370 lines
68 KiB
XML
370 lines
68 KiB
XML
<document id="2A9D95EC4F0D6A65F9AD5E8F5988DDF8" ID-DOI="10.11646/zootaxa.4803.2.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="3909195" ID-ZooBank="32379908-BED1-40A3-B77D-D43A8BD5FA33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1593158588459" checkinUser="plazi" docAuthor="Franzese, Sebastian & Ivanov, Veronica A." docDate="2020" docId="038C87BEDB5D7F4FFF735FDDCBB9FCA1" docLanguage="en" docName="zootaxa.4803.2.7.pdf" docOrigin="Zootaxa 4803 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Semiorbiseptum alfredoi Franzese & Ivanov 2020, sp. nov." docType="treatment" docUuid="685A2F80-698A-4420-98CC-62A8804F21A9" docUuidSource="ZooBank" docVersion="3" lastPageNumber="367" masterDocId="FFB5FFC6DB557F43FFE45E00C875FFB6" masterDocTitle="A new genus of Rhinebothriidea from species of Psammobatis (Rajiformes: Arhynchobatidae) off Argentina" masterLastPageNumber="372" masterPageNumber="355" pageNumber="363" updateTime="1700277165797" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="6F41D6F79503C871EE77BA370716AD17" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="9EB346818365E2D1EDA6831B8E0877AD">
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<mods:title id="7C5885FD0D0EFB156246FBC3E0565859">A new genus of Rhinebothriidea from species of Psammobatis (Rajiformes: Arhynchobatidae) off Argentina</mods:title>
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</mods:titleInfo>
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<mods:name id="EE569934612CC894E2D10A356BA2A5AB" type="personal">
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<mods:role id="1B903621A859117624ACD05DB8CE4955">
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<mods:roleTerm id="50619CB6F0CA889BA49F3D26D6C748D3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9892E8FA8A43CEFF27D4E77BDB456DDF">Franzese, Sebastian</mods:namePart>
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<mods:affiliation id="9934D961A8D9F0705929BF8C7EC6CA0F">Universidad de Buenos Aires, Facultad de Ciencias Exactas y Naturales, Departamento de Biodiversidad y Biología Experimental, Buenos Aires, Argentina. & CONICET-Universidad de Buenos Aires, Instituto de Biodiversidad y Biología Experimental y Aplicada (IBBEA), Buenos Aires, Argentina.</mods:affiliation>
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</mods:name>
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<mods:name id="195BB25243C914B2AEF886109AFF0673" type="personal">
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<mods:role id="D4D63F35CBDE6767759184631D840D14">
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<mods:roleTerm id="A87BF09283316A5E530DB3AED27E9FA1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E38FBDFEF635FF15D9365C85458ADBBA">Ivanov, Veronica A.</mods:namePart>
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<mods:affiliation id="8E5BC1ED5467759CC2B03BA0D92F8291">Universidad de Buenos Aires, Facultad de Ciencias Exactas y Naturales, Departamento de Biodiversidad y Biología Experimental, Buenos Aires, Argentina. & CONICET-Universidad de Buenos Aires, Instituto de Biodiversidad y Biología Experimental y Aplicada (IBBEA), Buenos Aires, Argentina.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="C2137CB32D11D81D489B639D1F8516E9">text</mods:typeOfResource>
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<mods:relatedItem id="C2814B3262EF7BB0965466F770F08998" type="host">
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<mods:titleInfo id="662AE50CFFAD9858C11CD091B81B3A1A">
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<mods:title id="7DDEA55CEE8A4B32D1B8948F6C8085E1">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="58DFF8438B9CEF495E65EE59BF34FE65">
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<mods:date id="371C86D063752A8EEBA013FE0FD1A32A">2020</mods:date>
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<mods:detail id="772DA211753A65E7CAAC4251D99B2A31" type="pubDate">
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<mods:number id="91F5FD52C750A85974D0E8816B457699">2020-06-26</mods:number>
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</mods:detail>
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<mods:detail id="823BFEDC2547F6BC72B124FDE605DA67" type="volume">
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<mods:number id="0A86CDB06BA56A7DB74F58DC32C0A5A1">4803</mods:number>
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</mods:detail>
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<mods:detail id="6810CA1BDCA32B5CF67E523CD685D24C" type="issue">
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<mods:number id="65AC4E1A971B5932E6E97125D1AF51F8">2</mods:number>
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</mods:detail>
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<mods:start id="2F65D0351645C2CFBBF9DB8DBCE7B7A9">355</mods:start>
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<mods:end id="A4934CD1B6C1EF5B67FFFA4AFC139ABF">372</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="D035EFDFE51955749175B51B436AAA34">journal article</mods:classification>
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<mods:identifier id="517A8ECE0F7134A113CE9BFA8C4F1FE8" type="DOI">10.11646/zootaxa.4803.2.7</mods:identifier>
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<mods:identifier id="02875BA762090B7B5F3F43535F1ADDC5" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="32AC1A7A6DBFB9ACB2818A9E0897739F" type="Zenodo-Dep">3909195</mods:identifier>
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<mods:identifier id="B121EBF7BBC367CA29BAAFE1463F297D" type="ZooBank">32379908-BED1-40A3-B77D-D43A8BD5FA33</mods:identifier>
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</mods:mods>
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<treatment id="038C87BEDB5D7F4FFF735FDDCBB9FCA1" LSID="urn:lsid:zoobank.org:act:685A2F80-698A-4420-98CC-62A8804F21A9" httpUri="http://treatment.plazi.org/id/038C87BEDB5D7F4FFF735FDDCBB9FCA1" lastPageId="12" lastPageNumber="367" pageId="8" pageNumber="363">
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<subSubSection id="C33F6523DB5D7F4BFF735FDDCA53FE41" box="[151,550,477,504]" pageId="8" pageNumber="363" type="nomenclature">
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<paragraph id="8B9A36A8DB5D7F4BFF735FDDCA53FE41" blockId="8.[151,550,477,540]" box="[151,550,477,504]" pageId="8" pageNumber="363">
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<heading id="D0D281C4DB5D7F4BFF735FDDCA53FE41" bold="true" box="[151,550,477,504]" fontSize="11" level="1" pageId="8" pageNumber="363" reason="1">
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<emphasis id="B951EABADB5D7F4BFF735FDDCA53FE41" bold="true" box="[151,550,477,504]" pageId="8" pageNumber="363">
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<taxonomicName id="4C254D2BDB5D7F4BFF735FDDC9B4FE4E" authority="Franzese & Ivanov, 2020" authorityName="Franzese & Ivanov" authorityYear="2020" box="[151,449,477,504]" class="Chondrichthyes" family="Arhynchobatidae" genus="Semiorbiseptum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="363" phylum="Chordata" rank="species" species="alfredoi" status="sp. nov.">
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||
<emphasis id="B951EABADB5D7F4BFF735FDDC9B4FE4E" bold="true" box="[151,449,477,504]" italics="true" pageId="8" pageNumber="363">Semiorbiseptum alfredoi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26257C1DB5D7F4BFE2C5FDDCA53FE41" box="[456,550,477,503]" pageId="8" pageNumber="363" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
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||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33F6523DB5D7F4BFF735C00C967FDAA" box="[151,274,512,540]" pageId="8" pageNumber="363" type="description">
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||
<paragraph id="8B9A36A8DB5D7F4BFF735C00C967FDAA" blockId="8.[151,550,477,540]" box="[151,274,512,540]" pageId="8" pageNumber="363">
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||
<emphasis id="B951EABADB5D7F4BFF735C00C8D5FDAD" bold="true" box="[151,160,512,539]" pageId="8" pageNumber="363">(</emphasis>
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||
<figureCitation id="131E2A2DDB5D7F4BFF455C01C97FFDAA" box="[161,266,513,540]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStartId-0="9.[151,250,1718,1743]" captionStartId-1="10.[151,250,1841,1866]" captionStartId-2="11.[151,250,1652,1677]" captionStartId-3="12.[151,250,1632,1657]" captionTargetBox-0="[242,1371,204,1680]" captionTargetBox-1="[151,1436,181,1816]" captionTargetBox-2="[151,1436,181,1626]" captionTargetBox-3="[242,1333,865,1555]" captionTargetId-0="figure@9.[200,1388,181,1693]" captionTargetId-1="figure@10.[151,1436,181,1816]" captionTargetId-2="figure@11.[151,1436,181,1626]" captionTargetId-3="figure@12.[170,1417,813,1607]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionText-0="FIGURE 6. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani. A. entire worm (holotype MACN-Pa 706); B. scolex (holotype MACN-Pa 706); C. bothridium, distal surface; D. terminal mature proglottid (paratype MACN-Pa 708/1); E. detail of terminal genitalia of a terminal mature proglottid (paratype MACN-Pa 708/1); F. detail of oötype region of a terminal mature proglottid (paratype MACN-Pa 708/1). Abbreviations: cs. cirrus sac; mg. Mehlis’ gland; oc. oocapt; od. oviduct; ov. ovary; t. testes; ud. utero duct; ut. uterus; vd. vas deferens; vf. vitelline follicle; vg. vagina; vod. ventral osmorregulatory duct. ▲loculi of the anterior region; * loculi of the central region; ● loculi of the posterior region" captionText-1="FIGURE 7. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani, scanning electron micrographs. A. scolex, small letters indicate locations of details shown in figures 7C-G; B. distal bothridial surface; C. detail of distal bothridial surface; D. botrhidial rim; E. proximal bothridial surface; F. surface of the cephalic peduncle; G. surface of a stalk; H. surface of a terminal mature proglottid." captionText-2="FIGURE 8. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani, septal musculature of bothridium. A. longitudinal section of bothridium (paratype MACN-Pa 707/4); B. light micrographs of bothridium (paratype MACN-Pa 707/3); C. transverse section of bothridium (paratype MACN-Pa 707/5). Abbreviations: l. loculus; lmb. longitudinal muscle bundle; ls. Longitudinal septum; rm. radial muscle; ss. semicircular septum; tg. triangular gap; tmb. transverse muscle bundles; ts. transverse septum" captionText-3="FIGURE 9. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani, cross sections of a terminal mature proglottid (paratype MACN-Pa 708/3). A. cross section at level of testes, anterior to cirrus sac; B. cross section at level of the genital pore; C. cross section at level of the cirrus sac; D. cross section at level of the ovarian isthmus. Abbreviations: cs. cirrus sac; dod. dorsal osmorregulatory duct; ov. ovary; t. testes; ut. uterus; vd. vas deferens; vf. vitelline follicle; vg. vagina; vod. ventral osmorregulatory duct" figureDoi-0="http://doi.org/10.5281/zenodo.3909207" figureDoi-1="http://doi.org/10.5281/zenodo.3909209" figureDoi-2="http://doi.org/10.5281/zenodo.3909211" figureDoi-3="http://doi.org/10.5281/zenodo.3909213" httpUri-0="https://zenodo.org/record/3909207/files/figure.png" httpUri-1="https://zenodo.org/record/3909209/files/figure.png" httpUri-2="https://zenodo.org/record/3909211/files/figure.png" httpUri-3="https://zenodo.org/record/3909213/files/figure.png" pageId="8" pageNumber="363">Figs. 6–9</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33F6523DB5D7F4BFF735C49CBBCFC32" pageId="8" pageNumber="363" type="materials_examined">
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||
<paragraph id="8B9A36A8DB5D7F4BFF735C49CC9CFDD2" blockId="8.[151,1437,585,1980]" box="[151,1257,585,612]" pageId="8" pageNumber="363">
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||
<emphasis id="B951EABADB5D7F4BFF735C49C967FDD5" bold="true" box="[151,274,585,611]" pageId="8" pageNumber="363">
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||
<typeStatus id="549E880ADB5D7F4BFF735C49C8A7FDD5" box="[151,210,585,611]" pageId="8" pageNumber="363">Type</typeStatus>
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||
host.
