194 lines
20 KiB
XML
194 lines
20 KiB
XML
<document id="E89AF73272D2B1DE307E86368857EAA6" ID-CLB-Dataset="51205" ID-DOI="10.11646/zootaxa.4061.5.3" ID-GBIF-Dataset="8e4c370b-2a1d-4bc4-a714-16f9f3a1bd65" ID-ISSN="1175-5326" ID-Zenodo-Dep="264294" ID-ZooBank="6EEE4C42-D292-490A-9549-CB3428430920" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459680617424" checkinUser="plazi" docAuthor="Häckel, Martin" docDate="2016" docId="038F87EE01543000FF0BD14BFADCFDF3" docLanguage="en" docName="zootaxa.4061.5.3.pdf" docOrigin="Zootaxa 4061 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Craspedophorus carbonarius Harold 1879" docType="treatment" docVersion="10" lastPageNumber="510" masterDocId="FFB6FF9601513006FF9CD433FFDDFFEB" masterDocTitle="A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 1. Revision of the Craspedophorus reflexus group (Coleoptera: Carabidae)" masterLastPageNumber="526" masterPageNumber="504" pageNumber="509" updateTime="1698207076058" updateUser="ExternalLinkService">
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<mods:title id="0F8723639CDE5BF77BC3D7986FDEEABB">A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 1. Revision of the Craspedophorus reflexus group (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="F6D9C1C43D65504EC6724D97507C9F4C">Häckel, Martin</mods:namePart>
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<mods:title id="7B2CBB6AD4988730C89E09C6031D92CF">Zootaxa</mods:title>
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<mods:date id="03C92AF00C1A0231AE36EDA48CE90E8D">2016</mods:date>
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<mods:number id="48CDE63203AA9045F76929C2494AD4EF">4061</mods:number>
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<treatment id="038F87EE01543000FF0BD14BFADCFDF3" ID-DOI="http://doi.org/10.5281/zenodo.6088129" ID-GBIF-Taxon="119333002" ID-Zenodo-Dep="6088129" LSID="urn:lsid:plazi:treatment:038F87EE01543000FF0BD14BFADCFDF3" httpUri="http://treatment.plazi.org/id/038F87EE01543000FF0BD14BFADCFDF3" lastPageId="6" lastPageNumber="510" pageId="5" pageNumber="509">
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<subSubSection id="C33C657301543003FF0BD14BFD0CFA79" box="[151,721,1400,1426]" pageId="5" pageNumber="509" type="nomenclature">
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<paragraph id="8B9936F801543003FF0BD14BFD0CFA79" blockId="5.[151,807,1400,1460]" box="[151,721,1400,1426]" pageId="5" pageNumber="509">
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<heading id="D0D1819401543003FF0BD14BFD0CFA79" bold="true" box="[151,721,1400,1426]" fontSize="11" level="1" pageId="5" pageNumber="509" reason="1">
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<emphasis id="B952EAEA01543003FF0BD14BFD0CFA79" bold="true" box="[151,721,1400,1426]" pageId="5" pageNumber="509">
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5.
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<taxonomicName id="4C264D7B01543003FF28D14BFD0CFA79" ID-CoL="6BC46" authority="Harold, 1879" authorityName="Harold" authorityYear="1879" box="[180,721,1400,1426]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="509" phylum="Arthropoda" rank="species" species="carbonarius">
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<emphasis id="B952EAEA01543003FF28D14BFDC8FA79" bold="true" box="[180,533,1400,1426]" italics="true" pageId="5" pageNumber="509">Craspedophorus carbonarius</emphasis>
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(
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<bibRefCitation id="EFB74B0901543003FDB8D14BFD14FA79" author="Harold" box="[548,713,1400,1426]" pageId="5" pageNumber="526" refString="Harold, E. von (1879) Bericht uber die von den Herren A. von Homeyer und P. Pogge in Angola und im Lunda-Reiche gesammelten Kafer. Coleopterologische Hefte, 16, 1 - 221." type="journal article" year="1879">Harold, 1879</bibRefCitation>
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)
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C657301543003FF0BD1A8FCFAFA58" box="[151,807,1435,1460]" pageId="5" pageNumber="509" type="description">
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<paragraph id="8B9936F801543003FF0BD1A8FCFAFA58" blockId="5.[151,807,1400,1460]" box="[151,807,1435,1460]" pageId="5" pageNumber="509">(Plate 2, Figs 15, 16, Plate 3, Fig. 23, Plate 4, Figs 30, 31.)</paragraph>
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</subSubSection>
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<subSubSection id="C33C657301543003FF0BD1D0FD16F9FC" pageId="5" pageNumber="509" type="reference_group">
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<paragraph id="8B9936F801543003FF0BD1D0FD16F9FC" blockId="5.[151,1437,1507,1559]" pageId="5" pageNumber="509">
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<treatmentCitationGroup id="AB3611D601543003FF0BD1D0FD16F9FC" pageId="5" pageNumber="509">
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<taxonomicName id="4C264D7B01543003FF0BD1D0FDF8FA12" authority="Harold 1879: 20" authorityName="Harold" authorityPageNumber="20" authorityYear="1879" box="[151,549,1507,1529]" class="Magnoliopsida" family="Brassicaceae" genus="Eudema" kingdom="Plantae" order="Brassicales" pageId="5" pageNumber="509" phylum="Tracheophyta" rank="species" species="carbonarium">
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<emphasis id="B952EAEA01543003FF0BD1D0FEA8FA12" box="[151,373,1507,1529]" italics="true" pageId="5" pageNumber="509">Eudema carbonarium</emphasis>
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<treatmentCitation id="0A8710E901543003FEE1D1D0FDF8FA12" author="Harold" box="[381,549,1507,1529]" page="20" pageId="5" pageNumber="509" year="1879">
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<bibRefCitation id="EFB74B0901543003FEE1D1D0FDDCFA12" author="Harold" box="[381,513,1507,1529]" pageId="5" pageNumber="526" refString="Harold, E. von (1879) Bericht uber die von den Herren A. von Homeyer und P. Pogge in Angola und im Lunda-Reiche gesammelten Kafer. Coleopterologische Hefte, 16, 1 - 221." type="journal article" year="1879">Harold 1879</bibRefCitation>
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: 20
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</treatmentCitation>
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</taxonomicName>
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(
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<typeStatus id="549D885A01543003FDAFD1D0FDBDFA12" box="[563,608,1507,1529]" pageId="5" pageNumber="509">type</typeStatus>
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loc. “
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<collectingCountry id="F331766801543003FD03D1D0FCDDFA12" box="[671,768,1507,1529]" name="Tanzania" pageId="5" pageNumber="509">Zanzibar</collectingCountry>
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”).
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<taxonomicName id="4C264D7B01543003FC87D1D0FA44FA12" ID-CoL="6BC46" authority="Burgeon 1930: 160" authorityName="Burgeon" authorityPageNumber="160" authorityYear="1930" box="[795,1433,1507,1529]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="509" phylum="Arthropoda" rank="species" species="carbonarius">
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<emphasis id="B952EAEA01543003FC87D1D0FB95FA12" box="[795,1096,1507,1529]" italics="true" pageId="5" pageNumber="509">Craspedophorus carbonarius</emphasis>
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<treatmentCitation id="0A8710E901543003FBD3D1D7FACBFA12" author="Burgeon" box="[1103,1302,1507,1529]" page="160" pageId="5" pageNumber="509" year="1930">
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<bibRefCitation id="EFB74B0901543003FBD3D1D7FB39FA12" author="Burgeon" box="[1103,1252,1507,1529]" pageId="5" pageNumber="526" refString="Burgeon, L. (1930) Les Panagaeini du Musee du Congo. Revue de Zoologie et de Botanique Africaines, 19, 151 - 166." type="journal article" year="1930">Burgeon 1930</bibRefCitation>
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: 160
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</treatmentCitation>
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, 1935b: 182
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</taxonomicName>
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.
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<treatmentCitation id="0A8710E901543003FF5BD232FE73F9FC" author="Basilewsky" box="[199,430,1537,1559]" page="172" pageId="5" pageNumber="509" year="1953">
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<bibRefCitation id="EFB74B0901543003FF5BD232FEA0F9FC" author="Basilewsky" box="[199,381,1537,1559]" pageId="5" pageNumber="525" refString="Basilewsky, P. (1953) Carabidae (Coleoptera, Adephaga). In: Exploration du Parc National de l´Upemba, Mission G. F. de Witte en collaboration avec W. Adam, A. Janssens, L. van Meel, et R. Verheyen (1946 - 1949), fascicules 10. Bruxelles (Institut de Parcs Nationaux du Congo-Belge), x + 252 pp." type="book chapter" year="1953">Basilewsky, 1953</bibRefCitation>
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: 172
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</treatmentCitation>
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,
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<treatmentCitation id="0A8710E901543003FE25D232FD1AF9FC" author="Hackel" box="[441,711,1537,1559]" page="80" pageId="5" pageNumber="509" year="2012">
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<bibRefCitation id="EFB74B0901543003FE25D232FD7FF9FC" author="Hackel" box="[441,674,1537,1559]" pageId="5" pageNumber="526" refString="Hackel, M. & Farkac, J. (2012) A check-list of the subfamily Panagaeinae Hope, 1838 of the World (Coleoptera: Carabidae). Studies and Reports, Taxonomical Series, 8 (1 - 2), 67 - 116." type="journal article" year="2012">Häckel et Farkač 2012</bibRefCitation>
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: 80
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C657301543000FF0BD277FC5EFE1E" lastPageId="6" lastPageNumber="510" pageId="5" pageNumber="509" type="materials_examined">
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<paragraph id="8B9936F801543003FF0BD277FF3CF923" blockId="5.[151,1437,1604,2025]" pageId="5" pageNumber="509">
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<emphasis id="B952EAEA01543003FF0BD277FE2AF9B6" bold="true" box="[151,503,1604,1629]" pageId="5" pageNumber="509">
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Material examined.
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<collectingCountry id="F331766801543003FE15D277FE2FF9B6" box="[393,498,1604,1629]" name="Ethiopia" pageId="5" pageNumber="509">Ethiopia</collectingCountry>
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.
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</emphasis>
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<specimenCount id="9D20FD7101543003FE62D276FDF9F9B6" box="[510,548,1605,1629]" pageId="5" pageNumber="509" type="generic">1♂</specimenCount>
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: “[Oromiya] Omo, Mago National Park, Jinka”, (Plate 3, Fig. 23, Plate 4, Fig. 30, cMH).
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<emphasis id="B952EAEA01543003FE89D25BFEB7F96A" bold="true" box="[277,362,1640,1665]" pageId="5" pageNumber="509">
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<collectingCountry id="F331766801543003FE89D25BFEB8F96A" box="[277,357,1640,1665]" name="Kenya" pageId="5" pageNumber="509">Kenya</collectingCountry>
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.
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</emphasis>
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1♀: “Indian Ocean,
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<collectingCountry id="F331766801543003FDC5D25BFD1AF96A" box="[601,711,1640,1665]" name="Kenya" pageId="5" pageNumber="509">Mombasa</collectingCountry>
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to Aldabra, on board m.v. “Maritime”, (
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<collectionCode id="ED37AE3D01543003FB0BD25BFB2CF96B" box="[1175,1265,1640,1664]" pageId="5" pageNumber="509">BMNH</collectionCode>
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);
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<specimenCount id="9D20FD7101543003FA9AD25BFAF1F96B" box="[1286,1324,1640,1664]" pageId="5" pageNumber="509" type="generic">1♂</specimenCount>
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: “
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<collectingCountry id="F331766801543003FAD8D25BFA50F96A" box="[1348,1421,1640,1665]" name="Kenya" pageId="5" pageNumber="509">Kenia</collectingCountry>
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”, (Plate 2, Fig. 15, Plate 4, Fig. 30,
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<collectionCode id="ED37AE3D01543003FD92D2BEFDB3F94E" box="[526,622,1677,1701]" httpUri="http://grbio.org/cool/e7dr-nxad" name="National Museum of Wales" pageId="5" pageNumber="509">NMWC</collectionCode>
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).
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<emphasis id="B952EAEA01543003FDE2D2BFFD2CF94E" bold="true" box="[638,753,1676,1701]" pageId="5" pageNumber="509">
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<collectingCountry id="F331766801543003FDE2D2BFFD34F94E" box="[638,745,1676,1701]" name="Tanzania" pageId="5" pageNumber="509">Tanzania</collectingCountry>
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.
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</emphasis>
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1♀: “Morogoro prov.,
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<quantity id="4CDE9B1D01543003FC69D2BFFBE1F94E" box="[1013,1084,1676,1701]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="509" unit="km" value="50.0">50 km</quantity>
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NE Mikumi”, (Plate 2, Fig. 16, cMH).
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</paragraph>
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<paragraph id="8B9936F801543000FF5BD2E7FEE2FEAB" blockId="5.[151,1437,1604,2025]" lastBlockId="6.[151,1437,151,536]" lastPageId="6" lastPageNumber="510" pageId="5" pageNumber="509">
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<emphasis id="B952EAEA01543003FF5BD2E7FEDBF906" bold="true" box="[199,262,1748,1773]" pageId="5" pageNumber="509">Note.</emphasis>
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The original description of this species was based on a single specimen, labelled “
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<collectingCountry id="F331766801543003FB3FD2E7FAD5F906" box="[1187,1288,1748,1773]" name="Tanzania" pageId="5" pageNumber="509">Sansibar</collectingCountry>
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” Description (in part, see
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<bibRefCitation id="EFB74B0901543003FEBFD2CBFE69F8FB" author="Harold" box="[291,436,1784,1809]" pageId="5" pageNumber="526" refString="Harold, E. von (1879) Bericht uber die von den Herren A. von Homeyer und P. Pogge in Angola und im Lunda-Reiche gesammelten Kafer. Coleopterologische Hefte, 16, 1 - 221." type="journal article" year="1879">Harold 1879</bibRefCitation>
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||
: 20). “Length
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<quantity id="4CDE9B1D01543003FDCBD2CBFD0EF8FB" box="[599,723,1784,1809]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="5" pageNumber="509" unit="mm" value="24.0" valueMax="25.0" valueMin="23.0">23–25 mm</quantity>
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. Body moderately convex, black, pronotal width hardly shorter [wider than long], rounded, almost hexagonal, elytra deeply striated and punctured, intervals fairly convex, quite densely punctured” [translated from Latin]. At the same time as that species [
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<taxonomicName id="4C264D7B01543003FB8CD372FAB8F8B3" authority="Boheman, 1848" authorityName="Boheman" authorityYear="1848" box="[1040,1381,1856,1881]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="509" phylum="Arthropoda" rank="species" species="impictus">
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<emphasis id="B952EAEA01543003FB8CD372FB4FF8B3" box="[1040,1170,1857,1880]" italics="true" pageId="5" pageNumber="509">C. impictus</emphasis>
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||
(
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<bibRefCitation id="EFB74B0901543003FB3ED373FA81F8B3" author="Boheman" box="[1186,1372,1856,1881]" pageId="5" pageNumber="525" refString="Boheman, C. H. (1848) Insecta Caffraria annis 1838 - 1845 a J. A. Wahlberg collecta amici auxillio suffultus. Pars I, Fasc. 1. (Carabici, Hydrocanthari, Gyrinii et Staphylinii). Fritze & Norstedt, Holmiae, viii + 625 pp. [1851]" type="book chapter" year="1848">Boheman, 1848</bibRefCitation>
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)
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</taxonomicName>
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], the same explorer [Pogge] collected also the other, very close species, which was originally identified also as
|
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<taxonomicName id="4C264D7B01543003FA1ED356FF2BF84B" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="509" phylum="Arthropoda" rank="species" species="impictus">
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<emphasis id="B952EAEA01543003FA1ED356FF2BF84B" italics="true" pageId="5" pageNumber="509">C. impictus</emphasis>
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</taxonomicName>
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, because of different sex. Only a single male and a single female were collected, each belonging to a different species. The other sexual differences are here hardly marked; I have not found any other differences except those of ventral setae in male protarsomeres, which are somewhat longer and stronger. The
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<collectingCountry id="F331766801543003FB7DD3E3FA98F802" box="[1249,1349,2000,2025]" name="Tanzania" pageId="5" pageNumber="509">Zanzibar</collectingCountry>
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species differs by pronotal shape, which although similar is relatively smaller than that of
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<taxonomicName id="4C264D7B01573000FBD5D4ABFB17FF5B" box="[1097,1226,152,176]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="510" phylum="Arthropoda" rank="species" species="impictus">
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||
<emphasis id="B952EAEA01573000FBD5D4ABFB81FF44" box="[1097,1116,152,175]" italics="true" pageId="6" pageNumber="510">C</emphasis>
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||
.
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<emphasis id="B952EAEA01573000FBF7D4AAFB17FF5B" box="[1131,1226,153,176]" italics="true" pageId="6" pageNumber="510">impictus</emphasis>
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</taxonomicName>
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, and hardly wider than long, maximum width at midlength, not behind midlength as in
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<taxonomicName id="4C264D7B01573000FC02D48EFBC3FF3F" box="[926,1054,189,212]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="510" phylum="Arthropoda" rank="species" species="impictus">
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<emphasis id="B952EAEA01573000FC02D48EFBC3FF3F" box="[926,1054,189,212]" italics="true" pageId="6" pageNumber="510">C. impictus</emphasis>
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</taxonomicName>
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, lateral margins forming an angle sharper than in
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<emphasis id="B952EAEA01573000FED0D4D3FE0BFF13" box="[332,470,224,248]" italics="true" pageId="6" pageNumber="510">
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<taxonomicName id="4C264D7B01573000FED0D4D3FE0FFF13" box="[332,466,224,248]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="510" phylum="Arthropoda" rank="species" species="impictus">C. impictus</taxonomicName>
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.
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</emphasis>
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Also elytral striae more coarsely and randomly punctured, punctuation transversely interrrupted and somewhat deeper than that on interval's surface. I add here diagnosis of this species” [translated from
|
||
<collectingCountry id="F331766801573000FF48D51BFEE8FEAB" box="[212,309,296,320]" name="Germany" pageId="6" pageNumber="510">German</collectingCountry>
|
||
].
|
||
</paragraph>
|
||
<paragraph id="8B9936F801573000FF5BD57EFC5EFE1E" blockId="6.[151,1437,151,536]" pageId="6" pageNumber="510">
|
||
<typeStatus id="549D885A01573000FF5BD57EFF22FE8E" box="[199,255,333,357]" pageId="6" pageNumber="510">Type</typeStatus>
|
||
not seen, and identified specimens are rare in collections. It appears though that specimens perfectly matching the original description belong to a species widespread along the eastern African shores, so far known from southern
|
||
<collectingCountry id="F331766801573000FEDED5A7FE79FE46" box="[322,420,404,429]" name="Ethiopia" pageId="6" pageNumber="510">Ethiopia</collectingCountry>
|
||
, eastern
|
||
<collectingCountry id="F331766801573000FD97D5A6FD89FE46" box="[523,596,405,429]" name="Kenya" pageId="6" pageNumber="510">Kenya</collectingCountry>
|
||
and eastern
|
||
<collectingCountry id="F331766801573000FD76D5A7FC92FE46" box="[746,847,404,429]" name="Tanzania" pageId="6" pageNumber="510">Tanzania</collectingCountry>
|
||
including
|
||
<collectingCountry id="F331766801573000FC51D5A7FBEEFE46" box="[973,1075,404,429]" name="Tanzania" pageId="6" pageNumber="510">Zanzibar</collectingCountry>
|
||
. The elytral coloration, namely macular reduction (especially apical) is quite variable even within the same sex (Plates 2, 3, Figs 15, 23,). More material is needed to better understand the intraspecific variability.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C657301573000FF5BD5CCFADCFDF3" box="[199,1281,511,536]" pageId="6" pageNumber="510" type="distribution">
|
||
<paragraph id="8B9936F801573000FF5BD5CCFADCFDF3" blockId="6.[151,1437,151,536]" box="[199,1281,511,536]" pageId="6" pageNumber="510">
|
||
<emphasis id="B952EAEA01573000FF5BD5CCFEBDFDF3" bold="true" box="[199,352,511,536]" pageId="6" pageNumber="510">Distribution.</emphasis>
|
||
<collectingCountry id="F331766801573000FEFBD5CCFE17FDF3" box="[359,458,511,536]" name="Ethiopia" pageId="6" pageNumber="510">Ethiopia</collectingCountry>
|
||
: Oromiya;
|
||
<collectingCountry id="F331766801573000FDDBD633FD49FDF3" box="[583,660,512,536]" name="Kenya" pageId="6" pageNumber="510">Kenya</collectingCountry>
|
||
: Coast Province;
|
||
<collectingCountry id="F331766801573000FCC4D5CCFC1CFDF3" box="[856,961,511,536]" name="Tanzania" pageId="6" pageNumber="510">Tanzania</collectingCountry>
|
||
: Morogoro,
|
||
<collectingCountry id="F331766801573000FBD7D5CCFB72FDF3" box="[1099,1199,511,536]" name="Tanzania" pageId="6" pageNumber="510">Zanzibar</collectingCountry>
|
||
Island.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |