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<document id="BB35193A485CD9524ECD5D1B4AB79585" ID-DOI="10.11646/zootaxa.5536.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="14239039" ID-ZooBank="DADA0F2F-C442-4B2F-8C08-D918CFFB8709" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1732798815797" checkinUser="plazi" docAuthor="De Brito, Chirlei D. &amp; Azevedo, Celso O." docDate="2024" docId="038CCD37FFA7BA6C3F8B75F378BBFD9B" docLanguage="en" docName="zootaxa.5536.3.2.pdf" docOrigin="Zootaxa 5536 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5536.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dissomphalus elzae De Brito &amp; Azevedo 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="410" masterDocId="FFB5B54FFFB3BA793F1C71017C6EFF82" masterDocTitle="Revision of the punctatus species-group of Dissomphalus Ashmead (Hymenoptera, Bethylidae)" masterLastPageNumber="431" masterPageNumber="389" pageNumber="409" updateTime="1732921489393" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="2D1995251844AE3B88EB5CCFF69178C5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C8038A093044C9DDFD0B8C7F8567420A">
<mods:title id="A791BA30542F5FA14119D66A7A95477E">Revision of the punctatus species-group of Dissomphalus Ashmead (Hymenoptera, Bethylidae)</mods:title>
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<mods:name id="AEAEF9EE2DC14C6E7FBC214CD2E2E3A2" type="personal">
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<mods:namePart id="77C54A2CB3DFE8FCC17942C323D978ED">De Brito, Chirlei D.</mods:namePart>
<mods:nameIdentifier id="A47CDA0788F917832663AB0E215F9448" type="ORCID">0000-0002-1081-744X</mods:nameIdentifier>
<mods:affiliation id="E560BC24C365038B280094733852E5AE">Universidade Federal do Espírito Santo, Avenida Fernando Ferrari, 514, Goiabeiras, 29075 - 910 Vitória, ES, Brazil.</mods:affiliation>
<mods:nameIdentifier id="03914882D41A545CEA18BF708D70D466" type="email">chirleidb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="BEE9164D49BD6EA3E136D246EFA79433" type="personal">
<mods:role id="DDAE64964300BBE8CEB0DD74828A82EE">
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<mods:namePart id="11E02932115B55F6015C59E4A7DBE83F">Azevedo, Celso O.</mods:namePart>
<mods:nameIdentifier id="1085EDDE3096BAA067874B35F3A089E3" type="ORCID">0000-0002-2423-9388</mods:nameIdentifier>
<mods:affiliation id="C1DC34FEE74FFA3E26B0F51AD520E670">Universidade Federal do Espírito Santo, Avenida Fernando Ferrari, 514, Goiabeiras, 29075 - 910 Vitória, ES, Brazil.</mods:affiliation>
<mods:nameIdentifier id="F2B090633531C62B950E427671FCC957" type="email">bethylidae@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="0A10C4984E8B36CE59F8C4CDCE74CC41">text</mods:typeOfResource>
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<mods:title id="C211276D55A6D96565AF49D04E9D352A">Zootaxa</mods:title>
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<mods:part id="5EFF1854C1BCAD3ECB877D48CF33A20A">
<mods:date id="EBBC150756B8B2421E114680FE17BB2D">2024</mods:date>
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<mods:number id="B0AEA4D969A98219FD6E0A3D25FF3904">2024-11-05</mods:number>
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<mods:number id="A8F8368310378DAEBC02CAA3148600E6">5536</mods:number>
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<mods:number id="804ABE5FECB0A2769DDEE3BE0AAC881E">3</mods:number>
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<mods:url id="FAB9DDC5E1ED16895B8231B92A323296">http://dx.doi.org/10.11646/zootaxa.5536.3.2</mods:url>
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<mods:identifier id="B50FD74BF1E9D5C18A55E4A38611F312" type="DOI">10.11646/zootaxa.5536.3.2</mods:identifier>
<mods:identifier id="28FCA68FE10D9700CF695AB00E3127AC" type="ISSN">1175-5326</mods:identifier>
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<treatment id="038CCD37FFA7BA6C3F8B75F378BBFD9B" ID-DOI="http://doi.org/10.5281/zenodo.14247967" ID-Zenodo-Dep="14247967" LSID="urn:lsid:plazi:treatment:038CCD37FFA7BA6C3F8B75F378BBFD9B" httpUri="http://treatment.plazi.org/id/038CCD37FFA7BA6C3F8B75F378BBFD9B" lastPageId="21" lastPageNumber="410" pageId="20" pageNumber="409">
<subSubSection id="C33F2FAAFFA7BA6D3F8B75F37EA4FA8F" box="[151,714,1266,1293]" pageId="20" pageNumber="409" type="nomenclature">
<paragraph id="8B9A7C21FFA7BA6D3F8B75F37EA4FA8F" blockId="20.[151,714,1266,1329]" box="[151,714,1266,1293]" pageId="20" pageNumber="409">
<heading id="D0D2CB4DFFA7BA6D3F8B75F37EA4FA8F" bold="true" box="[151,714,1266,1293]" fontSize="11" level="1" pageId="20" pageNumber="409" reason="1">
<emphasis id="B951A033FFA7BA6D3F8B75F37EA4FA8F" bold="true" box="[151,714,1266,1293]" pageId="20" pageNumber="409">
<taxonomicName id="4C2507A2FFA7BA6D3F8B75F37E0EFA8E" authority="Brito &amp; Azevedo" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[151,608,1266,1293]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="409" phylum="Arthropoda" rank="species" species="elzae" status="sp. nov.">
<emphasis id="B951A033FFA7BA6D3F8B75F37DE8FA8F" bold="true" box="[151,390,1266,1293]" italics="true" pageId="20" pageNumber="409">Dissomphalus elzae</emphasis>
Brito &amp; Azevedo
</taxonomicName>
,
<taxonomicNameLabel id="A2621D48FFA7BA6D3D7075F27EA4FA8F" box="[620,714,1267,1293]" pageId="20" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F2FAAFFA7BA6D3F8B74167D5CFAB3" box="[151,306,1303,1329]" pageId="20" pageNumber="409" type="description">
<paragraph id="8B9A7C21FFA7BA6D3F8B74167D5CFAB3" blockId="20.[151,714,1266,1329]" box="[151,306,1303,1329]" pageId="20" pageNumber="409">
(
<figureCitation id="131E60A4FFA7BA6D3F8374167C94FAB3" box="[159,250,1303,1329]" captionStart="FIGURE 1" captionStartId="12.[151,250,1639,1663]" captionTargetBox="[151,1435,181,1615]" captionTargetId="figure-18@12.[151,1435,181,1615]" captionTargetPageId="12" captionText="FIGURE 1. Dissomphalus sp. males. Patterns of metasomal tergal process in dorsal view. A. D. maryae sp. nov., paratype. B. D. punctatus. C. D. carolinae sp. nov., paratype. D. D. johannae sp. nov., paratype. E. D. deaneae sp. nov., paratype. F. D. margarethae sp. nov., paratype. Scale bars: 200 µm." figureDoi="http://doi.org/10.5281/zenodo.14239041" httpUri="https://zenodo.org/record/14239041/files/figure.png" pageId="20" pageNumber="409">Figs 1C</figureCitation>
,
<figureCitation id="131E60A4FFA7BA6D3E1A74167D44FAB3" box="[262,298,1303,1329]" captionStart="FIGURE 4" captionStartId="23.[151,250,1909,1933]" captionTargetBox="[228,1366,192,1872]" captionTargetId="figure-17@23.[214,1374,181,1884]" captionTargetPageId="23" captionText="FIGURE 4. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. elzae sp. nov., holotype. B. D. enedinae sp. nov., paratype. C. D. eulaliae sp. nov., holotype. D. D. evaristoae sp. nov., holotype. E. D. grazielae sp. nov., holotype. F. D. guidonae sp. nov., holotype. G. D. jaquelineae sp. nov., holotype. H. D. johannae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239047" httpUri="https://zenodo.org/record/14239047/files/figure.png" pageId="20" pageNumber="409">4A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33F2FAAFFA7BA6D3F8B745F7F97FA43" pageId="20" pageNumber="409" type="diagnosis">
<paragraph id="8B9A7C21FFA7BA6D3F8B745F7F97FA43" blockId="20.[151,1437,1374,2013]" pageId="20" pageNumber="409">
<emphasis id="B951A033FFA7BA6D3F8B745F7D64FAFA" bold="true" box="[151,266,1374,1400]" pageId="20" pageNumber="409">Diagnosis</emphasis>
. This species is easily recognized by having the aedeagal dorsal valve much smaller than harpe, gonostipe and aedeagal ventral valve, the apex of inner median filament short and directed dorsad, the basal cover plate very wide, and the aedeagal ventral valve much higher than aedeagal dorsal valve.
</paragraph>
</subSubSection>
<subSubSection id="C33F2FAAFFA7BA6D3FDB74CB7E81F8CF" pageId="20" pageNumber="409" type="description">
<paragraph id="8B9A7C21FFA7BA6D3FDB74CB7E81F8CF" blockId="20.[151,1437,1374,2013]" pageId="20" pageNumber="409">
<emphasis id="B951A033FFA7BA6D3FDB74CB7D3FFA66" bold="true" box="[199,337,1482,1508]" pageId="20" pageNumber="409">Description</emphasis>
. Head. Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina high in profile, incomplete apically, straight in profile or nearly so. Frons coriaceous, punctures large and dense. Vertex crest weakly incurved. Dorsal pronotal area with anterior margin coarse. Tergal process submedian, consisting of tuft with few setae, without depression. Posterior hypopygeal margin straight or nearly so. Genitalia. Harpe entirely wide, apical margin rounded, abaxial margin not projected; gonostipes shorter than harpe, ventral margin not excavated until outer margin of basivolsella; digitus short, basidorsal corner poorly projected; basivolsella without protuberance, without spines; aedeagal ventral valve with apex posterior to apex of dorsal one wide, progressively narrowing apicad; inner margin sinuous; outer margin sinuous, curved laterad; aedeagal dorsal valve narrow, with two pairs of apical lobe; outer lobe short, narrow, apical margin rounded, and mesad; apex of inner median filament aligned with apex of lobe, directed dorsad; basal cover plate entirely wide, posterior margin rounded; apodeme not extending beyond genital ring.
</paragraph>
</subSubSection>
<subSubSection id="C33F2FAAFFA7BA6C3FDB76577EB9FE9F" lastPageId="21" lastPageNumber="410" pageId="20" pageNumber="409" type="materials_examined">
<paragraph id="8B9A7C21FFA7BA6C3FDB76577EB9FE9F" blockId="20.[151,1437,1374,2013]" lastBlockId="21.[151,1436,151,537]" lastPageId="21" lastPageNumber="410" pageId="20" pageNumber="409">
<emphasis id="B951A033FFA7BA6D3FDB76577DC1F8F2" bold="true" box="[199,431,1878,1904]" pageId="20" pageNumber="409">Material examined</emphasis>
.
<materialsCitation id="3B4D767CFFA7BA6D3EDF76567E28F817" collectingDate="1982-07-09" collectingDateMax="1982-08-07" collectingDateMin="1982-07-09" collectionCode="CNCI" collectorName="B. Giil" country="Costa Rica" elevation="1200" location="San Vito de Coto Brus" municipality="Las Cruces" pageId="20" pageNumber="409" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="549EC283FFA7BA6D3EDF76567E40F8F3" box="[451,558,1879,1905]" pageId="20" pageNumber="409" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D23B7A8FFA7BA6D3D2376567E0BF8F3" box="[575,613,1879,1905]" count="1" pageId="20" pageNumber="409" type="male">1♂</specimenCount>
:
<collectingCountry id="F3323CB1FFA7BA6D3D6A76567F4FF8F3" box="[630,801,1879,1905]" name="Costa Rica" pageId="20" pageNumber="409">COSTA RICA</collectingCountry>
,
<emphasis id="B951A033FFA7BA6D3C2E7656782FF8F2" box="[818,1089,1879,1905]" italics="true" pageId="20" pageNumber="409">
<location id="8EFA2AFAFFA7BA6D3C2E7656782FF8F2" LSID="urn:lsid:plazi:treatment:038CCD37FFA7BA6C3F8B75F378BBFD9B:8EFA2AFAFFA7BA6D3C2E7656782FF8F2" box="[818,1089,1879,1905]" country="Costa Rica" municipality="Las Cruces" name="San Vito de Coto Brus" pageId="20" pageNumber="409">San Vito de Coto Brus</location>
</emphasis>
,
<collectingMunicipality id="6BFEE65BFFA7BA6D3B49765678B6F8F3" box="[1109,1240,1879,1905]" pageId="20" pageNumber="409">Las Cruces</collectingMunicipality>
<quantity id="4CDDD1C4FFA7BA6D3BF87656792FF8F3" box="[1252,1345,1879,1905]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="20" pageNumber="409" unit="m" value="1200.0">
<elevation id="00089B12FFA7BA6D3BF87656792FF8F3" box="[1252,1345,1879,1905]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="20" pageNumber="409" unit="m" value="1200.0">1200 m</elevation>
</quantity>
,
<date id="FF9B5AE1FFA7BA6D3A4E76567D70F817" pageId="20" pageNumber="409" value="1982-07-09" valueMax="1982-08-07" valueMin="1982-07-09">
<collectingDate id="EFDFA309FFA7BA6D3A4E76567D70F817" pageId="20" pageNumber="409" value="1982-07-09" valueMax="1982-08-07" valueMin="1982-07-09">9.VII– 7.VIII.1982</collectingDate>
</date>
,
<collectorName id="26D019F7FFA7BA6D3E37767A7D16F817" box="[299,376,1915,1941]" pageId="20" pageNumber="409">B. Giil</collectorName>
mal. trap (
<collectionCode id="ED34E4E4FFA7BA6D3EE9767A7E50F817" box="[501,574,1915,1941]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="20" pageNumber="409">CNCI</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4D767CFFA7BA6D3D4D767A7807F83B" collectingDate="1975-06-10" collectionCode="FSCA" country="Colombia" county="Dept. Valle" elevation="560" location="Buenaventura" municipality="Central de Anchicaya" pageId="20" pageNumber="409" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="549EC283FFA7BA6D3D4D767A7EACF817" box="[593,706,1915,1941]" pageId="20" pageNumber="409" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F3323CB1FFA7BA6D3DD2767A7F08F817" box="[718,870,1915,1941]" name="Colombia" pageId="20" pageNumber="409">COLOMBIA</collectingCountry>
,
<emphasis id="B951A033FFA7BA6D3C6F767A7F9EF817" box="[883,1008,1915,1941]" italics="true" pageId="20" pageNumber="409">
<collectingCounty id="62FB04ADFFA7BA6D3C6F767A7F9EF817" box="[883,1008,1915,1941]" pageId="20" pageNumber="409">Dept. Valle</collectingCounty>
</emphasis>
,
<specimenCount id="9D23B7A8FFA7BA6D3CE3767A784AF817" box="[1023,1060,1915,1941]" count="1" pageId="20" pageNumber="409" type="male">1♂</specimenCount>
,
<collectingMunicipality id="6BFEE65BFFA7BA6D3B2E767A7946F817" box="[1074,1320,1915,1941]" pageId="20" pageNumber="409">Central de Anchicaya</collectingMunicipality>
,
<quantity id="4CDDD1C4FFA7BA6D3A28767A7912F817" box="[1332,1404,1915,1941]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="20" pageNumber="409" unit="km" value="30.0">30 km</quantity>
E.
<location id="8EFA2AFAFFA7BA6D3F8B769E7D5BF83B" LSID="urn:lsid:plazi:treatment:038CCD37FFA7BA6C3F8B75F378BBFD9B:8EFA2AFAFFA7BA6D3F8B769E7D5BF83B" box="[151,309,1951,1977]" country="Colombia" county="Dept. Valle" municipality="Central de Anchicaya" name="Buenaventura" pageId="20" pageNumber="409">Buenaventura</location>
,
<location id="8EFA2AFAFFA7BA6D3E5E769E7E0CF83B" LSID="urn:lsid:plazi:treatment:038CCD37FFA7BA6C3F8B75F378BBFD9B:8EFA2AFAFFA7BA6D3E5E769E7E0CF83B" box="[322,610,1951,1977]" country="Colombia" county="Dept. Valle" municipality="Central de Anchicaya" name="Tropical Very Wet Forest" pageId="20" pageNumber="409">Tropical Very Wet Forest</location>
,
<quantity id="4CDDD1C4FFA7BA6D3D73769F7ED7F83B" box="[623,697,1950,1977]" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="20" pageNumber="409" unit="m" value="560.0">
<elevation id="00089B12FFA7BA6D3D73769F7ED7F83B" box="[623,697,1950,1977]" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="20" pageNumber="409" unit="m" value="560.0">560 m</elevation>
</quantity>
, R. C.
<location id="8EFA2AFAFFA7BA6D3DE2769E7F17F83B" LSID="urn:lsid:plazi:treatment:038CCD37FFA7BA6C3F8B75F378BBFD9B:8EFA2AFAFFA7BA6D3DE2769E7F17F83B" box="[766,889,1951,1977]" country="Colombia" county="Dept. Valle" municipality="Central de Anchicaya" name="Wilkerson" pageId="20" pageNumber="409">Wilkerson</location>
,
<date id="FF9B5AE1FFA7BA6D3C99769F786AF83B" box="[901,1028,1950,1977]" pageId="20" pageNumber="409" value="1975-06-10">
<collectingDate id="EFDFA309FFA7BA6D3C99769F786AF83B" box="[901,1028,1950,1977]" pageId="20" pageNumber="409" value="1975-06-10">10.VI.1975</collectingDate>
</date>
(
<collectionCode id="ED34E4E4FFA7BA6D3B09769E780FF83B" box="[1045,1121,1951,1977]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="20" pageNumber="409" type="Museum">FSCA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4D767CFFA7BA6D3B6A769E7F4FF85F" collectingDate="1991-03-10" collectingDateMax="1991-03-19" collectingDateMin="1991-03-10" collectionCode="EMUS" collectorName="F. D. Parker" country="Costa Rica" location="Upala" pageId="20" pageNumber="409" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectingCountry id="F3323CB1FFA7BA6D3B6A769E7976F83B" box="[1142,1304,1951,1977]" name="Costa Rica" pageId="20" pageNumber="409">COSTA RICA</collectingCountry>
<emphasis id="B951A033FFA7BA6D3A3D769E79F8F83B" box="[1313,1430,1951,1977]" italics="true" pageId="20" pageNumber="409">
<taxonomicName id="4C2507A2FFA7BA6D3A3D769E7926F83B" authorityName="Lockington" authorityYear="1877" box="[1313,1352,1951,1977]" class="Malacostraca" family="Mithracidae" genus="Ala" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="409" phylum="Arthropoda" rank="genus">Ala</taxonomicName>
[juela]
</emphasis>
,
<specimenCount id="9D23B7A8FFA7BA6D3F8B76C27CD2F85F" box="[151,188,1987,2013]" count="1" pageId="20" pageNumber="409" type="male">1♂</specimenCount>
,
<quantity id="4CDDD1C4FFA7BA6D3FD476C27D60F85F" box="[200,270,1987,2013]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="409" unit="km" value="20.0">20 km</quantity>
S
<location id="8EFA2AFAFFA7BA6D3E3776C27D1FF85F" LSID="urn:lsid:plazi:treatment:038CCD37FFA7BA6C3F8B75F378BBFD9B:8EFA2AFAFFA7BA6D3E3776C27D1FF85F" box="[299,369,1987,2013]" country="Costa Rica" name="Upala" pageId="20" pageNumber="409">Upala</location>
,
<date id="FF9B5AE1FFA7BA6D3E6776C27E48F85F" box="[379,550,1987,2013]" pageId="20" pageNumber="409" value="1991-03-10" valueMax="1991-03-19" valueMin="1991-03-10">
<collectingDate id="EFDFA309FFA7BA6D3E6776C27E48F85F" box="[379,550,1987,2013]" pageId="20" pageNumber="409" value="1991-03-10" valueMax="1991-03-19" valueMin="1991-03-10">10–19.III.1991</collectingDate>
</date>
,
<collectorName id="26D019F7FFA7BA6D3D2D76C27ED9F85F" box="[561,695,1987,2013]" pageId="20" pageNumber="409">F. D. Parker</collectorName>
(
<collectionCode id="ED34E4E4FFA7BA6D3DD976C27F77F85F" box="[709,793,1987,2013]" pageId="20" pageNumber="409">EMUS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4D767CFFA7BA6C3C3076C27D33FF33" collectingDate="1987-05-23" collectionCode="MUCR" collectorName="Col. Paul Hanson" country="Costa Rica" elevation="300" lastPageId="21" lastPageNumber="410" location="Iriquois" pageId="20" pageNumber="409" specimenCount="1" specimenCount-male="1" stateProvince="Limon" typeStatus="paratype">
<emphasis id="B951A033FFA7BA6D3C3076C27F1AF85F" box="[812,884,1987,2013]" italics="true" pageId="20" pageNumber="409">
<collectingRegion id="49E1B2C3FFA7BA6D3C3076C27F1AF85F" box="[812,884,1987,2013]" country="Costa Rica" name="Limon" pageId="20" pageNumber="409">Limón</collectingRegion>
</emphasis>
,
<specimenCount id="9D23B7A8FFA7BA6D3C9D76C27FC8F85F" box="[897,934,1987,2013]" count="1" pageId="20" pageNumber="409" type="male">1♂</specimenCount>
, sur de
<location id="8EFA2AFAFFA7BA6D3CE076C27838F85F" LSID="urn:lsid:plazi:treatment:038CCD37FFA7BA6C3F8B75F378BBFD9B:8EFA2AFAFFA7BA6D3CE076C27838F85F" box="[1020,1110,1987,2013]" country="Costa Rica" name="Iriquois" pageId="20" pageNumber="409" stateProvince="Limon">Iriquois</location>
,
<quantity id="4CDDD1C4FFA7BA6D3B7C76C278C7F85F" box="[1120,1193,1987,2013]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="20" pageNumber="409" unit="m" value="300.0">
<elevation id="00089B12FFA7BA6D3B7C76C278C7F85F" box="[1120,1193,1987,2013]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="20" pageNumber="409" unit="m" value="300.0">300 m</elevation>
</quantity>
,
<date id="FF9B5AE1FFA7BA6D3BA876C27947F85F" box="[1204,1321,1987,2013]" pageId="20" pageNumber="409" value="1987-05-23">
<collectingDate id="EFDFA309FFA7BA6D3BA876C27947F85F" box="[1204,1321,1987,2013]" pageId="20" pageNumber="409" value="1987-05-23">23.V.1987</collectingDate>
</date>
,
<collectorName id="26D019F7FFA7BA6C3A2876C27C83FF33" lastPageId="21" lastPageNumber="410" pageId="20" pageNumber="409">Col. Paul Hanson</collectorName>
(
<collectionCode id="ED34E4E4FFA6BA6C3FE071967D3BFF33" box="[252,341,151,177]" country="Costa Rica" httpUri="http://grbio.org/cool/ww9x-1v7b" name="Museo de Insectos" pageId="21" pageNumber="410">MUCR</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4D767CFFA6BA6C3E74719679F6FF33" box="[360,1432,151,177]" collectingDate="1988-12" collectionCode="MUCR" collectorName="San Vito &amp; Col. P. Hanson" country="Costa Rica" elevation="1200" location="Las Cruces" municipality="Jardin Bot." pageId="21" pageNumber="410" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
<emphasis id="B951A033FFA6BA6C3E7471967D86FF32" box="[360,488,151,176]" italics="true" pageId="21" pageNumber="410">
<collectingRegion id="49E1B2C3FFA6BA6C3E7471967D86FF32" box="[360,488,151,176]" country="Costa Rica" name="Puntarenas" pageId="21" pageNumber="410">Puntarenas</collectingRegion>
</emphasis>
,
<specimenCount id="9D23B7A8FFA6BA6C3EEA71967E74FF33" box="[502,538,151,177]" count="1" pageId="21" pageNumber="410" type="male">1♂</specimenCount>
,
<collectorName id="26D019F7FFA6BA6C3D3B71967EE5FF33" box="[551,651,151,177]" pageId="21" pageNumber="410">San Vito</collectorName>
,
<collectingMunicipality id="6BFEE65BFFA6BA6C3D8A71967F7AFF33" box="[662,788,151,177]" pageId="21" pageNumber="410">Jardin Bot.</collectingMunicipality>
,
<location id="8EFA2AFAFFA6BA6C3C0371967FCEFF33" LSID="urn:lsid:plazi:treatment:038CCD37FFA7BA6C3F8B75F378BBFD9B:8EFA2AFAFFA6BA6C3C0371967FCEFF33" box="[799,928,151,177]" country="Costa Rica" municipality="Jardin Bot." name="Las Cruces" pageId="21" pageNumber="410" stateProvince="Puntarenas">Las Cruces</location>
,
<date id="FF9B5AE1FFA6BA6C3CB67196787DFF33" box="[938,1043,151,177]" pageId="21" pageNumber="410" value="1988-12">
<collectingDate id="EFDFA309FFA6BA6C3CB67196787DFF33" box="[938,1043,151,177]" pageId="21" pageNumber="410" value="1988-12">XII.1988</collectingDate>
</date>
,
<quantity id="4CDDD1C4FFA6BA6C3B0271967818FF33" box="[1054,1142,151,177]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="21" pageNumber="410" unit="m" value="1200.0">
<elevation id="00089B12FFA6BA6C3B0271967818FF33" box="[1054,1142,151,177]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="21" pageNumber="410" unit="m" value="1200.0">1200 m</elevation>
</quantity>
,
<collectorName id="26D019F7FFA6BA6C3B9D71967947FF33" box="[1153,1321,151,177]" pageId="21" pageNumber="410">Col. P. Hanson</collectorName>
(
<collectionCode id="ED34E4E4FFA6BA6C3A2B719679FEFF33" box="[1335,1424,151,177]" country="Costa Rica" httpUri="http://grbio.org/cool/ww9x-1v7b" name="Museo de Insectos" pageId="21" pageNumber="410">MUCR</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4D767CFFA6BA6C3F8B71BA7840FF57" box="[151,1070,187,213]" collectingDate="1987-04" collectionCode="MUCR" collectorName="Col. W. Eberhard" country="Costa Rica" location="San Antonio de Escazu" pageId="21" pageNumber="410" specimenCount="2" specimenCount-male="2" stateProvince="San Jose" typeStatus="paratype">
<emphasis id="B951A033FFA6BA6C3F8B71BA7C95FF56" box="[151,251,187,213]" italics="true" pageId="21" pageNumber="410">
<collectingRegion id="49E1B2C3FFA6BA6C3F8B71BA7C95FF56" box="[151,251,187,213]" country="Costa Rica" name="San Jose" pageId="21" pageNumber="410">San Jose</collectingRegion>
</emphasis>
,
<specimenCount id="9D23B7A8FFA6BA6C3E1571BA7D40FF57" box="[265,302,187,213]" count="2" pageId="21" pageNumber="410" type="male">2♂</specimenCount>
,
<location id="8EFA2AFAFFA6BA6C3E2671BA7E2FFF57" LSID="urn:lsid:plazi:treatment:038CCD37FFA7BA6C3F8B75F378BBFD9B:8EFA2AFAFFA6BA6C3E2671BA7E2FFF57" box="[314,577,187,213]" country="Costa Rica" name="San Antonio de Escazu" pageId="21" pageNumber="410" stateProvince="San Jose">San Antonio de Escazu</location>
, 1300,
<date id="FF9B5AE1FFA6BA6C3D8F71BA7E81FF57" box="[659,751,187,213]" pageId="21" pageNumber="410" value="1987-04">
<collectingDate id="EFDFA309FFA6BA6C3D8F71BA7E81FF57" box="[659,751,187,213]" pageId="21" pageNumber="410" value="1987-04">IV.1987</collectingDate>
</date>
,
<collectorName id="26D019F7FFA6BA6C3DE771BA7FD0FF57" box="[763,958,187,213]" pageId="21" pageNumber="410">Col. W. Eberhard</collectorName>
(
<collectionCode id="ED34E4E4FFA6BA6C3CD171BA7848FF57" box="[973,1062,187,213]" country="Costa Rica" httpUri="http://grbio.org/cool/ww9x-1v7b" name="Museo de Insectos" pageId="21" pageNumber="410">MUCR</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4D767CFFA6BA6C3B2671BA7F79FF7B" collectingDate="1982-07-02" collectingDateMax="1982-07-09" collectingDateMin="1982-07-02" collectionCode="CNCI" collectorName="B. Gill" country="Costa Rica" elevation="1200" location="Las Cruces" municipality="San Vito de Coto Brus" pageId="21" pageNumber="410" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B951A033FFA6BA6C3B2671BA795AFF56" box="[1082,1332,187,213]" italics="true" pageId="21" pageNumber="410">
<collectingMunicipality id="6BFEE65BFFA6BA6C3B2671BA795AFF56" box="[1082,1332,187,213]" pageId="21" pageNumber="410">San Vito de Coto Brus</collectingMunicipality>
</emphasis>
,
<specimenCount id="9D23B7A8FFA6BA6C3A5E71BA7909FF57" box="[1346,1383,187,213]" count="1" pageId="21" pageNumber="410" type="male">1♂</specimenCount>
,
<location id="8EFA2AFAFFA6BA6C3A6871BA7C8BFF7B" LSID="urn:lsid:plazi:treatment:038CCD37FFA7BA6C3F8B75F378BBFD9B:8EFA2AFAFFA6BA6C3A6871BA7C8BFF7B" country="Costa Rica" municipality="San Vito de Coto Brus" name="Las Cruces" pageId="21" pageNumber="410">Las Cruces</location>
<quantity id="4CDDD1C4FFA6BA6C3FF271DE7D28FF7B" box="[238,326,223,249]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="21" pageNumber="410" unit="m" value="1200.0">
<elevation id="00089B12FFA6BA6C3FF271DE7D28FF7B" box="[238,326,223,249]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="21" pageNumber="410" unit="m" value="1200.0">1200 m</elevation>
</quantity>
,
<date id="FF9B5AE1FFA6BA6C3E4871DE7D83FF7B" box="[340,493,223,249]" pageId="21" pageNumber="410" value="1982-07-02" valueMax="1982-07-09" valueMin="1982-07-02">
<collectingDate id="EFDFA309FFA6BA6C3E4871DE7D83FF7B" box="[340,493,223,249]" pageId="21" pageNumber="410" value="1982-07-02" valueMax="1982-07-09" valueMin="1982-07-02">2–9.VII.1982</collectingDate>
</date>
,
<collectorName id="26D019F7FFA6BA6C3EE671DE7E26FF7B" box="[506,584,223,249]" pageId="21" pageNumber="410">B. Gill</collectorName>
mal. trap (
<collectionCode id="ED34E4E4FFA6BA6C3DDA71DE7F61FF7B" box="[710,783,223,249]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="21" pageNumber="410">CNCI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4D767CFFA6BA6C3C3871DE7EBDFE9F" collectingDate="1993-02-05" collectingDateMax="1993-02-28" collectingDateMin="1993-02-05" collectionCode="MIUP" collectorName="C. Mono &amp; Est. Inrenare &amp; Nac. &amp; R. Cambra. Malaise" country="Panama" location="Darien" pageId="21" pageNumber="410" specimenCount="1" specimenCount-male="1" stateProvince="Darien" typeStatus="paratype">
<collectingCountry id="F3323CB1FFA6BA6C3C3871DE7FF0FF7B" box="[804,926,223,249]" name="Panama" pageId="21" pageNumber="410">PANAMA</collectingCountry>
,
<emphasis id="B951A033FFA6BA6C3CB671DE7860FF7A" box="[938,1038,223,249]" italics="true" pageId="21" pageNumber="410">
<collectorName id="26D019F7FFA6BA6C3CB671DE7860FF7A" box="[938,1038,223,249]" pageId="21" pageNumber="410">C. Mono</collectorName>
</emphasis>
,
<collectorName id="26D019F7FFA6BA6C3B0271DE78DFFF7B" box="[1054,1201,223,249]" pageId="21" pageNumber="410">Est. Inrenare</collectorName>
,
<specimenCount id="9D23B7A8FFA6BA6C3BA271DE788CFF7B" box="[1214,1250,223,249]" count="1" pageId="21" pageNumber="410" type="male">1♂</specimenCount>
, P.
<collectorName id="26D019F7FFA6BA6C3A1071DE792FFF7B" box="[1292,1345,223,249]" pageId="21" pageNumber="410">Nac.</collectorName>
<collectingRegion id="49E1B2C3FFA6BA6C3A5571DE79F6FF7B" box="[1353,1432,223,249]" country="Panama" name="Darien" pageId="21" pageNumber="410">Darién</collectingRegion>
,
<collectingRegion id="49E1B2C3FFA6BA6C3F8B70027C88FE9F" box="[151,230,259,285]" country="Panama" name="Darien" pageId="21" pageNumber="410">Darién</collectingRegion>
,
<date id="FF9B5AE1FFA6BA6C3FED70037DEBFE9F" box="[241,389,258,285]" pageId="21" pageNumber="410" value="1993-02-05" valueMax="1993-02-28" valueMin="1993-02-05">
<collectingDate id="EFDFA309FFA6BA6C3FED70037DEBFE9F" box="[241,389,258,285]" pageId="21" pageNumber="410" value="1993-02-05" valueMax="1993-02-28" valueMin="1993-02-05">5–28.II.1993</collectingDate>
</date>
,
<collectorName id="26D019F7FFA6BA6C3E8C70027E1EFE9F" box="[400,624,259,285]" pageId="21" pageNumber="410">R. Cambra. Malaise</collectorName>
(
<collectionCode id="ED34E4E4FFA6BA6C3D6370027EA5FE9F" box="[639,715,259,285]" pageId="21" pageNumber="410">MIUP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F2FAAFFA6BA6C3FDB70277E3EFEC3" box="[199,592,294,321]" pageId="21" pageNumber="410" type="distribution">
<paragraph id="8B9A7C21FFA6BA6C3FDB70277E3EFEC3" blockId="21.[151,1436,151,537]" box="[199,592,294,321]" pageId="21" pageNumber="410">
<emphasis id="B951A033FFA6BA6C3FDB70277D37FEC2" bold="true" box="[199,345,294,320]" pageId="21" pageNumber="410">Distribution</emphasis>
.
<collectingCountry id="F3323CB1FFA6BA6C3E7B70267D8AFEC3" box="[359,484,295,321]" name="Costa Rica" pageId="21" pageNumber="410">Costa Rica</collectingCountry>
,
<collectingCountry id="F3323CB1FFA6BA6C3EEC70267E22FEC3" box="[496,588,295,321]" name="Panama" pageId="21" pageNumber="410">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F2FAAFFA6BA6C3FDB704A7D9EFE0B" pageId="21" pageNumber="410" type="etymology">
<paragraph id="8B9A7C21FFA6BA6C3FDB704A7D9EFE0B" blockId="21.[151,1436,151,537]" pageId="21" pageNumber="410">
<emphasis id="B951A033FFA6BA6C3FDB704A7D26FEE7" bold="true" box="[199,328,331,357]" pageId="21" pageNumber="410">Etymology</emphasis>
. This species is named after the mathematician Elza Furtado Gomide, one of the pioneers in the field of Mathematics in
<collectingCountry id="F3323CB1FFA6BA6C3EB8706E7D82FE0B" box="[420,492,367,393]" name="Brazil" pageId="21" pageNumber="410">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F2FAAFFA6BA6C3FDB709278BBFD9B" pageId="21" pageNumber="410" type="discussion">
<paragraph id="8B9A7C21FFA6BA6C3FDB709278BBFD9B" blockId="21.[151,1436,151,537]" pageId="21" pageNumber="410">
<emphasis id="B951A033FFA6BA6C3FDB70927D5AFE2F" bold="true" box="[199,308,403,429]" pageId="21" pageNumber="410">Remarks</emphasis>
. It is similar to
<taxonomicName id="4C2507A2FFA6BA6C3EF070927E29FE2E" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[492,583,403,428]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="410" phylum="Arthropoda" rank="species" species="napo" status="sp. nov.">
<emphasis id="B951A033FFA6BA6C3EF070927E29FE2E" box="[492,583,403,428]" italics="true" pageId="21" pageNumber="410">D. napo</emphasis>
</taxonomicName>
<emphasis id="B951A033FFA6BA6C3D4C70927EC4FE2F" bold="true" box="[592,682,403,429]" pageId="21" pageNumber="410">
<taxonomicNameLabel id="A2621D48FFA6BA6C3D4C70927EC4FE2F" box="[592,682,403,429]" pageId="21" pageNumber="410" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having the harpe entirely wide with dorsal margin straight, the aedeagal dorsal outer lobes short with apex of inner median filament dorsad. However,
<taxonomicName id="4C2507A2FFA6BA6C3B9E70B6788EFE53" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1154,1248,439,465]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="410" phylum="Arthropoda" rank="species" species="elzae" status="sp. nov.">
<emphasis id="B951A033FFA6BA6C3B9E70B6788EFE53" box="[1154,1248,439,465]" italics="true" pageId="21" pageNumber="410">D. elzae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFA6BA6C3BF470B6792DFE53" bold="true" box="[1256,1347,439,465]" pageId="21" pageNumber="410">
<taxonomicNameLabel id="A2621D48FFA6BA6C3BF470B6792DFE53" box="[1256,1347,439,465]" pageId="21" pageNumber="410" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has the basal cover plate very wide, the aedeagal ventral valve sinuous, and the digitus short and wide, whereas
<taxonomicName id="4C2507A2FFA6BA6C3A0B70DA791FFE76" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1303,1393,475,500]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="410" phylum="Arthropoda" rank="species" species="napo">
<emphasis id="B951A033FFA6BA6C3A0B70DA791FFE76" box="[1303,1393,475,500]" italics="true" pageId="21" pageNumber="410">D. napo</emphasis>
</taxonomicName>
has the basal cover plate narrow, the aedeagal ventral valve straight, and the digitus long and narrow.
</paragraph>
</subSubSection>
</treatment>
</document>
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