263 lines
32 KiB
XML
263 lines
32 KiB
XML
<document id="84C4E34C0C5C40517BDD200DE3E1C28C" ID-CLB-Dataset="28058" ID-DOI="10.11646/zootaxa.4508.1.4" ID-GBIF-Dataset="60d1bc89-5afc-4f3f-91c4-9b1c6da3f778" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606858" ID-ZooBank="DD50AB97-77F5-476F-9FFF-B33B96B43078" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553585449037" checkinUser="plazi" docAuthor="Grischenko, Andrei V., Gordon, Dennis P. & Morozov, Taras B." docDate="2018" docId="038C557E68071077FF564810FCB30382" docLanguage="en" docName="zootaxa.4508.1.4.pdf" docOrigin="Zootaxa 4508 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Fatkullinidae Grischenko, Gordon & Morozov 2018" docType="treatment" docVersion="9" masterDocId="FFB52D0668061075FFC14E45FFF9076A" masterDocTitle="Fatkullina imitata n. sp., second species of a unique cheilostome bryozoan genus with reversed-polarity zooidal budding, and new family Fatkullinidae" updateTime="1698712868930" updateUser="ExternalLinkService">
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<mods:mods id="110CE7DA52B963E2BA315680FFEDDE6C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="6CA960E843B089692B0B7CDC2B40C3AF">
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<mods:title id="ECB5C793DC6351585CF841AA44269C47">Fatkullina imitata n. sp., second species of a unique cheilostome bryozoan genus with reversed-polarity zooidal budding, and new family Fatkullinidae</mods:title>
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<mods:name id="1F3A443640A3A71F12F358DA0A4DDFF3" type="personal">
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<mods:roleTerm id="95828B8109CCEB198A9FDE36FFBE60A0">Author</mods:roleTerm>
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<mods:namePart id="17C1F6EF4DB6AC470618B8843D83535E">Grischenko, Andrei V.</mods:namePart>
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<mods:roleTerm id="35EF1127D1877745BBBB624A3C1CFBDF">Author</mods:roleTerm>
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<mods:namePart id="7392E1F948679CB1B3831CC2F8C7E424">Gordon, Dennis P.</mods:namePart>
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<mods:name id="B1258AFDB4A5454D2153FB882045CD2B" type="personal">
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<mods:roleTerm id="A8BDEADBAC8784F5634F6945B09827CF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="27A061C6511D84541A94B5C3F69408DC">Morozov, Taras B.</mods:namePart>
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<mods:typeOfResource id="C250D1272F8A3B7B5DC16015D791D1C1">text</mods:typeOfResource>
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<mods:title id="639A247F07FD6932A6BD3B8BDC1A7BDE">Zootaxa</mods:title>
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<mods:part id="7C9928FC7F9797FA5A14F93E53912230">
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<mods:date id="F9812D3D11D8BFC069252D0E8F4E7D80">2018</mods:date>
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<mods:number id="181509C18A65ADBFD2D2573CF674C0F0">2018-10-30</mods:number>
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<mods:detail id="BFFEC785F2024FF18CA4F702FF8481BD" type="volume">
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<mods:number id="83BB741D269C70F1882CBED825CB7A92">4508</mods:number>
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<mods:number id="4C30580139A5CA14EF548CA37300FF98">1</mods:number>
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<mods:classification id="8BFAF6E13C6A6BA1D70EE5D08C4D67A5">journal volume</mods:classification>
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<mods:identifier id="8ECEC34A0394FFBBD146A36821DCB4C3" type="CLB-Dataset">28058</mods:identifier>
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<mods:identifier id="EA1101326DB76B1E4E267714AECE4D65" type="DOI">10.11646/zootaxa.4508.1.4</mods:identifier>
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<treatment id="038C557E68071077FF564810FCB30382" ID-DOI="http://doi.org/10.5281/zenodo.5975682" ID-GBIF-Taxon="154495678" ID-Zenodo-Dep="5975682" LSID="urn:lsid:plazi:treatment:038C557E68071077FF564810FCB30382" httpUri="http://treatment.plazi.org/id/038C557E68071077FF564810FCB30382" lastPageId="2" pageId="1">
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<subSubSection id="C33FB7E368071074FF564810FE0C0105" box="[151,501,1621,1647]" pageId="1" type="nomenclature">
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<paragraph id="8B9AE46868071074FF564810FE0C0105" blockId="1.[151,501,1621,1647]" box="[151,501,1621,1647]" pageId="1">
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<heading id="D0D2530468071074FF564810FE0C0105" bold="true" box="[151,501,1621,1647]" fontSize="11" level="1" pageId="1" reason="1">
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<emphasis id="B951387A68071074FF564810FE0C0105" bold="true" box="[151,501,1621,1647]" pageId="1">
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Family
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<taxonomicName id="4C259FEB68071074FF354810FE600105" ID-CoL="84KF4" authorityName="Grischenko, Gordon & Morozov" authorityYear="2018" box="[244,409,1621,1647]" class="Gymnolaemata" family="Fatkullinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="1" phylum="Bryozoa" rank="family">Fatkullinidae</taxonomicName>
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n. fam.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33FB7E368071074FF5648D9FCFD01D8" box="[151,772,1692,1714]" pageId="1" type="materials_examined">
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<paragraph id="8B9AE46868071074FF5648D9FCFD01D8" blockId="1.[151,772,1692,1714]" box="[151,772,1692,1714]" pageId="1">
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<emphasis id="B951387A68071074FF5648D9FEE001D8" bold="true" box="[151,281,1692,1714]" pageId="1">
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<typeStatus id="549E5ACA68071074FF5648D9FF3501D8" box="[151,204,1692,1714]" pageId="1">Type</typeStatus>
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genus:
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</emphasis>
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<taxonomicName id="4C259FEB68071074FEDE48D9FCF801D8" authority="Grischenko, Gordon & Taylor, 1998" authorityName="Grischenko, Gordon & Taylor" authorityYear="1998" box="[287,769,1692,1714]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="1" phylum="Bryozoa" rank="genus">
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<emphasis id="B951387A68071074FEDE48D9FE7001D8" box="[287,393,1692,1714]" italics="true" pageId="1">Fatkullina</emphasis>
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<bibRefCitation id="EFB4999968071074FE4F48D9FCF801D8" author="Grischenko, A. V. & Gordon, D. P. & Taylor, P. D." box="[398,769,1692,1714]" pageId="1" pagination="105 - 117" refId="ref7852" refString="Grischenko, A. V., Gordon, D. P. & Taylor, P. D. (1998) A unique new genus of Cheilostomate Bryozoan with reversed-polarity zooidal budding. Asian Marine Biology, 15, 105 - 117." type="journal article" year="1998">Grischenko, Gordon & Taylor, 1998</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33FB7E368071074FF56489BFD0900C6" pageId="1" type="diagnosis">
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<paragraph id="8B9AE46868071074FF56489BFD0900C6" blockId="1.[151,1437,1758,2036]" pageId="1">
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<emphasis id="B951387A68071074FF56489BFEE8019D" bold="true" box="[151,273,1758,1783]" pageId="1">Diagnosis.</emphasis>
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Colony encrusting, unilaminar, darkly pigmented. Zooids with or without reversed-polarity budding. Frontal shield lepralioid, evenly pseudoporous. Primary orifice with median sinus and weakly developed condylar ridges. Oral spines and avicularia absent. Ooecia, if present, imperforate or with only a single pseudopore. Mural septula uniporous, set in buttressed recesses. Ancestrula externally resembling post-ancestrular zooids, but internally with reduced umbonuloid area suborally that also includes possible vestigial costal elements. Immediate periancestrular zooids part of an ancestrular complex.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33FB7E368071077FF0649F3FCB30382" lastPageId="2" pageId="1" type="discussion">
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<paragraph id="8B9AE46868071077FF0649F3FF11062A" blockId="1.[151,1437,1758,2036]" lastBlockId="2.[151,1437,151,1257]" lastPageId="2" pageId="1">
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<emphasis id="B951387A68071074FF0649F3FEC200A5" bold="true" box="[199,315,1974,1999]" pageId="1">Remarks.</emphasis>
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As noted by
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<bibRefCitation id="EFB4999968071074FE1049F3FCF100A5" author="Zagorsek, K. & Gordon, D. P." box="[465,776,1974,1999]" pageId="1" pagination="595 - 607" refId="ref9374" refString="Zagorsek, K. & Gordon, D. P. (2013) Late Tortonian bryozoans from Mut Basin, Central Anatolian Plateau, southern Turkey. Acta Paleontologica Polonica, 58, 595 - 607." type="journal article" year="2013">Zágoršek & Gordon (2013)</bibRefCitation>
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, the family
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<taxonomicName id="4C259FEB68071074FC4C49F3FB9F00A5" authorityName="Canu & Bassler" authorityYear="1917" box="[909,1126,1974,1999]" class="Gymnolaemata" family="Stomachetosellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="1" phylum="Bryozoa" rank="family">Stomachetosellidae</taxonomicName>
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is very heterogeneous. It is based on the
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<typeStatus id="549E5ACA68071074FE814999FE89009E" box="[320,368,2012,2036]" pageId="1">type</typeStatus>
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genus and species
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<taxonomicName id="4C259FEB68071074FDA4499EFB310099" authority="Canu & Bassler, 1917" authorityName="Canu & Bassler" authorityYear="1917" box="[613,1224,2011,2036]" class="Gymnolaemata" family="Stomachetosellidae" genus="Stomachetosella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="1" phylum="Bryozoa" rank="species" species="crassicollis">
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<emphasis id="B951387A68071074FDA4499EFC52009E" box="[613,939,2011,2036]" italics="true" pageId="1">Stomachetosella crassicollis</emphasis>
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<bibRefCitation id="EFB4999968071074FC78499EFB310099" author="Canu, F. & Bassler, R. S." box="[953,1224,2011,2036]" pageId="1" pagination="1 - 87" refId="ref6783" refString="Canu, F. & Bassler, R. S. (1917) A synopsis of American Early Tertiary cheilostome Bryozoa. United States National Museum Bulletin, 96, 1 - 87, 6 pls." type="book chapter" year="1917">Canu & Bassler, 1917</bibRefCitation>
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</taxonomicName>
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, early Oligocene,
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<collectingRegion id="49E12A8A68041077FF564ED2FEE307DA" box="[151,282,151,176]" country="United States of America" name="Mississippi" pageId="2">Mississippi</collectingRegion>
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,
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<collectingCountry id="F332A4F868041077FEE54EDDFEA607DA" box="[292,351,152,176]" name="United States of America" pageId="2">USA</collectingCountry>
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, which has erect stems that range from subvincularian to bilamellar/flabellate fronds in which the zooids, opening on all sides, have a regularly perforated pseudoporous lepralioid frontal shield. The orifice has a tapering rounded poster and condyles appear to be lacking. Ooecia are evenly pseudoporous, somewhat like the frontal shield. According to
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<bibRefCitation id="EFB4999968041077FE144F41FD230676" author="Canu, F. & Bassler, R. S." box="[469,730,260,285]" pageId="2" pagination="1 - 879" refId="ref6821" refString="Canu, F. & Bassler, R. S. (1920) North American Early Tertiary Bryozoa. United States National Museum Bulletin, 106, 1 - 879, 162 pls." type="book chapter" year="1920">Canu & Bassler (1920)</bibRefCitation>
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there is an inconspicuous avicularium near the orifice in some zooids.
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</paragraph>
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<paragraph id="8B9AE46868041077FF064F09FBCC050A" blockId="2.[151,1437,151,1257]" pageId="2">
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||
<bibRefCitation id="EFB4999968041077FF064F09FDFC060E" author="Winston, J. E. & Hayward, P. J." box="[199,517,332,357]" pageId="2" pagination="1 - 180" refId="ref9329" refString="Winston, J. E. & Hayward, P. J. (2012) The marine bryozoans of the northeast coast of the United States: Maine to Virginia. Virginia Museum of Natural History Memoir, 11, 1 - 180." type="journal article" year="2012">Winston & Hayward (2012)</bibRefCitation>
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compared the characters of
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<taxonomicName id="4C259FEB68041077FC894F08FC1F060F" authorityName="Canu & Bassler" authorityYear="1917" box="[840,998,332,357]" class="Gymnolaemata" family="Stomachetosellidae" genus="Stomachetosella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="species" species="crassicollis">
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<emphasis id="B951387A68041077FC894F08FC1F060F" box="[840,998,332,357]" italics="true" pageId="2">S. crassicollis</emphasis>
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</taxonomicName>
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to those of living species attributed to
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<taxonomicName id="4C259FEB68041077FF564F2AFEB706E2" authorityName="Canu & Bassler" authorityYear="1917" box="[151,334,367,392]" class="Gymnolaemata" family="Stomachetosellidae" genus="Stomachetosella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="genus">
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<emphasis id="B951387A68041077FF564F2AFEB706E2" box="[151,334,367,392]" italics="true" pageId="2">Stomachetosella</emphasis>
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</taxonomicName>
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, concluding that the latter are unrelated to the former. They established a new genus,
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<taxonomicName id="4C259FEB68041077FF564FD1FEC106C7" baseAuthorityName="Grischenko" baseAuthorityYear="2015" box="[151,312,404,429]" class="Gymnolaemata" family="Stomachetosellidae" genus="Stomacrustula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="genus">
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<emphasis id="B951387A68041077FF564FD1FEC106C7" box="[151,312,404,429]" italics="true" pageId="2">Stomacrustula</emphasis>
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</taxonomicName>
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, for all "the Recent species formerly attributed to
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<taxonomicName id="4C259FEB68041077FCB64FD1FBD706C7" authorityName="Canu & Bassler" authorityYear="1917" box="[887,1070,404,429]" class="Gymnolaemata" family="Stomachetosellidae" genus="Stomachetosella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="genus">
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||
<emphasis id="B951387A68041077FCB64FD1FBD706C7" box="[887,1070,404,429]" italics="true" pageId="2">Stomachetosella</emphasis>
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</taxonomicName>
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", making new combinations for three of them—
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<taxonomicName id="4C259FEB68041077FE8B4FF2FD1D06BA" authority="(Busk, 1854)" authorityName="Busk" authorityYear="1854" baseAuthorityName="Busk" baseAuthorityYear="1854" box="[330,740,439,464]" class="Gymnolaemata" family="Stomachetosellidae" genus="Stomacrustula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="species" species="cruenta">
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<emphasis id="B951387A68041077FE8B4FF2FDB006BA" box="[330,585,439,464]" italics="true" pageId="2">Stomacrustula cruenta</emphasis>
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(
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<bibRefCitation id="EFB4999968041077FD994FF2FD2206BA" author="Busk, G." box="[600,731,439,464]" pageId="2" pagination="69 - 124" refId="ref6640" refString="Busk, G. (1854) Catalogue of Marine Polyzoa in the Collection of the British Museum. Part II. Cheilostomata (part). Trustees of the British Museum, London, viii + 66 pp. + 56 pls. [pp. i-viii + 55 - 120, pls. 69 - 124]" type="book chapter" year="1854">Busk, 1854</bibRefCitation>
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)
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</taxonomicName>
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(the designated
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<typeStatus id="549E5ACA68041077FC5E4FFDFC3606BA" box="[927,975,440,464]" pageId="2">type</typeStatus>
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species),
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||
<taxonomicName id="4C259FEB68041077FB804FFDFAB606BA" authority="(Busk, 1860)" authorityName="Busk" authorityYear="1860" baseAuthorityName="Busk" baseAuthorityYear="1860" box="[1089,1359,439,464]" class="Gymnolaemata" family="Stomachetosellidae" genus="Stomacrustula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="species" species="sinuosa">
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<emphasis id="B951387A68041077FB804FFDFB4A06BA" box="[1089,1203,440,464]" italics="true" pageId="2">S. sinuosa</emphasis>
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(
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<bibRefCitation id="EFB4999968041077FB024FF2FABF06BA" author="Busk, G." box="[1219,1350,439,464]" pageId="2" pagination="123 - 125" refId="ref6742" refString="Busk, G. (1860) Descriptions of new species of Polyzoa. Collected by George Barlee Esq., in Shetland. Quarterly Journal of Microscopical Science, 8, 123 - 125, pls 24 - 25." type="journal article" year="1860">Busk, 1860</bibRefCitation>
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)
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</taxonomicName>
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and
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<taxonomicName id="4C259FEB68041077FA464FFDFE64069F" authority="(Powell, 1968)" authorityName="Powell" authorityYear="1968" baseAuthorityName="Powell" baseAuthorityYear="1968" class="Gymnolaemata" family="Stomachetosellidae" genus="Stomacrustula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="species" species="hincksi">
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<emphasis id="B951387A68041077FA464FFDFF1F069F" italics="true" pageId="2">S. hincksi</emphasis>
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(
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<bibRefCitation id="EFB4999968041077FF394F99FE6C069F" author="Powell, N. A." box="[248,405,476,501]" pageId="2" pagination="2269 - 2320" refId="ref9117" refString="Powell, N. A. (1968) Bryozoa (Polyzoa) of Arctic Canada. Journal of the Fisheries Research Board of Canada, 25, 2269 - 2320, 14 pls." type="book chapter" year="1968">Powell, 1968</bibRefCitation>
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)
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</taxonomicName>
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. Without explaining why, they did not make new combinations for the ten other Recent species.
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<taxonomicName id="4C259FEB68041077FF3F4FBAFE660572" baseAuthorityName="Grischenko" baseAuthorityYear="2015" box="[254,415,511,536]" class="Gymnolaemata" family="Stomachetosellidae" genus="Stomacrustula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="genus">
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<emphasis id="B951387A68041077FF3F4FBAFE660572" box="[254,415,511,536]" italics="true" pageId="2">Stomacrustula</emphasis>
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</taxonomicName>
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was not attributed to a family but left
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<emphasis id="B951387A68041077FC544C44FBC10572" box="[917,1080,511,536]" italics="true" pageId="2">incertae sedis</emphasis>
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.
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<taxonomicName id="4C259FEB68041077FB8F4FBAFB160572" baseAuthorityName="Grischenko" baseAuthorityYear="2015" box="[1102,1263,511,536]" class="Gymnolaemata" family="Stomachetosellidae" genus="Stomacrustula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="genus">
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||
<emphasis id="B951387A68041077FB8F4FBAFB160572" box="[1102,1263,511,536]" italics="true" pageId="2">Stomacrustula</emphasis>
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</taxonomicName>
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species differ particularly from
|
||
<taxonomicName id="4C259FEB68041077FE9C4C60FE000557" authorityName="Canu & Bassler" authorityYear="1917" box="[349,505,548,573]" class="Gymnolaemata" family="Stomachetosellidae" genus="Stomachetosella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="species" species="crassicollis">
|
||
<emphasis id="B951387A68041077FE9C4C60FE000557" box="[349,505,548,573]" italics="true" pageId="2">S. crassicollis</emphasis>
|
||
</taxonomicName>
|
||
in that the ooecium, if present, is imperforate or has only a central pore, and there are no avicularia. Note, however, that
|
||
<taxonomicName id="4C259FEB68041077FD834C0DFD4A050A" baseAuthorityName="Busk" baseAuthorityYear="1854" box="[578,691,584,608]" class="Gymnolaemata" family="Stomachetosellidae" genus="Stomacrustula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="species" species="cruenta">
|
||
<emphasis id="B951387A68041077FD834C0DFD4A050A" box="[578,691,584,608]" italics="true" pageId="2">S. cruenta</emphasis>
|
||
</taxonomicName>
|
||
lacks ooecia (
|
||
<bibRefCitation id="EFB4999968041077FC924C02FBD1050A" author="Dick, M. H. & Ross, J. R. P." box="[851,1064,583,608]" pageId="2" pagination="1 - 133" refId="ref7261" refString="Dick, M. H. & Ross, J. R. P. (1988) Intertidal Bryozoa (Cheilostomata) of the Kodiak vicinity, Alaska. Center for Pacific Northwest Studies, Western Washington University, Occasional Paper, 23, 1 - 133." type="journal article" year="1988">Dick & Ross 1988</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9AE46868041077FF064C28FA9C0437" blockId="2.[151,1437,151,1257]" pageId="2">
|
||
Because
|
||
<taxonomicName id="4C259FEB68041077FEED4C29FE3405EF" baseAuthorityName="Grischenko" baseAuthorityYear="2015" box="[300,461,620,645]" class="Gymnolaemata" family="Stomachetosellidae" genus="Stomacrustula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="B951387A68041077FEED4C29FE3405EF" box="[300,461,620,645]" italics="true" pageId="2">Stomacrustula</emphasis>
|
||
</taxonomicName>
|
||
has much in common with
|
||
<emphasis id="B951387A68041077FCCF4C29FC7005EF" box="[782,905,620,645]" italics="true" pageId="2">
|
||
<taxonomicName id="4C259FEB68041077FCCF4C29FC7F05EF" authorityName="Grischenko, Gordon & Taylor" authorityYear="1998" box="[782,902,620,645]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="genus">Fatkullina</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
we provisionally include it in
|
||
<taxonomicName id="4C259FEB68041077FB284C29FA8405EF" authorityName="Grischenko, Gordon & Morozov" authorityYear="2018" box="[1257,1405,620,645]" class="Gymnolaemata" family="Fatkullinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="family">Fatkullinidae</taxonomicName>
|
||
<emphasis id="B951387A68041077FA444C29FF3505C2" bold="true" pageId="2">n. fam.</emphasis>
|
||
pending confirmatory gene-sequence data.
|
||
<emphasis id="B951387A68041077FD154CD4FCBA05C2" box="[724,835,655,680]" italics="true" pageId="2">Inter alia</emphasis>
|
||
, species in both genera have a very similar darkpigmented, unilaminar encrusting colony form and zooidal frontal shield and they mutually lack oral spines and avicularia. Additionally, both have deeply recessed septular pores.
|
||
<bibRefCitation id="EFB4999968041077FC1E4C92FAEA059A" author="Grischenko, A. V. & Gordon, D. P. & Taylor, P. D." box="[991,1299,727,752]" pageId="2" pagination="105 - 117" refId="ref7852" refString="Grischenko, A. V., Gordon, D. P. & Taylor, P. D. (1998) A unique new genus of Cheilostomate Bryozoan with reversed-polarity zooidal budding. Asian Marine Biology, 15, 105 - 117." type="journal article" year="1998">
|
||
Grischenko
|
||
<emphasis id="B951387A68041077FBB24C9CFB56059A" box="[1139,1199,727,752]" italics="true" pageId="2">et al</emphasis>
|
||
. (1998)
|
||
</bibRefCitation>
|
||
previously hypothesized that an ooecium-lacking form like
|
||
<taxonomicName id="4C259FEB68041077FD104CB8FCB1047E" baseAuthorityName="Busk" baseAuthorityYear="1854" box="[721,840,765,788]" class="Gymnolaemata" family="Stomachetosellidae" genus="Stomacrustula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="species" species="cruenta">
|
||
<emphasis id="B951387A68041077FD104CB8FCB1047E" box="[721,840,765,788]" italics="true" pageId="2">S. cruenta</emphasis>
|
||
</taxonomicName>
|
||
may have been ancestral to
|
||
<taxonomicName id="4C259FEB68041077FB614CB9FAED047F" authorityName="Grischenko, Gordon & Taylor" authorityYear="1998" box="[1184,1300,764,789]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="B951387A68041077FB614CB9FAED047F" box="[1184,1300,764,789]" italics="true" pageId="2">Fatkullina</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C259FEB68041077FAE64CB9FA65047F" authorityName="Grischenko, Gordon & Taylor" authorityYear="1998" box="[1319,1436,764,789]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="B951387A68041077FAE64CB9FA65047F" box="[1319,1436,764,789]" italics="true" pageId="2">Fatkullina</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C259FEB68041077FEEC4D5AFE370452" baseAuthorityName="Grischenko" baseAuthorityYear="2015" box="[301,462,799,824]" class="Gymnolaemata" family="Stomachetosellidae" genus="Stomacrustula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="B951387A68041077FEEC4D5AFE370452" box="[301,462,799,824]" italics="true" pageId="2">Stomacrustula</emphasis>
|
||
</taxonomicName>
|
||
most obviously in having reversed-polarity zooidal budding, but there are also no ooecia in
|
||
<taxonomicName id="4C259FEB68041077FEC54D01FE800437" authorityName="Grischenko, Gordon & Taylor" authorityYear="1998" box="[260,377,836,861]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="B951387A68041077FEC54D01FE800437" box="[260,377,836,861]" italics="true" pageId="2">Fatkullina</emphasis>
|
||
</taxonomicName>
|
||
whereas they are well-developed in
|
||
<taxonomicName id="4C259FEB68041077FCD24D01FC4A0437" baseAuthorityName="Grischenko" baseAuthorityYear="2015" box="[787,947,836,861]" class="Gymnolaemata" family="Stomachetosellidae" genus="Stomacrustula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="B951387A68041077FCD24D01FC4A0437" box="[787,947,836,861]" italics="true" pageId="2">Stomacrustula</emphasis>
|
||
</taxonomicName>
|
||
(except, as noted, in the
|
||
<typeStatus id="549E5ACA68041077FB0D4D00FB050437" box="[1228,1276,837,861]" pageId="2">type</typeStatus>
|
||
species).
|
||
</paragraph>
|
||
<paragraph id="8B9AE46868041077FF064D2DFCB30382" blockId="2.[151,1437,151,1257]" pageId="2">
|
||
Some other living taxa attributed to the
|
||
<taxonomicName id="4C259FEB68041077FD7F4D22FC6E04EA" authorityName="Canu & Bassler" authorityYear="1917" box="[702,919,871,896]" class="Gymnolaemata" family="Stomachetosellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="family">Stomachetosellidae</taxonomicName>
|
||
can also be provisionally included in the
|
||
<taxonomicName id="4C259FEB68041077FF564DC9FED604CF" authorityName="Grischenko, Gordon & Morozov" authorityYear="2018" box="[151,303,908,933]" class="Gymnolaemata" family="Fatkullinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="family">Fatkullinidae</taxonomicName>
|
||
. For example, the monotypic Arctic genus
|
||
<taxonomicName id="4C259FEB68041077FCCC4DC9FBD504CF" authority="Kluge, 1962" authorityName="Kluge" authorityYear="1962" box="[781,1068,908,933]" class="Gymnolaemata" family="Stomachetosellidae" genus="Lepralioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="B951387A68041077FCCC4DC9FC6304CF" box="[781,922,908,933]" italics="true" pageId="2">Lepralioides</emphasis>
|
||
<bibRefCitation id="EFB4999968041077FC604DC9FBD504CF" author="Kluge, G. A." box="[929,1068,908,933]" pageId="2" pagination="1 - 584" refId="ref8537" refString="Kluge, G. A. (1962) Bryozoa of the northern seas of the USSR. Opredeliteli po Faune SSSR, 76, 1 - 584." type="journal article" year="1962">Kluge, 1962</bibRefCitation>
|
||
</taxonomicName>
|
||
is very similar to
|
||
<taxonomicName id="4C259FEB68041077FB354DC9FA6C04CF" baseAuthorityName="Grischenko" baseAuthorityYear="2015" box="[1268,1429,908,933]" class="Gymnolaemata" family="Stomachetosellidae" genus="Stomacrustula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="B951387A68041077FB354DC9FA6C04CF" box="[1268,1429,908,933]" italics="true" pageId="2">Stomacrustula</emphasis>
|
||
</taxonomicName>
|
||
, differing only in having a very broad, shallow orificial poster instead of a discrete sinus, and the single-pored ooecium is covered by secondary calcification (with sutures) formed by 2–3 neighboring zooids, which is not the case in
|
||
<taxonomicName id="4C259FEB68041077FF2C4DB2FE77037A" baseAuthorityName="Grischenko" baseAuthorityYear="2015" box="[237,398,1015,1040]" class="Gymnolaemata" family="Stomachetosellidae" genus="Stomacrustula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="B951387A68041077FF2C4DB2FE77037A" box="[237,398,1015,1040]" italics="true" pageId="2">Stomacrustula</emphasis>
|
||
</taxonomicName>
|
||
. The Boreal-Arctic, North Pacific genus
|
||
<taxonomicName id="4C259FEB68041077FCA24DB2FB8E037A" authority="Osburn, 1952" authorityName="Osburn" authorityYear="1952" box="[867,1143,1015,1040]" class="Gymnolaemata" family="Stomachetosellidae" genus="Pachyegis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="B951387A68041077FCA24DB2FC2F037A" box="[867,982,1015,1040]" italics="true" pageId="2">Pachyegis</emphasis>
|
||
<bibRefCitation id="EFB4999968041077FC1F4DB2FB8E037A" author="Osburn, R. C." box="[990,1143,1015,1040]" pageId="2" pagination="271 - 611" refId="ref9040" refString="Osburn, R. C. (1952) Bryozoa of the Pacific coast of America part 2, Cheilostomata-Ascophora. Allan Hancock Pacific Expeditions, 14, 271 - 611." type="journal article" year="1952">Osburn, 1952</bibRefCitation>
|
||
</taxonomicName>
|
||
also shares several of the characters of
|
||
<taxonomicName id="4C259FEB68041077FEEF4A59FE5A035F" authorityName="Grischenko, Gordon & Taylor" authorityYear="1998" box="[302,419,1052,1077]" class="Gymnolaemata" family="Stomachetosellidae" genus="Fatkullina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="B951387A68041077FEEF4A59FE5A035F" box="[302,419,1052,1077]" italics="true" pageId="2">Fatkullina</emphasis>
|
||
</taxonomicName>
|
||
—the
|
||
<typeStatus id="549E5ACA68041077FE284A58FDE0035F" box="[489,537,1053,1077]" pageId="2">type</typeStatus>
|
||
species,
|
||
<taxonomicName id="4C259FEB68041077FD414A58FC50035E" authority="Norman, 1903" authorityName="Norman" authorityYear="1903" box="[640,937,1052,1076]" class="Gymnolaemata" family="Stomachetosellidae" genus="Pachyegis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="species" species="princeps">
|
||
<emphasis id="B951387A68041077FD414A58FD05035E" box="[640,764,1053,1076]" italics="true" pageId="2">P. princeps</emphasis>
|
||
<bibRefCitation id="EFB4999968041077FCC54A58FC50035E" author="Norman, A. M." box="[772,937,1052,1076]" pageId="2" pagination="567 - 598" refId="ref8900" refString="Norman, A. M. (1903) Notes on the natural history of East Finmark, Polyzoa. Annals and Magazine of Natural History, Series 7, 11, 567 - 598, pl. 13." type="journal article" year="1903">Norman, 1903</bibRefCitation>
|
||
</taxonomicName>
|
||
, has reddish-brown to black pigmentation, a thickly calcified pseudoporous frontal shield, communication pores located in buttressed recesses and no oral spines. It is unclear if a suboral structure in some zooids is an avicularium or an opening pertaining to a gland. The ooecium resembles that in
|
||
<taxonomicName id="4C259FEB68041077FE384AC2FD7F03CA" authorityName="Kluge" authorityYear="1962" box="[505,646,1159,1184]" class="Gymnolaemata" family="Stomachetosellidae" genus="Lepralioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="B951387A68041077FE384AC2FD7F03CA" box="[505,646,1159,1184]" italics="true" pageId="2">Lepralioides</emphasis>
|
||
</taxonomicName>
|
||
(http://www.iopan.gda.pl/ekologia/Atlas_of_Arctic_
|
||
<taxonomicName id="4C259FEB68041077FB204AC2FAB903CA" box="[1249,1344,1159,1184]" higherTaxonomySource="GBIF" kingdom="Animalia" order="Bryozoa" pageId="2" phylum="Bryozoa" rank="order">Bryozoa</taxonomicName>
|
||
/robaki/ Porella_princeps/
|
||
<typeStatus id="549E5ACA68041077FE9E4AE9FE6003AF" box="[351,409,1196,1221]" pageId="2">Type</typeStatus>
|
||
/index.html). Further work needs to be carried out to determine what other putative stomachetosellids, if any, may be transferred to
|
||
<taxonomicName id="4C259FEB68041077FD6F4A8AFCBF0382" authorityName="Grischenko, Gordon & Morozov" authorityYear="2018" box="[686,838,1231,1256]" class="Gymnolaemata" family="Fatkullinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" phylum="Bryozoa" rank="family">Fatkullinidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |