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<document id="0A50C5661BA6CBC33905B68276997D6B" ID-CLB-Dataset="152185" ID-DOI="10.18590/euscorpius.2014.vol2014.iss191.1" ID-GBIF-Dataset="a6381f1d-22e1-4629-b369-07f5ab1d7834" ID-ISSN="1536-9307" ID-Zenodo-Dep="6505622" ID-ZooBank="E467B3C0-D693-4EAF-B5F0-759D8C63FE35" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638245227176" checkinUser="felipe" docAuthor="Lowe, Graeme, Yağmur, Ersen Aydın &amp; Kovařík, František" docDate="2014" docId="038C87BF2018191FFCBC8822DCDCF930" docLanguage="en" docName="Euscorpius.191.1-129.pdf" docOrigin="Euscorpius 191" docSource="https://doi.org/10.18590/euscorpius.2014.vol2014.iss191.1" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Leiurus hebraeus" docType="treatment" docVersion="13" lastPageNumber="79" masterDocId="FFB5FFC72052194EFFD78D2ADD40FF87" masterDocTitle="A review of the genus Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) with description of four new species from the Arabian Peninsula" masterLastPageNumber="129" masterPageNumber="1" pageNumber="73" updateTime="1699471136939" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="B91098BFD5BB88AA5D09917C50DBD187">A review of the genus Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) with description of four new species from the Arabian Peninsula</mods:title>
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<mods:namePart id="F4A9751AF1F1257D07E2AFF04014A790">Lowe, Graeme</mods:namePart>
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<mods:namePart id="8B74D71CB7807BFCAC53E340ED34AA4E">Yağmur, Ersen Aydın</mods:namePart>
<mods:affiliation id="D2C0F999680A8E204E1989E4F8048DF9">Alaşehir Vocational School, Celal Bayar University, Manisa, Turkey</mods:affiliation>
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<mods:namePart id="0132EE31F803933798C892DCB555A409">Kovařík, František</mods:namePart>
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<mods:title id="0E40145117F3D679FFCE98504E220A78">Euscorpius</mods:title>
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<treatment id="038C87BF2018191FFCBC8822DCDCF930" ID-DOI="http://doi.org/10.5281/zenodo.7117241" ID-GBIF-Taxon="201133229" ID-Zenodo-Dep="7117241" LSID="urn:lsid:plazi:treatment:038C87BF2018191FFCBC8822DCDCF930" httpUri="http://treatment.plazi.org/id/038C87BF2018191FFCBC8822DCDCF930" lastPageId="81" lastPageNumber="79" pageId="74" pageNumber="73">
<subSubSection id="C33F652220181904FCBC8822D833FAA2" box="[875,1395,1287,1317]" pageId="74" pageNumber="73" type="nomenclature">
<paragraph id="8B9A36A920181904FCBC8822D833FAA2" blockId="74.[859,1412,1287,1378]" box="[875,1395,1287,1317]" pageId="74" pageNumber="73">
<heading id="D0D281C520181904FCBC8822D833FAA2" box="[875,1395,1287,1317]" centered="true" fontSize="11" level="2" pageId="74" pageNumber="73" reason="2">
<taxonomicName id="4C254D2A20181904FCBC8822D850FAA3" ID-CoL="6P6WQ" authority="(Birula, 1908)" baseAuthorityName="Birula" baseAuthorityYear="1908" box="[875,1296,1287,1317]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="74" pageNumber="73" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB20181904FCBC8822D913FAA2" bold="true" box="[875,1107,1288,1317]" italics="true" pageId="74" pageNumber="73">Leiurus hebraeus</emphasis>
(
<bibRefCitation id="EFB44B5820181904FBB3882DD847FAA3" author="Birula" box="[1124,1287,1287,1316]" firstAuthor="Birula" journalOrPublisher="Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien" pageId="74" pageNumber="73" pagination="121 - 152" part="117" refId="ref59140" refString="BIRULA, A. 1908. Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 117 2 (1): 121 - 152." title="Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae" type="journal article" year="1908">Birula, 1908</bibRefCitation>
)
</taxonomicName>
<emphasis id="B951EABB20181904FACF8822D833FAA2" bold="true" box="[1304,1395,1288,1317]" pageId="74" pageNumber="73">stat. n.</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F652220181904FC8C8801D85BFAE5" pageId="74" pageNumber="73" type="description">
<paragraph id="8B9A36A920181904FC8C8801D85BFAE5" blockId="74.[859,1412,1287,1378]" pageId="74" pageNumber="73">
(
<figureCitation id="131E2A2C20181904FCB48801DEA4FAC4" box="[867,996,1323,1347]" captionStart-0="Figure 60" captionStart-1="Figure 61" captionStart-10="Figure 70" captionStart-11="Figure 71" captionStart-2="Figure 62" captionStart-3="Figure 63" captionStart-4="Figure 64" captionStart-5="Figure 65" captionStart-6="Figure 66" captionStart-7="Figure 67" captionStart-8="Figure 68" captionStart-9="Figure 69" captionStartId-0="78.[192,267,1592,1616]" captionStartId-1="79.[192,267,1593,1617]" captionStartId-10="88.[192,267,1055,1079]" captionStartId-11="89.[192,267,1598,1622]" captionStartId-2="80.[192,267,1303,1327]" captionStartId-3="81.[192,267,1470,1494]" captionStartId-4="82.[192,267,1712,1736]" captionStartId-5="83.[192,267,1697,1721]" captionStartId-6="84.[192,267,1300,1324]" captionStartId-7="85.[192,267,1640,1664]" captionStartId-8="86.[192,267,1809,1833]" captionStartId-9="87.[192,267,1103,1127]" captionTargetBox-0="[191,1438,167,1561]" captionTargetBox-1="[191,1439,167,1564]" captionTargetBox-10="[191,1440,191,1025]" captionTargetBox-11="[191,1439,191,1567]" captionTargetBox-2="[204,1425,204,1261]" captionTargetBox-3="[191,1440,133,1438]" captionTargetBox-4="[191,1437,205,1681]" captionTargetBox-5="[191,1437,191,1666]" captionTargetBox-6="[204,1425,204,1259]" captionTargetBox-7="[191,1440,133,1609]" captionTargetBox-8="[263,1361,215,1766]" captionTargetBox-9="[191,1440,167,1073]" captionTargetId-0="figure-173@78.[192,1437,169,1561]" captionTargetId-1="figure-161@79.[192,1438,169,1563]" captionTargetId-10="figure-490@88.[192,1439,192,1024]" captionTargetId-11="figure-114@89.[192,1438,192,1566]" captionTargetId-2="figure-333@80.[192,1437,192,1273]" captionTargetId-3="figure-166@81.[192,1439,144,1437]" captionTargetId-4="figure-87@82.[192,1436,206,1679]" captionTargetId-5="figure-114@83.[192,1436,192,1665]" captionTargetId-6="figure-353@84.[192,1437,192,1271]" captionTargetId-7="figure-121@85.[192,1439,153,1608]" captionTargetId-8="figure-43@86.[264,1360,216,1765]" captionTargetId-9="figure-471@87.[192,1439,168,1072]" captionTargetPageId-0="78" captionTargetPageId-1="79" captionTargetPageId-10="88" captionTargetPageId-11="89" captionTargetPageId-2="80" captionTargetPageId-3="81" captionTargetPageId-4="82" captionTargetPageId-5="83" captionTargetPageId-6="84" captionTargetPageId-7="85" captionTargetPageId-8="86" captionTargetPageId-9="87" captionText-0="Figure 60: Leiurus hebraeus (Birula, 1908) stat. n., male. Habitus, dorsal aspect. Israel (NHMB 17a)." captionText-1="Figure 61: Leiurus hebraeus (Birula, 1908) stat. n., male. Habitus, ventral aspect. Israel (NHMB 17a)." captionText-10="Figure 70: Leiurus hebraeus (Birula, 1908) stat. n., lectotype female, designated here (ZISP 578). Habitus. A. Dorsal aspect. B. Ventral aspect. Wadi ‘Arrud, Jordan." captionText-11="Figure 71: Leiurus hebraeus (Birula, 1908) stat. n., lectotype female and four paralectotypes, designated here (ZISP 578). Wadi ‘Arrud, Jordan. Label text in Russian: “Det. A. Birula, 5 ex. Coll. 25.IV.1897, Petro-Arabia, Wadi-Arrud, Southern shore of the Dead Sea, Davydov.”" captionText-2="Figure 62: Leiurus hebraeus (Birula, 1908) stat. n., male. A. Carapace and tergites. B. Coxosternal area and sternites. Israel (NHMB 17a)." captionText-3="Figure 63: Leiurus hebraeus (Birula, 1908) stat. n., male. Right pedipalp. A. Femur, dorsal aspect. B. Patella, dorsal aspect. C. Patella, external aspect. D. Chela, ventral aspect. E. Chela, dorsal aspect. F. Chela, external aspect. Israel (NHMB 17a)." captionText-4="Figure 64: Leiurus hebraeus (Birula, 1908) stat. n., female. Habitus, dorsal aspect. Israel (NHMB 17a)." captionText-5="Figure 65: Leiurus hebraeus (Birula, 1908) stat. n., female. Habitus, ventral aspect. Israel (NHMB 17a)." captionText-6="Figure 66: Leiurus hebraeus (Birula, 1908) stat. n., female. A. Carapace and tergites. B. Coxosternal area and sternites. Israel (NHMB 17a)." captionText-7="Figure 67: Leiurus hebraeus (Birula, 1908) stat. n., female. Right pedipalp. A. Femur, dorsal aspect. B. Patella, dorsal aspect. C. Patella, external aspect. D. Chela, ventral aspect. E. Chela, dorsal aspect. F. Fixed finger dentition. G. Movable finger dentition. H. Chela, external aspect. Israel (NHMB 17a)." captionText-8="Figure 68: Leiurus hebraeus (Birula, 1908) stat. n.. Metasoma. A. Male, lateral aspect. B. Male, ventral aspect. C. Female, lateral aspect. D. Female, ventral aspect. Male and female, Israel (NHMB 17a)." captionText-9="Figure 69: Leiurus hebraeus (Birula, 1908) stat. n., female. Tarsi. A-D. Left telotarsus and distal basitarsus, ventral aspect. A. Leg I. B. Leg II. C. Leg III. D. Leg IV. E. Right basitarsus III retrolateral aspect. A–D: female, Israel (NHMB 17a). E: female, Jericho, Israel (NHMB 17I-a)." figureDoi-0="http://doi.org/10.5281/zenodo.6505749" figureDoi-1="http://doi.org/10.5281/zenodo.6505751" figureDoi-10="http://doi.org/10.5281/zenodo.6505769" figureDoi-11="http://doi.org/10.5281/zenodo.6505771" figureDoi-2="http://doi.org/10.5281/zenodo.6505753" figureDoi-3="http://doi.org/10.5281/zenodo.6505755" figureDoi-4="http://doi.org/10.5281/zenodo.6505757" figureDoi-5="http://doi.org/10.5281/zenodo.6505759" figureDoi-6="http://doi.org/10.5281/zenodo.6505761" figureDoi-7="http://doi.org/10.5281/zenodo.6505763" figureDoi-8="http://doi.org/10.5281/zenodo.6505765" figureDoi-9="http://doi.org/10.5281/zenodo.6505767" httpUri-0="https://zenodo.org/record/6505749/files/figure.png" httpUri-1="https://zenodo.org/record/6505751/files/figure.png" httpUri-10="https://zenodo.org/record/6505769/files/figure.png" httpUri-11="https://zenodo.org/record/6505771/files/figure.png" httpUri-2="https://zenodo.org/record/6505753/files/figure.png" httpUri-3="https://zenodo.org/record/6505755/files/figure.png" httpUri-4="https://zenodo.org/record/6505757/files/figure.png" httpUri-5="https://zenodo.org/record/6505759/files/figure.png" httpUri-6="https://zenodo.org/record/6505761/files/figure.png" httpUri-7="https://zenodo.org/record/6505763/files/figure.png" httpUri-8="https://zenodo.org/record/6505765/files/figure.png" httpUri-9="https://zenodo.org/record/6505767/files/figure.png" pageId="74" pageNumber="73">Figs. 60–71</figureCitation>
,
<figureCitation id="131E2A2C20181904FC388801D95BFAC4" box="[1007,1051,1323,1347]" captionStart="Figure 87" captionStartId="107.[192,267,1798,1822]" captionTargetBox="[343,1333,196,1766]" captionTargetId="figure-90@107.[344,1333,197,1765]" captionTargetPageId="107" captionText="Figure 87: Metasoma V and telson of Leiurus spp. Males, lateral aspect. A. L. abdullahbayrami Yağmur, Koç et Kunt, 2009. B. L. arabicus sp. n.. C. L. brachycentrus (Ehrenberg, 1829) stat. n.. D. L. haenggii sp. n. E. L. heberti sp. n. F. L. hebraeus (Birula, 1908) stat. n.. G. L. macroctenus sp. n.. H. L. quinquestriatus (Ehrenberg, 1828). Locality data as in Figs. 9, 20, 31, 43, 55, 67, 79. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6505806" httpUri="https://zenodo.org/record/6505806/files/figure.png" pageId="74" pageNumber="73">87F</figureCitation>
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<figureCitation id="131E2A2C20181904FBF18801D912FAC4" box="[1062,1106,1323,1347]" captionStart="Figure 88" captionStartId="108.[192,267,1816,1840]" captionTargetBox="[318,1309,215,1785]" captionTargetId="figure-95@108.[319,1308,216,1784]" captionTargetPageId="108" captionText="Figure 88: Metasoma V and telson of Leiurus spp. Males, ventral aspect. A. L. abdullahbayrami Yağmur, Koç et Kunt, 2009. B. L. arabicus sp. n.. C. L. brachycentrus (Ehrenberg, 1829) stat. n.. D. L. haenggii sp. n.. E. L. heberti sp. n. F. L. hebraeus (Birula, 1908) stat. n.. G. L. macroctenus sp. n.. H. L. quinquestriatus (Ehrenberg, 1828). Locality data as in Figs. 9, 20, 31, 43, 55, 67, 79. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6505808" httpUri="https://zenodo.org/record/6505808/files/figure.png" pageId="74" pageNumber="73">88F</figureCitation>
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<figureCitation id="131E2A2C20181904FB8A8801D9C8FAC4" box="[1117,1160,1323,1347]" captionStart="Figure 89" captionStartId="109.[192,267,1809,1833]" captionTargetBox="[345,1336,208,1778]" captionTargetId="figure-94@109.[346,1335,209,1777]" captionTargetPageId="109" captionText="Figure 89: Metasoma V and telson of Leiurus spp. Females, lateral aspect. A. L. abdullahbayrami Yağmur, Koç et Kunt, 2009. B. L. arabicus sp. n. C. L. brachycentrus (Ehrenberg, 1829) stat. n. D. L. haenggii sp. n. E. L. heberti sp. n. F. L. hebraeus (Birula, 1908) stat. n. G. L. macroctenus sp. n. H. L. quinquestriatus (Ehrenberg, 1828). Locality data as in Figs. 9, 20, 31, 43, 55, 67, 79. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6505810" httpUri="https://zenodo.org/record/6505810/files/figure.png" pageId="74" pageNumber="73">89F</figureCitation>
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<figureCitation id="131E2A2C20181904FB438801D980FAC4" box="[1172,1216,1323,1347]" captionStart="Figure 90" captionStartId="110.[192,267,1808,1832]" captionTargetBox="[318,1309,207,1777]" captionTargetId="figure-95@110.[319,1308,208,1776]" captionTargetPageId="110" captionText="Figure 90: Metasoma V and telson of Leiurus spp. Females, ventral aspect. A. L. abdullahbayrami Yağmur, Koç et Kunt, 2009. B. L. arabicus sp. n. C. L. brachycentrus (Ehrenberg, 1829) stat. n.. D. L. haenggii sp. n. E. L. heberti sp. n. F. L. hebraeus (Birula, 1908) stat. n. G. L. macroctenus sp. n. H. L. quinquestriatus (Ehrenberg, 1828). Locality data as in Figs. 9, 20, 31, 43, 55, 67, 79. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6505812" httpUri="https://zenodo.org/record/6505812/files/figure.png" pageId="74" pageNumber="73">90F</figureCitation>
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<figureCitation id="131E2A2C20181904FB1D8801D9BAFAC4" box="[1226,1274,1323,1347]" captionStart="Figure 91" captionStartId="111.[192,267,1766,1790]" captionTargetBox="[303,1321,191,1734]" captionTargetId="figure-124@111.[304,1320,192,1733]" captionTargetPageId="111" captionText="Figure 91: Pigmentation patterns of metasoma IV, V of Leiurus species, ventral aspect. A. L. haenggii sp. n., holotype female, Ta’if, Saudi Arabia (NHMB 17k). B. L. macroctenus sp. n., holotype male, Thumrait, Oman. C. L. heberti sp. n., holotype male, Wadi Andur, Oman. D. L. quinquestriatus (Ehrenberg, 1828), female, Cairo, Egypt (NHMB 17d). E. L. brachycentrus (Ehrenberg, 1829) stat. n., male, Ad Darb, Saudi Arabia (NHMB 17ag). F. L. arabicus sp. n., holotype female, Kushm Dibi, Saudi Arabia (NHMB 17aq). G. L. hebraeus (Birula, 1908) stat. n., female, Israel (NHMB 17a). H. L. abdullahbayrami Yağmur, Koç et Kunt, 2009, male, Eski Sarkaya Village, Turkey. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6505814" httpUri="https://zenodo.org/record/6505814/files/figure.png" pageId="74" pageNumber="73">91G</figureCitation>
,
<figureCitation id="131E2A2C20181904FAD18801D876FAC4" box="[1286,1334,1323,1347]" captionStart="Figure 92" captionStartId="112.[192,267,1486,1510]" captionTargetBox="[224,1390,157,1461]" captionTargetId="figure-274@112.[212,1402,145,1473]" captionTargetPageId="112" captionText="Figure 92: Sternite III of Leiurus spp., female. A. L. abdullahbayrami Yağmur, Koç et Kunt, 2009, Caybasi Village, Turkey. B. L. arabicus sp. n., holotype, Kushm Dibi, Saudi Arabia (NHMB 17aq). C. L. arabicus sp. n., paratype, Al Amar, Bahrain (NHMB 17bg). D. L. heberti sp. n., paratype, Jabal Samhan. Oman. E. L. haenggii sp. n., paratype, Wadi Turabah, Saudi Arabia (NHMB 17al). F. L. haenggii sp. n., paratype, Adnan, Saudi Arabia (NHMB 17am). G. L. brachycentrus (Ehrenberg, 1829) stat. n., Al Mansuriah, Yemen. H. L. macroctenus sp. n., paratype, Masirah Island, Oman. I. L. macroctenus sp. n., paratype, Thumrait, Oman. J. L. quinquestriatus (Ehrenberg, 1828), Kharga Oasis, Egypt (NHMB 17i). K. L. hebraeus (Birula, 1908) stat. n., Israel (NHMB 17a). L. L. jordanensis Lourenço, Modry et Amr, 2002, al-Tawil, Saudi Arabia." figureDoi="http://doi.org/10.5281/zenodo.6505816" httpUri="https://zenodo.org/record/6505816/files/figure.png" pageId="74" pageNumber="73">92K</figureCitation>
,
<figureCitation id="131E2A2C20181904FA968801D83EFAC4" box="[1345,1406,1323,1347]" captionStart="Figure 93" captionStartId="113.[192,267,1572,1596]" captionTargetBox="[224,1390,195,1538]" captionTargetId="figure-226@113.[212,1402,183,1550]" captionTargetPageId="113" captionText="Figure 93: Medial intercarinal areas of tergite III of female Leiurus spp. A. L. arabicus sp. n., paratype, Jeddah, Saudi Arabia. B. L. arabicus sp. n., paratype, Wadi Khumrah, Saudi Arabia (NHMB 17bj). C. L. haenggii sp. n., paratype, Yemen. D. L. haenggii sp. n., paratype, Wadi Maraba, Saudi Arabia (NHMB 17al). E. L. heberti sp. n., paratype, Jabal Samhan. Oman. F. L. abdullahbayrami Yağmur, Koç et Kunt, 2009, Caybasi Village, Turkey. G. L. brachycentrus (Ehrenberg, 1829) stat. n., Al Mansuriah, Yemen. H. L. macroctenus sp. n., paratype, Masirah Island, Oman. I. L. hebraeus (Birula, 1908) stat. n., Israel (NHMB 17a). J. L. hebraeus Birula, 1908, Kurayyima, Jordan. K. L. quinquestriatus (Ehrenberg, 1828), Kharga Oasis, Egypt (NHMB 17i). L. L. jordanensis Lourenço, Modry et Amr, 2002, al-Tawil, Saudi Arabia." figureDoi="http://doi.org/10.5281/zenodo.6505818" httpUri="https://zenodo.org/record/6505818/files/figure.png" pageId="74" pageNumber="73">93I–J</figureCitation>
,
<figureCitation id="131E2A2C20181904FC138860D953FAE5" box="[964,1043,1354,1378]" captionStart="Figure 94" captionStartId="114.[192,260,765,786]" captionTargetBox="[204,1427,204,723]" captionTargetId="figure-776@114.[192,1439,192,745]" captionTargetPageId="114" captionText="Figure 94: Comparative analysis of granulation on tergite III of female Leiurus. A. UV fluorescence image of tergite III medial intercarinal surface of L. quinquestriatus (Egypt). B. Mask of granulation pattern on medial intercarinal surface from image in Figure 94A. C. Enlarged view of tergite III granulation in rectangular area indicated in Figure 94A. Granules were identified as local maxima of fluorescence intensity with contour shadows cast by directional UV illumination, and were modeled by elliptical regions-of-interest (ROIs). Parameters of granule ROIs were measured in ImageJ 1.44 (Rasband, 1997–2011). Area of granulometric analysis was restricted to surfaces bounded by the lateral and posterior marginal carinae, and a line passing through oblique, anterior transverse rows of enlarged granules (granules along carinae and bounding lines were omitted). D. Scatter plot of mean granule diameter vs. total granule area for 4 species of Leiurus (L. arabicus sp. n., L. haenggii sp. n., L. hebraeus stat. n. and L. quinquestriatus). Each point represents granulometric data derived from tergite III of one scorpion specimen (bilaterally, as shown in Figure 94B). Data were extracted from 18,423 ROIs from 36 scorpions. For each specimen, total granule area (a measure of density of granulation) was computed as sum of areas of all ROIs, and mean ROI diameter (a measure of coarseness of granulation) as the mean value of the maximum diameters of all granule ROIs. For comparative analysis, images from different size scorpions were resampled to equalize the distance between left and right posterior marginal granules of the lateral carinae (arbitrarily set to 4,000 units or pixels; linear dimensions expressed as [pixel], areas as [pixel2]). The total granule area separated the species into 2 groups: i.e. sparsely granulated (L. arabicus, L. haenggii) and densely granulated (L. hebraeus, L. quinquestriatus). In contrast, the distributions of ROI diameter were broadly overlapping. E. To obtain a more sensitive comparison of the coarseness of granulation, normalized cumulative distributions of single ROI areas for 3 species of Leiurus were computed (inset indicates number of scorpions analyzed). Relative horizontal positions of these curves indicated increasing coarseness of granulation, in rank order: L. haenggii &lt;L. quinquestriatus &lt;L. hebraeus (Ngranules = 2103, 5202, 2190 respectively). This ranking was confirmed by a Kolmogorov-Smirnov test which detected significant differences between the distributions of log (granule area) (P &lt;0.001)." figureDoi="http://doi.org/10.5281/zenodo.6505820" httpUri="https://zenodo.org/record/6505820/files/figure.png" pageId="74" pageNumber="73">94D–E</figureCitation>
,
<figureCitation id="131E2A2C20181904FBCA8860D97AFAE5" box="[1053,1082,1354,1378]" captionStart="Figure 95" captionStartId="118.[192,260,1217,1238]" captionTargetBox="[214,1409,159,1149]" captionTargetId="figure-476@118.[192,1437,128,1181]" captionTargetPageId="118" captionText="Figure 95: Variation in key diagnostic biometrics of adult Leiurus spp. A. Cumulative distributions of position of trichobothrium db relative to est on pedipalp fixed finger for: L. abdullahbayrami (53), L. brachycentrus stat. n. (11), L. hebraeus stat. n. (38), L. macroctenus sp. n. (176), L. quinquestriatus (61), L. haenggii sp. n. (158), L. arabicus sp. n. (87) (number of fingers measured in parentheses). Data for males and females were pooled. The db–est distances were normalized to pedipalp movable finger length (chord length from finger tip to external articular condyle), with positive values indicating db distal to est, negative values indicating db proximal to est. Distributions of L. haenggii (µ = 0.06367) and L. arabicus sp. n. (µ = 0.06671) were not significantly different (P = 0.405148, t test), indicating a close relationship between these species. Distribution of L. quinquestriatus (µ = 0.047313) was significantly different from those of L. haenggii and L. arabicus (P = 0.000102 and 0.000030 respectively). B. Scatter plot of morphometric ratios of mid-pectine sensillar margin L (MPSM, indicated in inset) to metasoma I W, and to carapace L, showing differences in relative pectinal tooth size in females of nine species of Leiurus. C. A subset of the female data in Figure 95B, plotted as cumulative distributions of morphometric ratio of mid-pectine sensillar margin L to metasoma I W. Distributions of L. haenggii (µ = 0.102268) and L. arabicus (µ = 0.10432) were not significantly different (P&gt; 0.1, K-S test). D. Scatter plot of morphometric ratios of mid-pectine sensillar margin L to metasoma I W (relative size of pectine teeth), and metasoma II L/W (slenderness of metasoma II) for females of nine species of Leiurus. A, C, D: symbol key as in Figure 95B." figureDoi="http://doi.org/10.5281/zenodo.6505822" httpUri="https://zenodo.org/record/6505822/files/figure.png" pageId="74" pageNumber="73">95</figureCitation>
,
<figureCitation id="131E2A2C20181904FB928860D9CAFAE5" box="[1093,1162,1354,1378]" captionStart-0="Figure 98" captionStart-1="Figure 99" captionStartId-0="121.[192,267,1251,1275]" captionStartId-1="122.[192,267,1014,1038]" captionTargetBox-0="[215,1377,224,1220]" captionTargetBox-1="[191,1438,222,978]" captionTargetId-0="figure-474@121.[192,1437,206,1246]" captionTargetId-1="figure-479@122.[192,1437,223,978]" captionTargetPageId-0="121" captionTargetPageId-1="122" captionText-0="Figure 98: Selected morphometrics of Leiurus, Cicileiurus, Cicileus and Compsobuthus compared to other Buthus group scorpions. A–B. Scatter plots of the fraction of fixed finger length distal to db (A) and est (B) vs. the ratio of movable finger length to carapace length. Each point represents one sex of one species. Larger ordinate values correspond to more basal positions of the trichobothria, and larger abscissa values to longer pedipalp fingers. There was a significant inverse correlation between relative length of the portion of the fixed finger distal to db and est (R = -0.6534, -0.5488, respectively), and the relative length of the movable finger (the latter being a measure of elongation of both fixed and movable fingers). Highlighted symbols show that Leiurus (light magenta circles), Cicileiurus (red triangle), Cicileus (green squares) and Compsobuthus (yellow circles) are located in the lower right halves of the plots, i.e. all have relatively elongated fingers and more distal placement of both db and est. Gray circles are data from other Buthus group species. C. Scatter plot of the fraction of fixed finger length distal to db vs. the fraction distal to est. The strong positive correlation (R = 0.8052) indicates a tendency for db and est to move together towards more distal locations as the fixed finger becomes more elongated. Species above the diagonal (solid blue) have db proximal to est, and those below have db distal to est. Solid gray lines in A–C are fits by least squares regression through all points. D. Scatter plot of pedipalp femur L/W (a measure of pedipalp elongation) vs. carapace L (a measure of body size). These two variables were not significantly correlated (R = 0.094). Data were compiled from the literature and specimens in the authors collections for 38 genera and 203 species representing the majority of taxa in the Buthus group, including both males (N = 124) and females (N = 97). Genera (and number of species) included: Afghanobuthus (1), Androctonus (11), Apistobuthus (2), Baloorthochirus (1), Birulatus (2), Buthacus (13), Butheoloides (11), Butheolus (5), Buthiscus (1), Buthus (26), Cicileiurus (1), Cicileus (2), Compsobuthus (33), Congobuthus (1), Darchenia (1), Gint (1), Hemibuthus (1), Hottentotta (31), Leiurus (10), Liobuthus (1), Lissothus (3), Mesobuthus (7), Neobuthus (2), Odontobuthus (6), Orthochirus (12), Pantobuthus (1), Pectinibuthus (1), Plesiobuthus (1), Polisius (1), Razianus (3), Saharobuthus (1), Somalibuthus (1), Vachoniolus (4), Vachonus (1)." captionText-1="Figure 99: Geographic plot of locality data of material examined for nine species of Leiurus distributed over northeast Africa, the Levant and Arabian Peninsula. Map colored by elevation with shaded relief. Additional locality data for: L. abdullahbayrami from Yağmur et al., 2009; for L. brachycentrus Ehrenberg, 1829 stat. n., from Simon, 1882 (Buthus beccarii)." figureDoi-0="http://doi.org/10.5281/zenodo.6505828" figureDoi-1="http://doi.org/10.5281/zenodo.6505832" httpUri-0="https://zenodo.org/record/6505828/files/figure.png" httpUri-1="https://zenodo.org/record/6505832/files/figure.png" pageId="74" pageNumber="73">98–99</figureCitation>
,
<tableCitation id="C6A7031220181904FB428860D9B9FAE5" box="[1173,1273,1354,1378]" captionStart="Table 3" captionStartId="105.[192,257,1349,1373]" captionTargetBox="[209,1890,232,1297]" captionTargetId="graphics-7@105.[194,1919,192,1298]" captionText="Table 3A: Summary of key characters separating four Leiurus species examined in this study (adult females unless otherwise specified). Numeric values indicate ranges with sample sizes in parentheses." httpUri="http://table.plazi.org/id/DF5A6621203B1927FF17886FDC8EFAF2" pageId="74" pageNumber="73" tableUuid="DF5A6621203B1927FF17886FDC8EFAF2">Tabs. 3A</tableCitation>
, 5)
</paragraph>
</subSubSection>
<subSubSection id="C33F652220181905FCE888ABDCE4FA37" lastPageId="75" lastPageNumber="74" pageId="74" pageNumber="73" type="reference_group">
<paragraph id="8B9A36A920181904FCE888ABD8DCFA1E" blockId="74.[831,1436,1409,1433]" box="[831,1436,1409,1433]" pageId="74" pageNumber="73">
<treatmentCitationGroup id="AB35118720181904FCE888ABD8DCFA1E" box="[831,1436,1409,1433]" pageId="74" pageNumber="73">
<treatmentCitation id="0A8410B820181904FCE888ABD8DCFA1E" author="Birula" box="[831,1436,1409,1433]" page="124" pageId="74" pageNumber="73" year="1908">
<taxonomicName id="4C254D2A20181904FCE888ABD8D8FA1E" authority="Birula, 1908: 124 - 129" authorityName="Birula" authorityPageNumber="124 - 129" authorityYear="1908" box="[831,1432,1409,1433]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="74" pageNumber="73" phylum="Arthropoda" rank="subSpecies" species="quinquestriatus" subSpecies="hebraeus">
<emphasis id="B951EABB20181904FCE888ABD9DEFA1E" box="[831,1182,1409,1433]" italics="true" pageId="74" pageNumber="73">Buthus quinquestriatus hebraeus</emphasis>
<bibRefCitation id="EFB44B5820181904FB7388ABD8D8FA1E" author="Birula" box="[1188,1432,1409,1433]" firstAuthor="Birula" journalOrPublisher="Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien" page="124 - 129" pageId="74" pageNumber="73" pagination="121 - 152" part="117" refId="ref59140" refString="BIRULA, A. 1908. Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 117 2 (1): 121 - 152." title="Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae" type="journal article" year="1908">Birula, 1908: 124–129</bibRefCitation>
</taxonomicName>
.
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</paragraph>
<paragraph id="8B9A36A920181904FCE88892DEA6FA57" blockId="74.[831,1441,1464,1918]" box="[831,998,1464,1488]" pageId="74" pageNumber="73">REFERENCES</paragraph>
<paragraph id="8B9A36A920181904FCE888FDD98FF9AB" blockId="74.[831,1441,1464,1918]" pageId="74" pageNumber="73">
<treatmentCitationGroup id="AB35118720181904FCE888FDD98FF9AB" pageId="74" pageNumber="73">
<taxonomicName id="4C254D2A20181904FCE888FDD942FA68" authority="Amoreux, 1789" authorityName="Amoreux" box="[831,1026,1495,1519]" class="Arachnida" family="Buthidae" genus="Scorpio" kingdom="Animalia" order="Scorpiones" pageId="74" pageNumber="73" phylum="Arthropoda" rank="species" species="occitanus">
<emphasis id="B951EABB20181904FCE888FDD942FA68" box="[831,1026,1495,1519]" italics="true" pageId="74" pageNumber="73">Scorpio occitanus</emphasis>
</taxonomicName>
:
<treatmentCitation id="0A8410B820181904FBC388FDD835FA68" author="AUDOUIN, V." box="[1044,1397,1495,1519]" page="173" pageId="74" pageNumber="73" year="1826">
<bibRefCitation id="EFB44B5820181904FBC388FDD87EFA68" author="AUDOUIN, V." box="[1044,1342,1495,1519]" firstAuthor="Audouin" journalOrPublisher="Histoire Naturelle, 1. C. L. P. Panekoucke, Paris" pageId="74" pageNumber="73" pagination="172 - 174" part="22" refId="ref58940" refString="AUDOUIN, V. 1826. Planche 8. Scorpions, Pinces, Solifuges. In: Explication sommaire des planches d'Arachnides de l'Egypte et de la Syrie, publiees par J. - C. Savigny. In: Description de l'Egypte, ou Recueil des Observations et des Recherches qui out ete Faites en Egypte Pendant l'Expedition de l'Armee Francaise. Histoire Naturelle, 1. C. L. P. Panekoucke, Paris, 22: 172 - 174." title="Planche 8. Scorpions, Pinces, Solifuges" type="book chapter" volumeTitle="Explication sommaire des planches d'Arachnides de l'Egypte et de la Syrie, publiees par J. - C. Savigny" year="1826">Audouin in Savigny, 1826</bibRefCitation>
: 173
</treatmentCitation>
, pl. VIII, fig. 1 (in part);
<treatmentCitation id="0A8410B820181904FBBA88DFDE8AF9AB" author="AUDOUIN, V." page="410" pageId="74" pageNumber="73" year="1827">
<bibRefCitation id="EFB44B5820181904FBBA88DFDE8AF9AB" author="AUDOUIN, V." firstAuthor="Audouin" journalOrPublisher="Paris" pageId="74" pageNumber="73" pagination="409 - 412" part="22" refId="ref59024" refString="AUDOUIN, V. 1827. Planche 8. Scorpions, Pinces, Solifuges. In: Explication sommaire des planches d'Arachnides de l'Egypte et de la Syrie, publiees par J. - C. SAVIGNY. In: Description de l'Egypte, ou Recueil des Observations et des Recherches qui out ete Faites en Egypte Pendant l'Expedition de l'Armee Francaise. Histoire Naturelle, 1. C. L. P. Panekoucke, Paris, 22: 409 - 412." title="Planche 8. Scorpions, Pinces, Solifuges" type="journal article" volumeTitle="Explication sommaire des planches d'Arachnides de l'Egypte et de la Syrie, publiees par J. - C. SAVIGNY." year="1827">Audouin in Savigny, 1827: 410-411</bibRefCitation>
</treatmentCitation>
, pl. VIII, fig. 1 (in part).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A36A920181904FCE88B19D984F884" blockId="74.[831,1441,1464,1918]" pageId="74" pageNumber="73">
<treatmentCitationGroup id="AB35118720181904FCE88B19D984F884" pageId="74" pageNumber="73">
<treatmentCitation id="0A8410B820181904FCE88B19D817F9CC" author="KRAEPELIN, K." box="[831,1367,1587,1611]" page="58" pageId="74" pageNumber="73" year="1891">
<taxonomicName id="4C254D2A20181904FCE88B19D817F9CC" authority=": Kraepelin, 1891: 58 - 60" authorityName=": Kraepelin" authorityPageNumber="58 - 60" authorityYear="1891" box="[831,1367,1587,1611]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="74" pageNumber="73" phylum="Arthropoda" rank="species" species="quinquestriatus">
<emphasis id="B951EABB20181904FCE88B19D97CF9CC" box="[831,1084,1587,1611]" italics="true" pageId="74" pageNumber="73">Buthus quinquestriatus</emphasis>
:
<bibRefCitation id="EFB44B5820181904FB878B19D817F9CC" author="Kraepelin" box="[1104,1367,1587,1611]" firstAuthor="Kraepelin" journalOrPublisher="Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten Hamburg" page="58 - 60" pageId="74" pageNumber="73" pagination="1 - 144" part="8" refId="ref61934" refString="KRAEPELIN, K. 1891. Revision der Skorpione. I. Die Familie der Androctonidae. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten Hamburg, 8: 1 - 144." title="Revision der Skorpione. I. Die Familie der Androctonidae" type="journal article" year="1891">Kraepelin, 1891: 58-60</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. I, fig. 7, pl. II, fig. 38 (in part);
<treatmentCitation id="0A8410B820181904FB148B78DED7F90F" author="POCOCK" page="242" pageId="74" pageNumber="73" year="1891">
<bibRefCitation id="EFB44B5820181904FB148B78DED7F90F" author="Pocock" firstAuthor="Pocock" page="242 - 243" pageId="74" pageNumber="73" pagination="241 - 247" refId="ref63303" refString="POCOCK, R. I. 1891. Notes on some scorpions collected by Mr. J. J. Walker, with descriptions of two new species and a new genus. Annals and Magazine of Natural History, (6), 8: 241 - 247." type="journal article" year="1891">Pocock, 1891: 242– 243</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B820181904FBDF8B5AD9F2F90F" author="PAVESI" box="[1032,1202,1648,1672]" page="4" pageId="74" pageNumber="73" year="1895">
<bibRefCitation id="EFB44B5820181904FBDF8B5AD9F2F90F" author="Pavesi" box="[1032,1202,1648,1672]" firstAuthor="Pavesi" page="4" pageId="74" pageNumber="73" pagination="1 - 11" refId="ref63148" refString="PAVESI, P. 1895. Viaggio del Dott. E. Festa in Palestina, nel Libano e regioni vicine. XIV. Aracnidi. Bollettino dei Musei di Zoologia ed Anatomia Comparata, 10 (216): 1 - 11." type="journal article" year="1895">Pavesi, 1895: 4</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B820181904FB688B5ADECAF920" author="KRAEPELIN, K." page="27" pageId="74" pageNumber="73" year="1899">
<bibRefCitation id="EFB44B5820181904FB688B5ADECAF920" author="Kraepelin" editor="Das Tierreich, R." firstAuthor="Kraepelin" journalOrPublisher="Friedlander und Sohn Verl Berlin" page="27 - 28" pageId="74" pageNumber="73" pagination="1 - 265" part="8" refId="ref61964" refString="KRAEPELIN, K. 1899. Scorpiones und Pedipalpi. In: Das Tierreich, R. Friedlander und Sohn Verl Berlin, 8: 1 - 265." title="Scorpiones und Pedipalpi" type="book chapter" year="1899">Kraepelin, 1899: 27- 28</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B820181904FC288BA5D9F8F920" author="SIMON" box="[1023,1208,1679,1703]" page="83" pageId="74" pageNumber="73" year="1892">
<bibRefCitation id="EFB44B5820181904FC288BA5D9F8F920" author="Simon" box="[1023,1208,1679,1703]" firstAuthor="Simon" page="83" pageId="74" pageNumber="73" pagination="80 - 84" refId="ref63953" refString="SIMON, E. 1892. Liste des Arachnides recueillis en Syrie par M. le docteur Theod. Barrois. Revue Biologique du Nord de la France, 5: 80 - 84." type="journal article" year="1892">Simon, 1892: 83</bibRefCitation>
</treatmentCitation>
(?);
<treatmentCitation id="0A8410B820181904FB268BA5DEF4F941" author="Davydov" page="41" pageId="74" pageNumber="73" year="1898">
<bibRefCitation id="EFB44B5820181904FB268BA5DEF4F941" author="Davydov" firstAuthor="Davydov" journalOrPublisher="Annuaire du Musee Zoologique de l'Academie Imperiale des Sciences de St. - Petersbourg" page="41 - 43" pageId="74" pageNumber="73" pagination="39 - 51" part="3" refId="ref59970" refString="DAVYDOV, K. N. 1898. (Notes on some scorpion species of the Palestinian fauna). Annuaire du Musee Zoologique de l'Academie Imperiale des Sciences de St. - Petersbourg, 3 (1 - 2): 39 - 51 (in Russian)." title="(Notes on some scorpion species of the Palestinian fauna)" type="journal article" year="1898">Davydov, 1898: 41–43</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B820181904FC168B84D9E2F941" author="SCHENKEL" box="[961,1186,1710,1734]" page="380" pageId="74" pageNumber="73" year="1932">
<bibRefCitation id="EFB44B5820181904FC168B84D9E2F941" author="Schenkel" box="[961,1186,1710,1734]" firstAuthor="Schenkel" page="380" pageId="74" pageNumber="73" pagination="375 - 396" refId="ref63734" refString="SCHENKEL, E. 1932. Notizen uber einige Scorpione und Solifugen. Revue Suisse de Zoologie, 39 (15): 375 - 396." type="journal article" year="1932">Schenkel, 1932: 380</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B820181904FB798B84D83AF941" author="WERNER" box="[1198,1402,1710,1734]" page="269" pageId="74" pageNumber="73" year="1934">
<bibRefCitation id="EFB44B5820181904FB798B84D83AF941" author="Werner" box="[1198,1402,1710,1734]" firstAuthor="Werner" page="269" pageId="74" pageNumber="73" pagination="8" refId="ref65242" refString="WERNER, F. 1934. Scorpiones, Pedipalpi. In: H. G. Bronns Klassen und Ordnungen des Tierreichs. Akademische Verlaggesellschaft, Leipzig. 5, IV, 8, Lief. 1 - 2, Scorpiones, pp. 1 - 316." type="journal article" year="1934">Werner, 1934: 269</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B820181904FC668BE7D9C0F963" author="WERNER" box="[945,1152,1740,1764]" page="211" pageId="74" pageNumber="73" year="1935">
<bibRefCitation id="EFB44B5820181904FC668BE7D9C0F963" author="Werner" box="[945,1152,1740,1764]" firstAuthor="Werner" page="211" pageId="74" pageNumber="73" pagination="211 - 216" refId="ref65290" refString="WERNER, F. 1935. Ueber Skorpione aus Palastina. Zoologisher Anzeiger, 109 (7 - 8): 211 - 216." type="journal article" year="1935">Werner, 1935: 211</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B820181904FB5B8BE6D8DBF963" author="Bodenheimer" box="[1164,1435,1740,1764]" page="235" pageId="74" pageNumber="73" year="1937">
<bibRefCitation id="EFB44B5820181904FB5B8BE6D8DBF963" author="Bodenheimer" box="[1164,1435,1740,1764]" firstAuthor="Bodenheimer" journalOrPublisher="Memoires Presentes a l'Institut d'Egypte" page="235" pageId="74" pageNumber="73" pagination="1 - 286" part="33" refId="ref59516" refString="BODENHEIMER, F. 1937. Prodromus Faunae Palestinae. Memoires Presentes a l'Institut d'Egypte, 33: 1 - 286." title="Prodromus Faunae Palestinae" type="journal article" year="1937">Bodenheimer, 1937: 235</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="EFB44B5820181904FCB88BC1D952F884" author="Whittick" box="[879,1042,1771,1795]" firstAuthor="Whittick" pageId="74" pageNumber="73" pagination="6" refId="ref65390" refString="WHITTICK, R. J. 1955. Scorpions from Palestine, Syria, Irak and Iran. In: H. Field (ed.), Contributions to the Fauna and Flora of Southwestern Asia. No 6. Henry Field, Coconut Grove, Florida." type="book chapter" year="1955">Whittick, 1955</bibRefCitation>
;
<bibRefCitation id="EFB44B5820181904FBCA8BC1D980F884" author="Whittick" box="[1053,1216,1771,1795]" firstAuthor="Whittick" pageId="74" pageNumber="73" refId="ref65439" refString="WHITTICK, R. 1970. Scorpions collected by Field Museum Near East expedition, 1934. Henry Field, Coconut Grove, Florida." type="book" year="1970">Whittick, 1970</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A36A920181904FCE88A20D82EF8A5" blockId="74.[831,1441,1464,1918]" box="[831,1390,1802,1826]" pageId="74" pageNumber="73">
<treatmentCitationGroup id="AB35118720181904FCE88A20D82EF8A5" box="[831,1390,1802,1826]" pageId="74" pageNumber="73">
<taxonomicName id="4C254D2A20181904FCE88A20D82AF8A5" authority=": Birula, 1910: 118" authorityName="Birula" authorityPageNumber="118" authorityYear="1910" box="[831,1386,1802,1826]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="74" pageNumber="73" phylum="Arthropoda" rank="subSpecies" species="quinquestriatus" subSpecies="hebraeus">
<emphasis id="B951EABB20181904FCE88A20D9DEF8A5" box="[831,1182,1802,1826]" italics="true" pageId="74" pageNumber="73">Buthus quinquestriatus hebraeus</emphasis>
:
<treatmentCitation id="0A8410B820181904FB7B8A20D82AF8A5" author="Birula" box="[1196,1386,1802,1826]" page="118" pageId="74" pageNumber="73" year="1910">
<bibRefCitation id="EFB44B5820181904FB7B8A20D82AF8A5" author="Birula" box="[1196,1386,1802,1826]" firstAuthor="Birula" journalOrPublisher="Horae Societatis Entomologicae Rossicae, St Petersburg" page="118" pageId="74" pageNumber="73" pagination="115 - 192" part="39" refId="ref59195" refString="BIRULA, A. 1910. Ueber Scorpio maurus Linne und seine Unterarten. Horae Societatis Entomologicae Rossicae, St Petersburg, 39: 115 - 192." title="Ueber Scorpio maurus Linne und seine Unterarten" type="journal article" year="1910">Birula, 1910: 118</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
<paragraph id="8B9A36A920181904FCE88A02D8D2F8C7" blockId="74.[831,1441,1464,1918]" box="[831,1426,1832,1856]" pageId="74" pageNumber="73">
<treatmentCitation id="0A8410B820181904FCE88A02D8D2F8C7" author="BIRULA, A." box="[831,1426,1832,1856]" page="170" pageId="74" pageNumber="73" year="1910">
<treatmentCitationGroup id="AB35118720181904FCE88A02D8CDF8C7" box="[831,1421,1832,1856]" pageId="74" pageNumber="73">
<taxonomicName id="4C254D2A20181904FCE88A02D8CDF8C7" authority="(Buthus)" authorityName="Birula" authorityPageNumber="170" authorityYear="1910" box="[831,1421,1832,1856]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="74" pageNumber="73" phylum="Arthropoda" rank="subGenus" subGenus="Buthus">
<emphasis id="B951EABB20181904FCE88A02DEC9F8C7" box="[831,905,1832,1856]" italics="true" pageId="74" pageNumber="73">Buthus</emphasis>
(
<emphasis id="B951EABB20181904FC4E8A02DEA3F8C7" box="[921,995,1832,1856]" italics="true" pageId="74" pageNumber="73">Buthus</emphasis>
) 5-
<emphasis id="B951EABB20181904FBDF8A03D981F8C7" box="[1032,1217,1832,1856]" italics="true" pageId="74" pageNumber="73">striatus hebraeus</emphasis>
:
<bibRefCitation id="EFB44B5820181904FB188A02D8CDF8C7" author="Birula" box="[1231,1421,1832,1856]" firstAuthor="Birula" journalOrPublisher="Horae Societatis Entomologicae Rossicae, St Petersburg" page="170" pageId="74" pageNumber="73" pagination="115 - 192" part="39" refId="ref59195" refString="BIRULA, A. 1910. Ueber Scorpio maurus Linne und seine Unterarten. Horae Societatis Entomologicae Rossicae, St Petersburg, 39: 115 - 192." title="Ueber Scorpio maurus Linne und seine Unterarten" type="journal article" year="1910">Birula, 1910: 170</bibRefCitation>
</taxonomicName>
</treatmentCitationGroup>
.
</treatmentCitation>
</paragraph>
<paragraph id="8B9A36A920181904FCE88A6DD840F8F9" blockId="74.[831,1441,1464,1918]" pageId="74" pageNumber="73">
<treatmentCitationGroup id="AB35118720181904FCE88A6DD840F8F9" pageId="74" pageNumber="73">
<treatmentCitation id="0A8410B820181904FCE88A6DD8E0F8D8" author="Birula" box="[831,1440,1863,1887]" page="23" pageId="74" pageNumber="73" year="1917">
<taxonomicName id="4C254D2A20181904FCE88A6DD82BF8D8" authority=": Birula, 1917 a: 23" authorityName=": Birula" authorityPageNumber="23" authorityYear="1917" box="[831,1387,1863,1887]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="74" pageNumber="73" phylum="Arthropoda" rank="species" species="quinquestriatus" subGenus="Buthus">
<emphasis id="B951EABB20181904FCE88A6DDEC9F8D8" box="[831,905,1863,1887]" italics="true" pageId="74" pageNumber="73">Buthus</emphasis>
(
<emphasis id="B951EABB20181904FC4D8A6DDEA4F8D8" box="[922,996,1863,1887]" italics="true" pageId="74" pageNumber="73">Buthus</emphasis>
)
<emphasis id="B951EABB20181904FC218A62D9DCF8D8" box="[1014,1180,1864,1887]" italics="true" pageId="74" pageNumber="73">quinquestriatus</emphasis>
:
<bibRefCitation id="EFB44B5820181904FB7C8A6DD82BF8D8" author="Birula" bookContentInfo="253 pp." box="[1195,1387,1863,1887]" firstAuthor="Birula" journalOrPublisher="Zapiski Kavkazskogo Muzeya (Memoires du Musee du Caucase), Tiflis, Imprimerie de la Chancellerie du Comite pour la Transcaucasie. Ser. A" page="23" pageId="74" pageNumber="73" part="5" refId="ref59223" refString="BIRULA, A. A. 1917 a. Arachnoidea Arthrogastra Caucasica. Pars I. Scorpiones. Zapiski Kavkazskogo Muzeya (Memoires du Musee du Caucase), Tiflis, Imprimerie de la Chancellerie du Comite pour la Transcaucasie. Ser. A, 5, 253 pp. (in Russian; published August 1917). English translation: Byalynitskii-Birulya, A. A. 1964. Arthrogastric Arachnids of Caucasia. 1. Scorpions. Israel Program for Scientific Translations, Jerusalem, 170 pp." title="Arachnoidea Arthrogastra Caucasica. Pars I. Scorpiones" type="book" year="1917" yearSuffix="a">Birula, 1917a: 23</bibRefCitation>
</taxonomicName>
, 213
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B820181904FC048A4CD9DCF8F9" author="WHITTICK" box="[979,1180,1894,1918]" page="43" pageId="74" pageNumber="73" year="1941">
<bibRefCitation id="EFB44B5820181904FC048A4CD9DCF8F9" author="Whittick" box="[979,1180,1894,1918]" firstAuthor="Whittick" page="43" pageId="74" pageNumber="73" pagination="43 - 49" refId="ref65348" refString="WHITTICK, R. J. 1941. Arachnida: Scorpiones, Pedipalpi and Solifugae. In: Expedition to South-West Arabia, 1937 - 1938. Bulletin of the British Museum, Natural History, 1: 43 - 49." type="journal article" year="1941">Whittick, 1941: 43</bibRefCitation>
</treatmentCitation>
(in part).
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</paragraph>
<paragraph id="8B9A36A920191905FCC68DE2D9A4FD01" pageId="75" pageNumber="74">
<table id="F925C4092019E6B1FEE88DE2D843FD01" box="[319,1283,200,646]" gridcols="4" gridrows="14" pageId="75" pageNumber="74">
<tr id="351534EB2019E6B1FEE88DE2D843FF67" box="[319,1283,200,224]" gridrow="0" pageId="75" pageNumber="74" rowspan-0="1">
<th id="76C45D972019E6B1FCC68DE2DECDFF67" box="[785,909,200,224]" gridcol="1" gridrow="0" pageId="75" pageNumber="74">
<typeStatus id="549E880B20191905FCC68DE2DE39FF67" box="[785,889,200,224]" pageId="75" pageNumber="74" type="paratype">Paratype</typeStatus>
1
</th>
<th id="76C45D972019E6B1FC068DE2D90DFF67" box="[977,1101,200,224]" gridcol="2" gridrow="0" pageId="75" pageNumber="74">
<specimenCount id="9D23FD2020191905FC068DE2D979FF67" box="[977,1081,200,224]" pageId="75" pageNumber="74" type="generic" typeStatus="Paratype">Paratype</specimenCount>
2
</th>
<th id="76C45D972019E6B1FB4C8DE2D843FF67" box="[1179,1283,200,224]" gridcol="3" gridrow="0" pageId="75" pageNumber="74">
<emphasis id="B951EABB20191905FB4C8DE2D843FF67" bold="true" box="[1179,1283,200,224]" pageId="75" pageNumber="74">
<specimenCount id="9D23FD2020191905FB4C8DE2D843FF67" box="[1179,1283,200,224]" pageId="75" pageNumber="74" type="generic" typeStatus="Holotype">Holotype</specimenCount>
</emphasis>
</th>
</tr>
<tr id="351534EB2019E6B1FEE88DD9D843FE8E" box="[319,1283,243,265]" gridrow="1" pageId="75" pageNumber="74">
<th id="76C45D972019E6B1FEE88DD9DF22FE8E" box="[319,610,243,265]" gridcol="0" gridrow="1" pageId="75" pageNumber="74">Carapace L</th>
<td id="76C45D972019E6B1FCC68DD9DECDFE8E" box="[785,909,243,265]" gridcol="1" gridrow="1" pageId="75" pageNumber="74">6.30</td>
<td id="76C45D972019E6B1FC068DD9D90DFE8E" box="[977,1101,243,265]" gridcol="2" gridrow="1" pageId="75" pageNumber="74">8.68</td>
<td id="76C45D972019E6B1FB4C8DD9D843FE8E" box="[1179,1283,243,265]" gridcol="3" gridrow="1" pageId="75" pageNumber="74">8.79</td>
</tr>
<tr id="351534EB2019E6B1FEE88C39D843FEAE" box="[319,1283,275,297]" gridrow="2" pageId="75" pageNumber="74">
<th id="76C45D972019E6B1FEE88C39DF22FEAE" box="[319,610,275,297]" gridcol="0" gridrow="2" pageId="75" pageNumber="74">Pedipalp femur L/W</th>
<td id="76C45D972019E6B1FCC68C39DECDFEAE" box="[785,909,275,297]" gridcol="1" gridrow="2" pageId="75" pageNumber="74">
<geoCoordinate id="EE11506E20191905FCED8C39DE24FEAE" box="[826,868,275,297]" degrees="3.86" direction="west" orientation="longitude" pageId="75" pageNumber="74" precision="555" value="-3.86">3.86</geoCoordinate>
</td>
<td id="76C45D972019E6B1FC068C39D90DFEAE" box="[977,1101,275,297]" gridcol="2" gridrow="2" pageId="75" pageNumber="74">4.28</td>
<td id="76C45D972019E6B1FB4C8C39D843FEAE" box="[1179,1283,275,297]" gridcol="3" gridrow="2" pageId="75" pageNumber="74">4.30</td>
</tr>
<tr id="351534EB2019E6B1FEE88C19D843FECE" box="[319,1283,307,329]" gridrow="3" pageId="75" pageNumber="74">
<th id="76C45D972019E6B1FEE88C19DF22FECE" box="[319,610,307,329]" gridcol="0" gridrow="3" pageId="75" pageNumber="74">Pedipalp patella L/W</th>
<td id="76C45D972019E6B1FCC68C19DECDFECE" box="[785,909,307,329]" gridcol="1" gridrow="3" pageId="75" pageNumber="74">
<geoCoordinate id="EE11506E20191905FCED8C19DE24FECE" box="[826,868,307,329]" degrees="3.52" direction="west" orientation="longitude" pageId="75" pageNumber="74" precision="555" value="-3.52">3.52</geoCoordinate>
</td>
<td id="76C45D972019E6B1FC068C19D90DFECE" box="[977,1101,307,329]" gridcol="2" gridrow="3" pageId="75" pageNumber="74">4.28</td>
<td id="76C45D972019E6B1FB4C8C19D843FECE" box="[1179,1283,307,329]" gridcol="3" gridrow="3" pageId="75" pageNumber="74">4.48</td>
</tr>
<tr id="351534EB2019E6B1FEE88C78D843FEEF" box="[319,1283,338,360]" gridrow="4" pageId="75" pageNumber="74">
<th id="76C45D972019E6B1FEE88C78DF22FEEF" box="[319,610,338,360]" gridcol="0" gridrow="4" pageId="75" pageNumber="74">Pedipalp chela L/manus W</th>
<td id="76C45D972019E6B1FCC68C78DECDFEEF" box="[785,909,338,360]" gridcol="1" gridrow="4" pageId="75" pageNumber="74">
<geoCoordinate id="EE11506E20191905FCED8C78DE24FEEF" box="[826,868,338,360]" degrees="7.74" direction="west" orientation="longitude" pageId="75" pageNumber="74" precision="555" value="-7.74">7.74</geoCoordinate>
</td>
<td id="76C45D972019E6B1FC068C78D90DFEEF" box="[977,1101,338,360]" gridcol="2" gridrow="4" pageId="75" pageNumber="74">7.76</td>
<td id="76C45D972019E6B1FB4C8C78D843FEEF" box="[1179,1283,338,360]" gridcol="3" gridrow="4" pageId="75" pageNumber="74">8.24</td>
</tr>
<tr id="351534EB2019E6B1FEE88C58D843FE0F" box="[319,1283,370,392]" gridrow="5" pageId="75" pageNumber="74">
<th id="76C45D972019E6B1FEE88C58DF22FE0F" box="[319,610,370,392]" gridcol="0" gridrow="5" pageId="75" pageNumber="74">Leg III femur L/ Carapace L</th>
<td id="76C45D972019E6B1FCC68C58DECDFE0F" box="[785,909,370,392]" gridcol="1" gridrow="5" pageId="75" pageNumber="74">1.15</td>
<td id="76C45D972019E6B1FC068C58D90DFE0F" box="[977,1101,370,392]" gridcol="2" gridrow="5" pageId="75" pageNumber="74">1.16</td>
<td id="76C45D972019E6B1FB4C8C58D843FE0F" box="[1179,1283,370,392]" gridcol="3" gridrow="5" pageId="75" pageNumber="74">1.27</td>
</tr>
<tr id="351534EB2019E6B1FEE88CB8D843FE2F" box="[319,1283,402,424]" gridrow="6" pageId="75" pageNumber="74">
<th id="76C45D972019E6B1FEE88CB8DF22FE2F" box="[319,610,402,424]" gridcol="0" gridrow="6" pageId="75" pageNumber="74">Leg III patella L/ Carapace L</th>
<td id="76C45D972019E6B1FCC68CB8DECDFE2F" box="[785,909,402,424]" gridcol="1" gridrow="6" pageId="75" pageNumber="74">0.97</td>
<td id="76C45D972019E6B1FC068CB8D90DFE2F" box="[977,1101,402,424]" gridcol="2" gridrow="6" pageId="75" pageNumber="74">0.99</td>
<td id="76C45D972019E6B1FB4C8CB8D843FE2F" box="[1179,1283,402,424]" gridcol="3" gridrow="6" pageId="75" pageNumber="74">1.07</td>
</tr>
<tr id="351534EB2019E6B1FEE88C98D843FE4F" box="[319,1283,434,456]" gridrow="7" pageId="75" pageNumber="74">
<th id="76C45D972019E6B1FEE88C98DF22FE4F" box="[319,610,434,456]" gridcol="0" gridrow="7" pageId="75" pageNumber="74">Leg III patella L/D</th>
<td id="76C45D972019E6B1FCC68C98DECDFE4F" box="[785,909,434,456]" gridcol="1" gridrow="7" pageId="75" pageNumber="74">4.23</td>
<td id="76C45D972019E6B1FC068C98D90DFE4F" box="[977,1101,434,456]" gridcol="2" gridrow="7" pageId="75" pageNumber="74">4.35</td>
<td id="76C45D972019E6B1FB4C8C98D843FE4F" box="[1179,1283,434,456]" gridcol="3" gridrow="7" pageId="75" pageNumber="74">4.54</td>
</tr>
<tr id="351534EB2019E6B1FEE88CFBD843FE60" box="[319,1283,465,487]" gridrow="8" pageId="75" pageNumber="74">
<th id="76C45D972019E6B1FEE88CFBDF22FE60" box="[319,610,465,487]" gridcol="0" gridrow="8" pageId="75" pageNumber="74">Pedipalp femur L/ Carapace L</th>
<td id="76C45D972019E6B1FCC68CFBDECDFE60" box="[785,909,465,487]" gridcol="1" gridrow="8" pageId="75" pageNumber="74">1.08</td>
<td id="76C45D972019E6B1FC068CFBD90DFE60" box="[977,1101,465,487]" gridcol="2" gridrow="8" pageId="75" pageNumber="74">1.09</td>
<td id="76C45D972019E6B1FB4C8CFBD843FE60" box="[1179,1283,465,487]" gridcol="3" gridrow="8" pageId="75" pageNumber="74">1.14</td>
</tr>
<tr id="351534EB2019E6B1FEE88CDBD843FD80" box="[319,1283,497,519]" gridrow="9" pageId="75" pageNumber="74">
<th id="76C45D972019E6B1FEE88CDBDF22FD80" box="[319,610,497,519]" gridcol="0" gridrow="9" pageId="75" pageNumber="74">Metasoma I L/W</th>
<td id="76C45D972019E6B1FCC68CDBDECDFD80" box="[785,909,497,519]" gridcol="1" gridrow="9" pageId="75" pageNumber="74">1.37</td>
<td id="76C45D972019E6B1FC068CDBD90DFD80" box="[977,1101,497,519]" gridcol="2" gridrow="9" pageId="75" pageNumber="74">1.41</td>
<td id="76C45D972019E6B1FB4C8CDBD843FD80" box="[1179,1283,497,519]" gridcol="3" gridrow="9" pageId="75" pageNumber="74">1.41</td>
</tr>
<tr id="351534EB2019E6B1FEE88F3BD843FDA0" box="[319,1283,529,551]" gridrow="10" pageId="75" pageNumber="74">
<th id="76C45D972019E6B1FEE88F3BDF22FDA0" box="[319,610,529,551]" gridcol="0" gridrow="10" pageId="75" pageNumber="74">
Metasoma
<collectionCode id="ED34AE6C20191905FE7E8F3BDCF9FDA1" box="[425,441,529,550]" pageId="75" pageNumber="74">II</collectionCode>
L/W
</th>
<td id="76C45D972019E6B1FCC68F3BDECDFDA0" box="[785,909,529,551]" gridcol="1" gridrow="10" pageId="75" pageNumber="74">1.87</td>
<td id="76C45D972019E6B1FC068F3BD90DFDA0" box="[977,1101,529,551]" gridcol="2" gridrow="10" pageId="75" pageNumber="74">1.95</td>
<td id="76C45D972019E6B1FB4C8F3BD843FDA0" box="[1179,1283,529,551]" gridcol="3" gridrow="10" pageId="75" pageNumber="74">1.91</td>
</tr>
<tr id="351534EB2019E6B1FEE88F1AD843FDC1" box="[319,1283,560,582]" gridrow="11" pageId="75" pageNumber="74">
<th id="76C45D972019E6B1FEE88F1ADF22FDC1" box="[319,610,560,582]" gridcol="0" gridrow="11" pageId="75" pageNumber="74">
Metasoma
<collectionCode id="ED34AE6C20191905FE7E8F1BDC81FDC1" box="[425,449,561,582]" pageId="75" pageNumber="74">III</collectionCode>
L/W
</th>
<td id="76C45D972019E6B1FCC68F1ADECDFDC1" box="[785,909,560,582]" gridcol="1" gridrow="11" pageId="75" pageNumber="74">2.00</td>
<td id="76C45D972019E6B1FC068F1AD90DFDC1" box="[977,1101,560,582]" gridcol="2" gridrow="11" pageId="75" pageNumber="74">2.19</td>
<td id="76C45D972019E6B1FB4C8F1AD843FDC1" box="[1179,1283,560,582]" gridcol="3" gridrow="11" pageId="75" pageNumber="74">2.30</td>
</tr>
<tr id="351534EB2019E6B1FEE88F7AD843FDE1" box="[319,1283,592,614]" gridrow="12" pageId="75" pageNumber="74">
<th id="76C45D972019E6B1FEE88F7ADF22FDE1" box="[319,610,592,614]" gridcol="0" gridrow="12" pageId="75" pageNumber="74">Metasoma IV L/W</th>
<td id="76C45D972019E6B1FCC68F7ADECDFDE1" box="[785,909,592,614]" gridcol="1" gridrow="12" pageId="75" pageNumber="74">2.50</td>
<td id="76C45D972019E6B1FC068F7AD90DFDE1" box="[977,1101,592,614]" gridcol="2" gridrow="12" pageId="75" pageNumber="74">2.70</td>
<td id="76C45D972019E6B1FB4C8F7AD843FDE1" box="[1179,1283,592,614]" gridcol="3" gridrow="12" pageId="75" pageNumber="74">2.94</td>
</tr>
<tr id="351534EB2019E6B1FEE88F5AD843FD01" box="[319,1283,624,646]" gridrow="13" pageId="75" pageNumber="74">
<th id="76C45D972019E6B1FEE88F5ADF22FD01" box="[319,610,624,646]" gridcol="0" gridrow="13" pageId="75" pageNumber="74">
Metasoma
<collectionCode id="ED34AE6C20191905FE7E8F5BDCFBFD01" box="[425,443,625,646]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="75" pageNumber="74" type="Museum">V</collectionCode>
L/W
</th>
<td id="76C45D972019E6B1FCC68F5ADECDFD01" box="[785,909,624,646]" gridcol="1" gridrow="13" pageId="75" pageNumber="74">2.75</td>
<td id="76C45D972019E6B1FC068F5AD90DFD01" box="[977,1101,624,646]" gridcol="2" gridrow="13" pageId="75" pageNumber="74">3.11</td>
<td id="76C45D972019E6B1FB4C8F5AD843FD01" box="[1179,1283,624,646]" gridcol="3" gridrow="13" pageId="75" pageNumber="74">3.23</td>
</tr>
</table>
</paragraph>
<caption id="DF5A662120191905FF178F80D909FD46" ID-Table-UUID="DF5A662120191905FF178F80D909FD46" box="[192,1097,682,706]" httpUri="http://table.plazi.org/id/DF5A662120191905FF178F80D909FD46" pageId="75" pageNumber="74" startId="75.[192,257,682,706]" targetBox="[319,1283,200,646]" targetIsTable="true" targetPageId="75">
<paragraph id="8B9A36A920191905FF178F80D909FD46" blockId="75.[192,1097,682,706]" box="[192,1097,682,706]" pageId="75" pageNumber="74">
<emphasis id="B951EABB20191905FF178F80DC5EFD45" bold="true" box="[192,286,682,706]" pageId="75" pageNumber="74">Table 2:</emphasis>
Variation in male morphometric ratios with increasing size in
<taxonomicName id="4C254D2A20191905FCAE8F86D94DFD46" ID-CoL="6P795" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" box="[889,1037,683,705]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="75" pageNumber="74" phylum="Arthropoda" rank="species" species="heberti" status="sp. nov.">
<emphasis id="B951EABB20191905FCAE8F86D94DFD46" box="[889,1037,683,705]" italics="true" pageId="75" pageNumber="74">Leiurus heberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C020191905FBC48F86D909FD46" box="[1043,1097,684,705]" pageId="75" pageNumber="74" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8B9A36A920191905FF178FDDDFB5FBD8" blockId="75.[192,802,758,1456]" pageId="75" pageNumber="74">
<treatmentCitationGroup id="AB35118720191905FF178FDDDFB5FBD8" pageId="75" pageNumber="74">
<treatmentCitation id="0A8410B820191905FF178FDDDFF5FC89" author="VACHON" box="[192,693,758,782]" page="197" pageId="75" pageNumber="74" year="1950">
<taxonomicName id="4C254D2A20191905FF178FDDDFF5FC89" ID-CoL="6P78R" authority=": Vachon, 1950 a: 197" authorityName="Vachon" authorityPageNumber="197" authorityYear="1950" box="[192,693,758,782]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="75" pageNumber="74" phylum="Arthropoda" rank="species" species="quinquestriatus">
<emphasis id="B951EABB20191905FF178FDDDC82FC89" box="[192,450,759,782]" italics="true" pageId="75" pageNumber="74">Leiurus quinquestriatus</emphasis>
:
<bibRefCitation id="EFB44B5820191905FE028FDCDFF5FC89" author="Vachon" box="[469,693,758,782]" firstAuthor="Vachon" page="197" pageId="75" pageNumber="74" pagination="152 - 216" refId="ref64629" refString="VACHON, M. 1950 a. Etudes sur les Scorpions. III (suite). Description des Scorpions du Nord de l'Afrique. Archives de l'Institut Pasteur d'Algerie, 28 (2): 152 - 216." type="journal article" year="1950" yearSuffix="a">Vachon, 1950a: 197</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B820191905FF278E3EDC83FCAB" author="STAHNKE" box="[240,451,788,812]" page="130" pageId="75" pageNumber="74" year="1972">
<bibRefCitation id="EFB44B5820191905FF278E3EDC83FCAB" author="Stahnke" box="[240,451,788,812]" firstAuthor="Stahnke" page="130" pageId="75" pageNumber="74" pagination="121 - 133" refId="ref64369" refString="STAHNKE, H. L. 1972. A key to the genera of Buthidae (Scorpionida). Entomological News, 83 (5): 121 - 133." type="journal article" year="1972">Stahnke, 1972: 130</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B820191905FDE18E3EDCD1FCCC" author="LAMORAL, B. H. &amp; S. C. REYNDERS" page="509" pageId="75" pageNumber="74" year="1975">
<bibRefCitation id="EFB44B5820191905FDE18E3EDCD1FCCC" author="Lamoral &amp; Reynders" firstAuthor="Lamoral" journalOrPublisher="Annals of the Natal Museum" page="509 - 510" pageId="75" pageNumber="74" pagination="489 - 576" part="22" refId="ref62059" refString="LAMORAL, B. H. &amp; S. C. REYNDERS. 1975. A catalogue of the scorpions described from the Ethiopian faunal region up to December 1973. Annals of the Natal Museum, 22 (2): 489 - 576." title="A catalogue of the scorpions described from the Ethiopian faunal region up to December 1973" type="journal article" year="1975">Lamoral &amp; Reynders, 1975: 509–510</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B820191905FE2A8E19DF85FCCC" author="WAHBEH" box="[509,709,819,843]" page="89" pageId="75" pageNumber="74" year="1976">
<bibRefCitation id="EFB44B5820191905FE2A8E19DF85FCCC" author="Wahbeh" box="[509,709,819,843]" firstAuthor="Wahbeh" page="89" pageId="75" pageNumber="74" pagination="84 - 92" refId="ref65138" refString="WAHBEH, Y. 1976. A study of Jordanian scorpions. Jordan Medical Journal, 11 (2): 84 - 92." type="journal article" year="1976">Wahbeh, 1976: 89</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B820191905FD078E1EDC90FCED" author="LEVY, G. &amp; P. AMITAI" page="47" pageId="75" pageNumber="74" year="1980">
<bibRefCitation id="EFB44B5820191905FD078E1EDC90FCED" author="Levy &amp; Amitai" bookContentInfo="130 pp" firstAuthor="Levy" journalOrPublisher="The Israel Academy of Sciences and Humanities. Jerusalem" page="47 - 48" pageId="75" pageNumber="74" refId="ref62185" refString="LEVY, G. &amp; P. AMITAI. 1980. Scorpiones. Fauna Palaestina. Arachnida I. The Israel Academy of Sciences and Humanities. Jerusalem, 130 pp .." title="Scorpiones. Fauna Palaestina. Arachnida I" type="book" year="1980">Levy &amp; Amitai, 1980: 47–48</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B820191905FDEB8E78DC5BFC0F" author="WARBURG" page="206" pageId="75" pageNumber="74" year="1980">
<bibRefCitation id="EFB44B5820191905FDEB8E78DC5BFC0F" author="Warburg" etAl="et al." firstAuthor="Warburg" page="206" pageId="75" pageNumber="74" pagination="205 - 213" refId="ref65163" refString="WARBURG, M. R., S. GOLDENBERG &amp; A. BEN- HORIN. 1980. Scorpion species diversity and distribution within the Mediterranean and arid regions of northern Israel. Journal of Arid Environments, 3: 205 - 213." type="journal article" year="1980">Warburg et al., 1980: 206</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B820191905FEF88E5ADF75FC0F" author="Kinzelbach" box="[303,565,880,904]" page="100" pageId="75" pageNumber="74" year="1984">
<bibRefCitation id="EFB44B5820191905FEF88E5ADF75FC0F" author="Kinzelbach" box="[303,565,880,904]" firstAuthor="Kinzelbach" journalOrPublisher="Mainzer Naturwisserschaftiches Archiv" page="100" pageId="75" pageNumber="74" pagination="97 - 106" part="22" refId="ref61492" refString="KINZELBACH, R. 1984. Die Skorpionssammlung des Naturhistorischen Museums der Stadt Mainz. - Teil II: Vorderasien. Mainzer Naturwisserschaftiches Archiv, 22: 97 - 106." title="Die Skorpionssammlung des Naturhistorischen Museums der Stadt Mainz. - Teil II: Vorderasien" type="journal article" year="1984">Kinzelbach, 1984: 100</bibRefCitation>
</treatmentCitation>
,
<figureCitation id="131E2A2C20191905FD9E8E5ADFCCFC0F" box="[585,652,880,904]" captionStart="Figure 2" captionStartId="8.[192,267,1666,1690]" captionTargetBox="[201,1405,193,1633]" captionTargetId="figure-120@8.[202,1404,194,1632]" captionTargetPageId="8" captionText="Figure 2: Leiurus brachycentrus (Ehrenberg, 1829) stat. n., male. Habitus, ventral aspect. Ad Darb, Saudi Arabia (NHMB 17ag)." figureDoi="http://doi.org/10.5281/zenodo.6505626" httpUri="https://zenodo.org/record/6505626/files/figure.png" pageId="75" pageNumber="74">Fig 2</figureCitation>
;
<bibRefCitation id="EFB44B5820191905FD768E5ADC6AFC20" author="Kinzelbach" firstAuthor="Kinzelbach" journalOrPublisher="Tubinger Atlas des Vorderen Orients (TAVO). Karte A VI 14.2. Tubingen" pageId="75" pageNumber="74" refId="ref61523" refString="KINZELBACH, R. 1985. Vorder Orient. Skorpione (Arachnida: Scorpiones). Tubinger Atlas des Vorderen Orients (TAVO). Karte A VI 14.2. Tubingen." title="Vorder Orient. Skorpione (Arachnida: Scorpiones)" type="book" year="1985">Kinzelbach, 1985</bibRefCitation>
: map II (in part);
<treatmentCitation id="0A8410B820191905FE2D8EA5DFB1FC20" author="AMR, Z. S. &amp; K. E. HILAND &amp; R. KINZELBACH &amp; S. S. AMR &amp; D. DEFOSSE" box="[506,753,911,935]" page="373" pageId="75" pageNumber="74" year="1988">
<bibRefCitation id="EFB44B5820191905FE2D8EA5DFB1FC20" author="AMR, Z. S. &amp; K. E. HILAND &amp; R. KINZELBACH &amp; S. S. AMR &amp; D. DEFOSSE" box="[506,753,911,935]" etAl="et al." firstAuthor="Amr" journalOrPublisher="Bulletin de la Societe de Pathologie Exotique" page="373" pageId="75" pageNumber="74" pagination="369 - 379" part="81" refId="ref58889" refString="AMR, Z. S., K. E. HILAND, R. KINZELBACH, S. S. AMR &amp; D. DEFOSSE. 1988. Scorpions et piqures de scorpions en Jordanie. Bulletin de la Societe de Pathologie Exotique, 81: 369 - 379." title="Scorpions et piqures de scorpions en Jordanie" type="journal article" year="1988">Amr et al., 1988: 373</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B820191905FCD68EA5DF72FC41" author="EL-HENNAWY, H. K." page="14" pageId="75" pageNumber="74" year="1988">
<bibRefCitation id="EFB44B5820191905FCD68EA5DCE7FC41" author="EL-HENNAWY, H. K." journalOrPublisher="Serket" pageId="75" pageNumber="74" pagination="13 - 20" part="1" refId="ref60078" refString="EL-HENNAWY, H. K. 1988. Scorpions of Jordan. Serket, 1 (2): 13 - 20." title="Scorpions of Jordan" type="journal article" year="1988">El- Hennawy, 1988</bibRefCitation>
: 14–15, 18
</treatmentCitation>
;
<treatmentCitation id="0A8410B820191905FD908E84DCC4FC63" author="EL-HENNAWY, H. K." page="101" pageId="75" pageNumber="74" year="1992">
<bibRefCitation id="EFB44B5820191905FD908E84DC5AFC63" author="El-Hennawy" firstAuthor="El-Hennawy" journalOrPublisher="Serket" page="101" pageId="75" pageNumber="74" pagination="95 - 153" part="2" refId="ref60101" refString="EL-HENNAWY, H. K. 1992. A catalogue of the scorpions described from the Arab Countries (1758 - 1990) (Arachnida: Scorpionida). Serket, 2 (4): 95 - 153." title="A catalogue of the scorpions described from the Arab Countries (1758 - 1990) (Arachnida: Scorpionida)" type="journal article" year="1992">El-Hennawy, 1992: 101</bibRefCitation>
, 125–126
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B820191905FE248EE6DE5DFC63" author="Amr &amp; El-Oran" box="[499,797,972,996]" page="186" pageId="75" pageNumber="74" year="1994">
<bibRefCitation id="EFB44B5820191905FE248EE6DE5DFC63" author="Amr &amp; El-Oran" box="[499,797,972,996]" firstAuthor="Amr" journalOrPublisher="Bolletino di zoologia" page="186" pageId="75" pageNumber="74" pagination="185 - 190" part="61" refId="ref58850" refString="AMR, Z. S. &amp; R. EL-ORAN. 1994. Systematics and distribution of scorpions (Arachnida, Scorpionida) in Jordan. Bolletino di zoologia, 61 (2): 185 - 190." title="Systematics and distribution of scorpions (Arachnida, Scorpionida) in Jordan" type="journal article" year="1994">Amr &amp; El-Oran, 1994: 186</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B820191905FF278EC1DC84FB84" author="KOVARIK, F." box="[240,452,1003,1027]" page="112" pageId="75" pageNumber="74" year="1998">
<bibRefCitation id="EFB44B5820191905FF278EC1DC84FB84" author="Kovarik" bookContentInfo="175 pp." box="[240,452,1003,1027]" firstAuthor="Kovarik" journalOrPublisher="Madagaskar, Jihlava" page="112" pageId="75" pageNumber="74" refId="ref61762" refString="KOVARIK, F. 1998. Stiri. Madagaskar, Jihlava, 175 pp." title="Stiri" type="book" year="1998">Kovařík, 1998: 112</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B820191905FDE28EC1DC4DFBA5" author="KABAKIBI, M. M. &amp; N. KHALIL &amp; Z. AMR" page="80" pageId="75" pageNumber="74" year="1999">
<bibRefCitation id="EFB44B5820191905FDE28EC1DC4DFBA5" author="KABAKIBI, M. M. &amp; N. KHALIL &amp; Z. AMR" etAl="et al." firstAuthor="Kabakibi" journalOrPublisher="Zoology in the Middle East" page="80" pageId="75" pageNumber="74" pagination="79 - 89" part="17" refId="ref61234" refString="KABAKIBI, M. M., N. KHALIL &amp; Z. AMR. 1999. Scorpions of southern Syria. Zoology in the Middle East, 17: 79 - 89." title="Scorpions of southern Syria" type="journal article" year="1999">Kabakibi et al., 1999: 80</bibRefCitation>
</treatmentCitation>
, fig. 1;
<treatmentCitation id="0A8410B820191905FEB38920DFA5FBA5" author="Amr &amp; Abu Baker" box="[356,741,1034,1058]" page="237" pageId="75" pageNumber="74" year="2004">
<bibRefCitation id="EFB44B5820191905FEB38920DFA5FBA5" author="Amr &amp; Abu Baker" box="[356,741,1034,1058]" firstAuthor="Amr" journalOrPublisher="Denisia" page="237 - 238" pageId="75" pageNumber="74" pagination="237 - 244" part="14" refId="ref58822" refString="AMR, Z. &amp; M. ABU BAKER. 2004. The scorpions of Jordan. Denisia, 14 (2): 237 - 244." title="The scorpions of Jordan" type="journal article" year="2004">Amr &amp; Abu Baker, 2004: 237–238</bibRefCitation>
</treatmentCitation>
, 241;
<treatmentCitation id="0A8410B820191905FF278902DF5EFBC7" author="YAGMUR" box="[240,542,1064,1088]" page="2" pageId="75" pageNumber="74" year="2009">
<bibRefCitation id="EFB44B5820191905FF278902DF5EFBC7" author="Yagmur" box="[240,542,1064,1088]" etAl="et al." firstAuthor="Yagmur" page="2 - 3" pageId="75" pageNumber="74" pagination="1 - 20" refId="ref65464" refString="YAGMUR, E. A., H. KOC &amp; K. B. KUNT. 2009. Des- cription of a new species of Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) from Southeastern Turkey. Euscorpius, 85: 1 - 20." type="journal article" year="2009">Yağmur et al., 2009: 2–3</bibRefCitation>
</treatmentCitation>
(in part), fig. 10–11;
<treatmentCitation id="0A8410B820191905FF27896DDF69FBD8" author="SHEHAB" box="[240,553,1095,1119]" page="335" pageId="75" pageNumber="74" year="2011">
<bibRefCitation id="EFB44B5820191905FF27896DDF69FBD8" author="Shehab" box="[240,553,1095,1119]" etAl="et al." firstAuthor="Shehab" page="335 - 337" pageId="75" pageNumber="74" pagination="333 - 341" refId="ref63761" refString="SHEHAB, A. H., Z. S. AMR &amp; J. A. LINDSELL. 2011. Ecology and biology of scorpions in Palmyra, Syria. Turkish Journal of Zoology, 35 (3): 333 - 341." type="journal article" year="2011">Shehab et al., 2011: 335–337</bibRefCitation>
</treatmentCitation>
, fig. 2D, tabs. 1–2.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A36A920191905FF17894DDE5CFA90" blockId="75.[192,802,758,1456]" pageId="75" pageNumber="74">
<treatmentCitationGroup id="AB35118720191905FF17894DDE5CFA90" pageId="75" pageNumber="74">
<treatmentCitation id="0A8410B820191905FF17894DDE5DFBF9" author="VACHON" box="[192,797,1126,1150]" page="212" pageId="75" pageNumber="74" year="1966">
<taxonomicName id="4C254D2A20191905FF17894DDE5DFBF9" ID-CoL="6P6WQ" authority=": Vachon, 1966: 212" authorityName="Vachon" authorityPageNumber="212" authorityYear="1966" box="[192,797,1126,1150]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="75" pageNumber="74" phylum="Arthropoda" rank="subSpecies" species="quinquestriatus" subSpecies="hebraeus">
<emphasis id="B951EABB20191905FF17894DDF71FBF9" box="[192,561,1126,1150]" italics="true" pageId="75" pageNumber="74">Leiurus quinquestriatus hebraeus</emphasis>
:
<bibRefCitation id="EFB44B5820191905FD93894CDE5DFBF9" author="Vachon" box="[580,797,1126,1150]" firstAuthor="Vachon" page="212" pageId="75" pageNumber="74" pagination="209 - 218" refId="ref64772" refString="VACHON, M. 1966. Liste des Scorpions connus en Egypte, Arabie, Israel, Liban, Syrie, Jordanie, Turquie, Irak, Iran. Toxicon, 4: 209 - 218." type="journal article" year="1966">Vachon, 1966: 212</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0A8410B820191905FF2789AFDFF3FB1A" author="Levy, Amitai &amp; Shulov" box="[240,691,1157,1181]" page="231" pageId="75" pageNumber="74" year="1970">
<bibRefCitation id="EFB44B5820191905FF2789AFDFF3FB1A" author="Levy, Amitai &amp; Shulov" box="[240,691,1157,1181]" firstAuthor="Levy" journalOrPublisher="Israel Journal of Zoology" page="231 - 242" pageId="75" pageNumber="74" pagination="231 - 242" part="19" refId="ref62217" refString="LEVY, G., P. AMITAI &amp; A. SHULOV. 1970. Leiurus quinquestriatus hebraeus (Birula, 1908) (Scorpiones, Buthidae) and its systematic position. Israel Journal of Zoology, 19 (4): 231 - 242." title="Leiurus quinquestriatus hebraeus (Birula, 1908) (Scorpiones, Buthidae) and its systematic position" type="journal article" year="1970">Levy, Amitai &amp; Shulov, 1970: 231–242</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B820191905FF27898EDF7EFB3C" author="Levy &amp; Amitai" box="[240,574,1187,1212]" page="48" pageId="75" pageNumber="74" year="1980">
<bibRefCitation id="EFB44B5820191905FF27898EDF7EFB3C" author="Levy &amp; Amitai" bookContentInfo="130 pp" box="[240,574,1187,1212]" firstAuthor="Levy" journalOrPublisher="The Israel Academy of Sciences and Humanities. Jerusalem" page="48 - 53" pageId="75" pageNumber="74" refId="ref62185" refString="LEVY, G. &amp; P. AMITAI. 1980. Scorpiones. Fauna Palaestina. Arachnida I. The Israel Academy of Sciences and Humanities. Jerusalem, 130 pp .." title="Scorpiones. Fauna Palaestina. Arachnida I" type="book" year="1980">Levy &amp; Amitai, 1980: 48–53</bibRefCitation>
</treatmentCitation>
, fig. 47–51, map 3, Appendix (in part);
<treatmentCitation id="0A8410B820191905FE0289E8DFA2FB5D" author="EL-HENNAWY, H. K." box="[469,738,1218,1242]" page="101" pageId="75" pageNumber="74" year="1992">
<bibRefCitation id="EFB44B5820191905FE0289E8DFA2FB5D" author="El-Hennawy" box="[469,738,1218,1242]" firstAuthor="El-Hennawy" journalOrPublisher="Serket" page="101" pageId="75" pageNumber="74" pagination="95 - 153" part="2" refId="ref60101" refString="EL-HENNAWY, H. K. 1992. A catalogue of the scorpions described from the Arab Countries (1758 - 1990) (Arachnida: Scorpionida). Serket, 2 (4): 95 - 153." title="A catalogue of the scorpions described from the Arab Countries (1758 - 1990) (Arachnida: Scorpionida)" type="journal article" year="1992">El-Hennawy, 1992: 101</bibRefCitation>
</treatmentCitation>
, 126;
<treatmentCitation id="0A8410B820191905FF2789CADCFBFB7F" author="SISSOM" box="[240,443,1248,1272]" page="23" pageId="75" pageNumber="74" year="1994">
<bibRefCitation id="EFB44B5820191905FF2789CADCFBFB7F" author="Sissom" box="[240,443,1248,1272]" firstAuthor="Sissom" page="23" pageId="75" pageNumber="74" pagination="3 - 39" refId="ref64059" refString="SISSOM, W. D. 1994. Descriptions of new and poorly known scorpions of Yemen (Scorpiones: Buthidae, Diplocentridae, Scorpionidae). Fauna of Saudi Arabia, 14: 3 - 39." type="journal article" year="1994">Sissom, 1994: 23</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B820191905FE1A89CBDE5EFB7F" author="FET, V. &amp; G. LOWE" box="[461,798,1248,1272]" page="156" pageId="75" pageNumber="74" year="2000">
<bibRefCitation id="EFB44B5820191905FE1A89CBDE5EFB7F" author="Fet &amp; Lowe" box="[461,798,1248,1272]" editor="FET, V. &amp; W. D. SISSOM &amp; G. LOWE &amp; M. E. BRAUNWALDER" firstAuthor="Fet" journalOrPublisher="The New York Entomological Society" page="156 - 157" pageId="75" pageNumber="74" pagination="54 - 286" refId="ref60347" refString="FET, V. &amp; G. LOWE. 2000. Family Buthidae. Pp. 54 - 286 in FET, V., W. D. SISSOM, G. LOWE &amp; M. E. BRAUNWALDER. Catalog of the Scorpions of the World (1758 - 1998). The New York Entomological Society." title="Family Buthidae" type="book chapter" volumeTitle="Catalog of the Scorpions of the World (1758 - 1998)" year="2000">Fet &amp; Lowe, 2000: 156–157</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B820191905FF2789D5DCB1FA90" author="YAGMUR" box="[240,497,1279,1303]" page="12" pageId="75" pageNumber="74" year="2009">
<bibRefCitation id="EFB44B5820191905FF2789D5DCB1FA90" author="Yagmur" box="[240,497,1279,1303]" etAl="et al." firstAuthor="Yagmur" page="12" pageId="75" pageNumber="74" pagination="1 - 20" refId="ref65464" refString="YAGMUR, E. A., H. KOC &amp; K. B. KUNT. 2009. Des- cription of a new species of Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) from Southeastern Turkey. Euscorpius, 85: 1 - 20." type="journal article" year="2009">Yağmur et al., 2009: 12</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B820191905FE2C89D5DE5CFA90" author="Khalil &amp; Yağmur" box="[507,796,1279,1303]" page="2" pageId="75" pageNumber="74" year="2010">
<bibRefCitation id="EFB44B5820191905FE2C89D5DE58FA90" author="Khalil &amp; Yagmur" box="[507,792,1279,1303]" firstAuthor="Khalil" journalOrPublisher="Serket" page="2" pageId="75" pageNumber="74" pagination="1 - 6" part="12" refId="ref61426" refString="KHALIL, N. &amp; E. A. YAGMUR. 2010. Leiurus abdullahbayrami (Scorpiones: Buthidae), a new species for the scorpion fauna of Syria. Serket, 12 (1): 1 - 6." title="Leiurus abdullahbayrami (Scorpiones: Buthidae), a new species for the scorpion fauna of Syria" type="journal article" year="2010">Khalil &amp; Yağmur, 2010: 2</bibRefCitation>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A36A920191905FF178835DCC2FAF4" blockId="75.[192,802,758,1456]" pageId="75" pageNumber="74">
<treatmentCitationGroup id="AB35118720191905FF178835DCC2FAF4" pageId="75" pageNumber="74">
<treatmentCitation id="0A8410B820191905FF178835DC5BFAD2" author="POHL" page="215" pageId="75" pageNumber="74" year="1967">
<taxonomicName id="4C254D2A20191905FF178835DC5BFAD2" authority=": Pohl, 1967: 209 - 215" authorityName=": Pohl" authorityPageNumber="209 - 215" authorityYear="1967" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="75" pageNumber="74" phylum="Arthropoda" rank="subSpecies" species="quinquestriatus" subSpecies="voelschowi">
<emphasis id="B951EABB20191905FF178835DF08FAB1" box="[192,584,1310,1334]" italics="true" pageId="75" pageNumber="74">Leiurus quinquestriatus voelschowi</emphasis>
:
<bibRefCitation id="EFB44B5820191905FD8A8834DC5BFAD2" author="Pohl" firstAuthor="Pohl" page="209 - 215" pageId="75" pageNumber="74" pagination="209 - 215" refId="ref63415" refString="POHL, A. 1967. Zuordnung der Art Buthus voelschowi Werner, 1902 zum Formenkreis Leiurus quinquestriatus H. und E., 1829 (Arachnida, Scorpiones). Annalen des Naturhistorischen Museums in Wien, 70: 209 - 215." type="journal article" year="1967">Pohl, 1967: 209– 215</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 1–4;
<treatmentCitation id="0A8410B820191905FE4C8817DF31FAD2" author="KOVARIK, F." box="[411,625,1341,1365]" page="180" pageId="75" pageNumber="74" year="1997">
<bibRefCitation id="EFB44B5820191905FE4C8817DF31FAD2" author="Kovarik" box="[411,625,1341,1365]" firstAuthor="Kovarik" journalOrPublisher="Annales Historiconaturales Musei Nationalis Hungarici" page="180" pageId="75" pageNumber="74" pagination="177 - 185" part="89" refId="ref61724" refString="KOVARIK, F. 1997. A check- list of scorpions (Arachnida) in the collection of the Hungarian Natural History Museum, Budapest. Annales Historiconaturales Musei Nationalis Hungarici, 89: 177 - 185." title="A check- list of scorpions (Arachnida) in the collection of the Hungarian Natural History Museum, Budapest" type="journal article" year="1997">Kovařík, 1997: 180</bibRefCitation>
</treatmentCitation>
(in part, record from
<collectingCountry id="F332763920191905FEFD8871DC36FAF4" box="[298,374,1371,1395]" name="Jordan" pageId="75" pageNumber="74">Jordan</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A36A920191905FF178851DCE4FA37" blockId="75.[192,802,758,1456]" pageId="75" pageNumber="74">
<treatmentCitationGroup id="AB35118720191905FF178851DCE4FA37" pageId="75" pageNumber="74">
<treatmentCitation id="0A8410B820191905FF178851DC6AFA37" author="LEVY, G. &amp; P. AMITAI" pageId="75" pageNumber="74" year="1980">
<taxonomicName id="4C254D2A20191905FF178851DF1DFA15" ID-CoL="6P6WQ" authority=": Vachon, 1966: 212" authorityName="Vachon" authorityPageNumber="212" authorityYear="1966" box="[192,605,1402,1426]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="75" pageNumber="74" phylum="Arthropoda" rank="subSpecies" species="quinquestriatus" subSpecies="hebraeus">
<emphasis id="B951EABB20191905FF178851DC44FA15" box="[192,260,1403,1426]" italics="true" pageId="75" pageNumber="74">Leirus</emphasis>
(sic)
<emphasis id="B951EABB20191905FE9D8851DF1DFA15" box="[330,605,1402,1426]" italics="true" pageId="75" pageNumber="74">quinquestriatus hebraeus</emphasis>
</taxonomicName>
:
<bibRefCitation id="EFB44B5820191905FDB88850DC6AFA37" author="Levy &amp; Amitai" bookContentInfo="130 pp" firstAuthor="Levy" journalOrPublisher="The Israel Academy of Sciences and Humanities. Jerusalem" pageId="75" pageNumber="74" refId="ref62185" refString="LEVY, G. &amp; P. AMITAI. 1980. Scorpiones. Fauna Palaestina. Arachnida I. The Israel Academy of Sciences and Humanities. Jerusalem, 130 pp .." title="Scorpiones. Fauna Palaestina. Arachnida I" type="book" year="1980">Levy &amp; Amitai, 1980</bibRefCitation>
</treatmentCitation>
: Appendix.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33F652220191905FF1788FDD93AFBC7" pageId="75" pageNumber="74" type="materials_examined">
<paragraph id="8B9A36A920191905FF1788FDDC83F9AC" blockId="75.[192,801,1494,1579]" pageId="75" pageNumber="74">
<materialsCitation id="3B4D3CF420191905FF1788FDDC83F9AC" ID-GBIF-Occurrence="3913646507" collectionCode="ZISP" country="Jordan" location="Wadi" municipality="Arrud" pageId="75" pageNumber="74" specimenCode="ZISP 578" specimenCount="5" specimenCount-female="1" specimenCount-immature="4" typeStatus="lectotype">
<collectionCode id="ED34AE6C20191905FF1788FDDCE0FA69" box="[192,416,1494,1518]" pageId="75" pageNumber="74">TYPE MATERIAL</collectionCode>
.
<typeStatus id="549E880B20191905FE6488FDDF5EFA68" box="[435,542,1495,1519]" pageId="75" pageNumber="74" type="lectotype">Lectotype</typeStatus>
adult
<specimenCount id="9D23FD2020191905FDA488FDDFC7FA69" box="[627,647,1495,1518]" pageId="75" pageNumber="74" type="female">♀</specimenCount>
,
<specimenCount id="9D23FD2020191905FD4988FCDE61FA69" box="[670,801,1494,1518]" pageId="75" pageNumber="74" type="immature">4 immature</specimenCount>
paralectoptypes,
<emphasis id="B951EABB20191905FEAF88DFDC8AF98A" bold="true" box="[376,458,1525,1549]" pageId="75" pageNumber="74">
<collectingCountry id="F332763920191905FEAF88DFDC8AF98A" box="[376,458,1525,1549]" name="Jordan" pageId="75" pageNumber="74">Jordan</collectingCountry>
</emphasis>
,
<location id="8EFA607220191905FE0C88DFDF55F98A" LSID="urn:lsid:plazi:treatment:038C87BF2018191FFCBC8822DCDCF930:8EFA607220191905FE0C88DFDF55F98A" box="[475,533,1525,1549]" country="Jordan" municipality="Arrud" name="Wadi" pageId="75" pageNumber="74">Wadi</location>
‘
<collectingMunicipality id="6BFEACD320191905FDF488DFDF27F98A" box="[547,615,1525,1549]" pageId="75" pageNumber="74">Arrud</collectingMunicipality>
(
<specimenCode id="DB839ED220191905FDAF88DFDFA5F98A" box="[632,741,1525,1549]" collectionCode="ZISP" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="75" pageNumber="74">ZISP 578</specimenCode>
) (not examined,
<figureCitation id="131E2A2C20191905FEE28B39DCF6F9AC" box="[309,438,1555,1579]" captionStart-0="Figure 70" captionStart-1="Figure 71" captionStartId-0="88.[192,267,1055,1079]" captionStartId-1="89.[192,267,1598,1622]" captionTargetBox-0="[191,1440,191,1025]" captionTargetBox-1="[191,1439,191,1567]" captionTargetId-0="figure-490@88.[192,1439,192,1024]" captionTargetId-1="figure-114@89.[192,1438,192,1566]" captionTargetPageId-0="88" captionTargetPageId-1="89" captionText-0="Figure 70: Leiurus hebraeus (Birula, 1908) stat. n., lectotype female, designated here (ZISP 578). Habitus. A. Dorsal aspect. B. Ventral aspect. Wadi ‘Arrud, Jordan." captionText-1="Figure 71: Leiurus hebraeus (Birula, 1908) stat. n., lectotype female and four paralectotypes, designated here (ZISP 578). Wadi ‘Arrud, Jordan. Label text in Russian: “Det. A. Birula, 5 ex. Coll. 25.IV.1897, Petro-Arabia, Wadi-Arrud, Southern shore of the Dead Sea, Davydov.”" figureDoi-0="http://doi.org/10.5281/zenodo.6505769" figureDoi-1="http://doi.org/10.5281/zenodo.6505771" httpUri-0="https://zenodo.org/record/6505769/files/figure.png" httpUri-1="https://zenodo.org/record/6505771/files/figure.png" pageId="75" pageNumber="74">Figs. 70–71</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="8B9A36A920191905FF178B7AD93AFBC7" blockId="75.[192,802,1616,1917]" lastBlockId="75.[831,1440,758,1088]" pageId="75" pageNumber="74">
<collectionCode id="ED34AE6C20191905FF178B7ADF02F9EF" box="[192,578,1616,1640]" pageId="75" pageNumber="74">OTHER MATERIAL EXAMINED</collectionCode>
.
<materialsCitation id="3B4D3CF420191905FD9B8B7BDF99F900" ID-GBIF-Occurrence="3913646556" collectionCode="NHMB" country="Israel" location="Palaestina (ev. Jordanien)" pageId="75" pageNumber="74" specimenCount="25" specimenCount-female="7" specimenCount-male="3">
<emphasis id="B951EABB20191905FD9B8B7BDFD6F9EE" bold="true" box="[588,662,1617,1641]" pageId="75" pageNumber="74">
<collectingCountry id="F332763920191905FD9B8B7BDFD1F9EE" box="[588,657,1617,1641]" name="Israel" pageId="75" pageNumber="74">Israel</collectingCountry>
:
</emphasis>
<specimenCount id="9D23FD2020191905FD4B8B7ADF86F9EF" box="[668,710,1616,1640]" pageId="75" pageNumber="74" type="male">3 ♂</specimenCount>
,
<specimenCount id="9D23FD2020191905FD058B7BDFBAF9EF" box="[722,762,1617,1640]" pageId="75" pageNumber="74" type="female">7 ♀</specimenCount>
,
<specimenCount id="9D23FD2020191905FCD08B7ADDAFF900" pageId="75" pageNumber="74">15 juvs</specimenCount>
,
<location id="8EFA607220191905FED48B45DF69F900" LSID="urn:lsid:plazi:treatment:038C87BF2018191FFCBC8822DCDCF930:8EFA607220191905FED48B45DF69F900" box="[259,553,1647,1671]" country="Israel" name="Palaestina (ev. Jordanien)" pageId="75" pageNumber="74">
<taxonomicName id="4C254D2A20191905FED48B45DC2FF900" authorityName="O.Pickard-Cambridge" authorityYear="1872" box="[259,367,1647,1671]" class="Arachnida" family="Zodariidae" genus="Palaestina" kingdom="Animalia" order="Araneae" pageId="75" pageNumber="126" phylum="Arthropoda" rank="genus">Palaestina</taxonomicName>
(ev. Jordanien)
</location>
(
<collectionCode id="ED34AE6C20191905FD978B45DFD2F900" box="[576,658,1647,1671]" pageId="75" pageNumber="74">NHMB</collectionCode>
17a);
</materialsCitation>
<materialsCitation id="3B4D3CF420191905FD3E8B45DFE4F942" ID-GBIF-Occurrence="3913646555" collectingDate="1927-04" collectionCode="NHMB" collectorName="O. Wohlberedt" latitude="31.51" location="Palaestina (ev. Jordanien)" longLatPrecision="121" longitude="35.28" pageId="75" pageNumber="74" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D23FD2020191905FD3E8B45DE5BF900" box="[745,795,1647,1671]" pageId="75" pageNumber="74" type="female">1 ♀</specimenCount>
,
<location id="8EFA607220191905FF178BA4DC97F921" LSID="urn:lsid:plazi:treatment:038C87BF2018191FFCBC8822DCDCF930:8EFA607220191905FF178BA4DC97F921" box="[192,471,1678,1702]" latitude="31.51" longLatPrecision="121" longitude="35.28" name="Palaestina (ev. Jordanien)" pageId="75" pageNumber="74">
<taxonomicName id="4C254D2A20191905FF178BA4DC6CF921" authorityName="O.Pickard-Cambridge" authorityYear="1872" box="[192,300,1678,1702]" class="Arachnida" family="Zodariidae" genus="Palaestina" kingdom="Animalia" order="Araneae" pageId="75" pageNumber="126" phylum="Arthropoda" rank="genus">Palaestina</taxonomicName>
(ev. Jordanien)
</location>
,
<location id="8EFA607220191905FE348BA4DF73F921" LSID="urn:lsid:plazi:treatment:038C87BF2018191FFCBC8822DCDCF930:8EFA607220191905FE348BA4DF73F921" box="[483,563,1678,1702]" country="Israel" latitude="31.51" longLatPrecision="121" longitude="35.28" municipality="Jordanien" name="Jericho" pageId="75" pageNumber="74">Jericho</location>
,
<geoCoordinate id="EE11506E20191905FDE98BA4DFEAF921" box="[574,682,1678,1702]" degrees="31" direction="north" minutes="30.6" orientation="latitude" pageId="75" pageNumber="74" precision="92" value="31.51">31°30.6'N</geoCoordinate>
<geoCoordinate id="EE11506E20191905FD658BA4DE5DF921" box="[690,797,1678,1702]" degrees="35" direction="east" minutes="16.8" orientation="longitude" pageId="75" pageNumber="74" precision="92" value="35.28">35°16.8'E</geoCoordinate>
,
<date id="FF9B106920191905FF178B87DC5AF942" box="[192,282,1709,1733]" pageId="75" pageNumber="74" value="1927-04">
<collectingDate id="EFDFE98120191905FF178B87DC5AF942" box="[192,282,1709,1733]" pageId="75" pageNumber="74" value="1927-04">IV.1927</collectingDate>
</date>
, leg.
<collectorName id="26D0537F20191905FE828B87DCB4F942" box="[341,500,1709,1733]" pageId="75" pageNumber="74">O. Wohlberedt</collectorName>
(
<collectionCode id="ED34AE6C20191905FDD58B87DF13F942" box="[514,595,1709,1733]" pageId="75" pageNumber="74">NHMB</collectionCode>
17-Ia);
</materialsCitation>
<materialsCitation id="3B4D3CF420191905FD7C8B87DC05F8A7" ID-GBIF-Occurrence="3913646535" collectingDate="1983-04-15" collectionCode="MEB" collectorName="A. M. de Saint Michel" country="Israel" elevation="350" latitude="31.103167" location="Hadarom" longLatPrecision="12" longitude="34.8235" pageId="75" pageNumber="74" specimenCode="MEB 034" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D23FD2020191905FD7C8B87DF94F943" box="[683,724,1709,1733]" pageId="75" pageNumber="74" type="female">2 ♀</specimenCount>
, ca.
<locationDeviation id="9A282F8B20191905FCD08B87DCDBF964" location="Beersheba" pageId="75" pageNumber="74">
<quantity id="4CDD9B4C20191905FCD08B87DDA2F964" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="75" pageNumber="74" unit="km" value="11.0">11 km</quantity>
S of Beersheba
</locationDeviation>
,
<location id="8EFA607220191905FE7D8BE1DF50F964" LSID="urn:lsid:plazi:treatment:038C87BF2018191FFCBC8822DCDCF930:8EFA607220191905FE7D8BE1DF50F964" box="[426,528,1739,1763]" country="Israel" latitude="31.103167" longLatPrecision="12" longitude="34.8235" name="Hadarom" pageId="75" pageNumber="74">Hadarom</location>
,
<geoCoordinate id="EE11506E20191905FDC88BE1DFD8F964" box="[543,664,1739,1763]" degrees="31" direction="north" minutes="06.19" orientation="latitude" pageId="75" pageNumber="74" precision="9" value="31.103167">31°06.19'N</geoCoordinate>
<geoCoordinate id="EE11506E20191905FD738BE1DE5DF964" box="[676,797,1739,1763]" degrees="34" direction="east" minutes="49.41" orientation="longitude" pageId="75" pageNumber="74" precision="9" value="34.8235">34°49.41'E</geoCoordinate>
,
<elevation id="0008D19A20191905FF178BC0DC0BF885" box="[192,331,1770,1794]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="75" pageNumber="74" unit="m" value="350.0">
<quantity id="4CDD9B4C20191905FF178BC0DC4AF886" box="[192,266,1770,1794]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="75" pageNumber="74" unit="m" value="350.0">350 m</quantity>
a.s.l.
</elevation>
,
<date id="FF9B106920191905FE888BC0DC9BF885" box="[351,475,1770,1794]" pageId="75" pageNumber="74" value="1983-04-15">
<collectingDate id="EFDFE98120191905FE888BC0DC9BF885" box="[351,475,1770,1794]" pageId="75" pageNumber="74" value="1983-04-15">15.IV.1983</collectingDate>
</date>
, leg.
<collectorName id="26D0537F20191905FDF48BC0DE60F885" box="[547,800,1770,1794]" pageId="75" pageNumber="74">A.M. de Saint Michel</collectorName>
(
<specimenCode id="DB839ED220191905FF108A22DC7DF8A7" box="[199,317,1800,1824]" collectionCode="MEB" pageId="75" pageNumber="74">MEB 034</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B4D3CF420191905FE808A22DFE5F8D9" ID-GBIF-Occurrence="3913646524" collectingDate="1984-09-20" collectionCode="MEB" collectorName="Nesher &amp; Yagur" country="Israel" elevation="27" latitude="32.753666" location="Haifa District" longLatPrecision="12" longitude="35.0615" pageId="75" pageNumber="74" specimenCode="MEB 197" specimenCount="1" specimenCount-male="1" stateProvince="Haifa">
<specimenCount id="9D23FD2020191905FE808A22DCC8F8A7" box="[343,392,1800,1824]" pageId="75" pageNumber="74" type="male">1 ♂</specimenCount>
, 2 juvs, between
<collectorName id="26D0537F20191905FD8B8A22DFE7F8A7" box="[604,679,1800,1824]" pageId="75" pageNumber="74">Nesher</collectorName>
&amp;
<collectorName id="26D0537F20191905FD008A23DE5DF8A6" box="[727,797,1801,1825]" pageId="75" pageNumber="74">Yagur</collectorName>
,
<collectingRegion id="49E1F84B20191905FF178A0DDC21F8B8" box="[192,353,1831,1855]" country="Israel" name="Haifa" pageId="75" pageNumber="74">Haifa District</collectingRegion>
,
<geoCoordinate id="EE11506E20191905FEA08A0DDCB1F8B8" box="[375,497,1831,1855]" degrees="32" direction="north" minutes="45.22" orientation="latitude" pageId="75" pageNumber="74" precision="9" value="32.753666">32°45.22'N</geoCoordinate>
<geoCoordinate id="EE11506E20191905FDD38A0DDF3AF8B8" box="[516,634,1831,1855]" degrees="35" direction="east" minutes="03.69" orientation="longitude" pageId="75" pageNumber="74" precision="9" value="35.0615">35°03.69'E</geoCoordinate>
,
<elevation id="0008D19A20191905FD438A0DDE5CF8B8" box="[660,796,1831,1855]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="75" pageNumber="74" unit="m" value="27.0">
<quantity id="4CDD9B4C20191905FD438A0DDF96F8B8" box="[660,726,1831,1855]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="75" pageNumber="74" unit="m" value="27.0">27 m</quantity>
a.s.l.
</elevation>
,
<date id="FF9B106920191905FF178A6CDC7CF8D9" box="[192,316,1862,1886]" pageId="75" pageNumber="74" value="1984-09-20">
<collectingDate id="EFDFE98120191905FF178A6CDC7CF8D9" box="[192,316,1862,1886]" pageId="75" pageNumber="74" value="1984-09-20">20.IX.1984</collectingDate>
</date>
, leg. M.
<collectionCode id="ED34AE6C20191905FE4D8A6DDCEEF8D9" box="[410,430,1863,1886]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="75" pageNumber="74">R</collectionCode>
.
<location id="8EFA607220191905FE6B8A6CDF5BF8D9" LSID="urn:lsid:plazi:treatment:038C87BF2018191FFCBC8822DCDCF930:8EFA607220191905FE6B8A6CDF5BF8D9" box="[444,539,1862,1886]" country="Israel" name="Warburg" pageId="75" pageNumber="74">Warburg</location>
(
<specimenCode id="DB839ED220191905FDFB8A6CDFDDF8D9" box="[556,669,1862,1886]" collectionCode="MEB" pageId="75" pageNumber="74">MEB 197</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B4D3CF420191905FD648A6CD9D0FC89" ID-GBIF-Occurrence="3913646480" collectingDate="1984-11-30" collectionCode="MEB" country="Israel" elevation="300" latitude="32.8" location="Haifa" longLatPrecision="1209" longitude="34.983334" pageId="75" pageNumber="74" specimenCode="MEB 198" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Haifa">
<specimenCount id="9D23FD2020191905FD648A6CDFA1F8D9" box="[691,737,1862,1886]" pageId="75" pageNumber="74" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D23FD2020191905FD388A6CDE5AF8D9" box="[751,794,1862,1886]" pageId="75" pageNumber="74" type="female">1 ♀</specimenCount>
,
<collectingRegion id="49E1F84B20191905FF178A4FDDBBF8FA" box="[192,251,1893,1917]" country="Israel" name="Haifa" pageId="75" pageNumber="74">Haifa</collectingRegion>
area,
<geoCoordinate id="EE11506E20191905FE948A4FDCDBF8FA" box="[323,411,1893,1917]" degrees="32" direction="north" minutes="48" orientation="latitude" pageId="75" pageNumber="74" precision="925" value="32.8">32°48'N</geoCoordinate>
<geoCoordinate id="EE11506E20191905FE718A4FDCBDF8FA" box="[422,509,1893,1917]" degrees="34" direction="east" minutes="59" orientation="longitude" pageId="75" pageNumber="74" precision="925" value="34.983334">34°59'E</geoCoordinate>
,
<elevation id="0008D19A20191905FDDB8A4FDFD0F8FA" box="[524,656,1893,1917]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="75" pageNumber="74" unit="m" value="300.0">
<quantity id="4CDD9B4C20191905FDDB8A4FDF13F8FB" box="[524,595,1893,1917]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="75" pageNumber="74" unit="m" value="300.0">300 m</quantity>
a.s.l.
</elevation>
,
<date id="FF9B106920191905FD768A4FDE5CF8FA" box="[673,796,1893,1917]" pageId="75" pageNumber="74" value="1984-11-30">
<collectingDate id="EFDFE98120191905FD768A4FDE5CF8FA" box="[673,796,1893,1917]" pageId="75" pageNumber="74" value="1984-11-30">30.XI.1984</collectingDate>
</date>
, leg. M.
<collectionCode id="ED34AE6C20191905FC5B8FDCDEE0FC8A" box="[908,928,758,781]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="75" pageNumber="74">R</collectionCode>
.
<location id="8EFA607220191905FC7B8FDCD94BFC89" LSID="urn:lsid:plazi:treatment:038C87BF2018191FFCBC8822DCDCF930:8EFA607220191905FC7B8FDCD94BFC89" box="[940,1035,758,782]" country="Israel" name="Warburg" pageId="75" pageNumber="74">Warburg</location>
(
<specimenCode id="DB839ED220191905FBCE8FDCD9C7FC89" box="[1049,1159,758,782]" collectionCode="MEB" pageId="75" pageNumber="74">MEB 198</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B4D3CF420191905FB4B8FDCD989FCCC" ID-GBIF-Occurrence="3913646538" collectionCode="FKCP" collectorName="J. Kral" country="Israel" location="Negev" pageId="75" pageNumber="74" specimenCount="4" specimenCount-female="3" specimenCount-male="1">
<specimenCount id="9D23FD2020191905FB4B8FDCD987FC8A" box="[1180,1223,758,782]" pageId="75" pageNumber="74" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D23FD2020191905FB048FDCD9BCFC8A" box="[1235,1276,758,782]" pageId="75" pageNumber="74" type="female">3 ♀</specimenCount>
, 3 juvs,
<location id="8EFA607220191905FA8D8FDCD8DFFC89" LSID="urn:lsid:plazi:treatment:038C87BF2018191FFCBC8822DCDCF930:8EFA607220191905FA8D8FDCD8DFFC89" box="[1370,1439,758,782]" country="Israel" name="Negev" pageId="75" pageNumber="74">Negev</location>
desert,
<collectorName id="26D0537F20191905FC478E3ED951FCAB" box="[912,1041,788,812]" pageId="75" pageNumber="74">Vadi Hazaz</collectorName>
near
<collectorName id="26D0537F20191905FB838E3ED994FCAB" box="[1108,1236,788,812]" pageId="75" pageNumber="74">Sede Boqer</collectorName>
(
<collectorName id="26D0537F20191905FB308E3ED8D4FCAB" box="[1255,1428,788,812]" pageId="75" pageNumber="74">Haluqim Ridge</collectorName>
),
<date id="FF9B106920191905FCE88E1EDED1FCCC" box="[831,913,819,843]" pageId="75" pageNumber="74" value="2004-11" valueMax="2004-12-31" valueMin="2004-11-00">
XI.-
<collectionCode id="ED34AE6C20191905FCBD8E19DED1FCCC" box="[874,913,819,843]" pageId="75" pageNumber="74">XII</collectionCode>
</date>
.2004, leg.
<collectorName id="26D0537F20191905FBC48E1ED920FCCC" box="[1043,1120,819,843]" pageId="75" pageNumber="74">J. Král</collectorName>
(
<collectionCode id="ED34AE6C20191905FBA28E19D9FEFCCC" box="[1141,1214,819,843]" pageId="75" pageNumber="74">FKCP</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B4D3CF420191905FB008E19DED7FC0F" ID-GBIF-Occurrence="3913646417" collectingDate="2007-09" collectionCode="FKCP" collectorName="J. Kral &amp; M. Forman" country="Israel" location="Sede Boqer" municipality="Negev" pageId="75" pageNumber="74" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<specimenCount id="9D23FD2020191905FB008E19D847FCCC" box="[1239,1287,819,843]" pageId="75" pageNumber="74" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D23FD2020191905FACE8E19D807FCCC" box="[1305,1351,819,843]" pageId="75" pageNumber="74" type="female">2 ♀</specimenCount>
,
<collectingMunicipality id="6BFEACD320191905FA8C8E1ED8E0FCCB" box="[1371,1440,820,844]" pageId="75" pageNumber="74">Negev</collectingMunicipality>
desert,
<location id="8EFA607220191905FC5B8E78D94BFCED" LSID="urn:lsid:plazi:treatment:038C87BF2018191FFCBC8822DCDCF930:8EFA607220191905FC5B8E78D94BFCED" box="[908,1035,850,874]" country="Israel" municipality="Negev" name="Sede Boqer" pageId="75" pageNumber="74">Sede Boqer</location>
,
<date id="FF9B106920191905FBC18E78D931FCED" box="[1046,1137,850,874]" pageId="75" pageNumber="74" value="2007-09">
<collectingDate id="EFDFE98120191905FBC18E78D931FCED" box="[1046,1137,850,874]" pageId="75" pageNumber="74" value="2007-09">IX.2007</collectingDate>
</date>
, leg.
<collectorName id="26D0537F20191905FB7D8E79D9B2FCED" box="[1194,1266,850,874]" pageId="75" pageNumber="74">J. Král</collectorName>
and
<collectorName id="26D0537F20191905FAF08E79D8E0FCED" box="[1319,1440,851,874]" pageId="75" pageNumber="74">M. Forman</collectorName>
(
<collectionCode id="ED34AE6C20191905FC918E5ADECFFC0F" box="[838,911,880,904]" pageId="75" pageNumber="74">FKCP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4D3CF420191905FC748E5BD8C5FC20" ID-GBIF-Occurrence="3913646491" collectionCode="MEB" collectorName="M. A. Jafar" country="Jordan" elevation="764" latitude="31.716667" location="Madaba" longLatPrecision="1214" longitude="35.8" pageId="75" pageNumber="74" specimenCode="MEB 346" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Madaba">
<emphasis id="B951EABB20191905FC748E5BDEBEFC0E" bold="true" box="[931,1022,881,905]" pageId="75" pageNumber="74">
<collectingCountry id="F332763920191905FC748E5BDEB8FC0E" box="[931,1016,881,905]" name="Jordan" pageId="75" pageNumber="74">Jordan</collectingCountry>
:
</emphasis>
<specimenCount id="9D23FD2020191905FBD08E5AD974FC0F" box="[1031,1076,880,904]" pageId="75" pageNumber="74" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D23FD2020191905FB958E5AD92DFC0F" box="[1090,1133,880,904]" pageId="75" pageNumber="74" type="female">4 ♀</specimenCount>
,
<collectingRegion id="49E1F84B20191905FBAA8E5AD996FC0F" box="[1149,1238,880,904]" country="Jordan" name="Madaba" pageId="75" pageNumber="74">Madaba</collectingRegion>
,
<geoCoordinate id="EE11506E20191905FB348E5AD87BFC0F" box="[1251,1339,880,904]" degrees="31" direction="north" minutes="43" orientation="latitude" pageId="75" pageNumber="74" precision="925" value="31.716667">31°43'N</geoCoordinate>
<geoCoordinate id="EE11506E20191905FA938E5AD8DBFC0F" box="[1348,1435,880,904]" degrees="35" direction="east" minutes="48" orientation="longitude" pageId="75" pageNumber="74" precision="925" value="35.8">35°48'E</geoCoordinate>
,
<elevation id="0008D19A20191905FCE88EA5DE80FC20" box="[831,960,911,935]" metricMagnitude="2" metricUnit="m" metricValue="7.64" pageId="75" pageNumber="74" unit="m" value="764.0">
<quantity id="4CDD9B4C20191905FCE88EA5DEC4FC20" box="[831,900,911,935]" metricMagnitude="2" metricUnit="m" metricValue="7.64" pageId="75" pageNumber="74" unit="m" value="764.0">764 m</quantity>
a.s.l.
</elevation>
,
<date id="FF9B106920191905FC188EA5D94DFC20" box="[975,1037,911,935]" pageId="75" pageNumber="74" value="1983-03-22">
22.
<collectionCode id="ED34AE6C20191905FC278EA5D94DFC20" box="[1008,1037,911,935]" pageId="75" pageNumber="74">III</collectionCode>
</date>
.1983, leg.
<collectorName id="26D0537F20191905FB508EA5D9BCFC20" box="[1159,1276,911,935]" pageId="75" pageNumber="74">M.A. Jafar</collectorName>
(
<specimenCode id="DB839ED220191905FADB8EA5D83DFC20" box="[1292,1405,911,935]" collectionCode="MEB" pageId="75" pageNumber="74">MEB 346</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B4D3CF420191905FA458EA5D94DFC63" ID-GBIF-Occurrence="3913646393" collectionCode="GL" country="Jordan" elevation="200" latitude="32.45" location="Tabaqat" longLatPrecision="1211" longitude="35.616665" pageId="75" pageNumber="74" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D23FD2020191905FA458EA5DE13FC42" pageId="75" pageNumber="74" type="female">1 ♀</specimenCount>
,
<location id="8EFA607220191905FCB38E84DEFDFC41" LSID="urn:lsid:plazi:treatment:038C87BF2018191FFCBC8822DCDCF930:8EFA607220191905FCB38E84DEFDFC41" box="[868,957,942,966]" country="Jordan" latitude="32.45" longLatPrecision="1211" longitude="35.616665" name="Tabaqat" pageId="75" pageNumber="74">Tabaqat</location>
,
<geoCoordinate id="EE11506E20191905FC1C8E84D963FC41" box="[971,1059,942,966]" degrees="32" direction="north" minutes="27" orientation="latitude" pageId="75" pageNumber="74" precision="925" value="32.45">32°27'N</geoCoordinate>
<geoCoordinate id="EE11506E20191905FBFA8E84D9C5FC41" box="[1069,1157,942,966]" degrees="35" direction="east" minutes="37" orientation="longitude" pageId="75" pageNumber="74" precision="925" value="35.616665">35°37'E</geoCoordinate>
, –
<elevation id="0008D19A20191905FB758E84D865FC41" box="[1186,1317,942,966]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="75" pageNumber="74" unit="m" value="200.0">
<quantity id="4CDD9B4C20191905FB758E84D9A8FC42" box="[1186,1256,942,966]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="75" pageNumber="74" unit="m" value="200.0">200 m</quantity>
a.s.l.
</elevation>
,
<date id="FF9B106920191905FAE18E84D81EFC41" box="[1334,1374,942,966]" pageId="75" pageNumber="74" value="1995-05-04">
4.
<collectionCode id="ED34AE6C20191905FA9F8E84D81EFC41" box="[1352,1374,942,966]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="75" pageNumber="74" type="Museum">V</collectionCode>
</date>
.1995, leg.
<collectionCode id="ED34AE6C20191905FCB88EE7DEC5FC63" box="[879,901,973,996]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="75" pageNumber="74" type="Museum">V</collectionCode>
.
<location id="8EFA607220191905FC448EE6DE8EFC63" LSID="urn:lsid:plazi:treatment:038C87BF2018191FFCBC8822DCDCF930:8EFA607220191905FC448EE6DE8EFC63" box="[915,974,972,996]" country="Jordan" name="Sejva" pageId="75" pageNumber="74">Sejva</location>
(GL)
</materialsCitation>
;
<materialsCitation id="3B4D3CF420191905FBC98EE6D9E0FB84" ID-GBIF-Occurrence="3913646349" collectionCode="GL" country="Jordan" latitude="32.279167" location="Kurayyima Udoli" longLatPrecision="12" longitude="35.59783" pageId="75" pageNumber="74" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D23FD2020191905FBC98EE6D909FC63" box="[1054,1097,972,996]" pageId="75" pageNumber="74" type="female">1 ♀</specimenCount>
,
<location id="8EFA607220191905FB8E8EE6D859FC63" LSID="urn:lsid:plazi:treatment:038C87BF2018191FFCBC8822DCDCF930:8EFA607220191905FB8E8EE6D859FC63" box="[1113,1305,972,996]" country="Jordan" latitude="32.279167" longLatPrecision="12" longitude="35.59783" name="Kurayyima Udoli" pageId="75" pageNumber="74">Kurayyima Udoli</location>
,
<geoCoordinate id="EE11506E20191905FAF08EE6D8E0FC63" box="[1319,1440,972,996]" degrees="32" direction="north" minutes="16.75" orientation="latitude" pageId="75" pageNumber="74" precision="9" value="32.279167">32°16.75'N</geoCoordinate>
<geoCoordinate id="EE11506E20191905FCE88EC1DEF8FB84" box="[831,952,1003,1027]" degrees="35" direction="east" minutes="35.87" orientation="longitude" pageId="75" pageNumber="74" precision="9" value="35.59783">35°35.87'E</geoCoordinate>
,
<date id="FF9B106920191905FC188EC1D955FB84" box="[975,1045,1003,1027]" pageId="75" pageNumber="74" value="2000-07-23">
23.
<collectionCode id="ED34AE6C20191905FC3A8EC1D955FB84" box="[1005,1045,1003,1027]" pageId="75" pageNumber="74">VII</collectionCode>
</date>
.2000 (GL).
</materialsCitation>
<materialsCitation id="3B4D3CF420191905FB638EC6D9F1FBA5" ID-GBIF-Occurrence="3913646473" collectionCode="FKCP" collectorName="Bosra" country="Syria" location="Syria" pageId="75" pageNumber="74" specimenCount="4" specimenCount-male="4">
<emphasis id="B951EABB20191905FB638EC6D9B9FB83" bold="true" box="[1204,1273,1004,1028]" pageId="75" pageNumber="74">
<collectingCountry id="F332763920191905FB638EC6D9B5FB83" box="[1204,1269,1004,1028]" name="Syria" pageId="75" pageNumber="74">Syria</collectingCountry>
:
</emphasis>
<specimenCount id="9D23FD2020191905FADA8EC1D803FB84" box="[1293,1347,1003,1027]" pageId="75" pageNumber="74" type="male">4 ♂</specimenCount>
,
<collectorName id="26D0537F20191905FA8C8EC6D8DCFB84" box="[1371,1436,1004,1027]" pageId="75" pageNumber="74">Bosra</collectorName>
,
<date id="FF9B106920191905FCE88920DE14FBA5" box="[831,852,1034,1058]" pageId="75" pageNumber="74" value="1994-05">
<collectionCode id="ED34AE6C20191905FCE88920DE14FBA5" box="[831,852,1034,1058]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="75" pageNumber="74" type="Museum">V</collectionCode>
</date>
.1994, leg.
<collectorName id="26D0537F20191905FC008920D907FBA5" box="[983,1095,1034,1058]" pageId="75" pageNumber="74">D. Modrý</collectorName>
(
<collectionCode id="ED34AE6C20191905FB8B8920D9E5FBA5" box="[1116,1189,1034,1058]" pageId="75" pageNumber="74">FKCP</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B4D3CF420191905FB688920D93AFBC7" ID-GBIF-Occurrence="3913646430" collectionCode="FKCP" collectorName="Seydmaya" country="Syria" pageId="75" pageNumber="74" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D23FD2020191905FB688920D9AEFBA6" box="[1215,1262,1034,1058]" pageId="75" pageNumber="74" type="female">1 ♀</specimenCount>
,
<collectorName id="26D0537F20191905FAD58920D833FBA5" box="[1282,1395,1034,1058]" pageId="75" pageNumber="74">Seydmaya</collectorName>
,
<date id="FF9B106920191905FA528920D8DCFBA6" box="[1413,1436,1034,1057]" pageId="75" pageNumber="74" value="1994-05">
<collectionCode id="ED34AE6C20191905FA528920D8DCFBA6" box="[1413,1436,1034,1057]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="75" pageNumber="74" type="Museum">V</collectionCode>
</date>
. 1994, leg.
<collectorName id="26D0537F20191905FC788903D958FBC7" box="[943,1048,1064,1088]" pageId="75" pageNumber="74">D. Modrý</collectorName>
(
<collectionCode id="ED34AE6C20191905FBF18902D92FFBC7" box="[1062,1135,1064,1088]" pageId="75" pageNumber="74">FKCP</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33F652220191902FCE8894CDC34F9E7" lastPageId="76" lastPageNumber="75" pageId="75" pageNumber="74" type="diagnosis">
<paragraph id="8B9A36A920191902FCE8894CDC34F9E7" blockId="75.[831,1441,1126,1917]" lastBlockId="76.[192,802,1332,1632]" lastPageId="76" lastPageNumber="75" pageId="75" pageNumber="74">
DIAGNOSIS (adults). Small to medium sized
<taxonomicName id="4C254D2A20191905FA9F894DD8D9FBF9" authorityName=": Vachon" authorityYear="1949" box="[1352,1433,1127,1150]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="75" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABB20191905FA9F894DD8D9FBF9" box="[1352,1433,1127,1150]" italics="true" pageId="75" pageNumber="74">Leiurus</emphasis>
</taxonomicName>
,
<quantity id="4CDD9B4C20191905FCE889AFDEF2FB1B" box="[831,946,1157,1181]" metricMagnitude="-2" metricUnit="m" metricValue="6.75" metricValueMax="7.7" metricValueMin="5.8" pageId="75" pageNumber="74" unit="mm" value="67.5" valueMax="77.0" valueMin="58.0">58–77 mm</quantity>
in length, carapace L
<quantity id="4CDD9B4C20191905FB7589AFD867FB1B" box="[1186,1319,1157,1181]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" metricValueMax="9.3" metricValueMin="6.5" pageId="75" pageNumber="74" unit="mm" value="7.9" valueMax="9.3" valueMin="6.5">6.5–9.3 mm</quantity>
; base color yellow to orange-brown, with variable fuscosity on carapace and tergites; metasoma V either clear or fuscous except for posterior end; carapace with area between anterior median carinae finely shagreened, with scattered medium to fine granules, area between posterior median carinae with shallow median furrow flanked by arcs of medium or fine granules; medial intercarinal surfaces of tergites II–III shagreened, with variable medium to fine granulation; posterior margin of coxa III smooth or with sparse fine granules; metasoma robust, metasoma II L/
<geoCoordinate id="EE11506E20191905FA51889DDE2EFA69" degrees="1.38" direction="west" orientation="longitude" pageId="75" pageNumber="74" precision="555" value="-1.38">W 1.38</geoCoordinate>
–1.64, metasoma III L/
<geoCoordinate id="EE11506E20191905FBB988FCD980FA69" box="[1134,1216,1494,1518]" degrees="1.49" direction="west" orientation="longitude" pageId="75" pageNumber="74" precision="555" value="-1.49">W 1.49</geoCoordinate>
–1.69, metasoma IV L/
<geoCoordinate id="EE11506E20191905FC8188DFDEEBF98A" box="[854,939,1525,1549]" degrees="1.71" direction="west" orientation="longitude" pageId="75" pageNumber="74" precision="555" value="-1.71">W 1.71</geoCoordinate>
–2.03; ventromedian carinae of metasoma II and III with 14–19 denticles (46/48 carinae); metasoma V with enlarged subtriangular or lobate denticles on ventrolateral carinae; pedipalps relatively stout, patella L/W
<specimenCount id="9D23FD2020191905FCA18B5ADECAF900" box="[886,906,1648,1671]" pageId="75" pageNumber="74" type="male">♂</specimenCount>
2.81–2.95,
<specimenCount id="9D23FD2020191905FBDE8B5AD95DF900" box="[1033,1053,1648,1671]" pageId="75" pageNumber="74" type="female">♀</specimenCount>
2.48–2.83; leg III patella L/D 3.13– 3.71; pectine teeth
<specimenCount id="9D23FD2020191905FBD98BA5D962F921" box="[1038,1058,1679,1702]" pageId="75" pageNumber="74" type="male">♂</specimenCount>
33–37,
<specimenCount id="9D23FD2020191905FBAC8BA5D9CFF921" box="[1147,1167,1679,1702]" pageId="75" pageNumber="74" type="female">♀</specimenCount>
27–32; pectines medium sized, pectine L/ carapace L
<specimenCount id="9D23FD2020191905FBAD8B87D9CEF943" box="[1146,1166,1709,1732]" pageId="75" pageNumber="74" type="male">♂</specimenCount>
1.14–1.29,
<specimenCount id="9D23FD2020191905FAC58B87D866F943" box="[1298,1318,1709,1732]" pageId="75" pageNumber="74" type="female">♀</specimenCount>
0.92–1.21, mid-pectine sensillar margin L/ metasoma I W
<specimenCount id="9D23FD2020191905FAEC8BE6D80FF964" box="[1339,1359,1740,1763]" pageId="75" pageNumber="74" type="male">♂</specimenCount>
<geoCoordinate id="EE11506E20191905FA818BE1D8D2F964" box="[1366,1426,1739,1763]" degrees="0.169" direction="north" orientation="latitude" pageId="75" pageNumber="74" precision="55" value="0.169">0.169</geoCoordinate>
<geoCoordinate id="EE11506E20191905FA438BE6DE3EF885" degrees="0.180" direction="west" orientation="longitude" pageId="75" pageNumber="74" precision="55" value="-0.18">– 0.180</geoCoordinate>
,
<specimenCount id="9D23FD2020191905FC5C8BC0DEDFF886" box="[907,927,1770,1793]" pageId="75" pageNumber="74" type="female">♀</specimenCount>
<geoCoordinate id="EE11506E20191905FC7F8BC0DEA4F885" box="[936,996,1770,1794]" degrees="0.100" direction="north" orientation="latitude" pageId="75" pageNumber="74" precision="55" value="0.1">0.100</geoCoordinate>
<geoCoordinate id="EE11506E20191905FC338BC0D970F885" box="[996,1072,1770,1794]" degrees="0.136" direction="west" orientation="longitude" pageId="75" pageNumber="74" precision="55" value="-0.136">–0.136</geoCoordinate>
; basal 1–2 pectine teeth of males overlap if anterior pectine margins aligned to posterior margins of coxae IV; pectine basal piece very lightly, finely shagreened, almost smooth in females, almost smooth or weakly granulated in males; leg III basitarsus with 7–12 retrosuperior setae; pedipalp chela fixed finger with trichobothrium
<emphasis id="B951EABB201E1902FE348879DCBEFAEC" box="[483,510,1363,1387]" italics="true" pageId="76" pageNumber="75">db</emphasis>
either distal or proximal to
<emphasis id="B951EABB201E1902FF178858DD9DFA0E" box="[192,221,1394,1417]" italics="true" pageId="76" pageNumber="75">est</emphasis>
; sternite VII with area between median carinae densely, finely microgranulated or shagreened; sternite carination:
<emphasis id="B951EABB201E1902FEEC8884DC39FA41" box="[315,377,1454,1478]" italics="true" pageId="76" pageNumber="75">males</emphasis>
, sternite III with median carinae moderate to strong, sternites IV–V with lateral carinae strong, median carinae weak to moderate;
<emphasis id="B951EABB201E1902FD4688C6DFA2F983" box="[657,738,1516,1540]" italics="true" pageId="76" pageNumber="75">females</emphasis>
, sternite III with median carinae weak or obsolete, sternites IV–V with lateral carinae weak to moderate, median carinae obsolete.
</paragraph>
</subSubSection>
<caption id="DF5A6621201E1902FF17899DD86FFA87" ID-DOI="http://doi.org/10.5281/zenodo.6505745" ID-Zenodo-Dep="6505745" httpUri="https://zenodo.org/record/6505745/files/figure.png" pageId="76" pageNumber="75" startId="76.[192,267,1207,1231]" targetBox="[191,1437,213,1177]" targetPageId="76">
<paragraph id="8B9A36A9201E1902FF17899DD86FFA87" blockId="76.[192,1441,1207,1280]" pageId="76" pageNumber="75">
<emphasis id="B951EABB201E1902FF17899DDC77FB48" bold="true" box="[192,311,1207,1231]" pageId="76" pageNumber="75">Figure 58:</emphasis>
Hemispermatophores of new
<taxonomicName id="4C254D2A201E1902FD8A8990DE54FB49" box="[605,788,1209,1231]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="76" pageNumber="75" phylum="Arthropoda" rank="species" species="undefined-a">
<emphasis id="B951EABB201E1902FD8A8990DFE7FB48" box="[605,679,1210,1231]" italics="true" pageId="76" pageNumber="75">Leiurus</emphasis>
species.
<emphasis id="B951EABB201E1902FCD68993DE54FB49" bold="true" box="[769,788,1209,1230]" pageId="76" pageNumber="75">A</emphasis>
</taxonomicName>
<emphasis id="B951EABB201E1902FCC48993DE58FB49" bold="true" box="[787,792,1209,1230]" pageId="76" pageNumber="75">.</emphasis>
<taxonomicName id="4C254D2A201E1902FCF78990DEF6FB48" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" box="[800,950,1210,1231]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="76" pageNumber="75" phylum="Arthropoda" rank="species" species="macroctenus" status="sp. nov.">
<emphasis id="B951EABB201E1902FCF78990DE6DFB48" box="[800,813,1210,1231]" italics="true" pageId="76" pageNumber="75">L</emphasis>
.
<emphasis id="B951EABB201E1902FCEC8990DEF6FB48" box="[827,950,1210,1231]" italics="true" pageId="76" pageNumber="75">macroctenus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C0201E1902FC6A8990DEB3FB48" box="[957,1011,1210,1231]" pageId="76" pageNumber="75" rank="species">sp. n.</taxonomicNameLabel>
, paratype male, Duhai, Oman.
<emphasis id="B951EABB201E1902FAF38993D87BFB49" bold="true" box="[1316,1339,1209,1230]" pageId="76" pageNumber="75">B.</emphasis>
<taxonomicName id="4C254D2A201E1902FA958990D8E1FB48" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" box="[1346,1441,1209,1231]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="76" pageNumber="75" phylum="Arthropoda" rank="species" species="heberti" status="sp. nov.">
<emphasis id="B951EABB201E1902FA958990D80FFB48" box="[1346,1359,1210,1231]" italics="true" pageId="76" pageNumber="75">L</emphasis>
.
<emphasis id="B951EABB201E1902FA8A8993D8E1FB48" box="[1373,1441,1209,1231]" italics="true" pageId="76" pageNumber="75">heberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C0201E1902FF1789F8DDBAFB60" box="[192,250,1234,1255]" pageId="76" pageNumber="75" rank="species">sp. n.</taxonomicNameLabel>
, holotype male, Wadi Andur, Oman.
<emphasis id="B951EABB201E1902FDA889F8DFD7FB60" bold="true" box="[639,663,1234,1255]" pageId="76" pageNumber="75">C.</emphasis>
<taxonomicName id="4C254D2A201E1902FD7589F8DE54FB60" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" box="[674,788,1233,1255]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="76" pageNumber="75" phylum="Arthropoda" rank="species" species="haenggii" status="sp. nov.">
<emphasis id="B951EABB201E1902FD7589F8DFEFFB60" box="[674,687,1234,1255]" italics="true" pageId="76" pageNumber="75">L</emphasis>
.
<emphasis id="B951EABB201E1902FD1789FBDE54FB60" box="[704,788,1233,1255]" italics="true" pageId="76" pageNumber="75">haenggii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C0201E1902FCC889F8DE18FB60" box="[799,856,1234,1255]" pageId="76" pageNumber="75" rank="species">sp. n.</taxonomicNameLabel>
, paratype male, Jabal Qara, Oman. Distal portions hemispermatophores illustrated to show flagellum and lobes (right hemispermatophores, convex aspect). Scale bars: 1 mm.
</paragraph>
</caption>
<subSubSection id="C33F6522201E1902FF178BAFDEC9F8F3" pageId="76" pageNumber="75" type="description">
<paragraph id="8B9A36A9201E1902FF178BAFDCE1F91A" blockId="76.[192,801,1669,1908]" box="[192,417,1669,1693]" pageId="76" pageNumber="75">MEASUREMENTS.</paragraph>
<paragraph id="8B9A36A9201E1902FF278B8ED945FA62" blockId="76.[192,801,1669,1908]" lastBlockId="76.[831,1441,1332,1908]" pageId="76" pageNumber="75">
<emphasis id="B951EABB201E1902FF278B8EDF39F93B" bold="true" box="[240,633,1700,1724]" pageId="76" pageNumber="75">
Male,
<collectingCountry id="F3327639201E1902FEE98B8EDC3FF93B" box="[318,383,1700,1724]" name="Israel" pageId="76" pageNumber="75">Israel</collectingCountry>
(
<specimenCode id="DB839ED2201E1902FE438B8EDF52F93B" box="[404,530,1700,1724]" collectionCode="NHMB" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="76" pageNumber="75">NHMB 17</specimenCode>
a) (mm).
</emphasis>
<collectorName id="26D0537F201E1902FD508B8EDF9EF93B" box="[647,734,1700,1724]" pageId="76" pageNumber="75">Total L</collectorName>
58.0; metasoma + telson L 36.0; carapace L 6.74,
<geoCoordinate id="EE11506E201E1902FD108BE9DE5DF95C" box="[711,797,1731,1755]" degrees="7.22" direction="west" orientation="longitude" pageId="76" pageNumber="75" precision="555" value="-7.22">W 7.22</geoCoordinate>
, carapace preocular L 3.05; metasomal segments (L/ D /W) I 4.98/ 4.01/4.34,
<specimenCode id="DB839ED2201E1902FE0E8A2BDF69F89F" box="[473,553,1792,1816]" collectionCode="II" pageId="76" pageNumber="75">II 5.82</specimenCode>
/ 3.63/3.97,
<specimenCode id="DB839ED2201E1902FD688A2BDE59F89F" box="[703,793,1792,1816]" collectionCode="III" pageId="76" pageNumber="75">III 6.08</specimenCode>
/ 3.56/3.85, IV 6.92/ 3.44/3.59,
<specimenCode id="DB839ED2201E1902FDD88A35DF1AF8B0" box="[527,602,1823,1847]" collectionCode="V" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="76" pageNumber="75" type="Museum">V 7.67</specimenCode>
/ 2.89/3.49; telson L 7.00; vesicle L 3.85, D 2.78,
<geoCoordinate id="EE11506E201E1902FDC68A14DF23F8D2" box="[529,611,1853,1877]" degrees="2.87" direction="west" orientation="longitude" pageId="76" pageNumber="75" precision="555" value="-2.87">W 2.87</geoCoordinate>
; pedipalp chela L 11.62, manus ventral L 3.51, manus
<geoCoordinate id="EE11506E201E1902FD8C8A77DFEFF8F3" box="[603,687,1884,1908]" degrees="1.93" direction="west" orientation="longitude" pageId="76" pageNumber="75" precision="555" value="-1.93">W 1.93</geoCoordinate>
, manus D 2.14, fixed finger L 6.79, movable finger L 7.98; pedipalp femur L 5.98,
<geoCoordinate id="EE11506E201E1902FB958879D9D5FAEC" box="[1090,1173,1363,1387]" degrees="2.03" direction="west" orientation="longitude" pageId="76" pageNumber="75" precision="555" value="-2.03">W 2.03</geoCoordinate>
, patella L 7.36,
<geoCoordinate id="EE11506E201E1902FA9F8879D8DBFAEC" box="[1352,1435,1363,1387]" degrees="2.55" direction="west" orientation="longitude" pageId="76" pageNumber="75" precision="555" value="-2.55">W 2.55</geoCoordinate>
; pectine L 7.83, mid-pectine sensillar margin L 0.735; leg III femur L 6.87; leg III patella L 5.79, D 1.825; chela
<emphasis id="B951EABB201E1902FCE88884DEC5FA41" box="[831,901,1454,1478]" italics="true" pageId="76" pageNumber="75">db–est</emphasis>
(left/ right) distance
<geoCoordinate id="EE11506E201E1902FBA08884D9F3FA41" box="[1143,1203,1454,1478]" degrees="0.292" direction="north" orientation="latitude" pageId="76" pageNumber="75" precision="55" value="0.292">0.292</geoCoordinate>
/
<geoCoordinate id="EE11506E201E1902FB118884D845FA41" box="[1222,1285,1454,1478]" degrees="0.958" direction="east" orientation="longitude" pageId="76" pageNumber="75" precision="55" value="0.958">0.958</geoCoordinate>
; pectine teeth (left/ right) 35/ 35.
</paragraph>
<paragraph id="8B9A36A9201E1902FCB888C6DEC9F8F3" blockId="76.[831,1441,1332,1908]" pageId="76" pageNumber="75">
<emphasis id="B951EABB201E1902FCB888C6D843F983" bold="true" box="[879,1283,1516,1540]" pageId="76" pageNumber="75">
Female,
<collectingCountry id="F3327639201E1902FC0588C6D953F983" box="[978,1043,1516,1540]" name="Israel" pageId="76" pageNumber="75">Israel</collectingCountry>
(
<specimenCode id="DB839ED2201E1902FBF388C6D9DEF983" box="[1060,1182,1516,1540]" collectionCode="NHMB" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="76" pageNumber="75">NHMB 17</specimenCode>
a) (mm).
</emphasis>
<collectorName id="26D0537F201E1902FADA88C6D820F983" box="[1293,1376,1516,1540]" pageId="76" pageNumber="75">Total L</collectorName>
76.0; metasoma + telson L 44.0; carapace L 8.09,
<geoCoordinate id="EE11506E201E1902FA928B21D8DBF9A4" box="[1349,1435,1547,1571]" degrees="8.91" direction="west" orientation="longitude" pageId="76" pageNumber="75" precision="555" value="-8.91">W 8.91</geoCoordinate>
, carapace preocular L 3.82; metasomal segments (L/ D /W) I 5.70/ 4.30/4.83,
<specimenCode id="DB839ED2201E1902FB808B63D9E8F9E7" box="[1111,1192,1608,1632]" collectionCode="II" pageId="76" pageNumber="75">II 6.44</specimenCode>
/ 4.15/4.30,
<specimenCode id="DB839ED2201E1902FAE98B63D8D8F9E7" box="[1342,1432,1608,1632]" collectionCode="III" pageId="76" pageNumber="75">III 6.48</specimenCode>
/ 4.05/4.10, IV 7.53/ 3.82/3.84,
<specimenCode id="DB839ED2201E1902FB598B4DD999F9F8" box="[1166,1241,1639,1663]" collectionCode="V" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="76" pageNumber="75" type="Museum">V 9.06</specimenCode>
/ 3.49/4.06; telson L 8.41; vesicle L 4.53, D 3.28,
<geoCoordinate id="EE11506E201E1902FB588BACD9A2F91A" box="[1167,1250,1669,1693]" degrees="3.49" direction="west" orientation="longitude" pageId="76" pageNumber="75" precision="555" value="-3.49">W 3.49</geoCoordinate>
; pedipalp chela L 14.34, manus ventral L 4.28, manus
<geoCoordinate id="EE11506E201E1902FB0E8B8FD86EF93B" box="[1241,1326,1700,1724]" degrees="2.41" direction="west" orientation="longitude" pageId="76" pageNumber="75" precision="555" value="-2.41">W 2.41</geoCoordinate>
, manus D 2.56, fixed finger L 8.20, movable finger L 10.03; pedipalp femur L 6.88,
<geoCoordinate id="EE11506E201E1902FB958BC8D9D5F97E" box="[1090,1173,1761,1785]" degrees="2.23" direction="west" orientation="longitude" pageId="76" pageNumber="75" precision="555" value="-2.23">W 2.23</geoCoordinate>
, patella L 8.32,
<geoCoordinate id="EE11506E201E1902FA9F8BC8D8DAF97E" box="[1352,1434,1761,1785]" degrees="3.05" direction="west" orientation="longitude" pageId="76" pageNumber="75" precision="555" value="-3.05">W 3.05</geoCoordinate>
; pectine L 8.83, mid-pectine sensillar margin L 0.653; leg III femur L 8.15; leg III patella L 6.92, D 2.00; chela
<emphasis id="B951EABB201E1902FCE88A17DEC5F8D2" box="[831,901,1853,1877]" italics="true" pageId="76" pageNumber="75">db–est</emphasis>
(left/ right)
<geoCoordinate id="EE11506E201E1902FBD08A17D903F8D2" box="[1031,1091,1853,1877]" degrees="0.354" direction="north" orientation="latitude" pageId="76" pageNumber="75" precision="55" value="0.354">0.354</geoCoordinate>
/
<geoCoordinate id="EE11506E201E1902FB868A17D9D0F8D2" box="[1105,1168,1853,1877]" degrees="0.417" direction="east" orientation="longitude" pageId="76" pageNumber="75" precision="55" value="0.417">0.417</geoCoordinate>
; pectine teeth (left/ right) 30/ 30.
</paragraph>
</subSubSection>
<caption id="DF5A6621201F1903FF17883CDF48FA17" ID-DOI="http://doi.org/10.5281/zenodo.6505747" ID-Zenodo-Dep="6505747" httpUri="https://zenodo.org/record/6505747/files/figure.png" pageId="77" pageNumber="76" startId="77.[192,267,1302,1326]" targetBox="[200,1431,198,1264]" targetPageId="77">
<paragraph id="8B9A36A9201F1903FF17883CDF48FA17" blockId="77.[192,1441,1302,1424]" pageId="77" pageNumber="76">
<emphasis id="B951EABB201F1903FF17883CDC76FAA9" bold="true" box="[192,310,1302,1326]" pageId="77" pageNumber="76">Figure 59:</emphasis>
Chelicerae of
<taxonomicName id="4C254D2A201F1903FE128832DF39FAAB" box="[453,633,1303,1325]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="77" pageNumber="76" phylum="Arthropoda" rank="species" species="undefined-a">
<emphasis id="B951EABB201F1903FE128832DF4EFAAA" box="[453,526,1304,1325]" italics="true" pageId="77" pageNumber="76">Leiurus</emphasis>
species.
<emphasis id="B951EABB201F1903FDB0883DDF39FAAB" bold="true" box="[615,633,1303,1324]" pageId="77" pageNumber="76">A</emphasis>
</taxonomicName>
<emphasis id="B951EABB201F1903FDAE883DDFDBFAAB" bold="true" box="[633,667,1303,1324]" pageId="77" pageNumber="76">, B.</emphasis>
<taxonomicName id="4C254D2A201F1903FD768832DE77FAAA" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" box="[673,823,1304,1325]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="77" pageNumber="76" phylum="Arthropoda" rank="species" species="macroctenus" status="sp. nov.">
<emphasis id="B951EABB201F1903FD768832DFEEFAAA" box="[673,686,1304,1325]" italics="true" pageId="77" pageNumber="76">L</emphasis>
.
<emphasis id="B951EABB201F1903FD6B8832DE77FAAA" box="[700,823,1304,1325]" italics="true" pageId="77" pageNumber="76">macroctenus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C0201F1903FCEA8832DE32FAAA" box="[829,882,1304,1325]" pageId="77" pageNumber="76" rank="species">sp. n.</taxonomicNameLabel>
, paratype male, Thumrait, Oman. Right chelicera, ventral (
<emphasis id="B951EABB201F1903FF1F881ADD99FAC2" bold="true" box="[200,217,1328,1349]" pageId="77" pageNumber="76">A</emphasis>
) and dorsal (
<emphasis id="B951EABB201F1903FE8C881ADC2BFAC2" bold="true" box="[347,363,1328,1349]" pageId="77" pageNumber="76">B</emphasis>
) aspect.
<emphasis id="B951EABB201F1903FE14881ADCB8FAC2" bold="true" box="[451,504,1328,1349]" pageId="77" pageNumber="76">C, D.</emphasis>
<taxonomicName id="4C254D2A201F1903FE28881BDF2DFAC1" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" box="[511,621,1328,1350]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="77" pageNumber="76" phylum="Arthropoda" rank="species" species="haenggii" status="sp. nov.">
<emphasis id="B951EABB201F1903FE28881BDF4CFAC1" box="[511,524,1329,1350]" italics="true" pageId="77" pageNumber="76">L</emphasis>
.
<emphasis id="B951EABB201F1903FDCE881ADF2DFAC1" box="[537,621,1328,1350]" italics="true" pageId="77" pageNumber="76">haenggii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C0201F1903FDA3881ADFE9FAC2" box="[628,681,1328,1349]" pageId="77" pageNumber="76" rank="species">sp. n.</taxonomicNameLabel>
, paratype male, Jabal Qara, Oman. Right chelicera, ventral (
<emphasis id="B951EABB201F1903FB23881AD845FAC2" bold="true" box="[1268,1285,1328,1349]" pageId="77" pageNumber="76">C</emphasis>
) and dorsal (
<emphasis id="B951EABB201F1903FA5F881AD8D9FAC2" bold="true" box="[1416,1433,1328,1349]" pageId="77" pageNumber="76">D</emphasis>
) aspect.
<emphasis id="B951EABB201F1903FEDC8863DC7FFAD9" bold="true" box="[267,319,1353,1374]" pageId="77" pageNumber="76">E, F.</emphasis>
<taxonomicName id="4C254D2A201F1903FE9E8863DFEEFAD9" authority="(Ehrenberg, 1829)" baseAuthorityName="Ehrenberg" baseAuthorityYear="1829" box="[329,686,1352,1374]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="77" pageNumber="76" phylum="Arthropoda" rank="species" species="brachycentrus">
<emphasis id="B951EABB201F1903FE9E8863DC16FAD9" box="[329,342,1353,1374]" italics="true" pageId="77" pageNumber="76">L</emphasis>
.
<emphasis id="B951EABB201F1903FEB18862DCB0FAD9" box="[358,496,1352,1374]" italics="true" pageId="77" pageNumber="76">brachycentrus</emphasis>
(Ehrenberg, 1829)
</taxonomicName>
, female, Al Mansuriah, Yemen. Right chelicera, ventral (
<emphasis id="B951EABB201F1903FB278863D840FAD9" bold="true" box="[1264,1280,1353,1374]" pageId="77" pageNumber="76">E</emphasis>
) and dorsal (
<emphasis id="B951EABB201F1903FA5D8863D8D9FAD9" bold="true" box="[1418,1433,1353,1374]" pageId="77" pageNumber="76">F</emphasis>
) aspect.
<emphasis id="B951EABB201F1903FEDE8848DC02FAF0" bold="true" box="[265,322,1378,1399]" pageId="77" pageNumber="76">G, H.</emphasis>
<taxonomicName id="4C254D2A201F1903FE9F8848DCE7FAF0" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" box="[328,423,1377,1399]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="77" pageNumber="76" phylum="Arthropoda" rank="species" species="heberti" status="sp. nov.">
<emphasis id="B951EABB201F1903FE9F8848DC15FAF0" box="[328,341,1378,1399]" italics="true" pageId="77" pageNumber="76">L</emphasis>
.
<emphasis id="B951EABB201F1903FEB4884BDCE7FAF0" box="[355,423,1377,1399]" italics="true" pageId="77" pageNumber="76">heberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C0201F1903FE798848DCA4FAF0" box="[430,484,1378,1399]" pageId="77" pageNumber="76" rank="species">sp. n.</taxonomicNameLabel>
, holotype male, Wadi Andur, Oman. Right chelicera, ventral (
<emphasis id="B951EABB201F1903FB918848D919FAF0" bold="true" box="[1094,1113,1378,1399]" pageId="77" pageNumber="76">G</emphasis>
) and dorsal (
<emphasis id="B951EABB201F1903FB0B8848D9AFFAF0" bold="true" box="[1244,1263,1378,1399]" pageId="77" pageNumber="76">H</emphasis>
) aspect. Scale bar in
<emphasis id="B951EABB201F1903FF0E8850DDAAFA08" bold="true" box="[217,234,1402,1423]" pageId="77" pageNumber="76">A</emphasis>
: 1 mm (also applies to
<emphasis id="B951EABB201F1903FE1C8850DCBBFA08" bold="true" box="[459,507,1402,1423]" pageId="77" pageNumber="76">B–H</emphasis>
).
</paragraph>
</caption>
<subSubSection id="C33F6522201F1903FF1788EEDF8EFA5C" box="[192,718,1476,1500]" pageId="77" pageNumber="76" type="distribution">
<paragraph id="8B9A36A9201F1903FF1788EEDF8EFA5C" blockId="77.[192,718,1476,1500]" box="[192,718,1476,1500]" pageId="77" pageNumber="76">
DISTRIBUTION.
<collectingCountry id="F3327639201F1903FE5F88EEDC86FA5B" box="[392,454,1476,1500]" name="Israel" pageId="77" pageNumber="76">Israel</collectingCountry>
,
<collectingCountry id="F3327639201F1903FE0688EEDF5AFA5B" box="[465,538,1476,1500]" name="Jordan" pageId="77" pageNumber="76">Jordan</collectingCountry>
,
<collectingCountry id="F3327639201F1903FDF288EEDF20FA5B" box="[549,608,1476,1500]" name="Syria" pageId="77" pageNumber="76">Syria</collectingCountry>
,
<collectingCountry id="F3327639201F1903FDBD88EEDF87FA5B" box="[618,711,1476,1500]" name="Lebanon" pageId="77" pageNumber="76">Lebanon</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F6522201F191FFF178B2BDCDCF930" lastPageId="81" lastPageNumber="80" pageId="77" pageNumber="76" type="discussion">
<paragraph id="8B9A36A9201F1903FF178B2BDF25F913" blockId="77.[192,802,1537,1898]" pageId="77" pageNumber="76">
REMARKS.
<bibRefCitation id="EFB44B58201F1903FE808B2BDCB2F99E" author="Birula" box="[343,498,1537,1561]" firstAuthor="Birula" journalOrPublisher="Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien" pageId="77" pageNumber="76" pagination="121 - 152" part="117" refId="ref59140" refString="BIRULA, A. 1908. Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 117 2 (1): 121 - 152." title="Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae" type="journal article" year="1908">Birula (1908)</bibRefCitation>
separated
<emphasis id="B951EABB201F1903FDA08B28DFC6F99E" box="[631,646,1538,1561]" italics="true" pageId="77" pageNumber="76">L</emphasis>
.
<emphasis id="B951EABB201F1903FD4B8B28DFE9F99E" box="[668,681,1538,1561]" italics="true" pageId="77" pageNumber="76">q</emphasis>
.
<taxonomicName id="4C254D2A201F1903FD178B2BDE62F99E" authorityName=": Levy &amp; Amitai" authorityYear="1980" box="[704,802,1537,1561]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="77" pageNumber="76" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB201F1903FD178B2BDE62F99E" box="[704,802,1537,1561]" italics="true" pageId="77" pageNumber="76">hebraeus</emphasis>
</taxonomicName>
from the nominotypic subspecies by 11 characters. We assessed these characters by comparing
<specimenCount id="9D23FD20201F1903FD4C8B14DFBBF9D1" box="[667,763,1598,1622]" pageId="77" pageNumber="76" type="adult">20 adult</specimenCount>
<taxonomicName id="4C254D2A201F1903FCDB8B15DC62F9F2" authorityName=": Levy &amp; Amitai" authorityYear="1980" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="77" pageNumber="76" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB201F1903FCDB8B15DE5BF9D1" box="[780,795,1599,1622]" italics="true" pageId="77" pageNumber="76">L</emphasis>
.
<emphasis id="B951EABB201F1903FF178B77DC62F9F2" box="[192,290,1629,1653]" italics="true" pageId="77" pageNumber="76">hebraeus</emphasis>
</taxonomicName>
from
<collectingCountry id="F3327639201F1903FEAE8B77DCF4F9F2" box="[377,436,1629,1653]" name="Israel" pageId="77" pageNumber="76">Israel</collectingCountry>
and
<collectingCountry id="F3327639201F1903FE298B77DF05F9F2" box="[510,581,1629,1653]" name="Jordan" pageId="77" pageNumber="76">Jordan</collectingCountry>
with
<specimenCount id="9D23FD20201F1903FD4F8B77DFBAF9F2" box="[664,762,1629,1653]" pageId="77" pageNumber="76" type="adult">26 adult</specimenCount>
<taxonomicName id="4C254D2A201F1903FCDB8B74DC26F913" class="Insecta" family="Carabidae" genus="Leirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="76" phylum="Arthropoda" rank="species" species="quinquestriatus">
<emphasis id="B951EABB201F1903FCDB8B74DE5BF9F2" box="[780,795,1630,1653]" italics="true" pageId="77" pageNumber="76">L</emphasis>
.
<emphasis id="B951EABB201F1903FF178B57DC26F913" box="[192,358,1661,1684]" italics="true" pageId="77" pageNumber="76">quinquestriatus</emphasis>
</taxonomicName>
from
<collectingCountry id="F3327639201F1903FE708B56DCA7F913" box="[423,487,1660,1684]" name="Egypt" pageId="77" pageNumber="76">Egypt</collectingCountry>
and
<collectingCountry id="F3327639201F1903FDCC8B56DF21F913" box="[539,609,1660,1684]" name="Sudan" pageId="77" pageNumber="76">Sudan</collectingCountry>
:
</paragraph>
<paragraph id="8B9A36A9201F1903FF278BB0DF8DF8AA" blockId="77.[192,802,1537,1898]" pageId="77" pageNumber="76">
(1) weaker carapace carination and granulation, (2) granulated ocular tubercle, (3) more coarsely shagreened tergites, (4) non-granulated intercarinal surfaces on the metasoma: (1)
<emphasis id="B951EABB201F1903FEBD8BD0DC37F889" box="[362,375,1786,1806]" italics="true" pageId="77" pageNumber="76">–</emphasis>
(4) were unreliable when granulation was visualized in fine detail by UV fluorescence;
</paragraph>
<paragraph id="8B9A36A9201F1903FF278A1EDEF3F99E" blockId="77.[192,802,1537,1898]" lastBlockId="77.[831,1441,1476,1898]" pageId="77" pageNumber="76">
(5) ventromedian carinae of metasoma II
<emphasis id="B951EABB201F1903FD698A1DDF8BF8CC" box="[702,715,1847,1867]" italics="true" pageId="77" pageNumber="76">–</emphasis>
III with denticles increasing in size posteriorly (vs. uniform in size): this was confirmed for
<taxonomicName id="4C254D2A201F1903FB7288EFD86DFA5B" authorityName=": Levy &amp; Amitai" authorityYear="1980" box="[1189,1325,1476,1500]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="77" pageNumber="76" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB201F1903FB7288EFD9F4FA5B" box="[1189,1204,1477,1500]" italics="true" pageId="77" pageNumber="76">L</emphasis>
.
<emphasis id="B951EABB201F1903FB1C88EED86DFA5B" box="[1227,1325,1476,1500]" italics="true" pageId="77" pageNumber="76">hebraeus</emphasis>
</taxonomicName>
, but also observed in some
<taxonomicName id="4C254D2A201F1903FBD388C9D988FA7D" box="[1028,1224,1507,1530]" class="Insecta" family="Carabidae" genus="Leirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="76" phylum="Arthropoda" rank="species" species="quinquestriatus">
<emphasis id="B951EABB201F1903FBD388C9D953FA7D" box="[1028,1043,1507,1530]" italics="true" pageId="77" pageNumber="76">L</emphasis>
.
<emphasis id="B951EABB201F1903FBF588C9D988FA7D" box="[1058,1224,1507,1530]" italics="true" pageId="77" pageNumber="76">quinquestriatus</emphasis>
</taxonomicName>
, and hence was not diagnostic;
</paragraph>
<paragraph id="8B9A36A9201F1903FCB88B0AD811F935" blockId="77.[831,1441,1476,1898]" pageId="77" pageNumber="76">
(6) smaller number of denticles (15
<emphasis id="B951EABB201F1903FAEA8B09D80AF9B0" box="[1341,1354,1571,1591]" italics="true" pageId="77" pageNumber="76">–</emphasis>
17) on ventromedian carinae of metasoma III (vs. 18
<emphasis id="B951EABB201F1903FAEF8B68D805F9D1" box="[1336,1349,1602,1622]" italics="true" pageId="77" pageNumber="76">–</emphasis>
27): this was well supported with no overlap, i.e. 46/46 carinae of
<taxonomicName id="4C254D2A201F1903FCE88B57D94AF913" box="[831,1034,1660,1684]" class="Insecta" family="Carabidae" genus="Leirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="76" phylum="Arthropoda" rank="species" species="quinquestriatus">
<emphasis id="B951EABB201F1903FCE88B57DE0EF913" box="[831,846,1661,1684]" italics="true" pageId="77" pageNumber="76">L</emphasis>
.
<emphasis id="B951EABB201F1903FCB38B57D94AF913" box="[868,1034,1661,1684]" italics="true" pageId="77" pageNumber="76">quinquestriatus</emphasis>
</taxonomicName>
with 19 or more denticles, 24/24 carinae of
<taxonomicName id="4C254D2A201F1903FC788BB1D96DF935" authorityName=": Levy &amp; Amitai" authorityYear="1980" box="[943,1069,1690,1714]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="77" pageNumber="76" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB201F1903FC788BB1DEFEF935" box="[943,958,1691,1714]" italics="true" pageId="77" pageNumber="76">L</emphasis>
.
<emphasis id="B951EABB201F1903FC1C8BB0D96DF935" box="[971,1069,1690,1714]" italics="true" pageId="77" pageNumber="76">hebraeus</emphasis>
</taxonomicName>
with 18 or fewer denticles;
</paragraph>
<paragraph id="8B9A36A9201F1903FCB88B93D840F889" blockId="77.[831,1441,1476,1898]" pageId="77" pageNumber="76">
(7) lower range of denticles (18
<emphasis id="B951EABB201F1903FB168B96D98EF957" box="[1217,1230,1724,1744]" italics="true" pageId="77" pageNumber="76">–</emphasis>
31) on ventrolateral carinae of metasoma V (vs. 23
<emphasis id="B951EABB201F1903FB778BF1D9EDF968" box="[1184,1197,1755,1775]" italics="true" pageId="77" pageNumber="76">–</emphasis>
38): due to the broad overlap of ranges, this was not diagnostic;
</paragraph>
<paragraph id="8B9A36A9201F1903FCB88A3FD809F8ED" blockId="77.[831,1441,1476,1898]" pageId="77" pageNumber="76">
(8) more stout pedipalp and metasomal segments: this was supported by smaller morphometric ratios, pedipalp patella L/W and metasoma III
<emphasis id="B951EABB201F1903FB368A7CD9AEF8ED" box="[1249,1262,1878,1898]" italics="true" pageId="77" pageNumber="76">–</emphasis>
IV L/W;
</paragraph>
<caption id="DF5A6621201C1900FF178B12D9DFF9C8" ID-DOI="http://doi.org/10.5281/zenodo.6505749" ID-Zenodo-Dep="6505749" box="[192,1183,1592,1616]" httpUri="https://zenodo.org/record/6505749/files/figure.png" pageId="78" pageNumber="77" startId="78.[192,267,1592,1616]" targetBox="[191,1438,167,1561]" targetPageId="78">
<paragraph id="8B9A36A9201C1900FF178B12D9DFF9C8" blockId="78.[192,1183,1592,1616]" box="[192,1183,1592,1616]" pageId="78" pageNumber="77">
<emphasis id="B951EABB201C1900FF178B12DC75F9D7" bold="true" box="[192,309,1592,1616]" pageId="78" pageNumber="77">Figure 60:</emphasis>
<taxonomicName id="4C254D2A201C1900FEEB8B10DF31F9C8" authority="(Birula, 1908)" baseAuthorityName="Birula" baseAuthorityYear="1908" box="[316,625,1593,1615]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="78" pageNumber="77" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB201C1900FEEB8B10DCA3F9C8" box="[316,483,1593,1615]" italics="true" pageId="78" pageNumber="77">Leiurus hebraeus</emphasis>
(
<bibRefCitation id="EFB44B58201C1900FE278B13DF2AF9C8" author="Birula" box="[496,618,1593,1615]" firstAuthor="Birula" journalOrPublisher="Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien" pageId="78" pageNumber="77" pagination="121 - 152" part="117" refId="ref59140" refString="BIRULA, A. 1908. Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 117 2 (1): 121 - 152." title="Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae" type="journal article" year="1908">Birula, 1908</bibRefCitation>
)
</taxonomicName>
stat. n., male. Habitus, dorsal aspect. Israel (NHMB 17a).
</paragraph>
</caption>
<paragraph id="8B9A36A9201C1900FF278BA8DDBFF892" blockId="78.[192,802,1666,1905]" pageId="78" pageNumber="77">
(9) metasoma I only barely wider than metasoma II (vs. significantly wider): this was not diagnostic, due to significant overlap of the ratio metasoma I W/ metasoma II W (
<taxonomicName id="4C254D2A201C1900FEDD8BF5DC91F971" box="[266,465,1759,1782]" class="Insecta" family="Carabidae" genus="Leirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="77" phylum="Arthropoda" rank="species" species="quinquestriatus">
<emphasis id="B951EABB201C1900FEDD8BF5DC59F971" box="[266,281,1759,1782]" italics="true" pageId="78" pageNumber="77">L</emphasis>
.
<emphasis id="B951EABB201C1900FEFC8BF5DC91F971" box="[299,465,1759,1782]" italics="true" pageId="78" pageNumber="77">quinquestriatus</emphasis>
</taxonomicName>
1.10
<emphasis id="B951EABB201C1900FDDC8BC8DF58F971" box="[523,536,1762,1782]" italics="true" pageId="78" pageNumber="77">–</emphasis>
1.21,
<taxonomicName id="4C254D2A201C1900FD8E8BF5DF9BF971" baseAuthorityName="Birula" baseAuthorityYear="1908" box="[601,731,1758,1782]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="78" pageNumber="77" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB201C1900FD8E8BF5DF28F971" box="[601,616,1759,1782]" italics="true" pageId="78" pageNumber="77">L</emphasis>
.
<emphasis id="B951EABB201C1900FDAE8BF4DF9BF971" box="[633,731,1758,1782]" italics="true" pageId="78" pageNumber="77">hebraeus</emphasis>
</taxonomicName>
1.09
<emphasis id="B951EABB201C1900FCC28BC8DE62F971" box="[789,802,1762,1782]" italics="true" pageId="78" pageNumber="77">–</emphasis>
1.14);
</paragraph>
<paragraph id="8B9A36A9201C1900FF278A36D873F892" blockId="78.[192,802,1666,1905]" lastBlockId="78.[831,1440,1666,1905]" pageId="78" pageNumber="77">
(10) telson vesicle longer than aculeus (chord length), vesicle width equal to aculeus length (vs. about the same length or shorter, and width &lt;aculeus length): there was a trend for
<taxonomicName id="4C254D2A201C1900FB868BA9D85CF91D" box="[1105,1308,1667,1690]" class="Insecta" family="Carabidae" genus="Leirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="77" phylum="Arthropoda" rank="species" species="quinquestriatus">
<emphasis id="B951EABB201C1900FB868BA9D920F91D" box="[1105,1120,1667,1690]" italics="true" pageId="78" pageNumber="77">L</emphasis>
.
<emphasis id="B951EABB201C1900FBA18BA9D85CF91D" box="[1142,1308,1667,1690]" italics="true" pageId="78" pageNumber="77">quinquestriatus</emphasis>
</taxonomicName>
to have a relatively longer aculeus and a shorter, more bulbous vesicle than
<taxonomicName id="4C254D2A201C1900FC0E8BEAD922F950" baseAuthorityName="Birula" baseAuthorityYear="1908" box="[985,1122,1727,1751]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="78" pageNumber="77" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB201C1900FC0E8BEADEA8F950" box="[985,1000,1728,1751]" italics="true" pageId="78" pageNumber="77">L</emphasis>
.
<emphasis id="B951EABB201C1900FBD78B95D922F950" box="[1024,1122,1727,1751]" italics="true" pageId="78" pageNumber="77">hebraeus</emphasis>
</taxonomicName>
, but there was significant overlap in the ratio of vesicle L/ telson L (
<taxonomicName id="4C254D2A201C1900FA938BF5DEF5F892" class="Insecta" family="Carabidae" genus="Leirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="77" phylum="Arthropoda" rank="species" species="quinquestriatus">
<emphasis id="B951EABB201C1900FA938BF5D813F971" box="[1348,1363,1759,1782]" italics="true" pageId="78" pageNumber="77">L</emphasis>
.
<emphasis id="B951EABB201C1900FABF8BF5DEF5F892" italics="true" pageId="78" pageNumber="77">quinquestriatus</emphasis>
</taxonomicName>
0.48
<emphasis id="B951EABB201C1900FC3C8A2ADEB8F893" box="[1003,1016,1792,1812]" italics="true" pageId="78" pageNumber="77">–</emphasis>
0.56,
<taxonomicName id="4C254D2A201C1900FBE38BD4D9F2F892" baseAuthorityName="Birula" baseAuthorityYear="1908" box="[1076,1202,1789,1813]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="78" pageNumber="77" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB201C1900FBE38BD4D903F892" box="[1076,1091,1790,1813]" italics="true" pageId="78" pageNumber="77">L</emphasis>
.
<emphasis id="B951EABB201C1900FB878BD7D9F2F892" box="[1104,1202,1789,1813]" italics="true" pageId="78" pageNumber="77">hebraeus</emphasis>
</taxonomicName>
0.52
<emphasis id="B951EABB201C1900FB308A2AD9B4F893" box="[1255,1268,1792,1812]" italics="true" pageId="78" pageNumber="77">–</emphasis>
0.61);
</paragraph>
<paragraph id="8B9A36A9201C1901FCB88A36DFA5F890" blockId="78.[831,1440,1666,1905]" lastBlockId="79.[192,802,1668,1907]" lastPageId="79" lastPageNumber="78" pageId="78" pageNumber="77">
(11) pedipalp movable finger short, less than twice manus length, with 12 denticle subrows (vs. long, slender,&gt; twice manus length, and 13 subrows): this was not diagnostic, because although the movable finger was on average relatively shorter in
<taxonomicName id="4C254D2A201D1901FD8F8B8EDF9AF93C" baseAuthorityName="Birula" baseAuthorityYear="1908" box="[600,730,1699,1723]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="79" pageNumber="78" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB201D1901FD8F8B8EDF27F93C" box="[600,615,1700,1723]" italics="true" pageId="79" pageNumber="78">L</emphasis>
.
<emphasis id="B951EABB201D1901FDAF8B89DF9AF93C" box="[632,730,1699,1723]" italics="true" pageId="79" pageNumber="78">hebraeus</emphasis>
</taxonomicName>
vs.
<taxonomicName id="4C254D2A201D1901FCDB8B8EDC26F95D" class="Insecta" family="Carabidae" genus="Leirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="78" phylum="Arthropoda" rank="species" species="quinquestriatus">
<emphasis id="B951EABB201D1901FCDB8B8EDE5BF93C" box="[780,795,1700,1723]" italics="true" pageId="79" pageNumber="78">L</emphasis>
.
<emphasis id="B951EABB201D1901FF178BE9DC26F95D" box="[192,358,1731,1754]" italics="true" pageId="79" pageNumber="78">quinquestriatus</emphasis>
</taxonomicName>
, there was broad overlap in the ratio movable finger L (chord)/ manus ventral L, and movable fingers had either 12 or 13 subrows in both species.
</paragraph>
<caption id="DF5A6621201D1901FF178B13D9E7F9D6" ID-DOI="http://doi.org/10.5281/zenodo.6505751" ID-Zenodo-Dep="6505751" box="[192,1191,1593,1617]" httpUri="https://zenodo.org/record/6505751/files/figure.png" pageId="79" pageNumber="78" startId="79.[192,267,1593,1617]" targetBox="[191,1439,167,1564]" targetPageId="79">
<paragraph id="8B9A36A9201D1901FF178B13D9E7F9D6" blockId="79.[192,1191,1593,1617]" box="[192,1191,1593,1617]" pageId="79" pageNumber="78">
<emphasis id="B951EABB201D1901FF178B13DC76F9D6" bold="true" box="[192,310,1593,1617]" pageId="79" pageNumber="78">Figure 61:</emphasis>
<taxonomicName id="4C254D2A201D1901FEEB8B16DF31F9D6" authority="(Birula, 1908)" baseAuthorityName="Birula" baseAuthorityYear="1908" box="[316,625,1595,1617]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="79" pageNumber="78" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB201D1901FEEB8B16DCA3F9D6" box="[316,483,1595,1617]" italics="true" pageId="79" pageNumber="78">Leiurus hebraeus</emphasis>
(
<bibRefCitation id="EFB44B58201D1901FE278B11DF2AF9D6" author="Birula" box="[496,618,1595,1617]" firstAuthor="Birula" journalOrPublisher="Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien" pageId="79" pageNumber="78" pagination="121 - 152" part="117" refId="ref59140" refString="BIRULA, A. 1908. Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 117 2 (1): 121 - 152." title="Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae" type="journal article" year="1908">Birula, 1908</bibRefCitation>
)
</taxonomicName>
stat. n., male. Habitus, ventral aspect. Israel (NHMB 17a).
</paragraph>
</caption>
<paragraph id="8B9A36A9201D1901FF178A34DCA9F8F4" blockId="79.[192,802,1668,1907]" pageId="79" pageNumber="78">
<bibRefCitation id="EFB44B58201D1901FF178A34DCCDF8B1" author="Levy" box="[192,397,1822,1846]" etAl="et al." firstAuthor="Levy" pageId="79" pageNumber="78" pagination="231 - 242" refId="ref62217" refString="LEVY, G., P. AMITAI &amp; A. SHULOV. 1970. Leiurus quinquestriatus hebraeus (Birula, 1908) (Scorpiones, Buthidae) and its systematic position. Israel Journal of Zoology, 19 (4): 231 - 242." type="journal article" year="1970">Levy et al. (1970)</bibRefCitation>
accepted characters (5) and (10) of
<bibRefCitation id="EFB44B58201D1901FF178A16DC18F8D3" author="Birula" box="[192,344,1852,1876]" firstAuthor="Birula" journalOrPublisher="Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien" pageId="79" pageNumber="78" pagination="121 - 152" part="117" refId="ref59140" refString="BIRULA, A. 1908. Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 117 2 (1): 121 - 152." title="Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae" type="journal article" year="1908">Birula (1908)</bibRefCitation>
, and proposed 3 additional characters to differentiate
<emphasis id="B951EABB201D1901FE9E8A76DC18F8F4" box="[329,344,1884,1907]" italics="true" pageId="79" pageNumber="78">L</emphasis>
.
<emphasis id="B951EABB201D1901FEB28A76DC32F8F4" box="[357,370,1884,1907]" italics="true" pageId="79" pageNumber="78">q</emphasis>
.
<taxonomicName id="4C254D2A201D1901FE578A71DCA2F8F4" baseAuthorityName="Birula" baseAuthorityYear="1908" box="[384,482,1883,1907]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="79" pageNumber="78" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB201D1901FE578A71DCA2F8F4" box="[384,482,1883,1907]" italics="true" pageId="79" pageNumber="78">hebraeus</emphasis>
</taxonomicName>
:
</paragraph>
<paragraph id="8B9A36A9201D1901FCB88BAED94CF890" blockId="79.[831,1441,1668,1907]" pageId="79" pageNumber="78">
(12) posterior median area of carapace granulated, with paired granule arcs (vs. smooth or sparsely granulated): this difference was confirmed for most specimens, although some
<taxonomicName id="4C254D2A201D1901FBFA8BCBD9B1F97F" box="[1069,1265,1761,1784]" class="Insecta" family="Carabidae" genus="Leirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="78" phylum="Arthropoda" rank="species" species="quinquestriatus">
<emphasis id="B951EABB201D1901FBFA8BCBD97CF97F" box="[1069,1084,1761,1784]" italics="true" pageId="79" pageNumber="78">L</emphasis>
.
<emphasis id="B951EABB201D1901FB9C8BCBD9B1F97F" box="[1099,1265,1761,1784]" italics="true" pageId="79" pageNumber="78">quinquestriatus</emphasis>
</taxonomicName>
have weak arcs of fine granulation;
</paragraph>
<paragraph id="8B9A36A9201D191EFCB88A34DF5FFA36" blockId="79.[831,1441,1668,1907]" lastBlockId="80.[192,802,1403,1918]" lastPageId="80" lastPageNumber="79" pageId="79" pageNumber="78">
(13) lateral flanks of tergite V with irregular granulation (vs. short rows of granules, resembling carinae): this rather subjective character was quite variable, and short granule rows could be discerned or not in both
<taxonomicName id="4C254D2A2002191EFCDB8856DC62FA35" baseAuthorityName="Birula" baseAuthorityYear="1908" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="80" pageNumber="79" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB2002191EFCDB8856DE5BFA14" box="[780,795,1404,1427]" italics="true" pageId="80" pageNumber="79">L</emphasis>
.
<emphasis id="B951EABB2002191EFF1788B0DC62FA35" box="[192,290,1434,1458]" italics="true" pageId="80" pageNumber="79">hebraeus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C254D2A2002191EFE8188B1DF58FA35" box="[342,536,1434,1458]" class="Insecta" family="Carabidae" genus="Leirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="79" phylum="Arthropoda" rank="species" species="quinquestriatus">
<emphasis id="B951EABB2002191EFE8188B1DC25FA35" box="[342,357,1435,1458]" italics="true" pageId="80" pageNumber="79">L</emphasis>
.
<emphasis id="B951EABB2002191EFEA588B1DF58FA35" box="[370,536,1435,1458]" italics="true" pageId="80" pageNumber="79">quinquestriatus</emphasis>
</taxonomicName>
;
</paragraph>
<caption id="DF5A66212002191EFF17883DDC08FAC0" ID-DOI="http://doi.org/10.5281/zenodo.6505753" ID-Zenodo-Dep="6505753" httpUri="https://zenodo.org/record/6505753/files/figure.png" pageId="80" pageNumber="79" startId="80.[192,267,1303,1327]" targetBox="[204,1425,204,1261]" targetPageId="80">
<paragraph id="8B9A36A92002191EFF17883DDC08FAC0" blockId="80.[192,1440,1303,1351]" pageId="80" pageNumber="79">
<emphasis id="B951EABB2002191EFF17883DDC78FAA8" bold="true" box="[192,312,1303,1327]" pageId="80" pageNumber="79">Figure 62:</emphasis>
<taxonomicName id="4C254D2A2002191EFE968830DF3EFAA8" authority="(Birula, 1908)" baseAuthorityName="Birula" baseAuthorityYear="1908" box="[321,638,1305,1327]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="80" pageNumber="79" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB2002191EFE968830DCABFAA8" box="[321,491,1305,1327]" italics="true" pageId="80" pageNumber="79">Leiurus hebraeus</emphasis>
(
<bibRefCitation id="EFB44B582002191EFE2D8833DF37FAA8" author="Birula" box="[506,631,1305,1327]" firstAuthor="Birula" journalOrPublisher="Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien" pageId="80" pageNumber="79" pagination="121 - 152" part="117" refId="ref59140" refString="BIRULA, A. 1908. Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 117 2 (1): 121 - 152." title="Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae" type="journal article" year="1908">Birula, 1908</bibRefCitation>
)
</taxonomicName>
stat. n., male.
<emphasis id="B951EABB2002191EFCC38833DE6CFAA9" bold="true" box="[788,812,1305,1326]" pageId="80" pageNumber="79">A.</emphasis>
Carapace and tergites.
<emphasis id="B951EABB2002191EFBC18833D96CFAA9" bold="true" box="[1046,1068,1305,1326]" pageId="80" pageNumber="79">B.</emphasis>
Coxosternal area and sternites. Israel (NHMB 17a).
</paragraph>
</caption>
<paragraph id="8B9A36A92002191EFF278892DF16F9EE" blockId="80.[192,802,1403,1918]" pageId="80" pageNumber="79">
(14) weak or obsolete carinae on sternites (slightly more distinct on posterior segments) (vs. distinct, elevated strong carinae on sternites IV–VII): this difference applies to males, however females have weaker carinae and some
<taxonomicName id="4C254D2A2002191EFEE48B1EDCB8F9CC" box="[307,504,1588,1611]" class="Insecta" family="Carabidae" genus="Leirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="79" phylum="Arthropoda" rank="species" species="quinquestriatus">
<emphasis id="B951EABB2002191EFEE48B1EDC02F9CC" box="[307,322,1588,1611]" italics="true" pageId="80" pageNumber="79">L</emphasis>
.
<emphasis id="B951EABB2002191EFE858B1EDCB8F9CC" box="[338,504,1588,1611]" italics="true" pageId="80" pageNumber="79">quinquestriatus</emphasis>
</taxonomicName>
females can have weak to obsolete carination on sternites IV–V.
</paragraph>
<paragraph id="8B9A36A92002191FFF278B5ADCDCF930" blockId="80.[192,802,1403,1918]" lastBlockId="81.[192,802,1569,1719]" lastPageId="81" lastPageNumber="80" pageId="80" pageNumber="79">
Thus, from the above list, only characters (6), (14) (males only), and perhaps (12), were potential diagnostic characters separating
<taxonomicName id="4C254D2A2002191EFE1E8B85DF19F941" baseAuthorityName="Birula" baseAuthorityYear="1908" box="[457,601,1710,1734]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="80" pageNumber="79" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB2002191EFE1E8B85DC98F941" box="[457,472,1711,1734]" italics="true" pageId="80" pageNumber="79">L</emphasis>
.
<emphasis id="B951EABB2002191EFE208B84DF19F941" box="[503,601,1710,1734]" italics="true" pageId="80" pageNumber="79">hebraeus</emphasis>
</taxonomicName>
from
<taxonomicName id="4C254D2A2002191EFD6C8B85DC76F962" class="Insecta" family="Carabidae" genus="Leirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="79" phylum="Arthropoda" rank="species" species="quinquestriatus">
<emphasis id="B951EABB2002191EFD6C8B85DF8AF941" box="[699,714,1711,1734]" italics="true" pageId="80" pageNumber="79">L</emphasis>
.
<emphasis id="B951EABB2002191EFD3E8B85DC76F962" italics="true" pageId="80" pageNumber="79">quinquestriatus</emphasis>
</taxonomicName>
. Four examined adult and subadult males from southern Sinai resembled
<taxonomicName id="4C254D2A2002191EFDE48BC6DFBEF884" box="[563,766,1772,1795]" class="Insecta" family="Carabidae" genus="Leirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="79" phylum="Arthropoda" rank="species" species="quinquestriatus">
<emphasis id="B951EABB2002191EFDE48BC6DF02F884" box="[563,578,1772,1795]" italics="true" pageId="80" pageNumber="79">L</emphasis>
.
<emphasis id="B951EABB2002191EFD8F8BC6DFBEF884" box="[600,766,1772,1795]" italics="true" pageId="80" pageNumber="79">quinquestriatus</emphasis>
</taxonomicName>
in having slender pedipalp and metasomal segments, and reduced granulation between the posterior median carinae of the carapace. However, they grouped with the northern populations from
<collectingCountry id="F33276392002191EFE238A4CDF6FF8F9" box="[500,559,1894,1918]" name="Israel" pageId="80" pageNumber="79">Israel</collectingCountry>
and
<collectingCountry id="F33276392002191EFDA58A4CDFF9F8F9" box="[626,697,1894,1918]" name="Jordan" pageId="80" pageNumber="79">Jordan</collectingCountry>
in other morphometrics (i.e. lower metasoma I W/ II W, lower pedipalp movable finger L/ manus ventral L, higher vesicle L/ telson L) and meristics (number of denticles on ventromedian carinae of metasoma II–III).
<bibRefCitation id="EFB44B582002191EFAE488F2DEC6F989" author="Levy" etAl="et al" firstAuthor="Levy" pageId="80" pageNumber="79" refId="ref62185" refString="LEVY, G. &amp; P. AMITAI. 1980. Scorpiones. Fauna Palaestina. Arachnida I. The Israel Academy of Sciences and Humanities. Jerusalem, 130 pp .." type="book" year="1980">Levy et al (1980)</bibRefCitation>
also noted that the southern populations of
<emphasis id="B951EABB2002191EFABA88DDD83CF989" box="[1389,1404,1527,1550]" italics="true" pageId="80" pageNumber="79">L</emphasis>
.
<emphasis id="B951EABB2002191EFA5B88DDD8D9F989" box="[1420,1433,1527,1550]" italics="true" pageId="80" pageNumber="79">q</emphasis>
.
<taxonomicName id="4C254D2A2002191EFCE88B3FDEE1F9AA" baseAuthorityName="Birula" baseAuthorityYear="1908" box="[831,929,1557,1581]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="80" pageNumber="79" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB2002191EFCE88B3FDEE1F9AA" box="[831,929,1557,1581]" italics="true" pageId="80" pageNumber="79">hebraeus</emphasis>
</taxonomicName>
had more slender appendages. Partial overlap of characters suggests that there could be two subspecies,
<emphasis id="B951EABB2002191EFC4D8B79DEE9F9ED" box="[922,937,1619,1642]" italics="true" pageId="80" pageNumber="79">L</emphasis>
.
<emphasis id="B951EABB2002191EFC6F8B79DE85F9ED" box="[952,965,1619,1642]" italics="true" pageId="80" pageNumber="79">q</emphasis>
.
<taxonomicName id="4C254D2A2002191EFC028B79D93BF9ED" box="[981,1147,1619,1642]" class="Insecta" family="Carabidae" genus="Leirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="79" phylum="Arthropoda" rank="species" species="quinquestriatus">
<emphasis id="B951EABB2002191EFC028B79D93BF9ED" box="[981,1147,1619,1642]" italics="true" pageId="80" pageNumber="79">quinquestriatus</emphasis>
</taxonomicName>
and
<emphasis id="B951EABB2002191EFB658B79D981F9ED" box="[1202,1217,1619,1642]" italics="true" pageId="80" pageNumber="79">L</emphasis>
.
<emphasis id="B951EABB2002191EFB078B79D99DF9ED" box="[1232,1245,1619,1642]" italics="true" pageId="80" pageNumber="79">q</emphasis>
.
<taxonomicName id="4C254D2A2002191EFB3B8B78D80EF9ED" baseAuthorityName="Birula" baseAuthorityYear="1908" box="[1260,1358,1618,1642]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="80" pageNumber="79" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB2002191EFB3B8B78D80EF9ED" box="[1260,1358,1618,1642]" italics="true" pageId="80" pageNumber="79">hebraeus</emphasis>
</taxonomicName>
, with a hybrid zone in southern Sinai. However, there appears to be a physical barrier preventing gene flow between these populations because
<taxonomicName id="4C254D2A2002191EFBF78B85D932F941" authorityName=": Vachon" authorityYear="1949" box="[1056,1138,1711,1734]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="80" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABB2002191EFBF78B85D932F941" box="[1056,1138,1711,1734]" italics="true" pageId="80" pageNumber="79">Leiurus</emphasis>
</taxonomicName>
is excluded from the sandy coastal plain connecting North Africa and the Sinai Peninsula (
<bibRefCitation id="EFB44B582002191EFC6F8BC1D9F3F884" author="Levy &amp; Amitai" bookContentInfo="130 pp" box="[952,1203,1771,1795]" firstAuthor="Levy" journalOrPublisher="The Israel Academy of Sciences and Humanities. Jerusalem" pageId="80" pageNumber="79" refId="ref62185" refString="LEVY, G. &amp; P. AMITAI. 1980. Scorpiones. Fauna Palaestina. Arachnida I. The Israel Academy of Sciences and Humanities. Jerusalem, 130 pp .." title="Scorpiones. Fauna Palaestina. Arachnida I" type="book" year="1980">Levy &amp; Amitai, 1980</bibRefCitation>
). Genetic isolation is supported by divergent evolution of distinct polypeptide toxins in their venoms (
<bibRefCitation id="EFB44B582002191EFB8F8A02D823F8C7" author="Smertenko" box="[1112,1379,1832,1856]" etAl="et al." firstAuthor="Smertenko" pageId="80" pageNumber="79" pagination="229 - 244" refId="ref64154" refString="SMERTENKO, A., M. A. A. OMRAN, P. J. HUSSEY &amp; A. McVEAN. 2001. Toxin evolution in scorpion venom: evidence for toxin divergence under strong negative selection in Leiurus quinquestriatus subspecies. Journal of Toxicology. - Toxin Reviews, 20 (3 &amp; 4): 229 - 244." type="journal article" year="2001">Smertenko et al., 2001</bibRefCitation>
) and different physicochemical profiles of venom proteins (
<bibRefCitation id="EFB44B582002191EFC918A4CD908F8F9" author="Nascimento" box="[838,1096,1894,1918]" etAl="et al." firstAuthor="Nascimento" pageId="80" pageNumber="79" pagination="628 - 639" refId="ref62846" refString="NASCIMENTO, D. G., B. RATES, D. M. SANTOS, T. VERANO-BRAGA, A. BARBOSA-SILVA, A. A. DUTRA, I. BIONDI, M. F. MARTIN-EAU- CLAIRE, M. E. DE LIMA &amp; A. M. C. PIMENTA. 2006. Moving pieces in a taxonomic puzzle: Venom 2 D-LC / MS and data clustering analyses to infer phylogenetic relationships in some scorpions from the Buthidae family (Scorpiones). Toxicon, 47: 628 - 639." type="journal article" year="2006">Nascimento et al., 2006</bibRefCitation>
). Here, we propose to diagnose the African and Middle Eastern populations as distinct species by a combination of morphological characters. The southern Sinai populations occur in a different habitat (i.e. rocky mountains), and may belong to other undescribed species.
</paragraph>
</subSubSection>
</treatment>
</document>
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