449 lines
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449 lines
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<bibRefCitation id="EFB44B20CE05FFBFFEBDFBF0FE52FBCA" author="Janson, O. E." box="[289,480,1048,1073]" pageId="2" pageNumber="93" pagination="603 - 611" refId="ref20457" refString="Janson, O. E. (1881) Notices of new or little known Cetoniidae No. 7. Cistula Entomologica, 2 (24), 603 - 611." type="journal article" year="1881">Janson, 1881: 609</bibRefCitation>
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<treatmentCitation id="0A8410C0CE05FFBFFE70FBF2FD00FBCA" author="Schoch, G." box="[492,690,1048,1073]" page="361" pageId="2" pageNumber="93" year="1896">
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<bibRefCitation id="EFB44B20CE05FFBFFE70FBF2FD00FBCA" author="Schoch, G." box="[492,690,1048,1073]" pageId="2" pageNumber="93" pagination="356 - 370" refId="ref21739" refString="Schoch, G. (1896 a) Einiges uber Cetoniden. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 9 (8), 356 - 370." type="journal article" year="1896">Schoch 1896a: 361</bibRefCitation>
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(in key);
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<treatmentCitation id="0A8410C0CE05FFBFFC87FBF2FC54FBCA" author="Schoch, G." box="[795,998,1048,1073]" page="394" pageId="2" pageNumber="93" year="1896">
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<bibRefCitation id="EFB44B20CE05FFBFFC87FBF2FC54FBCA" author="Schoch, G." box="[795,998,1048,1073]" pageId="2" pageNumber="93" pagination="378 - 395" refId="ref21766" refString="Schoch, G. (1896 b) Nachtrag II zu Schoch: Die Genera und Species meiner Cetoniden-sammlung. Mitteilungen der Schweizeri- schen Entomologischen Gesellschaft, 9 (9), 378 - 395." type="journal article" year="1896">Schoch 1896b: 394</bibRefCitation>
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;
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;
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<treatmentCitation id="0A8410C0CE05FFBFFB2FFBF2FA2AFBCA" author="Schenkling, S." box="[1203,1432,1048,1073]" page="364" pageId="2" pageNumber="93" year="1921">
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<bibRefCitation id="EFB44B20CE05FFBFFB2FFBF2FA2AFBCA" author="Schenkling, S." box="[1203,1432,1048,1073]" pageId="2" pageNumber="93" pagination="1 - 431" refId="ref21671" refString="Schenkling, S. (1921) Scarabaeidae: Cetoniinae. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Vol. XXI. Scarabaeidae III. Pars 72. W. Junk, Berlin, pp. 1 - 431 pp. https: // doi. org / 10.1007 / 978 - 94 - 011 - 9697 - 0 _ 1" type="book chapter" year="1921">Schenkling 1921: 364</bibRefCitation>
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<treatmentCitation id="0A8410C0CE05FFBFFF5BFBD2FEC3FBAB" author="Mao, Y. T." box="[199,369,1079,1104]" page="1096" pageId="2" pageNumber="93" year="1937">
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<bibRefCitation id="EFB44B20CE05FFBFFF5BFBD2FEC3FBAB" author="Mao, Y. T." box="[199,369,1079,1104]" pageId="2" pageNumber="93" pagination="961 - 1102" refId="ref20892" refString="Mao, Y. T. (1937) Family Scarabaeidae. In: Wu, C. F. (Ed.), Catalogus Insectorum Sinensium (Catalogue of Chinese Insects). Vol. III. The Fan Memorial Institute of Biology, Peiping, pp. 961 - 1102." type="book chapter" year="1937">Mao 1937: 1096</bibRefCitation>
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<treatmentCitation id="0A8410C0CE05FFBFFEE6FBDDFDA9FBAB" author="Paulian, R." box="[378,539,1079,1104]" page="6" pageId="2" pageNumber="93" year="1961">
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<bibRefCitation id="EFB44B20CE05FFBFFEE6FBDDFDA9FBAB" author="Paulian, R." box="[378,539,1079,1104]" pageId="2" pageNumber="93" pagination="1 - 47" refId="ref21277" refString="Paulian, R. (1961) Coleopteres Scarabeides de L'Indochine II (Rutelines et Cetonines) (suite et fin). Annales de la Societe En- tomologique de France, 130, 1 - 47." type="journal article" year="1961">Paulian 1961: 6</bibRefCitation>
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;
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<treatmentCitation id="0A8410C0CE05FFBFFDB9FBDDFD4CFBAB" author="Medvedev, S. I." box="[549,766,1079,1104]" page="333" pageId="2" pageNumber="93" year="1964">
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<bibRefCitation id="EFB44B20CE05FFBFFDB9FBDDFD4CFBAB" author="Medvedev, S. I." box="[549,766,1079,1104]" pageId="2" pageNumber="93" refId="ref20989" refString="Medvedev, S. I. (1964) Plastinchatousye (Scarabaeidae), podsem. Cetoniinae, Valginae, Fauna SSSR, zhestkokrylye. Tom 10. Vyp. 5. Izdatel'stvo Akademii Nauk SSSR, Moscow, 375 pp. [in Russian]" type="book" year="1964">Medvedev 1964: 333</bibRefCitation>
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<treatmentCitation id="0A8410C0CE05FFBFFC94FBDDFC0EFBAB" author="Krikken, J." box="[776,956,1079,1104]" page="55" pageId="2" pageNumber="93" year="1984">
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<bibRefCitation id="EFB44B20CE05FFBFFC94FBDDFC0EFBAB" author="Krikken, J." box="[776,956,1079,1104]" pageId="2" pageNumber="93" pagination="1 - 75" refId="ref20712" refString="Krikken, J. (1984) A new key to the suprageneric taxa in the beetle family Cetoniidae, with annotated lists of the known genera. Zoologische Verhandelingen, 210, 1 - 75." type="journal article" year="1984">Krikken 1984: 55</bibRefCitation>
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;
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<treatmentCitation id="0A8410C0CE05FFBFFC59FBDDFBDBFBAB" author="Antoine, P." box="[965,1129,1079,1104]" page="9" pageId="2" pageNumber="93" year="1991">
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(key);
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<treatmentCitation id="0A8410C0CE05FFBFFB2DFBD2FAF3FBAB" author="Ma, W. Z." box="[1201,1345,1079,1104]" page="153" pageId="2" pageNumber="93" year="1995">
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<bibRefCitation id="EFB44B20CE05FFBFFB2DFBD2FAF3FBAB" author="Ma, W. Z." box="[1201,1345,1079,1104]" pageId="2" pageNumber="93" refId="ref20850" refString="Ma, W. Z. (1995) Economic Insect Fauna of China. Vol. 46. Coleoptera: Cetoniidae, Trichiidae, and Valgidae. Science and Technology Press, Beijing, 210 pp. [in Chinese]" type="book" year="1995">Ma 1995: 153</bibRefCitation>
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<treatmentCitation id="0A8410C0CE05FFBFFAD7FBDDFEC1FB8F" author="Sakai, K & Nagai, S." page="155" pageId="2" pageNumber="93" year="1998">
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<bibRefCitation id="EFB44B20CE05FFBFFAD7FBDDFEC1FB8F" author="Sakai, K & Nagai, S." pageId="2" pageNumber="93" refId="ref21636" refString="Sakai, K & Nagai, S. (1998) The Cetoniine Beetles of the World. Mushi-Sha's Iconographic Series of Insects. Vol. 3. Mushi-Sha, Tokyo, 421 pp." type="book" year="1998">Sakai & Nagai 1998: 155</bibRefCitation>
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<treatmentCitation id="0A8410C0CE05FFBFFEE1FBB1FD9EFB8F" author="Krajcik, M." box="[381,556,1115,1140]" page="39" pageId="2" pageNumber="93" year="1999">
