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<document id="73F102168DBB9970401430ED58C86248" ID-CLB-Dataset="26574" ID-DOI="10.11646/zootaxa.4613.3.8" ID-GBIF-Dataset="67477ca5-246c-46b9-9387-63463c0b5970" ID-ISSN="1175-5326" ID-Zenodo-Dep="3239923" ID-ZooBank="F645124D-22BC-4D7E-AB19-94F5F59CA3BA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1559796749597" checkinUser="plazi" docAuthor="Viraktamath, C. A. &amp; Webb, M. D." docDate="2019" docId="038C87DFE07C0A37FF5DFB79FCD0FF4E" docLanguage="en" docName="zootaxa.4613.3.8.pdf" docOrigin="Zootaxa 4613 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Daimachus robustus Viraktamath &amp; Webb 2019, sp. nov." docType="treatment" docVersion="11" lastPageNumber="561" masterDocId="FFB5FFA7E0780A32FFCAFF93FFB6FFB5" masterDocTitle="Revision of the Ulopinae leafhoppers (Hemiptera: Cicadellidae) of the Indian subcontinent, I. Ulopini genera: Daimachus, Radhades and Ulopsina" masterLastPageNumber="577" masterPageNumber="557" pageNumber="561" updateTime="1698733371643" updateUser="ExternalLinkService">
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<mods:title id="5ABF8176EB8A47748AD35D3A6F4D515E">Revision of the Ulopinae leafhoppers (Hemiptera: Cicadellidae) of the Indian subcontinent, I. Ulopini genera: Daimachus, Radhades and Ulopsina</mods:title>
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<mods:name id="51A7DBF82749A373F02A2B1F8DA748A2" type="personal">
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<mods:namePart id="0B6BCA3547D7A2A221EE23DF834DB37B">Viraktamath, C. A.</mods:namePart>
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<mods:namePart id="1E70C610997A38666BDEF841D145AE4E">Webb, M. D.</mods:namePart>
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<mods:title id="3FBE010FF66F6198B05FA22548B2A7B0">Zootaxa</mods:title>
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<mods:part id="731E533B070F71E4A9E6966F22D08BD0">
<mods:date id="E35F3B0E42BAE16FA4B7A15D88D599D2">2019</mods:date>
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<mods:number id="E350280DAA95BCB1A3133048AAC06F4E">2019-06-06</mods:number>
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<treatment id="038C87DFE07C0A37FF5DFB79FCD0FF4E" ID-DOI="http://doi.org/10.5281/zenodo.5941764" ID-GBIF-Taxon="157422518" ID-Zenodo-Dep="5941764" LSID="urn:lsid:plazi:treatment:038C87DFE07C0A37FF5DFB79FCD0FF4E" httpUri="http://treatment.plazi.org/id/038C87DFE07C0A37FF5DFB79FCD0FF4E" lastPageId="5" lastPageNumber="561" pageId="4" pageNumber="561">
<subSubSection id="C33F6542E07C0A36FF5DFB79FE40FAB0" box="[151,502,1258,1285]" pageId="4" pageNumber="561" type="nomenclature">
<paragraph id="8B9A36C9E07C0A36FF5DFB79FE40FAB0" blockId="4.[151,519,1258,1321]" box="[151,502,1258,1285]" pageId="4" pageNumber="561">
<heading id="D0D281A5E07C0A36FF5DFB79FE40FAB0" bold="true" box="[151,502,1258,1285]" fontSize="11" level="1" pageId="4" pageNumber="561" reason="1">
<emphasis id="B951EADBE07C0A36FF5DFB79FE40FAB0" bold="true" box="[151,502,1258,1285]" pageId="4" pageNumber="561">
<taxonomicName id="4C254D4AE07C0A36FF5DFB79FE27FAB0" ID-CoL="8JRL5" authority="Viraktamath &amp; Webb, 2019" authorityName="Viraktamath &amp; Webb" authorityYear="2019" box="[151,401,1258,1285]" class="Insecta" family="Cicadellidae" genus="Daimachus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="561" phylum="Arthropoda" rank="species" species="robustus" status="sp. nov.">
<emphasis id="B951EADBE07C0A36FF5DFB79FE27FAB0" bold="true" box="[151,401,1258,1285]" italics="true" pageId="4" pageNumber="561">Daimachus robustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257A0E07C0A36FE53FB78FE40FAB0" box="[409,502,1259,1285]" pageId="4" pageNumber="561" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F6542E07C0A36FF5DFA9DFF7CF8DC" pageId="4" pageNumber="561" type="description">
<paragraph id="8B9A36C9E07C0A36FF5DFA9DFDB1FA9D" blockId="4.[151,519,1258,1321]" box="[151,519,1293,1321]" pageId="4" pageNumber="561">
<figureCitation id="131E2A4CE07C0A36FF5DFA9DFEB7FA9C" box="[151,257,1294,1321]" captionStart="FIGURES 1" captionStartId="10.[151,264,1914,1939]" captionTargetBox="[207,1373,192,1871]" captionTargetId="figure@10.[195,1392,177,1880]" captionTargetPageId="10" captionText="FIGURES 1A–L. Species of Daimachus, habitus.A–F, colour variation in D. exemplificatus Distant:ABE, males, CD, females; GH, D. matheranensis sp. nov., holotype female; IJ. D. robustus sp. nov., holotype male; KL, D. sirsiensis sp. nov., holotype male. ACE-GIK, dorsal view; BDHJL, lateral view." httpUri="https://zenodo.org/record/3239925/files/figure.png" pageId="4" pageNumber="561">
Figs
<date id="FF9B1009E07C0A36FF04FA9DFF51FA9D" box="[206,231,1294,1320]" pageId="4" pageNumber="561">1I</date>
, J
</figureCitation>
,
<figureCitation id="131E2A4CE07C0A36FEC7FA9DFE87FA9D" box="[269,305,1294,1320]" captionStart="FIGURES 3" captionStartId="12.[151,264,1872,1897]" captionTargetBox="[166,1426,191,1840]" captionTargetId="figure@12.[151,1436,181,1846]" captionTargetPageId="12" captionText="FIGURES 3A–M. Species of Ulopini: A–C, Radhades crassus Distant, holotype male, dorsal and lateral views and label data; D–H, Ulopsina (Indoulopa) himalayana sp. nov.: D–E, holotype male dorsal, lateral and ventral view; GH, forewing and hind wing of paratype male; I–M, species of Daimachus, faces: I. D. exemplificatus Distant; J. D. matheranensis sp. nov.; K. D. robustus sp. nov.; L. D. sirsiensis sp. nov.; M. D. sudindicus sp. nov." httpUri="https://zenodo.org/record/3239931/files/figure.png" pageId="4" pageNumber="561">3K</figureCitation>
,
<figureCitation id="131E2A4CE07C0A36FEF7FA9DFED6FA9D" box="[317,352,1294,1320]" captionStart="FIGURES 4" captionStartId="13.[151,275,1787,1814]" captionTargetBox="[166,1426,191,1748]" captionTargetId="figure@13.[151,1436,181,1763]" captionText="FIGURES 4A–H. Female pregenital sternites and valvulae of species of Daimachus: A–D, sternites VI and VII of (A) D. exemplificatus Distant, (B) D. matheranensis sp. nov., (C) D. robustus sp. nov. and (D) D. sudindicus sp. nov.; E–H, D. exemplificatus (EF) valvulae I and (GH) valvulae II." httpUri="https://zenodo.org/record/3406227/files/figure.png" pageId="4" pageNumber="561">4C</figureCitation>
,
<figureCitation id="131E2A4CE07C0A36FEA6FA9EFECFFA9D" box="[364,377,1293,1320]" captionStart="FIGURES 5" captionStartId="14.[151,275,1889,1916]" captionTargetBox="[156,1426,184,1853]" captionTargetId="figure@14.[151,1436,179,1864]" captionText="FIGURES 5A–L. Valvulae I and II of species of Daimachus: A–D, D. matheranensis sp. nov.; E–H, D. robustus sp. nov. I–L, D. sudindicus sp. nov." httpUri="https://zenodo.org/record/3406229/files/figure.png" pageId="4" pageNumber="561">5</figureCitation>
E–H, 7A–G.
</paragraph>
<paragraph id="8B9A36C9E07C0A36FF5DFAC5FED9FA21" blockId="4.[151,1437,1366,2041]" pageId="4" pageNumber="561">
Colouration and structure as in generic description but forewings shorter and broader, about
<date id="FF9B1009E07C0A36FB7DFAC5FB64FAC5" box="[1207,1234,1366,1392]" pageId="4" pageNumber="561">2x</date>
as long as broad. Hind wings absent.
</paragraph>
<paragraph id="8B9A36C9E07C0A36FF0DFA0DFEB3F900" blockId="4.[151,1437,1366,2041]" pageId="4" pageNumber="561">
<emphasis id="B951EADBE07C0A36FF0DFA0DFEDBFA0D" box="[199,365,1438,1464]" italics="true" pageId="4" pageNumber="561">Male genitalia</emphasis>
. Pygofer in lateral view more or less rectangular, both posterodorsal and posteroventral angles rounded, posterior margin more or less straight. Subgenital plate held vertically, broadened in distal region, exceeding posterior margin of pygofer. Style with apophysis dorsally curved subapically with apex broad, posterior angle of apex slightly longer than anterior angle. Connective with stem longer than arms, apex broad, anteromedial lobe vestigial. Aedeagus with shaft tapered in apical region in lateral view, without triangular projections on dorsal or ventral surface, apex of shaft bifid. Segment X and XI concealed within pygofer, ventral aspect of segment produced both anteriorly and posteriorly, but more so anteriorly forming tubular cover for apical part of aedeagal shaft in repose.
</paragraph>
<paragraph id="8B9A36C9E07C0A36FF0DF92DFC3CF949" blockId="4.[151,1437,1366,2041]" pageId="4" pageNumber="561">
<emphasis id="B951EADBE07C0A36FF0DF92DFE3EF96D" box="[199,392,1726,1752]" italics="true" pageId="4" pageNumber="561">Female genitalia</emphasis>
. Sternite VII posterior margin less broadly excavated in the form of V (
<figureCitation id="131E2A4CE07C0A36FB77F92DFAA1F96D" box="[1213,1303,1726,1752]" captionStart="FIGURES 4" captionStartId="13.[151,275,1787,1814]" captionTargetBox="[166,1426,191,1748]" captionTargetId="figure@13.[151,1436,181,1763]" captionText="FIGURES 4A–H. Female pregenital sternites and valvulae of species of Daimachus: A–D, sternites VI and VII of (A) D. exemplificatus Distant, (B) D. matheranensis sp. nov., (C) D. robustus sp. nov. and (D) D. sudindicus sp. nov.; E–H, D. exemplificatus (EF) valvulae I and (GH) valvulae II." httpUri="https://zenodo.org/record/3406227/files/figure.png" pageId="4" pageNumber="561">Fig. 4C</figureCitation>
). Valvula II with short, less prominent teeth with secondary dentition (
<figureCitation id="131E2A4CE07C0A36FCE9F971FCCBF949" box="[803,893,1761,1788]" captionStart="FIGURES 5" captionStartId="14.[151,275,1889,1916]" captionTargetBox="[156,1426,184,1853]" captionTargetId="figure@14.[151,1436,179,1864]" captionText="FIGURES 5A–L. Valvulae I and II of species of Daimachus: A–D, D. matheranensis sp. nov.; E–H, D. robustus sp. nov. I–L, D. sudindicus sp. nov." httpUri="https://zenodo.org/record/3406229/files/figure.png" pageId="4" pageNumber="561">Fig. 5H</figureCitation>
).
</paragraph>
<paragraph id="8B9A36C9E07C0A36FF0DF894FF7CF8DC" blockId="4.[151,1437,1366,2041]" pageId="4" pageNumber="561">
<emphasis id="B951EADBE07C0A36FF0DF894FEDFF895" box="[199,361,1799,1824]" italics="true" pageId="4" pageNumber="561">Measurements</emphasis>
. Male 3.0 mm long,
<quantity id="4CDD9B2CE07C0A36FD92F895FD19F894" box="[600,687,1798,1825]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="561" unit="mm" value="1.1">1.1 mm</quantity>
wide across eyes and 1.0 mm wide across posterolateral angles of pronotum. Female
<quantity id="4CDD9B2CE07C0A36FEA6F8B9FE75F8F0" box="[364,451,1834,1861]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="4" pageNumber="561" unit="mm" value="2.9">2.9 mm</quantity>
long,
<quantity id="4CDD9B2CE07C0A36FDC0F8B9FDD6F8F0" box="[522,608,1834,1861]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="561" unit="mm" value="1.1">1.1 mm</quantity>
wide across eyes and 1.0 mm wide across posterolateral angles of pronotum.
</paragraph>
</subSubSection>
<subSubSection id="C33F6542E07C0A36FF0DF8E1FC58F805" pageId="4" pageNumber="561" type="materials_examined">
<paragraph id="8B9A36C9E07C0A36FF0DF8E1FC58F805" blockId="4.[151,1437,1366,2041]" pageId="4" pageNumber="561">
<emphasis id="B951EADBE07C0A36FF0DF8E1FE2FF839" box="[199,409,1906,1932]" italics="true" pageId="4" pageNumber="561">Material examined</emphasis>
.
<materialsCitation id="3B4D3C94E07C0A36FE6FF8E1FE56F805" ID-GBIF-Occurrence="2248132640" collectingDate="2011-09-29" collectionCode="UASB" country="India" elevation="15" latitude="12.256383" location="Periya" longLatPrecision="9" longitude="75.1225" municipality="Kasargod" pageId="4" pageNumber="561" specimenCount="1" specimenCount-male="1" stateProvince="Kerala" typeStatus="holotype">
<typeStatus id="549E886BE07C0A36FE6FF8E1FDBBF839" box="[421,525,1906,1932]" pageId="4" pageNumber="561" type="holotype">Holotype</typeStatus>
<specimenCount id="9D23FD40E07C0A36FDD8F8E0FD9FF838" box="[530,553,1907,1933]" pageId="4" pageNumber="561" type="male">♂</specimenCount>
,
<collectingCountry id="F3327659E07C0A36FDF9F8E1FD31F839" box="[563,647,1906,1932]" name="India" pageId="4" pageNumber="561">INDIA</collectingCountry>
:
<collectingRegion id="49E1F82BE07C0A36FD44F8E1FD6BF839" box="[654,733,1906,1932]" country="India" name="Kerala" pageId="4" pageNumber="561">Kerala</collectingRegion>
:
<collectingMunicipality id="6BFEACB3E07C0A36FD2FF8E1FCE5F839" box="[741,851,1906,1932]" pageId="4" pageNumber="561">Kasargod</collectingMunicipality>
,
<location id="8EFA6012E07C0A36FC96F8E1FC10F839" LSID="urn:lsid:plazi:treatment:038C87DFE07C0A37FF5DFB79FCD0FF4E:8EFA6012E07C0A36FC96F8E1FC10F839" box="[860,934,1906,1932]" country="India" latitude="12.256383" longLatPrecision="9" longitude="75.1225" municipality="Kasargod" name="Periya" pageId="4" pageNumber="561" stateProvince="Kerala">Periya</location>
,
<elevation id="0008D1FAE07C0A36FC65F8E2FC55F839" box="[943,995,1905,1932]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="561" unit="m" value="15.0">
<quantity id="4CDD9B2CE07C0A36FC65F8E2FC55F839" box="[943,995,1905,1932]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="561" unit="m" value="15.0">15m</quantity>
</elevation>
,
<geoCoordinate id="EE11500EE07C0A36FC27F8E1FBC8F839" box="[1005,1150,1905,1932]" direction="north" orientation="latitude" pageId="4" pageNumber="561" precision="1" value="12.256383">
12
<superScript id="7C509B81E07C0A36FBC3F8E2FBA7F8CA" attach="none" box="[1033,1041,1905,1919]" fontSize="6" pageId="4" pageNumber="561">o</superScript>
15.383’N
</geoCoordinate>
, 0
<geoCoordinate id="EE11500EE07C0A36FB5FF8E2FAA0F839" box="[1173,1302,1905,1932]" direction="east" orientation="longitude" pageId="4" pageNumber="561" precision="9" value="75.1225">
75
<superScript id="7C509B81E07C0A36FB7BF8E2FB0FF8CA" attach="none" box="[1201,1209,1905,1919]" fontSize="6" pageId="4" pageNumber="561">o</superScript>
07.35’E
</geoCoordinate>
,
<collectingDate id="EFDFE9E1E07C0A36FAD4F8E1FA21F839" box="[1310,1431,1906,1932]" pageId="4" pageNumber="561" value="2011-09-29">
<date id="FF9B1009E07C0A36FAD4F8E1FA21F839" box="[1310,1431,1906,1932]" pageId="4" pageNumber="561" value="2011-09-29">29.ix.2011</date>
</collectingDate>
, Srikanth,
<collectingMethod id="52644EDEE07C0A36FECEF805FECEF804" box="[260,376,1942,1969]" pageId="4" pageNumber="561">Sweep net</collectingMethod>
(
<collectionCode id="ED34AE0CE07C0A36FE4DF805FE6EF805" box="[391,472,1942,1968]" country="India" httpUri="http://grbio.org/cool/yr7i-jnjm" name="University of Agricultural Sciences" pageId="4" pageNumber="561">UASB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4D3C94E07C0A36FE20F804FC5FF805" ID-GBIF-Occurrence="2248132644" box="[490,1001,1942,1969]" collectingDate="2011-09-29" collectionCode="UASB" country="India" elevation="15" latitude="12.256383" location="Periya" longLatPrecision="9" longitude="75.1225" municipality="Kasargod" pageId="4" pageNumber="561" specimenCount="1" specimenCount-female="1" stateProvince="Kerala" typeStatus="paratype">
<typeStatus id="549E886BE07C0A36FE20F804FDFAF804" box="[490,588,1943,1969]" pageId="4" pageNumber="561" type="paratype">Paratype</typeStatus>
<specimenCount id="9D23FD40E07C0A36FD99F805FDCBF804" box="[595,637,1942,1969]" pageId="4" pageNumber="561" type="female">1 ♀</specimenCount>
, data same as holotype (
<collectionCode id="ED34AE0CE07C0A36FC5BF805FC54F805" box="[913,994,1942,1968]" country="India" httpUri="http://grbio.org/cool/yr7i-jnjm" name="University of Agricultural Sciences" pageId="4" pageNumber="561">UASB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F6542E07C0A36FF0DF829FC41F861" box="[199,1015,1978,2005]" pageId="4" pageNumber="561" type="etymology">
<paragraph id="8B9A36C9E07C0A36FF0DF829FC41F861" blockId="4.[151,1437,1366,2041]" box="[199,1015,1978,2005]" pageId="4" pageNumber="561">
<emphasis id="B951EADBE07C0A36FF0DF829FE89F861" box="[199,319,1978,2004]" italics="true" pageId="4" pageNumber="561">Etymology</emphasis>
. The species epithet refers to the robust nature of this species.
</paragraph>
</subSubSection>
<subSubSection id="C33F6542E07C0A37FF0DF84DFCD0FF4E" lastPageId="5" lastPageNumber="562" pageId="4" pageNumber="561" type="discussion">
<paragraph id="8B9A36C9E07C0A37FF0DF84DFCD0FF4E" blockId="4.[151,1437,1366,2041]" lastBlockId="5.[151,1437,153,251]" lastPageId="5" lastPageNumber="562" pageId="4" pageNumber="561">
<emphasis id="B951EADBE07C0A36FF0DF84DFE82F84D" bold="true" box="[199,308,2014,2040]" pageId="4" pageNumber="561">Remarks</emphasis>
. This is the shortest and most robust species among
<taxonomicName id="4C254D4AE07C0A36FC4EF84DFBB2F84D" authorityName="Distant" authorityYear="1916" box="[900,1028,2014,2040]" class="Insecta" family="Cicadellidae" genus="Daimachus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="561" phylum="Arthropoda" rank="genus">
<emphasis id="B951EADBE07C0A36FC4EF84DFBB2F84D" box="[900,1028,2014,2040]" italics="true" pageId="4" pageNumber="561">Daimachus</emphasis>
</taxonomicName>
species so far known. It has a much longer and strongly anteriorly recurved aedeagal shaft. It resembles
<taxonomicName id="4C254D4AE07D0A37FC51FF09FB90FF06" authority="Viraktamath &amp; Webb, 2019" authorityName="Viraktamath &amp; Webb" authorityYear="2019" box="[923,1062,154,179]" class="Insecta" family="Cicadellidae" genus="Daimachus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="562" phylum="Arthropoda" rank="species" species="sirsiensis" status="sp. nov.">
<emphasis id="B951EADBE07D0A37FC51FF09FB90FF06" box="[923,1062,154,179]" italics="true" pageId="5" pageNumber="562">D. sirsiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257A0E07D0A37FBE5FF0AFB31FF06" box="[1071,1159,153,179]" pageId="5" pageNumber="562" rank="species">sp. nov.</taxonomicNameLabel>
but has short prong-like lateral projections on the apex of the aedeagal shaft compared to long thin processes in the latter species and also lacks the subapical ventral projection found in the latter species.
</paragraph>
</subSubSection>
</treatment>
</document>
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