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<document id="0D0A8B650AD9AF7861DC29D46E1FEEAE" ID-CLB-Dataset="306634" ID-DOI="10.1093/zoolinnean/zlae137" ID-GBIF-Dataset="b9ccfb40-9194-4228-abd2-acaf275f5749" ID-ISSN="0024-4082" ID-Zenodo-Dep="14516114" ID-ZooBank="9059AA2-5086-46AD-85C6-DBDA56CA72E0" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1734033959621" checkinUser="plazi" docAuthor="Simon-Pražák, Jan, Yamamoto, Shûhei, Lackner, Tomáš, Fikáček, Martin, Prokop, Jakub &amp; Caterino, Michael S" docDate="2024" docId="038C87BE170781794E4DFC45FF4B7C1F" docLanguage="en" docName="zlae137.pdf" docOrigin="Zoological Journal of the Linnean Society (zlae 137) 202 (3)" docSource="https://doi.org/10.1093/zoolinnean/zlae137" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.9:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="9" docTitle="Phasmister planatus Simon-Pražák, Yamamoto, Lackner, Fikáček, Prokop &amp; Caterino, 2024, sp. nov." docType="treatment" docVersion="5" lastPageNumber="16" masterDocId="FFB5FFC6170881764F58FFA0FFA07B0E" masterDocTitle="A bonanza of Cretaceous fossils provides insights into the evolution of antennal protection in clown beetles (Coleoptera: Histeridae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="16" updateTime="1734537818702" updateUser="ExternalLinkService">
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<mods:title id="345A5B60DD3166B5EBF8E05CF95AE754">A bonanza of Cretaceous fossils provides insights into the evolution of antennal protection in clown beetles (Coleoptera: Histeridae)</mods:title>
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<mods:namePart id="F60FC88C1B4DD2EBA3886EB249374C46">Simon-Pražák, Jan</mods:namePart>
<mods:affiliation id="1BFF5CC2470428A686E9CB0E2FC987E1">Charles University, Faculty of Science, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic &amp; Museum of Eastern Bohemia in Hradec Králové, Eliščino nábřeží 465, 500 03 Hradec Králové 3, Czech Republic</mods:affiliation>
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<mods:namePart id="0EA2789E876CBBFD61F4462601CD4859">Yamamoto, Shûhei</mods:namePart>
<mods:affiliation id="FB1AE21E331672CED821D84007252940">The Hokkaido University Museum, Hokkaido University, Kita 10, Nishi 8, Kita-ku, Sapporo 060 - 0810, Japan</mods:affiliation>
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<mods:namePart id="DEF3117164B660536BAFAB9971322511">Lackner, Tomáš</mods:namePart>
<mods:affiliation id="1D144D6A71BA07C920E5F8E1015B50BD">Department of Environmental Systems Science, ETH Zürich, Weinbergstrasse 56, 8092 Zürich, Switzerland</mods:affiliation>
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<mods:namePart id="B937F49BD57ED5CF8B39ACE1D8067C13">Fikáček, Martin</mods:namePart>
<mods:affiliation id="B7C36330F509B90F6067723269AE7806">Department of Biological Sciences, National Sun Yat-sen University, No. 70, Lienhai Rd., Kaohsiung 80424, Taiwan &amp; Department of Entomology, National Museum, Cirkusová 1740 CZ- 19300 Praha 9, Czech Republic</mods:affiliation>
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<mods:namePart id="86550B2B95897A575C44A67C36BD4EC5">Prokop, Jakub</mods:namePart>
<mods:affiliation id="A5013950958478161D753E002940B21E">Charles University, Faculty of Science, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic &amp; Department of Plant and Environmental Sciences, Clemson University, Clemson, SC 29634, United States</mods:affiliation>
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<mods:namePart id="9470F6DAEB8A4C967DD306C7E63CEB25">Caterino, Michael S</mods:namePart>
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<mods:title id="1FB3E1443CBE5CA248607F8D2DB81F61">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="CA911577D00EE5B8F49896DB97DA40F7">2024</mods:date>
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<mods:title id="1AB0CC6EDCC9DE8879C87E7A3E7A5B4E">zlae 137</mods:title>
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<mods:number id="426FF16BE02BE03028AEC61F7373E3DB">2024-11-20</mods:number>
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<treatment id="038C87BE170781794E4DFC45FF4B7C1F" ID-DOI="http://doi.org/10.5281/zenodo.14424567" ID-Zenodo-Dep="14424567" LSID="urn:lsid:plazi:treatment:038C87BE170781794E4DFC45FF4B7C1F" httpUri="http://treatment.plazi.org/id/038C87BE170781794E4DFC45FF4B7C1F" lastPageNumber="16" pageId="15" pageNumber="16" scope_family="Histeridae" scope_order="Coleoptera" scope_subFamily="Onthophilinae">
<subSubSection id="C33F6523170781794E4DFC45FDF77F0E" box="[277,599,997,1024]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8B9A36A8170781794E4DFC45FDF77F0E" blockId="15.[277,599,997,1024]" box="[277,599,997,1024]" pageId="15" pageNumber="16">
<heading id="D0D281C4170781794E4DFC45FDF77F0E" bold="true" box="[277,599,997,1024]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="16" reason="2">
<emphasis id="B951EABA170781794E4DFC45FDF77F0E" bold="true" box="[277,599,997,1024]" pageId="15" pageNumber="16">
<taxonomicName id="4C254D2B170781794E4DFC45FE5A7F0E" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[277,506,997,1024]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="planatus" status="sp. nov.">
<emphasis id="B951EABA170781794E4DFC45FE5A7F0E" bold="true" box="[277,506,997,1024]" italics="true" pageId="15" pageNumber="16">Phasmister planatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C1170781794D58FC46FDF77F0E" box="[512,599,998,1024]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F6523170781794E37FBB7FE5D7F3F" box="[367,509,1047,1073]" pageId="15" pageNumber="16" type="description">
<paragraph id="8B9A36A8170781794E37FBB7FE5D7F3F" blockId="15.[367,509,1047,1073]" box="[367,509,1047,1073]" pageId="15" pageNumber="16">
<emphasis id="B951EABA170781794E37FBB7FE5D7F3F" bold="true" box="[367,509,1047,1073]" pageId="15" pageNumber="16">
(
<figureCitation id="131E2A2D170781794E21FBB7FE527F3F" box="[377,498,1047,1073]" captionStart="Figure 10" captionStartId="14.[129,194,1835,1859]" captionTargetBox="[141,1461,143,1807]" captionTargetId="figure-60@14.[141,1461,143,1807]" captionTargetPageId="14" captionText="Figure 10. Phasmister planatus sp. nov. (AC) and Phasmister kraliceki sp. nov.(D, E)." figureDoi="http://doi.org/10.5281/zenodo.14516138" httpUri="https://zenodo.org/record/14516138/files/figure.png" pageId="15" pageNumber="16">Fig. 10AC</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33F6523170781794F29FBF8FD027FA1" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8B9A36A8170781794F29FBF8FD027FA1" blockId="15.[113,763,1112,1199]" pageId="15" pageNumber="16">
<emphasis id="B951EABA170781794F29FBF8FF6D7F7E" box="[113,205,1112,1136]" italics="true" pageId="15" pageNumber="16">Material:</emphasis>
<materialsCitation id="3B4D3CF5170781794F8DFBF8FD3E7FA1" collectionCode="HUM" location="Kachin" pageId="15" pageNumber="16" specimenCode="SEHU-0000121216" specimenCount="1" stateProvince="Kachin">
<collectingRegion id="49E1F84A170781794F8DFBF8FEBF7F7E" box="[213,287,1112,1136]" country="Myanmar" name="Kachin" pageId="15" pageNumber="16">Kachin</collectingRegion>
amber (
<emphasis id="B951EABA170781794E20FBF9FE217F7E" box="[376,385,1113,1136]" italics="true" pageId="15" pageNumber="16">c</emphasis>
. 99 Mya). Amber piece dimensions: 9 × 5 ×
<quantity id="4CDD9B4D170781794F9EFBD7FEB37F81" box="[198,275,1143,1167]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="16" unit="mm" value="1.5">1.5 mm</quantity>
. Amber clear with several insect fragments and air bubbles. Deposited at
<collectionCode id="ED34AE6D170781794E2DFB37FE147FA0" box="[373,436,1175,1198]" pageId="15" pageNumber="16">HUM</collectionCode>
(
<specimenCode id="DB839ED3170781794E98FB37FD377FA1" box="[448,663,1175,1199]" pageId="15" pageNumber="16">SEHU-0000121216</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F6523170781794F29FB71FD707E4B" pageId="15" pageNumber="16" type="description">
<paragraph id="8B9A36A8170781794F29FB71FD707E4B" blockId="15.[113,763,1232,1350]" pageId="15" pageNumber="16">
<emphasis id="B951EABA170781794F29FB71FF487FE6" box="[113,232,1233,1256]" italics="true" pageId="15" pageNumber="16">Dimensions.</emphasis>
Head width:
<quantity id="4CDD9B4D170781794ED3FB70FE467FE9" box="[395,486,1232,1256]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="15" pageNumber="16" unit="mm" value="0.48">0.48 mm</quantity>
; width between anterior pronotal angles:
<quantity id="4CDD9B4D170781794E45FB4FFECB7E09" box="[285,363,1263,1287]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="16" unit="mm" value="0.6">0.6 mm</quantity>
; width between posterior pronotal angles:
<quantity id="4CDD9B4D170781794FF1FAAFFF577E28" box="[169,247,1295,1319]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="15" pageNumber="16" unit="mm" value="1.1">1.1 mm</quantity>
; pronotal length:
<quantity id="4CDD9B4D170781794EE2FAAFFDA87E28" box="[442,520,1295,1319]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="16" unit="mm" value="0.6">0.6 mm</quantity>
; sutural elytral length:
<quantity id="4CDD9B4D170781794F29FA8EFF107E4B" box="[113,176,1326,1350]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="mm" value="1.0">1 mm</quantity>
; elytral width (in pair, across widest point):
<quantity id="4CDD9B4D170781794D37FA8EFD6D7E4B" box="[623,717,1326,1350]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="15" pageNumber="16" unit="mm" value="1.25">1.25 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F6523170781794F29FAC8FD537DB7" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="8B9A36A8170781794F29FAC8FD537DB7" blockId="15.[113,763,1383,1721]" pageId="15" pageNumber="16">
<emphasis id="B951EABA170781794F29FAC8FEEF7E71" box="[113,335,1383,1407]" italics="true" pageId="15" pageNumber="16">Diagnostic description.</emphasis>
The state of pronotal carinae is almost identical to
<taxonomicName id="4C254D2B170781794FB3FA27FD587E91" authority="Caterino and Yamamoto, 2023" authorityName="Caterino and Yamamoto" authorityYear="2023" box="[235,760,1415,1439]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B951EABA170781794FB3FA27FE157E91" box="[235,437,1415,1439]" italics="true" pageId="15" pageNumber="16">Phasmister parallelus</emphasis>
Caterino and Yamamoto, 2023
</taxonomicName>
. However,
<taxonomicName id="4C254D2B170781794F83FA07FEE77EB0" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[219,327,1446,1470]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="planatus">
<emphasis id="B951EABA170781794F83FA07FEE77EB0" box="[219,327,1446,1470]" italics="true" pageId="15" pageNumber="16">P. planatus</emphasis>
</taxonomicName>
bears a significantly more flattened body shape and several differential characteristics are linked with that— elytra are straight and flat (whereas in
<taxonomicName id="4C254D2B170781794EB3FA45FDFB7EF3" authorityName="Caterino and Yamamoto" authorityYear="2023" box="[491,603,1509,1533]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B951EABA170781794EB3FA45FDFB7EF3" box="[491,603,1509,1533]" italics="true" pageId="15" pageNumber="16">P. parallelus</emphasis>
</taxonomicName>
they are convex and curved downwards apically), spaces between first dorsal carina and upper epipleural carina and between the upper and lower epipleural carinae are narrower than in
<taxonomicName id="4C254D2B170781794D7DF9E3FD347D55" authorityName="Caterino and Yamamoto" authorityYear="2023" box="[549,660,1603,1627]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B951EABA170781794D7DF9E3FD347D55" box="[549,660,1603,1627]" italics="true" pageId="15" pageNumber="16">P.parallelus</emphasis>
</taxonomicName>
. Ventrally,
<taxonomicName id="4C254D2B170781794F29F9C3FF437D74" authorityName="Caterino and Yamamoto" authorityYear="2023" box="[113,227,1634,1658]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B951EABA170781794F29F9C3FF437D74" box="[113,227,1634,1658]" italics="true" pageId="15" pageNumber="16">P. parallelus</emphasis>
</taxonomicName>
has a long impunctate channel along the metaventral midline. In
<taxonomicName id="4C254D2B170781794FBDF922FEED7D97" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[229,333,1665,1689]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="planatus">
<emphasis id="B951EABA170781794FBDF922FEED7D97" box="[229,333,1665,1689]" italics="true" pageId="15" pageNumber="16">P. planatus</emphasis>
</taxonomicName>
the entire metaventral disc is covered with punctures and displays a finely carinate median metaventral stria.
</paragraph>
</subSubSection>
<subSubSection id="C33F6523170781794F29F97AFF4B7C1F" pageId="15" pageNumber="16" type="etymology">
<paragraph id="8B9A36A8170781794F29F97AFF4B7C1F" blockId="15.[113,762,1754,1809]" pageId="15" pageNumber="16">
<emphasis id="B951EABA170781794F29F97AFF417DFC" box="[113,225,1754,1778]" italics="true" pageId="15" pageNumber="16">Etymology:</emphasis>
The species name was chosen to signify its flattened body shape.
</paragraph>
</subSubSection>
</treatment>
</document>