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<mods:title id="C5834535011AA5238148B6C5D528583E">Redescription of the poorly known ghost shrimp species, Scallasis amboinae Bate, 1888, review of the genus, and description of a new species from the Ryukyu Islands, Japan (Decapoda: Axiidea: Callianassidae)</mods:title>
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Genus
<taxonomicName id="4C2A5ACD3476E4789B8DFE84FE36FEFB" ID-CoL="a1cb4223-1103-48a6-855c-a7b5a81c111d" authority="Bate, 1888" authorityName="Bate" authorityYear="1888" box="[238,482,297,324]" class="Malacostraca" family="Callianassidae" genus="Scallasis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="genus">
<emphasis id="B95EFD5C3476E4789B8DFE84FE8CFEFC" bold="true" box="[238,344,297,324]" italics="true" pageId="1" pageNumber="402">Scallasis</emphasis>
<bibRefCitation id="EFBB5CBF3476E4789A03FE87FE36FEFB" author="Bate, S. C." box="[352,482,297,324]" pageId="1" pageNumber="402" pagination="1 - 942" refId="ref11048" refString="Bate, S. C. (1888) Report on the Crustacea Macrura collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. &quot; Challenger &quot; during the Years 1873 - 76, Zoology, 24 (i-xc), 1 - 942, pls. 1 - 157." type="journal article" year="1888">Bate, 1888</bibRefCitation>
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<treatmentCitation id="0A8B075F3476E4789BF4FEC5FE58FE39" author="Bate, S. C." box="[151,396,360,385]" page="34" pageId="1" pageNumber="402" year="1888">
<taxonomicName id="4C2A5ACD3476E4789BF4FEC5FE58FE39" ID-CoL="a1cb4223-1103-48a6-855c-a7b5a81c111d" authority="Bate, 1888: 34" authorityName="Bate" authorityPageNumber="34" authorityYear="1888" box="[151,396,360,385]" class="Malacostraca" family="Callianassidae" genus="Scallasis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="genus">
<emphasis id="B95EFD5C3476E4789BF4FEC5FF25FE38" box="[151,241,360,384]" italics="true" pageId="1" pageNumber="402">Scallasis</emphasis>
<bibRefCitation id="EFBB5CBF3476E4789B9BFEC4FE58FE39" author="Bate, S. C." box="[248,396,360,385]" pageId="1" pageNumber="402" pagination="1 - 942" refId="ref11048" refString="Bate, S. C. (1888) Report on the Crustacea Macrura collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. &quot; Challenger &quot; during the Years 1873 - 76, Zoology, 24 (i-xc), 1 - 942, pls. 1 - 157." type="journal article" year="1888">Bate, 1888: 34</bibRefCitation>
</taxonomicName>
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.—
<treatmentCitation id="0A8B075F3476E4789ACAFEC5FD01FE38" author="Manning, R. B. &amp; Felder, D. L." box="[425,725,360,385]" page="780" pageId="1" pageNumber="402" year="1991">
<bibRefCitation id="EFBB5CBF3476E4789ACAFEC5FD01FE38" author="Manning, R. B. &amp; Felder, D. L." box="[425,725,360,385]" pageId="1" pageNumber="402" pagination="764 - 792" refId="ref11522" refString="Manning, R. B. &amp; Felder, D. L. (1991) Revision of the American Callianassidae (Crustacea, Decapoda, Thalassinidea). Proceedings of the Biological Society of Washington, 104, 764 - 792." type="journal article" year="1991">Manning &amp; Felder 1991: 780</bibRefCitation>
</treatmentCitation>
.—
<treatmentCitation id="0A8B075F3476E4789991FEC5FC1FFE38" author="Poore, G. C. B. &amp; Dworschak, P. C. &amp; Robles, R. &amp; Mantelatto, F. L. &amp; Felder, D. L." box="[754,971,360,384]" page="99" pageId="1" pageNumber="402" year="2019">
<bibRefCitation id="EFBB5CBF3476E4789991FEC5FC1FFE38" author="Poore, G. C. B. &amp; Dworschak, P. C. &amp; Robles, R. &amp; Mantelatto, F. L. &amp; Felder, D. L." box="[754,971,360,384]" pageId="1" pageNumber="402" pagination="73 - 146" refId="ref11625" refString="Poore, G. C. B., Dworschak, P. C., Robles, R., Mantelatto, F. L. &amp; Felder, D. L. (2019) A new classification of Callianassidae and related families (Crustacea: Decapoda: Axiidea) derived from a molecular phylogeny with morphological support. Memoirs of Museum Victoria, 78, 73 - 146. https: // doi. org / 10.24199 / j. mmv. 2019.78.05" type="journal article" year="2019">
Poore
<emphasis id="B95EFD5C3476E4789855FEC4FCB6FE38" box="[822,866,360,384]" italics="true" pageId="1" pageNumber="402">et al</emphasis>
. 2019: 99
</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8B95214E3476E4789BF4FE2AFCD8FE27" blockId="1.[151,975,360,446]" box="[151,780,390,415]" pageId="1" pageNumber="402">
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<taxonomicName id="4C2A5ACD3476E4789BF4FE2AFE54FE27" authority="" box="[151,384,390,415]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="subGenus" subGenus="Scallasis">
<emphasis id="B95EFD5C3476E4789BF4FE2AFEC4FE27" box="[151,272,391,415]" italics="true" pageId="1" pageNumber="402">Callianassa</emphasis>
(
<emphasis id="B95EFD5C3476E4789A7CFE2AFEADFE27" box="[287,377,391,415]" italics="true" pageId="1" pageNumber="402">Scallasis</emphasis>
)
</taxonomicName>
.—
<treatmentCitation id="0A8B075F3476E4789AFDFE2BFD78FE27" author="Borradaile, L. A." box="[414,684,390,415]" page="547" pageId="1" pageNumber="402" year="1903">
<bibRefCitation id="EFBB5CBF3476E4789AFDFE2BFD78FE27" author="Borradaile, L. A." box="[414,684,390,415]" pageId="1" pageNumber="402" pagination="534 - 551" refId="ref11120" refString="Borradaile, L. A. (1903) On the classification of the Thalassinidea. Annals and Magazine of Natural History, Series 7, 12, 534 - 551 + 638 (addendum) https: // doi. org / 10.1080 / 00222930309487049" type="journal article" year="1903">Borradaile 1903: 547548</bibRefCitation>
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(partim).
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</paragraph>
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<treatmentCitationGroup id="AB3A06603476E4789BF4FE0BFDB5FE06" box="[151,609,421,446]" pageId="1" pageNumber="402">
<emphasis id="B95EFD5C3476E4789BF4FE0BFED3FE06" box="[151,263,422,446]" italics="true" pageId="1" pageNumber="402">
<taxonomicName id="4C2A5ACD3476E4789BF4FE0BFED6FE06" ID-CoL="8f84ed4e-2899-4c8d-87f2-78cc7a7e5a75" authorityName="Spence Bate" authorityYear="1888" box="[151,258,422,446]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="genus">Cheramus</taxonomicName>
.
</emphasis>
<treatmentCitation id="0A8B075F3476E4789A43FE08FDD5FE06" author="Sakai, K." box="[288,513,421,446]" page="363" pageId="1" pageNumber="402" year="2011">
<bibRefCitation id="EFBB5CBF3476E4789A43FE08FDD5FE06" author="Sakai, K." box="[288,513,421,446]" pageId="1" pageNumber="402" pagination="1 - 616" refId="ref12046" refString="Sakai, K. (2011 a) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. https: // doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai 2011a: 363365</bibRefCitation>
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(partim).
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</paragraph>
</subSubSection>
<subSubSection id="C33072C53476E4789BF4FE4AFC6CFA85" pageId="1" pageNumber="402" type="discussion">
<paragraph id="8B95214E3476E4789BF4FE4AFBFDFDD5" blockId="1.[151,1437,487,1341]" pageId="1" pageNumber="402">
<emphasis id="B95EFD5C3476E4789BF4FE4AFED0FDB9" bold="true" box="[151,260,487,513]" pageId="1" pageNumber="402">Remarks</emphasis>
. The genus was rediagnosed by
<bibRefCitation id="EFBB5CBF3476E47899E4FE4AFC46FDB9" author="Poore, G. C. B. &amp; Dworschak, P. C. &amp; Robles, R. &amp; Mantelatto, F. L. &amp; Felder, D. L." box="[647,914,487,513]" pageId="1" pageNumber="402" pagination="73 - 146" refId="ref11625" refString="Poore, G. C. B., Dworschak, P. C., Robles, R., Mantelatto, F. L. &amp; Felder, D. L. (2019) A new classification of Callianassidae and related families (Crustacea: Decapoda: Axiidea) derived from a molecular phylogeny with morphological support. Memoirs of Museum Victoria, 78, 73 - 146. https: // doi. org / 10.24199 / j. mmv. 2019.78.05" type="journal article" year="2019">
Poore
<emphasis id="B95EFD5C3476E47899B0FE45FCDCFDB9" box="[723,776,487,513]" italics="true" pageId="1" pageNumber="402">et al</emphasis>
. (2019: 99)
</bibRefCitation>
after thorough molecular and morphological analysis (
<bibRefCitation id="EFBB5CBF3476E4789B9CFDA6FE04FD9D" author="Robles, R. &amp; Dworschak, P. C. &amp; Felder, D. L. &amp; Poore, G. C. B. &amp; Mantelatto, F. L." box="[255,464,523,549]" pageId="1" pageNumber="402" refId="ref11711" refString="Robles, R., Dworschak, P. C., Felder, D. L., Poore, G. C. B. &amp; Mantelatto, F. L. (2020) A molecular phylogeny of Callianassidae and related families (Crustacea: Decapoda: Axiidea) with morphological support. Invertebrate Systematics, 34. https: // doi. org / 10.1071 / IS 19021" type="book" year="2020">
Robles
<emphasis id="B95EFD5C3476E4789A36FDA1FE52FD9D" box="[341,390,523,549]" italics="true" pageId="1" pageNumber="402">et al</emphasis>
. 2020
</bibRefCitation>
). The genus is recognised by the combination of a narrow rostrum, eyestalks with acute apices, a narrow maxilliped 3 ischium-merus, a simple or bifid spine on the merus of the major cheliped, and strong spiniform setae on the face of the uropodal endopod and on the face of the telson.
</paragraph>
<paragraph id="8B95214E3476E4789BA4FDDAFE57FCFD" blockId="1.[151,1437,487,1341]" pageId="1" pageNumber="402">
As noted above,
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Poore
<emphasis id="B95EFD5C3476E4789AA8FDD5FE28FD29" box="[459,508,631,657]" italics="true" pageId="1" pageNumber="402">et al</emphasis>
. (2019
</bibRefCitation>
: tab. 1, 2) included seven species, all from the Indo-West Pacific:
<taxonomicName id="4C2A5ACD3476E4789E52FDDAFF6BFD0D" class="Malacostraca" family="Callianassidae" genus="Scallasis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="species" species="amboinae">
<emphasis id="B95EFD5C3476E4789E52FDDAFF6BFD0D" italics="true" pageId="1" pageNumber="402">S. amboinae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A5ACD3476E4789BADFD36FE47FD0D" baseAuthorityName="Sakai" baseAuthorityYear="2002" box="[206,403,667,693]" class="Malacostraca" family="Callianassidae" genus="Scallasis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="species" species="andamaniensis">
<emphasis id="B95EFD5C3476E4789BADFD36FE47FD0D" box="[206,403,667,693]" italics="true" pageId="1" pageNumber="402">S. andamaniensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A5ACD3476E4789AC1FD36FDEDFD0D" baseAuthorityName="Ngoc-Ho" baseAuthorityYear="1991" box="[418,569,667,693]" class="Malacostraca" family="Callianassidae" genus="Scallasis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="species" species="caledonica">
<emphasis id="B95EFD5C3476E4789AC1FD36FDEDFD0D" box="[418,569,667,693]" italics="true" pageId="1" pageNumber="402">S. caledonica</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A5ACD3476E478992BFD36FD15FD0D" baseAuthorityName="Sakai" baseAuthorityYear="2002" box="[584,705,667,693]" class="Malacostraca" family="Callianassidae" genus="Scallasis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="species" species="contipes">
<emphasis id="B95EFD5C3476E478992BFD36FD15FD0D" box="[584,705,667,693]" italics="true" pageId="1" pageNumber="402">S. contipes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A5ACD3476E47899B3FD36FC86FD0D" baseAuthorityName="de Man" baseAuthorityYear="1928" box="[720,850,667,693]" class="Malacostraca" family="Callianassidae" genus="Scallasis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="species" species="pygmaea">
<emphasis id="B95EFD5C3476E47899B3FD36FC86FD0D" box="[720,850,667,693]" italics="true" pageId="1" pageNumber="402">S. pygmaea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A5ACD3476E4789802FD36FBF0FD0D" baseAuthorityName="Sakai" baseAuthorityYear="1970" box="[865,1060,667,693]" class="Malacostraca" family="Callianassidae" genus="Scallasis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="species" species="spinophthalma">
<emphasis id="B95EFD5C3476E4789802FD36FBF0FD0D" box="[865,1060,667,693]" italics="true" pageId="1" pageNumber="402">S. spinophthalma</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2A5ACD3476E4789F3FFD36FB0CFD0D" baseAuthorityName="Grebenyuk" baseAuthorityYear="1975" box="[1116,1240,667,693]" class="Malacostraca" family="Callianassidae" genus="Scallasis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="species" species="tonkinae">
<emphasis id="B95EFD5C3476E4789F3FFD36FB0CFD0D" box="[1116,1240,667,693]" italics="true" pageId="1" pageNumber="402">S. tonkinae</emphasis>
</taxonomicName>
. Some, however, have been synonymised with others and later removed from synonymy (see
<taxonomicName id="4C2A5ACD3476E4789F72FD12FB4AFD61" box="[1041,1182,703,729]" class="Malacostraca" family="Callianassidae" genus="Scallasis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="species" species="amboinae">
<emphasis id="B95EFD5C3476E4789F72FD12FB4AFD61" box="[1041,1182,703,729]" italics="true" pageId="1" pageNumber="402">S. amboinae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A5ACD3476E4789FD3FD12FA9EFD61" baseAuthorityName="Ngoc-Ho" baseAuthorityYear="1991" box="[1200,1354,703,729]" class="Malacostraca" family="Callianassidae" genus="Scallasis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="species" species="caledonica">
<emphasis id="B95EFD5C3476E4789FD3FD12FA9EFD61" box="[1200,1354,703,729]" italics="true" pageId="1" pageNumber="402">S. caledonica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2A5ACD3476E4789EE4FD12FF23FD45" baseAuthorityName="Grebenyuk" baseAuthorityYear="1975" class="Malacostraca" family="Callianassidae" genus="Scallasis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="species" species="tonkinae">
<emphasis id="B95EFD5C3476E4789EE4FD12FF23FD45" italics="true" pageId="1" pageNumber="402">S. tonkinae</emphasis>
</taxonomicName>
below). In his major axiidean review,
<bibRefCitation id="EFBB5CBF3476E47899CCFD4EFC9FFD45" author="Sakai, K." box="[687,843,739,765]" pageId="1" pageNumber="402" pagination="1 - 616" refId="ref12046" refString="Sakai, K. (2011 a) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. https: // doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai (2011a)</bibRefCitation>
regarded the four specific taxa,
<taxonomicName id="4C2A5ACD3476E4789FD8FD4EFA41FD45" box="[1211,1429,739,765]" class="Malacostraca" family="Callianassidae" genus="Scallasis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="species" species="amboinae">
<emphasis id="B95EFD5C3476E4789FD8FD4EFA41FD45" box="[1211,1429,739,765]" italics="true" pageId="1" pageNumber="402">Scallasis amboinae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A5ACD3476E4789BF4FCAAFE4BFC99" authorityName="Ngoc-Ho" authorityYear="1991" box="[151,415,775,801]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="species" species="caledonica">
<emphasis id="B95EFD5C3476E4789BF4FCAAFE4BFC99" box="[151,415,775,801]" italics="true" pageId="1" pageNumber="402">Callianassa caledonica</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A5ACD3476E4789ACFFCAAFCFEFC99" baseAuthorityName="De Man" baseAuthorityYear="1928" box="[428,810,775,801]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="species" species="pygmaea" subGenus="Cheramus">
<emphasis id="B95EFD5C3476E4789ACFFCAAFDE6FC99" box="[428,562,775,801]" italics="true" pageId="1" pageNumber="402">Callianassa</emphasis>
(
<emphasis id="B95EFD5C3476E4789921FCAAFD61FC99" box="[578,693,775,801]" italics="true" pageId="1" pageNumber="402">Cheramus</emphasis>
)
<emphasis id="B95EFD5C3476E47899A6FCA5FCFEFC99" box="[709,810,776,801]" italics="true" pageId="1" pageNumber="402">pygmaea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2A5ACD3476E4789803FCAAFB1EFC99" authorityName="Grebenjuk" authorityYear="1975" box="[864,1226,775,801]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="species" species="tonkinae" subGenus="Scallasis">
<emphasis id="B95EFD5C3476E4789803FCAAFC32FC99" box="[864,998,775,801]" italics="true" pageId="1" pageNumber="402">Callianassa</emphasis>
(
<emphasis id="B95EFD5C3476E4789895FCAAFB8EFC99" box="[1014,1114,775,801]" italics="true" pageId="1" pageNumber="402">Scallasis</emphasis>
)
<emphasis id="B95EFD5C3476E4789F09FCAAFB1EFC99" box="[1130,1226,775,801]" italics="true" pageId="1" pageNumber="402">tonkinae</emphasis>
</taxonomicName>
as valid species of
<taxonomicName id="4C2A5ACD3476E4789BF4FC81FEAAFCFD" authority="Dana, 1852" authorityName="Dana" authorityYear="1852" box="[151,382,810,837]" class="Malacostraca" family="Callianassidae" genus="Trypaea" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="genus">
<emphasis id="B95EFD5C3476E4789BF4FC81FF27FCFD" box="[151,243,812,837]" italics="true" pageId="1" pageNumber="402">Trypaea</emphasis>
<bibRefCitation id="EFBB5CBF3476E4789B99FC86FEAAFCFD" author="Dana, J. D." box="[250,382,810,837]" pageId="1" pageNumber="402" pagination="10 - 28" refId="ref11166" refString="Dana, J. D. (1852) Macroura. Conspectus Crustaceorum &amp;. Conspectus of the Crustacea of the Exploring Expedition under Capt C. Wilkes, U. S. N. Proceedings of the Academy of Natural Sciences of Philadelphia, 6, 10 - 28." type="journal article" year="1852">Dana, 1852</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8B95214E3476E4789BA4FCE2FE5EFB15" blockId="1.[151,1437,487,1341]" pageId="1" pageNumber="402">
Characterisation of these species is limited without reference to
<typeStatus id="54919FEC3476E47898CEFCE2FC09FCD1" box="[941,989,847,873]" pageId="1" pageNumber="402">type</typeStatus>
material we have relied for most on published works but believe many descriptions and figures omit critical characters not usually noted in callianassid descriptions in the past and which have been often overlooked. For example, the facial spiniform setae on the uropodal rami were unexpected or considered unimportant by earlier authors and are difficult to observe without appropriate lighting or staining. The scaphocerite is acute and short in species we have observed but not mentioned in most descriptions. There appear few reliable differences between species in the rostrum (triangular, sharp, reaching the cornea) and eyestalks (tapering to acute anterior lobes) but we see species differences in maxilliped 3 and in the chelipeds. Adding to the difficulty differentiating species is the paucity of individuals, and differences between sexes, growth stages and individuals. Nevertheless, we have prepared short diagnoses of the seven species of
<taxonomicName id="4C2A5ACD3476E4789E00FBC2FF1EFB15" authorityName="Bate" authorityYear="1888" class="Malacostraca" family="Callianassidae" genus="Scallasis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="genus">
<emphasis id="B95EFD5C3476E4789E00FBC2FF1EFB15" italics="true" pageId="1" pageNumber="402">Scallasis</emphasis>
</taxonomicName>
recognised here.
</paragraph>
<paragraph id="8B95214E3476E4789BA4FB1AFC6CFA85" blockId="1.[151,1437,487,1341]" pageId="1" pageNumber="402">
<taxonomicName id="4C2A5ACD3476E4789BA4FB1AFE79FB69" baseAuthorityName="Ngoc-Ho" baseAuthorityYear="1991" box="[199,429,1207,1233]" class="Malacostraca" family="Callianassidae" genus="Scallasis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="species" species="caledonica">
<emphasis id="B95EFD5C3476E4789BA4FB1AFE79FB69" box="[199,429,1207,1233]" italics="true" pageId="1" pageNumber="402">Scallasis caledonica</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A5ACD3476E4789ADFFB1AFDF9FB69" authority="Komai &amp; Poore &amp; Fujita, 2020" authorityName="Komai &amp; Poore &amp; Fujita" authorityYear="2020" box="[444,557,1207,1233]" class="Malacostraca" family="Callianassidae" genus="Scallasis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="species" species="inermis" status="sp. nov.">
<emphasis id="B95EFD5C3476E4789ADFFB1AFDF9FB69" box="[444,557,1207,1233]" italics="true" pageId="1" pageNumber="402">S. inermis</emphasis>
</taxonomicName>
<emphasis id="B95EFD5C3476E4789957FB1AFDA0FB69" bold="true" box="[564,628,1207,1233]" pageId="1" pageNumber="402">
<taxonomicNameLabel id="A26D40273476E4789957FB1AFDA0FB69" box="[564,628,1207,1233]" pageId="1" pageNumber="402" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C2A5ACD3476E47899CFFB1AFCBBFB69" baseAuthorityName="Sakai" baseAuthorityYear="1970" box="[684,879,1207,1233]" class="Malacostraca" family="Callianassidae" genus="Scallasis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="species" species="spinophthalma">
<emphasis id="B95EFD5C3476E47899CFFB1AFCBBFB69" box="[684,879,1207,1233]" italics="true" pageId="1" pageNumber="402">S. spinophthalma</emphasis>
</taxonomicName>
are known from both sexes. Pleopod 1 is present in males of all three but variously developed, possibly due to the maturity of the specimens examined. Pleopod 2 is present in two species, not in
<taxonomicName id="4C2A5ACD3476E4789A8BFB52FDABFAA1" baseAuthorityName="Ngoc-Ho" baseAuthorityYear="1991" box="[488,639,1279,1305]" class="Malacostraca" family="Callianassidae" genus="Scallasis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="species" species="caledonica">
<emphasis id="B95EFD5C3476E4789A8BFB52FDABFAA1" box="[488,639,1279,1305]" italics="true" pageId="1" pageNumber="402">S. caledonica</emphasis>
</taxonomicName>
. Chelipeds are apparently sexually dimorphic only in
<taxonomicName id="4C2A5ACD3476E4789F87FB52FA81FAA1" authority="Komai &amp; Poore &amp; Fujita, 2020" authorityName="Komai &amp; Poore &amp; Fujita" authorityYear="2020" box="[1252,1365,1279,1305]" class="Malacostraca" family="Callianassidae" genus="Scallasis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="402" phylum="Arthropoda" rank="species" species="inermis" status="sp. nov.">
<emphasis id="B95EFD5C3476E4789F87FB52FA81FAA1" box="[1252,1365,1279,1305]" italics="true" pageId="1" pageNumber="402">S. inermis</emphasis>
</taxonomicName>
<emphasis id="B95EFD5C3476E4789E3FFB52FA48FAA1" bold="true" box="[1372,1436,1279,1305]" pageId="1" pageNumber="402">
<taxonomicNameLabel id="A26D40273476E4789E3FFB52FA48FAA1" box="[1372,1436,1279,1305]" pageId="1" pageNumber="402" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
but this species may be the only one where a fully adult male is known.
</paragraph>
</subSubSection>
</treatment>
</document>