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<bibRefCitation id="EFB44B20CE05FFBFFC9BFBB1FC07FB8F" author="Krajcik, M." box="[775,949,1115,1140]" pageId="2" pageNumber="93" pagination="1 - 113" refId="ref20569" refString="Krajcik, M. (2011) Illustrated Catalogue of Cetoniinae, Trichiinae and Valginae of China (Coleoptera, Cetoniidae). Animma. x, Supplement 1, 1 - 113." type="journal article" year="2011">Krajčík 2011: 69</bibRefCitation>
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<treatmentCitation id="0A8410C0CE05FFBFFC5CFBB1FBC8FB8F" author="Krajcik, M." box="[960,1146,1115,1140]" page="116" pageId="2" pageNumber="93" year="2012">
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<bibRefCitation id="EFB44B20CE05FFBFFC5CFBB1FBC8FB8F" author="Krajcik, M." box="[960,1146,1115,1140]" pageId="2" pageNumber="93" pagination="1 - 278" refId="ref20603" refString="Krajcik, M. (2012) Checklist of the World Scarabaeoidea. Animma. x, Supplement 5, 1 - 278." type="journal article" year="2012">Krajčík 2012: 116</bibRefCitation>
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species:
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(
|
||
<bibRefCitation id="EFB44B20CE05FFBFFD82FB90FD7EFB68" author="Westwood, J. O." box="[542,716,1146,1171]" pageId="2" pageNumber="93" refId="ref21916" refString="Westwood, J. O. (1873) Thesaurus Entomologicus Oxoniensis; or, Illustrations of new, rare, and interesting insects, for the most part contained in the collections presented to the University of Oxford by the Rev. F. W. Hope. Part I. Clarendon Press, Oxford, 56 pp., 10 pls. https: // doi. org / 10.5962 / bhl. title. 14077" type="book" year="1873">Westwood, 1873</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, subsequent designation by
|
||
<bibRefCitation id="EFB44B20CE05FFBFFC72FB91FBC8FB68" author="Arrow, G. J." box="[1006,1146,1147,1171]" pageId="2" pageNumber="93" refId="ref19451" refString="Arrow, G. J. (1910) The Fauna of British India (Including Ceylon and Burma) Coleoptera, Lamellicornia (Cetoniinae and Dy- nastinae). Taylor & Francis, London, 322 pp." type="book" year="1910">Arrow (1910)</bibRefCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33F655ACE05FFBFFF0BFB2AFD36F8E0" pageId="2" pageNumber="93" type="diagnosis">
|
||
<paragraph id="8B9A36D1CE05FFBFFF0BFB2AFEB6F954" blockId="2.[151,1437,1216,2035]" pageId="2" pageNumber="93">
|
||
<emphasis id="B951EAC3CE05FFBFFF0BFB2AFEA0FB20" bold="true" box="[151,274,1216,1243]" pageId="2" pageNumber="93">Diagnosis.</emphasis>
|
||
Body small sized, 11.0–13.5 mm. Body usually black, or brown; surfaces clad with sparse U-shaped, or drop shaped setiferous punctures; setae long, acicular, yellow; dorsal surface usually covered with small and large, tomentose, yellow spots; ventral surface covered with light yellow, gray, or khaki tomentum. Antennal scapus simply claviform. Anterior margin of clypeus arc, slightly raised. Frons tomentose. Prementum slightly expanded, more or less thickened or folded in front (
|
||
<figureCitation id="131E2A54CE05FFBFFDF8FABBFD1BFA90" box="[612,681,1361,1387]" captionStart="FIGURES 1–12" captionStartId="3.[151,264,1705,1730]" captionTargetBox="[151,1435,278,1670]" captionTargetId="figure@3.[151,1435,273,1680]" captionTargetPageId="3" captionText="FIGURES 1–12. Characters of Goliathopsis Janson, 1881 species. 1–4, mouthparts of Goliathopsis duponti Antoine, 1991: 1, epipharynx; 2, mandibles; 3, maxillae; 4, labium. 5–12, setiferous punctures on the disc of elytron: 5, G. cervus Janson, 1881; 6, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in BMNH; 7, G. duponti Antoine, 1991; 8, G. esquiroli Pouillaude, 1913; 9, G. ferreroi Antoine, 1991; 10, G. gressitti Ruter, 1978; 11, G. lameyi Fairmaire, 1893; 12, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884727" httpUri="https://zenodo.org/record/3884727/files/figure.png" pageId="2" pageNumber="93">Fig. 4</figureCitation>
|
||
). Pronotum elliptical; widest near the middle; disc with a tomentose longitudinal line in median, distinct, continuous, or discontinuous, even absent in some species; setiferous punctures rounded. Elytron usually with yellow maculae on posthumeral area, mediodiscal areas near sutural costa, median portion of lateral declivity, and postdiscal areas near sutural costa; few other small maculae scattered on the rest of disc. Disc of elytron flattened; apicosutural angle not pointed. Sides of abdominal sternites exposed in dorsal view; terminal spiracle convex. Propygidial surface with tomentum in some species. Pygidium conical; with an indistinct longitudinal ridge in median portion, ridge without tomentum in some species; dorsal side densely punctate, with tomentum or not; apical portion and ventral side with dense, setiferous punctures, without tomentum (
|
||
<figureCitation id="131E2A54CE05FFBFFA9BF9A7FA23F99C" box="[1287,1425,1613,1639]" captionStart="FIGURES 22–37" captionStartId="6.[151,264,1543,1568]" captionTargetBox="[156,1430,332,1511]" captionTargetId="figure@6.[151,1435,309,1518]" captionTargetPageId="6" captionText="FIGURES 22–37. Pygidium of Goliathopsis Janson, 1881 species (male in left, female in right; posterior view). 22–23, G. cer- vus Janson, 1881; 24–25, G. despectus (Westwood, 1873), lectotype and paralectotype of G. capreolus Gestro, 1888 in MSNG; 26–27, G. duponti Antoine, 1991; 28–29, G. esquiroli Pouillaude, 1913; 30–31, G. ferreroi Antoine, 1991; 32–33, G. gressitti Ruter, 1978; 34–35, G. lameyi Fairmaire, 1893; 36–37, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884731" httpUri="https://zenodo.org/record/3884731/files/figure.png" pageId="2" pageNumber="93">Figs. 22–37</figureCitation>
|
||
). Femora and tibiae usually partly clad with tomentum; two teeth on the outer margin of protibia; tarsi slender, unmodified.
|
||
</paragraph>
|
||
<paragraph id="8B9A36D1CE05FFBFFF5BF952FD36F8E0" blockId="2.[151,1437,1216,2035]" pageId="2" pageNumber="93">
|
||
<emphasis id="B951EAC3CE05FFBFFF5BF952FE0AF928" bold="true" box="[199,440,1720,1747]" pageId="2" pageNumber="93">Sexual dimorphism.</emphasis>
|
||
Genae of male with a pair of dendritic horns, outer surface glabrous, inner surface covered with tomentum or glabrous (
|
||
<figureCitation id="131E2A54CE05FFBFFD8AF937FD12F90C" box="[534,672,1757,1783]" captionStart="FIGURES 13–21" captionStartId="5.[151,264,1549,1574]" captionTargetBox="[177,1420,257,1500]" captionTargetId="figure@5.[151,1435,234,1521]" captionTargetPageId="5" captionText="FIGURES 13–21. Cephalic horns of Goliathopsis Janson, 1881 species (dorsal view). 13, G. cervus Janson, 1881; 14, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in MNHN; 15, G. duponti Antoine, 1991; 16, G. es- quiroli Pouillaude, 1913; 17–18, G. ferreroi Antoine, 1991; 19, G. gressitti Ruter, 1978; 20, G. lameyi Fairmaire, 1893; 21, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884729" httpUri="https://zenodo.org/record/3884729/files/figure.png" pageId="2" pageNumber="93">Figs. 13–21</figureCitation>
|
||
). Ventral surface of female covered with tomentum, but abdominal sternites VI–VII without tomentum in male.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33F655ACE05FFB9FF5BF8CEFA23FD08" lastPageId="4" lastPageNumber="95" pageId="2" pageNumber="93" type="distribution">
|
||
<paragraph id="8B9A36D1CE05FFBFFF5BF8CEFDABF8C4" blockId="2.[151,1437,1216,2035]" box="[199,537,1828,1855]" pageId="2" pageNumber="93">
|
||
<emphasis id="B951EAC3CE05FFBFFF5BF8CEFED3F8C4" bold="true" box="[199,353,1828,1855]" pageId="2" pageNumber="93">Distribution.</emphasis>
|
||
Oriental region.
|
||
</paragraph>
|
||
<paragraph id="8B9A36D1CE05FFBEFF5BF8A3FB03FF00" blockId="2.[151,1437,1216,2035]" lastBlockId="3.[151,1436,153,251]" lastPageId="3" lastPageNumber="94" pageId="2" pageNumber="93">
|
||
<emphasis id="B951EAC3CE05FFBFFF5BF8A3FEC5F898" bold="true" box="[199,375,1865,1891]" pageId="2" pageNumber="93">Nomenclature.</emphasis>
|
||
The genus
|
||
<taxonomicName id="4C254D52CE05FFBFFE60F8A3FD39F898" authorityName="Janson" authorityYear="1881" box="[508,651,1865,1891]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EAC3CE05FFBFFE60F8A3FD39F898" box="[508,651,1865,1891]" italics="true" pageId="2" pageNumber="93">Goliathopsis</emphasis>
|
||
</taxonomicName>
|
||
was established by
|
||
<bibRefCitation id="EFB44B20CE05FFBFFCECF8A3FBBCF898" author="Janson, O. E." box="[880,1038,1865,1891]" pageId="2" pageNumber="93" pagination="603 - 611" refId="ref20457" refString="Janson, O. E. (1881) Notices of new or little known Cetoniidae No. 7. Cistula Entomologica, 2 (24), 603 - 611." type="journal article" year="1881">Janson (1881)</bibRefCitation>
|
||
for his new species
|
||
<taxonomicName id="4C254D52CE05FFBFFB64F8A3FAD7F898" authorityName="in Janson" authorityYear="1881" box="[1272,1381,1865,1891]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="cervus">
|
||
<emphasis id="B951EAC3CE05FFBFFB64F8A3FAD7F898" box="[1272,1381,1865,1891]" italics="true" pageId="2" pageNumber="93">G. cervus</emphasis>
|
||
</taxonomicName>
|
||
, and type species of the genus was not fixed in the original publication. While one
|
||
<typeStatus id="549E8873CE05FFBFFC6FF887FBF8F87C" box="[1011,1098,1901,1927]" pageId="2" pageNumber="93" type="syntype">syntype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C254D52CE05FFBFFBF2F887FEAEF850" authority="Westwood, 1873" authorityName="Westwood" authorityYear="1873" class="Insecta" family="Cetoniidae" genus="Pilinurgus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="despectus">
|
||
<emphasis id="B951EAC3CE05FFBFFBF2F887FAE4F87C" box="[1134,1366,1901,1927]" italics="true" pageId="2" pageNumber="93">Pilinurgus despectus</emphasis>
|
||
<bibRefCitation id="EFB44B20CE05FFBFFAC0F887FEAEF850" author="Westwood, J. O." pageId="2" pageNumber="93" refId="ref21916" refString="Westwood, J. O. (1873) Thesaurus Entomologicus Oxoniensis; or, Illustrations of new, rare, and interesting insects, for the most part contained in the collections presented to the University of Oxford by the Rev. F. W. Hope. Part I. Clarendon Press, Oxford, 56 pp., 10 pls. https: // doi. org / 10.5962 / bhl. title. 14077" type="book" year="1873">Westwood, 1873</bibRefCitation>
|
||
</taxonomicName>
|
||
was examined by
|
||
<bibRefCitation id="EFB44B20CE05FFBFFE6DF87BFD3DF850" author="Janson, O. E." box="[497,655,1937,1963]" pageId="2" pageNumber="93" pagination="603 - 611" refId="ref20457" refString="Janson, O. E. (1881) Notices of new or little known Cetoniidae No. 7. Cistula Entomologica, 2 (24), 603 - 611." type="journal article" year="1881">Janson (1881)</bibRefCitation>
|
||
for comparison of female characters. It is suggested that both names can be treated as originally included species of the genus
|
||
<taxonomicName id="4C254D52CE05FFBFFCAEF85FFC73F834" authorityName="Janson" authorityYear="1881" box="[818,961,1973,1999]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EAC3CE05FFBFFCAEF85FFC73F834" box="[818,961,1973,1999]" italics="true" pageId="2" pageNumber="93">Goliathopsis</emphasis>
|
||
</taxonomicName>
|
||
(following Article 67.2 of ICZN).
|
||
<bibRefCitation id="EFB44B20CE05FFBFFACFF85FFF50F808" author="Arrow, G. J." pageId="2" pageNumber="93" refId="ref19451" refString="Arrow, G. J. (1910) The Fauna of British India (Including Ceylon and Burma) Coleoptera, Lamellicornia (Cetoniinae and Dy- nastinae). Taylor & Francis, London, 322 pp." type="book" year="1910">Arrow (1910)</bibRefCitation>
|
||
later designated
|
||
<taxonomicName id="4C254D52CE05FFBFFE3DF833FCB7F808" authority="(Westwood, 1873)" baseAuthorityName="Westwood" baseAuthorityYear="1873" box="[417,773,2009,2035]" class="Insecta" family="Cetoniidae" genus="Pilinurgus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="despectus">
|
||
<emphasis id="B951EAC3CE05FFBFFE3DF833FD82F808" box="[417,560,2009,2035]" italics="true" pageId="2" pageNumber="93">G. despectus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB44B20CE05FFBFFDA2F833FD4FF808" author="Westwood, J. O." box="[574,765,2009,2035]" pageId="2" pageNumber="93" refId="ref21916" refString="Westwood, J. O. (1873) Thesaurus Entomologicus Oxoniensis; or, Illustrations of new, rare, and interesting insects, for the most part contained in the collections presented to the University of Oxford by the Rev. F. W. Hope. Part I. Clarendon Press, Oxford, 56 pp., 10 pls. https: // doi. org / 10.5962 / bhl. title. 14077" type="book" year="1873">Westwood, 1873</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
as type species, and the accusation on subsequent designa- tion is apparently unreasonable (
|
||
<bibRefCitation id="EFB44B20CE04FFBEFD97FF73FD14FF48" author="Krajcik, M." box="[523,678,153,179]" pageId="3" pageNumber="94" pagination="1 - 113" refId="ref20569" refString="Krajcik, M. (2011) Illustrated Catalogue of Cetoniinae, Trichiinae and Valginae of China (Coleoptera, Cetoniidae). Animma. x, Supplement 1, 1 - 113." type="journal article" year="2011">Krajčík 2011</bibRefCitation>
|
||
). Additionally, regarding the name of the subtribe
|
||
<taxonomicName id="4C254D52CE04FFBEFB75FF73FA2EFF48" authorityName="Krikken" authorityYear="1984" box="[1257,1436,153,179]" family="Null" pageId="3" pageNumber="94" rank="subTribe" subTribe="Goliathopsidina">Goliathopsidina</taxonomicName>
|
||
(stem: “Goliathopsid-”),
|
||
<bibRefCitation id="EFB44B20CE04FFBEFE2EFF57FD06FF2C" author="Bouchard, P. & Bousquet, Y. & Davies, A. E. & Alonso-Zarazaga, M. A. & Lawrence, J. F. & Lyal, C. H. C. & Newton, A. F. & Reid, C. A. M. & Schmitt, M. & Slipinski, S. A. & Smith, A. B. T." box="[434,692,189,215]" pageId="3" pageNumber="94" pagination="1 - 972" refId="ref19586" refString="Bouchard, P., Bousquet, Y., Davies, A. E., Alonso-Zarazaga, M. A., Lawrence, J. F., Lyal, C. H. C., Newton, A. F., Reid, C. A. M., Schmitt, M., Slipinski, S. A., Smith, A. B. T. (2011) Family-group names in Coleoptera (Insecta). ZooKeys, 88, 1 - 972. https: // doi. org / 10.3897 / zookeys. 88.807" type="journal article" year="2011">
|
||
Bouchard
|
||
<emphasis id="B951EAC3CE04FFBEFDB4FF54FDD0FF2C" box="[552,610,189,215]" italics="true" pageId="3" pageNumber="94">et al.</emphasis>
|
||
(2011)
|
||
</bibRefCitation>
|
||
indicated that the correct stem should be “Goliathopse-” and the incorrect stem formation in prevailing usage was maintained under Article 29.5 (
|
||
<bibRefCitation id="EFB44B20CE04FFBEFB82FF0BFB17FF00" author="ICZN" box="[1054,1189,225,251]" pageId="3" pageNumber="94" refId="ref20403" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, The Natural History Museum, London, 306 pp. Available from: https: // www. iczn. org / the-code / the-international-code-of-zoological-nomenclature / the-code-online / (accessed 9 March 2020)" type="book" year="1999">ICZN 1999</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF5A6659CE04FFBEFF0BF943FE1EF8AA" ID-DOI="http://doi.org/10.5281/zenodo.3884727" ID-Zenodo-Dep="3884727" httpUri="https://zenodo.org/record/3884727/files/figure.png" pageId="3" pageNumber="94" startId="3.[151,264,1705,1730]" targetBox="[151,1435,278,1670]" targetPageId="3">
|
||
<paragraph id="8B9A36D1CE04FFBEFF0BF943FE1EF8AA" blockId="3.[151,1437,1705,1874]" pageId="3" pageNumber="94">
|
||
<emphasis id="B951EAC3CE04FFBEFF0BF943FEFAF939" bold="true" box="[151,328,1705,1730]" pageId="3" pageNumber="94">FIGURES 1–12.</emphasis>
|
||
Characters of
|
||
<taxonomicName id="4C254D52CE04FFBEFE41F943FD59F93A" authority="Janson, 1881" authorityName="Janson" authorityYear="1881" box="[477,747,1705,1729]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EAC3CE04FFBEFE41F943FDEDF93A" box="[477,607,1705,1729]" italics="true" pageId="3" pageNumber="94">Goliathopsis</emphasis>
|
||
<bibRefCitation id="EFB44B20CE04FFBEFDFAF940FD59F93A" author="Janson, O. E." box="[614,747,1705,1729]" pageId="3" pageNumber="94" pagination="603 - 611" refId="ref20457" refString="Janson, O. E. (1881) Notices of new or little known Cetoniidae No. 7. Cistula Entomologica, 2 (24), 603 - 611." type="journal article" year="1881">Janson, 1881</bibRefCitation>
|
||
</taxonomicName>
|
||
species. 1–4, mouthparts of
|
||
<taxonomicName id="4C254D52CE04FFBEFB93F943FA24F93A" authority="Antoine, 1991: 1" authorityName="Antoine" authorityPageNumber="1" authorityYear="1991" box="[1039,1430,1705,1730]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="duponti">
|
||
<emphasis id="B951EAC3CE04FFBEFB93F943FB56F93A" box="[1039,1252,1705,1729]" italics="true" pageId="3" pageNumber="94">Goliathopsis duponti</emphasis>
|
||
<bibRefCitation id="EFB44B20CE04FFBEFB76F943FA24F93A" author="Antoine, P." box="[1258,1430,1705,1730]" pageId="3" pageNumber="94" pagination="9 - 16" refId="ref19382" refString="Antoine, P. (1991) Quelques especes nouvelles ou peu connues de la famille des Cetoniidae II. Bulletin de la Societe Sciences Nat Compiegne, 69, 9 - 16." type="journal article" year="1991">Antoine, 1991: 1</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B951EAC3CE04FFBEFF0BF927FEBBF91E" box="[151,265,1741,1765]" italics="true" pageId="3" pageNumber="94">epipharynx</emphasis>
|
||
; 2,
|
||
<emphasis id="B951EAC3CE04FFBEFEADF927FE2BF91E" box="[305,409,1741,1765]" italics="true" pageId="3" pageNumber="94">mandibles</emphasis>
|
||
; 3,
|
||
<emphasis id="B951EAC3CE04FFBEFE5DF927FDAAF91E" box="[449,536,1741,1765]" italics="true" pageId="3" pageNumber="94">maxillae</emphasis>
|
||
; 4,
|
||
<emphasis id="B951EAC3CE04FFBEFDDCF927FD34F91E" box="[576,646,1741,1765]" italics="true" pageId="3" pageNumber="94">labium</emphasis>
|
||
. 5–12, setiferous punctures on the disc of elytron: 5,
|
||
<taxonomicName id="4C254D52CE04FFBEFB34F924FA2BF91E" authority="Janson, 1881" authorityName="in Janson" authorityYear="1881" box="[1192,1433,1741,1765]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="cervus">
|
||
<emphasis id="B951EAC3CE04FFBEFB34F924FABBF91E" box="[1192,1289,1742,1765]" italics="true" pageId="3" pageNumber="94">G. cervus</emphasis>
|
||
<bibRefCitation id="EFB44B20CE04FFBEFA8CF924FA2BF91E" author="Janson, O. E." box="[1296,1433,1741,1765]" pageId="3" pageNumber="94" pagination="603 - 611" refId="ref20457" refString="Janson, O. E. (1881) Notices of new or little known Cetoniidae No. 7. Cistula Entomologica, 2 (24), 603 - 611." type="journal article" year="1881">Janson, 1881</bibRefCitation>
|
||
</taxonomicName>
|
||
; 6,
|
||
<taxonomicName id="4C254D52CE04FFBEFF2DF918FE4BF8F2" authority="(Westwood, 1873)" baseAuthorityName="Westwood" baseAuthorityYear="1873" box="[177,505,1777,1802]" class="Insecta" family="Cetoniidae" genus="Pilinurgus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="despectus">
|
||
<emphasis id="B951EAC3CE04FFBEFF2DF918FE81F8F2" box="[177,307,1777,1801]" italics="true" pageId="3" pageNumber="94">G. despectus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB44B20CE04FFBEFEDEF91BFE40F8F2" author="Westwood, J. O." box="[322,498,1777,1802]" pageId="3" pageNumber="94" refId="ref21916" refString="Westwood, J. O. (1873) Thesaurus Entomologicus Oxoniensis; or, Illustrations of new, rare, and interesting insects, for the most part contained in the collections presented to the University of Oxford by the Rev. F. W. Hope. Part I. Clarendon Press, Oxford, 56 pp., 10 pls. https: // doi. org / 10.5962 / bhl. title. 14077" type="book" year="1873">Westwood, 1873</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, paralectotype of
|
||
<taxonomicName id="4C254D52CE04FFBEFD2DF91BFC0DF8F2" authority="Gestro, 1888" authorityName="Gestro" authorityYear="1888" box="[689,959,1777,1801]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="98" phylum="Arthropoda" rank="species" species="capreolus">
|
||
G.
|
||
<emphasis id="B951EAC3CE04FFBEFD4DF91BFC86F8F2" box="[721,820,1777,1801]" italics="true" pageId="3" pageNumber="94">capreolus</emphasis>
|
||
<bibRefCitation id="EFB44B20CE04FFBEFCA7F91BFC0DF8F2" author="Gestro, R." box="[827,959,1777,1801]" pageId="3" pageNumber="94" pagination="87 - 132" refId="ref20159" refString="Gestro, R. (1888) Viaggio di Leonardo Fea in Birmania regione vicine. IV. Nuove specie di Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 6, 87 - 132." type="journal article" year="1888">Gestro, 1888</bibRefCitation>
|
||
</taxonomicName>
|
||
in BMNH; 7,
|
||
<taxonomicName id="4C254D52CE04FFBEFBCAF918FAECF8F2" authority="Antoine, 1991" authorityName="Antoine" authorityYear="1991" box="[1110,1374,1777,1802]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="duponti">
|
||
<emphasis id="B951EAC3CE04FFBEFBCAF918FB71F8F2" box="[1110,1219,1777,1801]" italics="true" pageId="3" pageNumber="94">G. duponti</emphasis>
|
||
<bibRefCitation id="EFB44B20CE04FFBEFB55F91BFAECF8F2" author="Antoine, P." box="[1225,1374,1777,1802]" pageId="3" pageNumber="94" pagination="9 - 16" refId="ref19382" refString="Antoine, P. (1991) Quelques especes nouvelles ou peu connues de la famille des Cetoniidae II. Bulletin de la Societe Sciences Nat Compiegne, 69, 9 - 16." type="journal article" year="1991">Antoine, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
; 8,
|
||
<taxonomicName id="4C254D52CE04FFBEFA18F918FE18F8D6" authority="Pouillaude, 1913" authorityName="Pouillaude" authorityYear="1913" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="esquiroli">
|
||
<emphasis id="B951EAC3CE04FFBEFA18F918FF42F8D6" italics="true" pageId="3" pageNumber="94">G. esquiroli</emphasis>
|
||
<bibRefCitation id="EFB44B20CE04FFBEFF65F8FFFE18F8D6" author="Pouillaude, I." box="[249,426,1813,1838]" pageId="3" pageNumber="94" pagination="421 - 427" refId="ref21344" refString="Pouillaude, I. (1913) Description de trois nouvelles espece de Cetonides. Insecta, 3, 421 - 427." type="journal article" year="1913">Pouillaude, 1913</bibRefCitation>
|
||
</taxonomicName>
|
||
; 9,
|
||
<taxonomicName id="4C254D52CE04FFBEFE4EF8FCFD6FF8D6" authority="Antoine, 1991" authorityName="Antoine" authorityYear="1991" box="[466,733,1813,1838]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="ferreroi">
|
||
<emphasis id="B951EAC3CE04FFBEFE4EF8FCFDF2F8D6" box="[466,576,1813,1837]" italics="true" pageId="3" pageNumber="94">G. ferreroi</emphasis>
|
||
<bibRefCitation id="EFB44B20CE04FFBEFDDBF8FFFD6FF8D6" author="Antoine, P." box="[583,733,1813,1838]" pageId="3" pageNumber="94" pagination="9 - 16" refId="ref19382" refString="Antoine, P. (1991) Quelques especes nouvelles ou peu connues de la famille des Cetoniidae II. Bulletin de la Societe Sciences Nat Compiegne, 69, 9 - 16." type="journal article" year="1991">Antoine, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
; 10,
|
||
<taxonomicName id="4C254D52CE04FFBEFC8DF8FCFBBAF8D6" authority="Ruter, 1978" authorityName="Ruter" authorityYear="1978" box="[785,1032,1813,1837]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="gressitti">
|
||
<emphasis id="B951EAC3CE04FFBEFC8DF8FCFC31F8D6" box="[785,899,1814,1837]" italics="true" pageId="3" pageNumber="94">G. gressitti</emphasis>
|
||
<bibRefCitation id="EFB44B20CE04FFBEFC10F8FCFBBAF8D6" author="Ruter, G." box="[908,1032,1813,1837]" pageId="3" pageNumber="94" pagination="285 - 287" refId="ref21505" refString="Ruter, G. (1978) Description de Cetonides nouveaux. Nouvelle Revue d'Entomologie, 8 (3), 285 - 287." type="journal article" year="1978">Ruter, 1978</bibRefCitation>
|
||
</taxonomicName>
|
||
; 11,
|
||
<taxonomicName id="4C254D52CE04FFBEFBA7F8FCFAFDF8D6" authority="Fairmaire, 1893" authorityName="Fairmaire" authorityYear="1893" box="[1083,1359,1813,1838]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="lameyi">
|
||
<emphasis id="B951EAC3CE04FFBEFBA7F8FCFB12F8D6" box="[1083,1184,1813,1837]" italics="true" pageId="3" pageNumber="94">G. lameyi</emphasis>
|
||
Fairmaire, 1893
|
||
</taxonomicName>
|
||
; 12,
|
||
<taxonomicName id="4C254D52CE04FFBEFA18F8FCFE1AF8AA" authority="Pouillaude, 1913" authorityName="Pouillaude" authorityYear="1913" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="velutinus">
|
||
<emphasis id="B951EAC3CE04FFBEFA18F8FCFF40F8AA" italics="true" pageId="3" pageNumber="94">G. velutinus</emphasis>
|
||
<bibRefCitation id="EFB44B20CE04FFBEFF65F8D3FE1AF8AA" author="Pouillaude, I." box="[249,424,1849,1874]" pageId="3" pageNumber="94" pagination="421 - 427" refId="ref21344" refString="Pouillaude, I. (1913) Description de trois nouvelles espece de Cetonides. Insecta, 3, 421 - 427." type="journal article" year="1913">Pouillaude, 1913</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9A36D1CE04FFB9FF5BF86AFBAEFDC4" blockId="3.[151,1437,1919,2018]" lastBlockId="4.[151,1437,153,755]" lastPageId="4" lastPageNumber="95" pageId="3" pageNumber="94">
|
||
<emphasis id="B951EAC3CE04FFBEFF5BF86AFE35F861" bold="true" box="[199,391,1919,1946]" pageId="3" pageNumber="94">Natural history.</emphasis>
|
||
Adults of the tribe
|
||
<taxonomicName id="4C254D52CE04FFBEFDF4F86AFC95F861" authorityName="Burmeister & Schaum" authorityYear="1841" box="[616,807,1920,1946]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="tribe" tribe="Cremastocheilini">Cremastocheilini</taxonomicName>
|
||
were usually believed as predaceous (
|
||
<bibRefCitation id="EFB44B20CE04FFBEFB40F86AFA25F861" author="Westwood, J. O." box="[1244,1431,1920,1946]" pageId="3" pageNumber="94" refId="ref21916" refString="Westwood, J. O. (1873) Thesaurus Entomologicus Oxoniensis; or, Illustrations of new, rare, and interesting insects, for the most part contained in the collections presented to the University of Oxford by the Rev. F. W. Hope. Part I. Clarendon Press, Oxford, 56 pp., 10 pls. https: // doi. org / 10.5962 / bhl. title. 14077" type="book" year="1873">Westwood 1873</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB44B20CE04FFBEFF0BF84EFE96F845" author="Arrow, G. J." box="[151,292,1956,1982]" pageId="3" pageNumber="94" refId="ref19451" refString="Arrow, G. J. (1910) The Fauna of British India (Including Ceylon and Burma) Coleoptera, Lamellicornia (Cetoniinae and Dy- nastinae). Taylor & Francis, London, 322 pp." type="book" year="1910">Arrow 1910</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB44B20CE04FFBEFEAEF84EFE61F845" author="Krikken, J." box="[306,467,1956,1982]" pageId="3" pageNumber="94" pagination="1 - 75" refId="ref20712" refString="Krikken, J. (1984) A new key to the suprageneric taxa in the beetle family Cetoniidae, with annotated lists of the known genera. Zoologische Verhandelingen, 210, 1 - 75." type="journal article" year="1984">Krikken 1984</bibRefCitation>
|
||
). By dissecting mouthparts of an adult of
|
||
<taxonomicName id="4C254D52CE04FFBEFC2BF84EFB17F845" authorityName="Antoine" authorityYear="1991" box="[951,1189,1956,1982]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="duponti">
|
||
<emphasis id="B951EAC3CE04FFBEFC2BF84EFB17F845" box="[951,1189,1956,1982]" italics="true" pageId="3" pageNumber="94">Goliathopsis duponti</emphasis>
|
||
</taxonomicName>
|
||
, the slightly enlarged mentum, sparse setae on epipharynx and galea, and heavily sclerotized maxillae and mandibles were observed (
|
||
<figureCitation id="131E2A54CE03FFB9FF03FF73FEBFFF48" box="[159,269,153,179]" captionStart="FIGURES 1–12" captionStartId="3.[151,264,1705,1730]" captionTargetBox="[151,1435,278,1670]" captionTargetId="figure@3.[151,1435,273,1680]" captionTargetPageId="3" captionText="FIGURES 1–12. Characters of Goliathopsis Janson, 1881 species. 1–4, mouthparts of Goliathopsis duponti Antoine, 1991: 1, epipharynx; 2, mandibles; 3, maxillae; 4, labium. 5–12, setiferous punctures on the disc of elytron: 5, G. cervus Janson, 1881; 6, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in BMNH; 7, G. duponti Antoine, 1991; 8, G. esquiroli Pouillaude, 1913; 9, G. ferreroi Antoine, 1991; 10, G. gressitti Ruter, 1978; 11, G. lameyi Fairmaire, 1893; 12, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884727" httpUri="https://zenodo.org/record/3884727/files/figure.png" pageId="4" pageNumber="95">Figs. 1–4</figureCitation>
|
||
). These characters are generally congruous with the typical mouthparts of
|
||
<taxonomicName id="4C254D52CE03FFB9FBCAFF73FAA7FF48" authorityName="Burmeister & Schaum" authorityYear="1841" box="[1110,1301,153,179]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="tribe" tribe="Cremastocheilini">Cremastocheilini</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFB44B20CE03FFB9FAB9FF73FF60FF2C" author="Westwood, J. O." pageId="4" pageNumber="95" refId="ref21916" refString="Westwood, J. O. (1873) Thesaurus Entomologicus Oxoniensis; or, Illustrations of new, rare, and interesting insects, for the most part contained in the collections presented to the University of Oxford by the Rev. F. W. Hope. Part I. Clarendon Press, Oxford, 56 pp., 10 pls. https: // doi. org / 10.5962 / bhl. title. 14077" type="book" year="1873">Westwood 1873</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB44B20CE03FFB9FF43FF57FE70FF2C" author="Nel, A. & Scholtz, C. H." box="[223,450,189,215]" pageId="4" pageNumber="95" pagination="1 - 84" refId="ref21219" refString="Nel, A. & Scholtz, C. H. (1990) Comparative morphology of the mouthparts of adult Scarabaeoidea (Coleoptera). Entomology Memoir, 80, 1 - 84." type="journal article" year="1990">Nel & Scholtz 1990</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB44B20CE03FFB9FE53FF57FDDEFF2C" author="Li, J. K. & Zhang, X. P. & Gao, M. X. & Zhang, L. M. & Huang, L. R. & Flutsch, G." box="[463,620,189,215]" pageId="4" pageNumber="95" pagination="30 - 40" refId="ref20749" refString="Li, J. K., Zhang, X. P., Gao, M. X., Zhang, L. M., Huang, L. R. & Flutsch, G. (2013) Cetoniidae from Mount Gaoligongshan, Yunnan, China (Coleoptera, Scarabaeoidea, Cetoniidae). Cetoniimania, New Series, 5, 30 - 40." type="journal article" year="2013">
|
||
Li
|
||
<emphasis id="B951EAC3CE03FFB9FE73FF54FD9AFF2C" box="[495,552,189,215]" italics="true" pageId="4" pageNumber="95">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
). However, the apical portion of molar, lacinia, and galea are not shaped as that of the predaceous genera, such as
|
||
<taxonomicName id="4C254D52CE03FFB9FDFAFF0BFCDDFF00" authority="Hope, 1831" authorityName="Hope" authorityYear="1831" box="[614,879,225,251]" class="Insecta" family="Cetoniidae" genus="Campsiura" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EAC3CE03FFB9FDFAFF0BFD50FF00" box="[614,738,225,251]" italics="true" pageId="4" pageNumber="95">Campsiura</emphasis>
|
||
Hope, 1831
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C254D52CE03FFB9FCE7FF0BFB5AFF00" authority="Motschulsky, 1857" authorityName="Motschulsky" authorityYear="1857" box="[891,1256,224,251]" class="Insecta" family="Cetoniidae" genus="Clinterocera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EAC3CE03FFB9FCE7FF0BFBBBFF00" box="[891,1033,225,251]" italics="true" pageId="4" pageNumber="95">Clinterocera</emphasis>
|
||
Motschulsky, 1857
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C254D52CE03FFB9FABFFF0BFEE7FEE4" authority="Westwood, 1873" authorityName="Westwood" authorityYear="1873" class="Insecta" family="Cetoniidae" genus="Platysodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EAC3CE03FFB9FABFFF0BFA2EFF00" box="[1315,1436,225,251]" italics="true" pageId="4" pageNumber="95">Platysodes</emphasis>
|
||
<bibRefCitation id="EFB44B20CE03FFB9FF0BFEEFFEE7FEE4" author="Westwood, J. O." box="[151,341,261,287]" pageId="4" pageNumber="95" refId="ref21916" refString="Westwood, J. O. (1873) Thesaurus Entomologicus Oxoniensis; or, Illustrations of new, rare, and interesting insects, for the most part contained in the collections presented to the University of Oxford by the Rev. F. W. Hope. Part I. Clarendon Press, Oxford, 56 pp., 10 pls. https: // doi. org / 10.5962 / bhl. title. 14077" type="book" year="1873">Westwood, 1873</bibRefCitation>
|
||
</taxonomicName>
|
||
, but are blunt and somewhat similar to that of phytophagous scarabs (
|
||
<bibRefCitation id="EFB44B20CE03FFB9FBC5FEEFFA89FEE4" author="Nel, A. & Scholtz, C. H." box="[1113,1339,261,287]" pageId="4" pageNumber="95" pagination="1 - 84" refId="ref21219" refString="Nel, A. & Scholtz, C. H. (1990) Comparative morphology of the mouthparts of adult Scarabaeoidea (Coleoptera). Entomology Memoir, 80, 1 - 84." type="journal article" year="1990">Nel & Scholtz 1990</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB44B20CE03FFB9FADAFEEFFF60FEB8" author="Li, J. K. & Zhang, X. P. & Gao, M. X. & Zhang, L. M. & Huang, L. R. & Flutsch, G." pageId="4" pageNumber="95" pagination="30 - 40" refId="ref20749" refString="Li, J. K., Zhang, X. P., Gao, M. X., Zhang, L. M., Huang, L. R. & Flutsch, G. (2013) Cetoniidae from Mount Gaoligongshan, Yunnan, China (Coleoptera, Scarabaeoidea, Cetoniidae). Cetoniimania, New Series, 5, 30 - 40." type="journal article" year="2013">
|
||
Li
|
||
<emphasis id="B951EAC3CE03FFB9FAF9FEECFA2FFEE4" box="[1381,1437,261,287]" italics="true" pageId="4" pageNumber="95">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB44B20CE03FFB9FF42FEC3FE39FEB8" author="Qiu, J. Y. & Xu, H. & Chen, L." box="[222,395,296,323]" pageId="4" pageNumber="95" pagination="553 - 564" refId="ref21392" refString="Qiu, J. Y., Xu, H. & Chen, L. (2015) Review of the Oriental genus Platysodes Westwood (Coleoptera: Scarabaeidae: Cetoniinae: Cremastocheilini) with a redescription of Platysodes madoni Bourgoin. Zootaxa, 4021 (4), 553 - 564. https: // doi. org / 10.11646 / zootaxa. 4021.4.5" type="journal article" year="2015">
|
||
Qiu
|
||
<emphasis id="B951EAC3CE03FFB9FE8CFEC0FEFBFEB8" box="[272,329,297,323]" italics="true" pageId="4" pageNumber="95">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB44B20CE03FFB9FE0BFEC3FD8CFEB8" author="Xu, H. & Qiu, J. Y. & Huang, G. H." box="[407,574,297,323]" pageId="4" pageNumber="95" pagination="301 - 352" refId="ref22129" refString="Xu, H., Qiu, J. Y. & Huang, G. H. (2018) Taxonomy and natural history of the myrmecophilous genus Clinterocera Motschulsky, 1858 (Coleoptera: Scarabaeidae: Cetoniinae) from China and adjacent regions: revision of the C. jucunda species group. Zootaxa, 4531 (3), 301 - 352. https: // doi. org / 10.11646 / zootaxa. 4531.3.1" type="journal article" year="2018">
|
||
Xu
|
||
<emphasis id="B951EAC3CE03FFB9FE5DFEC0FE48FEB8" box="[449,506,297,323]" italics="true" pageId="4" pageNumber="95">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
). Adults of
|
||
<taxonomicName id="4C254D52CE03FFB9FD5CFEC3FCE2FEB8" baseAuthorityName="Westwood" baseAuthorityYear="1873" box="[704,848,297,323]" class="Insecta" family="Cetoniidae" genus="Pilinurgus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="despectus">
|
||
<emphasis id="B951EAC3CE03FFB9FD5CFEC3FCE2FEB8" box="[704,848,297,323]" italics="true" pageId="4" pageNumber="95">G. despectus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C254D52CE03FFB9FCC2FEC3FB33FEB8" authority="Antoine, 1991" authorityName="Antoine" authorityYear="1991" box="[862,1153,297,323]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="duponti">
|
||
<emphasis id="B951EAC3CE03FFB9FCC2FEC3FC65FEB8" box="[862,983,297,323]" italics="true" pageId="4" pageNumber="95">G. duponti</emphasis>
|
||
<bibRefCitation id="EFB44B20CE03FFB9FC42FEC3FB33FEB8" author="Antoine, P." box="[990,1153,297,323]" pageId="4" pageNumber="95" pagination="9 - 16" refId="ref19382" refString="Antoine, P. (1991) Quelques especes nouvelles ou peu connues de la famille des Cetoniidae II. Bulletin de la Societe Sciences Nat Compiegne, 69, 9 - 16." type="journal article" year="1991">Antoine, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C254D52CE03FFB9FB21FEC3FF7DFE9C" authority="Antoine, 1991" authorityName="Antoine" authorityYear="1991" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="ferreroi">
|
||
<emphasis id="B951EAC3CE03FFB9FB21FEC3FA87FEB8" box="[1213,1333,297,323]" italics="true" pageId="4" pageNumber="95">G. ferreroi</emphasis>
|
||
<bibRefCitation id="EFB44B20CE03FFB9FAA7FEC3FF7DFE9C" author="Antoine, P." pageId="4" pageNumber="95" pagination="9 - 16" refId="ref19382" refString="Antoine, P. (1991) Quelques especes nouvelles ou peu connues de la famille des Cetoniidae II. Bulletin de la Societe Sciences Nat Compiegne, 69, 9 - 16." type="journal article" year="1991">Antoine, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
have been reported as flower visitors indeed (
|
||
<bibRefCitation id="EFB44B20CE03FFB9FD52FEA7FCE9FE9C" author="Gestro, R." box="[718,859,333,359]" pageId="4" pageNumber="95" pagination="835 - 876" refId="ref20201" refString="Gestro, R. (1891) Viaggio di Leonardo Fea in Birmania regione vicine. XXXVII. Enumerazione delle Cetonie. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 10, 835 - 876." type="journal article" year="1891">Gestro 1891</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB44B20CE03FFB9FCFAFEA7FBF9FE9C" author="Sakai, K & Nagai, S." box="[870,1099,333,359]" pageId="4" pageNumber="95" refId="ref21636" refString="Sakai, K & Nagai, S. (1998) The Cetoniine Beetles of the World. Mushi-Sha's Iconographic Series of Insects. Vol. 3. Mushi-Sha, Tokyo, 421 pp." type="book" year="1998">Sakai & Nagai 1998</bibRefCitation>
|
||
), especially the latter two species are known from a large series of specimens (
|
||
<bibRefCitation id="EFB44B20CE03FFB9FD23FE9BFCEFFE70" author="Antoine, P." box="[703,861,369,395]" pageId="4" pageNumber="95" pagination="9 - 16" refId="ref19382" refString="Antoine, P. (1991) Quelques especes nouvelles ou peu connues de la famille des Cetoniidae II. Bulletin de la Societe Sciences Nat Compiegne, 69, 9 - 16." type="journal article" year="1991">Antoine 1991</bibRefCitation>
|
||
; see material examined in the present work). These facts suggested that adults of
|
||
<taxonomicName id="4C254D52CE03FFB9FE6DFE7FFD32FE54" authorityName="Janson" authorityYear="1881" box="[497,640,405,431]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EAC3CE03FFB9FE6DFE7FFD32FE54" box="[497,640,405,431]" italics="true" pageId="4" pageNumber="95">Goliathopsis</emphasis>
|
||
</taxonomicName>
|
||
are phytophagous species. Unfortunately, most species of this genus were rarely collected in the past three decades, and all the most recent collected adults (
|
||
<taxonomicName id="4C254D52CE03FFB9FB17FE53FF7DFE0C" authority="Pouillaude, 1913" authorityName="Pouillaude" authorityYear="1913" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="velutinus">
|
||
<emphasis id="B951EAC3CE03FFB9FB17FE53FAA6FE28" box="[1163,1300,441,467]" italics="true" pageId="4" pageNumber="95">G. velutinus</emphasis>
|
||
<bibRefCitation id="EFB44B20CE03FFB9FA81FE53FF7DFE0C" author="Pouillaude, I." pageId="4" pageNumber="95" pagination="421 - 427" refId="ref21344" refString="Pouillaude, I. (1913) Description de trois nouvelles espece de Cetonides. Insecta, 3, 421 - 427." type="journal article" year="1913">Pouillaude, 1913</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="49E1F833CE03FFB9FE89FE37FED9FE0C" box="[277,363,477,503]" country="China" name="Yunnan" pageId="4" pageNumber="95">Yunnan</collectingRegion>
|
||
and
|
||
<taxonomicName id="4C254D52CE03FFB9FE38FE37FD7CFE0C" authority="Fairmaire, 1893" authorityName="Fairmaire" authorityYear="1893" box="[420,718,477,503]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="lameyi">
|
||
<emphasis id="B951EAC3CE03FFB9FE38FE37FDA0FE0C" box="[420,530,477,503]" italics="true" pageId="4" pageNumber="95">G. lameyi</emphasis>
|
||
Fairmaire, 1893
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="49E1F833CE03FFB9FC89FE37FCC5FE03" box="[789,887,477,504]" country="China" name="Guizhou" pageId="4" pageNumber="95">Guizhou</collectingRegion>
|
||
) were caught on leaves of broadleaved trees, no any feeding behavior was observed (personal observation, authors,
|
||
<date id="FF9B1011CE03FFB9FC3AFDEBFBADFDE0" box="[934,1055,513,539]" pageId="4" pageNumber="95" value="2013-05">May 2013</date>
|
||
; Ji-Hui Liu,
|
||
<date id="FF9B1011CE03FFB9FB2BFDEBFA83FDE0" box="[1207,1329,513,539]" pageId="4" pageNumber="95" value="2014-06">June 2014</date>
|
||
; Ren-Zhi Zhang,
|
||
<date id="FF9B1011CE03FFB9FF71FDCFFEEFFDC4" box="[237,349,548,575]" pageId="4" pageNumber="95" value="2015-07">July 2015</date>
|
||
; Lu Qiu,
|
||
<date id="FF9B1011CE03FFB9FE5BFDCFFD8DFDC4" box="[455,575,549,575]" pageId="4" pageNumber="95" value="2019-06">June 2019</date>
|
||
). Their immature stages are still unknown.
|
||
</paragraph>
|
||
<paragraph id="8B9A36D1CE03FFB9FF5BFDA3FA23FD08" blockId="4.[151,1437,153,755]" pageId="4" pageNumber="95">
|
||
Some
|
||
<taxonomicName id="4C254D52CE03FFB9FE92FDA3FE2FFD98" authorityName="Janson" authorityYear="1881" box="[270,413,585,611]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EAC3CE03FFB9FE92FDA3FE2FFD98" box="[270,413,585,611]" italics="true" pageId="4" pageNumber="95">Goliathopsis</emphasis>
|
||
</taxonomicName>
|
||
species seem to mainly occur in Karst forests at low elevation (
|
||
<figureCitation id="131E2A54CE03FFB9FBF4FDA3FABFFD98" box="[1128,1293,584,611]" captionStart="FIGURES 165–168" captionStartId="34.[151,264,1845,1870]" captionTargetBox="[151,1435,499,1817]" captionTargetId="figure@34.[151,1435,486,1817]" captionTargetPageId="34" captionText="FIGURES 165–168. Habitat of Goliathopsis Janson, 1881 species. 165–166, a living male of G. lameyi Fairmaire, 1893 from Gengzao, Jia’ou, Libo County, near the Maolan Nature Reserve (Guizhou, China; photograph by Lu Qiu); 167–168, a living male of G. velutinus Pouillaude, 1913 from Liuku, Lushui County, in the valley of Nujiang River (Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.3884779" httpUri="https://zenodo.org/record/3884779/files/figure.png" pageId="4" pageNumber="95">Figs. 165–166</figureCitation>
|
||
), such as the
|
||
<typeStatus id="549E8873CE03FFB9FF0BFD87FF75FD7C" box="[151,199,621,647]" pageId="4" pageNumber="95">type</typeStatus>
|
||
specimens of
|
||
<taxonomicName id="4C254D52CE03FFB9FEF0FD87FE4EFD7C" box="[364,508,621,647]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="capreolus">
|
||
<emphasis id="B951EAC3CE03FFB9FEF0FD87FE4EFD7C" box="[364,508,621,647]" italics="true" pageId="4" pageNumber="95">G. capreolus</emphasis>
|
||
</taxonomicName>
|
||
, which were captured near a limestone cave in southern
|
||
<collectingCountry id="F3327641CE03FFB9FB1FFD87FB42FD7C" box="[1155,1264,621,647]" name="Myanmar" pageId="4" pageNumber="95">Myanmar</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFB44B20CE03FFB9FB63FD87FA3DFD7C" author="Gestro, R." box="[1279,1423,621,647]" pageId="4" pageNumber="95" pagination="87 - 132" refId="ref20159" refString="Gestro, R. (1888) Viaggio di Leonardo Fea in Birmania regione vicine. IV. Nuove specie di Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 6, 87 - 132." type="journal article" year="1888">Gestro 1888</bibRefCitation>
|
||
), and the known distributional localities of
|
||
<taxonomicName id="4C254D52CE03FFB9FDEDFD7BFD45FD50" authorityName="Pouillaude" authorityYear="1913" box="[625,759,657,683]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="esquiroli">
|
||
<emphasis id="B951EAC3CE03FFB9FDEDFD7BFD45FD50" box="[625,759,657,683]" italics="true" pageId="4" pageNumber="95">G. esquiroli</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C254D52CE03FFB9FCACFD7BFC2CFD50" authorityName="Fairmaire" authorityYear="1893" box="[816,926,657,683]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="lameyi">
|
||
<emphasis id="B951EAC3CE03FFB9FCACFD7BFC2CFD50" box="[816,926,657,683]" italics="true" pageId="4" pageNumber="95">G. lameyi</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<collectingCountry id="F3327641CE03FFB9FC58FD7BFBB5FD50" box="[964,1031,657,683]" name="China" pageId="4" pageNumber="95">China</collectingCountry>
|
||
are mostly located in the limestone area of
|
||
<collectingRegion id="49E1F833CE03FFB9FF76FD5FFEFFFD34" box="[234,333,693,719]" country="China" name="Guangxi" pageId="4" pageNumber="95">Guangxi</collectingRegion>
|
||
,
|
||
<collectingRegion id="49E1F833CE03FFB9FECBFD5FFE08FD34" box="[343,442,693,719]" country="China" name="Guizhou" pageId="4" pageNumber="95">Guizhou</collectingRegion>
|
||
, and
|
||
<collectingRegion id="49E1F833CE03FFB9FE68FD5FFD8DFD34" box="[500,575,693,719]" country="China" name="Hunan" pageId="4" pageNumber="95">Hunan</collectingRegion>
|
||
(see map,
|
||
<figureCitation id="131E2A54CE03FFB9FD2AFD5FFCA4FD2B" box="[694,790,693,720]" captionStart="FIGURE 164" captionStartId="33.[151,250,1616,1641]" captionTargetBox="[159,1427,209,1579]" captionTargetId="figure@33.[151,1436,181,1591]" captionTargetPageId="33" captionText="FIGURE 164. Known distribution of Goliathopsis Janson, 1881 species. Syncretic markers indicate identical or very close localities; type localities of the known species are marked, that of G. despectus (Westwood, 1873), G. cervus Janson, 1881, and G. esquiroli Pouillaude, 1913 are missing because the precise type localities are unknown." figureDoi="http://doi.org/10.5281/zenodo.3884777" httpUri="https://zenodo.org/record/3884777/files/figure.png" pageId="4" pageNumber="95">Fig. 164</figureCitation>
|
||
). While specimens data of
|
||
<taxonomicName id="4C254D52CE03FFB9FBDCFD5FFB75FD34" box="[1088,1223,693,719]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="velutinus">
|
||
<emphasis id="B951EAC3CE03FFB9FBDCFD5FFB75FD34" box="[1088,1223,693,719]" italics="true" pageId="4" pageNumber="95">G. velutinus</emphasis>
|
||
</taxonomicName>
|
||
show that this species lives at elevations of about
|
||
<quantity id="4CDD9B34CE03FFB9FE61FD33FD0EFD08" box="[509,700,728,755]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="4" pageNumber="95" unit="m" value="1.75" valueMax="2.5" valueMin="1.0">1,000 to 2,500 m</quantity>
|
||
in
|
||
<collectingRegion id="49E1F833CE03FFB9FD42FD33FC85FD08" box="[734,823,729,755]" country="China" name="Yunnan" pageId="4" pageNumber="95">Yunnan</collectingRegion>
|
||
. Ecological information on other species is unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |