560 lines
65 KiB
XML
560 lines
65 KiB
XML
<document id="CA61B6DAECE574E73D6973608578FA9B" ID-CLB-Dataset="45708" ID-DOI="10.5281/zenodo.279573" ID-GBIF-Dataset="7359830e-3896-4dc8-91bc-1c15c4ee2993" ID-ISSN="1175-5326" ID-Zenodo-Dep="279573" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460423643735" checkinUser="plazi" docAuthor="Holleman, Wouter & Bogorodsky, Sergey V." docDate="2012" docId="038C87863F69FFD8FF3661C64A2F6783" docLanguage="en" docName="zt03152p060.pdf" docOrigin="Zootaxa 3152" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Enneapterygius altipinnis Clark 1980" docType="treatment" docVersion="8" lastPageNumber="46" masterDocId="FFB5FFFE3F6CFFD2FFA16231482A6756" masterDocTitle="A review of the blennioid fish family Tripterygiidae (Perciformes) in the Red Sea, with description of Enneapterygius qirmiz, and reinstatement of Enneapterygius altipinnis Clark, 1980" masterLastPageNumber="60" masterPageNumber="36" pageNumber="41" updateTime="1698330352715" updateUser="ExternalLinkService">
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<mods:title id="ACAA0454E68DEEEBCD0A41FEE8D6A8F3">A review of the blennioid fish family Tripterygiidae (Perciformes) in the Red Sea, with description of Enneapterygius qirmiz, and reinstatement of Enneapterygius altipinnis Clark, 1980</mods:title>
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<mods:namePart id="7522D23F48EF2787B7F96B91E7C7EBFE">Holleman, Wouter</mods:namePart>
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<mods:namePart id="536B953EA5A3C56BE20C759FC179D3A8">Bogorodsky, Sergey V.</mods:namePart>
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<mods:title id="F94EB22A2D1C3E0662E5F91D71F60626">Zootaxa</mods:title>
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<mods:part id="C2F41FDFAF48B209421063482145012E">
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<mods:date id="030F16991F31B9B4E2E490A2E3A0BFCC">2012</mods:date>
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<treatment id="038C87863F69FFD8FF3661C64A2F6783" ID-DOI="http://doi.org/10.5281/zenodo.5626516" ID-GBIF-Taxon="119428003" ID-Zenodo-Dep="5626516" LSID="urn:lsid:plazi:treatment:038C87863F69FFD8FF3661C64A2F6783" httpUri="http://treatment.plazi.org/id/038C87863F69FFD8FF3661C64A2F6783" lastPageId="10" lastPageNumber="46" pageId="5" pageNumber="41">
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<subSubSection id="C33F651B3F69FFD7FF3661C64A656364" pageId="5" pageNumber="41" type="nomenclature">
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<paragraph id="8B9A36903F69FFD7FF3661C64A4D6347" blockId="5.[151,615,1015,1074]" box="[151,615,1015,1041]" pageId="5" pageNumber="41">
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<heading id="D0D281FC3F69FFD7FF3661C64A4D6347" bold="true" box="[151,615,1015,1041]" fontSize="11" level="1" pageId="5" pageNumber="41" reason="1">
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<taxonomicName id="4C254D133F69FFD7FF3661C64A4D6347" ID-CoL="39VYL" authority="Clark, 1980" authorityName="Clark" authorityYear="1980" box="[151,615,1015,1041]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="41" phylum="Chordata" rank="species" species="altipinnis">
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<emphasis id="B951EA823F69FFD7FF3661C64A4D6347" bold="true" box="[151,615,1015,1041]" pageId="5" pageNumber="41">
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<emphasis id="B951EA823F69FFD7FF3661C649E56347" bold="true" box="[151,463,1015,1041]" italics="true" pageId="5" pageNumber="41">Enneapterygius altipinnis</emphasis>
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<bibRefCitation id="EFB44B613F69FFD7FE7461C64A4D6347" author="Clark" box="[469,615,1015,1041]" pageId="5" pageNumber="41" refString="Clark, E. (1980) Red Sea fishes of the family Tripterygiidae, with descriptions of eight new species. Israel Journal of Zoology, 28 [for 1979], 65 - 113." type="journal article" year="1980">Clark, 1980</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8B9A36903F69FFD7FF36662B4A656364" blockId="5.[151,615,1015,1074]" box="[151,591,1050,1074]" pageId="5" pageNumber="41">
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<figureCitation id="131E2A153F69FFD7FF36662B48CE6364" box="[151,228,1050,1074]" captionStart="FIGURE 2" captionStartId="6.[151,250,1651,1674]" captionTargetBox="[184,1390,372,1603]" captionTargetId="figure@6.[151,1436,349,1608]" captionTargetPageId="6" captionText="FIGURE 2. Dentary pore patterns of Red Sea species of Enneapterygius: (A) E. abeli; (B) E. altipinnis; (C) E. clarkae; (D) E. destai; (E) E. obscurus; (F) E. pallidus (diagrammatic - J. T. Williams); (G) E. pusillus; (H) E. qirmiz; (I) E. ventermaculus." httpUri="https://zenodo.org/record/279575/files/figure.png" pageId="5" pageNumber="41">Figs. 2</figureCitation>
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B, 3A & B, 4A & B; table 1 & 2
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</paragraph>
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</subSubSection>
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<subSubSection id="C33F651B3F69FFD7FF3666524D676382" pageId="5" pageNumber="41" type="reference_group">
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<paragraph id="8B9A36903F69FFD7FF3666524CEC632F" blockId="5.[151,1436,1122,1237]" box="[151,1222,1122,1145]" pageId="5" pageNumber="41">
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<treatmentCitationGroup id="AB3511BE3F69FFD7FF3666524CEC632F" box="[151,1222,1122,1145]" pageId="5" pageNumber="41">
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<taxonomicName id="4C254D133F69FFD7FF3666524A68632E" ID-CoL="39VYL" authority="Clark, 1980: 99" authorityName="Clark" authorityPageNumber="99" authorityYear="1980" box="[151,578,1122,1144]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="41" phylum="Chordata" rank="species" species="altipinnis">
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<emphasis id="B951EA823F69FFD7FF36665249B6632E" box="[151,412,1122,1144]" italics="true" pageId="5" pageNumber="41">Enneapterygius altipinnis</emphasis>
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<treatmentCitation id="0A8410813F69FFD7FE0366534A68632E" author="Clark" box="[418,578,1122,1144]" page="99" pageId="5" pageNumber="41" year="1980">
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<bibRefCitation id="EFB44B613F69FFD7FE0366534A34632E" author="Clark" box="[418,542,1122,1144]" pageId="5" pageNumber="41" refString="Clark, E. (1980) Red Sea fishes of the family Tripterygiidae, with descriptions of eight new species. Israel Journal of Zoology, 28 [for 1979], 65 - 113." type="journal article" year="1980">Clark, 1980</bibRefCitation>
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: 99
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</treatmentCitation>
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</taxonomicName>
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–101,
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<figureCitation id="131E2A153F69FFD7FDDE66534AEE632E" box="[639,708,1122,1144]" captionStart="FIGURE 3" captionStartId="7.[151,250,1501,1524]" captionTargetBox="[151,1436,267,1477]" captionTargetId="figure@7.[151,1436,267,1477]" captionTargetPageId="7" captionText="FIGURE 3. Enneapterygius altipinnis, M (A) grey form; (B) green form. Photos by S. V. Bogorodsky." httpUri="https://zenodo.org/record/279576/files/figure.png" pageId="5" pageNumber="41">Figs. 3</figureCitation>
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b, 6c & 11 (Gulf of Aqaba and
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<collectingCountry id="F33276003F69FFD7FBA166534C72632E" box="[1024,1112,1122,1144]" name="Ethiopia" pageId="5" pageNumber="41">Ethiopia</collectingCountry>
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, Red Sea).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B9A36903F69FFD7FF3666B34CBE63E3" blockId="5.[151,1436,1122,1237]" pageId="5" pageNumber="41">
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<treatmentCitationGroup id="AB3511BE3F69FFD7FF3666B34CBE63E3" pageId="5" pageNumber="41">
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<taxonomicName id="4C254D133F69FFD7FF3666B349A063C1" ID-CoL="39W3G" box="[151,394,1153,1175]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="41" phylum="Chordata" rank="species" species="tutuilae">
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<emphasis id="B951EA823F69FFD7FF3666B349A063C1" box="[151,394,1153,1175]" italics="true" pageId="5" pageNumber="41">Enneapterygius tutuilae</emphasis>
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</taxonomicName>
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(non
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<bibRefCitation id="EFB44B613F69FFD7FE6766B04A8D63C1" author="Jordan" box="[454,679,1153,1175]" pageId="5" pageNumber="41" refString="Jordan, D. S. & Seale, A. (1906) Fishes of Samoa. Bulletin of the United States Bureau of Fisheries, 25, 173 - 455." type="journal article" year="1906">
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<collectingCountry id="F33276003F69FFD7FE6766B04A2063C1" box="[454,522,1153,1175]" name="Jordan" pageId="5" pageNumber="41">Jordan</collectingCountry>
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& Seale, 1906
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</bibRefCitation>
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):
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<treatmentCitation id="0A8410813F69FFD7FD1866B04B4463C1" author="Fricke" box="[697,878,1153,1175]" page="285" pageId="5" pageNumber="41" year="1994">
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<bibRefCitation id="EFB44B613F69FFD7FD1866B04B1763C1" author="Fricke" box="[697,829,1153,1175]" pageId="5" pageNumber="41" refString="Fricke, R. (1994) Tripterygiid Fishes of Australia, New Zealand and the Southwest Pacific Ocean. Koeltz Scientific Books, Koenigstein: ix + 585 pp." type="book" year="1994">Fricke, 1994</bibRefCitation>
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: 285
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</treatmentCitation>
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ff, Figs. 58 & 59; Randall, Allen & Steene, 1997: 364;
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<treatmentCitation id="0A8410813F69FFD7FF66669149B863E3" author="Holleman" box="[199,402,1184,1206]" page="20" pageId="5" pageNumber="41" year="2005">
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<bibRefCitation id="EFB44B613F69FFD7FF666691494763E0" author="Holleman" box="[199,365,1184,1206]" pageId="5" pageNumber="41" refString="Holleman, W. (2005) A review of the triplefin fish genus Enneapterygius (Blennioidei: Tripterygiidae) in the Western Indian Ocean, with descriptions of four new species. Smithiana Bulletin, 5, 27 pp, 2 pls." type="book" year="2005">Holleman, 2005</bibRefCitation>
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: 20
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</treatmentCitation>
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–21, Fig. 18, Pl.
