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<document id="D2D77B60AC5F8A7377BBA9B32A518807" ID-CLB-Dataset="152185" ID-DOI="10.18590/euscorpius.2014.vol2014.iss191.1" ID-GBIF-Dataset="a6381f1d-22e1-4629-b369-07f5ab1d7834" ID-ISSN="1536-9307" ID-Zenodo-Dep="6505622" ID-ZooBank="E467B3C0-D693-4EAF-B5F0-759D8C63FE35" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638245227176" checkinUser="felipe" docAuthor="Lowe, Graeme, Yağmur, Ersen Aydın &amp; Kovařík, František" docDate="2014" docId="038C87BF20031912FF0E8BCADC10F9EA" docLanguage="en" docName="Euscorpius.191.1-129.pdf" docOrigin="Euscorpius 191" docSource="https://doi.org/10.18590/euscorpius.2014.vol2014.iss191.1" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Leiurus quinquestriatus" docType="treatment" docVersion="12" lastPageNumber="88" masterDocId="FFB5FFC72052194EFFD78D2ADD40FF87" masterDocTitle="A review of the genus Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) with description of four new species from the Arabian Peninsula" masterLastPageNumber="129" masterPageNumber="1" pageNumber="80" updateTime="1699471136939" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="5D90C087FEE1A528D8E12B8D84665DC4">A review of the genus Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) with description of four new species from the Arabian Peninsula</mods:title>
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<taxonomicName id="4C254D2A2003191FFF0E8BCADE48F97A" ID-CoL="6P78R" authority="(Ehrenberg, 1828)" baseAuthorityName="Ehrenberg" baseAuthorityYear="1828" box="[217,776,1760,1789]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="81" pageNumber="80" phylum="Arthropoda" rank="species" species="quinquestriatus">
<emphasis id="B951EABB2003191FFF0E8BCADF56F97A" bold="true" box="[217,534,1760,1789]" italics="true" pageId="81" pageNumber="80">Leiurus quinquestriatus</emphasis>
(Ehrenberg, 1828)
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<figureCitation id="131E2A2C2003191FFF078A2DDC11F898" box="[208,337,1799,1823]" captionStart-0="Figure 72" captionStart-1="Figure 73" captionStart-10="Figure 82" captionStart-2="Figure 74" captionStart-3="Figure 75" captionStart-4="Figure 76" captionStart-5="Figure 77" captionStart-6="Figure 78" captionStart-7="Figure 79" captionStart-8="Figure 80" captionStart-9="Figure 81" captionStartId-0="90.[192,267,1856,1880]" captionStartId-1="91.[192,267,1874,1898]" captionStartId-10="100.[192,267,1070,1094]" captionStartId-2="92.[192,267,1305,1329]" captionStartId-3="93.[192,267,1606,1630]" captionStartId-4="94.[192,267,1880,1904]" captionStartId-5="95.[192,267,1867,1891]" captionStartId-6="96.[192,267,1229,1253]" captionStartId-7="97.[192,267,1623,1647]" captionStartId-8="98.[192,267,1856,1880]" captionStartId-9="99.[192,267,1185,1209]" captionTargetBox-0="[234,1369,198,1825]" captionTargetBox-1="[260,1357,215,1841]" captionTargetBox-10="[207,1412,188,1044]" captionTargetBox-2="[204,1425,196,1266]" captionTargetBox-3="[191,1441,191,1574]" captionTargetBox-4="[233,1393,212,1849]" captionTargetBox-5="[194,1441,217,1801]" captionTargetBox-6="[204,1427,215,1211]" captionTargetBox-7="[191,1440,160,1587]" captionTargetBox-8="[251,1361,191,1824]" captionTargetBox-9="[191,1439,191,1155]" captionTargetId-0="figure-24@90.[235,1368,199,1824]" captionTargetId-1="figure-22@91.[262,1356,216,1840]" captionTargetId-10="figure-431@100.[192,1439,168,1065]" captionTargetId-2="figure-245@92.[192,1437,169,1278]" captionTargetId-3="figure-148@93.[192,1440,192,1573]" captionTargetId-4="figure-24@94.[234,1392,214,1848]" captionTargetId-5="figure-22@95.[195,1440,219,1800]" captionTargetId-6="figure-350@96.[192,1439,203,1223]" captionTargetId-7="figure-131@97.[192,1439,161,1586]" captionTargetId-8="figure-39@98.[252,1360,192,1824]" captionTargetId-9="figure-417@99.[192,1438,192,1154]" captionTargetPageId-0="90" captionTargetPageId-1="91" captionTargetPageId-10="100" captionTargetPageId-2="92" captionTargetPageId-3="93" captionTargetPageId-4="94" captionTargetPageId-5="95" captionTargetPageId-6="96" captionTargetPageId-7="97" captionTargetPageId-8="98" captionTargetPageId-9="99" captionText-0="Figure 72: Leiurus quinquestriatus (Ehrenberg, 1828), male. Habitus, dorsal aspect. Kena, Egypt (NHMB 17g)." captionText-1="Figure 73: Leiurus quinquestriatus (Ehrenberg, 1828), male. Habitus, ventral aspect. Kena, Egypt (NHMB 17g)." captionText-10="Figure 82: Leiurus quinquestriatus (Ehrenberg, 1828), 5 of 8 syntypes (ZMHB No. 140). Females: A, #12C01-3; B, #12C04-6. Males: C, #12C07-9; D, #12C10-3. Juvenile: E, #12C14-6. Egypt; Sudan. Scale bars (10 mm) apply to the habitus photos. Body lengths: A, 90 mm; B, 88 mm; C, 63 mm; D, 69 mm; E, 65 mm. Not shown are three additional fragmented juvenile syntypes." captionText-2="Figure 74: Leiurus quinquestriatus (Ehrenberg, 1828), male. A. Carapace and tergites. B. Coxosternal area and sternites. Kena, Egypt (NHMB 17g)." captionText-3="Figure 75: Leiurus quinquestriatus (Ehrenberg, 1828), male. Right pedipalp. A. Femur, dorsal aspect. B. Patella, dorsal aspect. C. Patella, external aspect. D. Chela, ventral aspect. E. Chela, dorsal aspect. F. Chela, external aspect. Kena, Egypt (NHMB 17g)." captionText-4="Figure 76: Leiurus quinquestriatus (Ehrenberg, 1828), female. Habitus, dorsal aspect. Kena, Egypt (NHMB 17g)." captionText-5="Figure 77: Leiurus quinquestriatus (Ehrenberg, 1828), female. Habitus, ventral aspect. Kena, Egypt (NHMB 17g)." captionText-6="Figure 78: Leiurus quinquestriatus (Ehrenberg, 1828), female. A. Carapace and tergites. B. Coxosternal area and sternites. Kena, Egypt (NHMB 17g)." captionText-7="Figure 79: Leiurus quinquestriatus (Ehrenberg, 1828), female. Right pedipalp. A. Femur, dorsal aspect. B. Patella, dorsal aspect. C. Patella, external aspect. D. Chela, ventral aspect. E. Chela, dorsal aspect. F. Fixed finger dentition. G. Movable finger dentition. H. Chela, external aspect. Kena, Egypt (NHMB 17g)." captionText-8="Figure 80: Leiurus quinquestriatus (Ehrenberg, 1828). Metasoma. A. Male, lateral aspect. B. Male, ventral aspect. C. Female, lateral aspect. D. Female, ventral aspect. Kena, Egypt (NHMB 17g)." captionText-9="Figure 81: Leiurus quinquestriatus (Ehrenberg, 1828), female. Tarsi. A-D. Left telotarsus and distal basitarsus, ventral aspect. A. Leg I. B. Leg II. C. Leg III. D. Leg IV. E. Right basitarsus III retrolateral aspect. Kharge Oasis, Egypt (NHMB 17i)." figureDoi-0="http://doi.org/10.5281/zenodo.6505773" figureDoi-1="http://doi.org/10.5281/zenodo.6505776" figureDoi-10="http://doi.org/10.5281/zenodo.6505796" figureDoi-2="http://doi.org/10.5281/zenodo.6505778" figureDoi-3="http://doi.org/10.5281/zenodo.6505780" figureDoi-4="http://doi.org/10.5281/zenodo.6505782" figureDoi-5="http://doi.org/10.5281/zenodo.6505784" figureDoi-6="http://doi.org/10.5281/zenodo.6505786" figureDoi-7="http://doi.org/10.5281/zenodo.6505790" figureDoi-8="http://doi.org/10.5281/zenodo.6505792" figureDoi-9="http://doi.org/10.5281/zenodo.6505794" httpUri-0="https://zenodo.org/record/6505773/files/figure.png" httpUri-1="https://zenodo.org/record/6505776/files/figure.png" httpUri-10="https://zenodo.org/record/6505796/files/figure.png" httpUri-2="https://zenodo.org/record/6505778/files/figure.png" httpUri-3="https://zenodo.org/record/6505780/files/figure.png" httpUri-4="https://zenodo.org/record/6505782/files/figure.png" httpUri-5="https://zenodo.org/record/6505784/files/figure.png" httpUri-6="https://zenodo.org/record/6505786/files/figure.png" httpUri-7="https://zenodo.org/record/6505790/files/figure.png" httpUri-8="https://zenodo.org/record/6505792/files/figure.png" httpUri-9="https://zenodo.org/record/6505794/files/figure.png" pageId="81" pageNumber="80">Figs. 7282</figureCitation>
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<figureCitation id="131E2A2C2003191FFE8B8A2DDCCCF898" box="[348,396,1799,1823]" captionStart="Figure 87" captionStartId="107.[192,267,1798,1822]" captionTargetBox="[343,1333,196,1766]" captionTargetId="figure-90@107.[344,1333,197,1765]" captionTargetPageId="107" captionText="Figure 87: Metasoma V and telson of Leiurus spp. Males, lateral aspect. A. L. abdullahbayrami Yağmur, Koç et Kunt, 2009. B. L. arabicus sp. n.. C. L. brachycentrus (Ehrenberg, 1829) stat. n.. D. L. haenggii sp. n. E. L. heberti sp. n. F. L. hebraeus (Birula, 1908) stat. n.. G. L. macroctenus sp. n.. H. L. quinquestriatus (Ehrenberg, 1828). Locality data as in Figs. 9, 20, 31, 43, 55, 67, 79. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6505806" httpUri="https://zenodo.org/record/6505806/files/figure.png" pageId="81" pageNumber="80">87H</figureCitation>
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<figureCitation id="131E2A2C2003191FFE408A2DDC87F898" box="[407,455,1799,1823]" captionStart="Figure 88" captionStartId="108.[192,267,1816,1840]" captionTargetBox="[318,1309,215,1785]" captionTargetId="figure-95@108.[319,1308,216,1784]" captionTargetPageId="108" captionText="Figure 88: Metasoma V and telson of Leiurus spp. Males, ventral aspect. A. L. abdullahbayrami Yağmur, Koç et Kunt, 2009. B. L. arabicus sp. n.. C. L. brachycentrus (Ehrenberg, 1829) stat. n.. D. L. haenggii sp. n.. E. L. heberti sp. n. F. L. hebraeus (Birula, 1908) stat. n.. G. L. macroctenus sp. n.. H. L. quinquestriatus (Ehrenberg, 1828). Locality data as in Figs. 9, 20, 31, 43, 55, 67, 79. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6505808" httpUri="https://zenodo.org/record/6505808/files/figure.png" pageId="81" pageNumber="80">88H</figureCitation>
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<figureCitation id="131E2A2C2003191FFE058A2DDF43F898" box="[466,515,1799,1823]" captionStart="Figure 89" captionStartId="109.[192,267,1809,1833]" captionTargetBox="[345,1336,208,1778]" captionTargetId="figure-94@109.[346,1335,209,1777]" captionTargetPageId="109" captionText="Figure 89: Metasoma V and telson of Leiurus spp. Females, lateral aspect. A. L. abdullahbayrami Yağmur, Koç et Kunt, 2009. B. L. arabicus sp. n. C. L. brachycentrus (Ehrenberg, 1829) stat. n. D. L. haenggii sp. n. E. L. heberti sp. n. F. L. hebraeus (Birula, 1908) stat. n. G. L. macroctenus sp. n. H. L. quinquestriatus (Ehrenberg, 1828). Locality data as in Figs. 9, 20, 31, 43, 55, 67, 79. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6505810" httpUri="https://zenodo.org/record/6505810/files/figure.png" pageId="81" pageNumber="80">89H</figureCitation>
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<figureCitation id="131E2A2C2003191FFDDA8A2DDF7EF898" box="[525,574,1799,1823]" captionStart="Figure 90" captionStartId="110.[192,267,1808,1832]" captionTargetBox="[318,1309,207,1777]" captionTargetId="figure-95@110.[319,1308,208,1776]" captionTargetPageId="110" captionText="Figure 90: Metasoma V and telson of Leiurus spp. Females, ventral aspect. A. L. abdullahbayrami Yağmur, Koç et Kunt, 2009. B. L. arabicus sp. n. C. L. brachycentrus (Ehrenberg, 1829) stat. n.. D. L. haenggii sp. n. E. L. heberti sp. n. F. L. hebraeus (Birula, 1908) stat. n. G. L. macroctenus sp. n. H. L. quinquestriatus (Ehrenberg, 1828). Locality data as in Figs. 9, 20, 31, 43, 55, 67, 79. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6505812" httpUri="https://zenodo.org/record/6505812/files/figure.png" pageId="81" pageNumber="80">90H</figureCitation>
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<figureCitation id="131E2A2C2003191FFD9E8A2DDF39F898" box="[585,633,1799,1823]" captionStart="Figure 91" captionStartId="111.[192,267,1766,1790]" captionTargetBox="[303,1321,191,1734]" captionTargetId="figure-124@111.[304,1320,192,1733]" captionTargetPageId="111" captionText="Figure 91: Pigmentation patterns of metasoma IV, V of Leiurus species, ventral aspect. A. L. haenggii sp. n., holotype female, Taif, Saudi Arabia (NHMB 17k). B. L. macroctenus sp. n., holotype male, Thumrait, Oman. C. L. heberti sp. n., holotype male, Wadi Andur, Oman. D. L. quinquestriatus (Ehrenberg, 1828), female, Cairo, Egypt (NHMB 17d). E. L. brachycentrus (Ehrenberg, 1829) stat. n., male, Ad Darb, Saudi Arabia (NHMB 17ag). F. L. arabicus sp. n., holotype female, Kushm Dibi, Saudi Arabia (NHMB 17aq). G. L. hebraeus (Birula, 1908) stat. n., female, Israel (NHMB 17a). H. L. abdullahbayrami Yağmur, Koç et Kunt, 2009, male, Eski Sarkaya Village, Turkey. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6505814" httpUri="https://zenodo.org/record/6505814/files/figure.png" pageId="81" pageNumber="80">91D</figureCitation>
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<figureCitation id="131E2A2C2003191FFD538A2DDFECF898" box="[644,684,1799,1823]" captionStart="Figure 92" captionStartId="112.[192,267,1486,1510]" captionTargetBox="[224,1390,157,1461]" captionTargetId="figure-274@112.[212,1402,145,1473]" captionTargetPageId="112" captionText="Figure 92: Sternite III of Leiurus spp., female. A. L. abdullahbayrami Yağmur, Koç et Kunt, 2009, Caybasi Village, Turkey. B. L. arabicus sp. n., holotype, Kushm Dibi, Saudi Arabia (NHMB 17aq). C. L. arabicus sp. n., paratype, Al Amar, Bahrain (NHMB 17bg). D. L. heberti sp. n., paratype, Jabal Samhan. Oman. E. L. haenggii sp. n., paratype, Wadi Turabah, Saudi Arabia (NHMB 17al). F. L. haenggii sp. n., paratype, Adnan, Saudi Arabia (NHMB 17am). G. L. brachycentrus (Ehrenberg, 1829) stat. n., Al Mansuriah, Yemen. H. L. macroctenus sp. n., paratype, Masirah Island, Oman. I. L. macroctenus sp. n., paratype, Thumrait, Oman. J. L. quinquestriatus (Ehrenberg, 1828), Kharga Oasis, Egypt (NHMB 17i). K. L. hebraeus (Birula, 1908) stat. n., Israel (NHMB 17a). L. L. jordanensis Lourenço, Modry et Amr, 2002, al-Tawil, Saudi Arabia." figureDoi="http://doi.org/10.5281/zenodo.6505816" httpUri="https://zenodo.org/record/6505816/files/figure.png" pageId="81" pageNumber="80">92J</figureCitation>
,
<figureCitation id="131E2A2C2003191FFD618A2DDFA7F898" box="[694,743,1799,1823]" captionStart="Figure 93" captionStartId="113.[192,267,1572,1596]" captionTargetBox="[224,1390,195,1538]" captionTargetId="figure-226@113.[212,1402,183,1550]" captionTargetPageId="113" captionText="Figure 93: Medial intercarinal areas of tergite III of female Leiurus spp. A. L. arabicus sp. n., paratype, Jeddah, Saudi Arabia. B. L. arabicus sp. n., paratype, Wadi Khumrah, Saudi Arabia (NHMB 17bj). C. L. haenggii sp. n., paratype, Yemen. D. L. haenggii sp. n., paratype, Wadi Maraba, Saudi Arabia (NHMB 17al). E. L. heberti sp. n., paratype, Jabal Samhan. Oman. F. L. abdullahbayrami Yağmur, Koç et Kunt, 2009, Caybasi Village, Turkey. G. L. brachycentrus (Ehrenberg, 1829) stat. n., Al Mansuriah, Yemen. H. L. macroctenus sp. n., paratype, Masirah Island, Oman. I. L. hebraeus (Birula, 1908) stat. n., Israel (NHMB 17a). J. L. hebraeus Birula, 1908, Kurayyima, Jordan. K. L. quinquestriatus (Ehrenberg, 1828), Kharga Oasis, Egypt (NHMB 17i). L. L. jordanensis Lourenço, Modry et Amr, 2002, al-Tawil, Saudi Arabia." figureDoi="http://doi.org/10.5281/zenodo.6505818" httpUri="https://zenodo.org/record/6505818/files/figure.png" pageId="81" pageNumber="80">93K</figureCitation>
,
<figureCitation id="131E2A2C2003191FFD258A2DDC32F8C6" captionStart-0="Figure 94" captionStart-1="Figure 95" captionStartId-0="114.[192,260,765,786]" captionStartId-1="118.[192,260,1217,1238]" captionTargetBox-0="[204,1427,204,723]" captionTargetBox-1="[214,1409,159,1149]" captionTargetId-0="figure-776@114.[192,1439,192,745]" captionTargetId-1="figure-476@118.[192,1437,128,1181]" captionTargetPageId-0="114" captionTargetPageId-1="118" captionText-0="Figure 94: Comparative analysis of granulation on tergite III of female Leiurus. A. UV fluorescence image of tergite III medial intercarinal surface of L. quinquestriatus (Egypt). B. Mask of granulation pattern on medial intercarinal surface from image in Figure 94A. C. Enlarged view of tergite III granulation in rectangular area indicated in Figure 94A. Granules were identified as local maxima of fluorescence intensity with contour shadows cast by directional UV illumination, and were modeled by elliptical regions-of-interest (ROIs). Parameters of granule ROIs were measured in ImageJ 1.44 (Rasband, 19972011). Area of granulometric analysis was restricted to surfaces bounded by the lateral and posterior marginal carinae, and a line passing through oblique, anterior transverse rows of enlarged granules (granules along carinae and bounding lines were omitted). D. Scatter plot of mean granule diameter vs. total granule area for 4 species of Leiurus (L. arabicus sp. n., L. haenggii sp. n., L. hebraeus stat. n. and L. quinquestriatus). Each point represents granulometric data derived from tergite III of one scorpion specimen (bilaterally, as shown in Figure 94B). Data were extracted from 18,423 ROIs from 36 scorpions. For each specimen, total granule area (a measure of density of granulation) was computed as sum of areas of all ROIs, and mean ROI diameter (a measure of coarseness of granulation) as the mean value of the maximum diameters of all granule ROIs. For comparative analysis, images from different size scorpions were resampled to equalize the distance between left and right posterior marginal granules of the lateral carinae (arbitrarily set to 4,000 units or pixels; linear dimensions expressed as [pixel], areas as [pixel2]). The total granule area separated the species into 2 groups: i.e. sparsely granulated (L. arabicus, L. haenggii) and densely granulated (L. hebraeus, L. quinquestriatus). In contrast, the distributions of ROI diameter were broadly overlapping. E. To obtain a more sensitive comparison of the coarseness of granulation, normalized cumulative distributions of single ROI areas for 3 species of Leiurus were computed (inset indicates number of scorpions analyzed). Relative horizontal positions of these curves indicated increasing coarseness of granulation, in rank order: L. haenggii &lt;L. quinquestriatus &lt;L. hebraeus (Ngranules = 2103, 5202, 2190 respectively). This ranking was confirmed by a Kolmogorov-Smirnov test which detected significant differences between the distributions of log (granule area) (P &lt;0.001)." captionText-1="Figure 95: Variation in key diagnostic biometrics of adult Leiurus spp. A. Cumulative distributions of position of trichobothrium db relative to est on pedipalp fixed finger for: L. abdullahbayrami (53), L. brachycentrus stat. n. (11), L. hebraeus stat. n. (38), L. macroctenus sp. n. (176), L. quinquestriatus (61), L. haenggii sp. n. (158), L. arabicus sp. n. (87) (number of fingers measured in parentheses). Data for males and females were pooled. The dbest distances were normalized to pedipalp movable finger length (chord length from finger tip to external articular condyle), with positive values indicating db distal to est, negative values indicating db proximal to est. Distributions of L. haenggii (µ = 0.06367) and L. arabicus sp. n. (µ = 0.06671) were not significantly different (P = 0.405148, t test), indicating a close relationship between these species. Distribution of L. quinquestriatus (µ = 0.047313) was significantly different from those of L. haenggii and L. arabicus (P = 0.000102 and 0.000030 respectively). B. Scatter plot of morphometric ratios of mid-pectine sensillar margin L (MPSM, indicated in inset) to metasoma I W, and to carapace L, showing differences in relative pectinal tooth size in females of nine species of Leiurus. C. A subset of the female data in Figure 95B, plotted as cumulative distributions of morphometric ratio of mid-pectine sensillar margin L to metasoma I W. Distributions of L. haenggii (µ = 0.102268) and L. arabicus (µ = 0.10432) were not significantly different (P&gt; 0.1, K-S test). D. Scatter plot of morphometric ratios of mid-pectine sensillar margin L to metasoma I W (relative size of pectine teeth), and metasoma II L/W (slenderness of metasoma II) for females of nine species of Leiurus. A, C, D: symbol key as in Figure 95B." figureDoi-0="http://doi.org/10.5281/zenodo.6505820" figureDoi-1="http://doi.org/10.5281/zenodo.6505822" httpUri-0="https://zenodo.org/record/6505820/files/figure.png" httpUri-1="https://zenodo.org/record/6505822/files/figure.png" pageId="81" pageNumber="80">94 95</figureCitation>
,
<figureCitation id="131E2A2C2003191FFEAA8A03DCECF8C6" box="[381,428,1833,1857]" captionStart="Figure 96" captionStartId="119.[192,267,1530,1554]" captionTargetBox="[201,1430,216,1489]" captionTargetId="figure-220@119.[192,1440,206,1499]" captionTargetPageId="119" captionText="Figure 96: Carapace and tergites IIII of representatives of six genera of the Buthus group. A. Compsobuthus acutecarinatus (Simon, 1882), male, Jabal Qara, Oman. B. Cicileus exilis (Pallary, 1928), male, Hoggar, Algeria. C. Leiurus quinquestriatus (Ehrenberg, 1828), female, Egypt. D. Sassanidotus gracilis (Birula, 1900), female, Hormozgan Prov., Iran. E. Odontobuthus bidentatus Lourenço &amp; Pézier, 2002, male, Omidiyeh, Iran. F. Apistobuthus susanae Lourenço 1998, male, Albadji, Iran. Scale bars: A, B, D: 1 mm; C, E, F: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6505824" httpUri="https://zenodo.org/record/6505824/files/figure.png" pageId="81" pageNumber="80">96C</figureCitation>
,
<figureCitation id="131E2A2C2003191FFE608A03DCBCF8C6" box="[439,508,1833,1857]" captionStart-0="Figure 98" captionStart-1="Figure 99" captionStartId-0="121.[192,267,1251,1275]" captionStartId-1="122.[192,267,1014,1038]" captionTargetBox-0="[215,1377,224,1220]" captionTargetBox-1="[191,1438,222,978]" captionTargetId-0="figure-474@121.[192,1437,206,1246]" captionTargetId-1="figure-479@122.[192,1437,223,978]" captionTargetPageId-0="121" captionTargetPageId-1="122" captionText-0="Figure 98: Selected morphometrics of Leiurus, Cicileiurus, Cicileus and Compsobuthus compared to other Buthus group scorpions. AB. Scatter plots of the fraction of fixed finger length distal to db (A) and est (B) vs. the ratio of movable finger length to carapace length. Each point represents one sex of one species. Larger ordinate values correspond to more basal positions of the trichobothria, and larger abscissa values to longer pedipalp fingers. There was a significant inverse correlation between relative length of the portion of the fixed finger distal to db and est (R = -0.6534, -0.5488, respectively), and the relative length of the movable finger (the latter being a measure of elongation of both fixed and movable fingers). Highlighted symbols show that Leiurus (light magenta circles), Cicileiurus (red triangle), Cicileus (green squares) and Compsobuthus (yellow circles) are located in the lower right halves of the plots, i.e. all have relatively elongated fingers and more distal placement of both db and est. Gray circles are data from other Buthus group species. C. Scatter plot of the fraction of fixed finger length distal to db vs. the fraction distal to est. The strong positive correlation (R = 0.8052) indicates a tendency for db and est to move together towards more distal locations as the fixed finger becomes more elongated. Species above the diagonal (solid blue) have db proximal to est, and those below have db distal to est. Solid gray lines in AC are fits by least squares regression through all points. D. Scatter plot of pedipalp femur L/W (a measure of pedipalp elongation) vs. carapace L (a measure of body size). These two variables were not significantly correlated (R = 0.094). Data were compiled from the literature and specimens in the authors collections for 38 genera and 203 species representing the majority of taxa in the Buthus group, including both males (N = 124) and females (N = 97). Genera (and number of species) included: Afghanobuthus (1), Androctonus (11), Apistobuthus (2), Baloorthochirus (1), Birulatus (2), Buthacus (13), Butheoloides (11), Butheolus (5), Buthiscus (1), Buthus (26), Cicileiurus (1), Cicileus (2), Compsobuthus (33), Congobuthus (1), Darchenia (1), Gint (1), Hemibuthus (1), Hottentotta (31), Leiurus (10), Liobuthus (1), Lissothus (3), Mesobuthus (7), Neobuthus (2), Odontobuthus (6), Orthochirus (12), Pantobuthus (1), Pectinibuthus (1), Plesiobuthus (1), Polisius (1), Razianus (3), Saharobuthus (1), Somalibuthus (1), Vachoniolus (4), Vachonus (1)." captionText-1="Figure 99: Geographic plot of locality data of material examined for nine species of Leiurus distributed over northeast Africa, the Levant and Arabian Peninsula. Map colored by elevation with shaded relief. Additional locality data for: L. abdullahbayrami from Yağmur et al., 2009; for L. brachycentrus Ehrenberg, 1829 stat. n., from Simon, 1882 (Buthus beccarii)." figureDoi-0="http://doi.org/10.5281/zenodo.6505828" figureDoi-1="http://doi.org/10.5281/zenodo.6505832" httpUri-0="https://zenodo.org/record/6505828/files/figure.png" httpUri-1="https://zenodo.org/record/6505832/files/figure.png" pageId="81" pageNumber="80">9899</figureCitation>
,
<tableCitation id="C6A703122003191FFDD08A03DF2BF8C6" box="[519,619,1833,1857]" captionStart="Table 3" captionStartId="105.[192,257,1349,1373]" captionTargetBox="[209,1890,232,1297]" captionTargetId="graphics-7@105.[194,1919,192,1298]" captionText="Table 3A: Summary of key characters separating four Leiurus species examined in this study (adult females unless otherwise specified). Numeric values indicate ranges with sample sizes in parentheses." httpUri="http://table.plazi.org/id/DF5A6621203B1927FF17886FDC8EFAF2" pageId="81" pageNumber="80" tableUuid="DF5A6621203B1927FF17886FDC8EFAF2">Tabs. 3A</tableCitation>
, 5)
</paragraph>
</subSubSection>
<subSubSection id="C33F65222003191BFCE88B35D806F8B2" lastPageId="85" lastPageNumber="84" pageId="81" pageNumber="80" type="reference_group">
<paragraph id="8B9A36A92003191FFCE88B35D841F9D5" blockId="81.[831,1440,1567,1618]" pageId="81" pageNumber="80">
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<treatmentCitation id="0A8410B82003191FFCE88B35D93FF9D5" author="HEMPRICH, F. W. &amp; C. G. EHRENBERG" pageId="81" pageNumber="80" year="1828">
<taxonomicName id="4C254D2A2003191FFCE88B35D839F9B0" authority="Ehrenberg" authorityName="Ehrenberg" authorityYear="1828" box="[831,1401,1567,1591]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="81" pageNumber="80" phylum="Arthropoda" rank="species" species="quinquestriatus" subGenus="Leiurus">
<emphasis id="B951EABB2003191FFCE88B35DE87F9B0" box="[831,967,1567,1591]" italics="true" pageId="81" pageNumber="80">Androctonus</emphasis>
(
<emphasis id="B951EABB2003191FFC348B0AD974F9B0" box="[995,1076,1568,1591]" italics="true" pageId="81" pageNumber="80">Leiurus</emphasis>
)
<emphasis id="B951EABB2003191FFB868B0AD9B7F9B0" box="[1105,1271,1568,1591]" italics="true" pageId="81" pageNumber="80">quinquestriatus</emphasis>
Ehrenberg
</taxonomicName>
in
<bibRefCitation id="EFB44B582003191FFCE88B10D93FF9D5" author="Hemprich &amp; Ehrenberg" box="[831,1151,1594,1618]" firstAuthor="Hemprich" journalOrPublisher="Decas Prima. Berolini ex Officina Academica, Venditur a Mittlero: plate I Buthus; plate II: Androctonus" pageId="81" pageNumber="80" refId="ref60936" refString="HEMPRICH, F. W. &amp; C. G. EHRENBERG. 1828. Zoologica II. Arachnoidea. In: Symbolae Physicae seu Icones et Descriptiones Animalium Evertebratorum Sepositis Insectis quae ex Itinere per Africam Borealem et Asiam Occidentalem. Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg, Medicinae et Chirurgiae Doctorum, Studio Novae aut Illustratae Redierunt. Percensuit et Regis Iussu et Impensis Edidit Dr. C. G. Ehrenberg. Decas Prima. Berolini ex Officina Academica, Venditur a Mittlero: plate I Buthus; plate II: Androctonus." title="Zoologica II. Arachnoidea" type="book chapter" volumeTitle="Symbolae Physicae seu Icones et Descriptiones Animalium Evertebratorum Sepositis Insectis quae ex Itinere per Africam Borealem et Asiam Occidentalem. Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg, Medicinae et Chirurgiae Doctorum, Studio Novae aut Illustratae Redierunt. Percensuit et Regis Iussu et Impensis Edidit Dr. C. G. Ehrenberg" year="1828">Hemprich &amp; Ehrenberg, 1828</bibRefCitation>
</treatmentCitation>
, pl. I, fig. 5.
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</paragraph>
<paragraph id="8B9A36A92003191FFCE88B46DE94F903" blockId="81.[831,1440,1644,1728]" box="[831,980,1644,1668]" pageId="81" pageNumber="80">SYNONYMS</paragraph>
<paragraph id="8B9A36A92003191FFCE88BA0D996F947" blockId="81.[831,1440,1644,1728]" pageId="81" pageNumber="80">
<treatmentCitationGroup id="AB3511872003191FFCE88BA0D996F947" pageId="81" pageNumber="80">
<treatmentCitation id="0A8410B82003191FFCE88BA0D996F947" author="HEMPRICH, F. W. &amp; C. G. EHRENBERG" pageId="81" pageNumber="80" year="1831">
<taxonomicName id="4C254D2A2003191FFCE88BA0DE2EF947" authority="Ehrenberg" authorityName=": Ehrenberg" authorityYear="1831" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="81" pageNumber="80" phylum="Arthropoda" rank="subSpecies" species="quinquestriatus" subGenus="Liurus" subSpecies="aculeatus">
<emphasis id="B951EABB2003191FFCE88BA0DE87F925" box="[831,967,1674,1698]" italics="true" pageId="81" pageNumber="80">Androctonus</emphasis>
(
<emphasis id="B951EABB2003191FFC0C8BA1D960F925" box="[987,1056,1675,1698]" italics="true" pageId="81" pageNumber="80">Liurus</emphasis>
)
<emphasis id="B951EABB2003191FFBE28BA1D80CF925" box="[1077,1356,1674,1698]" italics="true" pageId="81" pageNumber="80">quinquestriatus aculeatus</emphasis>
Ehrenberg
</taxonomicName>
in
<bibRefCitation id="EFB44B582003191FFC478B82D992F947" author="Hemprich &amp; Ehrenberg" bookContentInfo="12 pp." box="[912,1234,1704,1728]" firstAuthor="Hemprich" journalOrPublisher="Berolini ex Officina Academica, Venditur a Mittlero" pageId="81" pageNumber="80" refId="ref61080" refString="HEMPRICH, F. W. &amp; C. G. EHRENBERG. 1831. Animalia articulata, Arachnoidea. Scorpiones Africani et Asiatici. In: Symbolae Physicae. Animalia Evertebrata Exclusis Insectis Percensuit Dr. C. G. Ehrenberg. Series Prima cum Tabularum Decade Prima. Continet Animalia Africana et Asiatica 162. Berolini ex Officina Academica, Venditur a Mittlero, 12 pp. (Latin text, pages unnumbered)." title="Animalia articulata, Arachnoidea. Scorpiones Africani et Asiatici" type="book" volumeTitle="Symbolae Physicae. Animalia Evertebrata Exclusis Insectis Percensuit Dr. C. G. Ehrenberg. Series Prima cum Tabularum Decade Prima. Continet Animalia Africana et Asiatica 162" year="1831">Hemprich &amp; Ehrenberg, 1831</bibRefCitation>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A36A92003191FFCE88BF0DEA6F975" blockId="81.[831,1440,1754,1868]" box="[831,998,1754,1778]" pageId="81" pageNumber="80">REFERENCES</paragraph>
<paragraph id="8B9A36A92003191FFCE88BD2D98FF8CB" blockId="81.[831,1440,1754,1868]" pageId="81" pageNumber="80">
<treatmentCitationGroup id="AB3511872003191FFCE88BD2D98FF8CB" pageId="81" pageNumber="80">
<taxonomicName id="4C254D2A2003191FFCE88BD2D942F897" authority="Amoreux, 1789" authorityName="Amoreux" box="[831,1026,1784,1808]" class="Arachnida" family="Buthidae" genus="Scorpio" kingdom="Animalia" order="Scorpiones" pageId="81" pageNumber="80" phylum="Arthropoda" rank="species" species="occitanus">
<emphasis id="B951EABB2003191FFCE88BD2D942F897" box="[831,1026,1784,1808]" italics="true" pageId="81" pageNumber="80">Scorpio occitanus</emphasis>
</taxonomicName>
:
<treatmentCitation id="0A8410B82003191FFBC38BD2D835F897" author="AUDOUIN, V." box="[1044,1397,1784,1808]" page="173" pageId="81" pageNumber="80" year="1826">
<bibRefCitation id="EFB44B582003191FFBC38BD2D87EF897" author="AUDOUIN, V." box="[1044,1342,1784,1808]" firstAuthor="Audouin" journalOrPublisher="Histoire Naturelle, 1. C. L. P. Panekoucke, Paris" pageId="81" pageNumber="80" pagination="172 - 174" part="22" refId="ref58940" refString="AUDOUIN, V. 1826. Planche 8. Scorpions, Pinces, Solifuges. In: Explication sommaire des planches d'Arachnides de l'Egypte et de la Syrie, publiees par J. - C. Savigny. In: Description de l'Egypte, ou Recueil des Observations et des Recherches qui out ete Faites en Egypte Pendant l'Expedition de l'Armee Francaise. Histoire Naturelle, 1. C. L. P. Panekoucke, Paris, 22: 172 - 174." title="Planche 8. Scorpions, Pinces, Solifuges" type="book chapter" volumeTitle="Explication sommaire des planches d'Arachnides de l'Egypte et de la Syrie, publiees par J. - C. Savigny" year="1826">Audouin in Savigny, 1826</bibRefCitation>
: 173
</treatmentCitation>
, pl. VIII, fig. 1 (in part);
<treatmentCitation id="0A8410B82003191FFBBB8A3CDE8AF8CB" author="AUDOUIN, V." page="410" pageId="81" pageNumber="80" year="1827">
<bibRefCitation id="EFB44B582003191FFBBB8A3CD8DCF8A9" author="AUDOUIN, V." box="[1132,1436,1814,1838]" firstAuthor="Audouin" journalOrPublisher="Paris" pageId="81" pageNumber="80" pagination="409 - 412" part="22" refId="ref59024" refString="AUDOUIN, V. 1827. Planche 8. Scorpions, Pinces, Solifuges. In: Explication sommaire des planches d'Arachnides de l'Egypte et de la Syrie, publiees par J. - C. SAVIGNY. In: Description de l'Egypte, ou Recueil des Observations et des Recherches qui out ete Faites en Egypte Pendant l'Expedition de l'Armee Francaise. Histoire Naturelle, 1. C. L. P. Panekoucke, Paris, 22: 409 - 412." title="Planche 8. Scorpions, Pinces, Solifuges" type="journal article" volumeTitle="Explication sommaire des planches d'Arachnides de l'Egypte et de la Syrie, publiees par J. - C. SAVIGNY." year="1827">Audouin in Savigny, 1827</bibRefCitation>
: 410-411
</treatmentCitation>
, pl. VIII, fig. 1 (in part).
</treatmentCitationGroup>
</paragraph>
<caption id="DF5A66212000191CFF178B9AD9F2F940" ID-DOI="http://doi.org/10.5281/zenodo.6505757" ID-Zenodo-Dep="6505757" box="[192,1202,1712,1736]" httpUri="https://zenodo.org/record/6505757/files/figure.png" pageId="82" pageNumber="81" startId="82.[192,267,1712,1736]" targetBox="[191,1437,205,1681]" targetPageId="82">
<paragraph id="8B9A36A92000191CFF178B9AD9F2F940" blockId="82.[192,1202,1712,1736]" box="[192,1202,1712,1736]" pageId="82" pageNumber="81">
<emphasis id="B951EABB2000191CFF178B9ADC75F94F" bold="true" box="[192,309,1712,1736]" pageId="82" pageNumber="81">Figure 64:</emphasis>
<taxonomicName id="4C254D2A2000191CFEEB8B98DF31F940" ID-CoL="6P6WQ" authority="(Birula, 1908)" baseAuthorityName="Birula" baseAuthorityYear="1908" box="[316,625,1713,1735]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="82" pageNumber="81" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB2000191CFEEB8B98DCA3F940" box="[316,483,1713,1735]" italics="true" pageId="82" pageNumber="81">Leiurus hebraeus</emphasis>
(
<bibRefCitation id="EFB44B582000191CFE278B9BDF2AF940" author="Birula" box="[496,618,1713,1735]" firstAuthor="Birula" journalOrPublisher="Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien" pageId="82" pageNumber="81" pagination="121 - 152" part="117" refId="ref59140" refString="BIRULA, A. 1908. Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 117 2 (1): 121 - 152." title="Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae" type="journal article" year="1908">Birula, 1908</bibRefCitation>
)
</taxonomicName>
stat. n., female. Habitus, dorsal aspect. Israel (NHMB 17a).
</paragraph>
</caption>
<paragraph id="8B9A36A92000191CFF178BC1DCDFF8A5" blockId="82.[192,802,1771,1887]" pageId="82" pageNumber="81">
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<treatmentCitation id="0A8410B82000191CFF178BC1DCDFF8A5" author="HEMPRICH, F. W. &amp; C. G. EHRENBERG" page="353" pageId="82" pageNumber="81" year="1829">
<taxonomicName id="4C254D2A2000191CFF178BC1DF6CF884" authority="" box="[192,556,1771,1795]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="82" pageNumber="81" phylum="Arthropoda" rank="subSpecies" species="quinquestriatus" subGenus="Leiurus" subSpecies="quinquestriatus">
<emphasis id="B951EABB2000191CFF178BC1DC09F884" box="[192,329,1771,1795]" italics="true" pageId="82" pageNumber="81">Androctonus</emphasis>
(
<emphasis id="B951EABB2000191CFE898BC6DCEFF884" box="[350,431,1772,1795]" italics="true" pageId="82" pageNumber="81">Leiurus</emphasis>
) 5-
<emphasis id="B951EABB2000191CFE0B8BC6DF6CF884" box="[476,556,1772,1795]" italics="true" pageId="82" pageNumber="81">striatus</emphasis>
</taxonomicName>
:
<bibRefCitation id="EFB44B582000191CFD978BC1DCDBF8A5" author="Hemprich &amp; Ehrenberg" firstAuthor="Hemprich" journalOrPublisher="Verhandungen der Gesellschaft der Naturforschender Freunde in Berlin" page="353" pageId="82" pageNumber="81" pagination="348 - 362" part="1" refId="ref61033" refString="HEMPRICH, F. W. &amp; C. G. EHRENBERG. 1829. Vorlaufige Uebersicht der in Nord-Afrika und West-Asien einheimischen Skorpione und deren geographischen Verbreitung. Verhandungen der Gesellschaft der Naturforschender Freunde in Berlin, 1 (6): 348 - 362." title="Vorlaufige Uebersicht der in Nord-Afrika und West-Asien einheimischen Skorpione und deren geographischen Verbreitung" type="journal article" year="1829">Hemprich &amp; Ehrenberg, 1829: 353</bibRefCitation>
.
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</paragraph>
<paragraph id="8B9A36A92000191CFF178A02DFC7F8D8" blockId="82.[192,802,1771,1887]" pageId="82" pageNumber="81">
<treatmentCitationGroup id="AB3511872000191CFF178A02DFC7F8D8" pageId="82" pageNumber="81">
<treatmentCitation id="0A8410B82000191CFF178A02DFC7F8D8" author="HEMPRICH, F. W. &amp; C. G. EHRENBERG" pageId="82" pageNumber="81" year="1831">
<taxonomicName id="4C254D2A2000191CFF178A02DC5FF8D8" authority=": Ehrenberg" authorityName=": Ehrenberg" authorityYear="1831" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="82" pageNumber="81" phylum="Arthropoda" rank="subSpecies" species="quinquestriatus" subGenus="Liurus" subSpecies="aculeatus">
<emphasis id="B951EABB2000191CFF178A02DC09F8C7" box="[192,329,1832,1856]" italics="true" pageId="82" pageNumber="81">Androctonus</emphasis>
(
<emphasis id="B951EABB2000191CFE8C8A03DCE0F8C7" box="[347,416,1833,1856]" italics="true" pageId="82" pageNumber="81">Liurus</emphasis>
)
<emphasis id="B951EABB2000191CFE648A03DF88F8C7" box="[435,712,1832,1856]" italics="true" pageId="82" pageNumber="81">quinquestriatus aculeatus</emphasis>
: Ehrenberg
</taxonomicName>
in
<bibRefCitation id="EFB44B582000191CFE968A6DDFC3F8D8" author="Hemprich &amp; Ehrenberg" bookContentInfo="12 pp." box="[321,643,1863,1887]" firstAuthor="Hemprich" journalOrPublisher="Berolini ex Officina Academica, Venditur a Mittlero" pageId="82" pageNumber="81" refId="ref61080" refString="HEMPRICH, F. W. &amp; C. G. EHRENBERG. 1831. Animalia articulata, Arachnoidea. Scorpiones Africani et Asiatici. In: Symbolae Physicae. Animalia Evertebrata Exclusis Insectis Percensuit Dr. C. G. Ehrenberg. Series Prima cum Tabularum Decade Prima. Continet Animalia Africana et Asiatica 162. Berolini ex Officina Academica, Venditur a Mittlero, 12 pp. (Latin text, pages unnumbered)." title="Animalia articulata, Arachnoidea. Scorpiones Africani et Asiatici" type="book" volumeTitle="Symbolae Physicae. Animalia Evertebrata Exclusis Insectis Percensuit Dr. C. G. Ehrenberg. Series Prima cum Tabularum Decade Prima. Continet Animalia Africana et Asiatica 162" year="1831">Hemprich &amp; Ehrenberg, 1831</bibRefCitation>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A36A92000191CFCE88BC1D802F8A5" blockId="82.[831,1440,1771,1887]" pageId="82" pageNumber="81">
<treatmentCitationGroup id="AB3511872000191CFCE88BC1D802F8A5" pageId="82" pageNumber="81">
<treatmentCitation id="0A8410B82000191CFCE88BC1D8DCF884" author="Koch" box="[831,1436,1771,1795]" page="43" pageId="82" pageNumber="81" year="1841">
<taxonomicName id="4C254D2A2000191CFCE88BC1D8DCF884" authority=": C. L. Koch, 1841 a: 43 - 44" authorityName="C. L. Koch" authorityPageNumber="43 - 44" authorityYear="1841" box="[831,1436,1771,1795]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="82" pageNumber="81" phylum="Arthropoda" rank="species" species="quinquestriatus">
<emphasis id="B951EABB2000191CFCE88BC1D933F884" box="[831,1139,1771,1795]" italics="true" pageId="82" pageNumber="81">Androctonus quinquestriatus</emphasis>
: C. L. Koch, 1841a: 4344
</taxonomicName>
</treatmentCitation>
, pl. CCLXV, fig. 622; C. L.
<treatmentCitation id="0A8410B82000191CFB418A20D87EF8A5" author="Koch" box="[1174,1342,1802,1826]" page="90" pageId="82" pageNumber="81" year="1850">
<bibRefCitation id="EFB44B582000191CFB418A20D87EF8A5" author="Koch" box="[1174,1342,1802,1826]" firstAuthor="Koch" journalOrPublisher="C. H. Zeh'sche Buchhandlung, Nurnberg" page="90" pageId="82" pageNumber="81" pagination="86 - 92" part="5" refId="ref61602" refString="KOCH, C. L. 1850. Scorpionen. In Uebersicht des Arachnidensystems. C. H. Zeh'sche Buchhandlung, Nurnberg, 5: 86 - 92." title="Scorpionen" type="book chapter" volumeTitle="Uebersicht des Arachnidensystems" year="1850">Koch, 1850: 90</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A36A92000191CFCE88A02D94CF8D8" blockId="82.[831,1440,1771,1887]" pageId="82" pageNumber="81">
<treatmentCitationGroup id="AB3511872000191CFCE88A02D94CF8D8" pageId="82" pageNumber="81">
<treatmentCitation id="0A8410B82000191CFCE88A02D94CF8D8" author="GERVAIS, P. M." page="46" pageId="82" pageNumber="81" year="1844">
<taxonomicName id="4C254D2A2000191CFCE88A02D948F8D8" authority=": Gervais, 1844: 46" authorityName="Gervais" authorityPageNumber="46" authorityYear="1844" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="82" pageNumber="81" phylum="Arthropoda" rank="subSpecies" species="quinquestriatus" subGenus="Liurus" subSpecies="aculeatus">
<emphasis id="B951EABB2000191CFCE88A02DE87F8C7" box="[831,967,1832,1856]" italics="true" pageId="82" pageNumber="81">Androctonus</emphasis>
(
<emphasis id="B951EABB2000191CFC378A03D965F8C7" box="[992,1061,1833,1856]" italics="true" pageId="82" pageNumber="81">Liurus</emphasis>
)
<emphasis id="B951EABB2000191CFBE98A03D818F8C7" box="[1086,1368,1832,1856]" italics="true" pageId="82" pageNumber="81">quinquestriatus aculeatus</emphasis>
:
<bibRefCitation id="EFB44B582000191CFAB88A02D948F8D8" author="Gervais" bookContentInfo="418 pp." editor="WALCKENAER, C. A" firstAuthor="Gervais" journalOrPublisher="Librarie Encyclopedique de Roret, Paris" page="46" pageId="82" pageNumber="81" pagination="14 - 74" part="3" refId="ref60650" refString="GERVAIS, P. M. 1844. Scorpions. Pp. 14 - 74 in WALCKENAER, C. A (Ed.). Histoire Naturelle des Insectes. Apteres. Librarie Encyclopedique de Roret, Paris, 3 (8), 418 pp." title="Scorpions" type="book chapter" volumeTitle="Histoire Naturelle des Insectes. Apteres" year="1844">Gervais, 1844: 46</bibRefCitation>
</taxonomicName>
.
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</treatmentCitationGroup>
</paragraph>
<caption id="DF5A66212001191DFF178B8BD9F9F93E" ID-DOI="http://doi.org/10.5281/zenodo.6505759" ID-Zenodo-Dep="6505759" box="[192,1209,1697,1721]" httpUri="https://zenodo.org/record/6505759/files/figure.png" pageId="83" pageNumber="82" startId="83.[192,267,1697,1721]" targetBox="[191,1437,191,1666]" targetPageId="83">
<paragraph id="8B9A36A92001191DFF178B8BD9F9F93E" blockId="83.[192,1209,1697,1721]" box="[192,1209,1697,1721]" pageId="83" pageNumber="82">
<emphasis id="B951EABB2001191DFF178B8BDC75F93E" bold="true" box="[192,309,1697,1721]" pageId="83" pageNumber="82">Figure 65:</emphasis>
<taxonomicName id="4C254D2A2001191DFEEB8B8EDF31F93E" ID-CoL="6P6WQ" authority="(Birula, 1908)" baseAuthorityName="Birula" baseAuthorityYear="1908" box="[316,625,1699,1721]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="83" pageNumber="82" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB2001191DFEEB8B8EDCA3F93E" box="[316,483,1699,1721]" italics="true" pageId="83" pageNumber="82">Leiurus hebraeus</emphasis>
(
<bibRefCitation id="EFB44B582001191DFE278B89DF2AF93E" author="Birula" box="[496,618,1699,1721]" firstAuthor="Birula" journalOrPublisher="Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien" pageId="83" pageNumber="82" pagination="121 - 152" part="117" refId="ref59140" refString="BIRULA, A. 1908. Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 117 2 (1): 121 - 152." title="Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae" type="journal article" year="1908">Birula, 1908</bibRefCitation>
)
</taxonomicName>
stat. n., female. Habitus, ventral aspect. Israel (NHMB 17a).
</paragraph>
</caption>
<paragraph id="8B9A36A92001191DFF178BC6DC89F8A4" blockId="83.[192,802,1772,1919]" pageId="83" pageNumber="82">
<treatmentCitationGroup id="AB3511872001191DFF178BC6DC89F8A4" pageId="83" pageNumber="82">
<treatmentCitation id="0A8410B82001191DFF178BC6DC89F8A4" author="GERVAIS, P. M." page="46" pageId="83" pageNumber="82" year="1844">
<taxonomicName id="4C254D2A2001191DFF178BC6DF2FF883" baseAuthorityName="Ehrenberg" box="[192,623,1772,1796]" class="Arachnida" family="Buthidae" genus="Scorpio" kingdom="Animalia" order="Scorpiones" pageId="83" pageNumber="82" phylum="Arthropoda" rank="species" species="quinquestriatus" subGenus="Androctonus">
<emphasis id="B951EABB2001191DFF178BC6DC53F883" box="[192,275,1772,1796]" italics="true" pageId="83" pageNumber="82">Scorpio</emphasis>
(
<emphasis id="B951EABB2001191DFEF18BC6DCEEF883" box="[294,430,1772,1796]" italics="true" pageId="83" pageNumber="82">Androctonus</emphasis>
)
<emphasis id="B951EABB2001191DFE168BC7DF2FF883" box="[449,623,1773,1796]" italics="true" pageId="83" pageNumber="82">quinque-striatus</emphasis>
</taxonomicName>
:
<bibRefCitation id="EFB44B582001191DFD568BC6DC4DF8A4" author="Gervais" bookContentInfo="418 pp." editor="WALCKENAER, C. A" firstAuthor="Gervais" journalOrPublisher="Librarie Encyclopedique de Roret, Paris" page="46" pageId="83" pageNumber="82" pagination="14 - 74" part="3" refId="ref60650" refString="GERVAIS, P. M. 1844. Scorpions. Pp. 14 - 74 in WALCKENAER, C. A (Ed.). Histoire Naturelle des Insectes. Apteres. Librarie Encyclopedique de Roret, Paris, 3 (8), 418 pp." title="Scorpions" type="book chapter" volumeTitle="Histoire Naturelle des Insectes. Apteres" year="1844">Gervais, 1844: 46</bibRefCitation>
, pl. XXIV, fig. 2.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A36A92001191DFF178A00DFCCF8C5" blockId="83.[192,802,1772,1919]" box="[192,652,1834,1858]" pageId="83" pageNumber="82">
<treatmentCitationGroup id="AB3511872001191DFF178A00DFCCF8C5" box="[192,652,1834,1858]" pageId="83" pageNumber="82">
<treatmentCitation id="0A8410B82001191DFF178A00DFCCF8C5" author="Koch" box="[192,652,1834,1858]" page="90" pageId="83" pageNumber="82" year="1850">
<taxonomicName id="4C254D2A2001191DFF178A00DFC8F8C5" authority=": C. L. Koch, 1850: 90" authorityName="C. L. Koch" authorityPageNumber="90" authorityYear="1850" box="[192,648,1834,1858]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="83" pageNumber="82" phylum="Arthropoda" rank="species" species="troilus">
<emphasis id="B951EABB2001191DFF178A00DCD4F8C5" box="[192,404,1834,1858]" italics="true" pageId="83" pageNumber="82">Androctonus troilus</emphasis>
: C. L.
<bibRefCitation id="EFB44B582001191DFE088A00DFC8F8C5" author="Koch" box="[479,648,1834,1858]" firstAuthor="Koch" journalOrPublisher="C. H. Zeh'sche Buchhandlung, Nurnberg" page="90" pageId="83" pageNumber="82" pagination="86 - 92" part="5" refId="ref61602" refString="KOCH, C. L. 1850. Scorpionen. In Uebersicht des Arachnidensystems. C. H. Zeh'sche Buchhandlung, Nurnberg, 5: 86 - 92." title="Scorpionen" type="book chapter" volumeTitle="Uebersicht des Arachnidensystems" year="1850">Koch, 1850: 90</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A36A92001191AFF178A62DF42F9E0" blockId="83.[192,802,1772,1919]" lastBlockId="84.[192,802,1400,1914]" lastPageId="84" lastPageNumber="83" pageId="83" pageNumber="82">
<treatmentCitationGroup id="AB3511872001191AFF178A62DF42F9E0" lastPageId="84" lastPageNumber="83" pageId="83" pageNumber="82">
<treatmentCitation id="0A8410B82001191DFF178A62DF39F8E7" author="Koch" box="[192,633,1864,1888]" page="7" pageId="83" pageNumber="82" year="1875">
<taxonomicName id="4C254D2A2001191DFF178A62DF39F8E7" authority=": Koch, 1875: 7" authorityName="Koch" authorityPageNumber="7" authorityYear="1875" box="[192,633,1864,1888]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="83" pageNumber="82" phylum="Arthropoda" rank="species" species="quinquestriatus">
<emphasis id="B951EABB2001191DFF178A62DCFDF8E7" box="[192,445,1864,1888]" italics="true" pageId="83" pageNumber="82">Buthus quinquestriatus</emphasis>
:
<bibRefCitation id="EFB44B582001191DFE058A62DF39F8E7" author="Koch" bookContentInfo="96 pp." box="[466,633,1864,1888]" firstAuthor="Koch" journalOrPublisher="Nurnberg" page="7" pageId="83" pageNumber="82" refId="ref61632" refString="KOCH, L. 1875. Aegyptische und Abyssinische Arachniden Gesammelt von Herrn C. Jickeli Beschrieben und Abgebildet. Nurnberg. 96 pp." title="Aegyptische und Abyssinische Arachniden Gesammelt von Herrn C. Jickeli Beschrieben und Abgebildet" type="book" year="1875">Koch, 1875: 7</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82001191DFD5C8A62DC5BF8F8" author="SIMON" page="100" pageId="83" pageNumber="82" year="1879">
<bibRefCitation id="EFB44B582001191DFD5C8A62DC5BF8F8" author="Simon" firstAuthor="Simon" page="100" pageId="83" pageNumber="82" pagination="79 - 115" refId="ref63806" refString="SIMON, E. 1879. 3 e Ordre. - Scorpiones. Pp. 79 - 115 in: Les Arachnides de France. VII. Contenant les Ordres des Chernetes, Scorpiones et Opiliones. Roret, Paris, 332 pp." type="book chapter" year="1879">Simon, 1879: 100</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82001191DFEF08A4DDCA4F8F8" author="Karsch" box="[295,484,1895,1919]" page="89" pageId="83" pageNumber="82" year="1881">
<bibRefCitation id="EFB44B582001191DFEF08A4DDCA4F8F8" author="Karsch" box="[295,484,1895,1919]" firstAuthor="Karsch" journalOrPublisher="Berliner Entomologische Zeitschrift" page="89" pageId="83" pageNumber="82" pagination="15 - 17" part="25" refId="ref61405" refString="KARSCH, F. 1881. Arachniden und Myriopoden Micronesiens. Berliner Entomologische Zeitschrift, 25: 15 - 17." title="Arachniden und Myriopoden Micronesiens" type="journal article" year="1881">Karsch, 1881: 89</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82001191DFE388A4DDFE6F8F8" author="PAVESI" box="[495,678,1895,1919]" page="96" pageId="83" pageNumber="82" year="1883">
<bibRefCitation id="EFB44B582001191DFE388A4DDFE6F8F8" author="Pavesi" box="[495,678,1895,1919]" firstAuthor="Pavesi" page="96" pageId="83" pageNumber="82" pagination="5 - 106" refId="ref63113" refString="PAVESI, P. 1883. Studi sugli Aracnidi Africani. III. Aracnidi del regno di Scioa. Annali del Museo Civico di Storia Naturale di Genova, 20: 5 - 106." type="journal article" year="1883">Pavesi, 1883: 96</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82001191DFD668A4DDEBCF883" author="KRAEPELIN, K." box="[689,1020,1772,1919]" page="58" pageId="83" pageNumber="82" year="1891">
<bibRefCitation id="EFB44B582001191DFD668A4DDEBCF883" author="Kraepelin" box="[689,1020,1772,1919]" firstAuthor="Kraepelin" journalOrPublisher="Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten Hamburg" page="58 - 60" pageId="83" pageNumber="82" pagination="1 - 144" part="8" refId="ref61934" refString="KRAEPELIN, K. 1891. Revision der Skorpione. I. Die Familie der Androctonidae. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten Hamburg, 8: 1 - 144." title="Revision der Skorpione. I. Die Familie der Androctonidae" type="journal article" year="1891">Kraepelin, 1891: 5860</bibRefCitation>
</treatmentCitation>
, pl. I, fig. 7, pl. II, fig. 38 (in part);
<treatmentCitation id="0A8410B82001191DFCB88A21D93CF8A4" author="POCOCK" box="[879,1148,1803,1827]" page="242" pageId="83" pageNumber="82" year="1891">
<bibRefCitation id="EFB44B582001191DFCB88A21D93CF8A4" author="Pocock" box="[879,1148,1803,1827]" firstAuthor="Pocock" page="242 - 243" pageId="83" pageNumber="82" pagination="241 - 247" refId="ref63303" refString="POCOCK, R. I. 1891. Notes on some scorpions collected by Mr. J. J. Walker, with descriptions of two new species and a new genus. Annals and Magazine of Natural History, (6), 8: 241 - 247." type="journal article" year="1891">Pocock, 1891: 242243</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B82001191DFB288A21DE8CF8C5" author="THORELL" page="359" pageId="83" pageNumber="82" year="1893">
<bibRefCitation id="EFB44B582001191DFB288A21DE8CF8C5" author="Thorell" firstAuthor="Thorell" page="359 - 360" pageId="83" pageNumber="82" pagination="356 - 387" refId="ref64493" refString="THORELL, T. 1893. Scorpiones exotici R. Musei Historiae Naturalis Florentini. Bollettino della Societa Entomologica Italiana, 25: 356 - 387." type="journal article" year="1893">Thorell, 1893: 359360</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B82001191DFBEC8A00D847F8C5" author="POCOCK" box="[1083,1287,1834,1858]" page="299" pageId="83" pageNumber="82" year="1895">
<bibRefCitation id="EFB44B582001191DFBEC8A00D847F8C5" author="Pocock" box="[1083,1287,1834,1858]" firstAuthor="Pocock" page="299" pageId="83" pageNumber="82" pagination="292 - 316" refId="ref63353" refString="POCOCK, R. I. 1895. On the Arachnida and Myriopoda obtained by Dr. Anderson's collector during Mr T. Brent's expedition to the Hadramaut, South Arabia, with a supplement upon the scorpions obtained by Dr. Anderson in Egypt and the Eastern Soudan. Journal of the Linnean Society, 25: 292 - 316." type="journal article" year="1895">Pocock, 1895: 299</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82001191DFAC58A00DEDAF8E7" author="BIRULA, A. A." page="134" pageId="83" pageNumber="82" year="1898">
<bibRefCitation id="EFB44B582001191DFAC58A00DEDAF8E7" author="Birula" firstAuthor="Birula" journalOrPublisher="Horae Societatis Entomologicae Rossicae" page="134" pageId="83" pageNumber="82" pagination="132 - 140" part="33" refId="ref59108" refString="BIRULA, A. A. 1898. Ein Beitrag zur Kenntniss der Skorpionenfauna Kleinasiens. Horae Societatis Entomologicae Rossicae, 33 (1 - 2): 132 - 140." title="Ein Beitrag zur Kenntniss der Skorpionenfauna Kleinasiens" type="journal article" year="1898">Birula, 1898: 134</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82001191DFC7C8A62D9F3F8E7" author="KRAEPELIN, K." box="[939,1203,1864,1888]" page="27" pageId="83" pageNumber="82" year="1899">
<bibRefCitation id="EFB44B582001191DFC7C8A62D9F3F8E7" author="Kraepelin" box="[939,1203,1864,1888]" editor="Das Tierreich, R." firstAuthor="Kraepelin" journalOrPublisher="Friedlander und Sohn Verl Berlin" page="27 - 28" pageId="83" pageNumber="82" pagination="1 - 265" part="8" refId="ref61964" refString="KRAEPELIN, K. 1899. Scorpiones und Pedipalpi. In: Das Tierreich, R. Friedlander und Sohn Verl Berlin, 8: 1 - 265." title="Scorpiones und Pedipalpi" type="book chapter" year="1899">Kraepelin, 1899: 2728</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B82001191DFAE78A62D953F8F8" author="KRAEPELIN, K." page="266" pageId="83" pageNumber="82" year="1901">
<bibRefCitation id="EFB44B582001191DFAE78A62D953F8F8" author="Kraepelin" firstAuthor="Kraepelin" journalOrPublisher="Bulletin du Museum National d'Histoire Naturelle, Paris" page="266 - 267" pageId="83" pageNumber="82" pagination="263 - 265" part="7" refId="ref61993" refString="KRAEPELIN, K. 1901. Catalogue des scorpions des collections du Museum d'histoire naturelle de Paris. Bulletin du Museum National d'Histoire Naturelle, Paris, 7: 263 - 265." title="Catalogue des scorpions des collections du Museum d'histoire naturelle de Paris" type="journal article" year="1901">Kraepelin, 1901: 266267</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82001191DFBC98A42D9A9F8F8" author="WERNER" box="[1054,1257,1895,1919]" page="598" pageId="83" pageNumber="82" year="1902">
<bibRefCitation id="EFB44B582001191DFBC98A42D9A9F8F8" author="Werner" box="[1054,1257,1895,1919]" firstAuthor="Werner" page="598" pageId="83" pageNumber="82" pagination="595 - 608" refId="ref65207" refString="WERNER, F. 1902. Die Scorpione, Pedipalpen und Solifugen in der zoologisch-vergleichend-anatomischen Sammlung der Universitat Wien. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 52: 595 - 608." type="journal article" year="1902">Werner, 1902: 598</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82001191AFB238A4DDDBFFA17" author="Tullgren" lastPageId="84" lastPageNumber="83" page="3" pageId="83" pageNumber="82" year="1909">
<bibRefCitation id="EFB44B582001191AFB238A4DDDBFFA17" author="Tullgren" firstAuthor="Tullgren" lastPageId="84" lastPageNumber="83" page="3" pageId="83" pageNumber="82" pagination="1 - 12" refId="ref64520" refString="TULLGREN, A. 1909. Solifugae, Scorpiones und Chelonethi aus Agypten und dem Sudan. In: Results of the Swedish Zoological Expedition to Egypt and the White Nile 1901 under the Direction of L. A. Jagerskiold. The Library of the Royal University of Uppsala, 2, 21 A: 1 - 12." type="journal article" year="1909">Tullgren, 1909: 3</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82006191AFEC38852DCA1FA17" author="BIRULA, A. A." box="[276,481,1400,1424]" page="118" pageId="84" pageNumber="83" year="1910">
<bibRefCitation id="EFB44B582006191AFEC38852DCA1FA17" author="Birula" box="[276,481,1400,1424]" firstAuthor="Birula" journalOrPublisher="Horae Societatis Entomologicae Rossicae, St Petersburg" page="118" pageId="84" pageNumber="83" pagination="115 - 192" part="39" refId="ref59195" refString="BIRULA, A. 1910. Ueber Scorpio maurus Linne und seine Unterarten. Horae Societatis Entomologicae Rossicae, St Petersburg, 39: 115 - 192." title="Ueber Scorpio maurus Linne und seine Unterarten" type="journal article" year="1910">Birula, 1910: 118</bibRefCitation>
</treatmentCitation>
, 119120 (in part);
<treatmentCitation id="0A8410B82006191AFD028852DCC2FA29" author="SIMON" page="70" pageId="84" pageNumber="83" year="1910">
<bibRefCitation id="EFB44B582006191AFD028852DCC2FA29" author="Simon" firstAuthor="Simon" page="70 - 71" pageId="84" pageNumber="83" pagination="57 - 87" refId="ref63988" refString="SIMON, E. 1910. Revision des Scorpions d'Egypte. Bulletin de la Societe Entomologique d'Egypte (Cairo), 1910: 57 - 87." type="journal article" year="1910">Simon, 1910: 7071</bibRefCitation>
</treatmentCitation>
, fig. 9;
<treatmentCitation id="0A8410B82006191AFE3A88BCDF84FA29" author="BORELLI, A." box="[493,708,1430,1454]" page="461" pageId="84" pageNumber="83" year="1915">
<bibRefCitation id="EFB44B582006191AFE3A88BCDF84FA29" author="Borelli" box="[493,708,1430,1454]" firstAuthor="Borelli" journalOrPublisher="Societia Italiana di Scienze Naturali" page="461" pageId="84" pageNumber="83" pagination="456 - 464" part="53" refId="ref59538" refString="BORELLI, A. 1915. Gli scorpioni del Museo Civico di Storia Naturale di Milano. Atti della Societia Italiana di Scienze Naturali e del Museo Civico di Storia Natural in Milano. Societia Italiana di Scienze Naturali, 53: 456 - 464." title="Gli scorpioni del Museo Civico di Storia Naturale di Milano. Atti della Societia Italiana di Scienze Naturali e del Museo Civico di Storia Natural in Milano" type="journal article" year="1915">Borelli, 1915: 461</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82006191AFD0F88BCDC30FA4A" author="BIRULA, A." page="4" pageId="84" pageNumber="83" year="1917">
<bibRefCitation id="EFB44B582006191AFD0F88BCDC30FA4A" author="Birula" bookContentInfo="xx + 227 pp." firstAuthor="Birula" journalOrPublisher="Arachnides (Arachnoidea)" page="4 - 5" pageId="84" pageNumber="83" part="1" refId="ref59313" refString="BIRULA, A. A. 1917 b. Faune de la Russie et des pays Limitrophes Fondee Principalement sur les Collections du Musee Zoologique de l'Academie des Sciences de Russie. Arachnides (Arachnoidea), 1 (1): xx + 227 pp. (in Russian; preface dated October 1917). English translation: Byalynitskii-Birulya, A. A. 1965. Fauna of Russia and Adjacent Countries. Arachnoidea. Vol. I. Scorpions. Israel Program for Scientific Translations, Jerusalem, xix, 154 pp." title="Faune de la Russie et des pays Limitrophes Fondee Principalement sur les Collections du Musee Zoologique de l'Academie des Sciences de Russie" type="book" year="1917" yearSuffix="b">Birula, 1917b: 45</bibRefCitation>
</treatmentCitation>
, figs. 1a, 2;
<treatmentCitation id="0A8410B82006191AFDD1889CDF95FA4A" author="Lampe" box="[518,725,1461,1485]" page="191" pageId="84" pageNumber="83" year="1917">
<bibRefCitation id="EFB44B582006191AFDD1889CDF95FA4A" author="Lampe" box="[518,725,1461,1485]" firstAuthor="Lampe" journalOrPublisher="Jahrbucher des Nassauischen Vereine fur Naturkunde" page="191" pageId="84" pageNumber="83" pagination="185 - 203" part="70" refId="ref62105" refString="LAMPE, E. 1917. Katalog der Skorpione, Pedipalpen und Solifugen des Naturhistorischen Museums der Residenzstadt Wiesbaden. Jahrbucher des Nassauischen Vereine fur Naturkunde, 70 (1): 185 - 203." title="Katalog der Skorpione, Pedipalpen und Solifugen des Naturhistorischen Museums der Residenzstadt Wiesbaden" type="journal article" year="1917">Lampe, 1917: 191</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82006191AFD32889FDC15FA6B" author="King" page="82" pageId="84" pageNumber="83" year="1925">
<bibRefCitation id="EFB44B582006191AFD32889FDC15FA6B" author="King" firstAuthor="King" journalOrPublisher="Sudan Notes and Records" page="82" pageId="84" pageNumber="83" pagination="79 - 84" part="8" refId="ref61468" refString="KING, H. H. 1925. Notes on Sudan Scorpions. Sudan Notes and Records, 8: 79 - 84." title="Notes on Sudan Scorpions" type="journal article" year="1925">King, 1925: 82</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82006191AFEB488FEDF26FA6B" author="BORELLI, A." box="[355,614,1492,1516]" page="351" pageId="84" pageNumber="83" year="1927">
<bibRefCitation id="EFB44B582006191AFEB488FEDF26FA6B" author="Borelli" box="[355,614,1492,1516]" firstAuthor="Borelli" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" page="351 - 352" pageId="84" pageNumber="83" pagination="346 - 355" part="52" refId="ref59584" refString="BORELLI, A. 1927. Risultati zoologici della Missione inviata dalla R. Societa Geografica Italiana per l'esplorazione dell'oasi di Giarabub. Scorpioni e Solfughi. Annali del Museo Civico di Storia Naturale di Genova, 52: 346 - 355." title="Risultati zoologici della Missione inviata dalla R. Societa Geografica Italiana per l'esplorazione dell'oasi di Giarabub. Scorpioni e Solfughi" type="journal article" year="1927">Borelli, 1927: 351352</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82006191AFDA288FEDC05F98D" author="Gough &amp; Hirst" page="5" pageId="84" pageNumber="83" year="1927">
<bibRefCitation id="EFB44B582006191AFDA288FEDC05F98D" author="Gough &amp; Hirst" firstAuthor="Gough" journalOrPublisher="Bulletin of the Ministry of Agriculture of Egypt, Technical Scientific Service" page="5" pageId="84" pageNumber="83" pagination="1 - 7" part="76" refId="ref60790" refString="GOUGH, L. H. &amp; S. HIRST. 1927. Key to the identification of Egyptian scorpions. A method of identifying Egyptian scorpions from the fifth caudal segment and stings. Bulletin of the Ministry of Agriculture of Egypt, Technical Scientific Service, 76: 1 - 7." title="Key to the identification of Egyptian scorpions. A method of identifying Egyptian scorpions from the fifth caudal segment and stings" type="journal article" year="1927">Gough &amp; Hirst, 1927: 5</bibRefCitation>
</treatmentCitation>
, fig. 8;
<treatmentCitation id="0A8410B82006191AFE4B88D8DF26F98D" author="BORELLI, A." box="[412,614,1522,1546]" page="297" pageId="84" pageNumber="83" year="1929">
<bibRefCitation id="EFB44B582006191AFE4B88D8DF26F98D" author="Borelli" box="[412,614,1522,1546]" firstAuthor="Borelli" journalOrPublisher="Annals and Magazine of Natural History" page="297" pageId="84" pageNumber="83" pagination="297 - 300" part="10" refId="ref59627" refString="BORELLI, A. 1929. Scorpions du Soudan. Annals and Magazine of Natural History, 10 (3): 297 - 300." title="Scorpions du Soudan" type="journal article" year="1929">Borelli, 1929: 297</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82006191AFDA488D8DC5BF9AE" author="Schenkel" page="381" pageId="84" pageNumber="83" year="1932">
<bibRefCitation id="EFB44B582006191AFDA488D8DC5BF9AE" author="Schenkel" firstAuthor="Schenkel" page="381" pageId="84" pageNumber="83" pagination="375 - 396" refId="ref63734" refString="SCHENKEL, E. 1932. Notizen uber einige Scorpione und Solifugen. Revue Suisse de Zoologie, 39 (15): 375 - 396." type="journal article" year="1932">Schenkel, 1932: 381</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82006191AFEFC8B3BDF47F9AE" author="Taborsky" box="[299,519,1553,1577]" page="40" pageId="84" pageNumber="83" year="1934">
<bibRefCitation id="EFB44B582006191AFEFC8B3BDF47F9AE" author="Taborsky" box="[299,519,1553,1577]" firstAuthor="Taborsky" page="40" pageId="84" pageNumber="83" pagination="39 - 40" refId="ref64423" refString="TABORSKY, K. 1934. Sur les especes du genre Buthus Leach du Museum National de Prague. Sbornik Zoologickeho Oddeleni Narodniho Muzea v Praze, 1: 39 - 40." type="journal article" year="1934">Táborsky, 1934: 40</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B82006191AFD558B38DC58F9CF" author="WERNER" page="269" pageId="84" pageNumber="83" year="1934">
<bibRefCitation id="EFB44B582006191AFD558B38DC58F9CF" author="Werner" firstAuthor="Werner" page="269" pageId="84" pageNumber="83" pagination="8" refId="ref65242" refString="WERNER, F. 1934. Scorpiones, Pedipalpi. In: H. G. Bronns Klassen und Ordnungen des Tierreichs. Akademische Verlaggesellschaft, Leipzig. 5, IV, 8, Lief. 1 - 2, Scorpiones, pp. 1 - 316." type="journal article" year="1934">Werner, 1934: 269</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B82006191AFE408B1ADF2BF9CF" author="Moriggi" box="[407,619,1584,1608]" page="84" pageId="84" pageNumber="83" year="1941">
<bibRefCitation id="EFB44B582006191AFE408B1ADF2BF9CF" author="Moriggi" box="[407,619,1584,1608]" firstAuthor="Moriggi" page="84" pageId="84" pageNumber="83" pagination="77 - 103" refId="ref62728" refString="MORIGGI, M. 1941. Gli scorpioni dell'Africa Orientale Italiana. Rivista di Biologia Coloniale, 4 (1 - 2): 77 - 103." type="journal article" year="1941">Moriggi, 1941: 84</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82006191AFDAB8B1ADCD3F9E0" author="Abd-el-Wahab" page="111" pageId="84" pageNumber="83" year="1957">
<bibRefCitation id="EFB44B582006191AFDAB8B1ADCD3F9E0" author="Abd-el-Wahab" firstAuthor="Abd-el-Wahab" journalOrPublisher="Quarterly Journal of Microscopical Science" page="111 - 121" pageId="84" pageNumber="83" pagination="111 - 122" part="98" refId="ref58763" refString="ABD-EL-WAHAB, A. 1957. The male genital system of the scorpion, Buthus quinquestriatus. Quarterly Journal of Microscopical Science, 98 (1): 111 - 122." title="The male genital system of the scorpion, Buthus quinquestriatus" type="journal article" year="1957">Abd-el-Wahab, 1957: 111121</bibRefCitation>
</treatmentCitation>
, figs. 15.
</treatmentCitationGroup>
</paragraph>
<caption id="DF5A66212006191AFF17883EDC08FAC3" ID-DOI="http://doi.org/10.5281/zenodo.6505761" ID-Zenodo-Dep="6505761" httpUri="https://zenodo.org/record/6505761/files/figure.png" pageId="84" pageNumber="83" startId="84.[192,267,1300,1324]" targetBox="[204,1425,204,1259]" targetPageId="84">
<paragraph id="8B9A36A92006191AFF17883EDC08FAC3" blockId="84.[192,1440,1300,1348]" pageId="84" pageNumber="83">
<emphasis id="B951EABB2006191AFF17883EDC77FAAB" bold="true" box="[192,311,1300,1324]" pageId="84" pageNumber="83">Figure 66:</emphasis>
<taxonomicName id="4C254D2A2006191AFEE8883CDF39FAAC" ID-CoL="6P6WQ" authority="(Birula, 1908)" baseAuthorityName="Birula" baseAuthorityYear="1908" box="[319,633,1301,1323]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="84" pageNumber="83" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB2006191AFEE8883CDCA8FAAC" box="[319,488,1301,1323]" italics="true" pageId="84" pageNumber="83">Leiurus hebraeus</emphasis>
(
<bibRefCitation id="EFB44B582006191AFE21883FDF32FAAC" author="Birula" box="[502,626,1301,1323]" firstAuthor="Birula" journalOrPublisher="Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien" pageId="84" pageNumber="83" pagination="121 - 152" part="117" refId="ref59140" refString="BIRULA, A. 1908. Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 117 2 (1): 121 - 152." title="Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae" type="journal article" year="1908">Birula, 1908</bibRefCitation>
)
</taxonomicName>
stat. n., female.
<emphasis id="B951EABB2006191AFCCA883CDE75FAAC" bold="true" box="[797,821,1302,1323]" pageId="84" pageNumber="83">A.</emphasis>
Carapace and tergites.
<emphasis id="B951EABB2006191AFBCC883CD971FAAC" bold="true" box="[1051,1073,1302,1323]" pageId="84" pageNumber="83">B.</emphasis>
Coxosternal area and sternites. Israel (NHMB 17a).
</paragraph>
</caption>
<paragraph id="8B9A36A92006191AFF178B47DF2EF923" blockId="84.[192,802,1400,1914]" pageId="84" pageNumber="83">
<treatmentCitationGroup id="AB3511872006191AFF178B47DF2EF923" pageId="84" pageNumber="83">
<treatmentCitation id="0A8410B82006191AFF178B47DCD4F923" author="Birula" page="124" pageId="84" pageNumber="83" year="1908">
<taxonomicName id="4C254D2A2006191AFF178B47DCD4F923" authority="quinquestriatus: Birula, 1908: 124 - 129" authorityName="Birula" authorityPageNumber="124 - 129" authorityYear="1908" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="84" pageNumber="83" phylum="Arthropoda" rank="subSpecies" species="quinquestriatus" subGenus="Buthus" subSpecies="quinquestriatus">
<emphasis id="B951EABB2006191AFF178B47DC4AF902" box="[192,266,1645,1669]" italics="true" pageId="84" pageNumber="83">Buthus</emphasis>
(
<emphasis id="B951EABB2006191AFECC8B47DC25F902" box="[283,357,1645,1669]" italics="true" pageId="84" pageNumber="83">Buthus</emphasis>
)
<emphasis id="B951EABB2006191AFEA18B44DF89F902" box="[374,713,1646,1669]" italics="true" pageId="84" pageNumber="83">quinquestriatus quinquestriatus</emphasis>
:
<bibRefCitation id="EFB44B582006191AFD0F8B47DCD4F923" author="Birula" firstAuthor="Birula" journalOrPublisher="Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien" page="124 - 129" pageId="84" pageNumber="83" pagination="121 - 152" part="117" refId="ref59140" refString="BIRULA, A. 1908. Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 117 2 (1): 121 - 152." title="Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae" type="journal article" year="1908">Birula, 1908: 124129</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82006191AFE488BA6DF2EF923" author="Birula" box="[415,622,1676,1700]" page="223" pageId="84" pageNumber="83" year="1917">
<bibRefCitation id="EFB44B582006191AFE488BA6DF29F923" author="Birula" bookContentInfo="253 pp." box="[415,617,1676,1700]" firstAuthor="Birula" journalOrPublisher="Zapiski Kavkazskogo Muzeya (Memoires du Musee du Caucase), Tiflis, Imprimerie de la Chancellerie du Comite pour la Transcaucasie. Ser. A" page="223" pageId="84" pageNumber="83" part="5" refId="ref59223" refString="BIRULA, A. A. 1917 a. Arachnoidea Arthrogastra Caucasica. Pars I. Scorpiones. Zapiski Kavkazskogo Muzeya (Memoires du Musee du Caucase), Tiflis, Imprimerie de la Chancellerie du Comite pour la Transcaucasie. Ser. A, 5, 253 pp. (in Russian; published August 1917). English translation: Byalynitskii-Birulya, A. A. 1964. Arthrogastric Arachnids of Caucasia. 1. Scorpions. Israel Program for Scientific Translations, Jerusalem, 170 pp." title="Arachnoidea Arthrogastra Caucasica. Pars I. Scorpiones" type="book" year="1917" yearSuffix="a">Birula, 1917a: 223</bibRefCitation>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A36A92006191AFF178B80DC5FF966" blockId="84.[192,802,1400,1914]" pageId="84" pageNumber="83">
<treatmentCitationGroup id="AB3511872006191AFF178B80DC5FF966" pageId="84" pageNumber="83">
<treatmentCitation id="0A8410B82006191AFF178B80DC5FF966" author="BIRULA, A." page="118" pageId="84" pageNumber="83" year="1910">
<taxonomicName id="4C254D2A2006191AFF178B80DC5BF966" authority="quinquestriatus: Birula, 1910: 118" authorityName="Birula" authorityPageNumber="118" authorityYear="1910" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="84" pageNumber="83" phylum="Arthropoda" rank="subSpecies" species="quinquestriatus" subSpecies="quinquestriatus">
<emphasis id="B951EABB2006191AFF178B80DF34F945" box="[192,628,1706,1730]" italics="true" pageId="84" pageNumber="83">Buthus quinquestriatus quinquestriatus</emphasis>
:
<bibRefCitation id="EFB44B582006191AFD5B8B80DC5BF966" author="Birula" firstAuthor="Birula" journalOrPublisher="Horae Societatis Entomologicae Rossicae, St Petersburg" page="118" pageId="84" pageNumber="83" pagination="115 - 192" part="39" refId="ref59195" refString="BIRULA, A. 1910. Ueber Scorpio maurus Linne und seine Unterarten. Horae Societatis Entomologicae Rossicae, St Petersburg, 39: 115 - 192." title="Ueber Scorpio maurus Linne und seine Unterarten" type="journal article" year="1910">Birula, 1910: 118</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A36A92006191AFF178BC2DFC5F8BA" blockId="84.[192,802,1400,1914]" pageId="84" pageNumber="83">
<treatmentCitationGroup id="AB3511872006191AFF178BC2DFC5F8BA" pageId="84" pageNumber="83">
<treatmentCitation id="0A8410B82006191AFF178BC2DE62F887" author="BIRULA, A." box="[192,802,1768,1792]" page="23" pageId="84" pageNumber="83" year="213">
<taxonomicName id="4C254D2A2006191AFF178BC2DFADF887" authority=": Birula, 1917 a: 23" authorityName=": Birula" authorityPageNumber="23" authorityYear="1917" box="[192,749,1768,1792]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="84" pageNumber="83" phylum="Arthropoda" rank="species" species="quinquestriatus" subGenus="Buthus">
<emphasis id="B951EABB2006191AFF178BC2DC4AF887" box="[192,266,1768,1792]" italics="true" pageId="84" pageNumber="83">Buthus</emphasis>
(
<emphasis id="B951EABB2006191AFECB8BC2DC26F887" box="[284,358,1768,1792]" italics="true" pageId="84" pageNumber="83">Buthus</emphasis>
)
<emphasis id="B951EABB2006191AFEA08BC3DF5DF887" box="[375,541,1769,1792]" italics="true" pageId="84" pageNumber="83">quinquestriatus</emphasis>
:
<bibRefCitation id="EFB44B582006191AFDFA8BC2DFADF887" author="Birula" bookContentInfo="253 pp." box="[557,749,1768,1792]" firstAuthor="Birula" journalOrPublisher="Zapiski Kavkazskogo Muzeya (Memoires du Musee du Caucase), Tiflis, Imprimerie de la Chancellerie du Comite pour la Transcaucasie. Ser. A" page="23" pageId="84" pageNumber="83" part="5" refId="ref59223" refString="BIRULA, A. A. 1917 a. Arachnoidea Arthrogastra Caucasica. Pars I. Scorpiones. Zapiski Kavkazskogo Muzeya (Memoires du Musee du Caucase), Tiflis, Imprimerie de la Chancellerie du Comite pour la Transcaucasie. Ser. A, 5, 253 pp. (in Russian; published August 1917). English translation: Byalynitskii-Birulya, A. A. 1964. Arthrogastric Arachnids of Caucasia. 1. Scorpions. Israel Program for Scientific Translations, Jerusalem, 170 pp." title="Arachnoidea Arthrogastra Caucasica. Pars I. Scorpiones" type="book" year="1917" yearSuffix="a">Birula, 1917a: 23</bibRefCitation>
</taxonomicName>
, 213
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B82006191AFE898A2DDF00F898" author="BIRULA, A." box="[350,576,1799,1823]" page="79" pageId="84" pageNumber="83" year="1927">
<bibRefCitation id="EFB44B582006191AFE898A2DDF00F898" author="Birula" box="[350,576,1799,1823]" firstAuthor="Birula" journalOrPublisher="Denkschriften der Akademie der Wissenschaften in Wien. Mathemathischnaturwissenschaftliche Klasse" page="79 - 80" pageId="84" pageNumber="83" pagination="79 - 88" part="101" refId="ref59409" refString="BIRULA, A. A. 1927. Wissenschaftliche Ergebnisse der mit Unterstutzung der Akademie der Wissenschaften in Wien aus der Erbschaft Treitl von F. Werner unternommenen zoologischen Expedition nach dem Anglo-Aegyptischen Sudan (Kordofan) 1914. XXV. Skorpione. Denkschriften der Akademie der Wissenschaften in Wien. Mathemathischnaturwissenschaftliche Klasse, 101: 79 - 88." title="Wissenschaftliche Ergebnisse der mit Unterstutzung der Akademie der Wissenschaften in Wien aus der Erbschaft Treitl von F. Werner unternommenen zoologischen Expedition nach dem Anglo-Aegyptischen Sudan (Kordofan) 1914. XXV. Skorpione" type="journal article" year="1927">Birula, 1927: 7980</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82006191AFD878A2DDE62F898" author="WHITTICK" box="[592,802,1799,1823]" page="43" pageId="84" pageNumber="83" year="1941">
<bibRefCitation id="EFB44B582006191AFD878A2DDE62F898" author="Whittick" box="[592,802,1799,1823]" firstAuthor="Whittick" page="43" pageId="84" pageNumber="83" pagination="43 - 49" refId="ref65348" refString="WHITTICK, R. J. 1941. Arachnida: Scorpiones, Pedipalpi and Solifugae. In: Expedition to South-West Arabia, 1937 - 1938. Bulletin of the British Museum, Natural History, 1: 43 - 49." type="journal article" year="1941">Whittick, 1941: 43</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B82006191AFE828A0CDF60F8BA" author="ROEWER" box="[341,544,1829,1853]" page="206" pageId="84" pageNumber="83" year="1943">
<bibRefCitation id="EFB44B582006191AFE828A0CDF60F8BA" author="Roewer" box="[341,544,1829,1853]" firstAuthor="Roewer" page="206" pageId="84" pageNumber="83" pagination="205 - 244" refId="ref63705" refString="ROEWER, C. F. 1943. Uber eine neuerworbene Sammlung von Skorpionen des Natur-Museums Senckenberg. Senckenbergiana, 26 (4): 205 - 244." type="journal article" year="1943">Roewer, 1943: 206</bibRefCitation>
</treatmentCitation>
(in part).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A36A92006191AFF178A6FD98CF8BA" blockId="84.[192,802,1400,1914]" lastBlockId="84.[831,1441,1400,1914]" pageId="84" pageNumber="83">
<treatmentCitationGroup id="AB3511872006191AFF178A6FD98CF8BA" pageId="84" pageNumber="83">
<treatmentCitation id="0A8410B82006191AFF178A6FDFE4F8DB" author="VACHON" box="[192,676,1860,1884]" page="83" pageId="84" pageNumber="83" year="1949">
<taxonomicName id="4C254D2A2006191AFF178A6FDFE4F8DB" ID-CoL="6P78R" authority=": Vachon, 1949: 83" authorityName="Vachon" authorityPageNumber="83" authorityYear="1949" box="[192,676,1860,1884]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="84" pageNumber="83" phylum="Arthropoda" rank="species" species="quinquestriatus">
<emphasis id="B951EABB2006191AFF178A6FDC84F8DB" box="[192,452,1861,1884]" italics="true" pageId="84" pageNumber="83">Leiurus quinquestriatus</emphasis>
:
<bibRefCitation id="EFB44B582006191AFE0F8A6EDFE4F8DB" author="Vachon" box="[472,676,1860,1884]" firstAuthor="Vachon" page="83" pageId="84" pageNumber="83" pagination="66 - 100" refId="ref64580" refString="VACHON, M. 1949. Etudes sur les Scorpions (suite). Chapitre III (suite). Description des scorpions du Nord de l'Afrique. Archives de l'Institut Pasteur d'Algerie, 27 (1): 66 - 100, figs. 242 - 292." type="journal article" year="1949">Vachon, 1949: 83</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 8893 (in part);
<treatmentCitation id="0A8410B82006191AFEE48A48DF4FF8FD" author="VACHON" box="[307,527,1890,1914]" page="197" pageId="84" pageNumber="83" year="1950">
<bibRefCitation id="EFB44B582006191AFEE48A48DF4FF8FD" author="Vachon" box="[307,527,1890,1914]" firstAuthor="Vachon" page="197" pageId="84" pageNumber="83" pagination="152 - 216" refId="ref64629" refString="VACHON, M. 1950 a. Etudes sur les Scorpions. III (suite). Description des Scorpions du Nord de l'Afrique. Archives de l'Institut Pasteur d'Algerie, 28 (2): 152 - 216." type="journal article" year="1950" yearSuffix="a">Vachon, 1950a: 197</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B82006191AFD558A48DE8FFA17" author="VACHON" box="[642,975,1400,1914]" page="208" pageId="84" pageNumber="83" year="1952">
<bibRefCitation id="EFB44B582006191AFD558A48DE8FFA17" author="Vachon" box="[642,975,1400,1914]" firstAuthor="Vachon" page="208 - 213" pageId="84" pageNumber="83" refId="ref64708" refString="VACHON, M. 1952. Etude sur les Scorpions. Institut Pasteur d'Algerie, Alger, 482 pp." type="book" year="1952">Vachon, 1952: 208213</bibRefCitation>
</treatmentCitation>
, 369, 411, 441, 445, 449452, figs. 639, 678, 686 (in part);
<treatmentCitation id="0A8410B82006191AFB8388BCD871FA29" author="VACHON" box="[1108,1329,1430,1454]" page="211" pageId="84" pageNumber="83" year="1966">
<bibRefCitation id="EFB44B582006191AFB8388BCD871FA29" author="Vachon" box="[1108,1329,1430,1454]" firstAuthor="Vachon" page="211" pageId="84" pageNumber="83" pagination="209 - 218" refId="ref64772" refString="VACHON, M. 1966. Liste des Scorpions connus en Egypte, Arabie, Israel, Liban, Syrie, Jordanie, Turquie, Irak, Iran. Toxicon, 4: 209 - 218." type="journal article" year="1966">Vachon, 1966: 211</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82006191AFA9488BCDE9BFA4A" author="STAHNKE" page="130" pageId="84" pageNumber="83" year="1972">
<bibRefCitation id="EFB44B582006191AFA9488BCDE9BFA4A" author="Stahnke" firstAuthor="Stahnke" page="130" pageId="84" pageNumber="83" pagination="121 - 133" refId="ref64369" refString="STAHNKE, H. L. 1972. A key to the genera of Buthidae (Scorpionida). Entomological News, 83 (5): 121 - 133." type="journal article" year="1972">Stahnke, 1972: 130</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B82006191AFB9D889FD84DFA4A" author="PROBST" box="[1098,1293,1461,1485]" page="329" pageId="84" pageNumber="83" year="1973">
<bibRefCitation id="EFB44B582006191AFB9D889FD84DFA4A" author="Probst" box="[1098,1293,1461,1485]" firstAuthor="Probst" page="329" pageId="84" pageNumber="83" pagination="312 - 335" refId="ref63630" refString="PROBST, P. J. 1973. A review of the scorpions of East Africa with special regard to Kenya and Tanzania. Acta Tropica, 30 (4): 312 - 335. RASBAND, W. S. 1997 - 2011. ImageJ, U. S. National Institutes of Health, Bethesda, Maryland, USA, http: // imagej. nih. gov / ij /." type="journal article" year="1973">Probst, 1973: 329</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82006191AFACE889FDECAFA6B" author="PEREZ" page="24" pageId="84" pageNumber="83" year="1974">
<bibRefCitation id="EFB44B582006191AFACE889FDECAFA6B" author="Perez" firstAuthor="Perez" page="24" pageId="84" pageNumber="83" pagination="1 - 45" refId="ref63192" refString="PEREZ, S. M. 1974. Un inventario preliminar de los escorpiones de la region Paleartica y claves para la identificacion de los generos de la region Paleartica Occidental. Madrid: Universidad Complutense de Madrid, Facultad de Ciencias, Departamento de Zoologia, Catedra de Artropodos, 7: 1 - 45." type="journal article" year="1974">Pérez, 1974: 24</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B82006191AFBD688FED8E1FA6B" author="Lamoral &amp; Reynders" box="[1025,1441,1492,1516]" page="509" pageId="84" pageNumber="83" year="1975">
<bibRefCitation id="EFB44B582006191AFBD688FED8E1FA6B" author="Lamoral &amp; Reynders" box="[1025,1441,1492,1516]" firstAuthor="Lamoral" journalOrPublisher="Annals of the Natal Museum" page="509 - 510" pageId="84" pageNumber="83" pagination="489 - 576" part="22" refId="ref62059" refString="LAMORAL, B. H. &amp; S. C. REYNDERS. 1975. A catalogue of the scorpions described from the Ethiopian faunal region up to December 1973. Annals of the Natal Museum, 22 (2): 489 - 576." title="A catalogue of the scorpions described from the Ethiopian faunal region up to December 1973" type="journal article" year="1975">Lamoral &amp; Reynders, 1975: 509510</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B82006191AFC3788D9D870F98D" author="LEVY, G. &amp; P. AMITAI" box="[992,1328,1522,1547]" page="47" pageId="84" pageNumber="83" year="1980">
<bibRefCitation id="EFB44B582006191AFC3788D9D870F98D" author="Levy &amp; Amitai" bookContentInfo="130 pp" box="[992,1328,1522,1547]" firstAuthor="Levy" journalOrPublisher="The Israel Academy of Sciences and Humanities. Jerusalem" page="47 - 48" pageId="84" pageNumber="83" refId="ref62185" refString="LEVY, G. &amp; P. AMITAI. 1980. Scorpiones. Fauna Palaestina. Arachnida I. The Israel Academy of Sciences and Humanities. Jerusalem, 130 pp .." title="Scorpiones. Fauna Palaestina. Arachnida I" type="book" year="1980">Levy &amp; Amitai, 1980: 4748</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B82006191AFCB88B3BD976F9AE" author="Kinzelbach" box="[879,1078,1553,1577]" pageId="84" pageNumber="83" year="1985">
<bibRefCitation id="EFB44B582006191AFCB88B3BD976F9AE" author="Kinzelbach" box="[879,1078,1553,1577]" firstAuthor="Kinzelbach" journalOrPublisher="Tubinger Atlas des Vorderen Orients (TAVO). Karte A VI 14.2. Tubingen" pageId="84" pageNumber="83" refId="ref61523" refString="KINZELBACH, R. 1985. Vorder Orient. Skorpione (Arachnida: Scorpiones). Tubinger Atlas des Vorderen Orients (TAVO). Karte A VI 14.2. Tubingen." title="Vorder Orient. Skorpione (Arachnida: Scorpiones)" type="book" year="1985">Kinzelbach, 1985</bibRefCitation>
</treatmentCitation>
: map II (in part);
<treatmentCitation id="0A8410B82006191AFAC58B3BDE90F9CF" author="EL-HENNAWY, H. K." page="18" pageId="84" pageNumber="83" year="1987">
<bibRefCitation id="EFB44B582006191AFAC58B3BDE90F9CF" author="El-Hennawy" firstAuthor="El-Hennawy" journalOrPublisher="Serket" page="18" pageId="84" pageNumber="83" pagination="15 - 17" part="1" refId="ref60046" refString="EL-HENNAWY, H. K. 1987. A simplified key to Egyptian scorpion species (Arachnida: Scorpionida). Serket, 1 (1): 15 - 17." title="A simplified key to Egyptian scorpion species (Arachnida: Scorpionida)" type="journal article" year="1987">El-Hennawy, 1987: 18</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82006191AFC0A8B1AD994F9CF" author="EL-HENNAWY, H. K." box="[989,1236,1584,1608]" page="98" pageId="84" pageNumber="83" year="1992">
<bibRefCitation id="EFB44B582006191AFC0A8B1AD994F9CF" author="El-Hennawy" box="[989,1236,1584,1608]" firstAuthor="El-Hennawy" journalOrPublisher="Serket" page="98" pageId="84" pageNumber="83" pagination="95 - 153" part="2" refId="ref60101" refString="EL-HENNAWY, H. K. 1992. A catalogue of the scorpions described from the Arab Countries (1758 - 1990) (Arachnida: Scorpionida). Serket, 2 (4): 95 - 153." title="A catalogue of the scorpions described from the Arab Countries (1758 - 1990) (Arachnida: Scorpionida)" type="journal article" year="1992">El-Hennawy, 1992: 98</bibRefCitation>
</treatmentCitation>
, 125-126 (in part);
<treatmentCitation id="0A8410B82006191AFCB88B65D905F9E0" author="KOVARIK, F." box="[879,1093,1615,1639]" page="183" pageId="84" pageNumber="83" year="1992">
<bibRefCitation id="EFB44B582006191AFCB88B65D905F9E0" author="Kovarik" box="[879,1093,1615,1639]" firstAuthor="Kovarik" journalOrPublisher="Acta Societatis Zoologicae Bohemoslovenicae" page="183" pageId="84" pageNumber="83" pagination="181 - 186" part="56" refId="ref61657" refString="KOVARIK, F. 1992. A check list of scorpions (Arachnida: Scorpiones) in the collections of the zoological department, National Museum in Prague. Acta Societatis Zoologicae Bohemoslovenicae, 56: 181 - 186." title="A check list of scorpions (Arachnida: Scorpiones) in the collections of the zoological department, National Museum in Prague" type="journal article" year="1992">Kovařík, 1992: 183</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82006191AFB848B65D878F9E0" author="SISSOM" box="[1107,1336,1615,1639]" page="20" pageId="84" pageNumber="83" year="1994">
<bibRefCitation id="EFB44B582006191AFB848B65D878F9E0" author="Sissom" box="[1107,1336,1615,1639]" firstAuthor="Sissom" page="20 - 23" pageId="84" pageNumber="83" pagination="3 - 39" refId="ref64059" refString="SISSOM, W. D. 1994. Descriptions of new and poorly known scorpions of Yemen (Scorpiones: Buthidae, Diplocentridae, Scorpionidae). Fauna of Saudi Arabia, 14: 3 - 39." type="journal article" year="1994">Sissom, 1994: 2023</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B82006191AFCB88B47D9FAF902" author="BRAUNWALDER, M. E. &amp; V. FET" box="[879,1210,1645,1669]" page="32" pageId="84" pageNumber="83" year="1998">
<bibRefCitation id="EFB44B582006191AFCB88B47D9FAF902" author="Braunwalder &amp; Fet" box="[879,1210,1645,1669]" firstAuthor="Braunwalder" journalOrPublisher="Bulletin of the British Arachnological Society" page="32" pageId="84" pageNumber="83" pagination="29 - 35" part="11" refId="ref59689" refString="BRAUNWALDER, M. E. &amp; V. FET. 1998. On publications about scorpions (Arachnida: Scorpiones) by Hemprich and Ehrenberg (1828 - 1831). Bulletin of the British Arachnological Society, 11 (1): 29 - 35." title="On publications about scorpions (Arachnida: Scorpiones) by Hemprich and Ehrenberg (1828 - 1831)" type="journal article" year="1998">Braunwalder &amp; Fet, 1998: 32</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82006191AFB1F8B47D8E0F902" author="KOVARIK, F." box="[1224,1440,1645,1669]" page="112" pageId="84" pageNumber="83" year="1998">
<bibRefCitation id="EFB44B582006191AFB1F8B47D8E0F902" author="Kovarik" bookContentInfo="175 pp." box="[1224,1440,1645,1669]" firstAuthor="Kovarik" journalOrPublisher="Madagaskar, Jihlava" page="112" pageId="84" pageNumber="83" refId="ref61762" refString="KOVARIK, F. 1998. Stiri. Madagaskar, Jihlava, 175 pp." title="Stiri" type="book" year="1998">Kovařík, 1998: 112</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B82006191AFC048BA6D98DF923" author="Fet &amp; Lowe" box="[979,1229,1676,1700]" page="155" pageId="84" pageNumber="83" year="2000">
<bibRefCitation id="EFB44B582006191AFC048BA6D98DF923" author="Fet &amp; Lowe" box="[979,1229,1676,1700]" editor="FET, V. &amp; W. D. SISSOM &amp; G. LOWE &amp; M. E. BRAUNWALDER" firstAuthor="Fet" journalOrPublisher="The New York Entomological Society" page="155" pageId="84" pageNumber="83" pagination="54 - 286" refId="ref60347" refString="FET, V. &amp; G. LOWE. 2000. Family Buthidae. Pp. 54 - 286 in FET, V., W. D. SISSOM, G. LOWE &amp; M. E. BRAUNWALDER. Catalog of the Scorpions of the World (1758 - 1998). The New York Entomological Society." title="Family Buthidae" type="book chapter" volumeTitle="Catalog of the Scorpions of the World (1758 - 1998)" year="2000">Fet &amp; Lowe, 2000: 155</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B82006191AFAEE8BA6D94DF945" author="LOURENCO" page="98" pageId="84" pageNumber="83" year="2006">
<bibRefCitation id="EFB44B582006191AFAEE8BA6D94DF945" author="Lourenco" etAl="et al." firstAuthor="Lourenco" page="98" pageId="84" pageNumber="83" pagination="97 - 101" refId="ref62489" refString="LOURENCO, W. R., J-X. QI &amp; J. L. CLOUDSLEY- THOMPSON. 2006. The African species of the genus Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) with the description of a new species. Boletin Sociedad Entomologica Aragonesa, 39: 97 - 101." type="journal article" year="2006">Lourenço et al. 2006: 98</bibRefCitation>
</treatmentCitation>
, 100 (in part), fig. 13;
<treatmentCitation id="0A8410B82006191AFAC48B80DE98F966" author="EL-HENNAWY, H. K." page="76" pageId="84" pageNumber="83" year="2005">
<bibRefCitation id="EFB44B582006191AFAC48B80DE98F966" author="El-Hennawy" firstAuthor="El-Hennawy" journalOrPublisher="Serket" page="76" pageId="84" pageNumber="83" pagination="73 - 84" part="9" refId="ref60141" refString="EL-HENNAWY, H. K. 2005. Arachnids in Mediterranean protected areas of Egypt. Serket, 9 (3): 73 - 84." title="Arachnids in Mediterranean protected areas of Egypt" type="journal article" year="2005">El-Hennawy, 2005: 76</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82006191AFC3C8BE3D9BEF966" author="EL-HENNAWY, H. K." box="[1003,1278,1737,1761]" page="118" pageId="84" pageNumber="83" year="2008">
<bibRefCitation id="EFB44B582006191AFC3C8BE3D9BEF966" author="El-Hennawy" box="[1003,1278,1737,1761]" firstAuthor="El-Hennawy" journalOrPublisher="Revista Iberica de Aracnologia" page="118" pageId="84" pageNumber="83" pagination="115 - 121" part="15" refId="ref60168" refString="EL-HENNAWY, H. K. 2008. Arachnids of Elba protected area in the southern part of the eastern desert of Egypt. Revista Iberica de Aracnologia, 15: 115 - 121." title="Arachnids of Elba protected area in the southern part of the eastern desert of Egypt" type="journal article" year="2008">El-Hennawy, 2008: 118</bibRefCitation>
</treatmentCitation>
, 121, tab IV;
<treatmentCitation id="0A8410B82006191AFCB88BC3D81FF887" author="LOURENCO, W. R. &amp; J. L. CLOUDSLEY- THOMPSON" box="[879,1375,1768,1792]" page="126" pageId="84" pageNumber="83" year="2009">
<bibRefCitation id="EFB44B582006191AFCB88BC3D868F887" author="LOURENCO, W. R. &amp; J. L. CLOUDSLEY- THOMPSON" box="[879,1320,1768,1792]" firstAuthor="Lourenco" journalOrPublisher="Boletin Sociedad Entomologica Aragonesa" pageId="84" pageNumber="83" pagination="123 - 126" part="45" refId="ref62301" refString="LOURENCO, W. R. &amp; J. L. CLOUDSLEY- THOMPSON. 2009. A new species of the genus Scorpio Linnaeus 1758 from Sudan (Scorpiones, Scorpionidae). Boletin Sociedad Entomologica Aragonesa, 45: 123 - 126." title="A new species of the genus Scorpio Linnaeus 1758 from Sudan (Scorpiones, Scorpionidae)" type="journal article" year="2009">Lourenço &amp; Cloudsley-Thompson, 2009</bibRefCitation>
: 126
</treatmentCitation>
;
<treatmentCitation id="0A8410B82006191AFABC8BC2D993F898" author="YAGMUR, E. A. &amp; H. KOC &amp; K. B. KUNT" page="2" pageId="84" pageNumber="83" year="2009">
<bibRefCitation id="EFB44B582006191AFABC8BC2D96FF898" author="YAGMUR, E. A. &amp; H. KOC &amp; K. B. KUNT" journalOrPublisher="Euscorpius" pageId="84" pageNumber="83" pagination="1 - 20" part="85" refId="ref65464" refString="YAGMUR, E. A., H. KOC &amp; K. B. KUNT. 2009. Des- cription of a new species of Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) from Southeastern Turkey. Euscorpius, 85: 1 - 20." title="Des- cription of a new species of Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) from Southeastern Turkey" type="journal article" year="2009">Yağmur et al., 2009:</bibRefCitation>
23 (in part);
</treatmentCitation>
<treatmentCitation id="0A8410B82006191AFB088A2DD98CF8BA" author="KOVARIK, F &amp; A. A. OJANGUREN AFFILASTRO" page="158" pageId="84" pageNumber="83" year="2013">
<bibRefCitation id="EFB44B582006191AFB088A2DDEB2F8BA" author="KOVARIK, F &amp; A. A. OJANGUREN AFFILASTRO" bookContentInfo="400 pp." firstAuthor="Kovarik" journalOrPublisher="Clairon Production, Prague" pageId="84" pageNumber="83" refId="ref61886" refString="KOVARIK, F &amp; A. A. OJANGUREN AFFILASTRO. 2013. Illustrated catalog of scorpions. Part II. Bothriuridae; Chaerilidae; Buthidae I., genera Compsobuthus, Hottentotta, Isometrus, Lychas and Sassanidotus. Clairon Production, Prague, 400 pp." title="Illustrated catalog of scorpions. Part II. Bothriuridae; Chaerilidae; Buthidae I., genera Compsobuthus, Hottentotta, Isometrus, Lychas and Sassanidotus" type="book" year="2013">Kovařík &amp; Ojanguren, 2013</bibRefCitation>
: 158, figs. 836838.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A36A92006191BFCE88A6FDF20F89A" blockId="84.[831,1441,1400,1914]" lastBlockId="85.[192,802,1764,1887]" lastPageId="85" lastPageNumber="84" pageId="84" pageNumber="83">
<treatmentCitationGroup id="AB3511872006191BFCE88A6FDF20F89A" lastPageId="85" lastPageNumber="84" pageId="84" pageNumber="83">
<treatmentCitation id="0A8410B82006191AFCE88A6FDEF2F8FD" author="VACHON" page="93" pageId="84" pageNumber="83" year="1949">
<taxonomicName id="4C254D2A2006191AFCE88A6FDEF2F8FD" ID-CoL="6P78R" authority="quinquestriatus: Vachon, 1949: 88 - 93" authorityName="Vachon" authorityPageNumber="88 - 93" authorityYear="1949" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="84" pageNumber="83" phylum="Arthropoda" rank="subSpecies" species="quinquestriatus" subSpecies="quinquestriatus">
<emphasis id="B951EABB2006191AFCE88A6FD9AFF8DB" box="[831,1263,1861,1884]" italics="true" pageId="84" pageNumber="83">Leiurus quinquestriatus quinquestriatus</emphasis>
:
<bibRefCitation id="EFB44B582006191AFAD78A6EDEF2F8FD" author="Vachon" firstAuthor="Vachon" page="88 - 93" pageId="84" pageNumber="83" pagination="66 - 100" refId="ref64580" refString="VACHON, M. 1949. Etudes sur les Scorpions (suite). Chapitre III (suite). Description des scorpions du Nord de l'Afrique. Archives de l'Institut Pasteur d'Algerie, 27 (1): 66 - 100, figs. 242 - 292." type="journal article" year="1949">Vachon, 1949: 8893</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(in part);
<treatmentCitation id="0A8410B82006191AFBCB8A48D986F8FD" author="PEREZ" box="[1052,1222,1890,1914]" page="24" pageId="84" pageNumber="83" year="1974">
<bibRefCitation id="EFB44B582006191AFBCB8A48D986F8FD" author="Perez" box="[1052,1222,1890,1914]" firstAuthor="Perez" page="24" pageId="84" pageNumber="83" pagination="1 - 45" refId="ref63192" refString="PEREZ, S. M. 1974. Un inventario preliminar de los escorpiones de la region Paleartica y claves para la identificacion de los generos de la region Paleartica Occidental. Madrid: Universidad Complutense de Madrid, Facultad de Ciencias, Departamento de Zoologia, Catedra de Artropodos, 7: 1 - 45." type="journal article" year="1974">Pérez, 1974: 24</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82006191BFB068A49DC5BF97B" author="FET, V. &amp; G. LOWE" lastPageId="85" lastPageNumber="84" page="155" pageId="84" pageNumber="83" year="2000">
<bibRefCitation id="EFB44B582006191BFB068A49DC5BF97B" author="Fet &amp; Lowe" editor="FET, V. &amp; W. D. SISSOM &amp; G. LOWE &amp; M. E. BRAUNWALDER" firstAuthor="Fet" journalOrPublisher="The New York Entomological Society" lastPageId="85" lastPageNumber="84" page="155" pageId="84" pageNumber="83" pagination="54 - 286" refId="ref60347" refString="FET, V. &amp; G. LOWE. 2000. Family Buthidae. Pp. 54 - 286 in FET, V., W. D. SISSOM, G. LOWE &amp; M. E. BRAUNWALDER. Catalog of the Scorpions of the World (1758 - 1998). The New York Entomological Society." title="Family Buthidae" type="book chapter" volumeTitle="Catalog of the Scorpions of the World (1758 - 1998)" year="2000">Fet &amp; Lowe, 2000: 155</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82007191BFEF98BCFDF20F97B" author="LOURENCO" box="[302,608,1764,1788]" page="641" pageId="85" pageNumber="84" year="2002">
<bibRefCitation id="EFB44B582007191BFEF98BCFDF20F97B" author="Lourenco" box="[302,608,1764,1788]" etAl="et al." firstAuthor="Lourenco" page="641" pageId="85" pageNumber="84" pagination="635 - 642" refId="ref62388" refString="LOURENCO, W. R., D. MODRY &amp; Z. AMR. 2002. Description of a new species of Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) from the south of Jordan. Revue suisse de Zoologie, 109 (3): 635 - 642." type="journal article" year="2002">Lourenço et al. 2002: 641</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82007191BFDA48BCFDC11F89A" author="LOURENCO" page="97" pageId="85" pageNumber="84" year="2006">
<bibRefCitation id="EFB44B582007191BFDA48BCFDC11F89A" author="Lourenco" etAl="et al." firstAuthor="Lourenco" page="97" pageId="85" pageNumber="84" pagination="97 - 101" refId="ref62489" refString="LOURENCO, W. R., J-X. QI &amp; J. L. CLOUDSLEY- THOMPSON. 2006. The African species of the genus Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) with the description of a new species. Boletin Sociedad Entomologica Aragonesa, 39: 97 - 101." type="journal article" year="2006">Lourenço et al. 2006: 97</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410B82007191BFE8B8A2FDF1CF89A" author="YAGMUR" box="[348,604,1797,1821]" page="12" pageId="85" pageNumber="84" year="2009">
<bibRefCitation id="EFB44B582007191BFE8B8A2FDF1CF89A" author="Yagmur" box="[348,604,1797,1821]" etAl="et al." firstAuthor="Yagmur" page="12" pageId="85" pageNumber="84" pagination="1 - 20" refId="ref65464" refString="YAGMUR, E. A., H. KOC &amp; K. B. KUNT. 2009. Des- cription of a new species of Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) from Southeastern Turkey. Euscorpius, 85: 1 - 20." type="journal article" year="2009">Yağmur et al., 2009: 12</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<caption id="DF5A66212007191BFF178B42DFA2F936" ID-DOI="http://doi.org/10.5281/zenodo.6505763" ID-Zenodo-Dep="6505763" httpUri="https://zenodo.org/record/6505763/files/figure.png" pageId="85" pageNumber="84" startId="85.[192,267,1640,1664]" targetBox="[191,1440,133,1609]" targetPageId="85">
<paragraph id="8B9A36A92007191BFF178B42DFA2F936" blockId="85.[192,1441,1640,1713]" pageId="85" pageNumber="84">
<emphasis id="B951EABB2007191BFF178B42DC77F907" bold="true" box="[192,311,1640,1664]" pageId="85" pageNumber="84">Figure 67:</emphasis>
<taxonomicName id="4C254D2A2007191BFEE88B40DF39F9F8" ID-CoL="6P6WQ" authority="(Birula, 1908)" baseAuthorityName="Birula" baseAuthorityYear="1908" box="[319,633,1641,1663]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="85" pageNumber="84" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB2007191BFEE88B40DCA8F9F8" box="[319,488,1641,1663]" italics="true" pageId="85" pageNumber="84">Leiurus hebraeus</emphasis>
(
<bibRefCitation id="EFB44B582007191BFE218B43DF31F9F8" author="Birula" box="[502,625,1641,1663]" firstAuthor="Birula" journalOrPublisher="Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien" pageId="85" pageNumber="84" pagination="121 - 152" part="117" refId="ref59140" refString="BIRULA, A. 1908. Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 117 2 (1): 121 - 152." title="Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae" type="journal article" year="1908">Birula, 1908</bibRefCitation>
)
</taxonomicName>
stat. n., female. Right pedipalp.
<emphasis id="B951EABB2007191BFC6D8B43DE91F9F9" bold="true" box="[954,977,1641,1662]" pageId="85" pageNumber="84">A.</emphasis>
Femur, dorsal aspect.
<emphasis id="B951EABB2007191BFB678B43D986F9F9" bold="true" box="[1200,1222,1641,1662]" pageId="85" pageNumber="84">B.</emphasis>
Patella, dorsal aspect.
<emphasis id="B951EABB2007191BFF178BA8DD97F910" bold="true" box="[192,215,1666,1687]" pageId="85" pageNumber="84">C.</emphasis>
Patella, external aspect.
<emphasis id="B951EABB2007191BFE0F8BA8DCB0F910" bold="true" box="[472,496,1666,1687]" pageId="85" pageNumber="84">D.</emphasis>
Chela, ventral aspect.
<emphasis id="B951EABB2007191BFD0A8BA8DFB3F910" bold="true" box="[733,755,1666,1687]" pageId="85" pageNumber="84">E.</emphasis>
Chela, dorsal aspect.
<emphasis id="B951EABB2007191BFC0E8BA8DEADF910" bold="true" box="[985,1005,1666,1687]" pageId="85" pageNumber="84">F.</emphasis>
Fixed finger dentition.
<emphasis id="B951EABB2007191BFB348BA8D9BCF910" bold="true" box="[1251,1276,1666,1687]" pageId="85" pageNumber="84">G.</emphasis>
Movable finger dentition.
<emphasis id="B951EABB2007191BFEF58BB1DC7BF937" bold="true" box="[290,315,1691,1712]" pageId="85" pageNumber="84">H.</emphasis>
Chela, external aspect. Israel (NHMB 17a).
</paragraph>
</caption>
<paragraph id="8B9A36A92007191BFF178A0CDCC5F8D8" blockId="85.[192,802,1764,1887]" pageId="85" pageNumber="84">
<treatmentCitationGroup id="AB3511872007191BFF178A0CDCC5F8D8" pageId="85" pageNumber="84">
<treatmentCitation id="0A8410B82007191BFF178A0CDCC5F8D8" author="Braunwalder &amp; Fet" page="33" pageId="85" pageNumber="84" year="1998">
<taxonomicName id="4C254D2A2007191BFF178A0CDCC1F8D8" authority=": Braunwalder &amp; Fet, 1998: 33" authorityName="Braunwalder &amp; Fet" authorityPageNumber="33" authorityYear="1998" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="85" pageNumber="84" phylum="Arthropoda" rank="species" species="quinquestriatus" subGenus="Leiurus">
<emphasis id="B951EABB2007191BFF178A0CDC08F8B9" box="[192,328,1830,1854]" italics="true" pageId="85" pageNumber="84">Androctonus</emphasis>
(
<emphasis id="B951EABB2007191BFE8F8A0DDCE9F8B9" box="[344,425,1831,1854]" italics="true" pageId="85" pageNumber="84">Leiurus</emphasis>
)
<emphasis id="B951EABB2007191BFE6E8A0DDF1EF8B9" box="[441,606,1831,1854]" italics="true" pageId="85" pageNumber="84">quinquestriatus</emphasis>
:
<bibRefCitation id="EFB44B582007191BFDBA8A0CDCC1F8D8" author="Braunwalder &amp; Fet" firstAuthor="Braunwalder" journalOrPublisher="Bulletin of the British Arachnological Society" page="33" pageId="85" pageNumber="84" pagination="29 - 35" part="11" refId="ref59689" refString="BRAUNWALDER, M. E. &amp; V. FET. 1998. On publications about scorpions (Arachnida: Scorpiones) by Hemprich and Ehrenberg (1828 - 1831). Bulletin of the British Arachnological Society, 11 (1): 29 - 35." title="On publications about scorpions (Arachnida: Scorpiones) by Hemprich and Ehrenberg (1828 - 1831)" type="journal article" year="1998">Braunwalder &amp; Fet, 1998: 33</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A36A92007191BFCE88BC9D806F8B2" blockId="85.[831,1439,1763,1845]" pageId="85" pageNumber="84">
NOTE: The above synonymy and reference listing omits records from
<collectingCountry id="F33276392007191BFC0C8A2AD95BF89F" box="[987,1051,1792,1816]" name="Libya" pageId="85" pageNumber="84">Libya</collectingCountry>
,
<collectingCountry id="F33276392007191BFBF98A2AD93EF89F" box="[1070,1150,1792,1816]" name="Algeria" pageId="85" pageNumber="84">Algeria</collectingCountry>
and the southern Sinai, as we did not revise the status of those populations.
</paragraph>
</subSubSection>
<subSubSection id="C33F652220071919FCE88A61DFB8F948" lastPageId="87" lastPageNumber="86" pageId="85" pageNumber="84" type="materials_examined">
<paragraph id="8B9A36A920071919FCE88A61DE49FAA6" blockId="85.[831,1433,1866,1891]" lastBlockId="87.[192,802,1228,1313]" lastPageId="87" lastPageNumber="86" pageId="85" pageNumber="84">
<collectionCode id="ED34AE6C2007191BFCE88A61D95CF8E5" box="[831,1052,1866,1890]" pageId="85" pageNumber="84">TYPE MATERIAL</collectionCode>
.
<materialsCitation id="3B4D3CF420071919FBFB8A60DE49FAA6" ID-GBIF-Occurrence="3913646503" collectionCode="ZMHB" lastPageId="87" lastPageNumber="86" pageId="85" pageNumber="84" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="syntype">
8 original
<typeStatus id="549E880B2007191BFB708A61D844F8E4" box="[1191,1284,1867,1891]" pageId="85" pageNumber="84" type="syntype">syntypes</typeStatus>
(
<specimenCount id="9D23FD202007191BFACC8A60D805F8E5" box="[1307,1349,1866,1890]" pageId="85" pageNumber="84" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D23FD202007191BFA808A60D83FF8E5" box="[1367,1407,1866,1890]" pageId="85" pageNumber="84" type="female">2 ♀</specimenCount>
, 4 juv):
<emphasis id="B951EABB20051919FF2B89E6DC00FB63" bold="true" box="[252,320,1228,1252]" pageId="87" pageNumber="86">
<collectingCountry id="F332763920051919FF2B89E6DC00FB63" box="[252,320,1228,1252]" name="Egypt" pageId="87" pageNumber="86">Egypt</collectingCountry>
</emphasis>
,
<collectingRegion id="49E1F84B20051919FE8789E6DCCAFB63" box="[336,394,1228,1252]" country="Egypt" name="Shamal Sina'" pageId="87" pageNumber="86">Sinai</collectingRegion>
; Arabian Desert;
<emphasis id="B951EABB20051919FD9B89E6DFD4FB63" bold="true" box="[588,660,1228,1252]" pageId="87" pageNumber="86">
<collectingCountry id="F332763920051919FD9B89E6DFD4FB63" box="[588,660,1228,1252]" name="Sudan" pageId="87" pageNumber="86">Sudan</collectingCountry>
</emphasis>
, Dongola (= Dunqulah, Nubia); 18231824, leg. F.W. Hemprich &amp; C.G. Ehrenberg (
<collectionCode id="ED34AE6C20051919FEA28823DC84FAA6" box="[373,452,1289,1313]" pageId="87" pageNumber="86">ZMHB</collectionCode>
No. 140) (examined,
<figureCitation id="131E2A2C20051919FD798823DFBDFAA6" box="[686,765,1289,1313]" captionStart="Figure 82" captionStartId="100.[192,267,1070,1094]" captionTargetBox="[207,1412,188,1044]" captionTargetId="figure-431@100.[192,1439,168,1065]" captionTargetPageId="100" captionText="Figure 82: Leiurus quinquestriatus (Ehrenberg, 1828), 5 of 8 syntypes (ZMHB No. 140). Females: A, #12C01-3; B, #12C04-6. Males: C, #12C07-9; D, #12C10-3. Juvenile: E, #12C14-6. Egypt; Sudan. Scale bars (10 mm) apply to the habitus photos. Body lengths: A, 90 mm; B, 88 mm; C, 63 mm; D, 69 mm; E, 65 mm. Not shown are three additional fragmented juvenile syntypes." figureDoi="http://doi.org/10.5281/zenodo.6505796" httpUri="https://zenodo.org/record/6505796/files/figure.png" pageId="87" pageNumber="86">Fig. 82</figureCitation>
).
</materialsCitation>
</paragraph>
<caption id="DF5A662120041918FF178A3BDEFBF8C6" ID-DOI="http://doi.org/10.5281/zenodo.6505765" ID-Zenodo-Dep="6505765" httpUri="https://zenodo.org/record/6505765/files/figure.png" pageId="86" pageNumber="85" startId="86.[192,267,1809,1833]" targetBox="[263,1361,215,1766]" targetPageId="86">
<paragraph id="8B9A36A920041918FF178A3BDEFBF8C6" blockId="86.[192,1441,1809,1857]" pageId="86" pageNumber="85">
<emphasis id="B951EABB20041918FF178A3BDC78F8AE" bold="true" box="[192,312,1809,1833]" pageId="86" pageNumber="85">Figure 68:</emphasis>
<taxonomicName id="4C254D2A20041918FE958A39DFC0F8AF" authority="(Birula, 1908)" baseAuthorityName="Birula" baseAuthorityYear="1908" box="[322,640,1810,1832]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="86" pageNumber="85" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB20041918FE958A39DCACF8AF" box="[322,492,1810,1832]" italics="true" pageId="86" pageNumber="85">Leiurus hebraeus</emphasis>
(
<bibRefCitation id="EFB44B5820041918FE2B8A38DF39F8AF" author="Birula" box="[508,633,1810,1832]" firstAuthor="Birula" journalOrPublisher="Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien" pageId="86" pageNumber="85" pagination="121 - 152" part="117" refId="ref59140" refString="BIRULA, A. 1908. Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 117 2 (1): 121 - 152." title="Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae" type="journal article" year="1908">Birula, 1908</bibRefCitation>
)
</taxonomicName>
stat. n.. Metasoma.
<emphasis id="B951EABB20041918FC9A8A38DE25F8A0" bold="true" box="[845,869,1810,1831]" pageId="86" pageNumber="85">A.</emphasis>
Male, lateral aspect.
<emphasis id="B951EABB20041918FBEA8A38D914F8A0" bold="true" box="[1085,1108,1810,1831]" pageId="86" pageNumber="85">B.</emphasis>
Male, ventral aspect.
<emphasis id="B951EABB20041918FAE48A38D80BF8A0" bold="true" box="[1331,1355,1810,1831]" pageId="86" pageNumber="85">C.</emphasis>
Female, lateral aspect.
<emphasis id="B951EABB20041918FE9D8A01DC22F8C7" bold="true" box="[330,354,1835,1856]" pageId="86" pageNumber="85">D.</emphasis>
Female, ventral aspect. Male and female, Israel (NHMB 17a).
</paragraph>
</caption>
<caption id="DF5A662120051919FF178965DCA4FB1F" ID-DOI="http://doi.org/10.5281/zenodo.6505767" ID-Zenodo-Dep="6505767" httpUri="https://zenodo.org/record/6505767/files/figure.png" pageId="87" pageNumber="86" startId="87.[192,267,1103,1127]" targetBox="[191,1440,167,1073]" targetPageId="87">
<paragraph id="8B9A36A920051919FF178965DCA4FB1F" blockId="87.[192,1441,1103,1176]" pageId="87" pageNumber="86">
<emphasis id="B951EABB20051919FF178965DC77FBE0" bold="true" box="[192,311,1103,1127]" pageId="87" pageNumber="86">Figure 69:</emphasis>
<taxonomicName id="4C254D2A20051919FEE88978DF38FBE0" authority="(Birula, 1908)" baseAuthorityName="Birula" baseAuthorityYear="1908" box="[319,632,1105,1127]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="87" pageNumber="86" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB20051919FEE88978DCA7FBE0" box="[319,487,1105,1127]" italics="true" pageId="87" pageNumber="86">Leiurus hebraeus</emphasis>
(
<bibRefCitation id="EFB44B5820051919FE21897BDF31FBE0" author="Birula" box="[502,625,1105,1127]" firstAuthor="Birula" journalOrPublisher="Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien" pageId="87" pageNumber="86" pagination="121 - 152" part="117" refId="ref59140" refString="BIRULA, A. 1908. Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 117 2 (1): 121 - 152." title="Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae" type="journal article" year="1908">Birula, 1908</bibRefCitation>
)
</taxonomicName>
stat. n., female. Tarsi.
<emphasis id="B951EABB20051919FC8D897BDECBFBE1" bold="true" box="[858,907,1105,1126]" pageId="87" pageNumber="86">A-D.</emphasis>
Left telotarsus and distal basitarsus, ventral aspect.
<emphasis id="B951EABB20051919FA5D897BD8E1FBE1" bold="true" box="[1418,1441,1105,1126]" pageId="87" pageNumber="86">A.</emphasis>
Leg I.
<emphasis id="B951EABB20051919FED68940DC58FBF8" bold="true" box="[257,280,1130,1151]" pageId="87" pageNumber="86">B.</emphasis>
Leg II.
<emphasis id="B951EABB20051919FEBF8940DC3FFBF8" bold="true" box="[360,383,1130,1151]" pageId="87" pageNumber="86">C.</emphasis>
Leg III.
<emphasis id="B951EABB20051919FE0F8940DCB0FBF8" bold="true" box="[472,496,1130,1151]" pageId="87" pageNumber="86">D.</emphasis>
Leg IV.
<emphasis id="B951EABB20051919FD9D8940DF20FBF8" bold="true" box="[586,608,1130,1151]" pageId="87" pageNumber="86">E.</emphasis>
Right basitarsus III retrolateral aspect.
<emphasis id="B951EABB20051919FC358940D958FBF8" bold="true" box="[994,1048,1130,1151]" pageId="87" pageNumber="86">AD:</emphasis>
female, Israel (NHMB 17a).
<emphasis id="B951EABB20051919FAED8940D811FBF8" bold="true" box="[1338,1361,1130,1151]" pageId="87" pageNumber="86">E:</emphasis>
female, Jericho, Israel (NHMB 17I-a).
</paragraph>
</caption>
<paragraph id="8B9A36A920051919FF17886CDFB8F948" blockId="87.[192,802,1350,1743]" pageId="87" pageNumber="86">
<collectionCode id="ED34AE6C20051919FF17886CDF19FAD9" box="[192,601,1350,1374]" pageId="87" pageNumber="86">OTHER MATERIAL EXAMINED</collectionCode>
.
<materialsCitation id="3B4D3CF420051919FDB8886DDC63FA1B" ID-GBIF-Occurrence="3913646373" collectionCode="OTHER, MATERIAL, EXAMINED, USNM" collectorName="A. Hrdlicka" country="Egypt" elevation="54" latitude="29.55" location="Lisht" longLatPrecision="1226" longitude="31.133333" pageId="87" pageNumber="86" specimenCount="1">
<emphasis id="B951EABB20051919FDB8886DDFFCFAD8" bold="true" box="[623,700,1351,1375]" pageId="87" pageNumber="86">
<collectingCountry id="F332763920051919FDB8886DDFF7FAD8" box="[623,695,1351,1375]" name="Egypt" pageId="87" pageNumber="86">Egypt</collectingCountry>
:
</emphasis>
2 juvs,
<location id="8EFA607220051919FF17884FDDB9FAFA" LSID="urn:lsid:plazi:treatment:038C87BF20031912FF0E8BCADC10F9EA:8EFA607220051919FF17884FDDB9FAFA" box="[192,249,1381,1405]" country="Egypt" latitude="29.55" longLatPrecision="1226" longitude="31.133333" name="Lisht" pageId="87" pageNumber="86">Lisht</location>
,
<geoCoordinate id="EE11506E20051919FEDC884FDC22FAFA" box="[267,354,1381,1405]" degrees="29" direction="north" minutes="33" orientation="latitude" pageId="87" pageNumber="86" precision="925" value="29.55">29°33'N</geoCoordinate>
<geoCoordinate id="EE11506E20051919FEA7884FDC87FAFA" box="[368,455,1381,1405]" degrees="31" direction="east" minutes="08" orientation="longitude" pageId="87" pageNumber="86" precision="925" value="31.133333">31°08'E</geoCoordinate>
,
<elevation id="0008D19A20051919FE0E884FDF15FAFA" box="[473,597,1381,1405]" metricMagnitude="1" metricUnit="m" metricValue="5.4" pageId="87" pageNumber="86" unit="m" value="54.0">
<quantity id="4CDD9B4C20051919FE0E884FDF55FAFA" box="[473,533,1381,1405]" metricMagnitude="1" metricUnit="m" metricValue="5.4" pageId="87" pageNumber="86" unit="m" value="54.0">54 m</quantity>
a.s.l.
</elevation>
, leg.
<collectorName id="26D0537F20051919FD4A884FDE61FAFA" box="[669,801,1381,1405]" pageId="87" pageNumber="86">A. Hrdlička</collectorName>
(
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(
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(
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, leg.
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, leg.
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(
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17d);
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17f);
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, 2007 (GL).
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:
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a.s.l.
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, ca
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, leg.
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.
</paragraph>
</subSubSection>
<subSubSection id="C33F652220051916FF178BDEDFB6FA17" lastPageId="88" lastPageNumber="87" pageId="87" pageNumber="86" type="diagnosis">
<paragraph id="8B9A36A920051916FF178BDEDFB6FA17" blockId="87.[192,802,1780,1896]" lastBlockId="88.[192,802,1155,1425]" lastPageId="88" lastPageNumber="87" pageId="87" pageNumber="86">
DIAGNOSIS (adults). Medium to large
<taxonomicName id="4C254D2A20051919FDAA8BDFDF8EF88B" authorityName=": Vachon" authorityYear="1949" box="[637,718,1781,1804]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="87" pageNumber="86" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABB20051919FDAA8BDFDF8EF88B" box="[637,718,1781,1804]" italics="true" pageId="87" pageNumber="86">Leiurus</emphasis>
</taxonomicName>
,
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in length, carapace L 7.811.0 mm, color uniformly yellow or with variable fuscous markings on carapace and tergites; metasoma V yellow, or darkened except for posterior end; carapace with area between anterior median carinae lacking dark pigment, smooth or slightly roughened, with few sparse granules, area between posterior median carinae flat or with shallow median depression, lateral arcs of granules either very fine or absent; medial intercarinal surfaces of tergites IIIII densely shagreened with fine granules, granulation weaker and sparser in females (
<figureCitation id="131E2A2C20051919FB538889D9B8FA3C" box="[1156,1272,1443,1467]" captionStart="Figure 93" captionStartId="113.[192,267,1572,1596]" captionTargetBox="[224,1390,195,1538]" captionTargetId="figure-226@113.[212,1402,183,1550]" captionTargetPageId="113" captionText="Figure 93: Medial intercarinal areas of tergite III of female Leiurus spp. A. L. arabicus sp. n., paratype, Jeddah, Saudi Arabia. B. L. arabicus sp. n., paratype, Wadi Khumrah, Saudi Arabia (NHMB 17bj). C. L. haenggii sp. n., paratype, Yemen. D. L. haenggii sp. n., paratype, Wadi Maraba, Saudi Arabia (NHMB 17al). E. L. heberti sp. n., paratype, Jabal Samhan. Oman. F. L. abdullahbayrami Yağmur, Koç et Kunt, 2009, Caybasi Village, Turkey. G. L. brachycentrus (Ehrenberg, 1829) stat. n., Al Mansuriah, Yemen. H. L. macroctenus sp. n., paratype, Masirah Island, Oman. I. L. hebraeus (Birula, 1908) stat. n., Israel (NHMB 17a). J. L. hebraeus Birula, 1908, Kurayyima, Jordan. K. L. quinquestriatus (Ehrenberg, 1828), Kharga Oasis, Egypt (NHMB 17i). L. L. jordanensis Lourenço, Modry et Amr, 2002, al-Tawil, Saudi Arabia." figureDoi="http://doi.org/10.5281/zenodo.6505818" httpUri="https://zenodo.org/record/6505818/files/figure.png" pageId="87" pageNumber="86">Figs. 93K</figureCitation>
,
<figureCitation id="131E2A2C20051919FADE8889D867FA3C" box="[1289,1319,1443,1467]" captionStart="Figure 94" captionStartId="114.[192,260,765,786]" captionTargetBox="[204,1427,204,723]" captionTargetId="figure-776@114.[192,1439,192,745]" captionTargetPageId="114" captionText="Figure 94: Comparative analysis of granulation on tergite III of female Leiurus. A. UV fluorescence image of tergite III medial intercarinal surface of L. quinquestriatus (Egypt). B. Mask of granulation pattern on medial intercarinal surface from image in Figure 94A. C. Enlarged view of tergite III granulation in rectangular area indicated in Figure 94A. Granules were identified as local maxima of fluorescence intensity with contour shadows cast by directional UV illumination, and were modeled by elliptical regions-of-interest (ROIs). Parameters of granule ROIs were measured in ImageJ 1.44 (Rasband, 19972011). Area of granulometric analysis was restricted to surfaces bounded by the lateral and posterior marginal carinae, and a line passing through oblique, anterior transverse rows of enlarged granules (granules along carinae and bounding lines were omitted). D. Scatter plot of mean granule diameter vs. total granule area for 4 species of Leiurus (L. arabicus sp. n., L. haenggii sp. n., L. hebraeus stat. n. and L. quinquestriatus). Each point represents granulometric data derived from tergite III of one scorpion specimen (bilaterally, as shown in Figure 94B). Data were extracted from 18,423 ROIs from 36 scorpions. For each specimen, total granule area (a measure of density of granulation) was computed as sum of areas of all ROIs, and mean ROI diameter (a measure of coarseness of granulation) as the mean value of the maximum diameters of all granule ROIs. For comparative analysis, images from different size scorpions were resampled to equalize the distance between left and right posterior marginal granules of the lateral carinae (arbitrarily set to 4,000 units or pixels; linear dimensions expressed as [pixel], areas as [pixel2]). The total granule area separated the species into 2 groups: i.e. sparsely granulated (L. arabicus, L. haenggii) and densely granulated (L. hebraeus, L. quinquestriatus). In contrast, the distributions of ROI diameter were broadly overlapping. E. To obtain a more sensitive comparison of the coarseness of granulation, normalized cumulative distributions of single ROI areas for 3 species of Leiurus were computed (inset indicates number of scorpions analyzed). Relative horizontal positions of these curves indicated increasing coarseness of granulation, in rank order: L. haenggii &lt;L. quinquestriatus &lt;L. hebraeus (Ngranules = 2103, 5202, 2190 respectively). This ranking was confirmed by a Kolmogorov-Smirnov test which detected significant differences between the distributions of log (granule area) (P &lt;0.001)." figureDoi="http://doi.org/10.5281/zenodo.6505820" httpUri="https://zenodo.org/record/6505820/files/figure.png" pageId="87" pageNumber="86">94</figureCitation>
); posterior margin of coxa III smooth, or with scattered fine granules; metasoma moderately slender, metasoma II L/
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1.96, metasoma III L/
<geoCoordinate id="EE11506E20051919FB4C88D4D9AAF991" box="[1179,1258,1534,1558]" degrees="1.67" direction="west" orientation="longitude" pageId="87" pageNumber="86" precision="555" value="-1.67">W 1.67</geoCoordinate>
2.22, metasoma IV L/
<geoCoordinate id="EE11506E20051919FCAE8B37DE88F9B2" box="[889,968,1565,1589]" degrees="2.07" direction="west" orientation="longitude" pageId="87" pageNumber="86" precision="555" value="-2.07">W 2.07</geoCoordinate>
2.63; ventromedian carinae of metasoma II and III with 1926 denticles (90/92 carinae); metasoma V with enlarged subtriangular or lobate denticles on ventrolateral carinae; pedipalps slender, patella L/W
<specimenCount id="9D23FD2020051919FA5B8B50D8E0F916" box="[1420,1440,1658,1681]" pageId="87" pageNumber="86" type="male"></specimenCount>
3.173.85,
<specimenCount id="9D23FD2020051919FC6C8BB3DE8FF937" box="[955,975,1689,1712]" pageId="87" pageNumber="86" type="female"></specimenCount>
2.883.61; leg III patella L/D 3.244.38; pectine teeth
<specimenCount id="9D23FD2020051919FC198B9DDEA2F949" box="[974,994,1719,1742]" pageId="87" pageNumber="86" type="male"></specimenCount>
3039,
<specimenCount id="9D23FD2020051919FBEE8B9DD90DF949" box="[1081,1101,1719,1742]" pageId="87" pageNumber="86" type="female"></specimenCount>
2638; pectines medium sized, pectine L/ carapace L
<specimenCount id="9D23FD2020051919FBEE8BFCD90DF96A" box="[1081,1101,1750,1773]" pageId="87" pageNumber="86" type="male"></specimenCount>
1.111.40,
<specimenCount id="9D23FD2020051919FB038BFCD9A8F96A" box="[1236,1256,1750,1773]" pageId="87" pageNumber="86" type="female"></specimenCount>
0.891.38, midpectine sensillar margin L/ metasoma I W
<specimenCount id="9D23FD2020051919FAE28BDFD809F88B" box="[1333,1353,1781,1804]" pageId="87" pageNumber="86" type="male"></specimenCount>
<geoCoordinate id="EE11506E20051919FA808BDED8D3F88B" box="[1367,1427,1780,1804]" degrees="0.142" direction="north" orientation="latitude" pageId="87" pageNumber="86" precision="55" value="0.142">0.142</geoCoordinate>
<geoCoordinate id="EE11506E20051919FA448BDFDE3EF8AC" degrees="0.184" direction="west" orientation="longitude" pageId="87" pageNumber="86" precision="55" value="-0.184"> 0.184</geoCoordinate>
,
<specimenCount id="9D23FD2020051919FC5C8A39DEDFF8AD" box="[907,927,1811,1834]" pageId="87" pageNumber="86" type="female"></specimenCount>
<geoCoordinate id="EE11506E20051919FC7F8A39DEA4F8AC" box="[936,996,1811,1835]" degrees="0.078" direction="north" orientation="latitude" pageId="87" pageNumber="86" precision="55" value="0.078">0.078</geoCoordinate>
<geoCoordinate id="EE11506E20051919FC338A39D970F8AC" box="[996,1072,1811,1835]" degrees="0.156" direction="west" orientation="longitude" pageId="87" pageNumber="86" precision="55" value="-0.156">0.156</geoCoordinate>
; basal 13 pectine teeth of males overlap if anterior pectine margins aligned to posterior margins of coxae IV; pectine basal piece densely, shagreened, usually with more coarse medial granules in males; leg III basitarsus with 816 retrosuperior setae; pedipalp chela fixed finger with trichobothrium usually
<emphasis id="B951EABB200A1916FF1789F5DD9BFB70" box="[192,219,1247,1271]" italics="true" pageId="88" pageNumber="87">db</emphasis>
distal to
<emphasis id="B951EABB200A1916FE8689CADC2EFB70" box="[337,366,1248,1271]" italics="true" pageId="88" pageNumber="87">est</emphasis>
; area of sternite VII between median carinae with dense, fine granulation or shagreened anteriorly, more heavily in males; sternite carination:
<emphasis id="B951EABB200A1916FF178811DCD7FAD4" box="[192,407,1339,1363]" italics="true" pageId="88" pageNumber="87">males and females</emphasis>
: sternite III with median carinae strong, densely granular or shagreened, sternites IVV with lateral carinae strong, median carinae moderate.
</paragraph>
</subSubSection>
<caption id="DF5A6621200A1916FF178935DF64FBC8" ID-DOI="http://doi.org/10.5281/zenodo.6505769" ID-Zenodo-Dep="6505769" httpUri="https://zenodo.org/record/6505769/files/figure.png" pageId="88" pageNumber="87" startId="88.[192,267,1055,1079]" targetBox="[191,1440,191,1025]" targetPageId="88">
<paragraph id="8B9A36A9200A1916FF178935DF64FBC8" blockId="88.[192,1441,1055,1103]" pageId="88" pageNumber="87">
<emphasis id="B951EABB200A1916FF178935DC76FBB0" bold="true" box="[192,310,1055,1079]" pageId="88" pageNumber="87">Figure 70:</emphasis>
<taxonomicName id="4C254D2A200A1916FEEA8908DF33FBB0" authority="(Birula, 1908)" baseAuthorityName="Birula" baseAuthorityYear="1908" box="[317,627,1057,1079]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="88" pageNumber="87" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB200A1916FEEA8908DCA4FBB0" box="[317,484,1057,1079]" italics="true" pageId="88" pageNumber="87">Leiurus hebraeus</emphasis>
(
<bibRefCitation id="EFB44B58200A1916FE25890BDF2CFBB0" author="Birula" box="[498,620,1057,1079]" firstAuthor="Birula" journalOrPublisher="Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien" pageId="88" pageNumber="87" pagination="121 - 152" part="117" refId="ref59140" refString="BIRULA, A. 1908. Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 117 2 (1): 121 - 152." title="Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae" type="journal article" year="1908">Birula, 1908</bibRefCitation>
)
</taxonomicName>
stat. n., lectotype female, designated here (ZISP 578). Habitus.
<emphasis id="B951EABB200A1916FB09890BD9AFFBB1" bold="true" box="[1246,1263,1057,1078]" pageId="88" pageNumber="87">A</emphasis>
. Dorsal aspect.
<emphasis id="B951EABB200A1916FA5D890BD8DAFBB1" bold="true" box="[1418,1434,1057,1078]" pageId="88" pageNumber="87">B</emphasis>
. Ventral aspect. Wadi Arrud, Jordan.
</paragraph>
</caption>
<subSubSection id="C33F6522200A1917FF17889CDCC7F898" lastPageId="89" lastPageNumber="88" pageId="88" pageNumber="87" type="description">
<paragraph id="8B9A36A9200A1916FF17889CDCE1FA49" blockId="88.[192,802,1462,1884]" box="[192,417,1462,1486]" pageId="88" pageNumber="87">MEASUREMENTS.</paragraph>
<paragraph id="8B9A36A9200A1916FF2788FFD902FB3D" blockId="88.[192,802,1462,1884]" lastBlockId="88.[831,1440,1155,1884]" pageId="88" pageNumber="87">
<emphasis id="B951EABB200A1916FF2788FFDF87FA6A" bold="true" box="[240,711,1493,1517]" pageId="88" pageNumber="87">
Male
<typeStatus id="549E880B200A1916FEE488FFDCCAFA6A" box="[307,394,1493,1517]" pageId="88" pageNumber="87" type="syntype">syntype</typeStatus>
(
<figureCitation id="131E2A2C200A1916FE4D88FFDCBFFA6A" box="[410,511,1493,1517]" captionStart="Figure 82" captionStartId="100.[192,267,1070,1094]" captionTargetBox="[207,1412,188,1044]" captionTargetId="figure-431@100.[192,1439,168,1065]" captionTargetPageId="100" captionText="Figure 82: Leiurus quinquestriatus (Ehrenberg, 1828), 5 of 8 syntypes (ZMHB No. 140). Females: A, #12C01-3; B, #12C04-6. Males: C, #12C07-9; D, #12C10-3. Juvenile: E, #12C14-6. Egypt; Sudan. Scale bars (10 mm) apply to the habitus photos. Body lengths: A, 90 mm; B, 88 mm; C, 63 mm; D, 69 mm; E, 65 mm. Not shown are three additional fragmented juvenile syntypes." figureDoi="http://doi.org/10.5281/zenodo.6505796" httpUri="https://zenodo.org/record/6505796/files/figure.png" pageId="88" pageNumber="87">Fig. 82C</figureCitation>
, subadult) (mm).
</emphasis>
<collectorName id="26D0537F200A1916FD0788FFDE62FA6B" box="[720,802,1493,1517]" pageId="88" pageNumber="87">Total L</collectorName>
63.0; metasoma + telson L 41.5; carapace L 6.6,
<geoCoordinate id="EE11506E200A1916FD0088DEDE5DF98C" box="[727,797,1523,1547]" degrees="7.8" direction="west" orientation="longitude" pageId="88" pageNumber="87" precision="5555" value="-7.8">W 7.8</geoCoordinate>
; metasomal segments (L/ W) I 5.4/ 4.2,
<specimenCode id="DB839ED2200A1916FDB18B38DFDFF9AD" box="[614,671,1554,1578]" collectionCode="II" pageId="88" pageNumber="87">II 6.3</specimenCode>
/ 3.6,
<specimenCode id="DB839ED2200A1916FD098B38DE61F9AD" box="[734,801,1554,1578]" collectionCode="III" pageId="88" pageNumber="87">III 6.3</specimenCode>
/ 3.5, IV 7.3 / 3.2,
<specimenCode id="DB839ED2200A1916FE4A8B1BDC9CF9CF" box="[413,476,1584,1608]" collectionCode="V" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="88" pageNumber="87" type="Museum">V 8.6</specimenCode>
/ 3.0; telson L 7.6; pedipalp chela L 13.1, manus
<geoCoordinate id="EE11506E200A1916FE688B7ADF4CF9E0" box="[447,524,1615,1639]" degrees="1.9" direction="west" orientation="longitude" pageId="88" pageNumber="87" precision="5555" value="-1.9">W 1.9</geoCoordinate>
, movable finger L 8.9; pedipalp femur L 7.2,
<geoCoordinate id="EE11506E200A1916FE1B8B45DF56F901" box="[460,534,1646,1670]" degrees="1.7" direction="west" orientation="longitude" pageId="88" pageNumber="87" precision="5555" value="-1.7">W 1.7</geoCoordinate>
, patella L 8.0,
<geoCoordinate id="EE11506E200A1916FD068B45DE5DF901" box="[721,797,1646,1670]" degrees="2.2" direction="west" orientation="longitude" pageId="88" pageNumber="87" precision="5555" value="-2.2">W 2.2</geoCoordinate>
; pectine L 8.5, mid-pectine sensillar margin L 0.85; pectine teeth 3030.
<emphasis id="B951EABB200A1916FE7E8B81DC44F965" bold="true" pageId="88" pageNumber="87">
Adult female
<typeStatus id="549E880B200A1916FD9D8B81DFE1F944" box="[586,673,1707,1731]" pageId="88" pageNumber="87" type="syntype">syntype</typeStatus>
(
<figureCitation id="131E2A2C200A1916FD648B81DE5AF944" box="[691,794,1707,1731]" captionStart="Figure 82" captionStartId="100.[192,267,1070,1094]" captionTargetBox="[207,1412,188,1044]" captionTargetId="figure-431@100.[192,1439,168,1065]" captionTargetPageId="100" captionText="Figure 82: Leiurus quinquestriatus (Ehrenberg, 1828), 5 of 8 syntypes (ZMHB No. 140). Females: A, #12C01-3; B, #12C04-6. Males: C, #12C07-9; D, #12C10-3. Juvenile: E, #12C14-6. Egypt; Sudan. Scale bars (10 mm) apply to the habitus photos. Body lengths: A, 90 mm; B, 88 mm; C, 63 mm; D, 69 mm; E, 65 mm. Not shown are three additional fragmented juvenile syntypes." figureDoi="http://doi.org/10.5281/zenodo.6505796" httpUri="https://zenodo.org/record/6505796/files/figure.png" pageId="88" pageNumber="87">Fig. 82A</figureCitation>
) (mm).
</emphasis>
<collectorName id="26D0537F200A1916FEDB8BE0DC1BF966" box="[268,347,1738,1762]" pageId="88" pageNumber="87">Total L</collectorName>
90.0; metasoma + telson L 56.7; carapace L 10.3,
<geoCoordinate id="EE11506E200A1916FECF8BC3DC2CF887" box="[280,364,1768,1792]" degrees="11.6" direction="west" orientation="longitude" pageId="88" pageNumber="87" precision="5555" value="-11.6">W 11.6</geoCoordinate>
; metasomal segments (L/ W) I 7.4/ 5.8,
<specimenCode id="DB839ED2200A1916FF178A22DDB9F898" box="[192,249,1799,1823]" collectionCode="II" pageId="88" pageNumber="87">II 8.5</specimenCode>
/ 5.3,
<specimenCode id="DB839ED2200A1916FEE08A22DC3AF898" box="[311,378,1799,1823]" collectionCode="III" pageId="88" pageNumber="87">III 8.6</specimenCode>
/ 5.2, IV 10.1 / 4.7,
<specimenCode id="DB839ED2200A1916FD838A22DFDDF898" box="[596,669,1799,1823]" collectionCode="V" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="88" pageNumber="87" type="Museum">V 11.4</specimenCode>
/ 4.4; telson L 10.7; pedipalp chela L 18.7, manus
<geoCoordinate id="EE11506E200A1916FDB88A0DDFF7F8B9" box="[623,695,1830,1854]" degrees="3.3" direction="west" orientation="longitude" pageId="88" pageNumber="87" precision="5555" value="-3.3">W 3.3</geoCoordinate>
, movable finger L 13.5; pedipalp femur L 10.1,
<geoCoordinate id="EE11506E200A1916FDBB8A6FDFF2F8DB" box="[620,690,1860,1884]" degrees="2.8" direction="west" orientation="longitude" pageId="88" pageNumber="87" precision="5555" value="-2.8">W 2.8</geoCoordinate>
, patella L 10.8,
<geoCoordinate id="EE11506E200A1916FCAB89AEDE81FB1C" box="[892,961,1155,1179]" degrees="3.5" direction="west" orientation="longitude" pageId="88" pageNumber="87" precision="5555" value="-3.5">W 3.5</geoCoordinate>
; pectine L 9.7, mid-pectine sensillar margin L 0.90; pectine teeth 30.
</paragraph>
<paragraph id="8B9A36A9200A1916FCB889EBD945F9CF" blockId="88.[831,1440,1155,1884]" pageId="88" pageNumber="87">
<emphasis id="B951EABB200A1916FCB889EBD800FB5E" bold="true" box="[879,1344,1217,1241]" pageId="88" pageNumber="87">
<collectingRegion id="49E1F84B200A1916FCB889EBDEEAFB5E" box="[879,938,1217,1241]" country="Egypt" name="Male" pageId="88" pageNumber="87">Male</collectingRegion>
,
<collectingRegion id="49E1F84B200A1916FC6D89EBDEB8FB5E" box="[954,1016,1217,1241]" country="Egypt" name="Qina" pageId="88" pageNumber="87">Kena</collectingRegion>
,
<collectingCountry id="F3327639200A1916FBDE89EBD90DFB5E" box="[1033,1101,1217,1241]" name="Egypt" pageId="88" pageNumber="87">Egypt</collectingCountry>
(NHMB
<quantity id="4CDD9B4C200A1916FB6889EBD9A9FB5E" box="[1215,1257,1217,1241]" metricMagnitude="-2" metricUnit="kg" metricValue="1.7" pageId="88" pageNumber="87" unit="g" value="17.0">17g</quantity>
) (mm).
</emphasis>
<collectorName id="26D0537F200A1916FA9B89EAD8E0FB5F" box="[1356,1440,1216,1240]" pageId="88" pageNumber="87">Total L</collectorName>
74.00; metasoma + telson L 48.00; carapace L 8.28,
<geoCoordinate id="EE11506E200A1916FA5189CADE30FA91" degrees="8.57" direction="west" orientation="longitude" pageId="88" pageNumber="87" precision="555" value="-8.57">W 8.57</geoCoordinate>
, carapace preocular L 3.69; metasomal segments (L/ D /W) I 6.31/ 4.30/5.15,
<specimenCode id="DB839ED2200A1916FBAE8837D982FAB2" box="[1145,1218,1309,1333]" collectionCode="II" pageId="88" pageNumber="87">II 7.50</specimenCode>
/ 4.57/4.55,
<specimenCode id="DB839ED2200A1916FA918837D8D6FAB2" box="[1350,1430,1309,1333]" collectionCode="III" pageId="88" pageNumber="87">III 7.71</specimenCode>
/ 3.78/4.28, IV 8.62/ 3.41/3.87,
<specimenCode id="DB839ED2200A1916FB598816D998FAD4" box="[1166,1240,1339,1363]" collectionCode="V" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="88" pageNumber="87" type="Museum">V 9.28</specimenCode>
/ 3.22/3.68; telson L 7.62; vesicle L 3.76, D 3.04,
<geoCoordinate id="EE11506E200A1916FB588870D9A2FAF5" box="[1167,1250,1370,1394]" degrees="3.04" direction="west" orientation="longitude" pageId="88" pageNumber="87" precision="555" value="-3.04">W 3.04</geoCoordinate>
; pedipalp chela L 15.60, manus ventral L 4.49, manus
<geoCoordinate id="EE11506E200A1916FB0E8853D86EFA17" box="[1241,1326,1400,1424]" degrees="2.35" direction="west" orientation="longitude" pageId="88" pageNumber="87" precision="555" value="-2.35">W 2.35</geoCoordinate>
, manus D 2.57, fixed finger L 9.90, movable finger L 11.40; pedipalp femur L 8.22,
<geoCoordinate id="EE11506E200A1916FB95889DD9D5FA49" box="[1090,1173,1462,1486]" degrees="2.31" direction="west" orientation="longitude" pageId="88" pageNumber="87" precision="555" value="-2.31">W 2.31</geoCoordinate>
, patella L 9.50,
<geoCoordinate id="EE11506E200A1916FA9F889DD8DAFA49" box="[1352,1434,1462,1486]" degrees="2.81" direction="west" orientation="longitude" pageId="88" pageNumber="87" precision="555" value="-2.81">W 2.81</geoCoordinate>
; pectine L 9.98, mid-pectine sensillar margin L 0.898; leg III femur L 8.77; leg III patella L 7.59, D 2.00; chela
<emphasis id="B951EABB200A1916FCE88B38DEC5F9AD" box="[831,901,1554,1578]" italics="true" pageId="88" pageNumber="87">dbest</emphasis>
distance (left/ right)
<geoCoordinate id="EE11506E200A1916FBA08B38D9F3F9AD" box="[1143,1203,1554,1578]" degrees="0.958" direction="north" orientation="latitude" pageId="88" pageNumber="87" precision="55" value="0.958">0.958</geoCoordinate>
/
<geoCoordinate id="EE11506E200A1916FB118B38D845F9AD" box="[1222,1285,1554,1578]" degrees="0.917" direction="east" orientation="longitude" pageId="88" pageNumber="87" precision="55" value="0.917">0.917</geoCoordinate>
; pectine teeth (left/ right) 35/ 35.
</paragraph>
<paragraph id="8B9A36A9200A1917FCB88B7ADCC7F898" blockId="88.[831,1440,1155,1884]" lastBlockId="89.[192,802,1707,1823]" lastPageId="89" lastPageNumber="88" pageId="88" pageNumber="87">
<emphasis id="B951EABB200A1916FCB88B7AD807F9EF" bold="true" box="[879,1351,1616,1640]" pageId="88" pageNumber="87">
<collectingRegion id="49E1F84B200A1916FCB88B7ADE83F9EF" box="[879,963,1616,1640]" country="Egypt" name="Female" pageId="88" pageNumber="87">Female</collectingRegion>
,
<collectingRegion id="49E1F84B200A1916FC188B7AD94DF9EF" box="[975,1037,1616,1640]" country="Egypt" name="Qina" pageId="88" pageNumber="87">Kena</collectingRegion>
,
<collectingCountry id="F3327639200A1916FBCC8B7AD91FF9EF" box="[1051,1119,1616,1640]" name="Egypt" pageId="88" pageNumber="87">Egypt</collectingCountry>
(NHMB
<quantity id="4CDD9B4C200A1916FB1D8B7AD9B4F9EF" box="[1226,1268,1616,1640]" metricMagnitude="-2" metricUnit="kg" metricValue="1.7" pageId="88" pageNumber="87" unit="g" value="17.0">17g</quantity>
) (mm).
</emphasis>
<collectorName id="26D0537F200A1916FA878B65D8E0F9E0" box="[1360,1440,1615,1639]" pageId="88" pageNumber="87">Total L</collectorName>
80.00; metasoma + telson L 53.00; carapace L 9.46,
<geoCoordinate id="EE11506E200A1916FA518B45DE3EF923" degrees="10.80" direction="west" orientation="longitude" pageId="88" pageNumber="87" precision="555" value="-10.8">W 10.80</geoCoordinate>
, carapace preocular L 4.44; metasomal segments (L/ D /W) I 6.62/ 5.18/5.75,
<specimenCode id="DB839ED2200A1916FBAE8B86D982F944" box="[1145,1218,1707,1731]" collectionCode="II" pageId="88" pageNumber="87">II 7.96</specimenCode>
/ 4.64/5.04,
<specimenCode id="DB839ED2200A1916FA918B86D8D8F944" box="[1350,1432,1707,1731]" collectionCode="III" pageId="88" pageNumber="87">III 8.16</specimenCode>
/ 4.49/4.79, IV 8.99/ 3.93/4.35,
<specimenCode id="DB839ED2200A1916FB518BE0D99CF965" box="[1158,1244,1738,1762]" collectionCode="V" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="88" pageNumber="87" type="Museum">V 10.11</specimenCode>
/ 3.59/4.34; telson L 9.33; vesicle L 4.63, D 3.51,
<geoCoordinate id="EE11506E200A1916FB588BC3D9A2F887" box="[1167,1250,1768,1792]" degrees="3.62" direction="west" orientation="longitude" pageId="88" pageNumber="87" precision="555" value="-3.62">W 3.62</geoCoordinate>
; pedipalp chela L 17.03, manus ventral L 5.14, manus
<geoCoordinate id="EE11506E200A1916FB0E8A22D86EF898" box="[1241,1326,1799,1823]" degrees="2.63" direction="west" orientation="longitude" pageId="88" pageNumber="87" precision="555" value="-2.63">W 2.63</geoCoordinate>
, manus D 3.14, fixed finger L 9.99, movable finger L 12.14; pedipalp femur L 8.88,
<geoCoordinate id="EE11506E200A1916FBED8A6FD9CBF8DB" box="[1082,1163,1860,1884]" degrees="2.75" direction="west" orientation="longitude" pageId="88" pageNumber="87" precision="555" value="-2.75">W 2.75</geoCoordinate>
, patella L 10.12,
<geoCoordinate id="EE11506E200A1916FA928A6FD8D6F8DB" box="[1349,1430,1860,1884]" degrees="3.44" direction="west" orientation="longitude" pageId="88" pageNumber="87" precision="555" value="-3.44">W 3.44</geoCoordinate>
; pectine L 9.53, mid-pectine sensillar margin L 0.653; leg III femur L 9.71; leg III patella L 8.29, D 2.17; chela
<emphasis id="B951EABB200B1917FF178BC2DC46F887" box="[192,262,1768,1792]" italics="true" pageId="89" pageNumber="88">dbest</emphasis>
distance (left/ right)
<geoCoordinate id="EE11506E200B1917FE2F8BC2DF74F887" box="[504,564,1768,1792]" degrees="0.917" direction="north" orientation="latitude" pageId="89" pageNumber="88" precision="55" value="0.917">0.917</geoCoordinate>
/
<geoCoordinate id="EE11506E200B1917FD908BC2DFC6F887" box="[583,646,1768,1792]" degrees="1.125" direction="east" orientation="longitude" pageId="89" pageNumber="88" precision="55" value="1.125">1.125</geoCoordinate>
; pectine teeth (left/ right) 28/ 28.
</paragraph>
</subSubSection>
<caption id="DF5A6621200B1917FF178B14DCD0F901" ID-DOI="http://doi.org/10.5281/zenodo.6505771" ID-Zenodo-Dep="6505771" httpUri="https://zenodo.org/record/6505771/files/figure.png" pageId="89" pageNumber="88" startId="89.[192,267,1598,1622]" targetBox="[191,1439,191,1567]" targetPageId="89">
<paragraph id="8B9A36A9200B1917FF178B14DCD0F901" blockId="89.[192,1440,1598,1670]" pageId="89" pageNumber="88">
<emphasis id="B951EABB200B1917FF178B14DC77F9D1" bold="true" box="[192,311,1598,1622]" pageId="89" pageNumber="88">Figure 71:</emphasis>
<taxonomicName id="4C254D2A200B1917FEEA8B6ADF33F9D2" authority="(Birula, 1908)" baseAuthorityName="Birula" baseAuthorityYear="1908" box="[317,627,1599,1621]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="89" pageNumber="88" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="B951EABB200B1917FEEA8B6ADCA4F9D2" box="[317,484,1599,1621]" italics="true" pageId="89" pageNumber="88">Leiurus hebraeus</emphasis>
(
<bibRefCitation id="EFB44B58200B1917FE268B15DF2BF9D2" author="Birula" box="[497,619,1599,1621]" firstAuthor="Birula" journalOrPublisher="Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien" pageId="89" pageNumber="88" pagination="121 - 152" part="117" refId="ref59140" refString="BIRULA, A. 1908. Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 117 2 (1): 121 - 152." title="Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae" type="journal article" year="1908">Birula, 1908</bibRefCitation>
)
</taxonomicName>
stat. n., lectotype female and four paralectotypes, designated here (ZISP 578). Wadi Arrud, Jordan. Label text in Russian: “Det. A. Birula, 5 ex. Coll. 25.IV.1897, Petro-Arabia, Wadi-Arrud, Southern shore of the Dead Sea, Davydov.”
</paragraph>
</caption>
<subSubSection id="C33F6522200B1917FCE88B81D9DDF944" box="[831,1181,1707,1731]" pageId="89" pageNumber="88" type="distribution">
<paragraph id="8B9A36A9200B1917FCE88B81D9DDF944" blockId="89.[831,1181,1707,1731]" box="[831,1181,1707,1731]" pageId="89" pageNumber="88">
DISTRIBUTION.
<collectingCountry id="F3327639200B1917FBD08B81D909F944" box="[1031,1097,1707,1731]" name="Egypt" pageId="89" pageNumber="88">Egypt</collectingCountry>
,
<collectingCountry id="F3327639200B1917FB848B81D9D9F944" box="[1107,1177,1707,1731]" name="Sudan" pageId="89" pageNumber="88">Sudan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F6522200B1912FCE88BC2DC10F9EA" lastPageId="92" lastPageNumber="91" pageId="89" pageNumber="88" type="discussion">
<paragraph id="8B9A36A9200B1912FCE88BC2DC10F9EA" blockId="89.[831,1440,1768,1823]" lastBlockId="92.[192,802,1411,1645]" lastPageId="92" lastPageNumber="91" pageId="89" pageNumber="88">
REMARKS. The
<typeStatus id="549E880B200B1917FBC58BC3D92FF886" box="[1042,1135,1769,1793]" pageId="89" pageNumber="88" type="syntype">syntypes</typeStatus>
include assorted material from the
<collectingRegion id="49E1F84B200B1917FC768A2DDE98F898" box="[929,984,1799,1823]" country="Egypt" name="Shamal Sina'" pageId="89" pageNumber="88">Sinai</collectingRegion>
, the
<location id="8EFA6072200B1917FBC48A2DD9CFF898" LSID="urn:lsid:plazi:treatment:038C87BF20031912FF0E8BCADC10F9EA:8EFA6072200B1917FBC48A2DD9CFF898" box="[1043,1167,1799,1823]" country="Egypt" municipality="Nile Valley" name="Nile Valley" pageId="89" pageNumber="88" stateProvince="Shamal Sina'">Nile Valley</location>
in
<collectingCountry id="F3327639200B1917FB668A2DD9B2F898" box="[1201,1266,1799,1823]" name="Egypt" pageId="89" pageNumber="88">Egypt</collectingCountry>
and
<collectingCountry id="F3327639200B1917FAF08A2DD82CF898" box="[1319,1388,1799,1823]" name="Sudan" pageId="89" pageNumber="88">Sudan</collectingCountry>
, and the desert region of
<collectingCountry id="F3327639200E1912FE4A88A9DC9DFA1C" box="[413,477,1411,1435]" name="Egypt" pageId="92" pageNumber="91">Egypt</collectingCountry>
east of the Nile. These could represent more than one species if the populations in the
<collectingRegion id="49E1F84B200E1912FF178894DDB7FA51" box="[192,247,1470,1494]" country="Egypt" name="Shamal Sina'" pageId="92" pageNumber="91">Sinai</collectingRegion>
are distinct from those of the
<collectingMunicipality id="6BFEACD3200E1912FD458894DE5DFA51" box="[658,797,1470,1494]" pageId="92" pageNumber="91">Nile Valley</collectingMunicipality>
. However, since individual specimens do not have locality labels, they cannot be referred to specific populations. We refrain from designating a
<typeStatus id="549E880B200E1912FD548B33DFA4F9B6" box="[643,740,1561,1585]" pageId="92" pageNumber="91" type="lectotype">lectotype</typeStatus>
until the taxonomic status of these populations is clari-fied by further study.
</paragraph>
</subSubSection>
</treatment>
</document>