionTargetPageId="30" captionText="Fig. 12. SEM images of skeletal elements of Ophioderma zibrowii sp. nov., from non-type specimen of 15.4 mm disc diameter (SMNH-178455), Crete, Greece. A. Radial shield, external aspect, white arrows point to larger peripheral pores. B. Radial shield, internal aspect. C. Scale-free part magnified from Fig. 12A, note the absence of pores on the denser area (naked in intact animal, see Fig. 11A). D. Dental plate fragments. E. Adradial genital plate, lateral aspect. F. Distal end of Fig. 12E. G. Abradial genital plate. H. Oral plate, abradial face. I. Oral plate, adradial face, black arrows point to oral papillae sockets and pores. J. Arm vertebra, proximal face. K. Arm vertebra, distal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; Sp = spurs; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.3637797" httpUri="https://zenodo.org/record/3637797/files/figure.png" pageId="30" pageNumber="31">Fig. 12</figureCitation>
L–M). Two spurs at proximal external edge and counterparts on internal distal edge of lateral arm plates. Internal lateral arm plate with vertical row of three pores just below the plate centre, distal to curved vertical ridge, fourth pore in tentacle opening edge, small ventral knob, protruding beyond plate edge. Spine articulations inset in the distal plate edge with one thickened lobe, separating muscle and nerve opening.
</paragraph>
</subSubSection>
<subSubSection id="C33C1AE68C55FFD283C9A3ADCF0997A5" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="discussion">
<paragraph id="8B99496D8C55FFD383C9A3ADCF3F958F" blockId="30.[189,303,474,500]" box="[189,303,474,500]" pageId="30" pageNumber="31">
<heading id="D0D1FE018C55FFD383C9A3ADCF3F958F" bold="true" box="[189,303,474,500]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B952957F8C55FFD383C9A3ADCF3F958F" bold="true" box="[189,303,474,500]" pageId="30" pageNumber="31">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B99496D8C55FFD283C9A07FCD9295F4" blockId="30.[189,1398,520,582]" lastBlockId="31.[189,1399,267,400]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
The species determination rests mainly on the genetic analyses. We chose the predominant genetic group (C5) to describe a new species, but due to its short reproduction period (May–June, 
<bibRefCitation id="EFB7349C8C55FFD387EFA05CCB7E963E" author="Weber" box="[1179,1390,555,581]" etAl="et al." firstAuthor="Weber" journalOrPublisher="Comptes Rendus Biologies" pageId="30" pageNumber="31" pagination="553 - 560" part="337" refId="ref19581" refString="Weber A. A. - T., Stohr S. &amp; Chenuil A. 2014. Genetic data, reproduction season and reproductive strategy data support the existence of biological species in Ophioderma longicauda. Comptes Rendus Biologies 337: 553 - 560. https: // doi. org / 10.1016 / j. crvi. 2014.07.007" title="Genetic data, reproduction season and reproductive strategy data support the existence of biological species in Ophioderma longicauda" type="journal article" year="2014">
Weber 
<emphasis id="B952957F8C55FFD3879BA05BCB3A963E" box="[1263,1322,555,581]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
2014
</bibRefCitation>
) most specimens of this cluster happen to have been collected outside the brooding time. Since brooding is one of the key characters of this species, we felt that the 
<typeStatus id="549DF7CF8C54FFD2800EA358CDF19532" box="[890,993,303,329]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
should be a brooding female. It is not ideal that the chosen animal has only a single intact arm remaining, but in a pentaradially symmetric animal one arm is sufficient and representative of the others.
</paragraph>
<caption id="DF5919E58C55FFD383C9A4C1CFDD9391" ID-DOI="http://doi.org/10.5281/zenodo.3637797" ID-Zenodo-Dep="3637797" httpUri="https://zenodo.org/record/3637797/files/figure.png" pageId="30" pageNumber="31" startId="30.[189,232,1718,1744]" targetBox="[189,1398,630,1678]" targetPageId="30">
<paragraph id="8B99496D8C55FFD383C9A4C1CFDD9391" blockId="30.[189,1399,1718,2027]" pageId="30" pageNumber="31">
<emphasis id="B952957F8C55FFD383C9A4C1CF0992AB" bold="true" box="[189,281,1718,1744]" pageId="30" pageNumber="31">Fig. 12.</emphasis>
SEM images of skeletal elements of 
<taxonomicName id="4C2632EE8C55FFD381BAA4C1CDD892AB" authority="Stöhr &amp; Weber &amp; Boissin &amp; Chenuil, 2020" authorityName="Stöhr &amp; Weber &amp; Boissin &amp; Chenuil" authorityYear="2020" box="[718,968,1718,1744]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="30" pageNumber="31" phylum="Echinodermata" rank="species" species="zibrowii" status="sp. nov.">
<emphasis id="B952957F8C55FFD381BAA4C1CDD892AB" box="[718,968,1718,1744]" italics="true" pageId="30" pageNumber="31">Ophioderma zibrowii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26128048C55FFD380A4A4C0CA3D92AA" box="[976,1069,1719,1745]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
, from non-type specimen of 15.4 mm disc diameter (SMNH-178455), Crete, Greece. 
<emphasis id="B952957F8C55FFD38023A4AECD7C9288" bold="true" box="[855,876,1753,1779]" pageId="30" pageNumber="31">A</emphasis>
. Radial shield, external aspect, white arrows point to larger peripheral pores. 
<emphasis id="B952957F8C55FFD38133A48ACC4B936C" bold="true" box="[583,603,1789,1815]" pageId="30" pageNumber="31">B</emphasis>
. Radial shield, internal aspect. 
<emphasis id="B952957F8C55FFD380A8A48ACDE1936C" bold="true" box="[988,1009,1789,1815]" pageId="30" pageNumber="31">C</emphasis>
. Scale-free part magnified from Fig. 12A, note the absence of pores on the denser area (naked in intact animal, see Fig. 11A). 
<emphasis id="B952957F8C55FFD38671A557CB0A9341" bold="true" box="[1285,1306,1824,1850]" pageId="30" pageNumber="31">D</emphasis>
. Dental plate fragments. 
<emphasis id="B952957F8C55FFD382F7A534CF879326" bold="true" box="[387,407,1859,1885]" pageId="30" pageNumber="31">E</emphasis>
. Adradial genital plate, lateral aspect. 
<emphasis id="B952957F8C55FFD38028A534CD7E9326" bold="true" box="[860,878,1859,1885]" pageId="30" pageNumber="31">F</emphasis>
. Distal end of Fig. 12E. 
<emphasis id="B952957F8C55FFD387E7A534CABA9326" bold="true" box="[1171,1194,1859,1885]" pageId="30" pageNumber="31">G</emphasis>
. Abradial genital plate. 
<emphasis id="B952957F8C55FFD38270A510CF0B93FA" bold="true" box="[260,283,1895,1921]" pageId="30" pageNumber="31">H</emphasis>
. Oral plate, abradial face. 
<emphasis id="B952957F8C55FFD3813AA510CC4993FA" bold="true" box="[590,601,1895,1921]" pageId="30" pageNumber="31">I</emphasis>
. Oral plate, adradial face, black arrows point to oral papillae sockets and pores. 
<emphasis id="B952957F8C55FFD3824BA5FDCF5E93DF" bold="true" box="[319,334,1930,1956]" pageId="30" pageNumber="31">J</emphasis>
. Arm vertebra, proximal face. 
<emphasis id="B952957F8C55FFD381CAA5FDCCC593DF" bold="true" box="[702,725,1930,1956]" pageId="30" pageNumber="31">K</emphasis>
. Arm vertebra, distal face. 
<emphasis id="B952957F8C55FFD3876EA5FDCA3E93DF" bold="true" box="[1050,1070,1930,1956]" pageId="30" pageNumber="31">L</emphasis>
. Lateral arm plate, external face. 
<emphasis id="B952957F8C55FFD38388A5DACF0893BC" bold="true" box="[252,280,1965,1991]" pageId="30" pageNumber="31">M</emphasis>
. Lateral arm plate, internal face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; Sp = spurs; vent = ventral.
</paragraph>
</caption>
<paragraph id="8B99496D8C54FFD283C9A3C0CB649774" blockId="31.[189,1399,439,784]" pageId="31" pageNumber="32">
Morphological differentiation from 
<taxonomicName id="4C2632EE8C54FFD2811AA3C0CD0A95AA" baseAuthorityName="Bruzelius" baseAuthorityYear="1805" box="[622,794,439,466]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="longicauda">
<emphasis id="B952957F8C54FFD2811AA3C0CC9395AA" box="[622,643,439,465]" italics="true" pageId="31" pageNumber="32">O</emphasis>
. 
<emphasis id="B952957F8C54FFD281E2A3C0CD0A95AA" box="[662,794,439,465]" italics="true" pageId="31" pageNumber="32">longicauda</emphasis>
</taxonomicName>
rests on the usually naked radial shields and the single, flat arm plates. Also, 
<taxonomicName id="4C2632EE8C54FFD2816BA3ACCCB6958F" authority="Stöhr &amp; Weber &amp; Boissin &amp; Chenuil, 2020" authorityName="Stöhr &amp; Weber &amp; Boissin &amp; Chenuil" authorityYear="2020" box="[543,678,474,501]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="zibrowii" status="sp. nov.">
<emphasis id="B952957F8C54FFD2816BA3ACCC24958E" box="[543,564,475,501]" italics="true" pageId="31" pageNumber="32">O</emphasis>
. 
<emphasis id="B952957F8C54FFD28133A3ADCCB6958F" box="[583,678,474,500]" italics="true" pageId="31" pageNumber="32">zibrowii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26128048C54FFD281C6A3ACCD01958E" box="[690,785,475,501]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
usually lacks red pigments which are common in 
<taxonomicName id="4C2632EE8C54FFD283C9A389CF749663" baseAuthorityName="Bruzelius" baseAuthorityYear="1805" box="[189,356,510,536]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="longicauda">
<emphasis id="B952957F8C54FFD283C9A389CEC29663" box="[189,210,510,536]" italics="true" pageId="31" pageNumber="32">O</emphasis>
. 
<emphasis id="B952957F8C54FFD28394A389CF749663" box="[224,356,510,536]" italics="true" pageId="31" pageNumber="32">longicauda</emphasis>
</taxonomicName>
, whereas green pigments are common in 
<taxonomicName id="4C2632EE8C54FFD28032A389CDD89663" authority="Stöhr &amp; Weber &amp; Boissin &amp; Chenuil, 2020" authorityName="Stöhr &amp; Weber &amp; Boissin &amp; Chenuil" authorityYear="2020" box="[838,968,510,536]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="zibrowii" status="sp. nov.">
<emphasis id="B952957F8C54FFD28032A389CDD89663" box="[838,968,510,536]" italics="true" pageId="31" pageNumber="32">O. zibrowii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26128048C54FFD280BAA389CA389663" box="[974,1064,510,536]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
but absent in 
<taxonomicName id="4C2632EE8C54FFD287BCA389CB7F9663" baseAuthorityName="Bruzelius" baseAuthorityYear="1805" box="[1224,1391,510,536]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="longicauda">
<emphasis id="B952957F8C54FFD287BCA389CB7F9663" box="[1224,1391,510,536]" italics="true" pageId="31" pageNumber="32">O. longicauda</emphasis>
</taxonomicName>
. In addition, 
<taxonomicName id="4C2632EE8C54FFD2823AA055CFC29640" authority="Stöhr &amp; Weber &amp; Boissin &amp; Chenuil, 2020" authorityName="Stöhr &amp; Weber &amp; Boissin &amp; Chenuil" authorityYear="2020" box="[334,466,545,572]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="zibrowii" status="sp. nov.">
<emphasis id="B952957F8C54FFD2823AA055CF739647" box="[334,355,546,572]" italics="true" pageId="31" pageNumber="32">O</emphasis>
. 
<emphasis id="B952957F8C54FFD28207A056CFC29640" box="[371,466,545,571]" italics="true" pageId="31" pageNumber="32">zibrowii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26128048C54FFD282AFA055CC279647" box="[475,567,546,572]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
commonly has a series of white and black spots along the distal edge of the dorsal arm plates, which are shared with 
<taxonomicName id="4C2632EE8C54FFD28193A032CD759625" authority="Stöhr &amp; Weber &amp; Boissin &amp; Chenuil, 2020" authorityName="Stöhr &amp; Weber &amp; Boissin &amp; Chenuil" authorityYear="2020" box="[743,869,580,607]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="hybrida" status="sp. nov.">
<emphasis id="B952957F8C54FFD28193A032CCEC9624" box="[743,764,581,607]" italics="true" pageId="31" pageNumber="32">O</emphasis>
. 
<emphasis id="B952957F8C54FFD2807EA033CD759625" box="[778,869,580,606]" italics="true" pageId="31" pageNumber="32">hybrida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26128048C54FFD28019A032CDD79624" box="[877,967,581,607]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C2632EE8C54FFD2808BA032CA889625" authorityName="Greeff" authorityYear="1882" box="[1023,1176,581,607]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="guineense">
<emphasis id="B952957F8C54FFD2808BA032CA049624" box="[1023,1044,581,607]" italics="true" pageId="31" pageNumber="32">O</emphasis>
. 
<emphasis id="B952957F8C54FFD28757A032CA889625" box="[1059,1176,581,606]" italics="true" pageId="31" pageNumber="32">guineense</emphasis>
</taxonomicName>
. The radial shields differ microscopically from those of 
<taxonomicName id="4C2632EE8C54FFD28107A01FCD0C96F9" baseAuthorityName="Bruzelius" baseAuthorityYear="1805" box="[627,796,616,642]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="longicauda">
<emphasis id="B952957F8C54FFD28107A01FCC9896F9" box="[627,648,616,642]" italics="true" pageId="31" pageNumber="32">O</emphasis>
. 
<emphasis id="B952957F8C54FFD281ECA01FCD0C96F9" box="[664,796,616,642]" italics="true" pageId="31" pageNumber="32">longicauda</emphasis>
</taxonomicName>
in the clear demarcation of an embossed part with finer meshed stereom, representing the naked visible part of the radial shields in the intact animal. The major part of the radial shield is overlapped and covered by disc scales, obscuring its true shape. Possibly, the more elongated oral plates and higher lateral arm plates in 
<taxonomicName id="4C2632EE8C54FFD280BDA0A5CA409697" authority="Stöhr &amp; Weber &amp; Boissin &amp; Chenuil, 2020" authorityName="Stöhr &amp; Weber &amp; Boissin &amp; Chenuil" authorityYear="2020" box="[969,1104,722,748]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="zibrowii" status="sp. nov.">
<emphasis id="B952957F8C54FFD280BDA0A5CDCE9697" box="[969,990,722,748]" italics="true" pageId="31" pageNumber="32">O</emphasis>
. 
<emphasis id="B952957F8C54FFD28085A0A5CA409697" box="[1009,1104,722,748]" italics="true" pageId="31" pageNumber="32">zibrowii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26128048C54FFD28728A0A5CAAB9697" box="[1116,1211,722,748]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
distinguish the species, but these small differences need to be studied with morphometric methods and larger samples.
</paragraph>
<paragraph id="8B99496D8C54FFD283C9A140CF0997A5" blockId="31.[189,1399,823,990]" pageId="31" pageNumber="32">
Brooding females have been collected from 
<collectingCountry id="F33109FD8C54FFD281A6A140CD37972A" box="[722,807,823,849]" name="Cyprus" pageId="31" pageNumber="32">Cyprus</collectingCountry>
and 
<collectingCountry id="F33109FD8C54FFD28010A140CDDE972A" box="[868,974,823,849]" name="Lebanon" pageId="31" pageNumber="32">Lebanon</collectingCountry>
, but they belong to another genetic cluster (C6). Morphologically they cannot be distinguished at this time and we consider them to be allopatric populations within the same species. They are excluded from the concept of the new species 
<taxonomicName id="4C2632EE8C54FFD283C9A1D6CF5297C0" authority="Stöhr &amp; Weber &amp; Boissin &amp; Chenuil, 2020" authorityName="Stöhr &amp; Weber &amp; Boissin &amp; Chenuil" authorityYear="2020" box="[189,322,929,955]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="zibrowii" status="sp. nov.">
<emphasis id="B952957F8C54FFD283C9A1D6CEC297C0" box="[189,210,929,955]" italics="true" pageId="31" pageNumber="32">O</emphasis>
. 
<emphasis id="B952957F8C54FFD28397A1D6CF5297C0" box="[227,322,929,955]" italics="true" pageId="31" pageNumber="32">zibrowii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26128048C54FFD28238A1D6CFBC97C0" box="[332,428,929,955]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
, as new evidence may in the future lead to them being recognized as a different species.
</paragraph>
</subSubSection>
</treatment>
</document>