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<date id="FF9B10503F69FFD7FD9866914A7F63E0" box="[569,597,1184,1206]" pageId="5" pageNumber="41">2 I</date>
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&
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<treatmentCitation id="0A8410813F69FFD7FDD466904B6863E0" author="Fricke" box="[629,834,1184,1206]" page="468" pageId="5" pageNumber="41" year="1999">
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<bibRefCitation id="EFB44B613F69FFD7FDD466904B2563E3" author="Fricke" box="[629,783,1184,1206]" pageId="5" pageNumber="41" refString="Fricke, R. (1999) Fishes of the Mascarene Islands (Reunion, Mauritius, Rodriguez). Koeltz Scientific Books, Koenigstein: viii + 759 pp." type="book" year="1999">J. Fricke, 1999</bibRefCitation>
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: 468
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</treatmentCitation>
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;
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<treatmentCitation id="0A8410813F69FFD7FCED66914CBA63E3" author="Golani" box="[844,1168,1184,1206]" page="44" pageId="5" pageNumber="41" year="2010">
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<bibRefCitation id="EFB44B613F69FFD7FCED66914C4063E3" author="Golani" box="[844,1130,1184,1206]" pageId="5" pageNumber="41" refString="Golani, D. & Bogorodsky, S. V. (2010) The Fishes of the Red Sea-Reappraisal and Updated Checklist. Zootaxa, 2463, 1 - 135." type="journal article" year="2010">Golani & Bogorodsky 2010</bibRefCitation>
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: 44
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B9A36903F69FFD7FF3666F14D676382" blockId="5.[151,1436,1122,1237]" box="[151,1357,1214,1237]" pageId="5" pageNumber="41">
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<taxonomicName id="4C254D133F69FFD7FF3666F149A16383" ID-CoL="6FBJQ" box="[151,395,1215,1237]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="41" phylum="Chordata" rank="species" species="pusillus">
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<emphasis id="B951EA823F69FFD7FF3666F149A16383" box="[151,395,1215,1237]" italics="true" pageId="5" pageNumber="41">Enneapterygius pusillus</emphasis>
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</taxonomicName>
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(non
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<bibRefCitation id="EFB44B613F69FFD7FE64668F4A736382" author="Ruppell" box="[453,601,1214,1236]" pageId="5" pageNumber="41" refString="Ruppell, E. (1835 - 1838) Fische des Rothen Meeres. Neue Wirbelthiere der Fauna Abyssiniens. Frankfurt am Main, [1835, 1 - 28, pls. 1 - 7]." type="book chapter" year="1835">Rüppell, 1835</bibRefCitation>
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): Holleman, 1986: 757, pl. 116, fig. 236: 5 (Great Barrier Reef,
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<collectingCountry id="F33276003F69FFD7FB41668F4D686382" box="[1248,1346,1214,1236]" name="Australia" pageId="5" pageNumber="41">Australia</collectingCountry>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C33F651B3F69FFD4FF3667304B3E664A" lastPageId="6" lastPageNumber="42" pageId="5" pageNumber="41" type="diagnosis">
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<paragraph id="8B9A36903F69FFD7FF3667304AA662D3" blockId="5.[151,1437,1281,1989]" pageId="5" pageNumber="41">
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<emphasis id="B951EA823F69FFD7FF366730493B624C" bold="true" box="[151,273,1281,1306]" pageId="5" pageNumber="41">Diagnosis.</emphasis>
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Dorsal fins III + XII–XIII + 8–10 (usually III + XII + 8 or 9, rarely 10); anal fin I, 15–18 (rarely 15, usually 17); pectoral-fin 14, all rays simple. LL with 10–12 (usually 12) pored scales and 21–23 (usually 21) notched scales; total lateral scales 27 or 28; dentary pores 2 + 1 or 2 + 2, usually 2 + 2 + 2 (
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<figureCitation id="131E2A153F69FFD7FB3867784CF76237" box="[1177,1245,1353,1377]" captionStart="FIGURE 2" captionStartId="6.[151,250,1651,1674]" captionTargetBox="[184,1390,372,1603]" captionTargetId="figure@6.[151,1436,349,1608]" captionTargetPageId="6" captionText="FIGURE 2. Dentary pore patterns of Red Sea species of Enneapterygius: (A) E. abeli; (B) E. altipinnis; (C) E. clarkae; (D) E. destai; (E) E. obscurus; (F) E. pallidus (diagrammatic - J. T. Williams); (G) E. pusillus; (H) E. qirmiz; (I) E. ventermaculus." httpUri="https://zenodo.org/record/279575/files/figure.png" pageId="5" pageNumber="41">Fig. 2</figureCitation>
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B). Head 3.3–4.0 [3.5] in SL; eye 2.6–3.2 [3.0] in head length.
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</paragraph>
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<paragraph id="8B9A36903F69FFD7FF6467A04CE7616F" blockId="5.[151,1437,1281,1989]" pageId="5" pageNumber="41">Head, chest and pectoral-fin base naked; nape scaled; abdomen naked, but scales extend to base of dorsal and anal fins; single row of scales at base of caudal fin; supratemporal sensory canal crescent-shaped; supraorbital cirrus a rounded, flat ‘paddle’, its length about equal to pupil diameter; first dorsal fin higher than second in adult males and equal to slightly higher in females and in juveniles; longest pectoral-fin ray reaches junction between second and third dorsal fins; longest pelvic-fin ray reaches first anal-fin ray in some individuals.</paragraph>
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<paragraph id="8B9A36903F69FFD7FF6464744C266093" blockId="5.[151,1437,1281,1989]" pageId="5" pageNumber="41">
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<emphasis id="B951EA823F69FFD7FF646474494E610B" bold="true" box="[197,356,1605,1629]" pageId="5" pageNumber="41">Fresh colour.</emphasis>
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<emphasis id="B951EA823F69FFD7FECF64744998610B" box="[366,434,1605,1629]" italics="true" pageId="5" pageNumber="41">Males</emphasis>
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: body cream above, white ventrally, many scales edged in orange, with five irregular patches of olive-brown with melanophores, first from front of second dorsal fin and below pectoral fin, second from mid-second dorsal fin, third from junction of second and third dorsal fins, fourth at mid-third dorsal fin and last across peduncle at base of caudal fin; head cream with pale green markings, eye greenish with small white spots; first dorsal fin mottled cream and green, and with red patch at base of membrane between spine 3 and dorsum, spine 1 with alternating dark brown and white bands; second dorsal fin red anteriorly with 2 white bands, first near base of spines 1 and 2, second from tip of spine 2 to base of spine 4, middle of fin with a broad band of melanophores on hyaline membrane, followed by band of cream spots along posterior margin; third dorsal fin similar, but paler and without the red; caudal fin hyaline with white flecks on rays; anal with alternating bands of white, clear and black; pectoral fins with irregular bands of brown, white and green, colour on rays only; anal fin with 6 or 7 black subcutaneous spots at base and black pre-anal spot; pelvic fins whitish.
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</paragraph>
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<paragraph id="8B9A36903F6AFFD4FF6462A94B5467F9" blockId="6.[151,1436,151,284]" box="[197,894,151,176]" pageId="6" pageNumber="42">
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<emphasis id="B951EA823F6AFFD4FF6462A9490E67E6" box="[197,292,152,176]" italics="true" pageId="6" pageNumber="42">Females</emphasis>
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: very similar to males, but without red on dorsal fins.
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</paragraph>
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<paragraph id="8B9A36903F6AFFD4FF64628A4B3E664A" blockId="6.[151,1436,151,284]" pageId="6" pageNumber="42">
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<emphasis id="B951EA823F6AFFD4FF64628A499B6782" bold="true" box="[197,433,187,212]" pageId="6" pageNumber="42">Colour underwater.</emphasis>
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Individuals cryptic on sand, body with pale and olive-green bands; long first spine of the first dorsal fin banded black and white, membranes white, second dorsal fin with white band close to posterior half of margin and other white marks, third dorsal fin similar.
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</paragraph>
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</subSubSection>
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<caption id="DF5A66183F6AFFD4FF3664424D5F61FE" httpUri="https://zenodo.org/record/279575/files/figure.png" pageId="6" pageNumber="42" targetBox="[184,1390,372,1603]" targetPageId="6">
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<paragraph id="8B9A36903F6AFFD4FF3664424D5F61FE" blockId="6.[151,1436,1651,1705]" pageId="6" pageNumber="42">
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<emphasis id="B951EA823F6AFFD4FF366442493961DE" bold="true" box="[151,275,1651,1674]" pageId="6" pageNumber="42">FIGURE 2.</emphasis>
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Dentary pore patterns of Red Sea species of
|
||
<taxonomicName id="4C254D133F6AFFD4FD7A64454B5061DF" box="[731,890,1652,1673]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B951EA823F6AFFD4FD7A64454B5061DF" box="[731,890,1652,1673]" italics="true" pageId="6" pageNumber="42">Enneapterygius</emphasis>
|
||
</taxonomicName>
|
||
: (A)
|
||
<taxonomicName id="4C254D133F6AFFD4FC1064454BD561DF" box="[945,1023,1651,1673]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="42" phylum="Chordata" rank="species" species="abeli">
|
||
<emphasis id="B951EA823F6AFFD4FC1064454BD561DF" box="[945,1023,1651,1673]" italics="true" pageId="6" pageNumber="42">E. abeli</emphasis>
|
||
</taxonomicName>
|
||
; (B)
|
||
<taxonomicName id="4C254D133F6AFFD4FB9464454C9861DF" box="[1077,1202,1651,1673]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="42" phylum="Chordata" rank="species" species="altipinnis">
|
||
<emphasis id="B951EA823F6AFFD4FB9464454C9861DF" box="[1077,1202,1651,1673]" italics="true" pageId="6" pageNumber="42">E. altipinnis</emphasis>
|
||
</taxonomicName>
|
||
; (C)
|
||
<taxonomicName id="4C254D133F6AFFD4FB4664454D6561DF" box="[1255,1359,1651,1673]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="42" phylum="Chordata" rank="species" species="clarkae">
|
||
<emphasis id="B951EA823F6AFFD4FB4664454D6561DF" box="[1255,1359,1651,1673]" italics="true" pageId="6" pageNumber="42">E. clarkae</emphasis>
|
||
</taxonomicName>
|
||
; (D)
|
||
<taxonomicName id="4C254D133F6AFFD4FA27644548F961FE" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="42" phylum="Chordata" rank="species" species="destai">
|
||
<emphasis id="B951EA823F6AFFD4FA27644548F961FE" italics="true" pageId="6" pageNumber="42">E. destai</emphasis>
|
||
</taxonomicName>
|
||
; (E)
|
||
<taxonomicName id="4C254D133F6AFFD4FEA664A2495461FE" box="[263,382,1682,1704]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="42" phylum="Chordata" rank="species" species="obscurus">
|
||
<emphasis id="B951EA823F6AFFD4FEA664A2495461FE" box="[263,382,1682,1704]" italics="true" pageId="6" pageNumber="42">E. obscurus</emphasis>
|
||
</taxonomicName>
|
||
; (F)
|
||
<taxonomicName id="4C254D133F6AFFD4FE1064A24A3561FE" box="[433,543,1682,1704]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="42" phylum="Chordata" rank="species" species="pallidus">
|
||
<emphasis id="B951EA823F6AFFD4FE1064A24A3561FE" box="[433,543,1682,1704]" italics="true" pageId="6" pageNumber="42">E. pallidus</emphasis>
|
||
</taxonomicName>
|
||
(diagrammatic - J.T. Williams); (G)
|
||
<taxonomicName id="4C254D133F6AFFD4FC3564A24BD561FE" box="[916,1023,1682,1704]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="42" phylum="Chordata" rank="species" species="pusillus">
|
||
<emphasis id="B951EA823F6AFFD4FC3564A24BD561FE" box="[916,1023,1682,1704]" italics="true" pageId="6" pageNumber="42">E. pusillus</emphasis>
|
||
</taxonomicName>
|
||
; (H)
|
||
<taxonomicName id="4C254D133F6AFFD4FB9464A24CB861FE" box="[1077,1170,1683,1704]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="42" phylum="Chordata" rank="species" species="qirmiz">
|
||
<emphasis id="B951EA823F6AFFD4FB9464A24CB861FE" box="[1077,1170,1683,1704]" italics="true" pageId="6" pageNumber="42">E. qirmiz</emphasis>
|
||
</taxonomicName>
|
||
; (I)
|
||
<taxonomicName id="4C254D133F6AFFD4FB1E64A24D4561FE" box="[1215,1391,1682,1704]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="42" phylum="Chordata" rank="species" species="ventermaculus">
|
||
<emphasis id="B951EA823F6AFFD4FB1E64A24D4561FE" box="[1215,1391,1682,1704]" italics="true" pageId="6" pageNumber="42">E. ventermaculus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33F651B3F6AFFD4FF6464E44CD56046" pageId="6" pageNumber="42" type="key">
|
||
<paragraph id="8B9A36903F6AFFD4FF6464E44CD56046" blockId="6.[151,1436,1748,2024]" pageId="6" pageNumber="42">
|
||
<emphasis id="B951EA823F6AFFD4FF6464E4494F61BB" bold="true" box="[197,357,1748,1773]" pageId="6" pageNumber="42">Key features.</emphasis>
|
||
First dorsal fin tall; crescent-shaped supraoccipital sensory canal; underwater – tall first dorsal fin; white to yellowish band across the posterior portion of second dorsal fin, band continuing to LL.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33F651B3F6AFFD4FF64652D49F8602B" pageId="6" pageNumber="42" type="distribution">
|
||
<paragraph id="8B9A36903F6AFFD4FF64652D4C716063" blockId="6.[151,1436,1748,2024]" box="[197,1115,1820,1846]" pageId="6" pageNumber="42">
|
||
<emphasis id="B951EA823F6AFFD4FF64652D497D6063" bold="true" box="[197,343,1820,1845]" pageId="6" pageNumber="42">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="F33276003F6AFFD4FEC4652F49816060" box="[357,427,1822,1846]" name="Egypt" pageId="6" pageNumber="42">Egypt</collectingCountry>
|
||
, Gulf Aqaba to
|
||
<collectingCountry id="F33276003F6AFFD4FDFC652C4A866063" box="[605,684,1821,1845]" name="Eritrea" pageId="6" pageNumber="42">Eritrea</collectingCountry>
|
||
, but not recorded outside the Red Sea.
|
||
</paragraph>
|
||
<paragraph id="8B9A36903F6AFFD4FF64657149F8602B" blockId="6.[151,1436,1748,2024]" pageId="6" pageNumber="42">
|
||
<emphasis id="B951EA823F6AFFD4FF6465714902600F" bold="true" box="[197,296,1856,1881]" pageId="6" pageNumber="42">Habitat.</emphasis>
|
||
In shallow water, collected material was taken from depth of
|
||
<quantity id="4CDD9B753F6AFFD4FC4765714C1A600E" box="[998,1072,1856,1880]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="6" pageNumber="42" unit="m" value="2.5" valueMax="3.0" valueMin="2.0">2–3 m</quantity>
|
||
, on dead corals near the base of reefs. Reaches
|
||
<quantity id="4CDD9B753F6AFFD4FEE16554498E602B" box="[320,420,1893,1917]" metricMagnitude="-2" metricUnit="m" metricValue="1.97" pageId="6" pageNumber="42" unit="mm" value="19.7">19.7 mm</quantity>
|
||
SL.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33F651B3F6AFFD5FF6665B84C056195" lastPageId="7" lastPageNumber="43" pageId="6" pageNumber="42" type="discussion">
|
||
<paragraph id="8B9A36903F6AFFD5FF6665B84DB76782" blockId="6.[151,1436,1748,2024]" lastBlockId="7.[151,1437,151,212]" lastPageId="7" lastPageNumber="43" pageId="6" pageNumber="42">
|
||
<emphasis id="B951EA823F6AFFD4FF6665B8491160F7" bold="true" box="[199,315,1929,1953]" pageId="6" pageNumber="42">Remarks.</emphasis>
|
||
The species was considered synonymous with
|
||
<taxonomicName id="4C254D133F6AFFD4FCFC65BB4C4660F7" box="[861,1132,1929,1953]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="42" phylum="Chordata" rank="species" species="tutuilae">
|
||
<emphasis id="B951EA823F6AFFD4FCFC65BB4C4660F7" box="[861,1132,1929,1953]" italics="true" pageId="6" pageNumber="42">Enneapterygius tutuilae</emphasis>
|
||
</taxonomicName>
|
||
<collectingCountry id="F33276003F6AFFD4FBD465B94CEA60F6" box="[1141,1216,1928,1952]" name="Jordan" pageId="6" pageNumber="42">Jordan</collectingCountry>
|
||
& Seale by
|
||
<bibRefCitation id="EFB44B613F6AFFD4FAF565B949306093" author="Fricke" pageId="6" pageNumber="42" refString="Fricke, R. (1994) Tripterygiid Fishes of Australia, New Zealand and the Southwest Pacific Ocean. Koeltz Scientific Books, Koenigstein: ix + 585 pp." type="book" year="1994">Fricke (1994: 285)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFB44B613F6AFFD4FEF0659C4A156093" author="Holleman" box="[337,575,1965,1989]" pageId="6" pageNumber="42" refString="Holleman, W. (2005) A review of the triplefin fish genus Enneapterygius (Blennioidei: Tripterygiidae) in the Western Indian Ocean, with descriptions of four new species. Smithiana Bulletin, 5, 27 pp, 2 pls." type="book" year="2005">Holleman (2005: 20)</bibRefCitation>
|
||
. In the Indian Ocean
|
||
<emphasis id="B951EA823F6AFFD4FC8E659F4BE86093" box="[815,962,1965,1989]" italics="true" pageId="6" pageNumber="42">
|
||
“
|
||
<taxonomicName id="4C254D133F6AFFD4FC99659F4B906093" box="[824,954,1965,1989]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="42" phylum="Chordata" rank="species" species="tutuilae">E. tutuilae</taxonomicName>
|
||
”
|
||
</emphasis>
|
||
— as then considered by these authors — is known from the coast of East Africa,
|
||
<collectingCountry id="F33276003F6AFFD4FDFC65E14ACC60BE" box="[605,742,2000,2024]" name="Madagascar" pageId="6" pageNumber="42">Madagascar</collectingCountry>
|
||
, and the Mascarene Islands, but not from the
|
||
<collectingCountry id="F33276003F6AFFD4FB4A65E14D7C60BE" box="[1259,1366,2000,2024]" name="Maldives" pageId="6" pageNumber="42">Maldives</collectingCountry>
|
||
,
|
||
<collectingCountry id="F33276003F6AFFD4FAC365E14DB660BE" box="[1378,1436,2000,2024]" name="India" pageId="6" pageNumber="42">India</collectingCountry>
|
||
or
|
||
<collectingCountry id="F33276003F6BFFD5FF1862A6490467F9" box="[185,302,151,175]" name="Sri Lanka" pageId="7" pageNumber="43">Sri Lanka</collectingCountry>
|
||
. Individuals of
|
||
<emphasis id="B951EA823F6BFFD5FE4662A94A5467E6" box="[487,638,152,176]" italics="true" pageId="7" pageNumber="43">
|
||
“
|
||
<taxonomicName id="4C254D133F6BFFD5FE5162A94A5C67E6" box="[496,630,152,176]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="43" phylum="Chordata" rank="species" species="tutuilae">E. tutuilae</taxonomicName>
|
||
”
|
||
</emphasis>
|
||
have been recorded from the Cocos-Keeling and Christmas Islands, throughout
|
||
<collectingCountry id="F33276003F6BFFD5FEB9628D49A26782" box="[280,392,188,212]" name="Indonesia" pageId="7" pageNumber="43">Indonesia</collectingCountry>
|
||
, tropical
|
||
<collectingCountry id="F33276003F6BFFD5FE4F628D4A726782" box="[494,600,188,212]" name="Australia" pageId="7" pageNumber="43">Australia</collectingCountry>
|
||
, and in the Western Pacific to
|
||
<collectingCountry id="F33276003F6BFFD5FC06628D4BE46782" box="[935,974,188,212]" name="Fiji" pageId="7" pageNumber="43">Fiji</collectingCountry>
|
||
and Rotuma (see
|
||
<bibRefCitation id="EFB44B613F6BFFD5FB37628D4D0A6782" author="Fricke" box="[1174,1312,188,212]" pageId="7" pageNumber="43" refString="Fricke, R. (1997) Tripterygiid Fishes of the Western and Central Pacific (Teleostei). Koeltz Scientific Books, Koenigstein: ix + 607 pp." type="book" year="1997">Fricke 1997</bibRefCitation>
|
||
: 339–349).
|
||
</paragraph>
|
||
<caption id="DF5A66183F6BFFD5FF3667EC4C8E62A5" box="[151,1188,1501,1524]" httpUri="https://zenodo.org/record/279576/files/figure.png" pageId="7" pageNumber="43" targetBox="[151,1436,267,1477]" targetPageId="7">
|
||
<paragraph id="8B9A36903F6BFFD5FF3667EC4C8E62A5" blockId="7.[151,1188,1501,1524]" box="[151,1188,1501,1524]" pageId="7" pageNumber="43">
|
||
<emphasis id="B951EA823F6BFFD5FF3667EC493862A4" bold="true" box="[151,274,1501,1524]" pageId="7" pageNumber="43">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C254D133F6BFFD5FEB867EF4A3462A5" box="[281,542,1501,1523]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="43" phylum="Chordata" rank="species" species="altipinnis">
|
||
<emphasis id="B951EA823F6BFFD5FEB867EF4A3462A5" box="[281,542,1501,1523]" italics="true" pageId="7" pageNumber="43">Enneapterygius altipinnis</emphasis>
|
||
</taxonomicName>
|
||
, M (A) grey form; (B) green form. Photos by S.V. Bogorodsky.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9A36903F6BFFD5FF64642A4C056195" blockId="7.[151,1436,1563,1984]" pageId="7" pageNumber="43">
|
||
<collectingCountry id="F33276003F6BFFD5FF64642A48CE6165" box="[197,228,1563,1587]" name="American Samoa" pageId="7" pageNumber="43">As</collectingCountry>
|
||
in the species of the
|
||
<taxonomicName id="4C254D133F6BFFD5FE70642A4AE56165" box="[465,719,1563,1587]" class="Actinopterygii" family="Tripterygiidae" genus="Helcogramma" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="43" phylum="Chordata" rank="species" species="steinitzi">
|
||
<emphasis id="B951EA823F6BFFD5FE70642A4AE56165" box="[465,719,1563,1587]" italics="true" pageId="7" pageNumber="43">Helcogramma steinitzi</emphasis>
|
||
</taxonomicName>
|
||
complex (
|
||
<bibRefCitation id="EFB44B613F6BFFD5FCE4642A4C026165" author="Holleman" box="[837,1064,1563,1587]" pageId="7" pageNumber="43" refString="Holleman, W. (2006) Fishes of the Helcogramma steinitzi species group (Blennioidei: Tripterygiidae) from the Indian Ocean with descriptions of two new species. aqua, Journal of Ichthyology and Aquatic Biology, 11 (3), 89 - 104." type="journal article" year="2006">Holleman, 2006: 91</bibRefCitation>
|
||
), there is little difference in morphometric counts between species of the
|
||
<taxonomicName id="4C254D133F6BFFD5FD2164714BB86101" box="[640,914,1599,1623]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="43" phylum="Chordata" rank="species" species="tutuilae">
|
||
<emphasis id="B951EA823F6BFFD5FD2164714BB86101" box="[640,914,1599,1623]" italics="true" pageId="7" pageNumber="43">Enneapterygius tutuilae</emphasis>
|
||
</taxonomicName>
|
||
complex distributed over this wide area (as shown in
|
||
<tableCitation id="C6A7032B3F6BFFD5FEA26452497F612D" box="[259,341,1635,1659]" captionStart="TABLE 2" captionStartId="8.[151,235,845,866]" captionTargetBox="[151,1384,895,1307]" captionTargetPageId="8" captionText="TABLE 2. Comparative counts for E. altipinnis sensu Clark and E. tutuilae sensu Jordan & Seale." httpUri="http://table.plazi.org/id/DF5A66183F64FFDAFF36617C4C506435" pageId="7" pageNumber="43" tableUuid="DF5A66183F64FFDAFF36617C4C506435">Table 2</tableCitation>
|
||
below). There are, however, pigmentation differences suggesting that the various ‘forms’ are part of a widespread species complex, similar to those already described for the
|
||
<taxonomicName id="4C254D133F6BFFD5FC4564B64D2C61C9" box="[996,1286,1671,1695]" class="Actinopterygii" family="Tripterygiidae" genus="Helcogramma" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="43" phylum="Chordata" rank="species" species="fuscopinna">
|
||
<emphasis id="B951EA823F6BFFD5FC4564B64D2C61C9" box="[996,1286,1671,1695]" italics="true" pageId="7" pageNumber="43">Helcogramma fuscopinna</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C254D133F6BFFD5FA9D64B948906195" class="Actinopterygii" family="Tripterygiidae" genus="Helcogramma" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="43" phylum="Chordata" rank="species" species="steinitzi">
|
||
<emphasis id="B951EA823F6BFFD5FA9D64B948906195" italics="true" pageId="7" pageNumber="43">H. steinitzi</emphasis>
|
||
</taxonomicName>
|
||
complexes (see also
|
||
<bibRefCitation id="EFB44B613F6BFFD5FE06649A4B2A6195" author="Williams" box="[423,768,1707,1731]" pageId="7" pageNumber="43" refString="Williams, J. T. & McCormick, C. J. (1990) Two new species of the triplefin fish genus Helcogramma (Tripterygiidae) from the Western Pacific Ocean. Copeia, 1990 (4), 1020 - 1030." type="journal article" year="1990">Williams & McCormick, 1990</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB44B613F6BFFD5FCAD649A4C096195" author="Williams" box="[780,1059,1707,1731]" pageId="7" pageNumber="43" refString="Williams, J. T. & Howe, J. C. (2003) Seven new species of the triplefin genus Helcogramma (Tripterygiidae) from the Indo - Pacific. Aqua, Journal of Ichthyology and Aquatic Biology, 7 (4), 151 - 176." type="journal article" year="2003">Williams & Howe, 2003</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33F651B3F6BFFDBFF6464E1491166FE" lastPageId="9" lastPageNumber="45" pageId="7" pageNumber="43" type="reference_group">
|
||
<paragraph id="8B9A36903F6BFFD5FF6464E14CE36096" blockId="7.[151,1436,1563,1984]" pageId="7" pageNumber="43">
|
||
<taxonomicName id="4C254D133F6BFFD5FF6464E1496461B1" ID-CoL="39VYL" box="[197,334,1743,1767]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="43" phylum="Chordata" rank="species" species="altipinnis">
|
||
<emphasis id="B951EA823F6BFFD5FF6464E1496461B1" box="[197,334,1743,1767]" italics="true" pageId="7" pageNumber="43">E. altipinnis</emphasis>
|
||
</taxonomicName>
|
||
from the Red Sea has no divided pectoral-fin rays (and Williams confirmed this for the
|
||
<typeStatus id="549E88323F6BFFD5FA8964FE4D4F61B1" box="[1320,1381,1743,1767]" pageId="7" pageNumber="43">types</typeStatus>
|
||
) and were incorrectly recorded by both
|
||
<bibRefCitation id="EFB44B613F6BFFD5FDBC64C24ACD605D" author="Clark" box="[541,743,1779,1803]" pageId="7" pageNumber="43" refString="Clark, E. (1980) Red Sea fishes of the family Tripterygiidae, with descriptions of eight new species. Israel Journal of Zoology, 28 [for 1979], 65 - 113." type="journal article" year="1980">Clark (1980: 100)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFB44B613F6BFFD5FC8164C24BDC605D" author="Fricke" box="[800,1014,1779,1803]" pageId="7" pageNumber="43" refString="Fricke, R. (1997) Tripterygiid Fishes of the Western and Central Pacific (Teleostei). Koeltz Scientific Books, Koenigstein: ix + 607 pp." type="book" year="1997">Fricke (1997: 338)</bibRefCitation>
|
||
. All the Western Indian Ocean specimens of “
|
||
<taxonomicName id="4C254D133F6BFFD5FEA4652949506079" ID-CoL="39W3G" box="[261,378,1815,1839]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="43" phylum="Chordata" rank="species" species="tutuilae">
|
||
<emphasis id="B951EA823F6BFFD5FEA4652949506079" box="[261,378,1815,1839]" italics="true" pageId="7" pageNumber="43">E. tutuilae</emphasis>
|
||
</taxonomicName>
|
||
” examined by the first author have only simple pectoral-fin rays (see
|
||
<bibRefCitation id="EFB44B613F6BFFD5FB3465264D7A6079" author="Holleman" box="[1173,1360,1815,1839]" pageId="7" pageNumber="43" refString="Holleman, W. (2005) A review of the triplefin fish genus Enneapterygius (Blennioidei: Tripterygiidae) in the Western Indian Ocean, with descriptions of four new species. Smithiana Bulletin, 5, 27 pp, 2 pls." type="book" year="2005">Holleman, 2005</bibRefCitation>
|
||
). Winterbottom (pers. comm.) confirmed that specimens from Raja Ampat also have only simple pectoral-fin rays. Photographs of a similar species
|
||
<taxonomicName id="4C254D133F6BFFD5FDA865514A936021" ID-CoL="4BXF" box="[521,697,1888,1911]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="43" phylum="Chordata" rank="genus">
|
||
<emphasis id="B951EA823F6BFFD5FDA865514A936021" box="[521,697,1888,1911]" italics="true" pageId="7" pageNumber="43">Enneapterygius</emphasis>
|
||
</taxonomicName>
|
||
from the Ogasawara Islands,
|
||
<collectingCountry id="F33276003F6BFFD5FBA665514C606021" box="[1031,1098,1888,1911]" name="Japan" pageId="7" pageNumber="43">Japan</collectingCountry>
|
||
, also show only simple pectoral-fin rays. This suggests simple pectoral-fin rays is a shared character for this species complex;
|
||
<taxonomicName id="4C254D133F6BFFD5FB7C65B5490B60E9" ID-CoL="6FBHY" authority="Holleman, 2005" authorityName="Holleman" authorityYear="2005" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="43" phylum="Chordata" rank="species" species="kosiensis">
|
||
<emphasis id="B951EA823F6BFFD5FB7C65B54D4B60CD" box="[1245,1377,1923,1947]" italics="true" pageId="7" pageNumber="43">E. kosiensis</emphasis>
|
||
<bibRefCitation id="EFB44B613F6BFFD5FAC865B2490B60E9" author="Holleman" pageId="7" pageNumber="43" refString="Holleman, W. (2005) A review of the triplefin fish genus Enneapterygius (Blennioidei: Tripterygiidae) in the Western Indian Ocean, with descriptions of four new species. Smithiana Bulletin, 5, 27 pp, 2 pls." type="book" year="2005">Holleman, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
is the only other
|
||
<taxonomicName id="4C254D133F6BFFD5FE4565994ABE60E9" ID-CoL="4BXF" box="[484,660,1960,1983]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="43" phylum="Chordata" rank="genus">
|
||
<emphasis id="B951EA823F6BFFD5FE4565994ABE60E9" box="[484,660,1960,1983]" italics="true" pageId="7" pageNumber="43">Enneapterygius</emphasis>
|
||
</taxonomicName>
|
||
species known with only simple pectoral-fin rays.
|
||
</paragraph>
|
||
<caption id="DF5A66183F64FFDAFF3660DA4C736477" httpUri="https://zenodo.org/record/279577/files/figure.png" pageId="8" pageNumber="44" targetBox="[151,1436,195,698]" targetPageId="8">
|
||
<paragraph id="8B9A36903F64FFDAFF3660DA4C736477" blockId="8.[151,1436,747,801]" pageId="8" pageNumber="44">
|
||
<emphasis id="B951EA823F64FFDAFF3660DA493E6457" bold="true" box="[151,276,747,770]" pageId="8" pageNumber="44">FIGURE 4.</emphasis>
|
||
(A, B)
|
||
<taxonomicName id="4C254D133F64FFDAFEC460DC4A416454" ID-CoL="39VYL" box="[357,619,748,770]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="44" phylum="Chordata" rank="species" species="altipinnis">
|
||
<emphasis id="B951EA823F64FFDAFEC460DC4A416454" box="[357,619,748,770]" italics="true" pageId="8" pageNumber="44">Enneapterygius altipinnis</emphasis>
|
||
</taxonomicName>
|
||
, M, Rabigh, Saudi Arabia; (B)
|
||
<taxonomicName id="4C254D133F64FFDAFC1260DC4C866454" box="[947,1196,748,770]" class="Actinopterygii" family="Tripterygiidae" genus="Enneaterygius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="44" phylum="Chordata" rank="species" species="altipinnis">
|
||
<emphasis id="B951EA823F64FFDAFC1260DC4C866454" box="[947,1196,748,770]" italics="true" pageId="8" pageNumber="44">Enneaterygius altipinnis</emphasis>
|
||
</taxonomicName>
|
||
, F, Marsa Alam; (C,D)
|
||
<emphasis id="B951EA823F64FFDAFF36613A498B6476" box="[151,417,778,800]" italics="true" pageId="8" pageNumber="44">
|
||
<taxonomicName id="4C254D133F64FFDAFF36613A491C6476" ID-CoL="4BXF" box="[151,310,779,800]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="44" phylum="Chordata" rank="genus">Enneapterygius</taxonomicName>
|
||
cf
|
||
<taxonomicName id="4C254D133F64FFDAFEF5613B498B6476" ID-CoL="39W3G" box="[340,417,778,800]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="44" phylum="Chordata" rank="species" species="tutuilae">tutuilae</taxonomicName>
|
||
</emphasis>
|
||
, Comoros. Photos by S. Bogorodsky (A,B); R. Winterbottom (C,D).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5A66183F64FFDAFF36617C4C506435" ID-Table-UUID="DF5A66183F64FFDAFF36617C4C506435" box="[151,1146,845,867]" httpUri="http://table.plazi.org/id/DF5A66183F64FFDAFF36617C4C506435" pageId="8" pageNumber="44" targetBox="[151,1384,895,1307]" targetIsTable="true" targetPageId="8">
|
||
<paragraph id="8B9A36903F64FFDAFF36617C4C506435" blockId="8.[151,1146,845,867]" box="[151,1146,845,867]" pageId="8" pageNumber="44">
|
||
<emphasis id="B951EA823F64FFDAFF36617C492E6434" bold="true" box="[151,260,845,866]" pageId="8" pageNumber="44">TABLE 2.</emphasis>
|
||
Comparative counts for
|
||
<taxonomicName id="4C254D133F64FFDAFDA1617F4AD26435" authority="Clark" authorityName="Clark" box="[512,760,845,867]" class="Actinopterygii" family="Tripterygiidae" genus="Enneaterygius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="44" phylum="Chordata" rank="species" species="altipinnis">
|
||
<emphasis id="B951EA823F64FFDAFDA1617F4A906435" box="[512,698,845,867]" italics="true" pageId="8" pageNumber="44">E. altipinnis sensu</emphasis>
|
||
Clark
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C254D133F64FFDAFC8B617F4BF86435" box="[810,978,845,867]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="44" phylum="Chordata" rank="subSpecies" species="tutuilae" subSpecies="sensu">
|
||
<emphasis id="B951EA823F64FFDAFC8B617F4BF86435" box="[810,978,845,867]" italics="true" pageId="8" pageNumber="44">E. tutuilae sensu</emphasis>
|
||
</taxonomicName>
|
||
Jordan & Seale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9A36903F64FFDAFEE661B14CBD63AA" pageId="8" pageNumber="44">
|
||
<table id="F925C4303F64002DFF36614E4D42624D" box="[151,1384,895,1307]" gridcols="3" gridrows="6" pageId="8" pageNumber="44">
|
||
<tr id="351534D23F64002DFF36614E4D4264C3" box="[151,1384,895,917]" gridrow="0" pageId="8" pageNumber="44" rowspan-0="1">
|
||
<th id="76C45DAE3F64002DFEE6614E4B1D64C3" box="[327,823,895,917]" gridcol="1" gridrow="0" pageId="8" pageNumber="44">
|
||
<taxonomicName id="4C254D133F64FFDAFEE661B149E964C3" box="[327,451,895,917]" class="Actinopterygii" family="Tripterygiidae" genus="Enneaterygius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="44" phylum="Chordata" rank="species" species="altipinnis">
|
||
<emphasis id="B951EA823F64FFDAFEE661B149E964C3" box="[327,451,895,917]" italics="true" pageId="8" pageNumber="44">E. altipinnis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="76C45DAE3F64002DFCC6614E4D4264C3" box="[871,1384,895,917]" gridcol="2" gridrow="0" pageId="8" pageNumber="44">
|
||
<taxonomicName id="4C254D133F64FFDAFCC661B14BFB64C3" ID-CoL="39W3G" box="[871,977,895,917]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="44" phylum="Chordata" rank="species" species="tutuilae">
|
||
<emphasis id="B951EA823F64FFDAFCC661B14BFB64C3" box="[871,977,895,917]" italics="true" pageId="8" pageNumber="44">E. tutuilae</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="351534D23F64002DFF3661994D4264AD" box="[151,1384,936,1019]" gridrow="1" pageId="8" pageNumber="44">
|
||
<th id="76C45DAE3F64002DFF366199490464AD" box="[151,302,936,1019]" gridcol="0" gridrow="1" pageId="8" pageNumber="44">Dorsal fins</th>
|
||
<td id="76C45DAE3F64002DFEE661994B1D64AD" box="[327,823,936,1019]" gridcol="1" gridrow="1" pageId="8" pageNumber="44">III + XII–XIII + 8–10, usually XII + 8 or 9, rarely 10 rays (Clark 1980; this study)</td>
|
||
<td id="76C45DAE3F64002DFCC661994D4264AD" box="[871,1384,936,1019]" gridcol="2" gridrow="1" pageId="8" pageNumber="44">III + XII + 9 (Jordan & Seale 1905) III + XI–XII + 9–10, usually III + XII + 10 (Holleman 2005)*</td>
|
||
</tr>
|
||
<tr id="351534D23F64002DFF36663F4D426315" box="[151,1384,1038,1091]" gridrow="2" pageId="8" pageNumber="44">
|
||
<th id="76C45DAE3F64002DFF36663F49046315" box="[151,302,1038,1091]" gridcol="0" gridrow="2" pageId="8" pageNumber="44">Anal fin</th>
|
||
<td id="76C45DAE3F64002DFEE6663F4B1D6315" box="[327,823,1038,1091]" gridcol="1" gridrow="2" pageId="8" pageNumber="44">I, 15–17, usually 17 rays, rarely 15 (Clark 1980); 17 or 18 (this study)</td>
|
||
<td id="76C45DAE3F64002DFCC6663F4D426315" box="[871,1384,1038,1091]" gridcol="2" gridrow="2" pageId="8" pageNumber="44">18 (I, 17?) (Jordan & Seale 1905) I, 15–17, usually 17 (Holleman 2005)</td>
|
||
</tr>
|
||
<tr id="351534D23F64002DFF3666674D4263DD" box="[151,1384,1110,1163]" gridrow="3" pageId="8" pageNumber="44">
|
||
<th id="76C45DAE3F64002DFF366667490463DD" box="[151,302,1110,1163]" gridcol="0" gridrow="3" pageId="8" pageNumber="44">Pectoral fins</th>
|
||
<td id="76C45DAE3F64002DFEE666674B1D63DD" box="[327,823,1110,1163]" gridcol="1" gridrow="3" pageId="8" pageNumber="44">13–15, usually14–15, usually 14, all simple (not from Clark — see below)</td>
|
||
<td id="76C45DAE3F64002DFCC666674D4263DD" box="[871,1384,1110,1163]" gridcol="2" gridrow="3" pageId="8" pageNumber="44">14, all simple (Holleman 2005)</td>
|
||
</tr>
|
||
<tr id="351534D23F64002DFF3666AF4D426382" box="[151,1384,1182,1236]" gridrow="4" pageId="8" pageNumber="44">
|
||
<th id="76C45DAE3F64002DFF3666AF49046382" box="[151,302,1182,1236]" gridcol="0" gridrow="4" pageId="8" pageNumber="44">Lateral line</th>
|
||
<td id="76C45DAE3F64002DFEE666AF4B1D6382" box="[327,823,1182,1236]" gridcol="1" gridrow="4" pageId="8" pageNumber="44">10–12 (usually 12) + 21–23 (usually 21) (Clark 1980)</td>
|
||
<td id="76C45DAE3F64002DFCC666AF4D426382" box="[871,1384,1182,1236]" gridcol="2" gridrow="4" pageId="8" pageNumber="44">33 lateral scales (Jordan & Seale 1905) 11–12 + 20–23 (usually 12 + 22) (Holleman 2005)</td>
|
||
</tr>
|
||
<tr id="351534D23F64002DFF3666D74D42624D" box="[151,1384,1254,1307]" gridrow="5" pageId="8" pageNumber="44">
|
||
<th id="76C45DAE3F64002DFF3666D74904624D" box="[151,302,1254,1307]" gridcol="0" gridrow="5" pageId="8" pageNumber="44">Dentary pores</th>
|
||
<td id="76C45DAE3F64002DFEE666D74B1D624D" box="[327,823,1254,1307]" gridcol="1" gridrow="5" pageId="8" pageNumber="44">2 + 2 + 2 (types, Williams pers. comm); occasionally 2 + 1 + 2 (this study)</td>
|
||
<td id="76C45DAE3F64002DFCC666D74D42624D" box="[871,1384,1254,1307]" gridcol="2" gridrow="5" pageId="8" pageNumber="44">2 + 1 / 2 + 2 (Holleman 2005)</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8B9A36903F64FFDAFF36670D49DF6226" blockId="8.[151,1436,1339,1392]" pageId="8" pageNumber="44">
|
||
* Data from
|
||
<bibRefCitation id="EFB44B613F64FFDAFEBD670A49966207" author="Holleman" box="[284,444,1339,1361]" pageId="8" pageNumber="44" refString="Holleman, W. (2005) A review of the triplefin fish genus Enneapterygius (Blennioidei: Tripterygiidae) in the Western Indian Ocean, with descriptions of four new species. Smithiana Bulletin, 5, 27 pp, 2 pls." type="book" year="2005">Holleman 2005</bibRefCitation>
|
||
, from specimens from E. Africa,
|
||
<collectingCountry id="F33276003F64FFDAFCB6670D4B966207" box="[791,956,1339,1361]" name="Comoros" pageId="8" pageNumber="44">Comoro Islands</collectingCountry>
|
||
,
|
||
<collectingCountry id="F33276003F64FFDAFC69670A4C076207" box="[968,1069,1339,1361]" name="Mauritius" pageId="8" pageNumber="44">Mauritius</collectingCountry>
|
||
, St Brandon Shoals, west
|
||
<collectingCountry id="F33276003F64FFDAFA9A670A4DB26207" box="[1339,1432,1339,1361]" name="Thailand" pageId="8" pageNumber="44">Thailand</collectingCountry>
|
||
, and a single specimen from
|
||
<collectingCountry id="F33276003F64FFDAFE15676A49DB6226" box="[436,497,1371,1392]" name="Japan" pageId="8" pageNumber="44">Japan</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9A36903F64FFDAFF6467AC49346177" blockId="8.[151,1436,1437,2036]" pageId="8" pageNumber="44">
|
||
Specimens from East African and Mascarene Islands and from
|
||
<collectingCountry id="F33276003F64FFDAFC2F67AC4BD162E3" box="[910,1019,1437,1461]" name="Indonesia" pageId="8" pageNumber="44">Indonesia</collectingCountry>
|
||
eastwards appear to vary in colouration, and species may be sexually dichromatic. Pertinent to this paper, however, is to distinguish between specimens from the Red Sea and those from elsewhere in the Western Indian Ocean, and this distinction lies in colouration.
|
||
</paragraph>
|
||
<paragraph id="8B9A36903F64FFDAFF64641C4CFA60DF" blockId="8.[151,1436,1437,2036]" pageId="8" pageNumber="44">
|
||
Presumed immature / sexually inactive males from the
|
||
<collectingCountry id="F33276003F64FFDAFC8A641C4BF66113" box="[811,988,1581,1605]" name="Comoros" pageId="8" pageNumber="44">Comoro Islands</collectingCountry>
|
||
have whitish bodies with dark brown to black markings, scale pockets edged in red, the first dorsal fin yellow (
|
||
<figureCitation id="131E2A153F64FFDAFC1164604BDF613F" box="[944,1013,1617,1641]" captionStart="FIGURE 4" captionStartId="8.[151,250,747,770]" captionTargetBox="[151,1436,195,698]" captionTargetId="figure@8.[151,1436,195,723]" captionTargetPageId="8" captionText="FIGURE 4. (A, B) Enneapterygius altipinnis, M, Rabigh, Saudi Arabia; (B) Enneaterygius altipinnis, F, Marsa Alam; (C, D) Enneapterygius cf tutuilae, Comoros. Photos by S. Bogorodsky (A, B); R. Winterbottom (C, D)." httpUri="https://zenodo.org/record/279577/files/figure.png" pageId="8" pageNumber="44">Fig. 4</figureCitation>
|
||
). Mature / sexually active males have the head and anterior part of the body suffused with bright green, the colour extending to the base and lower part if the first dorsal fin and on the belly as far as the pre-anal spot. Scale pockets on the body tend to be more intensely coloured, as are the rays of the pectoral fins, and the black spot between spines 2 and 4 of the second dorsal fin, with its ‘trailing tail’ of melanophores, rather like a black comet and tail. The orbital cirri are deep yellow. By comparison, an intensely coloured male
|
||
<taxonomicName id="4C254D133F64FFDAFD8F65374A9D604B" box="[558,695,1797,1821]" class="Actinopterygii" family="Tripterygiidae" genus="Enneaterygius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="44" phylum="Chordata" rank="species" species="altipinnis">
|
||
<emphasis id="B951EA823F64FFDAFD8F65374A9D604B" box="[558,695,1797,1821]" italics="true" pageId="8" pageNumber="44">E. altipinnis</emphasis>
|
||
</taxonomicName>
|
||
from the Red Sea lacks the green on the anterior of the body, has less intensely coloured body bars, but with the membrane between the third spine of the first dorsal fin and the dorsum with crimson and the anterior portion of the second dorsal fin crimson, as a triangle with the first spine and the bases of first 7 spines forming two of the sides, the third side is an irregular, narrow cream band.
|
||
</paragraph>
|
||
<paragraph id="8B9A36903F64FFDBFF6465A749436668" blockId="8.[151,1436,1437,2036]" lastBlockId="9.[151,1436,151,424]" lastPageId="9" lastPageNumber="45" pageId="8" pageNumber="44">
|
||
It is now well established that there are a number of tripterygiid species complexes, with closely related species often differing little in meristic counts and measurements, but in subtle ways in colour patterns. For example, the three currently recognised species of the
|
||
<taxonomicName id="4C254D133F64FFDAFDC965ED4B4C60A2" ID-CoL="3K482" box="[616,870,2012,2036]" class="Actinopterygii" family="Tripterygiidae" genus="Helcogramma" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="44" phylum="Chordata" rank="species" species="steinitzi">
|
||
<emphasis id="B951EA823F64FFDAFDC965ED4B4C60A2" box="[616,870,2012,2036]" italics="true" pageId="8" pageNumber="44">Helcogramma steinitzi</emphasis>
|
||
</taxonomicName>
|
||
species complex share the micromelanophores of the first dorsal fin and an overall red body colour in males (
|
||
<bibRefCitation id="EFB44B613F65FFDBFC8A62A64BF467F9" author="Holleman" box="[811,990,151,175]" pageId="9" pageNumber="45" refString="Holleman, W. (2006) Fishes of the Helcogramma steinitzi species group (Blennioidei: Tripterygiidae) from the Indian Ocean with descriptions of two new species. aqua, Journal of Ichthyology and Aquatic Biology, 11 (3), 89 - 104." type="journal article" year="2006">Holleman 2006</bibRefCitation>
|
||
; see also
|
||
<bibRefCitation id="EFB44B613F65FFDBFBE662A64DB667F9" author="Williams" box="[1095,1436,151,175]" pageId="9" pageNumber="45" refString="Williams, J. T. & McCormick, C. J. (1990) Two new species of the triplefin fish genus Helcogramma (Tripterygiidae) from the Western Pacific Ocean. Copeia, 1990 (4), 1020 - 1030." type="journal article" year="1990">Williams & McCormick, 1990</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFB44B613F65FFDBFF68628A49C06785" author="Williams" box="[201,490,187,211]" pageId="9" pageNumber="45" refString="Williams, J. T. & Howe, J. C. (2003) Seven new species of the triplefin genus Helcogramma (Tripterygiidae) from the Indo - Pacific. Aqua, Journal of Ichthyology and Aquatic Biology, 7 (4), 151 - 176." type="journal article" year="2003">Williams & Howe, 2003</bibRefCitation>
|
||
). The species of the
|
||
<taxonomicName id="4C254D133F65FFDBFD7F628D4BC46785" ID-CoL="39W3G" box="[734,1006,187,211]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="45" phylum="Chordata" rank="species" species="tutuilae">
|
||
<emphasis id="B951EA823F65FFDBFD7F628D4BC46785" box="[734,1006,187,211]" italics="true" pageId="9" pageNumber="45">Enneapterygius tutuilae</emphasis>
|
||
</taxonomicName>
|
||
species complex share the following characters: only simple pectoral-fin rays, a characteristic black area on the second dorsal fin, followed by a pale to golden band that runs from the fin onto the body, to or just beyond the midline, and six or seven dark or black bands across the anal fin.
|
||
</paragraph>
|
||
<paragraph id="8B9A36903F65FFDBFF646378491166FE" blockId="9.[151,1436,151,424]" pageId="9" pageNumber="45">
|
||
On the basis of the difference in colour pattern and distribution
|
||
<taxonomicName id="4C254D133F65FFDBFC2E637B4CDF6637" ID-CoL="39VYL" authority="Clark" authorityName="Clark" box="[911,1269,329,353]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="45" phylum="Chordata" rank="species" species="altipinnis">
|
||
<emphasis id="B951EA823F65FFDBFC2E637B4C9A6637" box="[911,1200,329,353]" italics="true" pageId="9" pageNumber="45">Enneapterygius altipinnis</emphasis>
|
||
Clark
|
||
</taxonomicName>
|
||
and is recognised as a valid species, distinct from similar fish from the east coast of Africa and westernmost Western Indian Ocean islands.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5A66183F65FFDBFF3664D74B93606C" httpUri="https://zenodo.org/record/279578/files/figure.png" pageId="9" pageNumber="45" targetBox="[151,1436,467,1753]" targetPageId="9">
|
||
<paragraph id="8B9A36903F65FFDBFF3664D74B93606C" blockId="9.[151,1436,1766,1850]" pageId="9" pageNumber="45">
|
||
<emphasis id="B951EA823F65FFDBFF3664D7493961AA" bold="true" box="[151,275,1766,1789]" pageId="9" pageNumber="45">FIGURE 5.</emphasis>
|
||
(A)
|
||
<emphasis id="B951EA823F65FFDBFEE564D94A0961AB" box="[324,547,1767,1789]" italics="true" pageId="9" pageNumber="45">
|
||
<taxonomicName id="4C254D133F65FFDBFEE564D94A3561AB" box="[324,543,1767,1789]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="45" phylum="Chordata" rank="species" species="abeli">Enneapterygius abeli</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
M, El Quseir; (B)
|
||
<taxonomicName id="4C254D133F65FFDBFD4464D94B9461AB" box="[741,958,1767,1789]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="45" phylum="Chordata" rank="species" species="abeli">
|
||
<emphasis id="B951EA823F65FFDBFD4464D94B9461AB" box="[741,958,1767,1789]" italics="true" pageId="9" pageNumber="45">Enneapterygius abeli</emphasis>
|
||
</taxonomicName>
|
||
, F, Marsa Alam; (C)
|
||
<emphasis id="B951EA823F65FFDBFB3864D94D5261AB" box="[1177,1400,1767,1789]" italics="true" pageId="9" pageNumber="45">
|
||
<taxonomicName id="4C254D133F65FFDBFB3864D94D5E61AB" box="[1177,1396,1767,1789]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="45" phylum="Chordata" rank="species" species="abeli">Enneapterygius abeli</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
M, El Quseir; (D)
|
||
<taxonomicName id="4C254D133F65FFDBFE9565374AB0604D" authority="El Quseir" authorityName="El Quseir" box="[308,666,1797,1819]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="45" phylum="Chordata" rank="species" species="clarkae">
|
||
<emphasis id="B951EA823F65FFDBFE9565374A0D604D" box="[308,551,1797,1819]" italics="true" pageId="9" pageNumber="45">Enneapterygius clarkae</emphasis>
|
||
, El Quseir
|
||
</taxonomicName>
|
||
; (E)
|
||
<emphasis id="B951EA823F65FFDBFD6E65374B93604D" box="[719,953,1797,1819]" italics="true" pageId="9" pageNumber="45">
|
||
<taxonomicName id="4C254D133F65FFDBFD6E65374B9F604D" box="[719,949,1797,1819]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="45" phylum="Chordata" rank="species" species="destai">Enneapterygius destai</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
M, Al Lith, Saudi Arabia; (F)
|
||
<taxonomicName id="4C254D133F65FFDBFB5C653748F9606C" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="45" phylum="Chordata" rank="species" species="destai">
|
||
<emphasis id="B951EA823F65FFDBFB5C653748F9606C" italics="true" pageId="9" pageNumber="45">Enneapterygius destai</emphasis>
|
||
</taxonomicName>
|
||
, F, Gulf of Aqaba. Photos by J. Herler (F); S.V. Bogorodsky (the others).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33F651B3F65FFD8FF64656E4A2F6783" lastPageId="10" lastPageNumber="46" pageId="9" pageNumber="45" type="materials_examined">
|
||
<paragraph id="8B9A36903F65FFDBFF64656E4D5860BE" blockId="9.[151,1436,1887,2024]" pageId="9" pageNumber="45">
|
||
<emphasis id="B951EA823F65FFDBFF64656E4A7E602E" bold="true" box="[197,596,1887,1912]" pageId="9" pageNumber="45">Comparative material examined.</emphasis>
|
||
<emphasis id="B951EA823F65FFDBFDFC65504BA3602E" box="[605,905,1888,1912]" italics="true" pageId="9" pageNumber="45">
|
||
<taxonomicName id="4C254D133F65FFDBFDFC65504BA9602E" box="[605,899,1888,1912]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="45" phylum="Chordata" rank="species" species="altipinnis">Enneapterygius altipinnis</taxonomicName>
|
||
:
|
||
</emphasis>
|
||
<collectionCode id="ED34AE553F65FFDBFC3265514BCD602E" LSID="urn:lsid:biocol.org:col:34984" box="[915,999,1888,1912]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="9" pageNumber="45">SAIAB</collectionCode>
|
||
88749 (3:
|
||
<quantity id="4CDD9B753F65FFDBFBC965514D24602E" box="[1128,1294,1888,1912]" metricMagnitude="-2" metricUnit="m" metricValue="1.855" metricValueMax="2.03" metricValueMin="1.68" pageId="9" pageNumber="45" unit="mm" value="18.55" valueMax="20.3" valueMin="16.8">16.8–20.3 mm</quantity>
|
||
SL), Shams Alam Reef, S.
|
||
<collectingCountry id="F33276003F65FFDBFEE265B749A360C8" box="[323,393,1926,1950]" name="Egypt" pageId="9" pageNumber="45">Egypt</collectingCountry>
|
||
, S. Bogorodsky, 0
|
||
<date id="FF9B10503F65FFDBFDFC65B44AC160CB" box="[605,747,1925,1949]" pageId="9" pageNumber="45" value="2008-06-08">8 June 2008</date>
|
||
;
|
||
<collectionCode id="ED34AE553F65FFDBFD5865B44B6760CB" LSID="urn:lsid:biocol.org:col:34984" box="[761,845,1925,1949]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="9" pageNumber="45">SAIAB</collectionCode>
|
||
88750 (3:
|
||
<quantity id="4CDD9B753F65FFDBFC6F65B44C5960CB" box="[974,1139,1925,1949]" metricMagnitude="-2" metricUnit="m" metricValue="1.895" metricValueMax="1.97" metricValueMin="1.8199999999999998" pageId="9" pageNumber="45" unit="mm" value="18.95" valueMax="19.7" valueMin="18.2">18.2–19.7 mm</quantity>
|
||
SL), El Quseir,
|
||
<collectingCountry id="F33276003F65FFDBFA9365B74D5260C8" box="[1330,1400,1926,1950]" name="Egypt" pageId="9" pageNumber="45">Egypt</collectingCountry>
|
||
, S. Bogorodsky, 0
|
||
<date id="FF9B10503F65FFDBFE9D659B49E26094" box="[316,456,1962,1986]" pageId="9" pageNumber="45" value="2008-06-08">8 June 2008</date>
|
||
;
|
||
<collectionCode id="ED34AE553F65FFDBFE74659B4A036094" LSID="urn:lsid:biocol.org:col:34984" box="[469,553,1962,1986]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="9" pageNumber="45">SAIAB</collectionCode>
|
||
88751 (
|
||
<quantity id="4CDD9B753F65FFDBFD29659B4AC46094" box="[648,750,1962,1986]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="9" pageNumber="45" unit="mm" value="16.3">16.3 mm</quantity>
|
||
SL), Shaab Suadi,
|
||
<collectingCountry id="F33276003F65FFDBFC6A659B4C3E6094" box="[971,1044,1962,1986]" name="Sudan" pageId="9" pageNumber="45">Sudan</collectingCountry>
|
||
, S. Bogorodsky,
|
||
<date id="FF9B10503F65FFDBFB74659B4DB26094" box="[1237,1432,1962,1986]" pageId="9" pageNumber="45" value="2009-10-12">12 October 2009</date>
|
||
;
|
||
<collectionCode id="ED34AE553F65FFDBFF3665E148E560BE" LSID="urn:lsid:biocol.org:col:34838" box="[151,207,2000,2024]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="9" pageNumber="45">SMF</collectionCode>
|
||
33608 (
|
||
<quantity id="4CDD9B753F65FFDBFE8A65E149BA60BE" box="[299,400,2000,2024]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="9" pageNumber="45" unit="mm" value="15.1">15.1 mm</quantity>
|
||
SL), Rabigh,
|
||
<collectingCountry id="F33276003F65FFDBFD8C65E14AEA60BE" box="[557,704,2000,2024]" name="Saudi Arabia" pageId="9" pageNumber="45">Saudi Arabia</collectingCountry>
|
||
(
|
||
<geoCoordinate id="EE1150573F65FFDBFD6E65E14B4260BE" box="[719,872,2000,2024]" direction="north" orientation="latitude" pageId="9" pageNumber="45" precision="1" value="23.047318">23°02.839' N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1150573F65FFDBFCD265E14C2D60BE" box="[883,1031,2000,2024]" direction="east" orientation="longitude" pageId="9" pageNumber="45" precision="1" value="38.777016">38°46.621' E</geoCoordinate>
|
||
), S. Bogorodsky, 0
|
||
<date id="FF9B10503F65FFDBFB7C65E14D4460BE" box="[1245,1390,2000,2024]" pageId="9" pageNumber="45" value="2011-04-09">9 April 2011</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9A36903F66FFD8FF6462A84A2F6783" blockId="10.[151,1436,151,213]" pageId="10" pageNumber="46">
|
||
<taxonomicName id="4C254D133F66FFD8FF6462A849DD67E6" box="[197,503,151,176]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="46" phylum="Chordata" rank="species" species="tutuilae">
|
||
<emphasis id="B951EA823F66FFD8FF6462A8495F67E6" box="[197,373,153,176]" italics="true" pageId="10" pageNumber="46">Enneapterygius</emphasis>
|
||
cf.
|
||
<emphasis id="B951EA823F66FFD8FE0062A949DD67E6" box="[417,503,152,176]" italics="true" pageId="10" pageNumber="46">tutuilae</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<collectionCode id="ED34AE553F66FFD8FDA762A64A7067F9" LSID="urn:lsid:biocol.org:col:34984" box="[518,602,151,175]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="10" pageNumber="46">SAIAB</collectionCode>
|
||
31515 (3: 15.0–
|
||
<quantity id="4CDD9B753F66FFD8FCB662A64B5167F9" box="[791,891,151,175]" metricMagnitude="-2" metricUnit="m" metricValue="1.86" pageId="10" pageNumber="46" unit="mm" value="18.6">18.6 mm</quantity>
|
||
SL), Pinda
|
||
<collectingCountry id="F33276003F66FFD8FBA262A64C6067F9" box="[1027,1098,151,175]" name="Iceland" pageId="10" pageNumber="46">Island</collectingCountry>
|
||
,
|
||
<collectingCountry id="F33276003F66FFD8FBF762A64CC467F9" box="[1110,1262,151,175]" name="Mozambique" pageId="10" pageNumber="46">Mozambique</collectingCountry>
|
||
;
|
||
<collectionCode id="ED34AE553F66FFD8FB5B62A64D6467F9" LSID="urn:lsid:biocol.org:col:34984" box="[1274,1358,151,175]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="10" pageNumber="46">SAIAB</collectionCode>
|
||
31520 (14.8 &
|
||
<quantity id="4CDD9B753F66FFD8FF54628C49736783" box="[245,345,189,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.52" pageId="10" pageNumber="46" unit="mm" value="15.2">15.2 mm</quantity>
|
||
SL)
|
||
<collectingCountry id="F33276003F66FFD8FE31628C4A2B6783" box="[400,513,189,213]" name="Mauritius" pageId="10" pageNumber="46">Mauritius</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |