306 lines
39 KiB
XML
306 lines
39 KiB
XML
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<mods:title id="58AFAE3C55FE8626CDB908BF019C5D87">Open-holed trapdoor spiders of the genus Teyl (Mygalomorphae: Nemesiidae Anamini) from Western Australia’s Pilbara bioregion: a new species and expanded phylogenetic assessment</mods:title>
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<mods:namePart id="29983FA997F01386ED41A30A3C49C548">Huey, Joel A.</mods:namePart>
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<mods:namePart id="E07CFA0550E616B2CD20EEB90D7E8761">Wilson, Jeremy D.</mods:namePart>
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<mods:namePart id="95B3D0B85B820838DBDF2CE7F54979C9">Hillyer, Mia J.</mods:namePart>
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Genus
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<taxonomicName id="4C2A68AF267CFFAEFF07B09960E81EF8" authority="Main, 1975" authorityName="Main" authorityYear="1975" box="[238,436,153,180]" class="Arachnida" family="Nemesiidae" genus="Teyl" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="genus">
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<emphasis id="B95ECF3E267CFFAEFF07B09960431EFF" bold="true" box="[238,287,153,180]" italics="true" pageId="7" pageNumber="356">Teyl</emphasis>
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<bibRefCitation id="EFBB6EDD267CFFAEFECEB09960E81EF8" author="Main, B. Y." box="[295,436,153,179]" pageId="7" pageNumber="356" pagination="73 - 78" refId="ref7346" refString="Main, B. Y. (1975) The citrine spider: a new genus of trapdoor spider (Mygalomorphae: Dipluridae). The Western Australian Naturalist, 13, 73 - 78." type="journal article" year="1975">Main, 1975</bibRefCitation>
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<taxonomicName id="4C2A68AF267CFFAEFF7EB0DE60391EBD" authority="Main, 1975: 74" authorityName="Main" authorityPageNumber="74" authorityYear="1975" box="[151,357,221,246]" class="Arachnida" family="Nemesiidae" genus="Teyl" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="genus">
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<emphasis id="B95ECF3E267CFFAEFF7EB0DE619C1EBD" box="[151,192,222,246]" italics="true" pageId="7" pageNumber="356">Teyl</emphasis>
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<treatmentCitation id="0A8B353D267CFFAEFF2EB0DD60391EBD" author="Main, B. Y." box="[199,357,221,246]" page="74" pageId="7" pageNumber="356" year="1975">
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<bibRefCitation id="EFBB6EDD267CFFAEFF2EB0DD60391EBD" author="Main, B. Y." box="[199,357,221,246]" pageId="7" pageNumber="356" pagination="73 - 78" refId="ref7346" refString="Main, B. Y. (1975) The citrine spider: a new genus of trapdoor spider (Mygalomorphae: Dipluridae). The Western Australian Naturalist, 13, 73 - 78." type="journal article" year="1975">Main, 1975: 74</bibRefCitation>
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.
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<typeStatus id="5491AD8E267CFFAEFE86B0DE60FD1EBD" box="[367,417,222,246]" pageId="7" pageNumber="356">Type</typeStatus>
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species by original designation
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<taxonomicName id="4C2A68AF267CFFAEFD0EB0DE62A11EBD" authority="Main, 1975" authorityName="Main" authorityYear="1975" box="[743,1021,221,246]" class="Arachnida" family="Nemesiidae" genus="Teyl" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="species" species="luculentus">
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<emphasis id="B95ECF3E267CFFAEFD0EB0DE62221EBD" box="[743,894,222,246]" italics="true" pageId="7" pageNumber="356">Teyl luculentus</emphasis>
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<bibRefCitation id="EFBB6EDD267CFFAEFC6CB0DD62A11EBD" author="Main, B. Y." box="[901,1021,221,246]" pageId="7" pageNumber="356" pagination="73 - 78" refId="ref7346" refString="Main, B. Y. (1975) The citrine spider: a new genus of trapdoor spider (Mygalomorphae: Dipluridae). The Western Australian Naturalist, 13, 73 - 78." type="journal article" year="1975">Main, 1975</bibRefCitation>
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.
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<taxonomicName id="4C2A68AF267CFFAEFF7EB0FC60E71F5F" authority="Main, 1983: 931" authorityName="Main" authorityPageNumber="931" authorityYear="1983" box="[151,443,252,277]" class="Arachnida" family="Nemesiidae" genus="Merredinia" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="genus">
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<emphasis id="B95ECF3E267CFFAEFF7EB0FC60551F5F" box="[151,265,252,276]" italics="true" pageId="7" pageNumber="356">Merredinia</emphasis>
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<treatmentCitation id="0A8B353D267CFFAEFEF9B0FC60E71F5F" author="Main, B. Y." box="[272,443,252,277]" page="931" pageId="7" pageNumber="356" year="1983">
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<bibRefCitation id="EFBB6EDD267CFFAEFEF9B0FC60E71F5F" author="Main, B. Y." box="[272,443,252,277]" pageId="7" pageNumber="356" pagination="923 - 949" refId="ref7444" refString="Main, B. Y. (1983) Further studies on the systematics of Australian Diplurinae (Chelicerata: Mygalomorphae: Dipluridae): two new genera from south western Australia. Journal of Natural History, 15, 923 - 949. https: // doi. org / 10.1080 / 00222938300770731" type="journal article" year="1983">Main, 1983: 931</bibRefCitation>
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.
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<typeStatus id="5491AD8E267CFFAEFE2DB0FD60AA1F5E" box="[452,502,253,277]" pageId="7" pageNumber="356">Type</typeStatus>
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species by original designation
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<taxonomicName id="4C2A68AF267CFFAEFCD5B0FC65E81F5F" authority="Main, 1983" authorityName="Main" authorityYear="1983" box="[828,1204,252,277]" class="Arachnida" family="Nemesiidae" genus="Merredinia" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="species" species="damsonoides">
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<emphasis id="B95ECF3E267CFFAEFCD5B0FC65651F5F" box="[828,1081,252,276]" italics="true" pageId="7" pageNumber="356">Merredinia damsonoides</emphasis>
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<bibRefCitation id="EFBB6EDD267CFFAEFBD6B0FC65E81F5F" author="Main, B. Y." box="[1087,1204,252,277]" pageId="7" pageNumber="356" pagination="923 - 949" refId="ref7444" refString="Main, B. Y. (1983) Further studies on the systematics of Australian Diplurinae (Chelicerata: Mygalomorphae: Dipluridae): two new genera from south western Australia. Journal of Natural History, 15, 923 - 949. https: // doi. org / 10.1080 / 00222938300770731" type="journal article" year="1983">Main, 1983</bibRefCitation>
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</taxonomicName>
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(synonymised by
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<treatmentCitation id="0A8B353D267CFFAEFA87B0FD60C71F78" author="Harvey, M. S. & Hillyer, M. J. & Main, B. Y. & Moulds, T. A. & Raven, R. J. & Rix, M. G. & Vink, C. J. & Huey, J. A." page="436" pageId="7" pageNumber="356" year="2018">
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<bibRefCitation id="EFBB6EDD267CFFAEFA87B0FD60C71F78" author="Harvey, M. S. & Hillyer, M. J. & Main, B. Y. & Moulds, T. A. & Raven, R. J. & Rix, M. G. & Vink, C. J. & Huey, J. A." pageId="7" pageNumber="356" pagination="407 - 452" refId="ref6824" refString="Harvey, M. S., Hillyer, M. J., Main, B. Y., Moulds, T. A., Raven, R. J., Rix, M. G., Vink, C. J. & Huey, J. A. (2018) Phylogenetic relationships of the Australasian open-holed trapdoor spiders (Araneae: Mygalomorphae: Nemesiidae: Anaminae): multilocus molecular analyses resolve the generic classification of a highly diverse fauna. Zoological Journal of the Linnean Society, 184, 407 - 452. https: // doi. org / 10.1093 / zoolinnean / zlx 111" type="journal article" year="2018">
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Harvey
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<emphasis id="B95ECF3E267CFFAEFF1BB11C60431F78" box="[242,287,283,307]" italics="true" pageId="7" pageNumber="356">et al</emphasis>
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., 2018: 436
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</bibRefCitation>
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</treatmentCitation>
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).
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</paragraph>
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<paragraph id="8B95132C267CFFAEFF7EB13A60201F3B" blockId="7.[151,1437,221,368]" pageId="7" pageNumber="356">
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<treatmentCitationGroup id="AB3A3402267CFFAEFF7EB13A60201F3B" pageId="7" pageNumber="356">
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<taxonomicName id="4C2A68AF267CFFAEFF7EB13A60EB1F19" authority="Main, 1985: 753" authorityName="Main" authorityPageNumber="753" authorityYear="1985" box="[151,439,313,338]" class="Arachnida" family="Nemesiidae" genus="Pseudoteyl" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="genus">
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<emphasis id="B95ECF3E267CFFAEFF7EB13A605A1F19" box="[151,262,314,338]" italics="true" pageId="7" pageNumber="356">Pseudoteyl</emphasis>
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<treatmentCitation id="0A8B353D267CFFAEFEE5B13960EB1F19" author="Main, B. Y." box="[268,439,313,338]" page="753" pageId="7" pageNumber="356" year="1985">
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<bibRefCitation id="EFBB6EDD267CFFAEFEE5B13960EB1F19" author="Main, B. Y." box="[268,439,313,338]" pageId="7" pageNumber="356" pagination="743 - 759" refId="ref7498" refString="Main, B. Y. (1985) Further studies on Australian Diplurinae: a review of the genera of the Teylini (Araneae: Mygalomorphae: Dipluridae). Australian Journal of Zoology, 33, 743 - 759. https: // doi. org / 10.1071 / ZO 9850743" type="journal article" year="1985">Main, 1985: 753</bibRefCitation>
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</treatmentCitation>
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.
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<typeStatus id="5491AD8E267CFFAEFE29B13A60AE1F19" box="[448,498,314,338]" pageId="7" pageNumber="356">Type</typeStatus>
|
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species by original designation
|
||
<taxonomicName id="4C2A68AF267CFFAEFCDEB13A65C41F19" authority="Main, 1985" authorityName="Main" authorityYear="1985" box="[823,1176,313,338]" class="Arachnida" family="Nemesiidae" genus="Pseudoteyl" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="species" species="vancouveri">
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||
<emphasis id="B95ECF3E267CFFAEFCDEB13A65421F1A" box="[823,1054,314,338]" italics="true" pageId="7" pageNumber="356">Pseudoteyl vancouveri</emphasis>
|
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<bibRefCitation id="EFBB6EDD267CFFAEFBCAB13965C41F19" author="Main, B. Y." box="[1059,1176,313,338]" pageId="7" pageNumber="356" pagination="743 - 759" refId="ref7498" refString="Main, B. Y. (1985) Further studies on Australian Diplurinae: a review of the genera of the Teylini (Araneae: Mygalomorphae: Dipluridae). Australian Journal of Zoology, 33, 743 - 759. https: // doi. org / 10.1071 / ZO 9850743" type="journal article" year="1985">Main, 1985</bibRefCitation>
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</taxonomicName>
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(synonymised by
|
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<treatmentCitation id="0A8B353D267CFFAEFABBB13A602D1F3B" author="Harvey, M. S. & Hillyer, M. J. & Main, B. Y. & Moulds, T. A. & Raven, R. J. & Rix, M. G. & Vink, C. J. & Huey, J. A." page="437" pageId="7" pageNumber="356" year="2018">
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<bibRefCitation id="EFBB6EDD267CFFAEFABBB13A602D1F3B" author="Harvey, M. S. & Hillyer, M. J. & Main, B. Y. & Moulds, T. A. & Raven, R. J. & Rix, M. G. & Vink, C. J. & Huey, J. A." pageId="7" pageNumber="356" pagination="407 - 452" refId="ref6824" refString="Harvey, M. S., Hillyer, M. J., Main, B. Y., Moulds, T. A., Raven, R. J., Rix, M. G., Vink, C. J. & Huey, J. A. (2018) Phylogenetic relationships of the Australasian open-holed trapdoor spiders (Araneae: Mygalomorphae: Nemesiidae: Anaminae): multilocus molecular analyses resolve the generic classification of a highly diverse fauna. Zoological Journal of the Linnean Society, 184, 407 - 452. https: // doi. org / 10.1093 / zoolinnean / zlx 111" type="journal article" year="2018">
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Harvey
|
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<emphasis id="B95ECF3E267CFFAEFF2EB15961A61F3B" box="[199,250,344,368]" italics="true" pageId="7" pageNumber="356">et al.</emphasis>
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, 2018: 437
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</bibRefCitation>
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</treatmentCitation>
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).
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</paragraph>
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<paragraph id="8B95132C267CFFAEFF7EB19E654E1CB6" blockId="7.[151,1437,414,765]" pageId="7" pageNumber="356">
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<emphasis id="B95ECF3E267CFFAEFF7EB19E604E1FF2" bold="true" box="[151,274,414,441]" pageId="7" pageNumber="356">Diagnosis.</emphasis>
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Most species of
|
||
<taxonomicName id="4C2A68AF267CFFAEFE34B19F63571FF2" authorityName="Main" authorityYear="1975" box="[477,523,415,441]" class="Arachnida" family="Nemesiidae" genus="Teyl" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95ECF3E267CFFAEFE34B19F63571FF2" box="[477,523,415,441]" italics="true" pageId="7" pageNumber="356">Teyl</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2A68AF267CFFAEFDA1B1A063CB1FF2" authorityName="Raven" authorityYear="1984" box="[584,663,416,441]" class="Arachnida" family="Nemesiidae" genus="Namea" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95ECF3E267CFFAEFDA1B1A063CB1FF2" box="[584,663,416,441]" italics="true" pageId="7" pageNumber="356">Namea</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from all other Anamini by the presence of a reflexed embolus arising on the lateral or proximal side of the pedipalpal bulb (
|
||
<figureCitation id="13110FA9267CFFAEFBC5B1C365ED1F95" box="[1068,1201,451,478]" captionStart="FIGURES 14–16" captionStartId="11.[151,264,855,880]" captionTargetBox="[151,1435,181,823]" captionTargetId="figure@11.[151,1435,181,829]" captionTargetPageId="11" captionText="FIGURES 14–16. Teyl heuretes sp. nov., male holotype (WAM T91918) from Weeli Wolli Creek region, Hamersley Range (Western Australia; PIL), pedipalp: 14, retrolateral view; 15, retro-ventral view; 16, prolateral view. Scale bar = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3460202" httpUri="https://zenodo.org/record/3460202/files/figure.png" pageId="7" pageNumber="356">Figs 14–16</figureCitation>
|
||
), combined with the absence of a megaspur on the male leg I tibia (
|
||
<figureCitation id="13110FA9267CFFAEFD7EB1E762441C4A" box="[663,792,487,513]" captionStart="FIGURES 4–13" captionStartId="10.[151,264,1902,1927]" captionTargetBox="[164,1423,166,1871]" captionTargetId="figure@10.[164,1423,166,1877]" captionTargetPageId="10" captionText="FIGURES 4–13. Teyl heuretes sp. nov., male holotype (WAM T91918) from Weeli Wolli Creek region, Hamersley Range (Western Australia; PIL), somatic morphology: 4–5, carapace and abdomen, dorsal view; 6, cephalothorax, lateral view; 7, eyes, dorsal view; 8, mouthparts, ventral view; 9–10, cephalothorax and abdomen, ventral view; 11, leg I, prolateral view; 12, leg I tibia, prolateral view; 13, leg I tibia, retrolateral view. Scale bars = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3460200" httpUri="https://zenodo.org/record/3460200/files/figure.png" pageId="7" pageNumber="356">Figs 11–13</figureCitation>
|
||
). However, a number of species of both
|
||
<taxonomicName id="4C2A68AF267CFFAEFB3CB1E7645F1C4A" authorityName="Main" authorityYear="1975" box="[1237,1283,487,513]" class="Arachnida" family="Nemesiidae" genus="Teyl" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95ECF3E267CFFAEFB3CB1E7645F1C4A" box="[1237,1283,487,513]" italics="true" pageId="7" pageNumber="356">Teyl</emphasis>
|
||
</taxonomicName>
|
||
(e.g.,
|
||
<taxonomicName id="4C2A68AF267CFFAEFAA1B1E861A51C6E" authorityName="Main" authorityYear="1983" class="Arachnida" family="Nemesiidae" genus="Merredinia" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="species" species="damsonoides">
|
||
<emphasis id="B95ECF3E267CFFAEFAA1B1E861A51C6E" italics="true" pageId="7" pageNumber="356">T. damsonoides</emphasis>
|
||
</taxonomicName>
|
||
) and
|
||
<taxonomicName id="4C2A68AF267CFFAEFEDEB20C60DA1C6E" authorityName="Raven" authorityYear="1984" box="[311,390,524,549]" class="Arachnida" family="Nemesiidae" genus="Namea" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95ECF3E267CFFAEFEDEB20C60DA1C6E" box="[311,390,524,549]" italics="true" pageId="7" pageNumber="356">Namea</emphasis>
|
||
</taxonomicName>
|
||
(e.g.,
|
||
<taxonomicName id="4C2A68AF267CFFAEFE22B20C625E1C6E" authority="Raven, 1984" authorityName="Raven" authorityYear="1984" box="[459,770,523,549]" class="Arachnida" family="Nemesiidae" genus="Namea" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="species" species="callemonda">
|
||
<emphasis id="B95ECF3E267CFFAEFE22B20C63311C6E" box="[459,621,523,549]" italics="true" pageId="7" pageNumber="356">N. callemonda</emphasis>
|
||
<bibRefCitation id="EFBB6EDD267CFFAEFD9AB20B625E1C6E" author="Raven, R. J." box="[627,770,523,549]" pageId="7" pageNumber="356" pagination="1 - 51" refId="ref7704" refString="Raven, R. J. (1984) A new diplurid genus from eastern Australia and a related Aname species (Diplurinae: Dipluridae: Araneae). Australian Journal of Zoology Supplementary Series, 96, 1 - 51. https: // doi. org / 10.1071 / AJZS 096" type="journal article" year="1984">Raven, 1984</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2A68AF267CFFAEFCE5B20C654C1C6E" authority="Raven, 1984" authorityName="Raven" authorityYear="1984" box="[780,1040,523,549]" class="Arachnida" family="Nemesiidae" genus="Namea" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="species" species="dahmsi">
|
||
<emphasis id="B95ECF3E267CFFAEFCE5B20C62211C6E" box="[780,893,523,549]" italics="true" pageId="7" pageNumber="356">N. dahmsi</emphasis>
|
||
<bibRefCitation id="EFBB6EDD267CFFAEFC6AB20B654C1C6E" author="Raven, R. J." box="[899,1040,523,549]" pageId="7" pageNumber="356" pagination="1 - 51" refId="ref7704" refString="Raven, R. J. (1984) A new diplurid genus from eastern Australia and a related Aname species (Diplurinae: Dipluridae: Araneae). Australian Journal of Zoology Supplementary Series, 96, 1 - 51. https: // doi. org / 10.1071 / AJZS 096" type="journal article" year="1984">Raven, 1984</bibRefCitation>
|
||
</taxonomicName>
|
||
) have a presumably autapomorphic morphology whereby the embolus is not reflexed, but rather positioned distally on the bulb; these species can be distinguished from similar Anamini by the absence of a leg I megaspur on males, or by the absence of incrassate tarsi on the leg I of males. Similarly, a number of species of
|
||
<taxonomicName id="4C2A68AF267CFFAEFCD9B27862231CDA" authorityName="Raven" authorityYear="1984" box="[816,895,632,657]" class="Arachnida" family="Nemesiidae" genus="Namea" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95ECF3E267CFFAEFCD9B27862231CDA" box="[816,895,632,657]" italics="true" pageId="7" pageNumber="356">Namea</emphasis>
|
||
</taxonomicName>
|
||
(e.g.
|
||
<taxonomicName id="4C2A68AF267CFFAEFC54B278658C1CDA" authority="Raven, 1984" authorityName="Raven" authorityYear="1984" box="[957,1232,631,657]" class="Arachnida" family="Nemesiidae" genus="Namea" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="species" species="calcaria">
|
||
<emphasis id="B95ECF3E267CFFAEFC54B27865661CDA" box="[957,1082,631,657]" italics="true" pageId="7" pageNumber="356">N. calcaria</emphasis>
|
||
<bibRefCitation id="EFBB6EDD267CFFAEFBA8B277658C1CDA" author="Raven, R. J." box="[1089,1232,631,657]" pageId="7" pageNumber="356" pagination="1 - 51" refId="ref7704" refString="Raven, R. J. (1984) A new diplurid genus from eastern Australia and a related Aname species (Diplurinae: Dipluridae: Araneae). Australian Journal of Zoology Supplementary Series, 96, 1 - 51. https: // doi. org / 10.1071 / AJZS 096" type="journal article" year="1984">Raven, 1984</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2A68AF267CFFAEFB32B278605C1CFE" authority="Raven 1984" authorityName="Raven" authorityYear="1984" class="Arachnida" family="Nemesiidae" genus="Namea" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="species" species="dicalcaria">
|
||
<emphasis id="B95ECF3E267CFFAEFB32B27864321CDA" box="[1243,1390,631,657]" italics="true" pageId="7" pageNumber="356">N. dicalcaria</emphasis>
|
||
<bibRefCitation id="EFBB6EDD267CFFAEFA9DB277605C1CFE" author="Raven, R. J." pageId="7" pageNumber="356" pagination="1 - 51" refId="ref7704" refString="Raven, R. J. (1984) A new diplurid genus from eastern Australia and a related Aname species (Diplurinae: Dipluridae: Araneae). Australian Journal of Zoology Supplementary Series, 96, 1 - 51. https: // doi. org / 10.1071 / AJZS 096" type="journal article" year="1984">Raven 1984</bibRefCitation>
|
||
</taxonomicName>
|
||
) have macrosetae on the male leg I tibia which are produced into a megaspur-like structure; these species can be distinguished from all other Anamini, except species of
|
||
<taxonomicName id="4C2A68AF267CFFAEFCB3B2BF65121C92" authority="Main, 1985" authorityName="Main" authorityYear="1985" box="[858,1102,702,729]" class="Arachnida" family="Nemesiidae" genus="Teyloides" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95ECF3E267CFFAEFCB3B2BF629F1C92" box="[858,963,703,729]" italics="true" pageId="7" pageNumber="356">Teyloides</emphasis>
|
||
<bibRefCitation id="EFBB6EDD267CFFAEFC23B2BF65121C92" author="Main, B. Y." box="[970,1102,702,729]" pageId="7" pageNumber="356" pagination="743 - 759" refId="ref7498" refString="Main, B. Y. (1985) Further studies on Australian Diplurinae: a review of the genera of the Teylini (Araneae: Mygalomorphae: Dipluridae). Australian Journal of Zoology, 33, 743 - 759. https: // doi. org / 10.1071 / ZO 9850743" type="journal article" year="1985">Main, 1985</bibRefCitation>
|
||
</taxonomicName>
|
||
, by the presence of a reflexed embolus; and from
|
||
<taxonomicName id="4C2A68AF267CFFAEFE9BB2E360871CB6" authorityName="Main" authorityYear="1985" box="[370,475,739,765]" class="Arachnida" family="Nemesiidae" genus="Teyloides" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95ECF3E267CFFAEFE9BB2E360871CB6" box="[370,475,739,765]" italics="true" pageId="7" pageNumber="356">Teyloides</emphasis>
|
||
</taxonomicName>
|
||
by the absence of coiled spermathecae in females.
|
||
</paragraph>
|
||
<caption id="DF5543A4267CFFAEFF7EB6A261BD196D" ID-DOI="http://doi.org/10.5281/zenodo.3460196" ID-Zenodo-Dep="3460196" httpUri="https://zenodo.org/record/3460196/files/figure.png" pageId="7" pageNumber="356" startId="7.[151,250,1698,1723]" targetBox="[159,1421,797,1662]" targetPageId="7">
|
||
<paragraph id="8B95132C267CFFAEFF7EB6A261BD196D" blockId="7.[151,1437,1698,1830]" pageId="7" pageNumber="356">
|
||
<emphasis id="B95ECF3E267CFFAEFF7EB6A2604A18F0" bold="true" box="[151,278,1698,1723]" pageId="7" pageNumber="356">FIGURE 2.</emphasis>
|
||
Map showing collection records of
|
||
<taxonomicName id="4C2A68AF267CFFAEFD7FB6A363E318F0" authorityName="Main" authorityYear="1975" box="[662,703,1699,1723]" class="Arachnida" family="Nemesiidae" genus="Teyl" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95ECF3E267CFFAEFD7FB6A363E318F0" box="[662,703,1699,1723]" italics="true" pageId="7" pageNumber="356">Teyl</emphasis>
|
||
</taxonomicName>
|
||
from the Pilbara bioregion (PIL) of Western Australia (see Fig. 1 for details). Black circles denote identified specimens of
|
||
<taxonomicName id="4C2A68AF267CFFAEFD57B6C762181894" authority="Huey & Rix & Wilson & Hillyer & Harvey, 2019" authorityName="Huey & Rix & Wilson & Hillyer & Harvey" authorityYear="2019" box="[702,836,1735,1759]" class="Arachnida" family="Nemesiidae" genus="Teyl" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="species" species="heuretes" status="sp. nov.">
|
||
<emphasis id="B95ECF3E267CFFAEFD57B6C762181894" box="[702,836,1735,1759]" italics="true" pageId="7" pageNumber="356">Teyl heuretes</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26D7245267CFFAEFCA2B6C762C21895" box="[843,926,1735,1758]" pageId="7" pageNumber="356" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; open circles denote unidentified juveniles. Other mapped IBRA 7.0 bioregions are as follows: CAR, Carnarvon; GAS, Gascoyne; GSD, Great Sandy Desert; LSD, Little Sandy Desert.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B95132C267CFFA1FF2EB75565EF1E9C" blockId="7.[151,1436,1877,2012]" lastBlockId="8.[151,1437,153,431]" lastPageId="8" lastPageNumber="357" pageId="7" pageNumber="356">
|
||
The diagnostic separation of
|
||
<taxonomicName id="4C2A68AF267CFFAEFDE5B75563661924" authorityName="Main" authorityYear="1975" box="[524,570,1877,1903]" class="Arachnida" family="Nemesiidae" genus="Teyl" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95ECF3E267CFFAEFDE5B75563661924" box="[524,570,1877,1903]" italics="true" pageId="7" pageNumber="356">Teyl</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2A68AF267CFFAEFD86B75663E21924" authorityName="Raven" authorityYear="1984" box="[623,702,1878,1903]" class="Arachnida" family="Nemesiidae" genus="Namea" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95ECF3E267CFFAEFD86B75663E21924" box="[623,702,1878,1903]" italics="true" pageId="7" pageNumber="356">Namea</emphasis>
|
||
</taxonomicName>
|
||
is currently problematic, due largely to the absence of a phylogenetic hypothesis for
|
||
<taxonomicName id="4C2A68AF267CFFAEFE93B77A609519D8" authorityName="Raven" authorityYear="1984" box="[378,457,1914,1939]" class="Arachnida" family="Nemesiidae" genus="Namea" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95ECF3E267CFFAEFE93B77A609519D8" box="[378,457,1914,1939]" italics="true" pageId="7" pageNumber="356">Namea</emphasis>
|
||
</taxonomicName>
|
||
, and the presence of a number of confounding diagnostic characters at the generic level (an issue discussed in detail by
|
||
<bibRefCitation id="EFBB6EDD267CFFAEFE15B79E63D419FC" author="Raven, R. J." box="[508,648,1949,1976]" pageId="7" pageNumber="356" pagination="1 - 51" refId="ref7704" refString="Raven, R. J. (1984) A new diplurid genus from eastern Australia and a related Aname species (Diplurinae: Dipluridae: Araneae). Australian Journal of Zoology Supplementary Series, 96, 1 - 51. https: // doi. org / 10.1071 / AJZS 096" type="journal article" year="1984">Raven 1984</bibRefCitation>
|
||
). At present,
|
||
<taxonomicName id="4C2A68AF267CFFAEFCF4B79D621719FC" authorityName="Main" authorityYear="1975" box="[797,843,1949,1975]" class="Arachnida" family="Nemesiidae" genus="Teyl" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="356" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95ECF3E267CFFAEFCF4B79D621719FC" box="[797,843,1949,1975]" italics="true" pageId="7" pageNumber="356">Teyl</emphasis>
|
||
</taxonomicName>
|
||
includes all of those species which occur in western and southern
|
||
<collectingCountry id="F33D53BC267CFFAEFEC4B7C160CB1990" box="[301,407,1985,2011]" name="Australia" pageId="7" pageNumber="356">Australia</collectingCountry>
|
||
, outside of the subtropical and tropical mesic zone of eastern
|
||
<collectingRegion id="49EEDDCE267CFFAEFBA4B7C1658D1990" box="[1101,1233,1985,2011]" country="Australia" name="Queensland" pageId="7" pageNumber="356">Queensland</collectingRegion>
|
||
and north-eastern
|
||
<collectingRegion id="49EEDDCE2673FFA1FF7EB099603D1EF8" box="[151,353,153,179]" country="Australia" name="New South Wales" pageId="8" pageNumber="357">New South Wales</collectingRegion>
|
||
. Whether
|
||
<taxonomicName id="4C2A68AF2673FFA1FE38B09A637C1EF8" authorityName="Raven" authorityYear="1984" box="[465,544,154,179]" class="Arachnida" family="Nemesiidae" genus="Namea" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="357" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95ECF3E2673FFA1FE38B09A637C1EF8" box="[465,544,154,179]" italics="true" pageId="8" pageNumber="357">Namea</emphasis>
|
||
</taxonomicName>
|
||
, as currently circumscribed (
|
||
<emphasis id="B95ECF3E2673FFA1FC8CB09A62FF1EF8" box="[869,931,154,179]" italics="true" pageId="8" pageNumber="357">sensu</emphasis>
|
||
<bibRefCitation id="EFBB6EDD2673FFA1FC43B09965681EF8" author="Raven, R. J." box="[938,1076,153,179]" pageId="8" pageNumber="357" pagination="1 - 51" refId="ref7704" refString="Raven, R. J. (1984) A new diplurid genus from eastern Australia and a related Aname species (Diplurinae: Dipluridae: Araneae). Australian Journal of Zoology Supplementary Series, 96, 1 - 51. https: // doi. org / 10.1071 / AJZS 096" type="journal article" year="1984">Raven 1984</bibRefCitation>
|
||
), represents one or more genera, and how it differs morphologically from
|
||
<taxonomicName id="4C2A68AF2673FFA1FD88B0BD63D31E9C" authority="Huey & Rix & Wilson & Hillyer & Harvey, 2019" authorityName="Huey & Rix & Wilson & Hillyer & Harvey" authorityYear="2019" box="[609,655,189,215]" class="Arachnida" family="Nemesiidae" genus="Teyl" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="357" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B95ECF3E2673FFA1FD88B0BD63D31E9C" box="[609,655,189,215]" italics="true" pageId="8" pageNumber="357">Teyl</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26D72452673FFA1FD7FB0BD63E71E9C" box="[662,699,189,215]" pageId="8" pageNumber="357" sensu="stricto">s.s.</taxonomicNameLabel>
|
||
, is currently the subject of ongoing research.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33040A72673FFA1FF2EB0E0644F1FE4" pageId="8" pageNumber="357" type="distribution">
|
||
<paragraph id="8B95132C2673FFA1FF2EB0E062AE1F2C" blockId="8.[151,1437,153,431]" pageId="8" pageNumber="357">
|
||
<emphasis id="B95ECF3E2673FFA1FF2EB0E0603D1EB0" bold="true" box="[199,353,224,251]" pageId="8" pageNumber="357">Distribution.</emphasis>
|
||
Species of
|
||
<taxonomicName id="4C2A68AF2673FFA1FE0AB0E1634D1EB0" authorityName="Huey & Rix & Wilson & Hillyer & Harvey" authorityYear="2019" box="[483,529,225,251]" class="Arachnida" family="Nemesiidae" genus="Teyl" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="357" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95ECF3E2673FFA1FE0AB0E1634D1EB0" box="[483,529,225,251]" italics="true" pageId="8" pageNumber="357">Teyl</emphasis>
|
||
</taxonomicName>
|
||
occur throughout the semi-arid and arid zones of southern
|
||
<collectingCountry id="F33D53BC2673FFA1FB40B0E1644F1EB0" box="[1193,1299,225,251]" name="Australia" pageId="8" pageNumber="357">Australia</collectingCountry>
|
||
, from
|
||
<collectingRegion id="49EEDDCE2673FFA1FAB5B0E160741F54" country="Australia" name="Western Australia" pageId="8" pageNumber="357">Western Australia</collectingRegion>
|
||
(
|
||
<figureCitation id="13110FA92673FFA1FEDEB105602B1F54" box="[311,375,261,287]" captionStart="FIGURE 1" captionStartId="6.[151,250,1984,2009]" captionTargetBox="[303,1280,748,1951]" captionTargetId="figure@6.[292,1294,736,1964]" captionTargetPageId="6" captionText="FIGURE 1. Map showing collection records of Teyl from Western Australia, including specimens from the Pilbara bioregion (inset box; see Fig. 2). Black circles denote male specimens; open circles denote female or juvenile specimens." figureDoi="http://doi.org/10.5281/zenodo.3460194" httpUri="https://zenodo.org/record/3460194/files/figure.png" pageId="8" pageNumber="357">Fig. 1</figureCitation>
|
||
) east to southern
|
||
<collectingRegion id="49EEDDCE2673FFA1FDA8B10563B31F54" box="[577,751,261,287]" country="Australia" name="South Australia" pageId="8" pageNumber="357">South Australia</collectingRegion>
|
||
and western
|
||
<collectingRegion id="49EEDDCE2673FFA1FC6AB10562BD1F54" box="[899,993,261,287]" country="Australia" name="Victoria" pageId="8" pageNumber="357">Victoria</collectingRegion>
|
||
. The genus reaches its highest diversity in south-western
|
||
<collectingRegion id="49EEDDCE2673FFA1FEB3B129637A1F08" box="[346,550,297,323]" country="Australia" name="Western Australia" pageId="8" pageNumber="357">Western Australia</collectingRegion>
|
||
, although a relatively few species can be found as far north as the Murchison, Gascoyne and Pilbara bioregions in the western-central arid zone (
|
||
<figureCitation id="13110FA92673FFA1FC96B14D629A1F2C" box="[895,966,333,359]" captionStart="FIGURE 1" captionStartId="6.[151,250,1984,2009]" captionTargetBox="[303,1280,748,1951]" captionTargetId="figure@6.[292,1294,736,1964]" captionTargetPageId="6" captionText="FIGURE 1. Map showing collection records of Teyl from Western Australia, including specimens from the Pilbara bioregion (inset box; see Fig. 2). Black circles denote male specimens; open circles denote female or juvenile specimens." figureDoi="http://doi.org/10.5281/zenodo.3460194" httpUri="https://zenodo.org/record/3460194/files/figure.png" pageId="8" pageNumber="357">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="13110FA92673FFA1FC3DB14D62B91F2C" box="[980,997,333,359]" captionStart="FIGURE 2" captionStartId="7.[151,250,1698,1723]" captionTargetBox="[159,1421,797,1662]" captionTargetId="figure@7.[151,1436,786,1673]" captionTargetPageId="7" captionText="FIGURE 2. Map showing collection records of Teyl from the Pilbara bioregion (PIL) of Western Australia (see Fig. 1 for details). Black circles denote identified specimens of Teyl heuretes sp. nov.; open circles denote unidentified juveniles. Other mapped IBRA 7.0 bioregions are as follows: CAR, Carnarvon; GAS, Gascoyne; GSD, Great Sandy Desert; LSD, Little Sandy Desert." figureDoi="http://doi.org/10.5281/zenodo.3460196" httpUri="https://zenodo.org/record/3460196/files/figure.png" pageId="8" pageNumber="357">2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B95132C2673FFA1FF2EB171644F1FE4" blockId="8.[151,1437,153,431]" pageId="8" pageNumber="357">
|
||
<emphasis id="B95ECF3E2673FFA1FF2EB17160C81FC0" bold="true" box="[199,404,368,395]" pageId="8" pageNumber="357">Included species.</emphasis>
|
||
The genus
|
||
<taxonomicName id="4C2A68AF2673FFA1FDF5B17163161FC0" authorityName="Huey & Rix & Wilson & Hillyer & Harvey" authorityYear="2019" box="[540,586,369,395]" class="Arachnida" family="Nemesiidae" genus="Teyl" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="357" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95ECF3E2673FFA1FDF5B17163161FC0" box="[540,586,369,395]" italics="true" pageId="8" pageNumber="357">Teyl</emphasis>
|
||
</taxonomicName>
|
||
includes seven described species, one of which is newly described in this study. Numerous undescribed species are also known from throughout western and southern
|
||
<collectingCountry id="F33D53BC2673FFA1FB4CB19564531FE4" box="[1189,1295,405,431]" name="Australia" pageId="8" pageNumber="357">Australia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |