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<document id="4F61E0853467BE380551F0DCD67A6EE9" ID-DOI="10.5281/zenodo.171388" ID-GBIF-Dataset="164b0a72-f861-4de7-a939-a3ad42b5c71d" ID-ISSN="1175­5326" ID-Zenodo-Dep="171388" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459721013543" checkinUser="plazi" docAuthor="Boucher, Stéphanie" docDate="2005" docId="03839D7E230CFFEDFEBEFD71FC96271F" docLanguage="en" docName="zt00993.pdf" docOrigin="Zootaxa 993" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Cerodontha (Dizygomyza) luctuosa Meigen" docType="treatment" docVersion="10" lastPageNumber="6" masterDocId="FFBAE5062308FFE8FFB6FFDFFFDB222B" masterDocTitle="Description of an unusual new Costa Rican species of Cerodontha (Dizygomyza) with additional notes on Neotropical species of Dizygomyza (Diptera: Agromyzidae)" masterLastPageNumber="8" masterPageNumber="1" pageNumber="5" updateTime="1698209392311" updateUser="plazi">
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<mods:title id="95AA40717243DADB40DC5412BB782EC3">Description of an unusual new Costa Rican species of Cerodontha (Dizygomyza) with additional notes on Neotropical species of Dizygomyza (Diptera: Agromyzidae)</mods:title>
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<mods:namePart id="C09625875F8D01B15411041EAD46B1E8">Boucher, Stéphanie</mods:namePart>
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<mods:date id="767FA4C3066482B5004AEB2985031138">2005</mods:date>
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<treatment id="03839D7E230CFFEDFEBEFD71FC96271F" ID-DOI="http://doi.org/10.5281/zenodo.6265863" ID-GBIF-Taxon="119335631" ID-Zenodo-Dep="6265863" LSID="urn:lsid:plazi:treatment:03839D7E230CFFEDFEBEFD71FC96271F" httpUri="http://treatment.plazi.org/id/03839D7E230CFFEDFEBEFD71FC96271F" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<subSubSection id="C3307FE3230CFFECFEBEFD71FE4320C0" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8B952C68230CFFECFEBEFD71FCF220E3" blockId="4.[264,809,686,747]" box="[264,809,686,712]" pageId="4" pageNumber="5">
<heading id="D0DD9B04230CFFECFEBEFD71FCF220E3" bold="true" box="[264,809,686,712]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<taxonomicName id="4C2A57EB230CFFECFEBEFD71FCF220E3" ID-CoL="8BDZT" authority="Meigen" authorityName="Meigen" box="[264,809,686,712]" class="Insecta" family="Agromyzidae" genus="Cerodontha" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="luctuosa" subGenus="Dizygomyza">
<emphasis id="B95EF07A230CFFECFEBEFD71FCF220E3" bold="true" box="[264,809,686,712]" pageId="4" pageNumber="5">
<emphasis id="B95EF07A230CFFECFEBEFD71FD6B20E3" bold="true" box="[264,688,686,712]" italics="true" pageId="4" pageNumber="5">Cerodontha (Dizygomyza) luctuosa</emphasis>
(Meigen)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B952C68230CFFECFEBEFD0EFE4320C0" blockId="4.[264,809,686,747]" box="[264,408,721,747]" pageId="4" pageNumber="5">
(
<figureCitation id="131130ED230CFFECFEA7FD0EFE5520C0" box="[273,398,721,747]" captionStart="FIGURES 8 12" captionStartId="5.[264,382,802,826]" captionTargetBox="[403,1194,290,776]" captionTargetId="figure@5.[379,1208,282,797]" captionTargetPageId="5" captionText="FIGURES 8 12. Cerodontha (Dizygomyza) luctuosa (Meigen). 8, phallus ventral view; 9, phallus lateral view; 10, epandrium with surstylus (ventral, left); 11, epandrium lateral view showing anal projection (ap); 12, ejaculatory apodeme." httpUri="https://zenodo.org/record/171390/files/figure.png" pageId="4" pageNumber="5">Figs. 812</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3307FE3230CFFEDFEBEFCC8FC96271F" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="description">
<paragraph id="8B952C68230CFFECFEBEFCC8FE1C21FA" blockId="4.[264,1324,791,1902]" pageId="4" pageNumber="5">Description. Head including antenna, brown, gena, lunule and inner margin of orbit paler, yellowish. Thorax uniformly brown, only anepisternum narrowly yellow on dorsal and posterior margins; calypter including margin and fringe yellow. Halter white. Legs completely brown, except foreknee yellow. Abdomen with anterior tergites yellow laterally, brown medially.</paragraph>
<paragraph id="8B952C68230CFFECFE8EFC00FD6324E5" blockId="4.[264,1324,791,1902]" pageId="4" pageNumber="5">
Frons width
<quantity id="4CD2818D230CFFECFE7DFC00FDAC21D2" box="[459,631,991,1017]" metricMagnitude="-4" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="4" pageNumber="5" unit="mm" value="0.425" valueMax="0.45" valueMin="0.4">0.400.45 mm</quantity>
; ratio of frons width to eye width 2.02.2; orbit broad: 0.2 0.25 width of frons at midpoint; frons slightly projecting above and in front of eye in profile, orbit forming only narrow ring below eye; eye bare; lunule with patch of silvery pubescence between antennae best seen in posterodorsal view; frons with microtomentum; 2 inclinate
<emphasis id="B95EF07A230CFFECFE3CFBA0FE7726B3" box="[394,428,1151,1176]" italics="true" pageId="4" pageNumber="5">ori</emphasis>
and 2 reclinate
<emphasis id="B95EF07A230CFFECFDD3FBA0FD5126B3" box="[613,650,1151,1176]" italics="true" pageId="4" pageNumber="5">ors</emphasis>
, anterior
<emphasis id="B95EF07A230CFFECFD4DFBA0FCC526B3" box="[763,798,1151,1176]" italics="true" pageId="4" pageNumber="5">ori</emphasis>
only slightly shorter than posterior
<emphasis id="B95EF07A230CFFECFB09FBA0FB3A26B3" box="[1215,1249,1151,1176]" italics="true" pageId="4" pageNumber="5">ori</emphasis>
; anterior
<emphasis id="B95EF07A230CFFECFE8DFB78FEBB26EB" box="[315,352,1191,1216]" italics="true" pageId="4" pageNumber="5">ors</emphasis>
0.750.80 length of posterior
<emphasis id="B95EF07A230CFFECFD7DFB78FD2B26EB" box="[715,752,1191,1216]" italics="true" pageId="4" pageNumber="5">ors</emphasis>
; 45 upright or reclinate orbital setulae, normal size, in one row, present anteriorly up to level of anterior
<emphasis id="B95EF07A230CFFECFC15FB10FC1326C3" box="[931,968,1231,1256]" italics="true" pageId="4" pageNumber="5">ors</emphasis>
; first flagellomere enlarged in male, covered with white pubescence; arista as long or shorter than eye height; arista with distinct pubescence and swollen basally for 40% of its length; gena slightly deeper at rear; gena height 0.20 times eye height at midpoint; lunule broad, in form of a semicircle, or a little higher; bases of antennae widely separated. Three postsutural dc, and 1 presutural dc decreasing in size: first (posterior) dc approximately twice the size of fourth (anterior) dc; acrostichals irregular, in about 45 rows; prsc absent. Midtibia with moderately strong apico­ventral bristle, about 0.3 length of first tarsomere. Wing length
<quantity id="4CD2818D230CFFECFBC2FA38FADC242A" box="[1140,1287,1511,1537]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.8" metricValueMin="2.7" pageId="4" pageNumber="5" unit="mm" value="2.75" valueMax="2.8" valueMin="2.7">2.72.8 mm</quantity>
in males; M1+2 ending closer to wing tip than R4+5; costa extending to M1+2; second costal sector 4.5 times length of fourth; last section of CuA1 equal or slightly longer than penultimate. Cross­vein r­m located at 0.50.6 the length of cell dm. Abdomen with stridulatory mechanism present in male (a line of scales along the center of the membrane connecting the abdominal tergites and sternites).
</paragraph>
<paragraph id="8B952C68230CFFECFE8EF903FDBB2545" blockId="4.[264,1324,791,1902]" pageId="4" pageNumber="5">
Male genitalia. Distiphallus with tubules diverging in ventral view (
<figureCitation id="131130ED230CFFECFBD0F903FB7424DD" box="[1126,1199,1756,1782]" captionStart="FIGURES 8 12" captionStartId="5.[264,382,802,826]" captionTargetBox="[403,1194,290,776]" captionTargetId="figure@5.[379,1208,282,797]" captionTargetPageId="5" captionText="FIGURES 8 12. Cerodontha (Dizygomyza) luctuosa (Meigen). 8, phallus ventral view; 9, phallus lateral view; 10, epandrium with surstylus (ventral, left); 11, epandrium lateral view showing anal projection (ap); 12, ejaculatory apodeme." httpUri="https://zenodo.org/record/171390/files/figure.png" pageId="4" pageNumber="5">Fig. 8</figureCitation>
), curving ventrally at apex in lateral view (
<figureCitation id="131130ED230CFFECFD20F8DBFD3B2535" box="[662,736,1796,1822]" captionStart="FIGURES 8 12" captionStartId="5.[264,382,802,826]" captionTargetBox="[403,1194,290,776]" captionTargetId="figure@5.[379,1208,282,797]" captionTargetPageId="5" captionText="FIGURES 8 12. Cerodontha (Dizygomyza) luctuosa (Meigen). 8, phallus ventral view; 9, phallus lateral view; 10, epandrium with surstylus (ventral, left); 11, epandrium lateral view showing anal projection (ap); 12, ejaculatory apodeme." httpUri="https://zenodo.org/record/171390/files/figure.png" pageId="4" pageNumber="5">Fig. 9</figureCitation>
). Surstylus with two stout anterodorsal, medially directed spines and 45 smaller ventral, medially directed spines (
<figureCitation id="131130ED230CFFECFBBFF8F3FBBB256D" box="[1033,1120,1836,1862]" captionStart="FIGURES 8 12" captionStartId="5.[264,382,802,826]" captionTargetBox="[403,1194,290,776]" captionTargetId="figure@5.[379,1208,282,797]" captionTargetPageId="5" captionText="FIGURES 8 12. Cerodontha (Dizygomyza) luctuosa (Meigen). 8, phallus ventral view; 9, phallus lateral view; 10, epandrium with surstylus (ventral, left); 11, epandrium lateral view showing anal projection (ap); 12, ejaculatory apodeme." httpUri="https://zenodo.org/record/171390/files/figure.png" pageId="4" pageNumber="5">Fig. 10</figureCitation>
). Anal projection of epandrium small (
<figureCitation id="131130ED230CFFECFE4AF88BFD892545" box="[508,594,1876,1902]" captionStart="FIGURES 8 12" captionStartId="5.[264,382,802,826]" captionTargetBox="[403,1194,290,776]" captionTargetId="figure@5.[379,1208,282,797]" captionTargetPageId="5" captionText="FIGURES 8 12. Cerodontha (Dizygomyza) luctuosa (Meigen). 8, phallus ventral view; 9, phallus lateral view; 10, epandrium with surstylus (ventral, left); 11, epandrium lateral view showing anal projection (ap); 12, ejaculatory apodeme." httpUri="https://zenodo.org/record/171390/files/figure.png" pageId="4" pageNumber="5">Fig. 11</figureCitation>
).
</paragraph>
<caption id="DF557CE0230DFFEDFEBEFCFDFD1B2155" httpUri="https://zenodo.org/record/171390/files/figure.png" pageId="5" pageNumber="6" targetBox="[403,1194,290,776]" targetPageId="5">
<paragraph id="8B952C68230DFFEDFEBEFCFDFD1B2155" blockId="5.[264,1323,802,894]" pageId="5" pageNumber="6">
<emphasis id="B95EF07A230DFFEDFEBEFCFDFE192111" bold="true" box="[264,450,802,826]" pageId="5" pageNumber="6">FIGURES 812.</emphasis>
<taxonomicName id="4C2A57EB230DFFEDFE7CFCFDFC6F2112" authority="(Meigen)" authorityName="Meigen" baseAuthorityName="Meigen" box="[458,948,802,825]" class="Insecta" family="Agromyzidae" genus="Cerodontha" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="luctuosa" subGenus="Dizygomyza">
<emphasis id="B95EF07A230DFFEDFE7CFCFDFC9E2112" box="[458,837,802,825]" italics="true" pageId="5" pageNumber="6">Cerodontha (Dizygomyza) luctuosa</emphasis>
(Meigen)
</taxonomicName>
. 8, phallus ventral view; 9, phallus lateral view; 10, epandrium with surstylus (ventral, left); 11, epandrium lateral view showing anal projection (ap); 12, ejaculatory apodeme.
</paragraph>
</caption>
<paragraph id="8B952C68230DFFEDFE8EFC6DFC96271F" blockId="5.[264,1325,946,1332]" pageId="5" pageNumber="6">
Comments. This Holarctic species was previously known in the Nearctic region from the following localities: California, Colorado, New York, Oregon, Washington, Alaska, Northwest Territories (
<bibRefCitation id="EFBB5199230DFFEDFDA2FBDDFC812637" author="Spencer" box="[532,858,1026,1052]" pageId="5" pageNumber="8" refString="Spencer, K. A. &amp; Steyskal, G. C. (1986) Manual of the Agromyzidae (Diptera) of the United States, U. S. Department of Agriculture, Agriculture Handbook, 638, 478 pp." type="book" year="1986">Spencer and Steyskal 1986</bibRefCitation>
;
<bibRefCitation id="EFBB5199230DFFEDFCD1FBDDFBD62637" author="Spencer" box="[871,1037,1026,1052]" pageId="5" pageNumber="8" refString="Spencer, K. A. (1969) The Agromyzidae of Canada and Alaska. Memoirs of the Entomological Society of Canada, 64, 1 - 311." type="journal article" year="1969">Spencer 1969</bibRefCitation>
). New localities include Texas (
<collectingCountry id="F33D6CF8230DFFEDFED7FBF5FE7A266F" box="[353,417,1066,1092]" name="United States of America" pageId="5" pageNumber="6">USA</collectingCountry>
: Texas, Davis Mts. State Park,
<geoCoordinate id="EE1E4AAF230DFFEDFC9DFBF5FC65266F" box="[811,958,1066,1092]" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="15" value="30.584446">30°3504”N</geoCoordinate>
;
<geoCoordinate id="EE1E4AAF230DFFEDFC66FBF5FBAF266F" box="[976,1140,1066,1092]" direction="west" orientation="longitude" pageId="5" pageNumber="6" precision="15" value="-103.92194">103°5519”W</geoCoordinate>
,
<date id="FF940AA8230DFFEDFB33FBF5FAF3266F" box="[1157,1320,1066,1092]" pageId="5" pageNumber="6" value="2001-05-29" valueMax="2001-05-30" valueMin="2001-05-29">2930.v.2001</date>
, Malaise trap, G.A. Dahlem,
<specimenCount id="9D2CE7E1230DFFEDFDE2FB8DFD7C2647" box="[596,679,1106,1132]" pageId="5" pageNumber="6" type="male">1 male</specimenCount>
, LEM) and Chiapas,
<collectingCountry id="F33D6CF8230DFFEDFC12FB8DFC252647" box="[932,1022,1106,1132]" name="Mexico" pageId="5" pageNumber="6">Mexico</collectingCountry>
(
<collectingCountry id="F33D6CF8230DFFEDFBB8FB8DFB532647" box="[1038,1160,1106,1132]" name="Mexico" pageId="5" pageNumber="6">MEXICO</collectingCountry>
: Chiapas,
<collectingCountry id="F33D6CF8230DFFEDFAB7FB8DFEA826BF" name="Saint Kitts and Nevis" pageId="5" pageNumber="6">San Cristobal</collectingCountry>
7000,
<date id="FF940AA8230DFFEDFE79FBA5FD9E26BF" box="[463,581,1146,1172]" pageId="5" pageNumber="6" value="1969-05-10">10.v.1969</date>
, H.J. Teskey,
<specimenCount id="9D2CE7E1230DFFEDFD5FFBA5FCE726BF" box="[745,828,1146,1172]" pageId="5" pageNumber="6" type="male">1 male</specimenCount>
, CNC). It could be expected that this species may be more widespread in the Neotropical region. Although the presence of the stridulatory organ in the male of
<taxonomicName id="4C2A57EB230DFFEDFD1EFB15FC0426C8" box="[680,991,1226,1251]" class="Insecta" family="Agromyzidae" genus="Cerodontha" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="luctuosa" subGenus="Dizygomyza">
<emphasis id="B95EF07A230DFFEDFD1EFB15FC0426C8" box="[680,991,1226,1251]" italics="true" pageId="5" pageNumber="6">Cerodontha (D.) luctuosa</emphasis>
</taxonomicName>
was noted by
<bibRefCitation id="EFBB5199230DFFEDFB2BFB15FE822727" author="Tschirnhaus" pageId="5" pageNumber="8" refString="Tschirnhaus, M. von. (1972) Unbekannte Stridulationsorgane bei Dipteren und ihre Bedeutung fur Taxonomie und Phylogenetik der Agromyziden (Diptera: Agromyzidae et Chamaemyiidae). Beitrage zur Entomologie, 21 (1971), 551 - 579." type="journal article" year="1972">Tschirnhaus (1972)</bibRefCitation>
, it was not mentioned in most other previous descriptions (e.g.
<bibRefCitation id="EFBB5199230DFFEDFBF3FB2DFAFE2727" author="Nowakowski" box="[1093,1317,1266,1292]" pageId="5" pageNumber="8" refString="Nowakowski, J. T. (1973) Monographie der europaischen Arten der Gattung Cerodontha Rond. (Diptera: Agromyzidae). Annales Zoologici (Warszawa), 31, 1 - 327." type="journal article" year="1973">Nowakowski 1973</bibRefCitation>
,
<bibRefCitation id="EFBB5199230DFFEDFEBEFAC5FE71271F" author="Spencer" box="[264,426,1306,1332]" pageId="5" pageNumber="8" refString="Spencer, K. A. (1969) The Agromyzidae of Canada and Alaska. Memoirs of the Entomological Society of Canada, 64, 1 - 311." type="journal article" year="1969">Spencer 1969</bibRefCitation>
,
<bibRefCitation id="EFBB5199230DFFEDFE01FAC5FE2F271F" author="Spencer" box="[439,500,1306,1332]" pageId="5" pageNumber="8" refString="Spencer, K. A. (1981) A revisionary study of the leaf-mining flies (Agromyzidae) of California. University of California, Division of Agricultural Sciences. Special Publication, 3273, 1 - 489." type="journal article" year="1981">1981</bibRefCitation>
,
<bibRefCitation id="EFBB5199230DFFEDFDB6FAC5FC9B271F" author="Spencer" box="[512,832,1306,1332]" pageId="5" pageNumber="8" refString="Spencer, K. A. &amp; Steyskal, G. C. (1986) Manual of the Agromyzidae (Diptera) of the United States, U. S. Department of Agriculture, Agriculture Handbook, 638, 478 pp." type="book" year="1986">Spencer and Steyskal 1986</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>