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||
</emphasis>
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||
<taxonomicName id="4C254D2BDB5D7F4BFEFD5C49CAD5FDD5" authority="McEachran" authorityName="McEachran. These" authorityYear="1983" box="[281,672,585,611]" class="Chondrichthyes" family="Arhynchobatidae" genus="Psammobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="363" phylum="Chordata" rank="species" species="normani">
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||
<emphasis id="B951EABADB5D7F4BFEFD5C49CA60FDD5" box="[281,533,585,611]" italics="true" pageId="8" pageNumber="363">Psammobatis normani</emphasis>
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||
McEachran
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||
</taxonomicName>
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||
, shortfin sand skate (
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||
<taxonomicName id="4C254D2BDB5D7F4BFC685C49CC7AFDD5" box="[908,1039,585,611]" class="Chondrichthyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="363" phylum="Chordata" rank="order">Rajiformes</taxonomicName>
|
||
:
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||
<taxonomicName id="4C254D2BDB5D7F4BFBF35C49CCA8FDD2" box="[1047,1245,585,612]" class="Chondrichthyes" family="Arhynchobatidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="363" phylum="Chordata" rank="family">Arhynchobatidae</taxonomicName>
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||
).
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||
</paragraph>
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||
<paragraph id="8B9A36A8DB5D7F4BFF235C6DCCCAFD31" blockId="8.[151,1437,585,1980]" box="[199,1215,620,647]" pageId="8" pageNumber="363">
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||
<materialsCitation id="3B4D3CF5DB5D7F4BFF235C6DCCCEFD31" box="[199,1211,620,647]" collectorName="de Ajo" latitude="-36.566666" location="Mar de Ajo" longLatPrecision="1186" longitude="-54.65" pageId="8" pageNumber="363" specimenCount="1" stateProvince="Buenos Aires" typeStatus="holotype">
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||
<emphasis id="B951EABADB5D7F4BFF235C6DC913FD31" bold="true" box="[199,358,620,647]" pageId="8" pageNumber="363">
|
||
<typeStatus id="549E880ADB5D7F4BFF235C6DC977FD31" box="[199,258,621,647]" pageId="8" pageNumber="363">Type</typeStatus>
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||
locality.
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||
</emphasis>
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||
Coastal waters off
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||
<location id="8EFA6073DB5D7F4BFDDB5C6DCAB4FD31" LSID="urn:lsid:plazi:treatment:038C87BEDB5D7F4FFF735FDDCBB9FCA1:8EFA6073DB5D7F4BFDDB5C6DCAB4FD31" box="[575,705,621,647]" latitude="-36.566666" longLatPrecision="1186" longitude="-54.65" name="Mar de Ajo" pageId="8" pageNumber="363" stateProvince="Buenos Aires">
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||
Mar
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||
<collectorName id="26D0537EDB5D7F4BFD905C6DCAB4FD31" box="[628,705,621,647]" pageId="8" pageNumber="363">de Ajó</collectorName>
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||
</location>
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||
,
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<collectingRegion id="49E1F84ADB5D7F4BFD285C6DCBB8FD31" box="[716,973,621,647]" country="Argentina" name="Buenos Aires" pageId="8" pageNumber="363">Buenos Aires Province</collectingRegion>
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||
(
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<geoCoordinate id="EE11506FDB5D7F4BFC395C6DCC49FD31" box="[989,1084,621,647]" degrees="36" direction="south" minutes="34" orientation="latitude" pageId="8" pageNumber="363" precision="925" value="-36.566666">36°34’S</geoCoordinate>
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||
,
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||
<geoCoordinate id="EE11506FDB5D7F4BFBA35C6CCCC7FD31" box="[1095,1202,620,647]" degrees="54" direction="west" minutes="39" orientation="longitude" pageId="8" pageNumber="363" precision="925" value="-54.65">54°39’W</geoCoordinate>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="8B9A36A8DB5D7F4BFF235C90CAB6FD66" blockId="8.[151,1437,585,1980]" pageId="8" pageNumber="363">
|
||
<emphasis id="B951EABADB5D7F4BFF235C90C9D8FD1D" bold="true" box="[199,429,656,683]" pageId="8" pageNumber="363">Additional locality.</emphasis>
|
||
<materialsCitation id="3B4D3CF5DB5D7F4BFE525C91CACAFD66" latitude="-46.383335" location="Pinamar" longLatPrecision="1124" longitude="-64.333336" municipality="Coastal" pageId="8" pageNumber="363" specimenCount="1" stateProvince="Buenos Aires">
|
||
<collectingMunicipality id="6BFEACD2DB5D7F4BFE525C91CA7FFD1D" box="[438,522,657,683]" pageId="8" pageNumber="363">Coastal</collectingMunicipality>
|
||
waters off
|
||
<location id="8EFA6073DB5D7F4BFD6B5C91CA9BFD1D" LSID="urn:lsid:plazi:treatment:038C87BEDB5D7F4FFF735FDDCBB9FCA1:8EFA6073DB5D7F4BFD6B5C91CA9BFD1D" box="[655,750,657,683]" latitude="-46.383335" longLatPrecision="1124" longitude="-64.333336" municipality="Coastal" name="Pinamar" pageId="8" pageNumber="363" stateProvince="Buenos Aires">Pinamar</location>
|
||
,
|
||
<collectingRegion id="49E1F84ADB5D7F4BFD185C91CC74FD1D" box="[764,1025,657,683]" country="Argentina" name="Buenos Aires" pageId="8" pageNumber="363">Buenos Aires Province</collectingRegion>
|
||
(
|
||
<geoCoordinate id="EE11506FDB5D7F4BFBF75C91CC07FD1D" box="[1043,1138,657,683]" degrees="37" direction="south" minutes="12" orientation="latitude" pageId="8" pageNumber="363" precision="925" value="-37.2">37°12’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE11506FDB5D7F4BFB645C90CC9DFD1D" box="[1152,1256,656,683]" degrees="54" direction="west" minutes="53" orientation="longitude" pageId="8" pageNumber="363" precision="925" value="-54.883335">54°53’W</geoCoordinate>
|
||
) and off Caleta Olivia,
|
||
<collectingRegion id="49E1F84ADB5D7F4BFF0F5CB5C9A7FD79" box="[235,466,693,719]" country="Argentina" name="Santa Cruz" pageId="8" pageNumber="363">Santa Cruz Province</collectingRegion>
|
||
(
|
||
<geoCoordinate id="EE11506FDB5D7F4BFE055CB5CA35FD79" box="[481,576,693,719]" degrees="46" direction="south" minutes="23" orientation="latitude" pageId="8" pageNumber="363" precision="925" value="-46.383335">46°23’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE11506FDB5D7F4BFDA85CB5CAC2FD66" box="[588,695,693,720]" degrees="64" direction="west" minutes="20" orientation="longitude" pageId="8" pageNumber="363" precision="925" value="-64.333336">64°20’W</geoCoordinate>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9A36A8DB5D7F4BFF235CD8CA35FD45" blockId="8.[151,1437,585,1980]" box="[199,576,728,755]" pageId="8" pageNumber="363">
|
||
<emphasis id="B951EABADB5D7F4BFF235CD8C9FCFD45" bold="true" box="[199,393,728,755]" pageId="8" pageNumber="363">Site of infection.</emphasis>
|
||
Spiral intestine.
|
||
</paragraph>
|
||
<paragraph id="8B9A36A8DB5D7F4BFF235CFDCA4AFCE9" blockId="8.[151,1437,585,1980]" pageId="8" pageNumber="363">
|
||
<emphasis id="B951EABADB5D7F4BFF235CFDC900FCA1" bold="true" box="[199,373,764,791]" pageId="8" pageNumber="363">Type material.</emphasis>
|
||
<materialsCitation id="3B4D3CF5DB5D7F4BFE995CFDCA4AFCE9" collectionCode="IPCAS" pageId="8" pageNumber="363" specimenCode="C-837" specimenCount="16" typeStatus="holotype">
|
||
<typeStatus id="549E880ADB5D7F4BFE995CFDC990FCA1" box="[381,485,765,791]" pageId="8" pageNumber="363" type="holotype">Holotype</typeStatus>
|
||
MACN-Pa 706 (entire worm),
|
||
<specimenCount id="9D23FD21DB5D7F4BFCAC5CFDCBA3FCA1" box="[840,982,765,791]" pageId="8" pageNumber="363" type="generic" typeStatus="paratypes">12 paratypes</specimenCount>
|
||
MACN-Pa 707/1–5, 708/1–3, 709, 710, 711/1–2 (9 entire worms, histological sections of 1 mature proglottid, histological sections of 2 scoleces);
|
||
<specimenCount id="9D23FD21DB5D7F4BFADC5D21C8C9FCE9" pageId="8" pageNumber="363" type="generic" typeStatus="paratypes">4 paratypes</specimenCount>
|
||
<collectionCode id="ED34AE6DDB5D7F4BFF205D45C966FCE9" box="[196,275,837,863]" pageId="8" pageNumber="363">IPCAS</collectionCode>
|
||
<specimenCode id="DB839ED3DB5D7F4BFEFE5D45C916FCE9" box="[282,355,837,863]" collectionCode="C-" pageId="8" pageNumber="363">C-837</specimenCode>
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||
/1 (4 entire worms).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B9A36A8DB5D7F4BFF235D69CBBCFC32" blockId="8.[151,1437,585,1980]" box="[199,969,872,900]" pageId="8" pageNumber="363">
|
||
<emphasis id="B951EABADB5D7F4BFF235D69C9ABFC35" bold="true" box="[199,478,872,899]" pageId="8" pageNumber="363">Prevalence of infection.</emphasis>
|
||
50% (5 hosts infected out of 10 examined).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33F6523DB5D7F4BFF235D8DCD6BFC11" box="[199,1310,909,935]" pageId="8" pageNumber="363" type="etymology">
|
||
<paragraph id="8B9A36A8DB5D7F4BFF235D8DCD6BFC11" blockId="8.[151,1437,585,1980]" box="[199,1310,909,935]" pageId="8" pageNumber="363">
|
||
<emphasis id="B951EABADB5D7F4BFF235D8DC93BFC11" bold="true" box="[199,334,909,935]" pageId="8" pageNumber="363">Etymology.</emphasis>
|
||
This species is named in memory of Alfredo Marino Franzese, the first author’s father.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33F6523DB5D7F4FFF235DB0CCCFFDAD" lastPageId="12" lastPageNumber="367" pageId="8" pageNumber="363" type="description">
|
||
<paragraph id="8B9A36A8DB5D7F4BFF235DB0CC19F909" blockId="8.[151,1437,585,1980]" pageId="8" pageNumber="363">
|
||
<emphasis id="B951EABADB5D7F4BFF235DB0C92CFC7D" bold="true" box="[199,345,944,971]" pageId="8" pageNumber="363">Description.</emphasis>
|
||
Based on
|
||
<specimenCount id="9D23FD21DB5D7F4BFE2A5DB1CA12FC7D" box="[462,615,945,971]" pageId="8" pageNumber="363" type="generic">24 specimens</specimenCount>
|
||
prepared as follows: whole mounts of 14 entire worms, serial sections of 1 mature proglottid and 2 scoleces, 7 worms examined with SEM. Worms euapolytic, 5.37–26.81 mm (13.03 mm ± 6.67 mm, 13) long, 420–705 (495 ± 95, 7) wide, maximum width at level of scolex. Proglottids craspedote, 85–187 (143 ± 32, 11) per worm, 84–172 (137 ± 29, 8) immature proglottids, 1–4 (3 ± 1, 8) mature proglottids (
|
||
<figureCitation id="131E2A2DDB5D7F4BFAD05A1DCDFAFB8E" box="[1332,1423,1053,1080]" captionStart="FIGURE 6" captionStartId="9.[151,250,1718,1743]" captionTargetBox="[242,1371,204,1680]" captionTargetId="figure@9.[200,1388,181,1693]" captionTargetPageId="9" captionText="FIGURE 6. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani. A. entire worm (holotype MACN-Pa 706); B. scolex (holotype MACN-Pa 706); C. bothridium, distal surface; D. terminal mature proglottid (paratype MACN-Pa 708/1); E. detail of terminal genitalia of a terminal mature proglottid (paratype MACN-Pa 708/1); F. detail of oötype region of a terminal mature proglottid (paratype MACN-Pa 708/1). Abbreviations: cs. cirrus sac; mg. Mehlis’ gland; oc. oocapt; od. oviduct; ov. ovary; t. testes; ud. utero duct; ut. uterus; vd. vas deferens; vf. vitelline follicle; vg. vagina; vod. ventral osmorregulatory duct. ▲loculi of the anterior region; * loculi of the central region; ● loculi of the posterior region" figureDoi="http://doi.org/10.5281/zenodo.3909207" httpUri="https://zenodo.org/record/3909207/files/figure.png" pageId="8" pageNumber="363">Fig. 6A</figureCitation>
|
||
). Scolex 710–1,435 (1059 ± 278, 10) long, composed of 4 stalked bothridia and cephalic peduncle; myzorhynchus absent in adult stage (
|
||
<figureCitation id="131E2A2DDB5D7F4BFE745A65CA7AFBC9" box="[400,527,1125,1151]" captionStart="FIGURE 6" captionStartId="9.[151,250,1718,1743]" captionTargetBox="[242,1371,204,1680]" captionTargetId="figure@9.[200,1388,181,1693]" captionTargetPageId="9" captionText="FIGURE 6. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani. A. entire worm (holotype MACN-Pa 706); B. scolex (holotype MACN-Pa 706); C. bothridium, distal surface; D. terminal mature proglottid (paratype MACN-Pa 708/1); E. detail of terminal genitalia of a terminal mature proglottid (paratype MACN-Pa 708/1); F. detail of oötype region of a terminal mature proglottid (paratype MACN-Pa 708/1). Abbreviations: cs. cirrus sac; mg. Mehlis’ gland; oc. oocapt; od. oviduct; ov. ovary; t. testes; ud. utero duct; ut. uterus; vd. vas deferens; vf. vitelline follicle; vg. vagina; vod. ventral osmorregulatory duct. ▲loculi of the anterior region; * loculi of the central region; ● loculi of the posterior region" figureDoi="http://doi.org/10.5281/zenodo.3909207" httpUri="https://zenodo.org/record/3909207/files/figure.png" pageId="8" pageNumber="363">Figs. 6A,B</figureCitation>
|
||
,
|
||
<figureCitation id="131E2A2DDB5D7F4BFDF95A65CA37FBC9" box="[541,578,1125,1151]" captionStart="FIGURE 7" captionStartId="10.[151,250,1841,1866]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure@10.[151,1436,181,1816]" captionTargetPageId="10" captionText="FIGURE 7. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani, scanning electron micrographs. A. scolex, small letters indicate locations of details shown in figures 7C-G; B. distal bothridial surface; C. detail of distal bothridial surface; D. botrhidial rim; E. proximal bothridial surface; F. surface of the cephalic peduncle; G. surface of a stalk; H. surface of a terminal mature proglottid." figureDoi="http://doi.org/10.5281/zenodo.3909209" httpUri="https://zenodo.org/record/3909209/files/figure.png" pageId="8" pageNumber="363">7A</figureCitation>
|
||
). Bothridia facially loculate, 280–445 (363 ± 117, 2) long, 230–360 (267 ± 39, 12) wide. Total number of loculi 13–16 (14 ± 1, 16) (
|
||
<figureCitation id="131E2A2DDB5D7F4BFCCB5A89CBE1FB15" box="[815,916,1161,1187]" captionStart="FIGURE 6" captionStartId="9.[151,250,1718,1743]" captionTargetBox="[242,1371,204,1680]" captionTargetId="figure@9.[200,1388,181,1693]" captionTargetPageId="9" captionText="FIGURE 6. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani. A. entire worm (holotype MACN-Pa 706); B. scolex (holotype MACN-Pa 706); C. bothridium, distal surface; D. terminal mature proglottid (paratype MACN-Pa 708/1); E. detail of terminal genitalia of a terminal mature proglottid (paratype MACN-Pa 708/1); F. detail of oötype region of a terminal mature proglottid (paratype MACN-Pa 708/1). Abbreviations: cs. cirrus sac; mg. Mehlis’ gland; oc. oocapt; od. oviduct; ov. ovary; t. testes; ud. utero duct; ut. uterus; vd. vas deferens; vf. vitelline follicle; vg. vagina; vod. ventral osmorregulatory duct. ▲loculi of the anterior region; * loculi of the central region; ● loculi of the posterior region" figureDoi="http://doi.org/10.5281/zenodo.3909207" httpUri="https://zenodo.org/record/3909207/files/figure.png" pageId="8" pageNumber="363">Figs. 6C</figureCitation>
|
||
,
|
||
<figureCitation id="131E2A2DDB5D7F4BFC465A89CBB3FB15" box="[930,966,1161,1187]" captionStart="FIGURE 7" captionStartId="10.[151,250,1841,1866]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure@10.[151,1436,181,1816]" captionTargetPageId="10" captionText="FIGURE 7. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani, scanning electron micrographs. A. scolex, small letters indicate locations of details shown in figures 7C-G; B. distal bothridial surface; C. detail of distal bothridial surface; D. botrhidial rim; E. proximal bothridial surface; F. surface of the cephalic peduncle; G. surface of a stalk; H. surface of a terminal mature proglottid." figureDoi="http://doi.org/10.5281/zenodo.3909209" httpUri="https://zenodo.org/record/3909209/files/figure.png" pageId="8" pageNumber="363">7B</figureCitation>
|
||
). Anterior region of bothridia divided by transverse septa in 4–5 (5 ± 0.4, 19) loculi arranged in a single column, central region divided in 2–3 (2 ± 0.24, 17) loculi delimited by 2–3 semicircular septa with convex surface facing anteriorly, and posterior region divided by transverse septa in 0–3 (2 ± 1, 17) loculi arranged in a single column, followed by 1–4 pairs of loculi delimited by transverse and medial longitudinal septa, arranged in 2 columns, ending in single posteriormost loculus. In some specimens, semicircular septa extend posteriorly, reaching first loculus of posterior part of botridium. Anteriormost loculus 28–55 (40 ± 8, 13) long, 35–83 (54 ± 13, 13) wide, muscular wall 8–10 (9 ± 1, 8) thick. Posteriormost loculus 20–33 (28 ± 4, 9) long, 20–48 (30 ± 9, 9) wide, muscular wall 5–10 (7 ± 2, 8) thick. Transverse, semicircular and longitudinal septa formed distally by transverse muscle bundles with underlying layer of radial muscle, proximal portion formed by radial muscles with proximal fibres ending adjacent to each other, forming triangular gap (
|
||
<figureCitation id="131E2A2DDB5D7F4BFA8B5BCDC8D2F9BD" captionStart="FIGURE 8" captionStartId="11.[151,250,1652,1677]" captionTargetBox="[151,1436,181,1626]" captionTargetId="figure@11.[151,1436,181,1626]" captionTargetPageId="11" captionText="FIGURE 8. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani, septal musculature of bothridium. A. longitudinal section of bothridium (paratype MACN-Pa 707/4); B. light micrographs of bothridium (paratype MACN-Pa 707/3); C. transverse section of bothridium (paratype MACN-Pa 707/5). Abbreviations: l. loculus; lmb. longitudinal muscle bundle; ls. Longitudinal septum; rm. radial muscle; ss. semicircular septum; tg. triangular gap; tmb. transverse muscle bundles; ts. transverse septum" figureDoi="http://doi.org/10.5281/zenodo.3909211" httpUri="https://zenodo.org/record/3909211/files/figure.png" pageId="8" pageNumber="363">Fig. 8</figureCitation>
|
||
). Bothridial stalks 70–150 (104 ± 27, 6) long, 78–98 (88 ± 8, 6) wide, attached to centre of bothridium. Cephalic peduncle 450–1,068 (779 ± 211, 12) long, 88–135 (109 ± 15, 12) wide at posterior end. Apex of scolex covered with acicular to capilliform filitriches. Proximal bothridial surface with acicular filitriches along most of its length, posterior portion also with small gladiate spinitriches (
|
||
<figureCitation id="131E2A2DDB5D7F4BFD1A585DCB23F9C1" box="[766,854,1629,1655]" captionStart="FIGURE 7" captionStartId="10.[151,250,1841,1866]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure@10.[151,1436,181,1816]" captionTargetPageId="10" captionText="FIGURE 7. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani, scanning electron micrographs. A. scolex, small letters indicate locations of details shown in figures 7C-G; B. distal bothridial surface; C. detail of distal bothridial surface; D. botrhidial rim; E. proximal bothridial surface; F. surface of the cephalic peduncle; G. surface of a stalk; H. surface of a terminal mature proglottid." figureDoi="http://doi.org/10.5281/zenodo.3909209" httpUri="https://zenodo.org/record/3909209/files/figure.png" pageId="8" pageNumber="363">Fig. 7E</figureCitation>
|
||
); proximal bothridial edge covered with acicular filitriches (
|
||
<figureCitation id="131E2A2DDB5D7F4BFEE55881C92EF92D" box="[257,347,1665,1691]" captionStart="FIGURE 7" captionStartId="10.[151,250,1841,1866]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure@10.[151,1436,181,1816]" captionTargetPageId="10" captionText="FIGURE 7. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani, scanning electron micrographs. A. scolex, small letters indicate locations of details shown in figures 7C-G; B. distal bothridial surface; C. detail of distal bothridial surface; D. botrhidial rim; E. proximal bothridial surface; F. surface of the cephalic peduncle; G. surface of a stalk; H. surface of a terminal mature proglottid." figureDoi="http://doi.org/10.5281/zenodo.3909209" httpUri="https://zenodo.org/record/3909209/files/figure.png" pageId="8" pageNumber="363">Fig. 7D</figureCitation>
|
||
). Distal bothridial surface with coniform spinitriches and acicular filitriches (
|
||
<figureCitation id="131E2A2DDB5D7F4BFB5A5881CD62F92D" box="[1214,1303,1665,1691]" captionStart="FIGURE 7" captionStartId="10.[151,250,1841,1866]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure@10.[151,1436,181,1816]" captionTargetPageId="10" captionText="FIGURE 7. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani, scanning electron micrographs. A. scolex, small letters indicate locations of details shown in figures 7C-G; B. distal bothridial surface; C. detail of distal bothridial surface; D. botrhidial rim; E. proximal bothridial surface; F. surface of the cephalic peduncle; G. surface of a stalk; H. surface of a terminal mature proglottid." figureDoi="http://doi.org/10.5281/zenodo.3909209" httpUri="https://zenodo.org/record/3909209/files/figure.png" pageId="8" pageNumber="363">Fig. 7C</figureCitation>
|
||
). Bothridial stalks and cephalic peduncle covered with acicular to capilliform filitriches (
|
||
<figureCitation id="131E2A2DDB5D7F4BFC1558A5CC2AF909" box="[1009,1119,1701,1727]" captionStart="FIGURE 7" captionStartId="10.[151,250,1841,1866]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure@10.[151,1436,181,1816]" captionTargetPageId="10" captionText="FIGURE 7. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani, scanning electron micrographs. A. scolex, small letters indicate locations of details shown in figures 7C-G; B. distal bothridial surface; C. detail of distal bothridial surface; D. botrhidial rim; E. proximal bothridial surface; F. surface of the cephalic peduncle; G. surface of a stalk; H. surface of a terminal mature proglottid." figureDoi="http://doi.org/10.5281/zenodo.3909209" httpUri="https://zenodo.org/record/3909209/files/figure.png" pageId="8" pageNumber="363">Fig. 7F,G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9A36A8DB5D7F4FFF2358C9CCCFFDAD" blockId="8.[151,1437,585,1980]" lastBlockId="12.[151,1437,152,791]" lastPageId="12" lastPageNumber="367" pageId="8" pageNumber="363">
|
||
Immature proglottids initially wider than long, becoming longer than wide with maturity. Mature proglottids (excluding terminal proglottid) longer than wide, 353–818 (573 ± 227, 6) long (velum not included in proglottid length), 295–355 (323 ± 26, 6) wide, velum 38–63 (50 ± 10, 6) long, covering 4–8% (6 ± 1, 6) of adjacent proglottid. Terminal mature proglottid 575–1,060 (824 ± 159, 13) long, 183–310 (277 ± 35, 13) wide, length to width ratio 1: 2–6 (3 ± 1, 11) (
|
||
<figureCitation id="131E2A2DDB5D7F4BFE8E5959C9B1F8C2" box="[362,452,1881,1908]" captionStart="FIGURE 6" captionStartId="9.[151,250,1718,1743]" captionTargetBox="[242,1371,204,1680]" captionTargetId="figure@9.[200,1388,181,1693]" captionTargetPageId="9" captionText="FIGURE 6. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani. A. entire worm (holotype MACN-Pa 706); B. scolex (holotype MACN-Pa 706); C. bothridium, distal surface; D. terminal mature proglottid (paratype MACN-Pa 708/1); E. detail of terminal genitalia of a terminal mature proglottid (paratype MACN-Pa 708/1); F. detail of oötype region of a terminal mature proglottid (paratype MACN-Pa 708/1). Abbreviations: cs. cirrus sac; mg. Mehlis’ gland; oc. oocapt; od. oviduct; ov. ovary; t. testes; ud. utero duct; ut. uterus; vd. vas deferens; vf. vitelline follicle; vg. vagina; vod. ventral osmorregulatory duct. ▲loculi of the anterior region; * loculi of the central region; ● loculi of the posterior region" figureDoi="http://doi.org/10.5281/zenodo.3909207" httpUri="https://zenodo.org/record/3909207/files/figure.png" pageId="8" pageNumber="363">Fig. 6D</figureCitation>
|
||
). Mature proglottids covered by capilliform filitriches (
|
||
<figureCitation id="131E2A2DDB5D7F4BFBD55959CCFEF8C5" box="[1073,1163,1881,1907]" captionStart="FIGURE 7" captionStartId="10.[151,250,1841,1866]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure@10.[151,1436,181,1816]" captionTargetPageId="10" captionText="FIGURE 7. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani, scanning electron micrographs. A. scolex, small letters indicate locations of details shown in figures 7C-G; B. distal bothridial surface; C. detail of distal bothridial surface; D. botrhidial rim; E. proximal bothridial surface; F. surface of the cephalic peduncle; G. surface of a stalk; H. surface of a terminal mature proglottid." figureDoi="http://doi.org/10.5281/zenodo.3909209" httpUri="https://zenodo.org/record/3909209/files/figure.png" pageId="8" pageNumber="363">Fig. 7H</figureCitation>
|
||
). Testes oval, 22–33 (26 ± 3, 9) per proglottid, 18–50 (33 ± 8, 8, 40) long, 32–73 (54 ± 10, 8, 40) wide, arranged in 2 columns, anterior to cirrus sac (
|
||
<figureCitation id="131E2A2DDB5D7F4BFEF459A1C91FF80A" box="[272,362,1953,1980]" captionStart="FIGURE 6" captionStartId="9.[151,250,1718,1743]" captionTargetBox="[242,1371,204,1680]" captionTargetId="figure@9.[200,1388,181,1693]" captionTargetPageId="9" captionText="FIGURE 6. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani. A. entire worm (holotype MACN-Pa 706); B. scolex (holotype MACN-Pa 706); C. bothridium, distal surface; D. terminal mature proglottid (paratype MACN-Pa 708/1); E. detail of terminal genitalia of a terminal mature proglottid (paratype MACN-Pa 708/1); F. detail of oötype region of a terminal mature proglottid (paratype MACN-Pa 708/1). Abbreviations: cs. cirrus sac; mg. Mehlis’ gland; oc. oocapt; od. oviduct; ov. ovary; t. testes; ud. utero duct; ut. uterus; vd. vas deferens; vf. vitelline follicle; vg. vagina; vod. ventral osmorregulatory duct. ▲loculi of the anterior region; * loculi of the central region; ● loculi of the posterior region" figureDoi="http://doi.org/10.5281/zenodo.3909207" httpUri="https://zenodo.org/record/3909207/files/figure.png" pageId="8" pageNumber="363">Fig. 6D</figureCitation>
|
||
), 1 layer deep in cross-section (
|
||
<figureCitation id="131E2A2DDB5D7F4BFD2D59A1CB56F80D" box="[713,803,1953,1979]" captionStart="FIGURE 9" captionStartId="12.[151,250,1632,1657]" captionTargetBox="[242,1333,865,1555]" captionTargetId="figure@12.[170,1417,813,1607]" captionTargetPageId="12" captionText="FIGURE 9. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani, cross sections of a terminal mature proglottid (paratype MACN-Pa 708/3). A. cross section at level of testes, anterior to cirrus sac; B. cross section at level of the genital pore; C. cross section at level of the cirrus sac; D. cross section at level of the ovarian isthmus. Abbreviations: cs. cirrus sac; dod. dorsal osmorregulatory duct; ov. ovary; t. testes; ut. uterus; vd. vas deferens; vf. vitelline follicle; vg. vagina; vod. ventral osmorregulatory duct" figureDoi="http://doi.org/10.5281/zenodo.3909213" httpUri="https://zenodo.org/record/3909213/files/figure.png" pageId="8" pageNumber="363">Fig. 9A</figureCitation>
|
||
). Cirrus sac oval to pyriform, 91–140 (125 ± 19, 5) long, 58–78 (69 ± 8, 5) wide, containing coiled cirrus covered with spinitriches; vas deferens dorsal, highly coiled, entering dorsally at proximal half of cirrus sac (
|
||
<figureCitation id="131E2A2DDB597F4FFD655EBDCA8BFF61" box="[641,766,189,215]" captionStart="FIGURE 6" captionStartId="9.[151,250,1718,1743]" captionTargetBox="[242,1371,204,1680]" captionTargetId="figure@9.[200,1388,181,1693]" captionTargetPageId="9" captionText="FIGURE 6. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani. A. entire worm (holotype MACN-Pa 706); B. scolex (holotype MACN-Pa 706); C. bothridium, distal surface; D. terminal mature proglottid (paratype MACN-Pa 708/1); E. detail of terminal genitalia of a terminal mature proglottid (paratype MACN-Pa 708/1); F. detail of oötype region of a terminal mature proglottid (paratype MACN-Pa 708/1). Abbreviations: cs. cirrus sac; mg. Mehlis’ gland; oc. oocapt; od. oviduct; ov. ovary; t. testes; ud. utero duct; ut. uterus; vd. vas deferens; vf. vitelline follicle; vg. vagina; vod. ventral osmorregulatory duct. ▲loculi of the anterior region; * loculi of the central region; ● loculi of the posterior region" figureDoi="http://doi.org/10.5281/zenodo.3909207" httpUri="https://zenodo.org/record/3909207/files/figure.png" pageId="12" pageNumber="367">Figs. 6D,E</figureCitation>
|
||
,
|
||
<figureCitation id="131E2A2DDB597F4FFCEF5EBDCB5AFF61" box="[779,815,189,215]" captionStart="FIGURE 9" captionStartId="12.[151,250,1632,1657]" captionTargetBox="[242,1333,865,1555]" captionTargetId="figure@12.[170,1417,813,1607]" captionTargetPageId="12" captionText="FIGURE 9. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani, cross sections of a terminal mature proglottid (paratype MACN-Pa 708/3). A. cross section at level of testes, anterior to cirrus sac; B. cross section at level of the genital pore; C. cross section at level of the cirrus sac; D. cross section at level of the ovarian isthmus. Abbreviations: cs. cirrus sac; dod. dorsal osmorregulatory duct; ov. ovary; t. testes; ut. uterus; vd. vas deferens; vf. vitelline follicle; vg. vagina; vod. ventral osmorregulatory duct" figureDoi="http://doi.org/10.5281/zenodo.3909213" httpUri="https://zenodo.org/record/3909213/files/figure.png" pageId="12" pageNumber="367">9C</figureCitation>
|
||
). Genital pores marginal, 33–60% (44 ± 8, 10) of proglottid length from posterior end. Vagina thick-walled, surrounded by darkly stained cells distally, extending from oötype along midline of proglottid to anterior margin of cirrus sac, then laterally along anterior margin of cirrus sac to common genital atrium; vaginal sphincter absent (
|
||
<figureCitation id="131E2A2DDB597F4FFD105F29CB07FEF5" box="[756,882,297,323]" captionStart="FIGURE 6" captionStartId="9.[151,250,1718,1743]" captionTargetBox="[242,1371,204,1680]" captionTargetId="figure@9.[200,1388,181,1693]" captionTargetPageId="9" captionText="FIGURE 6. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani. A. entire worm (holotype MACN-Pa 706); B. scolex (holotype MACN-Pa 706); C. bothridium, distal surface; D. terminal mature proglottid (paratype MACN-Pa 708/1); E. detail of terminal genitalia of a terminal mature proglottid (paratype MACN-Pa 708/1); F. detail of oötype region of a terminal mature proglottid (paratype MACN-Pa 708/1). Abbreviations: cs. cirrus sac; mg. Mehlis’ gland; oc. oocapt; od. oviduct; ov. ovary; t. testes; ud. utero duct; ut. uterus; vd. vas deferens; vf. vitelline follicle; vg. vagina; vod. ventral osmorregulatory duct. ▲loculi of the anterior region; * loculi of the central region; ● loculi of the posterior region" figureDoi="http://doi.org/10.5281/zenodo.3909207" httpUri="https://zenodo.org/record/3909207/files/figure.png" pageId="12" pageNumber="367">Figs. 6D,E</figureCitation>
|
||
,
|
||
<figureCitation id="131E2A2DDB597F4FFC645F29CBD1FEF5" box="[896,932,297,323]" captionStart="FIGURE 9" captionStartId="12.[151,250,1632,1657]" captionTargetBox="[242,1333,865,1555]" captionTargetId="figure@12.[170,1417,813,1607]" captionTargetPageId="12" captionText="FIGURE 9. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani, cross sections of a terminal mature proglottid (paratype MACN-Pa 708/3). A. cross section at level of testes, anterior to cirrus sac; B. cross section at level of the genital pore; C. cross section at level of the cirrus sac; D. cross section at level of the ovarian isthmus. Abbreviations: cs. cirrus sac; dod. dorsal osmorregulatory duct; ov. ovary; t. testes; ut. uterus; vd. vas deferens; vf. vitelline follicle; vg. vagina; vod. ventral osmorregulatory duct" figureDoi="http://doi.org/10.5281/zenodo.3909213" httpUri="https://zenodo.org/record/3909213/files/figure.png" pageId="12" pageNumber="367">9B</figureCitation>
|
||
). Ovary near posterior end of proglottid, Hshaped in dorsoventral view (
|
||
<figureCitation id="131E2A2DDB597F4FFE385F4DCA3EFEDE" box="[476,587,333,360]" captionStart="FIGURE 6" captionStartId="9.[151,250,1718,1743]" captionTargetBox="[242,1371,204,1680]" captionTargetId="figure@9.[200,1388,181,1693]" captionTargetPageId="9" captionText="FIGURE 6. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani. A. entire worm (holotype MACN-Pa 706); B. scolex (holotype MACN-Pa 706); C. bothridium, distal surface; D. terminal mature proglottid (paratype MACN-Pa 708/1); E. detail of terminal genitalia of a terminal mature proglottid (paratype MACN-Pa 708/1); F. detail of oötype region of a terminal mature proglottid (paratype MACN-Pa 708/1). Abbreviations: cs. cirrus sac; mg. Mehlis’ gland; oc. oocapt; od. oviduct; ov. ovary; t. testes; ud. utero duct; ut. uterus; vd. vas deferens; vf. vitelline follicle; vg. vagina; vod. ventral osmorregulatory duct. ▲loculi of the anterior region; * loculi of the central region; ● loculi of the posterior region" figureDoi="http://doi.org/10.5281/zenodo.3909207" httpUri="https://zenodo.org/record/3909207/files/figure.png" pageId="12" pageNumber="367">Fig. 6D,F</figureCitation>
|
||
), symmetric, tetralobed in cross-section (
|
||
<figureCitation id="131E2A2DDB597F4FFBF75F4DCC19FED1" box="[1043,1132,333,359]" captionStart="FIGURE 9" captionStartId="12.[151,250,1632,1657]" captionTargetBox="[242,1333,865,1555]" captionTargetId="figure@12.[170,1417,813,1607]" captionTargetPageId="12" captionText="FIGURE 9. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani, cross sections of a terminal mature proglottid (paratype MACN-Pa 708/3). A. cross section at level of testes, anterior to cirrus sac; B. cross section at level of the genital pore; C. cross section at level of the cirrus sac; D. cross section at level of the ovarian isthmus. Abbreviations: cs. cirrus sac; dod. dorsal osmorregulatory duct; ov. ovary; t. testes; ut. uterus; vd. vas deferens; vf. vitelline follicle; vg. vagina; vod. ventral osmorregulatory duct" figureDoi="http://doi.org/10.5281/zenodo.3909213" httpUri="https://zenodo.org/record/3909213/files/figure.png" pageId="12" pageNumber="367">Fig. 9D</figureCitation>
|
||
), 118–188 (150 ± 22, 8, 16) long, 78–163 (137 ± 32, 6) wide at ovarian isthmus. Mehlis’ gland posterior to ovarian isthmus, 38–55 (46 ± 8, 5) in diameter (
|
||
<figureCitation id="131E2A2DDB597F4FFEE35F95C929FE06" box="[263,348,405,432]" captionStart="FIGURE 6" captionStartId="9.[151,250,1718,1743]" captionTargetBox="[242,1371,204,1680]" captionTargetId="figure@9.[200,1388,181,1693]" captionTargetPageId="9" captionText="FIGURE 6. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani. A. entire worm (holotype MACN-Pa 706); B. scolex (holotype MACN-Pa 706); C. bothridium, distal surface; D. terminal mature proglottid (paratype MACN-Pa 708/1); E. detail of terminal genitalia of a terminal mature proglottid (paratype MACN-Pa 708/1); F. detail of oötype region of a terminal mature proglottid (paratype MACN-Pa 708/1). Abbreviations: cs. cirrus sac; mg. Mehlis’ gland; oc. oocapt; od. oviduct; ov. ovary; t. testes; ud. utero duct; ut. uterus; vd. vas deferens; vf. vitelline follicle; vg. vagina; vod. ventral osmorregulatory duct. ▲loculi of the anterior region; * loculi of the central region; ● loculi of the posterior region" figureDoi="http://doi.org/10.5281/zenodo.3909207" httpUri="https://zenodo.org/record/3909207/files/figure.png" pageId="12" pageNumber="367">Fig. 6F</figureCitation>
|
||
). Vitelline follicles 10–22 (16 ± 3, 6, 30) long, 15–35 (25 ± 4, 6, 30) wide, in 1 dorsal and 1 ventral column on each lateral margin of proglottid, extending along entire proglottid, interrupted by vagina and cirrus sac ventrally (
|
||
<figureCitation id="131E2A2DDB597F4FFEED5FDDC9F1FE41" box="[265,388,477,503]" captionStart="FIGURE 6" captionStartId="9.[151,250,1718,1743]" captionTargetBox="[242,1371,204,1680]" captionTargetId="figure@9.[200,1388,181,1693]" captionTargetPageId="9" captionText="FIGURE 6. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani. A. entire worm (holotype MACN-Pa 706); B. scolex (holotype MACN-Pa 706); C. bothridium, distal surface; D. terminal mature proglottid (paratype MACN-Pa 708/1); E. detail of terminal genitalia of a terminal mature proglottid (paratype MACN-Pa 708/1); F. detail of oötype region of a terminal mature proglottid (paratype MACN-Pa 708/1). Abbreviations: cs. cirrus sac; mg. Mehlis’ gland; oc. oocapt; od. oviduct; ov. ovary; t. testes; ud. utero duct; ut. uterus; vd. vas deferens; vf. vitelline follicle; vg. vagina; vod. ventral osmorregulatory duct. ▲loculi of the anterior region; * loculi of the central region; ● loculi of the posterior region" figureDoi="http://doi.org/10.5281/zenodo.3909207" httpUri="https://zenodo.org/record/3909207/files/figure.png" pageId="12" pageNumber="367">Figs. 6D,E</figureCitation>
|
||
,
|
||
<figureCitation id="131E2A2DDB597F4FFE6B5FDDC9C6FE41" box="[399,435,477,503]" captionStart="FIGURE 9" captionStartId="12.[151,250,1632,1657]" captionTargetBox="[242,1333,865,1555]" captionTargetId="figure@12.[170,1417,813,1607]" captionTargetPageId="12" captionText="FIGURE 9. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani, cross sections of a terminal mature proglottid (paratype MACN-Pa 708/3). A. cross section at level of testes, anterior to cirrus sac; B. cross section at level of the genital pore; C. cross section at level of the cirrus sac; D. cross section at level of the ovarian isthmus. Abbreviations: cs. cirrus sac; dod. dorsal osmorregulatory duct; ov. ovary; t. testes; ut. uterus; vd. vas deferens; vf. vitelline follicle; vg. vagina; vod. ventral osmorregulatory duct" figureDoi="http://doi.org/10.5281/zenodo.3909213" httpUri="https://zenodo.org/record/3909213/files/figure.png" pageId="12" pageNumber="367">9B</figureCitation>
|
||
). Uterus sacciform, occupying midline of proglottid, from level of ovarian isthmus to near anterior margin of proglottid (
|
||
<figureCitation id="131E2A2DDB597F4FFE035C01CA34FDAA" box="[487,577,513,540]" captionStart="FIGURE 6" captionStartId="9.[151,250,1718,1743]" captionTargetBox="[242,1371,204,1680]" captionTargetId="figure@9.[200,1388,181,1693]" captionTargetPageId="9" captionText="FIGURE 6. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani. A. entire worm (holotype MACN-Pa 706); B. scolex (holotype MACN-Pa 706); C. bothridium, distal surface; D. terminal mature proglottid (paratype MACN-Pa 708/1); E. detail of terminal genitalia of a terminal mature proglottid (paratype MACN-Pa 708/1); F. detail of oötype region of a terminal mature proglottid (paratype MACN-Pa 708/1). Abbreviations: cs. cirrus sac; mg. Mehlis’ gland; oc. oocapt; od. oviduct; ov. ovary; t. testes; ud. utero duct; ut. uterus; vd. vas deferens; vf. vitelline follicle; vg. vagina; vod. ventral osmorregulatory duct. ▲loculi of the anterior region; * loculi of the central region; ● loculi of the posterior region" figureDoi="http://doi.org/10.5281/zenodo.3909207" httpUri="https://zenodo.org/record/3909207/files/figure.png" pageId="12" pageNumber="367">Fig. 6D</figureCitation>
|
||
). Two pairs of osmoregulatory ducts, 1 dorsal, 1 ventral.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5A6620DB5C7F4AFF7358B6CB95F8DF" ID-DOI="http://doi.org/10.5281/zenodo.3909207" ID-Zenodo-Dep="3909207" httpUri="https://zenodo.org/record/3909207/files/figure.png" pageId="9" pageNumber="364" startId="9.[151,250,1718,1743]" targetBox="[242,1371,204,1680]" targetPageId="9">
|
||
<paragraph id="8B9A36A8DB5C7F4AFF7358B6CB95F8DF" blockId="9.[151,1437,1718,1897]" pageId="9" pageNumber="364">
|
||
<emphasis id="B951EABADB5C7F4AFF7358B6C967F979" bold="true" box="[151,274,1718,1743]" pageId="9" pageNumber="364">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C254D2BDB5C7F4AFEF358B7CA7BF979" authority="Franzese & Ivanov, 2020" authorityName="Franzese & Ivanov" authorityYear="2020" box="[279,526,1719,1743]" class="Chondrichthyes" family="Arhynchobatidae" genus="Semiorbiseptum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="364" phylum="Chordata" rank="species" species="alfredoi" status="sp. nov.">
|
||
<emphasis id="B951EABADB5C7F4AFEF358B7CA7BF979" box="[279,526,1719,1743]" italics="true" pageId="9" pageNumber="364">Semiorbiseptum alfredoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951EABADB5C7F4AFDF058B7CA16F979" bold="true" box="[532,611,1719,1743]" pageId="9" pageNumber="364">
|
||
<taxonomicNameLabel id="A26257C1DB5C7F4AFDF058B7CA16F979" box="[532,611,1719,1743]" pageId="9" pageNumber="364" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from
|
||
<taxonomicName id="4C254D2BDB5C7F4AFD7B58B7CBF6F979" authorityName="McEachran. These" authorityYear="1983" box="[671,899,1719,1743]" class="Chondrichthyes" family="Arhynchobatidae" genus="Psammobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="364" phylum="Chordata" rank="species" species="normani">
|
||
<emphasis id="B951EABADB5C7F4AFD7B58B7CBF6F979" box="[671,899,1719,1743]" italics="true" pageId="9" pageNumber="364">Psammobatis normani</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B951EABADB5C7F4AFC6A58B6CBD2F979" bold="true" box="[910,935,1718,1743]" pageId="9" pageNumber="364">A.</emphasis>
|
||
entire worm (holotype MACN-Pa 706);
|
||
<emphasis id="B951EABADB5C7F4AFADB58B7CD23F979" bold="true" box="[1343,1366,1719,1743]" pageId="9" pageNumber="364">B.</emphasis>
|
||
scolex (holotype MACN-Pa 706);
|
||
<emphasis id="B951EABADB5C7F4AFE5558D4C9BFF95B" bold="true" box="[433,458,1748,1773]" pageId="9" pageNumber="364">C.</emphasis>
|
||
bothridium, distal surface;
|
||
<emphasis id="B951EABADB5C7F4AFD0158D4CA8BF95A" bold="true" box="[741,766,1748,1772]" pageId="9" pageNumber="364">D.</emphasis>
|
||
terminal mature proglottid (paratype MACN-Pa 708/1);
|
||
<emphasis id="B951EABADB5C7F4AFAA158D4CD28F95A" bold="true" box="[1349,1373,1748,1772]" pageId="9" pageNumber="364">E.</emphasis>
|
||
detail of terminal genitalia of a terminal mature proglottid (paratype MACN-Pa 708/1);
|
||
<emphasis id="B951EABADB5C7F4AFC2958F2CB95F8BC" bold="true" box="[973,992,1778,1802]" pageId="9" pageNumber="364">F.</emphasis>
|
||
detail of oötype region of a terminal mature proglottid (paratype MACN-Pa 708/1).
|
||
<emphasis id="B951EABADB5C7F4AFDCB590FCAC9F891" box="[559,700,1807,1831]" italics="true" pageId="9" pageNumber="364">Abbreviations</emphasis>
|
||
: cs. cirrus sac; mg. Mehlis’ gland; oc. oocapt; od. oviduct; ov. ovary; t. testes; ud. utero duct; ut. uterus; vd. vas deferens; vf. vitelline follicle; vg. vagina; vod. ventral osmorregulatory duct. ▲loculi of the anterior region; * loculi of the central region; ● loculi of the posterior region
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5A6620DB5F7F49FF735931C93AF800" ID-DOI="http://doi.org/10.5281/zenodo.3909209" ID-Zenodo-Dep="3909209" httpUri="https://zenodo.org/record/3909209/files/figure.png" pageId="10" pageNumber="365" startId="10.[151,250,1841,1866]" targetBox="[151,1436,181,1816]" targetPageId="10">
|
||
<paragraph id="8B9A36A8DB5F7F49FF735931C93AF800" blockId="10.[151,1436,1841,1974]" pageId="10" pageNumber="365">
|
||
<emphasis id="B951EABADB5F7F49FF735931C960F8FC" bold="true" box="[151,277,1841,1866]" pageId="10" pageNumber="365">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C254D2BDB5F7F49FEF95932CA6DF8FC" authority="Franzese & Ivanov, 2020" authorityName="Franzese & Ivanov" authorityYear="2020" box="[285,536,1842,1866]" class="Chondrichthyes" family="Arhynchobatidae" genus="Semiorbiseptum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="365" phylum="Chordata" rank="species" species="alfredoi" status="sp. nov.">
|
||
<emphasis id="B951EABADB5F7F49FEF95932CA6DF8FC" box="[285,536,1842,1866]" italics="true" pageId="10" pageNumber="365">Semiorbiseptum alfredoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951EABADB5F7F49FDC55932CA06F8FC" bold="true" box="[545,627,1842,1866]" pageId="10" pageNumber="365">
|
||
<taxonomicNameLabel id="A26257C1DB5F7F49FDC55932CA06F8FC" box="[545,627,1842,1866]" pageId="10" pageNumber="365" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from
|
||
<taxonomicName id="4C254D2BDB5F7F49FD525932CBE8F8FC" authorityName="McEachran. These" authorityYear="1983" box="[694,925,1842,1866]" class="Chondrichthyes" family="Arhynchobatidae" genus="Psammobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="365" phylum="Chordata" rank="species" species="normani">
|
||
<emphasis id="B951EABADB5F7F49FD525932CBE8F8FC" box="[694,925,1842,1866]" italics="true" pageId="10" pageNumber="365">Psammobatis normani</emphasis>
|
||
</taxonomicName>
|
||
, scanning electron micrographs.
|
||
<emphasis id="B951EABADB5F7F49FB115931CD7BF8FC" bold="true" box="[1269,1294,1841,1866]" pageId="10" pageNumber="365">A.</emphasis>
|
||
scolex, small letters indicate locations of details shown in figures 7C-G;
|
||
<emphasis id="B951EABADB5F7F49FD145956CB72F8D8" bold="true" box="[752,775,1878,1902]" pageId="10" pageNumber="365">B.</emphasis>
|
||
distal bothridial surface;
|
||
<emphasis id="B951EABADB5F7F49FBE85955CC51F8D8" bold="true" box="[1036,1060,1877,1902]" pageId="10" pageNumber="365">C.</emphasis>
|
||
detail of distal bothridial surface;
|
||
<emphasis id="B951EABADB5F7F49FA605956CDE9F8D8" bold="true" box="[1412,1436,1878,1902]" pageId="10" pageNumber="365">D.</emphasis>
|
||
botrhidial rim;
|
||
<emphasis id="B951EABADB5F7F49FECB597AC932F824" bold="true" box="[303,327,1914,1938]" pageId="10" pageNumber="365">E.</emphasis>
|
||
proximal bothridial surface;
|
||
<emphasis id="B951EABADB5F7F49FD88597ACA0AF824" bold="true" box="[620,639,1914,1938]" pageId="10" pageNumber="365">F.</emphasis>
|
||
surface of the cephalic peduncle;
|
||
<emphasis id="B951EABADB5F7F49FC325979CB85F824" bold="true" box="[982,1008,1913,1938]" pageId="10" pageNumber="365">G.</emphasis>
|
||
surface of a stalk;
|
||
<emphasis id="B951EABADB5F7F49FB49597ACCB2F824" bold="true" box="[1197,1223,1914,1938]" pageId="10" pageNumber="365">H.</emphasis>
|
||
surface of a terminal mature proglottid.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5A6620DB5E7F48FF735874C895F8BE" ID-DOI="http://doi.org/10.5281/zenodo.3909211" ID-Zenodo-Dep="3909211" httpUri="https://zenodo.org/record/3909211/files/figure.png" pageId="11" pageNumber="366" startId="11.[151,250,1652,1677]" targetBox="[151,1436,181,1626]" targetPageId="11">
|
||
<paragraph id="8B9A36A8DB5E7F48FF735874C895F8BE" blockId="11.[151,1437,1652,1800]" pageId="11" pageNumber="366">
|
||
<emphasis id="B951EABADB5E7F48FF735874C961F93B" bold="true" box="[151,276,1652,1677]" pageId="11" pageNumber="366">FIGURE 8.</emphasis>
|
||
<taxonomicName id="4C254D2BDB5E7F48FEFF5875CA60F93B" authority="Franzese & Ivanov, 2020" authorityName="Franzese & Ivanov" authorityYear="2020" box="[283,533,1653,1677]" class="Chondrichthyes" family="Arhynchobatidae" genus="Semiorbiseptum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="366" phylum="Chordata" rank="species" species="alfredoi" status="sp. nov.">
|
||
<emphasis id="B951EABADB5E7F48FEFF5875CA60F93B" box="[283,533,1653,1677]" italics="true" pageId="11" pageNumber="366">Semiorbiseptum alfredoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951EABADB5E7F48FDF85875CA1BF93B" bold="true" box="[540,622,1653,1677]" pageId="11" pageNumber="366">
|
||
<taxonomicNameLabel id="A26257C1DB5E7F48FDF85875CA1BF93B" box="[540,622,1653,1677]" pageId="11" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from
|
||
<taxonomicName id="4C254D2BDB5E7F48FD4A5875CBE1F93A" authorityName="McEachran. These" authorityYear="1983" box="[686,916,1653,1677]" class="Chondrichthyes" family="Arhynchobatidae" genus="Psammobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="366" phylum="Chordata" rank="species" species="normani">
|
||
<emphasis id="B951EABADB5E7F48FD4A5875CBE1F93A" box="[686,916,1653,1677]" italics="true" pageId="11" pageNumber="366">Psammobatis normani</emphasis>
|
||
</taxonomicName>
|
||
, septal musculature of bothridium.
|
||
<emphasis id="B951EABADB5E7F48FAE65874CD6EF93B" bold="true" box="[1282,1307,1652,1677]" pageId="11" pageNumber="366">A.</emphasis>
|
||
longitudinal section of bothridium (paratype MACN-Pa 707/4);
|
||
<emphasis id="B951EABADB5E7F48FD435892CACBF91C" bold="true" box="[679,702,1682,1706]" pageId="11" pageNumber="366">B.</emphasis>
|
||
light micrographs of bothridium (paratype MACN-Pa 707/3);
|
||
<emphasis id="B951EABADB5E7F48FAA75891CD2EF91C" bold="true" box="[1347,1371,1681,1706]" pageId="11" pageNumber="366">C.</emphasis>
|
||
transverse section of bothridium (paratype MACN-Pa 707/5).
|
||
<emphasis id="B951EABADB5E7F48FD0358AFCB01F971" box="[743,884,1711,1735]" italics="true" pageId="11" pageNumber="366">Abbreviations</emphasis>
|
||
: l. loculus; lmb. longitudinal muscle bundle; ls. Longitudinal septum; rm. radial muscle; ss. semicircular septum; tg. triangular gap; tmb. transverse muscle bundles; ts. transverse septum
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33F6523DB597F4FFF235C25CBB9FCA1" pageId="12" pageNumber="367" type="discussion">
|
||
<paragraph id="8B9A36A8DB597F4FFF235C25CBB9FCA1" blockId="12.[151,1437,152,791]" pageId="12" pageNumber="367">
|
||
<emphasis id="B951EABADB597F4FFF235C25C94EFD89" bold="true" box="[199,315,549,575]" pageId="12" pageNumber="367">Remarks.</emphasis>
|
||
<taxonomicName id="4C254D2BDB597F4FFEA15C25CA2EFD89" authority="Franzese & Ivanov, 2020" authorityName="Franzese & Ivanov" authorityYear="2020" box="[325,603,549,575]" class="Chondrichthyes" family="Arhynchobatidae" genus="Semiorbiseptum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="12" pageNumber="367" phylum="Chordata" rank="species" species="alfredoi" status="sp. nov.">
|
||
<emphasis id="B951EABADB597F4FFEA15C25CA2EFD89" box="[325,603,549,575]" italics="true" pageId="12" pageNumber="367">Semiorbiseptum alfredoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951EABADB597F4FFD805C25CAB4FD89" bold="true" box="[612,705,549,575]" pageId="12" pageNumber="367">
|
||
<taxonomicNameLabel id="A26257C1DB597F4FFD805C25CAB4FD89" box="[612,705,549,575]" pageId="12" pageNumber="367" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be mainly distinguished from
|
||
<emphasis id="B951EABADB597F4FFBB25C25CCA5FD89" box="[1110,1232,549,575]" italics="true" pageId="12" pageNumber="367">
|
||
Se.
|
||
<taxonomicName id="4C254D2BDB597F4FFB655C26CCA5FD89" authorityName="Franzese & Ivanov" authorityYear="2020" box="[1153,1232,550,575]" class="Chondrichthyes" family="Arhynchobatidae" genus="Semiorbiseptum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="12" pageNumber="367" phylum="Chordata" rank="species" species="mariae">mariae</taxonomicName>
|
||
</emphasis>
|
||
in the number of facial loculi, especially in the anterior region of bothridia. Whereas
|
||
<emphasis id="B951EABADB597F4FFC7B5C49CBCCFDD5" box="[927,953,585,611]" italics="true" pageId="12" pageNumber="367">Se</emphasis>
|
||
.
|
||
<taxonomicName id="4C254D2BDB597F4FFC2D5C49CC54FDD5" authority="Franzese & Ivanov, 2020" authorityName="Franzese & Ivanov" authorityYear="2020" box="[969,1057,585,611]" class="Chondrichthyes" family="Arhynchobatidae" genus="Semiorbiseptum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="12" pageNumber="367" phylum="Chordata" rank="species" species="alfredoi" status="sp. nov.">
|
||
<emphasis id="B951EABADB597F4FFC2D5C49CC54FDD5" box="[969,1057,585,611]" italics="true" pageId="12" pageNumber="367">alfredoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951EABADB597F4FFBCE5C49CCF0FDD5" bold="true" box="[1066,1157,585,611]" pageId="12" pageNumber="367">
|
||
<taxonomicNameLabel id="A26257C1DB597F4FFBCE5C49CCF0FDD5" box="[1066,1157,585,611]" pageId="12" pageNumber="367" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has bothridia divided in 13–16 loculi, with 4–5 loculi in the anterior region,
|
||
<emphasis id="B951EABADB597F4FFD0B5C6DCB1DFD31" box="[751,872,621,647]" italics="true" pageId="12" pageNumber="367">
|
||
Se.
|
||
<taxonomicName id="4C254D2BDB597F4FFCFD5C6ECB1DFD31" authorityName="Franzese & Ivanov" authorityYear="2020" box="[793,872,622,647]" class="Chondrichthyes" family="Arhynchobatidae" genus="Semiorbiseptum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="12" pageNumber="367" phylum="Chordata" rank="species" species="mariae">mariae</taxonomicName>
|
||
</emphasis>
|
||
has bothridia divided in 19–23 loculi, with 7–10 loculi in the anterior region (
|
||
<figureCitation id="131E2A2DDB597F4FFE325C91CA4DFD1D" box="[470,568,657,683]" captionStart="FIGURE 2" captionStartId="4.[151,250,1832,1857]" captionTargetBox="[164,1425,185,1799]" captionTargetId="figure@4.[158,1428,181,1806]" captionTargetPageId="4" captionText="FIGURE 2. Semiorbiseptum mariae sp. nov. from Psammobatis rudis. A. entire worm (holotype MACN-Pa 701); B. scolex (paratype MACN-Pa 704/1); C. bothridium, distal surface; D. terminal mature proglottid (paratype MACN-Pa 702/1); E. detail of the oötype region of a terminal mature proglottid (paratype MACN-Pa 702/1); F. detail of terminal genitalia of a terminal mature proglottid (paratype MACN-Pa 702/1); G. egg. Abbreviations: cs. cirrus sac; mg. Mehlis’ gland; oc. oocapt; od. oviduct; ov. ovary; t. testes; ut. uterus; vf. vitelline follicle; vg. vagina; vd. vas deferens; vod. ventral osmorregulatory duct. ▲loculi of the anterior region; * loculi of the central region; ● loculi of the posterior region" figureDoi="http://doi.org/10.5281/zenodo.3909199" httpUri="https://zenodo.org/record/3909199/files/figure.png" pageId="12" pageNumber="367">Figs. 2C</figureCitation>
|
||
,
|
||
<figureCitation id="131E2A2DDB597F4FFDA75C91CA12FD1A" box="[579,615,657,684]" captionStart="FIGURE 6" captionStartId="9.[151,250,1718,1743]" captionTargetBox="[242,1371,204,1680]" captionTargetId="figure@9.[200,1388,181,1693]" captionTargetPageId="9" captionText="FIGURE 6. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani. A. entire worm (holotype MACN-Pa 706); B. scolex (holotype MACN-Pa 706); C. bothridium, distal surface; D. terminal mature proglottid (paratype MACN-Pa 708/1); E. detail of terminal genitalia of a terminal mature proglottid (paratype MACN-Pa 708/1); F. detail of oötype region of a terminal mature proglottid (paratype MACN-Pa 708/1). Abbreviations: cs. cirrus sac; mg. Mehlis’ gland; oc. oocapt; od. oviduct; ov. ovary; t. testes; ud. utero duct; ut. uterus; vd. vas deferens; vf. vitelline follicle; vg. vagina; vod. ventral osmorregulatory duct. ▲loculi of the anterior region; * loculi of the central region; ● loculi of the posterior region" figureDoi="http://doi.org/10.5281/zenodo.3909207" httpUri="https://zenodo.org/record/3909207/files/figure.png" pageId="12" pageNumber="367">6C</figureCitation>
|
||
). The microthrix pattern also differed between the two species. Filitriches on the proximal bothridial surface of
|
||
<emphasis id="B951EABADB597F4FFDA05CB5CA2BFD79" box="[580,606,693,719]" italics="true" pageId="12" pageNumber="367">Se</emphasis>
|
||
.
|
||
<taxonomicName id="4C254D2BDB597F4FFD8B5CB6CACBFD79" authorityName="Franzese & Ivanov" authorityYear="2020" box="[623,702,694,719]" class="Chondrichthyes" family="Arhynchobatidae" genus="Semiorbiseptum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="12" pageNumber="367" phylum="Chordata" rank="species" species="mariae">
|
||
<emphasis id="B951EABADB597F4FFD8B5CB6CACBFD79" box="[623,702,694,719]" italics="true" pageId="12" pageNumber="367">mariae</emphasis>
|
||
</taxonomicName>
|
||
are longer than in
|
||
<emphasis id="B951EABADB597F4FFC7E5CB5CBC1FD79" box="[922,948,693,719]" italics="true" pageId="12" pageNumber="367">Se</emphasis>
|
||
.
|
||
<taxonomicName id="4C254D2BDB597F4FFC205CB5CC69FD79" authority="Franzese & Ivanov, 2020" authorityName="Franzese & Ivanov" authorityYear="2020" box="[964,1052,693,719]" class="Chondrichthyes" family="Arhynchobatidae" genus="Semiorbiseptum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="12" pageNumber="367" phylum="Chordata" rank="species" species="alfredoi" status="sp. nov.">
|
||
<emphasis id="B951EABADB597F4FFC205CB5CC69FD79" box="[964,1052,693,719]" italics="true" pageId="12" pageNumber="367">alfredoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951EABADB597F4FFBC05CB5CCF5FD79" bold="true" box="[1060,1152,693,719]" pageId="12" pageNumber="367">
|
||
<taxonomicNameLabel id="A26257C1DB597F4FFBC05CB5CCF5FD79" box="[1060,1152,693,719]" pageId="12" pageNumber="367" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Gladiate spinitriches are densely arranged along the entire proximal bothridial surface in
|
||
<emphasis id="B951EABADB597F4FFCBE5CD9CB01FD45" box="[858,884,729,755]" italics="true" pageId="12" pageNumber="367">Se</emphasis>
|
||
.
|
||
<taxonomicName id="4C254D2BDB597F4FFC665CDACBA4FD45" authorityName="Franzese & Ivanov" authorityYear="2020" box="[898,977,730,755]" class="Chondrichthyes" family="Arhynchobatidae" genus="Semiorbiseptum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="12" pageNumber="367" phylum="Chordata" rank="species" species="mariae">
|
||
<emphasis id="B951EABADB597F4FFC665CDACBA4FD45" box="[898,977,730,755]" italics="true" pageId="12" pageNumber="367">mariae</emphasis>
|
||
</taxonomicName>
|
||
, whereas they are scattered and restricted to the posterior region of bothridia in
|
||
<emphasis id="B951EABADB597F4FFDD95CFDCA22FCA1" box="[573,599,765,791]" italics="true" pageId="12" pageNumber="367">Se</emphasis>
|
||
.
|
||
<taxonomicName id="4C254D2BDB597F4FFD825CFDCACBFCA1" authority="Franzese & Ivanov, 2020" authorityName="Franzese & Ivanov" authorityYear="2020" box="[614,702,765,791]" class="Chondrichthyes" family="Arhynchobatidae" genus="Semiorbiseptum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="12" pageNumber="367" phylum="Chordata" rank="species" species="alfredoi" status="sp. nov.">
|
||
<emphasis id="B951EABADB597F4FFD825CFDCACBFCA1" box="[614,702,765,791]" italics="true" pageId="12" pageNumber="367">alfredoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951EABADB597F4FFD205CFDCB6BFCA1" bold="true" box="[708,798,765,791]" pageId="12" pageNumber="367">
|
||
<taxonomicNameLabel id="A26257C1DB597F4FFD205CFDCB6BFCA1" box="[708,798,765,791]" pageId="12" pageNumber="367" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="131E2A2DDB597F4FFCC95CFDCBE4FCA1" box="[813,913,765,791]" captionStart="FIGURE 3" captionStartId="5.[151,250,1841,1866]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure@5.[151,1436,181,1816]" captionTargetPageId="5" captionText="FIGURE 3. Semiorbiseptum mariae sp. nov. from Psammobatis rudis, scanning electron micrographs. A. scolex, small letters indicate locations of details shown in figures 3B-G; B. distal bothridial surface; C. detail of bothridial rim; D. proximal bothridial surface; E. surface of the apex of the scolex; F. surface of a stalk; G. surface of the cephalic peduncle; H. surface of a terminal mature proglottid." figureDoi="http://doi.org/10.5281/zenodo.3909201" httpUri="https://zenodo.org/record/3909201/files/figure.png" pageId="12" pageNumber="367">Figs. 3D</figureCitation>
|
||
,
|
||
<figureCitation id="131E2A2DDB597F4FFC795CFDCBB5FCA1" box="[925,960,765,791]" captionStart="FIGURE 7" captionStartId="10.[151,250,1841,1866]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure@10.[151,1436,181,1816]" captionTargetPageId="10" captionText="FIGURE 7. Semiorbiseptum alfredoi sp. nov. from Psammobatis normani, scanning electron micrographs. A. scolex, small letters indicate locations of details shown in figures 7C-G; B. distal bothridial surface; C. detail of distal bothridial surface; D. botrhidial rim; E. proximal bothridial surface; F. surface of the cephalic peduncle; G. surface of a stalk; H. surface of a terminal mature proglottid." figureDoi="http://doi.org/10.5281/zenodo.3909209" httpUri="https://zenodo.org/record/3909209/files/figure.png" pageId="12" pageNumber="367">7E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |