263 lines
34 KiB
XML
263 lines
34 KiB
XML
<document id="ED5E1BA485825056B0347396DD6A285D" ID-CLB-Dataset="306634" ID-DOI="10.1093/zoolinnean/zlae137" ID-GBIF-Dataset="b9ccfb40-9194-4228-abd2-acaf275f5749" ID-ISSN="0024-4082" ID-Zenodo-Dep="14516114" ID-ZooBank="9059AA2-5086-46AD-85C6-DBDA56CA72E0" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1734033959621" checkinUser="plazi" docAuthor="Simon-Pražák, Jan, Yamamoto, Shûhei, Lackner, Tomáš, Fikáček, Martin, Prokop, Jakub & Caterino, Michael S" docDate="2024" docId="038C87BE170D81714CEAFCFAFF7B7E6D" docLanguage="en" docName="zlae137.pdf" docOrigin="Zoological Journal of the Linnean Society (zlae 137) 202 (3)" docSource="https://doi.org/10.1093/zoolinnean/zlae137" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.9:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="9" docTitle="Cretomalus tibiodentatus Simon-Pražák, Yamamoto, Lackner, Fikáček, Prokop & Caterino, 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="8" masterDocId="FFB5FFC6170881764F58FFA0FFA07B0E" masterDocTitle="A bonanza of Cretaceous fossils provides insights into the evolution of antennal protection in clown beetles (Coleoptera: Histeridae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="6" updateTime="1734537818702" updateUser="ExternalLinkService">
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<mods:titleInfo id="B6238C5D884EAEC08A1082150625B2F6">
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<mods:title id="E0FC8C3F99989428408B0BF0DA6A5931">A bonanza of Cretaceous fossils provides insights into the evolution of antennal protection in clown beetles (Coleoptera: Histeridae)</mods:title>
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<mods:name id="CF39EAAFE1CEBA133C2176A5DA47086A" type="personal">
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<mods:roleTerm id="FB789849E28FE2EBA2B8C6ED294CF357">Author</mods:roleTerm>
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<mods:namePart id="E1EE70BA52BF94F183E567E07C84AED8">Simon-Pražák, Jan</mods:namePart>
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<mods:affiliation id="7466EFB430BE0E35A36142F32FD6832B">Charles University, Faculty of Science, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic & Museum of Eastern Bohemia in Hradec Králové, Eliščino nábřeží 465, 500 03 Hradec Králové 3, Czech Republic</mods:affiliation>
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<mods:nameIdentifier id="1AD62399C51E0CCE757617BB7C449B52" type="email">j.prazak@muzeumhk.cz</mods:nameIdentifier>
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<mods:roleTerm id="C980D60BD7B426C9A381A6350AFADA55">Author</mods:roleTerm>
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<mods:namePart id="7D5A07EEAC646D9D6F0B0764A7897C17">Yamamoto, Shûhei</mods:namePart>
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<mods:affiliation id="6EAD1A9A2C3F69E314D28566C9262492">The Hokkaido University Museum, Hokkaido University, Kita 10, Nishi 8, Kita-ku, Sapporo 060 - 0810, Japan</mods:affiliation>
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<mods:name id="ADCBB8AFCAB7277A785ED0C9B747A7D6" type="personal">
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<mods:roleTerm id="5377F849CEF83F295125F6BC0D900B92">Author</mods:roleTerm>
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<mods:namePart id="8A59CA83923638EA44490E1F9813CBBE">Lackner, Tomáš</mods:namePart>
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<mods:affiliation id="C2B0DCA4509AEACE2A3D9713383ED1D5">Department of Environmental Systems Science, ETH Zürich, Weinbergstrasse 56, 8092 Zürich, Switzerland</mods:affiliation>
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</mods:name>
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<mods:name id="7C1F59D1FA3CD11715ECB28DC7C399EB" type="personal">
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<mods:namePart id="9B88277DE70B8B46123B70713043ACA9">Fikáček, Martin</mods:namePart>
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<mods:affiliation id="0023EE99B40E9657EAE2329398999C2F">Department of Biological Sciences, National Sun Yat-sen University, No. 70, Lienhai Rd., Kaohsiung 80424, Taiwan & Department of Entomology, National Museum, Cirkusová 1740 CZ- 19300 Praha 9, Czech Republic</mods:affiliation>
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<mods:namePart id="4D7831FAFF6699CBCCAEAB3DD6A8B881">Prokop, Jakub</mods:namePart>
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<mods:affiliation id="7A9101F0DBFF869BE736388129BC02E6">Charles University, Faculty of Science, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic & Department of Plant and Environmental Sciences, Clemson University, Clemson, SC 29634, United States</mods:affiliation>
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<mods:namePart id="85624712CCD08BA4405AF1EEED902CD2">Caterino, Michael S</mods:namePart>
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<mods:typeOfResource id="D1B80602918698CAFCA56E7C143DDA12">text</mods:typeOfResource>
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<mods:title id="17E5E5F184E247359C915CBE021CEA48">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="42D80758540348229FAE6BA10F1B4A68">
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<mods:date id="97B3F07BFC0BD9FE88BEA7C5533C03AE">2024</mods:date>
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<mods:detail id="DE7E425DFC0406EB7D8B258AC24A8965" type="series">
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<mods:title id="367E56DA1293D588CBBDAB3BA9745CE9">zlae 137</mods:title>
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<mods:number id="5F40E662566A7EFE8B199425D77EC6AA">2024-11-20</mods:number>
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<mods:number id="01F6D4519E70967FC4BAEACD378752FC">202</mods:number>
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<mods:detail id="412A6F974AA4BB7C1ED0D33D096FB2E4" type="issue">
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<mods:number id="8867354D2727490AA07921CFB049C553">3</mods:number>
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<mods:start id="C76D91186979BBC2826A50A972835D29">1</mods:start>
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<mods:end id="2363262CA7889A4E07AF56255279C849">28</mods:end>
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<mods:location id="15C107747CA62D8496FBDEAD4FDFC835">
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<mods:url id="A378E658C48C205CBD5D3BB8DE196565">https://doi.org/10.1093/zoolinnean/zlae137</mods:url>
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<mods:classification id="F14DBB3D1461EBD4D1C92378CC344930">journal article</mods:classification>
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<mods:identifier id="7057A871A8D54CBC99843F276EDA3F91" type="CLB-Dataset">306634</mods:identifier>
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<mods:identifier id="DC0578BC6BFE9818983F41D24A2F6035" type="DOI">10.1093/zoolinnean/zlae137</mods:identifier>
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<mods:identifier id="9016640EABC344C7477910C65E6CC504" type="GBIF-Dataset">b9ccfb40-9194-4228-abd2-acaf275f5749</mods:identifier>
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<mods:identifier id="593BCC22AFD67A3090FDC205F46581A9" type="ISSN">0024-4082</mods:identifier>
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<mods:identifier id="C75752167C80D527D08A9874895CC4C3" type="Zenodo-Dep">14516114</mods:identifier>
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<mods:identifier id="3481FD44B5BFDD5650F8B0A943BCA8C9" type="ZooBank">9059AA2-5086-46AD-85C6-DBDA56CA72E0</mods:identifier>
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<treatment id="038C87BE170D81714CEAFCFAFF7B7E6D" ID-DOI="http://doi.org/10.5281/zenodo.14516116" ID-Zenodo-Dep="14516116" LSID="urn:lsid:plazi:treatment:038C87BE170D81714CEAFCFAFF7B7E6D" httpUri="http://treatment.plazi.org/id/038C87BE170D81714CEAFCFAFF7B7E6D" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6" scope_family="Histeridae" scope_order="Coleoptera" scope_subFamily="Dendrophilinae" scope_tribe="Paromalini">
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<subSubSection id="C33F6523170D81734CEAFCFAFA8B787B" box="[946,1323,858,885]" pageId="5" pageNumber="6" type="nomenclature">
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<paragraph id="8B9A36A8170D81734CEAFCFAFA8B787B" blockId="5.[946,1323,858,885]" box="[946,1323,858,885]" pageId="5" pageNumber="6">
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<heading id="D0D281C4170D81734CEAFCFAFA8B787B" bold="true" box="[946,1323,858,885]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="6" reason="2">
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<emphasis id="B951EABA170D81734CEAFCFAFA8B787B" bold="true" box="[946,1323,858,885]" pageId="5" pageNumber="6">
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<taxonomicName id="4C254D2B170D81734CEAFCFAFB6F787B" authorityName="Jan & ºimon-Pražák & Yamamoto & Lackner & Fikáček & Prokop & Michael & Caterino" authorityYear="2024" box="[946,1231,858,885]" class="Insecta" family="Histeridae" genus="Cretomalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="tibiodentatus" status="sp. nov.">
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<emphasis id="B951EABA170D81734CEAFCFAFB6F787B" bold="true" box="[946,1231,858,885]" italics="true" pageId="5" pageNumber="6">Cretomalus tibiodentatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26257C1170D81734B8CFCFBFA8B787B" box="[1236,1323,859,885]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33F6523170D81734B1FFC2CFB3678A8" box="[1095,1174,908,935]" pageId="5" pageNumber="6" type="description">
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<paragraph id="8B9A36A8170D81734B1FFC2CFB3678A8" blockId="5.[1095,1174,908,935]" box="[1095,1174,908,935]" pageId="5" pageNumber="6">
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||
<emphasis id="B951EABA170D81734B1FFC2CFB3678A8" bold="true" box="[1095,1174,908,935]" pageId="5" pageNumber="6">
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(
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<figureCitation id="131E2A2D170D81734B0AFC2CFB2B78A9" box="[1106,1163,908,935]" captionStart="Figure 5" captionStartId="6.[129,194,1370,1394]" captionTargetBox="[129,1473,144,1342]" captionTargetId="figure-262@6.[129,1473,144,1342]" captionTargetPageId="6" captionText="Figure 5. Cretomalus tibiodentatus gen. et sp. nov. A, dorsal view.B, ventral view; C, postmetacoxal area. D, postmesocoxal area. E, front leg and antenna. Abbreviations: fs, frontal stria; las, lateral stria on first abdominal ventrite; lms, lateral metaventral stria; pl, prosternal lobe; pms, prosternomesoventral stria; pmss, postmesocoxal stria; pmts, postmetacoxal stria." figureDoi="http://doi.org/10.5281/zenodo.14516126" httpUri="https://zenodo.org/record/14516126/files/figure.png" pageId="5" pageNumber="6">Fig. 5</figureCitation>
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)
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</emphasis>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C33F6523170D81734C72FC68FAF97F11" pageId="5" pageNumber="6" type="materials_examined">
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<paragraph id="8B9A36A8170D81734C72FC68FAF97F11" blockId="5.[810,1460,968,1055]" pageId="5" pageNumber="6">
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<emphasis id="B951EABA170D81734C72FC68FC2678EE" box="[810,902,968,992]" italics="true" pageId="5" pageNumber="6">Material:</emphasis>
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<materialsCitation id="3B4D3CF5170D81734CD5FC68FAF57F11" collectionCode="HUM" location="Kachin" pageId="5" pageNumber="6" specimenCode="SEHU-0000121215" specimenCount="1" stateProvince="Kachin">
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<collectingRegion id="49E1F84A170D81734CD5FC68FC7778EE" box="[909,983,968,992]" country="Myanmar" name="Kachin" pageId="5" pageNumber="6">Kachin</collectingRegion>
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amber (
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<emphasis id="B951EABA170D81734B69FC69FB9A78EE" box="[1073,1082,969,992]" italics="true" pageId="5" pageNumber="6">c</emphasis>
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. 99 Mya). Amber piece dimensions: 13 × 8 ×
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<quantity id="4CDD9B4D170D81734CD2FC47FC7878F1" box="[906,984,999,1023]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="6" unit="mm" value="1.5">1.5 mm</quantity>
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. Amber clear with several insect fragments and air bubbles. Deposited at
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<collectionCode id="ED34AE6D170D81734B74FBA7FBCB7F10" box="[1068,1131,1031,1054]" pageId="5" pageNumber="6">HUM</collectionCode>
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(
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<specimenCode id="DB839ED3170D81734B2FFBA7FAEE7F11" box="[1143,1358,1031,1055]" pageId="5" pageNumber="6">SEHU-0000121215</specimenCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33F6523170D81704C72FBE1FA9B7DE3" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
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<paragraph id="8B9A36A8170D81704C72FBE1FA9B7DE3" blockId="5.[810,1461,1088,1958]" lastBlockId="6.[825,1475,1498,1773]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
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<emphasis id="B951EABA170D81734C72FBE1FC187F56" box="[810,952,1089,1112]" italics="true" pageId="5" pageNumber="6">Measurements:</emphasis>
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Head width:
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<quantity id="4CDD9B4D170D81734B0BFBE0FB0A7F56" box="[1107,1194,1088,1112]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="5" pageNumber="6" unit="mm" value="0.32">0.32 mm</quantity>
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; width between anterior pronotal angles:
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<quantity id="4CDD9B4D170D81734C8AFBFFFB8A7F79" box="[978,1066,1119,1143]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="6" unit="mm" value="0.35">0.35 mm</quantity>
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; width between posterior pronotal angles:
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<quantity id="4CDD9B4D170D81734C39FBDFFC197F98" box="[865,953,1151,1175]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="6" unit="mm" value="0.55">0.55 mm</quantity>
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; pronotal length:
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<quantity id="4CDD9B4D170D81734B22FBDFFB657F98" box="[1146,1221,1151,1175]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="6" unit="mm" value="0.5">0.5 mm</quantity>
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; sutural elytral length:
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<quantity id="4CDD9B4D170D81734C72FB3EFC227FB8" box="[810,898,1182,1206]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="5" pageNumber="6" unit="mm" value="0.73">0.73 mm</quantity>
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; elytral width (in pair, across widest point):
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<quantity id="4CDD9B4D170D81734A62FB3EFA327FB8" box="[1338,1426,1182,1206]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="5" pageNumber="6" unit="mm" value="0.53">0.53 mm</quantity>
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.—
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<emphasis id="B951EABA170D81734C72FB1DFCC07FDB" bold="true" box="[810,864,1213,1237]" italics="true" pageId="5" pageNumber="6">Body</emphasis>
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elongate, rectangular, parallel-sided, and flattened. Cuticle sparsely punctuate dorsally, smooth ventrally. Colour black to chestnut brown.—
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<emphasis id="B951EABA170D81734CBFFB5CFB877E1A" bold="true" box="[999,1063,1276,1300]" italics="true" pageId="5" pageNumber="6">Head:</emphasis>
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Frons weakly convex, finely punctuate, punctures separated by three times their diameter. Frontoclypeal suture not evident; frontal stria thin, complete.Labrum short, trapezoidal, with several pairs of prominent setae.Mouthparts retracted. Eyes not visible from above.Scape short, expanded apically, pedicel simple, as long as antennomeres 3–4 combined. Antennomeres 3–8 wide and short (width: length ratio 1.5: 1), almost rectangular. Antennal club round, dorsoventrally flattened, annuli well visible.—
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<emphasis id="B951EABA170D81734CD1FA57FC7C7D01" bold="true" box="[905,988,1527,1551]" italics="true" pageId="5" pageNumber="6">Thorax:</emphasis>
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Pronotum as wide as long (width: length ratio 1: 1), median portion of disk flat, sides curved ventrad. Anterior emarginationdeep,anterioranglesacute,convergentanteriad.Basal pronotal margin rounded. Marginal pronotal stria complete along lateral and anterior margins. Pronotum punctuate, punctures separated by two to three times their diameter.Scutellum not visible (obscured by bubbles). Elytra long and narrow, punctuation density as in pronotum. Sutural area slightly depressed (possibly deformed postmortem), then elevated medially forming a keel. Dorsal elytral striation absent; elytral epipleuron obscured. Prosternal lobe deflexed, long, wide, rounded anteriorly. Prosternal keel with fine punctuation.Prosternalstriationindeterminableduetoairbubbles. Basal prosternal margin rounded, projecting posteriorly. Antennal cavities vaguely defined and widely open, located posterolaterally (covered by the front leg when retracted). Mesoventrite very short and wide (1: 3 ratio). Anterior margin curved posteriorly to fit the basal prosternal margin. Punctuation not apparent. Marginal mesoventral stria complete, continuous with lateral metaventral stria. Transverse mesoventral stria not visible. Metaventrite large, weakly concave, slightly elevated anterad metacoxae, without apparent punctuation. Lateral metaventral striae present and extending posterolaterad on at least anterior half. Postmesocoxal striae present and extending posteriorly. Sides of meso- and metaventrites (and abdominal ventrites) ascending laterad coxae, oblique.—
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<emphasis id="B951EABA170E81704FB3F8B4FE817C25" bold="true" box="[235,289,1812,1835]" italics="true" pageId="6" pageNumber="7">Legs:</emphasis>
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Profemur flattened and broad. Protibia broad and strongly curved, withsix to seven small, symmetrical teeth along the outer margin. Protarsal groove weakly developed. Small inner and curved median protibial spurs present. Protarsomere 5 as long as protarsomeres 1–4 combined. Protarsal claws short. Mesofemur narrower than profemur, mesotibia smooth, spur present. Terminal mesotarsomere as long as mesotarsomeres 3–4 combined. Hindleg equivalent to middle leg.—
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<emphasis id="B951EABA170E81704B04F9B8FB667D3E" bold="true" box="[1116,1222,1560,1584]" italics="true" pageId="6" pageNumber="7">Abdomen:</emphasis>
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First visible abdominal sternite large, intercoxal disc almost square-shaped, with very fine, sparse punctuation (punctures separated by at least 10 times their diameter). Lateral striae strongly impressed, complete, running posterolaterad. Postmetacoxal striae present, curved along metacoxae. Other sternites with identical punctuation pattern. Pygidium strongly convex, apex subtruncate, rugose.
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</paragraph>
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</subSubSection>
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|
||
<paragraph id="8B9A36A8170E81704FD9FAFAFC217EA4" blockId="6.[129,1453,1370,1450]" pageId="6" pageNumber="7">
|
||
<emphasis id="B951EABA170E81704FD9FAFAFF787E7C" bold="true" box="[129,216,1370,1394]" pageId="6" pageNumber="7">Figure 5.</emphasis>
|
||
<taxonomicName id="4C254D2B170E81704F85FAFAFE1E7E7C" authorityName="Jan & ºimon-Pražák & Yamamoto & Lackner & Fikáček & Prokop & Michael & Caterino" authorityYear="2024" box="[221,446,1370,1394]" class="Insecta" family="Histeridae" genus="Cretomalus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tibiodentatus" status="gen. et sp. nov.">
|
||
<emphasis id="B951EABA170E81704F85FAFAFE1E7E7C" box="[221,446,1370,1394]" italics="true" pageId="6" pageNumber="7">Cretomalus tibiodentatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26257C1170E81704E9BFAFBFDEC7E7C" box="[451,588,1371,1395]" pageId="6" pageNumber="7" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
A, dorsal view. B, ventral view; C, postmetacoxal area. D, postmesocoxal area. E, front leg and antenna. Abbreviations: fs, frontal stria; las, lateral stria on first abdominal ventrite; lms, lateral metaventral stria; pl, prosternal lobe; pms, prosternomesoventral stria; pmss, postmesocoxal stria; pmts, postmetacoxal stria.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33F6523170E81714C61F8B2FE487950" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="discussion">
|
||
<paragraph id="8B9A36A8170E81714C61F8B2FE487950" blockId="6.[825,1474,1809,1958]" lastBlockId="7.[113,763,144,607]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
|
||
<emphasis id="B951EABA170E81704C61F8B2FB807C27" box="[825,1056,1810,1833]" italics="true" pageId="6" pageNumber="7">Taxonomic assignment:</emphasis>
|
||
We place this species in the
|
||
<taxonomicName id="4C254D2B170E81704A0BF8B1FA627C27" authorityName="Reitter" authorityYear="1909" box="[1363,1474,1809,1833]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="tribe" tribe="Paromalini">Paromalini</taxonomicName>
|
||
tribe based on the combination of the following characters (
|
||
<bibRefCitation id="EFB44B59170E81704C1BF8F1FC1E7C66" author="Ohara M" box="[835,958,1872,1900]" pageId="6" pageNumber="7" pagination="1 - 283" refId="ref18649" refString="Ohara M. A Revision of the ºuperfamily Histeroidea of Japan [Coleoptera]. Insecta Matsumurana 1994; 51: 1 - 283." type="journal article" year="1994">Ôhara 1994</bibRefCitation>
|
||
): broad, weakly convex frons (
|
||
<figureCitation id="131E2A2D170E81704BA6F8F0FAE97C66" box="[1278,1353,1872,1896]" captionStart="Figure 5" captionStartId="6.[129,194,1370,1394]" captionTargetBox="[129,1473,144,1342]" captionTargetId="figure-262@6.[129,1473,144,1342]" captionTargetPageId="6" captionText="Figure 5. Cretomalus tibiodentatus gen. et sp. nov. A, dorsal view.B, ventral view; C, postmetacoxal area. D, postmesocoxal area. E, front leg and antenna. Abbreviations: fs, frontal stria; las, lateral stria on first abdominal ventrite; lms, lateral metaventral stria; pl, prosternal lobe; pms, prosternomesoventral stria; pmss, postmesocoxal stria; pmts, postmetacoxal stria." figureDoi="http://doi.org/10.5281/zenodo.14516126" httpUri="https://zenodo.org/record/14516126/files/figure.png" pageId="6" pageNumber="7">Fig. 5A</figureCitation>
|
||
), short and wide labrum with several pairs of prominent setae, short and wide prosternal lobe, vaguely defined antennal cavities located at prosternal hypomeron, basal prosternal margin rounded and projecting posteriorly, wide, curving protibia with a laterally directed median apical spur. The position of this genus in
|
||
<taxonomicName id="4C254D2B170F81714F29FF4EFF407A08" authorityName="Reitter" authorityYear="1909" box="[113,224,238,262]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="tribe" tribe="Paromalini">Paromalini</taxonomicName>
|
||
is also supported by all phylogenetic analyses performed in this study. Within extant
|
||
<taxonomicName id="4C254D2B170F81714EA9FEADFDC27A2B" authorityName="Reitter" authorityYear="1909" box="[497,610,269,293]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="tribe" tribe="Paromalini">Paromalini</taxonomicName>
|
||
, possession of dorsal elytral striae in several genera (e.g.
|
||
<taxonomicName id="4C254D2B170F81714D4DFE8DFD6F7A4B" authority="Marseul" authorityName="Marseul" authorityYear="1855" box="[533,719,301,325]" class="Insecta" family="Histeridae" genus="Carcinops" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EABA170F81714D4DFE8DFDD77A4A" box="[533,631,301,324]" italics="true" pageId="7" pageNumber="8">Carcinops</emphasis>
|
||
Marseul
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C254D2B170F81714F29FEECFE9A7A6A" authority="Bickhardt" authorityName="Bickhardt" authorityYear="1921" box="[113,314,332,356]" class="Insecta" family="Histeridae" genus="Diplostix" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EABA170F81714F29FEECFF6A7A6A" box="[113,202,332,356]" italics="true" pageId="7" pageNumber="8">Diplostix</emphasis>
|
||
Bickhardt
|
||
</taxonomicName>
|
||
) is probably plesiomorphic, while their absence is apomorphic in others (
|
||
<taxonomicName id="4C254D2B170F81714EF3FECBFDD47A8D" authority="Erichson" authorityName="Erichson" authorityYear="1834" box="[427,628,363,387]" class="Insecta" family="Histeridae" genus="Paromalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EABA170F81714EF3FECBFDB37A8D" box="[427,531,363,387]" italics="true" pageId="7" pageNumber="8">Paromalus</emphasis>
|
||
Erichson
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C254D2B170F81714D23FECBFF6D7AAD" authority="Cooman" authorityName="Cooman" authorityYear="1948" class="Insecta" family="Histeridae" genus="Platylomalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EABA170F81714D23FECBFD5A7A8D" box="[635,762,363,387]" italics="true" pageId="7" pageNumber="8">Platylomalus</emphasis>
|
||
Cooman
|
||
</taxonomicName>
|
||
) (
|
||
<bibRefCitation id="EFB44B59170F81714FB2FE2BFE3D7AAD" author="Zhou Y & Ren D" box="[234,413,395,419]" pageId="7" pageNumber="8" pagination="521 - 39" refId="ref19290" refString="Zhou Y, Caterino Mº, Ren D et al. Phylogeny and evolution of Mesozoic and extant lineages of Histeridae (Coleoptera), with discovery of a new subfamily Antigracilinae from the Lower Cretaceous. Cladistics 2020; 36: 521 - 39. hups: // doi. org / 10.1111 / cla. 12418" type="journal article" year="2020">
|
||
Zhou
|
||
<emphasis id="B951EABA170F81714E74FE2BFEC07AAD" box="[300,352,395,419]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C254D2B170F81714EE0FE2BFD107AAD" authorityName="Jan & ºimon-Pražák & Yamamoto & Lackner & Fikáček & Prokop & Michael & Caterino" authorityYear="2024" box="[440,688,395,419]" class="Insecta" family="Histeridae" genus="Cretomalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tibiodentatus">
|
||
<emphasis id="B951EABA170F81714EE0FE2BFD107AAD" box="[440,688,395,419]" italics="true" pageId="7" pageNumber="8">Cretomalus tibiodentatus</emphasis>
|
||
</taxonomicName>
|
||
would appear to be more closely related to the latter, particularly to
|
||
<taxonomicName id="4C254D2B170F81714F29FE69FF797AEF" authorityName="Erichson" authorityYear="1834" box="[113,217,457,481]" class="Insecta" family="Histeridae" genus="Paromalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EABA170F81714F29FE69FF797AEF" box="[113,217,457,481]" italics="true" pageId="7" pageNumber="8">Paromalus</emphasis>
|
||
</taxonomicName>
|
||
, with which it shares several characters: absence of prosternal striation and complete frontal stria (
|
||
<figureCitation id="131E2A2D170F81714D0AFE49FD3C790E" box="[594,668,488,513]" captionStart="Figure 6" captionStartId="8.[129,194,1677,1701]" captionTargetBox="[129,1473,144,1650]" captionTargetId="figure-114@8.[129,1473,144,1650]" captionTargetPageId="8" captionText="Figure 6. Anapleus punctulatus sp.nov.. A, dorsal view.B, elytral epipleuron. C, ventral view.D, frontal view.Abbreviations:ees, elytral epipleural striae; lps, lateral pronotal stria; mes, marginal elytral stria; mps, marginal prosternal stria; ms, metaventral stria." figureDoi="http://doi.org/10.5281/zenodo.14516128" httpUri="https://zenodo.org/record/14516128/files/figure.png" pageId="7" pageNumber="8">Fig. 6A</figureCitation>
|
||
) (
|
||
<bibRefCitation id="EFB44B59170F81714DE0FE49FEAE792E" author="Zhang YJ & Zhou HZ" pageId="7" pageNumber="8" pagination="1 - 40" refId="ref19205" refString="Zhang YJ, Zhou HZ. Taxonomy of the tribe Paromalini Reiuer (Coleoptera: Histeridae, Dendrophilinae) from China. Zootaxa 2007; 1544: 1 - 40. hups: // doi. org / 10.11646 / zootaxa. 1544.1.1" type="journal article" year="2007">Zhang and Zhou 2007</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C254D2B170F81714E7CFDA8FDB6792E" authorityName="Jan & ºimon-Pražák & Yamamoto & Lackner & Fikáček & Prokop & Michael & Caterino" authorityYear="2024" box="[292,534,520,544]" class="Insecta" family="Histeridae" genus="Cretomalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tibiodentatus">
|
||
<emphasis id="B951EABA170F81714E7CFDA8FDB6792E" box="[292,534,520,544]" italics="true" pageId="7" pageNumber="8">Cretomalus tibiodentatus</emphasis>
|
||
</taxonomicName>
|
||
is significantly smaller than all
|
||
<taxonomicName id="4C254D2B170F81714F90FD87FE287931" box="[200,392,551,575]" class="Insecta" family="Histeridae" genus="Paromalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B951EABA170F81714F90FD87FE907931" box="[200,304,551,575]" italics="true" pageId="7" pageNumber="8">Paromalus</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
The serrated protibial margin (
|
||
<figureCitation id="131E2A2D170F81714D8CFD87FF2D7950" captionStart="Figure 5" captionStartId="6.[129,194,1370,1394]" captionTargetBox="[129,1473,144,1342]" captionTargetId="figure-262@6.[129,1473,144,1342]" captionTargetPageId="6" captionText="Figure 5. Cretomalus tibiodentatus gen. et sp. nov. A, dorsal view.B, ventral view; C, postmetacoxal area. D, postmesocoxal area. E, front leg and antenna. Abbreviations: fs, frontal stria; las, lateral stria on first abdominal ventrite; lms, lateral metaventral stria; pl, prosternal lobe; pms, prosternomesoventral stria; pmss, postmesocoxal stria; pmts, postmetacoxal stria." figureDoi="http://doi.org/10.5281/zenodo.14516126" httpUri="https://zenodo.org/record/14516126/files/figure.png" pageId="7" pageNumber="8">Fig. 5E</figureCitation>
|
||
) is unique among the entire tribe.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33F6523170F81714F29FD21FF697E05" pageId="7" pageNumber="8" type="diagnosis">
|
||
<paragraph id="8B9A36A8170F81714F29FD21FF697E05" blockId="7.[113,764,640,1291]" pageId="7" pageNumber="8">
|
||
<emphasis id="B951EABA170F81714F29FD21FF777996" box="[113,215,641,664]" italics="true" pageId="7" pageNumber="8">Diagnosis:</emphasis>
|
||
Frons weakly convex and finely punctuate, punctures separated by three times their diameter. Pronotum relatively long, punctuated with punctures separated by two to three times their diameter. Elytra without striation, punctuation with the same pattern as in pronotum. Prosternal keel with fine punctuation. Marginal mesoventral stria complete, continuous with lateral metaventral stria. Transverse mesoventral stria not visible. Lateral metaventral striae present, extending posterolaterad on at least anterior half. Postmesocoxal striae present, running posteriorly. Protibia wide and strongly curved, with six to seven small, symmetrical teeth along the outer margin. Protarsal groove weakly developed. Small inner and curved median protibial spurs present. Similarly shaped
|
||
<taxonomicName id="4C254D2B170F81714E9DFC58FD907F1E" box="[453,560,1016,1040]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
|
||
found in Burmese amber are
|
||
<taxonomicName id="4C254D2B170F81714FB2FBB7FE687F21" authorityName="Simon Prazak & Lackner" authorityYear="2023" box="[234,456,1047,1071]" class="Insecta" family="Histeridae" genus="Platycretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="muscularis">
|
||
<emphasis id="B951EABA170F81714FB2FBB7FE687F21" box="[234,456,1047,1071]" italics="true" pageId="7" pageNumber="8">Platycretus muscularis</emphasis>
|
||
</taxonomicName>
|
||
Simon-Pražák and
|
||
<bibRefCitation id="EFB44B59170F81714DFAFBB7FF077F41" author="Lackner T & Zhang YM & Kindler C" pageId="7" pageNumber="8" pagination="48 - 71" refId="ref18368" refString="Lackner T, Zhang YM, Kindler C et al. ºaprininae: phylogeny, biogeography and a new classification of the subfamily (Coleoptera: Histeridae). Systematic Entomology 2023; 49: 48 - 71. hups: // doi. org / 10.1111 / syen. 12606" type="journal article" year="2023">Lackner, 2023</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C254D2B170F81714FECFB96FE317F40" authorityName="Simon Prazak & Lackner" authorityYear="2023" box="[180,401,1078,1102]" class="Insecta" family="Histeridae" genus="Olexum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="complanatum">
|
||
<emphasis id="B951EABA170F81714FECFB96FE317F40" box="[180,401,1078,1102]" italics="true" pageId="7" pageNumber="8">Olexum complanatum</emphasis>
|
||
</taxonomicName>
|
||
Simon-Pražák and
|
||
<bibRefCitation id="EFB44B59170F81714D38FB96FD567F41" author="Lackner T & Zhang YM & Kindler C" box="[608,758,1078,1103]" pageId="7" pageNumber="8" pagination="48 - 71" refId="ref18368" refString="Lackner T, Zhang YM, Kindler C et al. ºaprininae: phylogeny, biogeography and a new classification of the subfamily (Coleoptera: Histeridae). Systematic Entomology 2023; 49: 48 - 71. hups: // doi. org / 10.1111 / syen. 12606" type="journal article" year="2023">Lackner, 2023</bibRefCitation>
|
||
, and
|
||
<taxonomicName id="4C254D2B170F81714FF9FBF6FE567F60" authority="Caterino, 2021" authorityName="Caterino" authorityYear="2021" box="[161,502,1110,1134]" class="Insecta" family="Histeridae" genus="Druantia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="aeterna">
|
||
<emphasis id="B951EABA170F81714FF9FBF6FEED7F63" box="[161,333,1110,1133]" italics="true" pageId="7" pageNumber="8">Druantia aeterna</emphasis>
|
||
<bibRefCitation id="EFB44B59170F81714E0FFBF6FE567F60" author="Caterino" box="[343,502,1110,1134]" pageId="7" pageNumber="8" pagination="211 - 21" refId="ref17393" refString="Caterino Mº. New fossil histerid species from Cretaceous Burmese amber (Coleoptera: Histeridae). Ne Coleopterists Bulletin 2021; 75: 211 - 21." type="journal article" year="2021">Caterino, 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C254D2B170F81714D5BFBF6FD5A7F60" authorityName="Jan & ºimon-Pražák & Yamamoto & Lackner & Fikáček & Prokop & Michael & Caterino" authorityYear="2024" box="[515,762,1110,1134]" class="Insecta" family="Histeridae" genus="Cretomalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tibiodentatus">
|
||
<emphasis id="B951EABA170F81714D5BFBF6FD5A7F60" box="[515,762,1110,1134]" italics="true" pageId="7" pageNumber="8">Cretomalus tibiodentatus</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from all of them by combination of following characters: long and narrow body (length: width 5: 2), shape of the prosterno-mesoventral suture (with a curved anteriorly pointed projection) and the state of regularly toothed protibia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33F6523170F81714F29FA8CFF7B7E6D" pageId="7" pageNumber="8" type="etymology">
|
||
<paragraph id="8B9A36A8170F81714F29FA8CFF7B7E6D" blockId="7.[113,763,1324,1379]" pageId="7" pageNumber="8">
|
||
<emphasis id="B951EABA170F81714F29FA8CFF417E4A" box="[113,225,1324,1348]" italics="true" pageId="7" pageNumber="8">Etymology:</emphasis>
|
||
The specific name refers to the unique protibial dentation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |