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<document id="C27DF2B8A1D9491F85D5C37E4CD0A33E" ID-DOI="10.11646/zootaxa.5536.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="14239039" ID-ZooBank="DADA0F2F-C442-4B2F-8C08-D918CFFB8709" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1732798815797" checkinUser="plazi" docAuthor="De Brito, Chirlei D. &amp; Azevedo, Celso O." docDate="2024" docId="038CCD37FF90BA5C3F8B77C77E42FBB7" docLanguage="en" docName="zootaxa.5536.3.2.pdf" docOrigin="Zootaxa 5536 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5536.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dissomphalus ritae De Brito &amp; Azevedo 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="426" masterDocId="FFB5B54FFFB3BA793F1C71017C6EFF82" masterDocTitle="Revision of the punctatus species-group of Dissomphalus Ashmead (Hymenoptera, Bethylidae)" masterLastPageNumber="431" masterPageNumber="389" pageNumber="424" updateTime="1732921489393" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="AF5FD2E3E47520907D9C0625E8F16E4D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="08A3CB221C0B1B09D51760847E933662">
<mods:title id="D3D98655651937A8C9F8D8C04CA30037">Revision of the punctatus species-group of Dissomphalus Ashmead (Hymenoptera, Bethylidae)</mods:title>
</mods:titleInfo>
<mods:name id="70DC6356185EBD0C9922C40494CEAC70" type="personal">
<mods:role id="EAA29826E5404924E4D4ADCE12504032">
<mods:roleTerm id="DB5D3631955091BC5E83C5FC11F3C437">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CD787B7AA6799C726C8CFB7F58881831">De Brito, Chirlei D.</mods:namePart>
<mods:nameIdentifier id="EA38D2B042C0B4D99F2A03BEB24F5C56" type="ORCID">0000-0002-1081-744X</mods:nameIdentifier>
<mods:affiliation id="561462685DA5AFE10B5CB1251B0A1F52">Universidade Federal do Espírito Santo, Avenida Fernando Ferrari, 514, Goiabeiras, 29075 - 910 Vitória, ES, Brazil.</mods:affiliation>
<mods:nameIdentifier id="D2021204E8881BD00DA22579EBBA5313" type="email">chirleidb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="184F790620A2D46689BC9EC8602C2717" type="personal">
<mods:role id="F2BC43CFF27512F877D2D82A13F04B38">
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</mods:role>
<mods:namePart id="ADE421841E8BD09F06235CEBFA6F9EEF">Azevedo, Celso O.</mods:namePart>
<mods:nameIdentifier id="688FB95718390A0CAF27BEA00181657E" type="ORCID">0000-0002-2423-9388</mods:nameIdentifier>
<mods:affiliation id="2EC699EB8A9E16D0DC21CEA602F409A8">Universidade Federal do Espírito Santo, Avenida Fernando Ferrari, 514, Goiabeiras, 29075 - 910 Vitória, ES, Brazil.</mods:affiliation>
<mods:nameIdentifier id="803353826C055BC5BCFAF36D8CF82557" type="email">bethylidae@gmail.com</mods:nameIdentifier>
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<mods:titleInfo id="58F696AF8E1B39CDAAE60CC9B8FF7CB6">
<mods:title id="FF44F170884A3FC824BB57CBBE622C50">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="773E10E475550840F06410F5CD76517F">
<mods:date id="167E7E043C72AC4180200F362ADF4FC8">2024</mods:date>
<mods:detail id="4534217E42ADFD78D26789CBDACE3B1B" type="pubDate">
<mods:number id="F6DE748DCC6E0B5331F5F0F50A880295">2024-11-05</mods:number>
</mods:detail>
<mods:detail id="B68B6198F5A3ABE4F65A5E955930CB38" type="volume">
<mods:number id="F1566D5D971E4402CE21C186E6169D52">5536</mods:number>
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<mods:detail id="30BF57AEE90B2A84570AA0E5D7F46B75" type="issue">
<mods:number id="A60A80F71391EAAD2BF516A345ECA63D">3</mods:number>
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<mods:url id="E0D473F293A5F6A75831F795D189E925">http://dx.doi.org/10.11646/zootaxa.5536.3.2</mods:url>
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<mods:classification id="86A89505128AE8477D49C2660D98173A">journal article</mods:classification>
<mods:identifier id="048E29F05873D8D0EAE0DB979A0D47FC" type="DOI">10.11646/zootaxa.5536.3.2</mods:identifier>
<mods:identifier id="BBD5EE6352C6CECE969424B77D0828AF" type="ISSN">1175-5326</mods:identifier>
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<treatment id="038CCD37FF90BA5C3F8B77C77E42FBB7" ID-DOI="http://doi.org/10.5281/zenodo.14248009" ID-Zenodo-Dep="14248009" LSID="urn:lsid:plazi:treatment:038CCD37FF90BA5C3F8B77C77E42FBB7" httpUri="http://treatment.plazi.org/id/038CCD37FF90BA5C3F8B77C77E42FBB7" lastPageId="37" lastPageNumber="426" pageId="35" pageNumber="424">
<subSubSection id="C33F2FAAFF90BA5A3F8B77C77EABF963" box="[151,709,1734,1761]" pageId="35" pageNumber="424" type="nomenclature">
<paragraph id="8B9A7C21FF90BA5A3F8B77C77EABF963" blockId="35.[151,709,1734,1797]" box="[151,709,1734,1761]" pageId="35" pageNumber="424">
<heading id="D0D2CB4DFF90BA5A3F8B77C77EABF963" bold="true" box="[151,709,1734,1761]" fontSize="11" level="1" pageId="35" pageNumber="424" reason="1">
<emphasis id="B951A033FF90BA5A3F8B77C77EABF963" bold="true" box="[151,709,1734,1761]" pageId="35" pageNumber="424">
<taxonomicName id="4C2507A2FF90BA5A3F8B77C77E35F962" authority="Brito &amp; Azevedo" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[151,603,1734,1761]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="424" phylum="Arthropoda" rank="species" species="ritae" status="sp. nov.">
<emphasis id="B951A033FF90BA5A3F8B77C77DEFF963" bold="true" box="[151,385,1734,1761]" italics="true" pageId="35" pageNumber="424">Dissomphalus ritae</emphasis>
Brito &amp; Azevedo
</taxonomicName>
,
<taxonomicNameLabel id="A2621D48FF90BA5A3D7477C67EABF963" box="[616,709,1735,1761]" pageId="35" pageNumber="424" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F2FAAFF90BA5A3F8B77EA7D43F887" box="[151,301,1771,1797]" pageId="35" pageNumber="424" type="description">
<paragraph id="8B9A7C21FF90BA5A3F8B77EA7D43F887" blockId="35.[151,709,1734,1797]" box="[151,301,1771,1797]" pageId="35" pageNumber="424">
(
<figureCitation id="131E60A4FF90BA5A3F8377EA7C94F887" box="[159,250,1771,1797]" captionStart="FIGURE 1" captionStartId="12.[151,250,1639,1663]" captionTargetBox="[151,1435,181,1615]" captionTargetId="figure-18@12.[151,1435,181,1615]" captionTargetPageId="12" captionText="FIGURE 1. Dissomphalus sp. males. Patterns of metasomal tergal process in dorsal view. A. D. maryae sp. nov., paratype. B. D. punctatus. C. D. carolinae sp. nov., paratype. D. D. johannae sp. nov., paratype. E. D. deaneae sp. nov., paratype. F. D. margarethae sp. nov., paratype. Scale bars: 200 µm." figureDoi="http://doi.org/10.5281/zenodo.14239041" httpUri="https://zenodo.org/record/14239041/files/figure.png" pageId="35" pageNumber="424">Figs 1C</figureCitation>
,
<figureCitation id="131E60A4FF90BA5A3E1A77EA7D4BF887" box="[262,293,1771,1797]" captionStart="FIGURE 6" captionStartId="36.[151,250,1905,1929]" captionTargetBox="[225,1364,190,1872]" captionTargetId="figure-18@36.[214,1373,181,1881]" captionTargetPageId="36" captionText="FIGURE 6. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. maryae sp. nov., holotype. B. D. meirelesae sp. nov., holotype. C. D. mirandae sp. nov., holotype. D. D. nisiae sp. nov., holotype. E. D. penhae sp. nov., holotype. F. D. ritae sp. nov. holotype. G. D. silveirae sp. nov., paratype. H. D. soniae sp. nov., holotype. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239051" httpUri="https://zenodo.org/record/14239051/files/figure.png" pageId="35" pageNumber="424">6F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33F2FAAFF90BA5A3F8B76337E33F817" pageId="35" pageNumber="424" type="diagnosis">
<paragraph id="8B9A7C21FF90BA5A3F8B76337E33F817" blockId="35.[151,1437,1842,1941]" pageId="35" pageNumber="424">
<emphasis id="B951A033FF90BA5A3F8B76337D64F8CE" bold="true" box="[151,266,1842,1868]" pageId="35" pageNumber="424">Diagnosis</emphasis>
. This species is easily recognized by having the tergal process with a thin tuft of setae, the harpe wide with posterior margin straight, the inner median filament short, the aedeagal dorsal valve robust and the basivolsellar protuberance narrow and well projected.
</paragraph>
</subSubSection>
<caption id="DF5A2CA9FF97BA5D3F8B76707C8EF874" ID-DOI="http://doi.org/10.5281/zenodo.14239051" ID-Zenodo-Dep="14239051" httpUri="https://zenodo.org/record/14239051/files/figure.png" pageId="36" pageNumber="425" startId="36.[151,250,1905,1929]" targetBox="[225,1364,190,1872]" targetPageId="36" targetType="figure">
<paragraph id="8B9A7C21FF97BA5D3F8B76707C8EF874" blockId="36.[151,1437,1905,2038]" pageId="36" pageNumber="425">
<emphasis id="B951A033FF97BA5D3F8B76707D7AF80B" bold="true" box="[151,276,1905,1929]" pageId="36" pageNumber="425">FIGURE 6.</emphasis>
Genitalia of
<taxonomicName id="4C2507A2FF97BA5D3E8776707E21F80B" box="[411,591,1905,1929]" class="Insecta" family="Bethylidae" genus="Dissomphalus" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="425" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B951A033FF97BA5D3E8776707E45F80B" box="[411,555,1905,1929]" italics="true" pageId="36" pageNumber="425">Dissomphalus</emphasis>
sp.
</taxonomicName>
males in ventral (in the left) and dorsal view (in the right).
<emphasis id="B951A033FF97BA5D3BAE767078AAF80B" bold="true" box="[1202,1220,1905,1929]" pageId="36" pageNumber="425">A</emphasis>
.
<taxonomicName id="4C2507A2FF97BA5D3BCD76737953F80B" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1233,1341,1906,1929]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="425" phylum="Arthropoda" rank="species" species="maryae" status="sp. nov.">
<emphasis id="B951A033FF97BA5D3BCD76737953F80B" box="[1233,1341,1906,1929]" italics="true" pageId="36" pageNumber="425">D. maryae</emphasis>
</taxonomicName>
<emphasis id="B951A033FF97BA5D3A58767079FBF80B" bold="true" box="[1348,1429,1905,1929]" pageId="36" pageNumber="425">
<taxonomicNameLabel id="A2621D48FF97BA5D3A58767079FBF80B" box="[1348,1429,1905,1929]" pageId="36" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype.
<emphasis id="B951A033FF97BA5D3FE076947D63F82F" bold="true" box="[252,269,1941,1965]" pageId="36" pageNumber="425">B</emphasis>
.
<taxonomicName id="4C2507A2FF97BA5D3E0676977DCBF82F" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[282,421,1941,1965]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="425" phylum="Arthropoda" rank="species" species="meirelesae" status="sp. nov.">
<emphasis id="B951A033FF97BA5D3E0676977DCBF82F" box="[282,421,1941,1965]" italics="true" pageId="36" pageNumber="425">D. meirelesae</emphasis>
</taxonomicName>
<emphasis id="B951A033FF97BA5D3EB776947D92F82F" bold="true" box="[427,508,1941,1965]" pageId="36" pageNumber="425">
<taxonomicNameLabel id="A2621D48FF97BA5D3EB776947D92F82F" box="[427,508,1941,1965]" pageId="36" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype.
<emphasis id="B951A033FF97BA5D3D7276947EEEF82F" bold="true" box="[622,640,1941,1965]" pageId="36" pageNumber="425">C</emphasis>
.
<taxonomicName id="4C2507A2FF97BA5D3D9076977F62F82F" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[652,780,1941,1965]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="425" phylum="Arthropoda" rank="species" species="mirandae" status="sp. nov.">
<emphasis id="B951A033FF97BA5D3D9076977F62F82F" box="[652,780,1941,1965]" italics="true" pageId="36" pageNumber="425">D. mirandae</emphasis>
</taxonomicName>
<emphasis id="B951A033FF97BA5D3C0E76947F0DF82F" bold="true" box="[786,867,1941,1965]" pageId="36" pageNumber="425">
<taxonomicNameLabel id="A2621D48FF97BA5D3C0E76947F0DF82F" box="[786,867,1941,1965]" pageId="36" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype.
<emphasis id="B951A033FF97BA5D3CC976947F89F82F" bold="true" box="[981,999,1941,1965]" pageId="36" pageNumber="425">D</emphasis>
.
<taxonomicName id="4C2507A2FF97BA5D3CEF76977821F82F" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1011,1103,1942,1965]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="425" phylum="Arthropoda" rank="species" species="nisiae" status="sp. nov.">
<emphasis id="B951A033FF97BA5D3CEF76977821F82F" box="[1011,1103,1942,1965]" italics="true" pageId="36" pageNumber="425">D. nisiae</emphasis>
</taxonomicName>
<emphasis id="B951A033FF97BA5D3B49769478CBF82F" bold="true" box="[1109,1189,1941,1965]" pageId="36" pageNumber="425">
<taxonomicNameLabel id="A2621D48FF97BA5D3B49769478CBF82F" box="[1109,1189,1941,1965]" pageId="36" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype.
<emphasis id="B951A033FF97BA5D3A0B76947946F82F" bold="true" box="[1303,1320,1941,1965]" pageId="36" pageNumber="425">E</emphasis>
.
<taxonomicName id="4C2507A2FF97BA5D3A28769779F2F82F" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1332,1436,1941,1965]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="425" phylum="Arthropoda" rank="species" species="penhae" status="sp. nov.">
<emphasis id="B951A033FF97BA5D3A28769779F2F82F" box="[1332,1436,1941,1965]" italics="true" pageId="36" pageNumber="425">D. penhae</emphasis>
</taxonomicName>
<emphasis id="B951A033FF97BA5D3F8B76B87C86F853" bold="true" box="[151,232,1977,2001]" pageId="36" pageNumber="425">
<taxonomicNameLabel id="A2621D48FF97BA5D3F8B76B87C86F853" box="[151,232,1977,2001]" pageId="36" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype.
<emphasis id="B951A033FF97BA5D3E4776B87D04F853" bold="true" box="[347,362,1977,2001]" pageId="36" pageNumber="425">F</emphasis>
.
<taxonomicName id="4C2507A2FF97BA5D3E6B76BB7DA9F853" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[375,455,1978,2001]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="425" phylum="Arthropoda" rank="species" species="ritae" status="sp. nov.">
<emphasis id="B951A033FF97BA5D3E6B76BB7DA9F853" box="[375,455,1978,2001]" italics="true" pageId="36" pageNumber="425">D. ritae</emphasis>
</taxonomicName>
<emphasis id="B951A033FF97BA5D3ED176B87E70F853" bold="true" box="[461,542,1977,2001]" pageId="36" pageNumber="425">
<taxonomicNameLabel id="A2621D48FF97BA5D3ED176B87E70F853" box="[461,542,1977,2001]" pageId="36" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype.
<emphasis id="B951A033FF97BA5D3D9776B87EF1F853" bold="true" box="[651,671,1977,2001]" pageId="36" pageNumber="425">G</emphasis>
.
<taxonomicName id="4C2507A2FF97BA5D3DB776BB7F4CF853" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[683,802,1977,2001]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="425" phylum="Arthropoda" rank="species" species="silveirae" status="sp. nov.">
<emphasis id="B951A033FF97BA5D3DB776BB7F4CF853" box="[683,802,1977,2001]" italics="true" pageId="36" pageNumber="425">D. silveirae</emphasis>
</taxonomicName>
<emphasis id="B951A033FF97BA5D3C3476B87F17F853" bold="true" box="[808,889,1977,2001]" pageId="36" pageNumber="425">
<taxonomicNameLabel id="A2621D48FF97BA5D3C3476B87F17F853" box="[808,889,1977,2001]" pageId="36" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype.
<emphasis id="B951A033FF97BA5D3CF776B87F91F853" bold="true" box="[1003,1023,1977,2001]" pageId="36" pageNumber="425">H</emphasis>
.
<taxonomicName id="4C2507A2FF97BA5D3B1776BB7803F853" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1035,1133,1978,2001]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="425" phylum="Arthropoda" rank="species" species="soniae" status="sp. nov.">
<emphasis id="B951A033FF97BA5D3B1776BB7803F853" box="[1035,1133,1978,2001]" italics="true" pageId="36" pageNumber="425">D. soniae</emphasis>
</taxonomicName>
<emphasis id="B951A033FF97BA5D3B6876B878AAF853" bold="true" box="[1140,1220,1977,2001]" pageId="36" pageNumber="425">
<taxonomicNameLabel id="A2621D48FF97BA5D3B6876B878AAF853" box="[1140,1220,1977,2001]" pageId="36" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype. Scale bar: 20 µm.
</paragraph>
</caption>
<subSubSection id="C33F2FAAFF96BA5C3FDB71977FF1FDBF" pageId="37" pageNumber="426" type="description">
<paragraph id="8B9A7C21FF96BA5C3FDB71977902FD9B" blockId="37.[151,1437,150,1077]" pageId="37" pageNumber="426">
<emphasis id="B951A033FF96BA5C3FDB71977D3FFF32" bold="true" box="[199,337,150,176]" pageId="37" pageNumber="426">Description</emphasis>
. Head. Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina low in profile, incomplete apically, straight in profile or nearly so. Frons weakly coriaceous, punctures small and sparse. Vertex crest weakly outcurved. Dorsal pronotal area with anterior margin coarse. Tergal process submedian, consisting of tuft with few setae, without depression. Posterior hypopygeal margin straight or nearly so. Genitalia. Harpe entirely wide, apical margin angulate, abaxial margin not projected; gonostipes shorter than harpe, ventral margin not excavated until outer margin of basivolsella; digitus long, basidorsal corner poorly projected; basivolsella well delimited with narrow protuberance, without spines; aedeagal ventral valve with apex anterior to apex of dorsal one, wide, progressively narrowing apicad; inner margin outcurved; outer margin outcurved, curved laterad; aedeagal dorsal valve robust, with two pairs of apical lobe; outer lobe very long, narrow, apical margin angled and posterad; apex of inner median filament anterior to apex of lobe, directed slightly laterad; basal cover plate entirely wide, posterior margin straight or nearly so; apodeme extending beyond genital ring.
</paragraph>
<paragraph id="8B9A7C21FF96BA5C3FDB73237FF1FDBF" blockId="37.[151,1437,150,1077]" box="[199,927,546,573]" pageId="37" pageNumber="426">
<emphasis id="B951A033FF96BA5C3FDB73237D56FDBE" bold="true" box="[199,312,546,572]" pageId="37" pageNumber="426">Variation</emphasis>
. Tuft of setae of tergal process very short, hardly seen.
</paragraph>
</subSubSection>
<subSubSection id="C33F2FAAFF96BA5C3FDB73477D83FD73" pageId="37" pageNumber="426" type="materials_examined">
<paragraph id="8B9A7C21FF96BA5C3FDB73477D83FD73" blockId="37.[151,1437,150,1077]" pageId="37" pageNumber="426">
<emphasis id="B951A033FF96BA5C3FDB73477DC7FDE2" bold="true" box="[199,425,582,608]" pageId="37" pageNumber="426">Material examined</emphasis>
.
<materialsCitation id="3B4D767CFF96BA5C3EAA73467EF8FD07" collectingDate="2001-09" collectionCode="INPA" collectorName="Rod AM &amp; Malaise, J. F &amp; J. F. Vidal" country="Brazil" county="Km" location="Amazonas" municipality="Reserva Ducke" pageId="37" pageNumber="426" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="549EC283FF96BA5C3EAA73467E4FFDE3" box="[438,545,583,609]" pageId="37" pageNumber="426" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D23B7A8FF96BA5C3D3073467E3EFDE3" box="[556,592,583,609]" count="1" pageId="37" pageNumber="426" type="male">1♂</specimenCount>
,
<collectingCountry id="F3323CB1FF96BA5C3D4073467EA8FDE3" box="[604,710,583,609]" name="Brazil" pageId="37" pageNumber="426">BRAZIL</collectingCountry>
:
<emphasis id="B951A033FF96BA5C3DCC73467F2DFDE3" box="[720,835,583,609]" italics="true" pageId="37" pageNumber="426">
<location id="8EFA2AFAFF96BA5C3DCC73467F2DFDE3" LSID="urn:lsid:plazi:treatment:038CCD37FF90BA5C3F8B77C77E42FBB7:8EFA2AFAFF96BA5C3DCC73467F2DFDE3" box="[720,835,583,609]" country="Brazil" county="Km" municipality="Reserva Ducke" name="Amazonas" pageId="37" pageNumber="426">Amazonas</location>
</emphasis>
,
<location id="8EFA2AFAFF96BA5C3C4C73467FC5FDE3" LSID="urn:lsid:plazi:treatment:038CCD37FF90BA5C3F8B77C77E42FBB7:8EFA2AFAFF96BA5C3C4C73467FC5FDE3" box="[848,939,583,609]" country="Brazil" county="Km" municipality="Reserva Ducke" name="Manaus" pageId="37" pageNumber="426">Manaus</location>
,
<collectingMunicipality id="6BFEE65BFF96BA5C3CAA7346780DFDE3" box="[950,1123,583,609]" pageId="37" pageNumber="426">Reserva Ducke</collectingMunicipality>
,
<collectorName id="26D019F7FF96BA5C3B71734678A0FDE3" box="[1133,1230,583,609]" pageId="37" pageNumber="426">Rod AM</collectorName>
010
<collectingCounty id="62FB04ADFF96BA5C3A1973467941FDE3" box="[1285,1327,583,609]" pageId="37" pageNumber="426">Km</collectingCounty>
26, mata,
<date id="FF9B5AE1FF96BA5C3F8B736A7C9AFD07" box="[151,244,619,645]" pageId="37" pageNumber="426" value="2001-09">
<collectingDate id="EFDFA309FF96BA5C3F8B736A7C9AFD07" box="[151,244,619,645]" pageId="37" pageNumber="426" value="2001-09">IX.2001</collectingDate>
</date>
arm.
<collectorName id="26D019F7FF96BA5C3E25736A7DAEFD07" box="[313,448,619,645]" pageId="37" pageNumber="426">Malaise, J.F</collectorName>
.Vidal col. (
<collectionCode id="ED34E4E4FF96BA5C3D54736A7EE0FD07" box="[584,654,619,645]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="37" pageNumber="426">INPA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4D767CFF96BA5C3DBE736A795FFD07" box="[674,1329,619,645]" collectorName="Manaus &amp; Reserva Ducke" country="Brazil" location="Amazonas" pageId="37" pageNumber="426" specimenCount="1" typeStatus="paratype">
<typeStatus id="549EC283FF96BA5C3DBE736A7F7DFD07" box="[674,787,619,645]" pageId="37" pageNumber="426" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F3323CB1FF96BA5C3C3C736A7FE6FD07" box="[800,904,619,645]" name="Brazil" pageId="37" pageNumber="426">BRAZIL</collectingCountry>
,
<emphasis id="B951A033FF96BA5C3C89736A7866FD07" box="[917,1032,619,645]" italics="true" pageId="37" pageNumber="426">
<location id="8EFA2AFAFF96BA5C3C89736A7866FD07" LSID="urn:lsid:plazi:treatment:038CCD37FF90BA5C3F8B77C77E42FBB7:8EFA2AFAFF96BA5C3C89736A7866FD07" box="[917,1032,619,645]" country="Brazil" name="Amazonas" pageId="37" pageNumber="426">Amazonas</location>
</emphasis>
,
<collectorName id="26D019F7FF96BA5C3B04736A781DFD07" box="[1048,1139,619,645]" pageId="37" pageNumber="426">Manaus</collectorName>
,
<collectorName id="26D019F7FF96BA5C3B9D736A795FFD07" box="[1153,1329,619,645]" pageId="37" pageNumber="426">Reserva Ducke</collectorName>
</materialsCitation>
,
<materialsCitation id="3B4D767CFF96BA5C3A22736A7EF7FD2B" collectionCode="INPA" collectorName="Adesiva &amp; J. Vidal" country="Brazil" elevation="1" pageId="37" pageNumber="426" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D23B7A8FF96BA5C3A22736A790DFD07" box="[1342,1379,619,645]" count="1" pageId="37" pageNumber="426" type="male">1♂</specimenCount>
,
<quantity id="4CDDD1C4FF96BA5C3A6D736A79F9FD07" box="[1393,1431,619,645]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="426" unit="m" value="1.0">
<elevation id="00089B12FF96BA5C3A6D736A79F9FD07" box="[1393,1431,619,645]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="426" unit="m" value="1.0">1m</elevation>
</quantity>
, 8–15.IV, 1992,
<collectorName id="26D019F7FF96BA5C3E5B738E7DC8FD2B" box="[327,422,655,681]" pageId="37" pageNumber="426">Adesiva</collectorName>
,
<collectorName id="26D019F7FF96BA5C3EAF738E7E6AFD2B" box="[435,516,655,681]" pageId="37" pageNumber="426">J.Vidal</collectorName>
, col. (
<collectionCode id="ED34E4E4FF96BA5C3D50738E7EFCFD2B" box="[588,658,655,681]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="37" pageNumber="426">INPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4D767CFF96BA5C3DBA738E790BFD2B" box="[678,1381,654,681]" collectingDate="1992-07-17" collectionCode="INPA" collectorName="Adesiva &amp; Vidal" country="Brazil" elevation="500" pageId="37" pageNumber="426" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D23B7A8FF96BA5C3DBA738E7EA5FD2B" box="[678,715,655,681]" count="1" pageId="37" pageNumber="426" type="male">1♂</specimenCount>
,
<quantity id="4CDDD1C4FF96BA5C3DC5738F7F72FD2B" box="[729,796,654,681]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="426" unit="m" value="500.0">
<elevation id="00089B12FF96BA5C3DC5738F7F72FD2B" box="[729,796,654,681]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="426" unit="m" value="500.0">500m</elevation>
</quantity>
,
<date id="FF9B5AE1FF96BA5C3C35738E7FDBFD2B" box="[809,949,655,681]" pageId="37" pageNumber="426" value="1992-07-17">
<collectingDate id="EFDFA309FF96BA5C3C35738E7FDBFD2B" box="[809,949,655,681]" pageId="37" pageNumber="426" value="1992-07-17">17.VII.1992</collectingDate>
</date>
,
<collectorName id="26D019F7FF96BA5C3CDD738E784EFD2B" box="[961,1056,655,681]" pageId="37" pageNumber="426">Adesiva</collectorName>
,
<collectorName id="26D019F7FF96BA5C3B30738E7807FD2B" box="[1068,1129,655,681]" pageId="37" pageNumber="426">Vidal</collectorName>
&amp;
<collectorName id="26D019F7FF96BA5C3B8D738E78BEFD2B" box="[1169,1232,655,681]" pageId="37" pageNumber="426">Vidal</collectorName>
, col. (
<collectionCode id="ED34E4E4FF96BA5C3A0B738E7933FD2B" box="[1303,1373,655,681]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="37" pageNumber="426">INPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4D767CFF96BA5C3A6E738E7851FD4F" collectingDate="2001-10" collectionCode="INPA" collectorName="Rod AM &amp; Malaise &amp; J. F. Vidal" country="Brazil" location="Km" municipality="Igarape Acarara" pageId="37" pageNumber="426" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D23B7A8FF96BA5C3A6E738E79F9FD2B" box="[1394,1431,655,681]" count="1" pageId="37" pageNumber="426" type="male">1♂</specimenCount>
,
<collectorName id="26D019F7FF96BA5C3F8B73B27C97FD4F" box="[151,249,691,717]" pageId="37" pageNumber="426">Rod AM</collectorName>
010
<location id="8EFA2AFAFF96BA5C3E2D73B27D35FD4F" LSID="urn:lsid:plazi:treatment:038CCD37FF90BA5C3F8B77C77E42FBB7:8EFA2AFAFF96BA5C3E2D73B27D35FD4F" box="[305,347,691,717]" country="Brazil" municipality="Igarape Acarara" name="Km" pageId="37" pageNumber="426">Km</location>
26,
<collectingMunicipality id="6BFEE65BFF96BA5C3E9173B27E2DFD4F" box="[397,579,691,717]" pageId="37" pageNumber="426">Igarapé Acarará</collectingMunicipality>
,
<date id="FF9B5AE1FF96BA5C3D5173B27ECDFD4F" box="[589,675,691,717]" pageId="37" pageNumber="426" value="2001-10">
<collectingDate id="EFDFA309FF96BA5C3D5173B27ECDFD4F" box="[589,675,691,717]" pageId="37" pageNumber="426" value="2001-10">X.2001</collectingDate>
</date>
, arm.
<collectorName id="26D019F7FF96BA5C3DF573B27F2AFD4F" box="[745,836,691,717]" pageId="37" pageNumber="426">Malaise</collectorName>
,
<collectorName id="26D019F7FF96BA5C3C4C73B27FDCFD4F" box="[848,946,691,717]" pageId="37" pageNumber="426">J.F.Vidal</collectorName>
col. (
<collectionCode id="ED34E4E4FF96BA5C3CEE73B27856FD4F" box="[1010,1080,691,717]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="37" pageNumber="426">INPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4D767CFF96BA5C3B5773B27D87FD73" collectingDate="1986-09-08" collectionCode="INPA" collectorName="Malaise &amp; B. Klein" country="Brazil" location="Reserva" pageId="37" pageNumber="426" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D23B7A8FF96BA5C3B5773B27801FD4F" box="[1099,1135,691,717]" count="1" pageId="37" pageNumber="426" type="male">1♂</specimenCount>
,
<location id="8EFA2AFAFF96BA5C3B6073B278B8FD4F" LSID="urn:lsid:plazi:treatment:038CCD37FF90BA5C3F8B77C77E42FBB7:8EFA2AFAFF96BA5C3B6073B278B8FD4F" box="[1148,1238,691,717]" country="Brazil" name="Reserva" pageId="37" pageNumber="426">Reserva</location>
1208,
<date id="FF9B5AE1FF96BA5C3A3873B279F6FD4F" box="[1316,1432,691,717]" pageId="37" pageNumber="426" value="1986-09-08">
<collectingDate id="EFDFA309FF96BA5C3A3873B279F6FD4F" box="[1316,1432,691,717]" pageId="37" pageNumber="426" value="1986-09-08">8.IX.1986</collectingDate>
</date>
,
<collectorName id="26D019F7FF96BA5C3F8B73D67C9CFD73" box="[151,242,727,753]" pageId="37" pageNumber="426">Malaise</collectorName>
,
<collectorName id="26D019F7FF96BA5C3FE273D67D37FD73" box="[254,345,727,753]" pageId="37" pageNumber="426">B.Klein</collectorName>
, col. (
<collectionCode id="ED34E4E4FF96BA5C3E8073D67D8CFD73" box="[412,482,727,753]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="37" pageNumber="426">INPA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F2FAAFF96BA5C3FDB73FB7E2DFC97" box="[199,579,762,789]" pageId="37" pageNumber="426" type="distribution">
<paragraph id="8B9A7C21FF96BA5C3FDB73FB7E2DFC97" blockId="37.[151,1437,150,1077]" box="[199,579,762,789]" pageId="37" pageNumber="426">
<emphasis id="B951A033FF96BA5C3FDB73FB7D37FC96" bold="true" box="[199,345,762,788]" pageId="37" pageNumber="426">Distribution</emphasis>
.
<collectingCountry id="F3323CB1FF96BA5C3E7B73FA7DC5FC97" box="[359,427,763,789]" name="Brazil" pageId="37" pageNumber="426">Brazil</collectingCountry>
(Amazonas).
</paragraph>
</subSubSection>
<subSubSection id="C33F2FAAFF96BA5C3FDB721E7D92FCDF" pageId="37" pageNumber="426" type="etymology">
<paragraph id="8B9A7C21FF96BA5C3FDB721E7D92FCDF" blockId="37.[151,1437,150,1077]" pageId="37" pageNumber="426">
<emphasis id="B951A033FF96BA5C3FDB721E7D26FCBB" bold="true" box="[199,328,799,825]" pageId="37" pageNumber="426">Etymology</emphasis>
. This species is named after the astrophysicist Rita de Cassia dos Anjos, a black woman who overcame inequalities in
<collectingCountry id="F3323CB1FF96BA5C3EAC72427D96FCDF" box="[432,504,835,861]" name="Brazil" pageId="37" pageNumber="426">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F2FAAFF96BA5C3FDB72667E42FBB7" pageId="37" pageNumber="426" type="discussion">
<paragraph id="8B9A7C21FF96BA5C3FDB72667E42FBB7" blockId="37.[151,1437,150,1077]" pageId="37" pageNumber="426">
<emphasis id="B951A033FF96BA5C3FDB72667D5AFC03" bold="true" box="[199,308,871,897]" pageId="37" pageNumber="426">Remarks</emphasis>
. It is similar to
<taxonomicName id="4C2507A2FF96BA5C3EFA72667E32FC02" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[486,604,871,896]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="426" phylum="Arthropoda" rank="species" species="maryae" status="sp. nov.">
<emphasis id="B951A033FF96BA5C3EFA72667E32FC02" box="[486,604,871,896]" italics="true" pageId="37" pageNumber="426">D. maryae</emphasis>
</taxonomicName>
<emphasis id="B951A033FF96BA5C3D7F72667ED3FC03" bold="true" box="[611,701,871,897]" pageId="37" pageNumber="426">
<taxonomicNameLabel id="A2621D48FF96BA5C3D7F72667ED3FC03" box="[611,701,871,897]" pageId="37" pageNumber="426" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having the aedeagal dorsal valve rounded, the aedeagal dorsal lobes long, the apex of inner median filament anterior to apex aedeagal dorsal lobes, and the apex of aedeagal dorsal valve narrow. However,
<taxonomicName id="4C2507A2FF96BA5C3EB772AE7E6CFC4A" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[427,514,943,968]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="426" phylum="Arthropoda" rank="species" species="ritae" status="sp. nov.">
<emphasis id="B951A033FF96BA5C3EB772AE7E6CFC4A" box="[427,514,943,968]" italics="true" pageId="37" pageNumber="426">D. ritae</emphasis>
</taxonomicName>
<emphasis id="B951A033FF96BA5C3D1672AE7E0DFC4B" bold="true" box="[522,611,943,969]" pageId="37" pageNumber="426">
<taxonomicNameLabel id="A2621D48FF96BA5C3D1672AE7E0DFC4B" box="[522,611,943,969]" pageId="37" pageNumber="426" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has the dorsal margin of harpe straight and entirely wide, the basal cover plate wide, the apex of outer lobe with a constriction and the basivolsellar protuberance small and well demarcated, whereas
<taxonomicName id="4C2507A2FF96BA5C3FE672F67D1EFB92" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[250,368,1015,1040]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="426" phylum="Arthropoda" rank="species" species="maryae" status="sp. nov.">
<emphasis id="B951A033FF96BA5C3FE672F67D1EFB92" box="[250,368,1015,1040]" italics="true" pageId="37" pageNumber="426">D. maryae</emphasis>
</taxonomicName>
<emphasis id="B951A033FF96BA5C3E6B72F67DBEFB93" bold="true" box="[375,464,1015,1041]" pageId="37" pageNumber="426">
<taxonomicNameLabel id="A2621D48FF96BA5C3E6B72F67DBEFB93" box="[375,464,1015,1041]" pageId="37" pageNumber="426" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has the harpe with dorsal margin incurved anteriorly, the basal cover plate narrow, and the basivolsellar protuberance wide.
</paragraph>
</subSubSection>
</treatment>
</document>
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