180 lines
19 KiB
XML
180 lines
19 KiB
XML
<document id="D0822D98431629D61D2D9E920B91C417" ID-DOI="10.5281/zenodo.5322469" ID-ISSN="0374-1036" ID-Zenodo-Dep="5322469" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630137619977" checkinUser="felipe" docAuthor="Hlaváč, Peter" docDate="2009" docId="038C5E69506DFF973DBD77B498CBFE52" docLanguage="en" docName="ActaEntMusNatPra.2009.49.2.651-659.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 49 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Bryaxis cepliki Hlaváč 2009, sp. nov." docType="treatment" docVersion="3" lastPageNumber="655" masterDocId="FFB52611506FFF933C42776F9A7CFFE6" masterDocTitle="Taxonomic notes on the Bryaxis splendidus species group (Coleoptera: Staphylinidae: Pselaphinae), with the description of a new species from the Ukraine" masterLastPageNumber="659" masterPageNumber="651" pageNumber="653" updateTime="1699163717770" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="863B5382E446B3A490C2BBF064D1FE19">Taxonomic notes on the Bryaxis splendidus species group (Coleoptera: Staphylinidae: Pselaphinae), with the description of a new species from the Ukraine</mods:title>
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<mods:name id="C23D192907E0D4793CE1DD1F10429484" type="personal">
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<mods:namePart id="59A17B65B79B8FEABE18696145DB3745">Hlaváč, Peter</mods:namePart>
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<mods:title id="8F100B4749388A28F49CE3ADC9394814">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:part id="7A4C1B0834FE62ABE5F2218C330A743E">
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<mods:date id="9BC2F5BE9D914382B982AA7ECAF54072">2009</mods:date>
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<mods:number id="B11429DF8035D7576A05C32634F19058">2009-12-15</mods:number>
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<mods:number id="D998D93BE91FF41FACA090A04132D156">49</mods:number>
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<mods:identifier id="65CC680EAFD0DE4B808143E96ED4AA55" type="DOI">10.5281/zenodo.5322469</mods:identifier>
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<treatment id="038C5E69506DFF973DBD77B498CBFE52" LSID="urn:lsid:plazi:treatment:038C5E69506DFF973DBD77B498CBFE52" httpUri="http://treatment.plazi.org/id/038C5E69506DFF973DBD77B498CBFE52" lastPageId="4" lastPageNumber="655" pageId="2" pageNumber="653">
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<subSubSection id="C33FBCF4506DFF913DBD77B498C4FEEA" pageId="2" pageNumber="653" type="nomenclature">
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<paragraph id="8B9AEF7F506DFF913DBD77B49883FF15" blockId="2.[511,767,218,243]" box="[511,767,218,243]" pageId="2" pageNumber="653">
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<heading id="D0D25813506DFF913DBD77B49883FF15" bold="true" box="[511,767,218,243]" centered="true" fontSize="8" level="2" pageId="2" pageNumber="653" reason="3">
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<emphasis id="B951336D506DFF913DBD77B49883FF15" bold="true" box="[511,767,218,243]" pageId="2" pageNumber="653">
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<taxonomicName id="4C2594FC506DFF913DBD77B498DFFF15" ID-CoL="NGFT" authority="Hlaváč, 2009" authorityName="Hlaváč" authorityYear="2009" box="[511,675,218,243]" class="Insecta" family="Staphylinidae" genus="Bryaxis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="653" phylum="Arthropoda" rank="species" species="cepliki" status="sp. nov.">
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<emphasis id="B951336D506DFF913DBD77B498DFFF15" bold="true" box="[511,675,218,243]" italics="true" pageId="2" pageNumber="653">Bryaxis cepliki</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2628E16506DFF913EEB77B49883FF15" box="[681,767,219,243]" pageId="2" pageNumber="653" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9AEF7F506DFF913E07779698C4FEEA" blockId="2.[581,696,249,269]" box="[581,696,249,269]" pageId="2" pageNumber="653">(Figs. 1–4, 8)</paragraph>
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</subSubSection>
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<subSubSection id="C33FBCF4506DFF973CD5764C99D4FF12" lastPageId="4" lastPageNumber="655" pageId="2" pageNumber="653" type="materials_examined">
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<paragraph id="8B9AEF7F506DFF913CD5764C9902FED1" blockId="2.[151,1127,290,398]" box="[151,894,290,311]" pageId="2" pageNumber="653">
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<emphasis id="B951336D506DFF913CD5764C9B6CFED0" bold="true" box="[151,272,290,310]" pageId="2" pageNumber="653">
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<typeStatus id="549E51DD506DFF913CD5764C9AB8FED0" box="[151,196,291,310]" pageId="2" pageNumber="653">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F332AFEF506DFF913D57764C9B20FED0" box="[277,348,291,310]" name="Ukraine" pageId="2" pageNumber="653">Ukraine</collectingCountry>
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, Transcarpathian Ruthenia, Mala Ugolka, Zhemchuzhnaya cave
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</paragraph>
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<paragraph id="8B9AEF7F506DFF913CD576299969FE68" blockId="2.[151,1127,290,398]" pageId="2" pageNumber="653">
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<emphasis id="B951336D506DFF913CD576299B61FEBF" bold="true" box="[151,285,325,345]" pageId="2" pageNumber="653">Type material.</emphasis>
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<materialsCitation id="3B4DE522506DFF913D6176299ACCFE95" collectingDate="2007-07-16" collectorName="D. Ceplik" country="Ukraine" location="Zemcuznaja cave" municipality="Mala Ugolka" pageId="2" pageNumber="653" specimenCount="1" stateProvince="Zakarpattya" typeStatus="holotype">
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<typeStatus id="549E51DD506DFF913D6176299B0AFEB1" box="[291,374,326,345]" pageId="2" pageNumber="653" type="holotype">HOLOTYPE</typeStatus>
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||
: J, ‘
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<collectingCountry id="F332AFEF506DFF913DE27629987BFEBF" box="[416,519,326,345]" name="Ukraine" pageId="2" pageNumber="653">UKRAINE</collectingCountry>
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:
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<collectingRegion id="49E1219D506DFF913E52762A9804FEBE" box="[528,632,325,344]" country="Ukraine" name="Zakarpattya" pageId="2" pageNumber="653">Zakarpattya</collectingRegion>
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,
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<collectingMunicipality id="6BFE7505506DFF913EC3762A9888FEBE" box="[641,756,325,344]" pageId="2" pageNumber="653">Mala Ugolka</collectingMunicipality>
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,
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<location id="8EFAB9A4506DFF913EBC762A99E9FEBF" LSID="urn:lsid:plazi:treatment:038C5E69506DFF973DBD77B498CBFE52:8EFAB9A4506DFF913EBC762A99E9FEBF" box="[766,917,325,345]" country="Ukraine" municipality="Mala Ugolka" name="Zemcuznaja cave" pageId="2" pageNumber="653" stateProvince="Zakarpattya">Žemčužnaja cave</location>
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,
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<date id="FF9BC9BF506DFF913FDD762A9E75FEBE" box="[927,1033,325,344]" pageId="2" pageNumber="653" value="2007-07-16">
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<collectingDate id="EFDF3057506DFF913FDD762A9E75FEBE" box="[927,1033,325,344]" pageId="2" pageNumber="653" value="2007-07-16">16.VII.2007</collectingDate>
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</date>
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,
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<collectorName id="26D08AA9506DFF91385076299E1BFEBE" box="[1042,1127,325,345]" pageId="2" pageNumber="653">D. Čeplík</collectorName>
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lgt
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</materialsCitation>
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. /
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<materialsCitation id="3B4DE522506DFF913C86760F9971FE95" box="[196,781,352,371]" collectionCode="NMPC" country="Ukraine" determinerName="P. Hlavac" pageId="2" pageNumber="653" specimenCount="1" stateProvince="Zakarpattya" typeStatus="holotype">
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<typeStatus id="549E51DD506DFF913C86760F9B49FE95" box="[196,309,352,371]" pageId="2" pageNumber="653" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="4C2594FC506DFF913D7B760E9BC5FE95" authority="Hlaváč, 2009" authorityName="Hlaváč" authorityYear="2009" box="[313,441,353,371]" class="Insecta" family="Staphylinidae" genus="Bryaxis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="653" phylum="Arthropoda" rank="species" species="cepliki" status="sp. nov.">
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<emphasis id="B951336D506DFF913D7B760E9BC5FE95" box="[313,441,353,371]" italics="true" pageId="2" pageNumber="653">Bryaxis cepliki</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2628E16506DFF913DFC760F987FFE95" box="[446,515,352,371]" pageId="2" pageNumber="653" rank="species">sp. nov.</taxonomicNameLabel>
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,
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<determinerName id="7E4603BB506DFF913E48760F9826FE95" box="[522,602,352,371]" pageId="2" pageNumber="653">P. Hlaváč</determinerName>
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det., 2008’ (
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<collectionCode id="ED3477BA506DFF913E84760F997BFE95" box="[710,775,352,371]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="653">NMPC</collectionCode>
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)
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</materialsCitation>
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.
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<typeStatus id="549E51DD506DFF913F57760F9918FE94" box="[789,868,352,371]" pageId="2" pageNumber="653" type="paratype">PARATYPE</typeStatus>
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: J, ‘
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<collectingCountry id="F332AFEF506DFF913FCE760F998FFE95" box="[908,1011,352,371]" name="Ukraine" pageId="2" pageNumber="653">UKRAINE</collectingCountry>
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:
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<collectingRegion id="49E1219D506DFF913FB9760F9E1FFE95" box="[1019,1123,352,371]" country="Ukraine" name="Zakarpattya" pageId="2" pageNumber="653">Zakarpattya</collectingRegion>
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, Velika Ugolka, cave Moločnyj kameň,
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<date id="FF9BC9BF506DFF913DA776149832FE68" box="[485,590,379,398]" pageId="2" pageNumber="653" value="2007-07-15">15.VII.2007</date>
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, D. Čeplík lgt.’ (
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<collectionCode id="ED3477BA506DFF913E9F76149977FE68" box="[733,779,379,398]" country="USA" lsid="urn:lsid:biocol.org:col:15612" name="University of the Pacific" pageId="2" pageNumber="653" type="Herbarium">CPH</collectionCode>
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).
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</paragraph>
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<caption id="DF5ABFF7506CFF903C3372969B85F9BA" ID-DOI="http://doi.org/10.5281/zenodo.5322471" ID-Zenodo-Dep="5322471" httpUri="https://zenodo.org/record/5322471/files/figure.png" pageId="3" pageNumber="654" startId="3.[113,152,1529,1548]" targetBox="[151,1044,219,1484]" targetPageId="3">
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||
<paragraph id="8B9AEF7F506CFF903C3372969B85F9BA" blockId="3.[113,1089,1529,1628]" pageId="3" pageNumber="654">
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Figs. 2–7. 2–4 –
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<taxonomicName id="4C2594FC506CFF903D4172959BFFF9EA" authority="Hlaváč, 2009" authorityName="Hlaváč" authorityYear="2009" box="[259,387,1530,1548]" class="Insecta" family="Staphylinidae" genus="Bryaxis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="654" phylum="Arthropoda" rank="species" species="cepliki" status="sp. nov.">
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<emphasis id="B951336D506CFF903D4172959BFFF9EA" box="[259,387,1530,1548]" italics="true" pageId="3" pageNumber="654">Bryaxis cepliki</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2628E16506CFF903DCB72969BB0F9EA" box="[393,460,1529,1548]" pageId="3" pageNumber="654" rank="species">sp. nov.</taxonomicNameLabel>
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2 – scape and pedicel; 3 – aedeagus, dorsal aspect; 4 – aedeagus, ventral aspect. 5 –
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<taxonomicName id="4C2594FC506CFF903C92717B9BD7F9C0" authority="(Jeannel, 1922)" baseAuthorityName="Jeannel" baseAuthorityYear="1922" box="[208,427,1555,1574]" class="Insecta" family="Staphylinidae" genus="Bryaxis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="654" phylum="Arthropoda" rank="species" species="goliath">
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<emphasis id="B951336D506CFF903C92717B9B5FF9C0" box="[208,291,1556,1574]" italics="true" pageId="3" pageNumber="654">B. goliath</emphasis>
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(
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<bibRefCitation id="EFB4928E506CFF903D6C717C9BD9F9C0" author="JEANNEL R." box="[302,421,1555,1574]" pageId="3" pageNumber="654" pagination="232 - 237" refId="ref3447" refString="JEANNEL R. 1922: Megalobythus Goliath Pselaphide cavernicole nouveau des Monts Bihor. Buletinul Societatii de Stiinte din Cluj 1: 232 - 237." type="journal article" year="1922">Jeannel, 1922</bibRefCitation>
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)
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</taxonomicName>
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, aedeagus, dorsal aspect (reproduced from
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<bibRefCitation id="EFB4928E506CFF903F58717B99CCF9C0" author="JEANNEL R." box="[794,944,1555,1575]" pageId="3" pageNumber="654" refId="ref3473" refString="JEANNEL R. 1955: L'edeage. Initiation aux recherches sur la systematique des Coleopteres. Publications du Museum National d'Histoire Naturelle No. 16, Paris, 155 pp." type="book" year="1955">JEANNEL 1955: 40</bibRefCitation>
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). 6 –
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<taxonomicName id="4C2594FC506CFF903F9F717B9B49F9A7" authority="(Stolz, 1923)" baseAuthorityName="Stolz" baseAuthorityYear="1923" class="Insecta" family="Staphylinidae" genus="Bryaxis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="654" phylum="Arthropoda" rank="species" species="monstrosetibialis">
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<emphasis id="B951336D506CFF903F9F717B9AC2F9A7" italics="true" pageId="3" pageNumber="654">B. monstrosetibialis</emphasis>
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(
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<bibRefCitation id="EFB4928E506CFF903C8871419B53F9A7" author="STOLZ" box="[202,303,1582,1601]" pageId="3" pageNumber="654" pagination="71 - 72" refId="ref3960" refString="STOLZ, H. 1923: Bythinus monstrosetibialis Stolz nov. spec. [Beschreibung eines neuen Bythinus aus Nordungarn]. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien 73: 71 - 72." type="journal article" year="1923">Stolz, 1923</bibRefCitation>
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||
)
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</taxonomicName>
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, aedeagus, dorsal aspect (from
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<bibRefCitation id="EFB4928E506CFF903E00714198ABF9A7" author="HLAVAC P." box="[578,727,1582,1601]" pageId="3" pageNumber="654" pagination="37 - 40" refId="ref3419" refString="HLAVAC P. 2000: Distribution of the species Bryaxis monstrosetibialis in Slovakia (Coleoptera: Staphylinidae: Pselaphinae). Klapalekiana 36: 37 - 40." type="journal article" year="2000">HLAVÁČ 2000: 38</bibRefCitation>
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||
). 7 –
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<taxonomicName id="4C2594FC506CFF903F4471409E41F9A7" authority="(Croissandeau, 1891)" baseAuthorityName="Croissandeau" baseAuthorityYear="1891" box="[774,1085,1582,1601]" class="Insecta" family="Staphylinidae" genus="Bryaxis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="654" phylum="Arthropoda" rank="species" species="splendidus">
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<emphasis id="B951336D506CFF903F4471409905F9A7" box="[774,889,1583,1601]" italics="true" pageId="3" pageNumber="654">B. splendidus</emphasis>
|
||
(
|
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<bibRefCitation id="EFB4928E506CFF903FC771419E4AF9A7" author="CROISSANDEAU J." box="[901,1078,1582,1601]" pageId="3" pageNumber="654" pagination="130 - 144" refId="ref3272" refString="CROISSANDEAU J. 1891: Etude sur les Bythinus gallo-rhenans et corses et description de plusieurs especes nouvelles. Coleopteriste 9: 130 - 144, pls. 3 - 4." type="journal article" year="1891">Croissandeau, 1891</bibRefCitation>
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||
)
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</taxonomicName>
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, aedeagus, dorsal aspect. Scale bars = 0.2 mm.
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</paragraph>
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</caption>
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<paragraph id="8B9AEF7F506BFF973CD577B399D4FF12" blockId="4.[151,1127,220,436]" box="[151,936,220,244]" pageId="4" pageNumber="655">
|
||
the
|
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<taxonomicName id="4C2594FC506BFF973CFC77B29B32FF12" baseAuthorityName="Croissandeau" baseAuthorityYear="1891" box="[190,334,221,244]" class="Insecta" family="Staphylinidae" genus="Bryaxis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="655" phylum="Arthropoda" rank="species" species="splendidus">
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<emphasis id="B951336D506BFF973CFC77B29B32FF12" box="[190,334,221,244]" italics="true" pageId="4" pageNumber="655">B. splendidus</emphasis>
|
||
</taxonomicName>
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species group by the shape of the aedeagus (
|
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<figureCitation id="131EF3FA506BFF973F7677B399E7FF12" box="[820,923,220,244]" captionStart="Figs" captionStartId="3.[113,152,1529,1548]" captionTargetBox="[151,1044,219,1484]" captionTargetId="figure-14@3.[113,1055,216,1501]" captionTargetPageId="3" captionText="Figs. 2–7. 2–4 – Bryaxis cepliki sp. nov. 2 – scape and pedicel; 3 – aedeagus, dorsal aspect; 4 – aedeagus, ventral aspect. 5 – B. goliath (Jeannel, 1922), aedeagus, dorsal aspect (reproduced from JEANNEL 1955: 40). 6 – B. monstrosetibialis (Stolz, 1923), aedeagus, dorsal aspect (from HLAVÁČ 2000: 38). 7 – B. splendidus (Croissandeau, 1891), aedeagus, dorsal aspect. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5322471" httpUri="https://zenodo.org/record/5322471/files/figure.png" pageId="4" pageNumber="655">Figs. 3, 4</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C33FBCF4506BFF973CD5779398C1FED2" pageId="4" pageNumber="655" type="etymology">
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<paragraph id="8B9AEF7F506BFF973CD5779398C1FED2" blockId="4.[151,1127,220,436]" pageId="4" pageNumber="655">
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<emphasis id="B951336D506BFF973CD577939B64FEF2" bold="true" box="[151,280,252,276]" pageId="4" pageNumber="655">Etymology.</emphasis>
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Named after my friend David Čeplík, Košice, a very enthusiastic biospeleologist and collector of both specimens of the new species.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33FBCF4506BFF973CD5765499E8FE75" pageId="4" pageNumber="655" type="biology_ecology">
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<paragraph id="8B9AEF7F506BFF973CD5765499E8FE75" blockId="4.[151,1127,220,436]" pageId="4" pageNumber="655">
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<emphasis id="B951336D506BFF973CD576549A8FFEB5" bold="true" box="[151,243,315,339]" pageId="4" pageNumber="655">Biology.</emphasis>
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The
|
||
<typeStatus id="549E51DD506BFF973D6876539BFBFEB2" box="[298,391,316,340]" pageId="4" pageNumber="655" type="holotype">holotype</typeStatus>
|
||
was collected in wet scree near the wall at the end of the Zhemchuzhnaya cave, about
|
||
<quantity id="4CDD429A506BFF973D3076349BD4FE95" box="[370,424,347,371]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="655" unit="m" value="50.0">50 m</quantity>
|
||
from the entrance. The
|
||
<typeStatus id="549E51DD506BFF973EE87633997AFE92" box="[682,774,348,372]" pageId="4" pageNumber="655" type="paratype">paratype</typeStatus>
|
||
was collected under large stones at the end of the Molochniy kamen cave, about
|
||
<quantity id="4CDD429A506BFF973ED1761398B6FE75" box="[659,714,380,404]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="655" unit="m" value="80.0">80 m</quantity>
|
||
from the entrance.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33FBCF4506BFF973CD576F498CBFE52" box="[151,695,411,436]" pageId="4" pageNumber="655" type="distribution">
|
||
<paragraph id="8B9AEF7F506BFF973CD576F498CBFE52" blockId="4.[151,1127,220,436]" box="[151,695,411,436]" pageId="4" pageNumber="655">
|
||
<emphasis id="B951336D506BFF973CD576F49B56FE55" bold="true" box="[151,298,411,435]" pageId="4" pageNumber="655">Distribution.</emphasis>
|
||
<collectingCountry id="F332AFEF506BFF973D7276F39BFAFE52" box="[304,390,412,436]" name="Ukraine" pageId="4" pageNumber="655">Ukraine</collectingCountry>
|
||
(Transcarpathian Ruthenia).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |