treatments-xml/data/03/87/76/038776236263FFC42DEFB058FAD95E4F.xml
2024-06-21 12:22:17 +02:00

583 lines
72 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="8614C428D89344C5BAA92395B2211161" ID-CLB-Dataset="263342" ID-DOI="10.11646/zootaxa.5286.1.1" ID-GBIF-Dataset="67e7a105-1f62-414a-83d9-f758039eb726" ID-ISSN="1175-5326" ID-Zenodo-Dep="7958937" ID-ZooBank="F9834684-24D3-4795-B5EB-77B451DF856D" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1684793492880" checkinUser="plazi" docAuthor="Shi, Hongliang &amp; Liang, Hongbin" docDate="2023" docId="038776236263FFC42DEFB058FAD95E4F" docLanguage="en" docName="zootaxa.5286.1.1.pdf" docOrigin="Zootaxa 5286 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5286.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Parena (Crossoglossa) testacea" docType="treatment" docVersion="8" lastPageNumber="40" masterDocId="FFBE0E5B6246FFE32D78B27BFFE75D5A" masterDocTitle="Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)" masterLastPageNumber="144" masterPageNumber="1" pageNumber="38" updateTime="1699506634024" updateUser="plazi">
<mods:mods id="0BC22048B5A301EE07323F9E445DB306" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FCC67D01E12ED52719C4D4F637211ED6">
<mods:title id="9D10D47B23313E43A577AAD884B5479E">Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)</mods:title>
</mods:titleInfo>
<mods:name id="4293F1C749FA8A5A635EE2A94451919B" type="personal">
<mods:role id="89350F5E812F33BAE561AA20EEAF07BD">
<mods:roleTerm id="D03DEA6A90F518AF96767B37E7E96D28">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0890F96EC1AED8B04D1415FE428C63A8">Shi, Hongliang</mods:namePart>
<mods:nameIdentifier id="5527B6A396A470DF4780904D816524A9" type="ORCID">0000-0002-9989-5830</mods:nameIdentifier>
<mods:nameIdentifier id="ED8BDFA8CA37A400FBDD9C85999CB169" type="email">shihl@bjfu.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="285E655EF6565556C01655EA3B386E86" type="personal">
<mods:role id="922288031C64C10CF00C76417211D620">
<mods:roleTerm id="7AD344F6CFC76818E2A8D5B505C64B25">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="730BCCB01EC798BCB0AF7B3D9B591F85">Liang, Hongbin</mods:namePart>
<mods:nameIdentifier id="C7B230CB7580B8178A8450F37F5E8E68" type="ORCID">0000-0002-9989-5830</mods:nameIdentifier>
<mods:nameIdentifier id="2985DB419F06D46B1C2EC5222BFEC29E" type="email">shihl@bjfu.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="53E6A926B8FE92FF494B3E05123700E0">text</mods:typeOfResource>
<mods:relatedItem id="B4CAB492664C3FD15CD34B5FEA06B4A9" type="host">
<mods:titleInfo id="CDB9F275CD9EA076A2C4EE84D17B6DAD">
<mods:title id="4426CA91F8EF2EDB442B266DCED8D33C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5DA2B8D7FB4A8BCD28142AEBDE44321A">
<mods:date id="F2AFD8D6DA664C758E5867248FA69096">2023</mods:date>
<mods:detail id="B92DF50699ECC12A135F50D75DB323E7" type="pubDate">
<mods:number id="25A75513EE78F751CE3BAF26535A9924">2023-05-18</mods:number>
</mods:detail>
<mods:detail id="090A0F840EA5032244C3C0DB48F7E34B" type="volume">
<mods:number id="7E78144ED0BA91960387F75FFF0A273E">5286</mods:number>
</mods:detail>
<mods:detail id="B3457F30C3C7DDC8C1C4A8E9A062EDA5" type="issue">
<mods:number id="916001FC3B4868901BBC3190C1E9FA14">1</mods:number>
</mods:detail>
<mods:extent id="5A9C601B54953C311810784ACA8605EB" unit="page">
<mods:start id="D1B705E8D5C51948DEFEB1FCC9C724D0">1</mods:start>
<mods:end id="A3FABFC365C1A56A44AD8A322849C7BC">144</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="64DF75EF05A4971709D78E1D49082AED">
<mods:url id="93189B96E3B9E5E7008CD3E3270B7A3D">http://dx.doi.org/10.11646/zootaxa.5286.1.1</mods:url>
</mods:location>
<mods:classification id="CCD7B09098AB392D94D85D7537291EC0">journal article</mods:classification>
<mods:identifier id="8F43CECA9FAE2A5E937DA36259C47792" type="CLB-Dataset">263342</mods:identifier>
<mods:identifier id="9B6DBD5562C574076B20096D74C5B739" type="DOI">10.11646/zootaxa.5286.1.1</mods:identifier>
<mods:identifier id="613F6C938463D11B8E87BBC76231BE1C" type="GBIF-Dataset">67e7a105-1f62-414a-83d9-f758039eb726</mods:identifier>
<mods:identifier id="9F1E1006280D23E010CB994D3C2191D3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BB5943A1DBBD76CB0EA65C19B2FB8065" type="Zenodo-Dep">7958937</mods:identifier>
<mods:identifier id="3A8ECBBA6C278D0FFA3A2746073315BA" type="ZooBank">F9834684-24D3-4795-B5EB-77B451DF856D</mods:identifier>
</mods:mods>
<treatment id="038776236263FFC42DEFB058FAD95E4F" ID-DOI="http://doi.org/10.5281/zenodo.7963725" ID-GBIF-Taxon="209629020" ID-Zenodo-Dep="7963725" LSID="urn:lsid:plazi:treatment:038776236263FFC42DEFB058FAD95E4F" httpUri="http://treatment.plazi.org/id/038776236263FFC42DEFB058FAD95E4F" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38">
<subSubSection id="C33494BE6263FFC62DEFB058FCF45F67" box="[151,787,547,574]" pageId="37" pageNumber="38" type="nomenclature">
<paragraph id="8B91C7356263FFC62DEFB058FCF45F67" blockId="37.[151,1108,547,609]" box="[151,787,547,574]" pageId="37" pageNumber="38">
<emphasis id="B95A1B276263FFC62DEFB058FCF45F67" bold="true" box="[151,787,547,574]" pageId="37" pageNumber="38">
[5]
<taxonomicName id="4C2EBCB66263FFC62DB9B058FCF45F67" ID-CoL="4DKQR" authority="(Chaudoir, 1872)" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[193,787,547,574]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="testacea" subGenus="Crossoglossa">
<emphasis id="B95A1B276263FFC62DB9B058FEFE5F67" bold="true" box="[193,281,547,573]" italics="true" pageId="37" pageNumber="38">Parena</emphasis>
(
<emphasis id="B95A1B276263FFC62C52B058FE2F5F64" bold="true" box="[298,456,547,574]" italics="true" pageId="37" pageNumber="38">Crossoglossa</emphasis>
)
<emphasis id="B95A1B276263FFC62CA1B058FDDE5F67" bold="true" box="[473,569,547,573]" italics="true" pageId="37" pageNumber="38">testacea</emphasis>
(
<bibRefCitation id="EFBFBAC46263FFC62F30B058FCEC5F67" author="Chaudoir, B. M." box="[584,779,547,573]" pageId="37" pageNumber="38" pagination="97 - 204" refId="ref110142" refString="Chaudoir, B. M. (1872) Monographie des Callidides. Annales de Societe Entomologique de Belgique, 16, 97 - 204." type="journal article" year="1872">Chaudoir, 1872</bibRefCitation>
)
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6263FFC62DEFB03CFBB35F3B" box="[151,1108,583,609]" pageId="37" pageNumber="38" type="description">
<paragraph id="8B91C7356263FFC62DEFB03CFBB35F3B" blockId="37.[151,1108,547,609]" box="[151,1108,583,609]" pageId="37" pageNumber="38">
Habitus:
<figureCitation id="1315DBB06263FFC62D85B03CFE435F3B" box="[253,420,583,609]" captionStart="FIGURE 21" captionStartId="38.[151,250,1850,1874]" captionTargetBox="[219,1375,184,1823]" captionTargetId="figure-28@38.[212,1375,181,1827]" captionTargetPageId="38" captionText="FIGURE 21. Habitus of Parena (Crossoglossa) testacea species group. A Lectotype of Parena testacea (Chaudoir). B P. testacea (Chaudoir), male from Guangdong. C Holotype of Parena sulawesiensis Kirschenhofer. D P. sulawesiensis Kirschenhofer, male from Sulawesi. E Holotype of Parena mellea (Chaudoir). F P. mellea (Chaudoir), male from Maluku. G P. mellea (Chaudoir), female from Philippine. H Holotype of Parena cruralis Andrewes. I P. cruralis Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958979" httpUri="https://zenodo.org/record/7958979/files/figure.png" pageId="37" pageNumber="38">Figs 21A, 21B</figureCitation>
. Male genitalia:
<figureCitation id="1315DBB06263FFC62F27B03CFD565F3B" box="[607,689,583,609]" captionStart="FIGURE 22" captionStartId="39.[151,250,1383,1407]" captionTargetBox="[159,1428,819,1352]" captionTargetId="figure-316@39.[151,1436,813,1359]" captionTargetPageId="39" captionText="FIGURE 22. Median lobe of male genitalia for Parena (Crossoglossa) testacea (Chaudoir), right-lateral (r), ventral (v), leftlateral (l), and dorsal (d) views. A India. B Shanghai. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958981" httpUri="https://zenodo.org/record/7958981/files/figure.png" pageId="37" pageNumber="38">Fig. 22</figureCitation>
. Gonocoxites of ovipositor:
<figureCitation id="1315DBB06263FFC62E88B03CFBD95F3B" box="[1008,1086,583,609]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). AJ subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. KX subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Yγ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="37" pageNumber="38">Fig. 11</figureCitation>
α.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6263FFC62DEFB0F7FA7A5FBB" pageId="37" pageNumber="38" type="reference_group">
<paragraph id="8B91C7356263FFC62DEFB0F7FA7A5FBB" blockId="37.[151,1437,652,737]" pageId="37" pageNumber="38">
<bibRefCitation id="EFBFBAC46263FFC62DEFB0F7FE805FFE" author="Chaudoir, B. M." box="[151,359,652,676]" pageId="37" pageNumber="38" pagination="97 - 204" refId="ref110142" refString="Chaudoir, B. M. (1872) Monographie des Callidides. Annales de Societe Entomologique de Belgique, 16, 97 - 204." type="journal article" year="1872">Chaudoir, 1872: 178</bibRefCitation>
(original:
<taxonomicName id="4C2EBCB66263FFC62CA9B0F7FDBE5FFE" ID-CoL="8KWFM" authority="Chaudoir, 1873" box="[465,601,652,676]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B95A1B276263FFC62CA9B0F7FDBE5FFE" box="[465,601,652,676]" italics="true" pageId="37" pageNumber="38">Crossoglossa</emphasis>
</taxonomicName>
; type locality:
<materialsCitation id="3B46CD686263FFC62F89B0F7FBFF5FFE" ID-GBIF-Occurrence="4115871423" box="[753,1048,652,676]" pageId="37" pageNumber="38" typeStatus="lectotype">
Deccan;
<typeStatus id="549579976263FFC62E32B0F7FC405FFE" box="[842,935,652,676]" pageId="37" pageNumber="38" type="lectotype">lectotype</typeStatus>
in MNHN
</materialsCitation>
);
<bibRefCitation id="EFBFBAC46263FFC62950B0F7FAE65FFE" author="Andrewes, H. E." box="[1064,1281,652,676]" pageId="37" pageNumber="38" pagination="303 - 340" refId="ref109622" refString="Andrewes, H. E. (1929) The Fauna of Sumatrensis. Tijdschrift voor Entomologie, 72, 303 - 340." type="journal article" year="1929">Andrewes, 1929: 314</bibRefCitation>
(misidentification of
<taxonomicName id="4C2EBCB66263FFC62C6DB0D0FE4C5F98" ID-CoL="4DKPT" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[277,427,682,706]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="mellea">
<emphasis id="B95A1B276263FFC62C6DB0D0FE4C5F98" box="[277,427,682,706]" italics="true" pageId="37" pageNumber="38">Parena mellea</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFBFBAC46263FFC62CB9B0D1FD785F98" author="Andrewes, H. E." box="[449,671,682,706]" pageId="37" pageNumber="38" refId="ref109646" refString="Andrewes, H. E. (1930) Catalogue of Indian Insects, pars 18 (carabidae), Carabidae. Calcutta: Government of India Central Publication Branch, xxii + 389 pp." type="book" year="1930">Andrewes, 1930: 258</bibRefCitation>
(
<collectingCountry id="F33987A56263FFC62FD6B0D1FD175F98" box="[686,752,682,706]" name="Nepal" pageId="37" pageNumber="38">Nepal</collectingCountry>
);
<bibRefCitation id="EFBFBAC46263FFC62E7CB0D1FC365F98" author="Jedlicka, A." box="[772,977,682,706]" pageId="37" pageNumber="38" pagination="207 - 211" refId="ref111420" refString="Jedlicka, A. (1952) Les Carabides nouveaux de la zone palaearctique (Col.). Sbornik Entomologickeho Oddeleni Narodniho Musea v Praze, 27 [1951], 207 - 211." type="journal article" year="1952">Jedlička, 1952: 210</bibRefCitation>
;
<bibRefCitation id="EFBFBAC46263FFC62EA5B0D1FB255F98" author="Darlington, P. J." box="[989,1218,682,706]" pageId="37" pageNumber="38" pagination="1 - 253" refId="ref110388" refString="Darlington, P. J. (1968) The carabid beetles of New Guinea. Part 3. Harpalinae (Continued): Perigonine to Pseudomorphini. Bulletin of the Museum of Comparative Zoology at Harvard College, 137 (1), 1 - 253." type="journal article" year="1968">Darlington, 1968: 139</bibRefCitation>
(misidentification of
<taxonomicName id="4C2EBCB66263FFC62DBFB0B1FEBC5FBB" ID-CoL="4DKPT" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[199,347,713,737]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="mellea">
<emphasis id="B95A1B276263FFC62DBFB0B1FEBC5FBB" box="[199,347,713,737]" italics="true" pageId="37" pageNumber="38">Parena mellea</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFBFBAC46263FFC62C17B0B2FDC75FBB" author="Shibata, T." box="[367,544,713,737]" pageId="37" pageNumber="38" pagination="63 - 67" refId="ref112910" refString="Shibata, T. (1987) Three new species and a new record of Carabidae from Taiwan (Truncatipennes) (Coleoptera). Entomological Review of Japan, 42 (1), 63 - 67." type="journal article" year="1987">Shibata, 1987: 63</bibRefCitation>
(
<collectingRegion id="49EA09D76263FFC62F55B0B2FD635FBB" box="[557,644,713,737]" country="China" name="Taiwan)" pageId="37" pageNumber="38">Taiwan)</collectingRegion>
;
<bibRefCitation id="EFBFBAC46263FFC62FF6B0B2FCB95FBB" author="Xie, W. P. &amp; Yu, P. Y." box="[654,862,713,737]" pageId="37" pageNumber="38" pagination="185 - 195" refId="ref113326" refString="Xie, W. P. &amp; Yu, P. Y. (1993) On the Chinese species of Parena Motschulsky (Coleoptera: Carabidae). Sinozoologia, 10, 185 - 195." type="journal article" year="1993">Xie &amp; Yu, 1993: 194</bibRefCitation>
(
<collectingCountry id="F33987A56263FFC62E13B0B2FC4A5FBB" box="[875,941,713,737]" name="China" pageId="37" pageNumber="38">China</collectingCountry>
:
<collectingRegion id="49EA09D76263FFC62ECDB0B2FBE35FBB" box="[949,1028,713,737]" country="China" name="Jiangsu" pageId="37" pageNumber="38">Jiangsu</collectingRegion>
,
<collectingRegion id="49EA09D76263FFC62975B0B2FBB85FBB" box="[1037,1119,713,737]" country="China" name="Yunnan" pageId="37" pageNumber="38">Yunnan</collectingRegion>
);
<bibRefCitation id="EFBFBAC46263FFC62917B0B2FA815FBB" author="Kirschenhofer, E." box="[1135,1382,713,737]" pageId="37" pageNumber="38" pagination="87 - 103" refId="ref111612" refString="Kirschenhofer, E. (2006) Neue Arten der Gattung Parena Motschulsky aus Japan, China Und Indonesien (Sulawesi) (Coleoptera: Carabidae). Mitteilungen des Internationalen Entomologischen Vereins E. V. Frankfurt, 31 (3 - 4), 87 - 103." type="journal article" year="2006">Kirschenhofer, 2006: 89</bibRefCitation>
, 102.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6263FFC62DEFB171FE795EEE" pageId="37" pageNumber="38" type="materials_examined">
<paragraph id="8B91C7356263FFC62DEFB171FE795EEE" blockId="37.[151,1437,777,2028]" pageId="37" pageNumber="38">
<emphasis id="B95A1B276263FFC62DEFB171FE595E79" bold="true" box="[151,446,777,804]" pageId="37" pageNumber="38">Type material examined</emphasis>
.
<emphasis id="B95A1B276263FFC62CB7B172FC355E7E" bold="true" box="[463,978,777,804]" pageId="37" pageNumber="38">
<taxonomicName id="4C2EBCB66263FFC62CB7B172FCAB5E79" authority="Chaudoir" authorityName="Chaudoir" authorityYear="1872" box="[463,844,777,804]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="B95A1B276263FFC62CB7B172FD2B5E7E" bold="true" box="[463,716,777,804]" italics="true" pageId="37" pageNumber="38">Crossoglossa testacea</emphasis>
Chaudoir
</taxonomicName>
:
<typeStatus id="549579976263FFC62E24B171FC355E7E" box="[860,978,778,804]" pageId="37" pageNumber="38" type="lectotype">Lectotype</typeStatus>
</emphasis>
(MNHN,
<figureCitation id="1315DBB06263FFC62934B171FB515E7E" box="[1100,1206,778,804]" captionStart="FIGURE 21" captionStartId="38.[151,250,1850,1874]" captionTargetBox="[219,1375,184,1823]" captionTargetId="figure-28@38.[212,1375,181,1827]" captionTargetPageId="38" captionText="FIGURE 21. Habitus of Parena (Crossoglossa) testacea species group. A Lectotype of Parena testacea (Chaudoir). B P. testacea (Chaudoir), male from Guangdong. C Holotype of Parena sulawesiensis Kirschenhofer. D P. sulawesiensis Kirschenhofer, male from Sulawesi. E Holotype of Parena mellea (Chaudoir). F P. mellea (Chaudoir), male from Maluku. G P. mellea (Chaudoir), female from Philippine. H Holotype of Parena cruralis Andrewes. I P. cruralis Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958979" httpUri="https://zenodo.org/record/7958979/files/figure.png" pageId="37" pageNumber="38">Fig. 21A</figureCitation>
), designated herein: female, body length =
<quantity id="4CD66AD06263FFC62CE7B155FDEF5E12" box="[415,520,814,840]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="37" pageNumber="38" unit="mm" value="10.1">10.1 mm</quantity>
, pin mounted, &quot;TYPE&quot; [red label, added by Mateu], &quot;
<taxonomicName id="4C2EBCB66263FFC62903B154FB305E12" authorityName="Chaudoir" authorityYear="1872" box="[1147,1239,815,840]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="B95A1B276263FFC62903B154FB305E12" box="[1147,1239,815,840]" italics="true" pageId="37" pageNumber="38">testacea</emphasis>
</taxonomicName>
/
<emphasis id="B95A1B276263FFC6298AB155FAA25E12" box="[1266,1349,814,840]" italics="true" pageId="37" pageNumber="38">Chaud.</emphasis>
/
<emphasis id="B95A1B276263FFC62818B155FE925E36" italics="true" pageId="37" pageNumber="38">Indes Orient. / S. Stevens</emphasis>
&quot; [ex Chaudoirs box label, pinned under specimen now], &quot;MUSEUM PARIS 1952, Coll. R. OBERTHUR&quot;.
<emphasis id="B95A1B276263FFC62C3CB10DFE0E5ECA" bold="true" box="[324,489,886,912]" pageId="37" pageNumber="38">
<typeStatus id="549579976263FFC62C3CB10DFE0E5ECA" box="[324,489,886,912]" pageId="37" pageNumber="38" type="paralectotype">Paralectotype</typeStatus>
</emphasis>
(MNHN):
<specimenCount id="9D280CBC6263FFC62F1DB10DFD565ECA" box="[613,689,886,912]" pageId="37" pageNumber="38" type="male">1 male</specimenCount>
, without head, &quot;
<typeStatus id="549579976263FFC62E27B10DFC155ECA" box="[863,1010,886,912]" pageId="37" pageNumber="38" type="paratype">PARATYPE</typeStatus>
&quot; [red label], &quot;MUSEUM PARIS 1952, Coll. R. OBERTHUR&quot;.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6263FFC62DBFB1C5FD14597A" pageId="37" pageNumber="38" type="discussion">
<paragraph id="8B91C7356263FFC62DBFB1C5FD14597A" blockId="37.[151,1437,777,2028]" pageId="37" pageNumber="38">
<emphasis id="B95A1B276263FFC62DBFB1C5FE945E82" bold="true" box="[199,371,958,984]" pageId="37" pageNumber="38">Notes on types</emphasis>
. This species was originally described from
<specimenCount id="9D280CBC6263FFC62E1EB1C5FC2C5E82" box="[870,971,958,984]" pageId="37" pageNumber="38" type="male">one male</specimenCount>
and
<specimenCount id="9D280CBC6263FFC62979B1C5FB9B5E82" box="[1025,1148,958,984]" pageId="37" pageNumber="38" type="female">one female</specimenCount>
from &quot;Deccan&quot;, perfectly in accord with the
<specimenCount id="9D280CBC6263FFC62C1CB199FE235EA6" box="[356,452,994,1020]" pageId="37" pageNumber="38" type="generic">two type</specimenCount>
specimens in MNHN that we examined. Because the male is incomplete (head missing), we herein designate the complete female as
<typeStatus id="549579976263FFC62FFDB67DFD08597A" box="[645,751,1030,1056]" pageId="37" pageNumber="38" type="lectotype">lectotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6263FFC62DBFB651FE8C5B42" pageId="37" pageNumber="38" type="materials_examined">
<paragraph id="8B91C7356263FFC62DBFB651FE8C5B42" blockId="37.[151,1437,777,2028]" pageId="37" pageNumber="38">
<materialsCitation id="3B46CD686263FFC62DBFB651FAFB591E" ID-GBIF-Occurrence="4115871305" box="[199,1308,1065,1092]" collectingDate="1912-06-24" country="China" location="China" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1">
<emphasis id="B95A1B276263FFC62DBFB651FDC75919" bold="true" box="[199,544,1065,1092]" pageId="37" pageNumber="38">Non-type material examined</emphasis>
.
<emphasis id="B95A1B276263FFC62F49B652FD9D5919" bold="true" box="[561,634,1065,1091]" pageId="37" pageNumber="38">
<collectingCountry id="F33987A56263FFC62F49B652FD9D5919" box="[561,634,1065,1091]" name="China" pageId="37" pageNumber="38">China</collectingCountry>
</emphasis>
:
<specimenCount id="9D280CBC6263FFC62FF3B651FD3F591E" box="[651,728,1066,1092]" pageId="37" pageNumber="38" type="male">1 male</specimenCount>
(IZAS), &quot;Zi-ka-wei,
<date id="FF90E1F56263FFC62EB5B651FBB7591E" box="[973,1104,1066,1092]" pageId="37" pageNumber="38" value="1912-06-24">
<collectingDate id="EFD4181D6263FFC62EB5B651FBB7591E" box="[973,1104,1066,1092]" pageId="37" pageNumber="38" value="1912-06-24">1912.VI.24</collectingDate>
</date>
&quot; &lt;
<figureCitation id="1315DBB06263FFC62909B651FB3B591E" box="[1137,1244,1066,1092]" captionStart="FIGURE 10" captionStartId="16.[151,250,1909,1933]" captionTargetBox="[163,1429,1250,1879]" captionTargetId="figure-182@16.[152,1436,1241,1885]" captionTargetPageId="16" captionText="FIGURE 10. Parameres of male genitalia for Parena spp. A P. (Crossoglossa) cavipennis (Bates), Beijing. B P. (Crossoglossa) testacea (Chaudoir), Shanghai. C P. (Bothynoptera) dorsigera (Schaum), Zhejiang. D P. (Bothynoptera) taiwana Hua, Taiwan. E P. (Bothynoptera) tripunctata (Bates), Shaanxi. F P. (Parena) rubripicta Andrewes, Xizang. G P. (Parena) fasciata (Chaudoir), Taiwan. H P. (Parena) nigrolineata (Chaudoir), Taiwan. I P. (Parena) latecincta (Bates), Yunnan. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958957" httpUri="https://zenodo.org/record/7958957/files/figure.png" pageId="37" pageNumber="38">Figs 10B</figureCitation>
,
<figureCitation id="1315DBB06263FFC62992B651FAFB591E" box="[1258,1308,1066,1092]" captionStart="FIGURE 22" captionStartId="39.[151,250,1383,1407]" captionTargetBox="[159,1428,819,1352]" captionTargetId="figure-316@39.[151,1436,813,1359]" captionTargetPageId="39" captionText="FIGURE 22. Median lobe of male genitalia for Parena (Crossoglossa) testacea (Chaudoir), right-lateral (r), ventral (v), leftlateral (l), and dorsal (d) views. A India. B Shanghai. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958981" httpUri="https://zenodo.org/record/7958981/files/figure.png" pageId="37" pageNumber="38">22B</figureCitation>
</materialsCitation>
&gt;.
<materialsCitation id="3B46CD686263FFC62841B651FCCC5932" ID-GBIF-Occurrence="4115871401" collectingDate="1974-10-09" country="China" location="Putuoshan" pageId="37" pageNumber="38" specimenCount="1" specimenCount-female="1" stateProvince="Zhejiang">
<specimenCount id="9D280CBC6263FFC62841B651FA7B591E" box="[1337,1436,1066,1092]" pageId="37" pageNumber="38" type="female">1 female</specimenCount>
(IZAS), &quot;
<collectingRegion id="49EA09D76263FFC62D87B635FE8E5932" box="[255,361,1102,1128]" country="China" name="Zhejiang" pageId="37" pageNumber="38">Zhejiang</collectingRegion>
,
<location id="8EF191EE6263FFC62C0EB635FE095932" LSID="urn:lsid:plazi:treatment:038776236263FFC42DEFB058FAD95E4F:8EF191EE6263FFC62C0EB635FE095932" box="[374,494,1102,1128]" country="China" name="Putuoshan" pageId="37" pageNumber="38" stateProvince="Zhejiang">Putuoshan</location>
,
<date id="FF90E1F56263FFC62C82B635FD835932" box="[506,612,1102,1128]" pageId="37" pageNumber="38" value="1974-10-09">
<collectingDate id="EFD4181D6263FFC62C82B635FD835932" box="[506,612,1102,1128]" pageId="37" pageNumber="38" value="1974-10-09">1974.X.9</collectingDate>
</date>
, Li Fasheng lgt.&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD686263FFC62E40B635FA655932" ID-GBIF-Occurrence="4115871315" box="[824,1410,1102,1128]" collectingDate="1981-08-28" country="China" location="Pinghu" pageId="37" pageNumber="38" specimenCount="1" specimenCount-female="1" stateProvince="Zhejiang">
<specimenCount id="9D280CBC6263FFC62E40B635FC7C5932" box="[824,923,1102,1128]" pageId="37" pageNumber="38" type="female">1 female</specimenCount>
(IZAS), &quot;
<collectingRegion id="49EA09D76263FFC62973B635FB925932" box="[1035,1141,1102,1128]" country="China" name="Zhejiang" pageId="37" pageNumber="38">Zhejiang</collectingRegion>
,
<location id="8EF191EE6263FFC629FAB635FB335932" LSID="urn:lsid:plazi:treatment:038776236263FFC42DEFB058FAD95E4F:8EF191EE6263FFC629FAB635FB335932" box="[1154,1236,1102,1128]" country="China" name="Pinghu" pageId="37" pageNumber="38" stateProvince="Zhejiang">Pinghu</location>
,
<date id="FF90E1F56263FFC62999B635FA9E5932" box="[1249,1401,1102,1128]" pageId="37" pageNumber="38" value="1981-08-28">
<collectingDate id="EFD4181D6263FFC62999B635FA9E5932" box="[1249,1401,1102,1128]" pageId="37" pageNumber="38" value="1981-08-28">1981.VIII.28</collectingDate>
</date>
&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD686263FFC628F6B635FCBB59D6" ID-GBIF-Occurrence="4115871478" collectingDate="1982-07-11" country="China" location="Longquan" pageId="37" pageNumber="38" specimenCount="1" specimenCount-female="1" stateProvince="Zhejiang">
<specimenCount id="9D280CBC6263FFC628F6B635FF0459D6" pageId="37" pageNumber="38" type="female">1 female</specimenCount>
(IZAS), &quot;
<collectingRegion id="49EA09D76263FFC62C2DB609FE5859D6" box="[341,447,1138,1164]" country="China" name="Zhejiang" pageId="37" pageNumber="38">Zhejiang</collectingRegion>
,
<location id="8EF191EE6263FFC62CB5B609FDA659D6" LSID="urn:lsid:plazi:treatment:038776236263FFC42DEFB058FAD95E4F:8EF191EE6263FFC62CB5B609FDA659D6" box="[461,577,1138,1164]" country="China" name="Longquan" pageId="37" pageNumber="38" stateProvince="Zhejiang">Longquan</location>
,
<date id="FF90E1F56263FFC62F37B609FD3D59D6" box="[591,730,1138,1164]" pageId="37" pageNumber="38" value="1982-07-11">
<collectingDate id="EFD4181D6263FFC62F37B609FD3D59D6" box="[591,730,1138,1164]" pageId="37" pageNumber="38" value="1982-07-11">1982.VII.11</collectingDate>
</date>
&quot; &lt;
<figureCitation id="1315DBB06263FFC62F83B609FCAC59D6" box="[763,843,1138,1164]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). AJ subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. KX subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Yγ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="37" pageNumber="38">Fig. 11</figureCitation>
α
</materialsCitation>
&gt;.
<materialsCitation id="3B46CD686263FFC62E02B609FB6B59EA" ID-GBIF-Occurrence="4115871488" collectingDate="2020-03-15" country="China" county="Shenzhen" elevation="233" latitude="22.56" location="Meilinhoushan" longLatPrecision="756" longitude="114.02" municipality="Futian" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1" stateProvince="Guangdong">
<specimenCount id="9D280CBC6263FFC62E02B609FC2059D6" box="[890,967,1138,1164]" pageId="37" pageNumber="38" type="male">1 male</specimenCount>
(IZAS), &quot;
<collectingRegion id="49EA09D76263FFC62942B609FB2459D6" box="[1082,1219,1138,1164]" country="China" name="Guangdong" pageId="37" pageNumber="38">Guangdong</collectingRegion>
,
<collectingCounty id="62F0BFB96263FFC629A8B609FAA759D6" box="[1232,1344,1138,1164]" pageId="37" pageNumber="38">Shenzhen</collectingCounty>
,
<collectingMunicipality id="6BF55D4F6263FFC62836B609FA7F59D6" box="[1358,1432,1138,1164]" pageId="37" pageNumber="38">Futian</collectingMunicipality>
,
<location id="8EF191EE6263FFC62DEFB6EDFEA559EA" LSID="urn:lsid:plazi:treatment:038776236263FFC42DEFB058FAD95E4F:8EF191EE6263FFC62DEFB6EDFEA559EA" box="[151,322,1174,1200]" country="China" county="Shenzhen" latitude="22.56" longLatPrecision="756" longitude="114.02" municipality="Futian" name="Meilinhoushan" pageId="37" pageNumber="38" stateProvince="Guangdong">Meilinhoushan</location>
,
<geoCoordinate id="EE1AA1F26263FFC62C35B6EDFE4459EB" box="[333,419,1174,1201]" degrees="22.56" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="555" value="22.56">N22.56</geoCoordinate>
,
<geoCoordinate id="EE1AA1F26263FFC62CD7B6EDFDE959EA" box="[431,526,1174,1200]" degrees="114.02" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="555" value="114.02">E114.02</geoCoordinate>
,
<quantity id="4CD66AD06263FFC62F62B6EDFD8359EA" box="[538,612,1174,1200]" metricMagnitude="2" metricUnit="m" metricValue="2.33" pageId="37" pageNumber="38" unit="m" value="233.0">
<elevation id="000320066263FFC62F62B6EDFD8359EA" box="[538,612,1174,1200]" metricMagnitude="2" metricUnit="m" metricValue="2.33" pageId="37" pageNumber="38" unit="m" value="233.0">233 m</elevation>
</quantity>
,
<location id="8EF191EE6263FFC62F08B6EDFD3E59EA" LSID="urn:lsid:plazi:treatment:038776236263FFC42DEFB058FAD95E4F:8EF191EE6263FFC62F08B6EDFD3E59EA" box="[624,729,1174,1200]" country="China" county="Shenzhen" latitude="22.56" longLatPrecision="756" longitude="114.02" municipality="Futian" name="Mai Zuqi" pageId="37" pageNumber="38" stateProvince="Guangdong">Mai Zuqi</location>
lgt.,
<date id="FF90E1F56263FFC62E6BB6EDFC6C59EB" box="[787,907,1174,1201]" pageId="37" pageNumber="38" value="2020-03-15">
<collectingDate id="EFD4181D6263FFC62E6BB6EDFC6C59EB" box="[787,907,1174,1201]" pageId="37" pageNumber="38" value="2020-03-15">2020-3-15</collectingDate>
</date>
&quot; &lt;
<figureCitation id="1315DBB06263FFC62ED2B6EDFBAA59EA" box="[938,1101,1174,1200]" captionStart="FIGURE 14" captionStartId="25.[151,250,1584,1608]" captionTargetBox="[151,1436,181,1561]" captionTargetId="figure-19@25.[151,1436,181,1561]" captionTargetPageId="25" captionText="FIGURE 14. Elytra apex of Parena (Crossoglossa) spp. AH Elytra sutural angle (left), scale bar = 0.2mm. A P. cavipennis (Bates), Shaanxi. B P. laesipennis (Bates), Vietnam. C P. levata Andrewes, Borneo. D P. testacea (Chaudoir), Guangdong. E P. sulawesiensis Kirschenhofer, Sulawesi. F P. cruralis Andrewes, Borneo. G P. mellea (Chaudoir), Philippine. H P. sciakyi sp. n., holotype. IL Elytra outer apical angle (left), scale bar = 0.5mm. I P. cavipennis (Bates), Shaanxi. J P. laesipennis (Bates), Taiwan. K P. sciakyi sp. n., holotype. L P. testacea (Chaudoir), Guangdong." figureDoi="http://doi.org/10.5281/zenodo.7958963" httpUri="https://zenodo.org/record/7958963/files/figure.png" pageId="37" pageNumber="38">Figs 14D, 14L</figureCitation>
,
<figureCitation id="1315DBB06263FFC62924B6EDFB6B59EA" box="[1116,1164,1174,1200]" captionStart="FIGURE 21" captionStartId="38.[151,250,1850,1874]" captionTargetBox="[219,1375,184,1823]" captionTargetId="figure-28@38.[212,1375,181,1827]" captionTargetPageId="38" captionText="FIGURE 21. Habitus of Parena (Crossoglossa) testacea species group. A Lectotype of Parena testacea (Chaudoir). B P. testacea (Chaudoir), male from Guangdong. C Holotype of Parena sulawesiensis Kirschenhofer. D P. sulawesiensis Kirschenhofer, male from Sulawesi. E Holotype of Parena mellea (Chaudoir). F P. mellea (Chaudoir), male from Maluku. G P. mellea (Chaudoir), female from Philippine. H Holotype of Parena cruralis Andrewes. I P. cruralis Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958979" httpUri="https://zenodo.org/record/7958979/files/figure.png" pageId="37" pageNumber="38">21B</figureCitation>
</materialsCitation>
&gt;.
<materialsCitation id="3B46CD686263FFC629D0B6EDFC5B598E" ID-GBIF-Occurrence="4115871436" collectingDate="1958-04-18" country="China" elevation="650" location="Damenglong" municipality="Xishuangbanna" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
<specimenCount id="9D280CBC6263FFC629D0B6EDFB1359EA" box="[1192,1268,1174,1200]" pageId="37" pageNumber="38" type="male">1 male</specimenCount>
(IZAS), &quot;
<collectingRegion id="49EA09D76263FFC6281AB6EDFF25598E" country="China" name="Yunnan" pageId="37" pageNumber="38">Yunnan</collectingRegion>
,
<collectingMunicipality id="6BF55D4F6263FFC62DB5B6C1FE99598E" box="[205,382,1210,1236]" pageId="37" pageNumber="38">Xishuangbanna</collectingMunicipality>
,
<location id="8EF191EE6263FFC62CF0B6C1FDFB598E" LSID="urn:lsid:plazi:treatment:038776236263FFC42DEFB058FAD95E4F:8EF191EE6263FFC62CF0B6C1FDFB598E" box="[392,540,1210,1236]" country="China" municipality="Xishuangbanna" name="Damenglong" pageId="37" pageNumber="38" stateProvince="Yunnan">Damenglong</location>
,
<quantity id="4CD66AD06263FFC62F5EB6C1FD89598E" box="[550,622,1210,1237]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="37" pageNumber="38" unit="m" value="650.0">
<elevation id="000320066263FFC62F5EB6C1FD89598E" box="[550,622,1210,1237]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="37" pageNumber="38" unit="m" value="650.0">650 m</elevation>
</quantity>
,
<date id="FF90E1F56263FFC62F01B6C1FD10598E" box="[633,759,1210,1237]" pageId="37" pageNumber="38" value="1958-04-18">
<collectingDate id="EFD4181D6263FFC62F01B6C1FD10598E" box="[633,759,1210,1237]" pageId="37" pageNumber="38" value="1958-04-18">1958.IV.18</collectingDate>
</date>
, Zheng Leyi lgt.&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD686263FFC62EBFB6C1FBCD59A2" ID-GBIF-Occurrence="4115871419" collectorName="Ex. Musaeo H. W." country="China" location="Yang-tsze Kiang" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D280CBC6263FFC62EBFB6C1FBF6598E" box="[967,1041,1210,1236]" pageId="37" pageNumber="38" type="male">1 male</specimenCount>
(MNHN), &quot;
<location id="8EF191EE6263FFC629EDB6C1FABF598E" LSID="urn:lsid:plazi:treatment:038776236263FFC42DEFB058FAD95E4F:8EF191EE6263FFC629EDB6C1FABF598E" box="[1173,1368,1210,1236]" country="China" name="Yang-tsze Kiang" pageId="37" pageNumber="38">Yang-tsze Kiang</location>
&quot;, &quot;
<collectorName id="26DBA2E36263FFC6280BB6C1FED459A2" pageId="37" pageNumber="38">Ex. Musaeo H.W.</collectorName>
BATES, 1892&quot;, &quot;MUSEUM PARIS 1952, Coll. R. OBERTHUR&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD686263FFC6294EB6A5FE3A5846" ID-GBIF-Occurrence="4115871516" collectorName="Leg. Y. Yano" country="China" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D280CBC6263FFC6294EB6A5FB6459A2" box="[1078,1155,1246,1272]" pageId="37" pageNumber="38" type="male">1 male</specimenCount>
(NHML), &quot;Kuraru. Formosa. 1932 5.
<collectorName id="26DBA2E36263FFC62C45B779FE315846" box="[317,470,1282,1308]" pageId="37" pageNumber="38">Leg. Y. Yano</collectorName>
&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD686263FFC62C90B779FD255865" ID-GBIF-Occurrence="4115871378" collectingDate="1938-05-10" collectorName="Yoshio Yano &amp; H. E. Andrewes Coll." country="Vietnam" location="Formosa." municipality="Kasapin. Near" pageId="37" pageNumber="38" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D280CBC6263FFC62C90B779FDB25846" box="[488,597,1282,1308]" pageId="37" pageNumber="38" type="female">2 females</specimenCount>
(NHML), &quot;
<collectingMunicipality id="6BF55D4F6263FFC62FA0B779FC9B5846" box="[728,892,1282,1308]" pageId="37" pageNumber="38">Kasapin. Near</collectingMunicipality>
avi,
<location id="8EF191EE6263FFC62ECDB779FBF85846" LSID="urn:lsid:plazi:treatment:038776236263FFC42DEFB058FAD95E4F:8EF191EE6263FFC62ECDB779FBF85846" box="[949,1055,1282,1308]" country="Vietnam" municipality="Kasapin. Near" name="Formosa." pageId="37" pageNumber="38">Formosa.</location>
<date id="FF90E1F56263FFC6295EB779FB435846" box="[1062,1188,1282,1308]" pageId="37" pageNumber="38" value="1938-05-10">
<collectingDate id="EFD4181D6263FFC6295EB779FB435846" box="[1062,1188,1282,1308]" pageId="37" pageNumber="38" value="1938-05-10">10-V-1938</collectingDate>
</date>
&quot;, &quot;coll.
<collectorName id="26DBA2E36263FFC62983B779FA775846" box="[1275,1424,1282,1308]" pageId="37" pageNumber="38">Yoshio Yano</collectorName>
&quot;, &quot;
<collectorName id="26DBA2E36263FFC62DD8B75DFE74581A" box="[160,403,1318,1344]" pageId="37" pageNumber="38">H.E. Andrewes Coll.</collectorName>
B.M. 1945-97&quot;.
<emphasis id="B95A1B276263FFC62F24B75EFD255865" bold="true" box="[604,706,1317,1343]" pageId="37" pageNumber="38">
<collectingCountry id="F33987A56263FFC62F24B75EFD255865" box="[604,706,1317,1343]" name="Vietnam" pageId="37" pageNumber="38">Vietnam</collectingCountry>
</emphasis>
</materialsCitation>
:
<materialsCitation id="3B46CD686263FFC62FACB75DFAA55839" ID-GBIF-Occurrence="4115871332" collectingDate="1985-06-26" collectorName="Coll. Ch." country="India" county="Chu" location="Buonmathuot" municipality="Vietnam" pageId="37" pageNumber="38" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D280CBC6263FFC62FACB75DFCC1581A" box="[724,806,1318,1344]" pageId="37" pageNumber="38" type="male">1 male</specimenCount>
,
<specimenCount id="9D280CBC6263FFC62E4DB75DFC7D581A" box="[821,922,1318,1344]" pageId="37" pageNumber="38" type="female">1 female</specimenCount>
(MNHN), &quot;
<collectingCounty id="62F0BFB96263FFC6295FB75DFBBE581A" box="[1063,1113,1318,1344]" pageId="37" pageNumber="38">Chu</collectingCounty>
ganh, 5.1914&quot;, &quot;MUSEUM PARIS,
<collectorName id="26DBA2E36263FFC62D8AB731FEBF583E" box="[242,344,1354,1380]" pageId="37" pageNumber="38">Coll. Ch.</collectorName>
ALLUAUD&quot;. 1 ex (CMB), &quot;
<collectingMunicipality id="6BF55D4F6263FFC62FE7B731FCE5583E" box="[671,770,1354,1380]" pageId="37" pageNumber="38">Vietnam</collectingMunicipality>
:
<location id="8EF191EE6263FFC62E75B731FC4F583E" LSID="urn:lsid:plazi:treatment:038776236263FFC42DEFB058FAD95E4F:8EF191EE6263FFC62E75B731FC4F583E" box="[781,936,1354,1380]" country="India" county="Chu" municipality="Vietnam" name="Buonmathuot" pageId="37" pageNumber="38">Buonmathuot</location>
,
<location id="8EF191EE6263FFC62ECBB731FBAF583E" LSID="urn:lsid:plazi:treatment:038776236263FFC42DEFB058FAD95E4F:8EF191EE6263FFC62ECBB731FBAF583E" box="[947,1096,1354,1380]" country="India" county="Chu" municipality="Vietnam" name="Prov. Daklak" pageId="37" pageNumber="38">Prov. Daklak</location>
, 23-
<date id="FF90E1F56263FFC6290FB731FB17583F" box="[1143,1264,1354,1381]" pageId="37" pageNumber="38" value="1985-06-26">
<collectingDate id="EFD4181D6263FFC6290FB731FB17583F" box="[1143,1264,1354,1381]" pageId="37" pageNumber="38" value="1985-06-26">26.6.1985</collectingDate>
</date>
&quot;.
<emphasis id="B95A1B276263FFC6287AB732FAA55839" bold="true" box="[1282,1346,1353,1379]" pageId="37" pageNumber="38">
<collectingCountry id="F33987A56263FFC6287AB732FAA55839" box="[1282,1346,1353,1379]" name="India" pageId="37" pageNumber="38">India</collectingCountry>
</emphasis>
</materialsCitation>
:
<materialsCitation id="3B46CD686263FFC62829B731FB63588B" ID-GBIF-Occurrence="4115871530" collectorName="H. E. Andrewes Coll." country="India" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D280CBC6263FFC62829B731FA7B583E" box="[1361,1436,1354,1380]" pageId="37" pageNumber="38" type="male">1 male</specimenCount>
(MNHN), &quot;
<collectorName id="26DBA2E36263FFC62C6EB715FEAC58D2" box="[278,331,1390,1416]" pageId="37" pageNumber="38">Inde</collectorName>
coll. Boucard 1919&quot;, &quot;MUSEUM PARIS, Ex Coll. M. MAINDRON, Coll. G. BABAULT, 1930&quot;, &quot;
<taxonomicName id="4C2EBCB66263FFC62DE7B7E9FE0858F6" authority="Chaud." authorityName="Chaudoir" authorityYear="1872" box="[159,495,1426,1452]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="testacea">Crossoglossa testacea Chaud.</taxonomicName>
&quot; &lt;
<figureCitation id="1315DBB06263FFC62F75B7E9FD9458F6" box="[525,627,1426,1452]" captionStart="FIGURE 22" captionStartId="39.[151,250,1383,1407]" captionTargetBox="[159,1428,819,1352]" captionTargetId="figure-316@39.[151,1436,813,1359]" captionTargetPageId="39" captionText="FIGURE 22. Median lobe of male genitalia for Parena (Crossoglossa) testacea (Chaudoir), right-lateral (r), ventral (v), leftlateral (l), and dorsal (d) views. A India. B Shanghai. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958981" httpUri="https://zenodo.org/record/7958981/files/figure.png" pageId="37" pageNumber="38">Fig. 22A</figureCitation>
&gt;. 1 ex (MHNL), &quot;Belgaum / Bombay/
<collectorName id="26DBA2E36263FFC6295CB7E9FB3258F6" box="[1060,1237,1426,1452]" pageId="37" pageNumber="38">H.E. Andrewes</collectorName>
&quot;; &quot;
<emphasis id="B95A1B276263FFC6298AB7E9FE9F588A" italics="true" pageId="37" pageNumber="38">
<taxonomicName id="4C2EBCB66263FFC6298AB7E9FA7B58F6" authorityName="Macleay" authorityYear="1871" box="[1266,1436,1426,1452]" class="Insecta" family="Carabidae" genus="Phloeodromius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">Phloeodromius</taxonomicName>
/ testaceus/ Chaud. /
</emphasis>
compared with /
<typeStatus id="549579976263FFC62F4FB7CDFD80588A" box="[567,615,1462,1488]" pageId="37" pageNumber="38">type</typeStatus>
H.E.A.&quot;; &quot;
<collectorName id="26DBA2E36263FFC62FA5B7CDFC21588A" box="[733,966,1462,1488]" pageId="37" pageNumber="38">H.E. Andrewes Coll.</collectorName>
/ B.M. 1945-97&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD686263FFC629F6B7CDFACE58AE" ID-GBIF-Occurrence="4115871313" collectingDate="1897-06" collectorName="H. E. Andrewes Coll." country="Nepal" location="Shimoga" municipality="Maissour" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D280CBC6263FFC629F6B7CDFB30588A" box="[1166,1239,1462,1488]" pageId="37" pageNumber="38" type="male">1 male</specimenCount>
(MHNL), &quot;
<collectingMunicipality id="6BF55D4F6263FFC6282FB7CDFF2058AE" pageId="37" pageNumber="38">Maissour</collectingMunicipality>
/
<location id="8EF191EE6263FFC62D99B7A1FEA258AE" LSID="urn:lsid:plazi:treatment:038776236263FFC42DEFB058FAD95E4F:8EF191EE6263FFC62D99B7A1FEA258AE" box="[225,325,1498,1524]" country="Nepal" municipality="Maissour" name="Shimoga" pageId="37" pageNumber="38">Shimoga</location>
/
<date id="FF90E1F56263FFC62C26B7A1FE3458AE" box="[350,467,1498,1524]" pageId="37" pageNumber="38" value="1897-06">
<collectingDate id="EFD4181D6263FFC62C26B7A1FE3458AE" box="[350,467,1498,1524]" pageId="37" pageNumber="38" value="1897-06">Juin 1897</collectingDate>
</date>
&quot;; &quot;
<collectorName id="26DBA2E36263FFC62C8AB7A1FDBC58AE" box="[498,603,1498,1524]" pageId="37" pageNumber="38">Ex. Coll.</collectorName>
/
<collectorName id="26DBA2E36263FFC62F0CB7A1FCE558AE" box="[628,770,1498,1524]" pageId="37" pageNumber="38">R. Oberthur</collectorName>
&quot;; &quot;
<collectorName id="26DBA2E36263FFC62E58B7A1FBE858AE" box="[800,1039,1498,1524]" pageId="37" pageNumber="38">H.E. Andrewes Coll.</collectorName>
/ B.M. 1945-97&quot;.
<emphasis id="B95A1B276263FFC6299BB7A1FACE58AE" bold="true" box="[1251,1321,1498,1524]" pageId="37" pageNumber="38">
<collectingCountry id="F33987A56263FFC6299BB7A1FACE58AE" box="[1251,1321,1498,1524]" name="Nepal" pageId="37" pageNumber="38">Nepal</collectingCountry>
</emphasis>
</materialsCitation>
:
<specimenCount id="9D280CBC6263FFC62841B7A1FA7B58AE" box="[1337,1436,1498,1524]" pageId="37" pageNumber="38" type="female">1 female</specimenCount>
(MHNL), &quot;Nepal&quot;.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6263FFC62DBFB45AFB095AFE" pageId="37" pageNumber="38" type="diagnosis">
<paragraph id="8B91C7356263FFC62DBFB45AFD7B5BF2" blockId="37.[151,1437,777,2028]" pageId="37" pageNumber="38">
<emphasis id="B95A1B276263FFC62DBFB45AFE815B61" bold="true" box="[199,358,1569,1595]" pageId="37" pageNumber="38">Comparisons</emphasis>
.
<taxonomicName id="4C2EBCB66263FFC62C00B458FE135B66" authorityName="Chaudoir" authorityYear="1872" box="[376,500,1571,1596]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="B95A1B276263FFC62C00B458FE135B66" box="[376,500,1571,1596]" italics="true" pageId="37" pageNumber="38">P. testacea</emphasis>
</taxonomicName>
is distinguishable from the other four species in this group by the combination of: only antennomere 1 yellow; elytral intervals with dense punctures, strial puncture rows poorly defined; elytra sutural angles sharply pointed, forming long spines; outer apical angles distinctly angulate; terminal sternite with apical margin straight at middle in both sexes.
</paragraph>
<paragraph id="8B91C7356263FFC62DBFB4C9FCE25A06" blockId="37.[151,1437,777,2028]" pageId="37" pageNumber="38">
This species is similar to
<taxonomicName id="4C2EBCB66263FFC62C97B4C8FD425B96" authorityName="Kirschenhofer. D" authorityYear="2006" box="[495,677,1714,1740]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="sulawesiensis">
<emphasis id="B95A1B276263FFC62C97B4C8FD425B96" box="[495,677,1714,1740]" italics="true" pageId="37" pageNumber="38">P. sulawesiensis</emphasis>
</taxonomicName>
in the shape of elytra sutural angles (
<figureCitation id="1315DBB06263FFC62923B4C9FAE05B96" box="[1115,1287,1714,1740]" captionStart="FIGURE 14" captionStartId="25.[151,250,1584,1608]" captionTargetBox="[151,1436,181,1561]" captionTargetId="figure-19@25.[151,1436,181,1561]" captionTargetPageId="25" captionText="FIGURE 14. Elytra apex of Parena (Crossoglossa) spp. AH Elytra sutural angle (left), scale bar = 0.2mm. A P. cavipennis (Bates), Shaanxi. B P. laesipennis (Bates), Vietnam. C P. levata Andrewes, Borneo. D P. testacea (Chaudoir), Guangdong. E P. sulawesiensis Kirschenhofer, Sulawesi. F P. cruralis Andrewes, Borneo. G P. mellea (Chaudoir), Philippine. H P. sciakyi sp. n., holotype. IL Elytra outer apical angle (left), scale bar = 0.5mm. I P. cavipennis (Bates), Shaanxi. J P. laesipennis (Bates), Taiwan. K P. sciakyi sp. n., holotype. L P. testacea (Chaudoir), Guangdong." figureDoi="http://doi.org/10.5281/zenodo.7958963" httpUri="https://zenodo.org/record/7958963/files/figure.png" pageId="37" pageNumber="38">Figs 14D, 14E</figureCitation>
), but the latter species is different in: (1) basal three antennomeres yellow; (2) punctures on elytra intervals slightly finer and much sparser than in
<taxonomicName id="4C2EBCB66263FFC62CFEB480FE1A5A4E" authorityName="Chaudoir" authorityYear="1872" box="[390,509,1787,1812]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="B95A1B276263FFC62CFEB480FE1A5A4E" box="[390,509,1787,1812]" italics="true" pageId="37" pageNumber="38">P. testacea</emphasis>
</taxonomicName>
, so that strial puncture rows well defined; (3) elytra outer apical angles completely rounded, not forming distinct angles as in
<taxonomicName id="4C2EBCB66263FFC62F16B564FD025A62" authorityName="Chaudoir" authorityYear="1872" box="[622,741,1823,1848]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="B95A1B276263FFC62F16B564FD025A62" box="[622,741,1823,1848]" italics="true" pageId="37" pageNumber="38">P. testacea</emphasis>
</taxonomicName>
; (4) apical lamella of aedeagus slightly longer; and (5) female terminal sternite strongly projected at middle (
<figureCitation id="1315DBB06263FFC62FE7B539FD105A06" box="[671,759,1858,1884]" captionStart="FIGURE 8" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[198,1388,1017,1840]" captionTargetId="figure-59@14.[198,1388,1017,1841]" captionTargetPageId="14" captionText="FIGURE 8. Apex of sternite VII of Parena spp. AD females: A P. (Crossoglossa) sciakyi sp. n., holotype. B P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. C P. (Crossoglossa) mellea (Chaudoir), Philippine. D P. (Crossoglossa) cruralis Andrewes, Borneo. EJ males: E P. (Crossoglossa) cavipennis (Bates), Tianjin. F P. (Crossoglossa) mellea (Chaudoir), Philippine. G P. (Parena) monticola Shibata, Jiangxi. H P. (Parena) picipes sp. n., Borneo, paratype. I P. (Parena) fasciata (Chaudoir), Philippine. J P. (Parena) andrewesi Jedlička, Philippine. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.7958953" httpUri="https://zenodo.org/record/7958953/files/figure.png" pageId="37" pageNumber="38">Fig. 8B</figureCitation>
).
</paragraph>
<paragraph id="8B91C7356263FFC62DBFB51DFB095AFE" blockId="37.[151,1437,777,2028]" pageId="37" pageNumber="38">
In the color of antennae, some specimens of
<taxonomicName id="4C2EBCB66263FFC62FBAB51CFCCD5ADA" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[706,810,1894,1920]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="mellea">
<emphasis id="B95A1B276263FFC62FBAB51CFCCD5ADA" box="[706,810,1894,1920]" italics="true" pageId="37" pageNumber="38">P. mellea</emphasis>
</taxonomicName>
are similar to
<taxonomicName id="4C2EBCB66263FFC62EB7B51CFBA05ADA" authorityName="Chaudoir" authorityYear="1872" box="[975,1095,1895,1920]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="B95A1B276263FFC62EB7B51CFBA05ADA" box="[975,1095,1895,1920]" italics="true" pageId="37" pageNumber="38">P. testacea</emphasis>
</taxonomicName>
. These two species are distinguishable by the different shape of elytral sutural denticles and the apical lamella of male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6263FFC42DBFB5D6FB795F43" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38" type="description">
<paragraph id="8B91C7356263FFC42DBFB5D6FB795F43" blockId="37.[151,1437,777,2028]" lastBlockId="39.[151,1437,151,789]" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38">
<emphasis id="B95A1B276263FFC62DBFB5D6FEB65A9D" bold="true" box="[199,337,1965,1991]" pageId="37" pageNumber="38">Description</emphasis>
. Body length
<quantity id="4CD66AD06263FFC62C8FB5D5FD765A92" box="[503,657,1966,1993]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" metricValueMax="10.1" metricValueMin="9.5" pageId="37" pageNumber="38" unit="mm" value="9.8" valueMax="10.1" valueMin="9.5">9.510.1 mm</quantity>
. Dorsum pale yellow; antennomere 1 yellow, antennomeres 2 and 3 black or dark brown, remainder of antennae black; apices of mandibles black; terminal labial palpomere black, terminal and penultimate maxillary palpomeres black; tarsomeres black, in strong contrast with light yellow tibiae; venter pale yellow. Pronotum strongly transverse, PW/PL = 1.701.80, much wider than head, PW/HW = 1.131.22; widest at anterior third, lateral explanations very wide. Elytra without microsculpture; striae not incised, intervals completely flat, striae and intervals with similar very fine punctures, thus striae difficult to distinguish; disc depressed near middle of intervals 3 to 6, depressions large and shallow; apical truncation distinct, straight or slightly curved inward, outer apical angles sharp, forming clear obtuse angles; sutural angles sharply pointed, forming long spines. Apex of abdominal sternite VII straight in both sexes. Median lobe of aedeagus gradually attenuate to apex, apical lamella weakly bent toward dorsum, length subequal to its basal width (LL/LW = 1.01.1), apex rounded or slightly truncate. Endophallus densely scaled on basal sheath, base of apical sheath ovate in dorsal view; squamate sac not divided, on basal fifth of median lobe, right to squamate sheath (
<figureCitation id="1315DBB06261FFC42EABB3A0FBCF5CAF" box="[979,1064,475,501]" captionStart="FIGURE 22" captionStartId="39.[151,250,1383,1407]" captionTargetBox="[159,1428,819,1352]" captionTargetId="figure-316@39.[151,1436,813,1359]" captionTargetPageId="39" captionText="FIGURE 22. Median lobe of male genitalia for Parena (Crossoglossa) testacea (Chaudoir), right-lateral (r), ventral (v), leftlateral (l), and dorsal (d) views. A India. B Shanghai. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958981" httpUri="https://zenodo.org/record/7958981/files/figure.png" pageId="39" pageNumber="40">Fig. 22</figureCitation>
). Gonocoxite II of ovipositor dichotomous, inner and outer branches each with two short ensiform setae apically (
<figureCitation id="1315DBB06261FFC42948B384FB985F43" box="[1072,1151,511,537]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). AJ subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. KX subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Yγ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="39" pageNumber="40">Fig. 11</figureCitation>
α).
</paragraph>
</subSubSection>
<caption id="DF5197BD6260FFC52DEFB541FEBC5AB8" ID-DOI="http://doi.org/10.5281/zenodo.7958979" ID-Zenodo-Dep="7958979" httpUri="https://zenodo.org/record/7958979/files/figure.png" pageId="38" pageNumber="39" startId="38.[151,250,1850,1874]" targetBox="[219,1375,184,1823]" targetPageId="38" targetType="figure">
<paragraph id="8B91C7356260FFC52DEFB541FEBC5AB8" blockId="38.[151,1437,1850,2019]" pageId="38" pageNumber="39">
<emphasis id="B95A1B276260FFC52DEFB541FEFB5A08" bold="true" box="[151,284,1850,1874]" pageId="38" pageNumber="39">FIGURE 21</emphasis>
. Habitus of
<taxonomicName id="4C2EBCB66260FFC52CDCB540FD0B5A08" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[420,748,1851,1875]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="testacea" subGenus="Crossoglossa">
<emphasis id="B95A1B276260FFC52CDCB540FE095A08" box="[420,494,1851,1874]" italics="true" pageId="38" pageNumber="39">Parena</emphasis>
(
<emphasis id="B95A1B276260FFC52C87B540FD605A09" box="[511,647,1851,1875]" italics="true" pageId="38" pageNumber="39">Crossoglossa</emphasis>
)
<emphasis id="B95A1B276260FFC52FE1B540FD0B5A08" box="[665,748,1851,1874]" italics="true" pageId="38" pageNumber="39">testacea</emphasis>
</taxonomicName>
species group.
<emphasis id="B95A1B276260FFC52EEBB541FC425A08" bold="true" box="[915,933,1850,1874]" pageId="38" pageNumber="39">A</emphasis>
Lectotype of
<taxonomicName id="4C2EBCB66260FFC52945B540FAB85A09" authority="(Chaudoir)" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[1085,1375,1851,1875]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="B95A1B276260FFC52945B540FB045A08" box="[1085,1251,1851,1874]" italics="true" pageId="38" pageNumber="39">Parena testacea</emphasis>
(Chaudoir)
</taxonomicName>
.
<emphasis id="B95A1B276260FFC52814B540FA9A5A09" bold="true" box="[1388,1405,1851,1875]" pageId="38" pageNumber="39">B</emphasis>
<taxonomicName id="4C2EBCB66260FFC528FEB540FE855A2D" authority="(Chaudoir)" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="B95A1B276260FFC528FEB540FA725A08" box="[1414,1429,1851,1874]" italics="true" pageId="38" pageNumber="39">P</emphasis>
.
<emphasis id="B95A1B276260FFC52DEFB524FF0D5A2C" box="[151,234,1887,1910]" italics="true" pageId="38" pageNumber="39">testacea</emphasis>
(Chaudoir)
</taxonomicName>
, male from Guangdong.
<emphasis id="B95A1B276260FFC52F27B525FD965A2C" bold="true" box="[607,625,1886,1910]" pageId="38" pageNumber="39">C</emphasis>
Holotype of
<taxonomicName id="4C2EBCB66260FFC52F81B524FB6F5A2D" authority="Kirschenhofer. D" authorityName="Kirschenhofer. D" authorityYear="2006" box="[761,1160,1887,1911]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="sulawesiensis">
<emphasis id="B95A1B276260FFC52F81B524FC345A2D" box="[761,979,1887,1911]" italics="true" pageId="38" pageNumber="39">Parena sulawesiensis</emphasis>
Kirschenhofer.
<emphasis id="B95A1B276260FFC5290EB524FB6F5A2D" bold="true" box="[1142,1160,1887,1911]" pageId="38" pageNumber="39">D</emphasis>
</taxonomicName>
<taxonomicName id="4C2EBCB66260FFC529F7B524FF295AC1" authority="Kirschenhofer" authorityName="Kirschenhofer. D" authorityYear="2006" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="sulawesiensis">
<emphasis id="B95A1B276260FFC529F7B524FAD55A2D" box="[1167,1330,1887,1911]" italics="true" pageId="38" pageNumber="39">P. sulawesiensis</emphasis>
Kirschenhofer
</taxonomicName>
, male from Sulawesi.
<emphasis id="B95A1B276260FFC52CCAB5F8FE245AC1" bold="true" box="[434,451,1923,1947]" pageId="38" pageNumber="39">E</emphasis>
Holotype of
<taxonomicName id="4C2EBCB66260FFC52F33B5F8FCBF5AC1" authority="(Chaudoir)" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[587,856,1923,1947]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="mellea">
<emphasis id="B95A1B276260FFC52F33B5F8FD385AC1" box="[587,735,1923,1947]" italics="true" pageId="38" pageNumber="39">Parena mellea</emphasis>
(Chaudoir)
</taxonomicName>
.
<emphasis id="B95A1B276260FFC52E1AB5F8FC965AC1" bold="true" box="[866,881,1923,1947]" pageId="38" pageNumber="39">F</emphasis>
<taxonomicName id="4C2EBCB66260FFC52E00B5F8FBA95AC1" authority="(Chaudoir)" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[888,1102,1923,1947]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="mellea">
<emphasis id="B95A1B276260FFC52E00B5F8FC315AC1" box="[888,982,1923,1947]" italics="true" pageId="38" pageNumber="39">P. mellea</emphasis>
(Chaudoir)
</taxonomicName>
, male from Maluku.
<emphasis id="B95A1B276260FFC5285DB5F9FADE5AC0" bold="true" box="[1317,1337,1922,1946]" pageId="38" pageNumber="39">G</emphasis>
<taxonomicName id="4C2EBCB66260FFC52847B5F8FEEE5AE5" authority="(Chaudoir)" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="mellea">
<emphasis id="B95A1B276260FFC52847B5F8FA7A5AC1" box="[1343,1437,1923,1947]" italics="true" pageId="38" pageNumber="39">P. mellea</emphasis>
(Chaudoir)
</taxonomicName>
, female from Philippine.
<emphasis id="B95A1B276260FFC52F69B5DCFDC25AE5" bold="true" box="[529,549,1959,1983]" pageId="38" pageNumber="39">H</emphasis>
Holotype of
<taxonomicName id="4C2EBCB66260FFC52FC8B5DCFC305AE5" authority="Andrewes. I" authorityName="Andrewes. I" box="[688,983,1959,1983]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="cruralis">
<emphasis id="B95A1B276260FFC52FC8B5DCFCB55AE5" box="[688,850,1959,1983]" italics="true" pageId="38" pageNumber="39">Parena cruralis</emphasis>
Andrewes.
<emphasis id="B95A1B276260FFC52EB5B5DCFC305AE5" bold="true" box="[973,983,1959,1983]" pageId="38" pageNumber="39">I</emphasis>
</taxonomicName>
<taxonomicName id="4C2EBCB66260FFC52EA7B5DCFB5E5AE5" authority="Andrewes" authorityName="Andrewes, Borneo. E" authorityYear="1933" box="[991,1209,1959,1983]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="cruralis">
<emphasis id="B95A1B276260FFC52EA7B5DCFBAD5AE5" box="[991,1098,1959,1983]" italics="true" pageId="38" pageNumber="39">P. cruralis</emphasis>
Andrewes
</taxonomicName>
, female from Borneo. Scale bars = 2 mm.
</paragraph>
</caption>
<subSubSection id="C33494BE6261FFC42DBFB059FAD95E4F" pageId="39" pageNumber="40" type="distribution">
<paragraph id="8B91C7356261FFC42DBFB059FA775F67" blockId="39.[151,1437,151,789]" box="[199,1424,546,573]" pageId="39" pageNumber="40">
<emphasis id="B95A1B276261FFC42DBFB059FEBE5F66" bold="true" box="[199,345,546,572]" pageId="39" pageNumber="40">Distribution</emphasis>
(
<figureCitation id="1315DBB06261FFC42C10B058FE535F67" box="[360,436,547,573]" captionStart="MAP 4" captionStartId="36.[152,211,1895,1919]" captionTargetBox="[158,1430,1002,1871]" captionTargetId="figure-71@36.[158,1430,1002,1871]" captionTargetPageId="36" captionText="MAP 4. Distributions for species in the Parena (Crossoglossa) testacea group." figureDoi="http://doi.org/10.5281/zenodo.7963663" httpUri="https://zenodo.org/record/7963663/files/figure.png" pageId="39" pageNumber="40">Map 4</figureCitation>
, red).
<collectingCountry id="F33987A56261FFC42C83B058FDD95F67" box="[507,574,547,573]" name="China" pageId="39" pageNumber="40">China</collectingCountry>
(
<collectingRegion id="49EA09D76261FFC42F35B058FD5D5F67" box="[589,698,547,573]" country="China" name="Shanghai" pageId="39" pageNumber="40">Shanghai</collectingRegion>
,
<collectingRegion id="49EA09D76261FFC42FBDB058FCCB5F67" box="[709,812,547,573]" country="China" name="Zhejiang" pageId="39" pageNumber="40">Zhejiang</collectingRegion>
,
<collectingRegion id="49EA09D76261FFC42E4EB058FC5B5F67" box="[822,956,547,573]" country="China" name="Guangdong" pageId="39" pageNumber="40">Guangdong</collectingRegion>
,
<collectingRegion id="49EA09D76261FFC42EBEB058FBF85F67" box="[966,1055,547,573]" country="China" name="Yunnan" pageId="39" pageNumber="40">Yunnan</collectingRegion>
,
<collectingRegion id="49EA09D76261FFC42951B058FB605F67" box="[1065,1159,547,573]" country="China" name="Taiwan)" pageId="39" pageNumber="40">Taiwan)</collectingRegion>
,
<collectingCountry id="F33987A56261FFC429EAB058FB135F67" box="[1170,1268,547,573]" name="Vietnam" pageId="39" pageNumber="40">Vietnam</collectingCountry>
,
<collectingCountry id="F33987A56261FFC42986B058FADD5F67" box="[1278,1338,547,573]" name="India" pageId="39" pageNumber="40">India</collectingCountry>
,
<collectingCountry id="F33987A56261FFC4283EB058FA6A5F67" box="[1350,1421,547,573]" name="Nepal" pageId="39" pageNumber="40">Nepal</collectingCountry>
.
</paragraph>
<paragraph id="8B91C7356261FFC42DBFB03CFDD35FF3" blockId="39.[151,1437,151,789]" pageId="39" pageNumber="40">
<taxonomicName id="4C2EBCB66261FFC42DBFB03CFED85F3A" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[199,319,583,608]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="B95A1B276261FFC42DBFB03CFED85F3A" box="[199,319,583,608]" italics="true" pageId="39" pageNumber="40">P. testacea</emphasis>
</taxonomicName>
was recorded from
<collectingCountry id="F33987A56261FFC42F67B03CFD105F3B" box="[543,759,583,609]" name="Papua New Guinea" pageId="39" pageNumber="40">Papua New Guinea</collectingCountry>
by
<bibRefCitation id="EFBFBAC46261FFC42E59B03CFC0B5F3B" author="Darlington, P. J." box="[801,1004,583,609]" pageId="39" pageNumber="40" pagination="1 - 253" refId="ref110388" refString="Darlington, P. J. (1968) The carabid beetles of New Guinea. Part 3. Harpalinae (Continued): Perigonine to Pseudomorphini. Bulletin of the Museum of Comparative Zoology at Harvard College, 137 (1), 1 - 253." type="journal article" year="1968">Darlington (1968)</bibRefCitation>
without a detailed description. We did not examine any materials from
<collectingCountry id="F33987A56261FFC42F7EB010FD045FDF" box="[518,739,619,645]" name="Papua New Guinea" pageId="39" pageNumber="40">Papua New Guinea</collectingCountry>
, but according to the distribution, this record should be based on the misidentification of
<taxonomicName id="4C2EBCB66261FFC42CBEB0F4FDCA5FF3" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[454,557,655,681]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mellea">
<emphasis id="B95A1B276261FFC42CBEB0F4FDCA5FF3" box="[454,557,655,681]" italics="true" pageId="39" pageNumber="40">P. mellea</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B91C7356261FFC42DBFB0C9FAD95E4F" blockId="39.[151,1437,151,789]" pageId="39" pageNumber="40">
<emphasis id="B95A1B276261FFC42DBFB0C9FE3A5F96" bold="true" box="[199,477,690,716]" pageId="39" pageNumber="40">Geographical variation</emphasis>
. We studied male genitalia of specimens from
<collectingCountry id="F33987A56261FFC42E9FB0C8FBC65F97" box="[999,1057,691,717]" name="India" pageId="39" pageNumber="40">India</collectingCountry>
and
<collectingCountry id="F33987A56261FFC4292EB0C8FB7E5F97" box="[1110,1177,691,717]" name="China" pageId="39" pageNumber="40">China</collectingCountry>
(
<collectingRegion id="49EA09D76261FFC429DFB0C8FAF15F97" box="[1191,1302,691,717]" country="China" name="Shanghai" pageId="39" pageNumber="40">Shanghai</collectingRegion>
). They have very slight differences in the shape of apical lamella, with the apex slightly truncate in specimens from
<collectingCountry id="F33987A56261FFC4285DB0ACFAB85FAB" box="[1317,1375,727,753]" name="India" pageId="39" pageNumber="40">India</collectingCountry>
(
<figureCitation id="1315DBB06261FFC42817B0ACFF2C5E4F" captionStart="FIGURE 22" captionStartId="39.[151,250,1383,1407]" captionTargetBox="[159,1428,819,1352]" captionTargetId="figure-316@39.[151,1436,813,1359]" captionTargetPageId="39" captionText="FIGURE 22. Median lobe of male genitalia for Parena (Crossoglossa) testacea (Chaudoir), right-lateral (r), ventral (v), leftlateral (l), and dorsal (d) views. A India. B Shanghai. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958981" httpUri="https://zenodo.org/record/7958981/files/figure.png" pageId="39" pageNumber="40">Fig. 22A</figureCitation>
), and completely rounded in specimens from
<collectingCountry id="F33987A56261FFC42FB2B080FCEA5E4F" box="[714,781,763,789]" name="China" pageId="39" pageNumber="40">China</collectingCountry>
(
<figureCitation id="1315DBB06261FFC42E64B080FC645E4F" box="[796,899,763,789]" captionStart="FIGURE 22" captionStartId="39.[151,250,1383,1407]" captionTargetBox="[159,1428,819,1352]" captionTargetId="figure-316@39.[151,1436,813,1359]" captionTargetPageId="39" captionText="FIGURE 22. Median lobe of male genitalia for Parena (Crossoglossa) testacea (Chaudoir), right-lateral (r), ventral (v), leftlateral (l), and dorsal (d) views. A India. B Shanghai. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958981" httpUri="https://zenodo.org/record/7958981/files/figure.png" pageId="39" pageNumber="40">Fig. 22B</figureCitation>
). They are identical in other characters.
</paragraph>
</subSubSection>
</treatment>
</document>