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<document id="2D5BC98E33CBA26CA4CD3925B38D4DAC" ID-CLB-Dataset="50356" ID-DOI="10.5281/zenodo.273633" ID-GBIF-Dataset="91494720-244e-4dd6-8b81-24cc0ba4aee7" ID-ISSN="1175-5326" ID-Zenodo-Dep="273633" checkinTime="1459787272321" checkinUser="plazi" docAuthor="Monguillot, Julio C., Cabrera, Mario R., Acosta, Juan C. &amp; Villavicencio, Jose" docDate="2006" docId="03838C1E5B1F3355FEB540534482FDBA" docLanguage="en" docName="zt01361p043.pdf" docOrigin="Zootaxa 1361" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@632cf7d3" docStyleName="zootaxa.0000.journal_article" docTitle="Liolaemus cinereus Monguillot, Cabrera, Acosta &amp; Villavicencio, 2006, sp. nov." docType="treatment" docVersion="7" lastPageNumber="40" masterDocId="FFBAF4665B1E335DFFBD46254516FF9C" masterDocTitle="A new species of Liolaemus (Reptilia: Iguanidae) from San Guillermo National Park, western Argentina" masterLastPageNumber="43" masterPageNumber="33" pageNumber="34" updateTime="1732908241336" updateUser="ExternalLinkService">
<mods:mods id="F0C6C286C60BC701F251DD7736FB6E6C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="790C2FC32672952FBECCE12415DC482B">
<mods:title id="3C3E0E9759CFAD0836565A686247E7A7">A new species of Liolaemus (Reptilia: Iguanidae) from San Guillermo National Park, western Argentina</mods:title>
</mods:titleInfo>
<mods:name id="1FD925AAD10A46C0879461077708A3F5" type="personal">
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<mods:namePart id="2311FA93329757D43921FE75AB53C8EF">Monguillot, Julio C.</mods:namePart>
</mods:name>
<mods:name id="9959F0DA088BA87D39D908FC10B58656" type="personal">
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<mods:namePart id="C4A176C72226D4BF8184B89E400009C4">Cabrera, Mario R.</mods:namePart>
</mods:name>
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<mods:role id="D85B7BE97696BC7488D772690D7C27BC">
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<mods:namePart id="D4C3A96A2B918E53BFDBF06BE5B32FD4">Acosta, Juan C.</mods:namePart>
</mods:name>
<mods:name id="5B5FDCACFDA892C2A4259D62972A0D4E" type="personal">
<mods:role id="6C86EE6B268EB93063617CAF8B2A88A8">
<mods:roleTerm id="AA779968471F0BC57A93F7074EDA3C73">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6442E9EBAD91CDEF23933FB4CB2519E4">Villavicencio, Jose</mods:namePart>
</mods:name>
<mods:typeOfResource id="7B4E1FF16C90134C93F622A0579F2143">text</mods:typeOfResource>
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<mods:title id="B46FCEB38A1EF4589D585A894F99EF84">Zootaxa</mods:title>
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<mods:part id="19A6F693650EE1E480A8A4CEB99726F0">
<mods:date id="7900D0C622169C48E27FFBFA7C0EC6A6">2006</mods:date>
<mods:detail id="931E31EC449703C45853DADDFEC91F8B" type="volume">
<mods:number id="97A543E82308634D7E3E900CEF15D1C3">1361</mods:number>
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<mods:identifier id="8B0C92C0174B9541991C2EA6BE8CB9D0" type="DOI">10.5281/zenodo.273633</mods:identifier>
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<mods:identifier id="541B04634F04F71AE1C6C36C8B838AEB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="51A0A49DCAE26123990A6786A4895A66" type="Zenodo-Dep">273633</mods:identifier>
</mods:mods>
<treatment id="03838C1E5B1F3355FEB540534482FDBA" ID-GBIF-Taxon="119349435" LSID="urn:lsid:plazi:treatment:03838C1E5B1F3355FEB540534482FDBA" httpUri="http://treatment.plazi.org/id/03838C1E5B1F3355FEB540534482FDBA" lastPageId="8" lastPageNumber="40" pageId="1" pageNumber="34" scope_family="Iguanidae">
<subSubSection id="C3306E835B1F335CFEB5405344B1F92F" pageId="1" pageNumber="34" type="nomenclature">
<paragraph id="8B953D085B1F335CFEB540534777F90C" blockId="1.[264,609,1654,1715]" box="[264,609,1654,1680]" pageId="1" pageNumber="34">
<heading id="D0DD8A645B1F335CFEB540534777F90C" bold="true" box="[264,609,1654,1680]" fontSize="11" level="1" pageId="1" pageNumber="34" reason="1">
<emphasis id="B95EE11A5B1F335CFEB540534777F90C" bold="true" box="[264,609,1654,1680]" pageId="1" pageNumber="34">
<taxonomicName id="4C2A468B5B1F335CFEB5405344E3F90C" box="[264,501,1654,1680]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="34" phylum="Chordata" rank="species" species="cinereus" status="sp. nov.">
<emphasis id="B95EE11A5B1F335CFEB5405344E3F90C" bold="true" box="[264,501,1654,1680]" italics="true" pageId="1" pageNumber="34">Liolaemus cinereus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D5C615B1F335CFDBE40534777F90C" box="[515,609,1654,1680]" pageId="1" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B953D085B1F335CFEB540BC44B1F92F" blockId="1.[264,609,1654,1715]" box="[264,423,1689,1715]" pageId="1" pageNumber="34">
(
<figureCitation id="1311218D5B1F335CFEAD40BC448BF92F" box="[272,413,1689,1715]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="4.[264,367,1454,1478]" captionStartId-1="5.[264,367,1335,1359]" captionStartId-2="6.[264,367,1380,1404]" captionTargetBox-0="[267,1319,794,1430]" captionTargetBox-1="[267,1319,630,1327]" captionTargetBox-2="[268,1318,310,1372]" captionTargetId-0="figure@4.[267,1319,794,1430]" captionTargetId-1="figure@5.[267,1319,628,1329]" captionTargetId-2="figure@6.[268,1319,308,1374]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 1. The holotype of Liolaemus cinereus, adult male, in life. The specimen is actually grey (see Figure 2); the brownish shade is an artifact due to lighting used to portray it." captionText-1="FIGURE 2. The holotype of Liolaemus cinereus in life, showing the grey lining into the prehumeral fold (here extended) and the dark prescapular spot." captionText-2="FIGURE 3. Comparative ventral view in life of the female paratype MLP. R 5140 (left) and the male holotype (right) of Liolaemus cinereus." figureDoi-0="http://doi.org/10.5281/zenodo.273634" figureDoi-1="http://doi.org/10.5281/zenodo.273635" figureDoi-2="http://doi.org/10.5281/zenodo.273636" httpUri-0="https://zenodo.org/record/273634/files/figure.png" httpUri-1="https://zenodo.org/record/273635/files/figure.png" httpUri-2="https://zenodo.org/record/273636/files/figure.png" pageId="1" pageNumber="34">
Figures
<number id="3DE22F2F5B1F335CFECC40BC448BF92F" box="[369,413,1689,1715]" pageId="1" pageNumber="34" value="2.0" valueMax="3.0" valueMin="1.0">1–3</number>
</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3306E835B1F335FFEB540FB46D3FE62" lastPageId="2" lastPageNumber="35" pageId="1" pageNumber="34" type="materials_examined">
<paragraph id="8B953D085B1F335FFEB540FB44FCFEAA" blockId="1.[264,1323,1758,1905]" lastBlockId="2.[264,1323,284,1870]" lastPageId="2" lastPageNumber="35" pageId="1" pageNumber="34">
<emphasis id="B95EE11A5B1F335CFEB540FB44ABF964" bold="true" box="[264,445,1758,1784]" pageId="1" pageNumber="34">
<typeStatus id="549183AA5B1F335CFEB540FB4455F964" box="[264,323,1758,1784]" pageId="1" pageNumber="34">Type</typeStatus>
material.
</emphasis>
<typeStatus id="549183AA5B1F335CFE7440FA472DF965" box="[457,571,1759,1785]" pageId="1" pageNumber="34">Holotype</typeStatus>
:
<collectionCode id="ED3BA5CD5B1F335CFDF540FA4793F965" box="[584,645,1759,1785]" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="1" pageNumber="34">MLP</collectionCode>
.R
<number id="3DE22F2F5B1F335CFD1840FA47F5F965" box="[677,739,1759,1785]" pageId="1" pageNumber="34" value="5139.0">5139</number>
, an adult male collected at the flood plain of the northern margin of the Blanco River, Quebrada Alcaparrosa (
<geoCoordinate id="EE1E5BCF5B1F335CFC45412241D6F8BD" box="[1016,1216,1799,1825]" direction="south" orientation="latitude" pageId="1" pageNumber="34" precision="1" value="-29.513195">
<number id="3DE22F2F5B1F335CFC4541224100F8BD" box="[1016,1046,1799,1825]" pageId="1" pageNumber="34" value="29.0">29</number>
º
<number id="3DE22F2F5B1F335CFB964122415BF8BD" box="[1067,1101,1799,1825]" pageId="1" pageNumber="34" value="30.0">30</number>
’
<number id="3DE22F2F5B1F335CFBE341224181F8BD" box="[1118,1175,1799,1825]" pageId="1" pageNumber="34" value="47.5">47.5</number>
’’ S
</geoCoordinate>
,
<geoCoordinate id="EE1E5BCF5B1F335CFB6F41224462F8D5" direction="west" orientation="longitude" pageId="1" pageNumber="34" precision="1" value="-69.17425">
<number id="3DE22F2F5B1F335CFB6F412241F9F8BD" box="[1234,1263,1799,1825]" pageId="1" pageNumber="34" value="69.0">69</number>
º
<number id="3DE22F2F5B1F335CFABE41224033F8BD" box="[1283,1317,1799,1825]" pageId="1" pageNumber="34" value="10.0">10</number>
’
<number id="3DE22F2F5B1F335CFEB5410A4457F8D5" box="[264,321,1839,1865]" pageId="1" pageNumber="34" value="27.3">27.3</number>
’’ W
</geoCoordinate>
),
<quantity id="4CD290ED5B1F335CFE36410A44F3F8D5" box="[395,485,1839,1865]" metricMagnitude="3" metricUnit="m" metricValue="2.27" pageId="1" pageNumber="34" unit="m" value="2270.0">2270 m</quantity>
above sea level, southern section of the San Guillermo National Park, Iglesia Department, San Juan province,
<collectingCountry id="F33D7D985B1F335CFD6141724642F8ED" box="[732,852,1879,1905]" name="Argentina" pageId="1" pageNumber="34">Argentina</collectingCountry>
,
<date id="FF941BC85B1F335CFCDD4172412FF8ED" box="[864,1081,1879,1905]" pageId="1" pageNumber="34" value="2005-02-11">February 11, 2005</date>
, by J. C. Acosta and J. H. Villavicencio.
</paragraph>
<paragraph id="8B953D085B1C335FFE85476146D3FE62" blockId="2.[264,1323,284,1870]" pageId="2" pageNumber="35">
<typeStatus id="549183AA5B1C335FFE85476144B8FEC2" box="[312,430,324,350]" pageId="2" pageNumber="35">Paratypes</typeStatus>
:
<collectionCode id="ED3BA5CD5B1C335FFE03476144EDFEC2" box="[446,507,324,350]" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="2" pageNumber="35">MLP</collectionCode>
.R
<number id="3DE22F2F5B1C335FFDA34761474AFEC2" box="[542,604,324,350]" pageId="2" pageNumber="35" value="5140.0">5140</number>
, an adult female, same data of collection as the
<typeStatus id="549183AA5B1C335FFB0147614030FEC2" box="[1212,1318,324,350]" pageId="2" pageNumber="35">holotype</typeStatus>
;
<collectionCode id="ED3BA5CD5B1C335FFEB547494476FE1A" box="[264,352,364,390]" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="2" pageNumber="35">MACN</collectionCode>
<number id="3DE22F2F5B1C335FFED7474944A5FE1A" box="[362,435,364,390]" pageId="2" pageNumber="35" value="38798.0">38798</number>
and
<number id="3DE22F2F5B1C335FFE4F47494728FE1A" box="[498,574,364,390]" pageId="2" pageNumber="35" value="38799.0">38799</number>
, two adult females collected
<quantity id="4CD290ED5B1C335FFC1D474946E3FE1A" box="[928,1013,364,390]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="35" unit="km" value="1.5">1.5 km</quantity>
S Quebrada Alcaparrosa, among this and the confluence of La Palca and Blanco rivers (
<geoCoordinate id="EE1E5BCF5B1C335FFC5247B14182FE32" box="[1007,1172,404,430]" direction="south" orientation="latitude" pageId="2" pageNumber="35" precision="15" value="-29.524445">
<number id="3DE22F2F5B1C335FFC5247B1411BFE32" box="[1007,1037,404,430]" pageId="2" pageNumber="35" value="29.0">29</number>
º
<number id="3DE22F2F5B1C335FFBA247B14157FE32" box="[1055,1089,404,430]" pageId="2" pageNumber="35" value="31.0">31</number>
’
<number id="3DE22F2F5B1C335FFBF247B14178FE32" box="[1103,1134,404,430]" pageId="2" pageNumber="35" value="28.0">28</number>
” S
</geoCoordinate>
,
<geoCoordinate id="EE1E5BCF5B1C335FFB1F47B14431FE4A" direction="west" orientation="longitude" pageId="2" pageNumber="35" precision="15" value="-69.185555">
<number id="3DE22F2F5B1C335FFB1F47B141A9FE32" box="[1186,1215,404,430]" pageId="2" pageNumber="35" value="69.0">69</number>
º
<number id="3DE22F2F5B1C335FFB6C47B141E7FE32" box="[1233,1265,404,430]" pageId="2" pageNumber="35" value="11.0">11</number>
’ 08” W
</geoCoordinate>
),
<quantity id="4CD290ED5B1C335FFE804799448EFE4A" box="[317,408,444,470]" metricMagnitude="3" metricUnit="m" metricValue="2.288" pageId="2" pageNumber="35" unit="m" value="2288.0">2288 m</quantity>
above sea level, San Guillermo National Park, Iglesia Department, San Juan province,
<collectingCountry id="F33D7D985B1C335FFEC047C144E3FE62" box="[381,501,484,510]" name="Argentina" pageId="2" pageNumber="35">Argentina</collectingCountry>
,
<date id="FF941BC85B1C335FFDBD47C147DAFE62" box="[512,716,484,510]" pageId="2" pageNumber="35" value="2005-01-24">January 24, 2005</date>
, by J. C. Monguillot.
</paragraph>
</subSubSection>
<subSubSection id="C3306E835B1C335FFE85442E4466F8D2" pageId="2" pageNumber="35" type="diagnosis">
<paragraph id="8B953D085B1C335FFE85442E4112FC42" blockId="2.[264,1323,284,1870]" pageId="2" pageNumber="35">
<emphasis id="B95EE11A5B1C335FFE85442E44A1FDB9" bold="true" box="[312,439,523,549]" pageId="2" pageNumber="35">Diagnosis.</emphasis>
<taxonomicName id="4C2A468B5B1C335FFE7F442947BEFDB9" box="[450,680,524,549]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="cinereus" status="sp. nov.">
<emphasis id="B95EE11A5B1C335FFE7F442947BEFDB9" box="[450,680,524,549]" italics="true" pageId="2" pageNumber="35">Liolaemus cinereus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D5C615B1C335FFD0E44294607FDBA" box="[691,785,524,550]" pageId="2" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
is a small iguanid lizard (
<quantity id="4CD290ED5B1C335FFBEF442941BFFDBA" box="[1106,1193,524,550]" metricMagnitude="-2" metricUnit="m" metricValue="6.3" pageId="2" pageNumber="35" unit="mm" value="63.0">63 mm</quantity>
maximum SVL in females), with a relatively long tail (
<number id="3DE22F2F5B1C335FFCB44411463EFDD2" box="[777,808,564,590]" pageId="2" pageNumber="35" value="57.0">57</number>
% of total length) and low number of scales around midbody (
<number id="3DE22F2F5B1C335FFE654479473EFDEA" box="[472,552,604,630]" pageId="2" pageNumber="35" value="60.0" valueMax="62.0" valueMin="58.0">58–62</number>
); dorsal scales moderately keeled; precloacal pores only in males; a bulged patch of enlarged scales on the proximal posterior surface of the thigh in both sexes, more evident in males; lacking of light vertebral or dorsolateral stripes; ash-grey dorsal background color with small transverse brown spots in two paravertebral series, and white scales roughly ordered on the trunk; antehumeral fold without black pigment (only faint greyish in the male, absent in females); scapular region pattern represented only by a prescapular dark spot dorsal to the antehumeral fold; throat and ventral neck brown in males; white, slightly dotted, in females. The new species is a member of the monophyletic
<taxonomicName id="4C2A468B5B1C335FFE0845B94722FC29" box="[437,564,924,949]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="B95EE11A5B1C335FFE0845B94722FC29" box="[437,564,924,949]" italics="true" pageId="2" pageNumber="35">boulengeri</emphasis>
</taxonomicName>
group of the genus
<taxonomicName id="4C2A468B5B1C335FFC8845B946B9FC29" box="[821,943,924,949]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="genus">
<emphasis id="B95EE11A5B1C335FFC8845B946B9FC29" box="[821,943,924,949]" italics="true" pageId="2" pageNumber="35">Liolaemus</emphasis>
</taxonomicName>
. Within this group
<taxonomicName id="4C2A468B5B1C335FFB1845B9403CFC29" box="[1189,1322,924,949]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="cinereus">
<emphasis id="B95EE11A5B1C335FFB1845B9403CFC29" box="[1189,1322,924,949]" italics="true" pageId="2" pageNumber="35">L. cinereus</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="4C2A468B5B1C335FFE0945E14701FC41" box="[436,535,964,989]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="darwinii">
<emphasis id="B95EE11A5B1C335FFE0945E14701FC41" box="[436,535,964,989]" italics="true" pageId="2" pageNumber="35">darwinii</emphasis>
</taxonomicName>
complex, as defined by
<bibRefCitation id="EFBB40F95B1C335FFC8B45E14116FC42" author="Etheridge" box="[822,1024,964,990]" pageId="2" pageNumber="43" refString="Etheridge, R. (1993) Lizards of the Liolaemus darwinii complex (Squamata: Iguania: Tropiduridae) in northern Argentina. Bolletino del Museo regionale di Scienze naturali Torino, 11 (1), 137 - 199." type="journal article" year="1993">
Etheridge (
<number id="3DE22F2F5B1C335FFC0A45E146E1FC42" box="[951,1015,964,990]" pageId="2" pageNumber="35" value="1993.0">1993</number>
)
</bibRefCitation>
.
</paragraph>
<paragraph id="8B953D085B1C335FFE8545C94466F8D2" blockId="2.[264,1323,284,1870]" pageId="2" pageNumber="35">
<emphasis id="B95EE11A5B1C335FFE8545C947C0FB9A" bold="true" box="[312,726,1004,1030]" pageId="2" pageNumber="35">Comparison with related species.</emphasis>
<taxonomicName id="4C2A468B5B1C335FFD6345C946D4FB99" box="[734,962,1004,1029]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="cinereus">
<emphasis id="B95EE11A5B1C335FFD6345C946D4FB99" box="[734,962,1004,1029]" italics="true" pageId="2" pageNumber="35">Liolaemus cinereus</emphasis>
</taxonomicName>
differs from any other species of the
<taxonomicName id="4C2A468B5B1C335FFEEA423144C0FBB1" box="[343,470,1044,1069]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="B95EE11A5B1C335FFEEA423144C0FBB1" box="[343,470,1044,1069]" italics="true" pageId="2" pageNumber="35">boulengeri</emphasis>
</taxonomicName>
group, except for those in the
<taxonomicName id="4C2A468B5B1C335FFCF6423146B8FBB1" box="[843,942,1044,1069]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="darwinii">
<emphasis id="B95EE11A5B1C335FFCF6423146B8FBB1" box="[843,942,1044,1069]" italics="true" pageId="2" pageNumber="35">darwinii</emphasis>
</taxonomicName>
complex, in having moderately cusped, straight-sided to slightly expanded crowns in their posterior marginal teeth. Within the
<taxonomicName id="4C2A468B5B1C335FFEF4424144BAFBE1" box="[329,428,1124,1149]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="darwinii">
<emphasis id="B95EE11A5B1C335FFEF4424144BAFBE1" box="[329,428,1124,1149]" italics="true" pageId="2" pageNumber="35">darwinii</emphasis>
</taxonomicName>
complex, the new species differs from
<taxonomicName id="4C2A468B5B1C335FFC1342414121FBE1" box="[942,1079,1124,1149]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="darwinii">
<emphasis id="B95EE11A5B1C335FFC1342414121FBE1" box="[942,1079,1124,1149]" italics="true" pageId="2" pageNumber="35">L. darwinii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A468B5B1C335FFBF0424141EBFBE1" box="[1101,1277,1124,1149]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="grosseorum">
<emphasis id="B95EE11A5B1C335FFBF0424141EBFBE1" box="[1101,1277,1124,1149]" italics="true" pageId="2" pageNumber="35">L. grosseorum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A468B5B1C335FFAAE42414472FB39" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="laurenti">
<emphasis id="B95EE11A5B1C335FFAAE42414472FB39" italics="true" pageId="2" pageNumber="35">L. laurenti</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A468B5B1C335FFEC542A94702FB39" box="[376,532,1164,1189]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="montanezi">
<emphasis id="B95EE11A5B1C335FFEC542A94702FB39" box="[376,532,1164,1189]" italics="true" pageId="2" pageNumber="35">L. montanezi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2A468B5B1C335FFDEA42A947F8FB39" box="[599,750,1164,1189]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="olongasta">
<emphasis id="B95EE11A5B1C335FFDEA42A947F8FB39" box="[599,750,1164,1189]" italics="true" pageId="2" pageNumber="35">L. olongasta</emphasis>
</taxonomicName>
in lacking intensely black pigment within the antehumeral fold, and by its different dorsal pattern. It differs from
<taxonomicName id="4C2A468B5B1C335FFBF7429141C4FB51" box="[1098,1234,1204,1229]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="abaucan">
<emphasis id="B95EE11A5B1C335FFBF7429141C4FB51" box="[1098,1234,1204,1229]" italics="true" pageId="2" pageNumber="35">L. abaucan</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2A468B5B1C335FFAAE42914491FB69" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="koslowskyi">
<emphasis id="B95EE11A5B1C335FFAAE42914491FB69" italics="true" pageId="2" pageNumber="35">L. koslowskyi</emphasis>
</taxonomicName>
in lacking large postscapular spot, and from
<taxonomicName id="4C2A468B5B1C335FFC2842F94100FB69" box="[917,1046,1244,1269]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="albiceps">
<emphasis id="B95EE11A5B1C335FFC2842F94100FB69" box="[917,1046,1244,1269]" italics="true" pageId="2" pageNumber="35">L. albiceps</emphasis>
</taxonomicName>
by its smaller body (up to
<quantity id="4CD290ED5B1C335FFE984321446EFA82" box="[293,376,1284,1310]" metricMagnitude="-2" metricUnit="m" metricValue="9.4" pageId="2" pageNumber="35" unit="mm" value="94.0">94 mm</quantity>
maximum SVL in males of the latter).
<taxonomicName id="4C2A468B5B1C335FFCFD43214135FA81" box="[832,1059,1284,1309]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="cinereus">
<emphasis id="B95EE11A5B1C335FFCFD43214135FA81" box="[832,1059,1284,1309]" italics="true" pageId="2" pageNumber="35">Liolaemus cinereus</emphasis>
</taxonomicName>
further differs from
<taxonomicName id="4C2A468B5B1C335FFAAE4321447CFAD9" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="albiceps">
<emphasis id="B95EE11A5B1C335FFAAE4321447CFAD9" italics="true" pageId="2" pageNumber="35">L. albiceps</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A468B5B1C335FFEC6430944E9FAD9" box="[379,511,1324,1349]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="darwinii">
<emphasis id="B95EE11A5B1C335FFEC6430944E9FAD9" box="[379,511,1324,1349]" italics="true" pageId="2" pageNumber="35">L. darwinii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A468B5B1C335FFDAC430947A7FAD9" box="[529,689,1324,1349]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="irregularis">
<emphasis id="B95EE11A5B1C335FFDAC430947A7FAD9" box="[529,689,1324,1349]" italics="true" pageId="2" pageNumber="35">L. irregularis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A468B5B1C335FFD7F43094641FAD9" box="[706,855,1324,1349]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="olongasta">
<emphasis id="B95EE11A5B1C335FFD7F43094641FAD9" box="[706,855,1324,1349]" italics="true" pageId="2" pageNumber="35">L. olongasta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2A468B5B1C335FFC2843094106FAD9" box="[917,1040,1324,1349]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="ornatus">
<emphasis id="B95EE11A5B1C335FFC2843094106FAD9" box="[917,1040,1324,1349]" italics="true" pageId="2" pageNumber="35">L. ornatus</emphasis>
</taxonomicName>
(unknown condition in female
<taxonomicName id="4C2A468B5B1C335FFEDC4371471DFAF1" box="[353,523,1364,1389]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="grosseorum">
<emphasis id="B95EE11A5B1C335FFEDC4371471DFAF1" box="[353,523,1364,1389]" italics="true" pageId="2" pageNumber="35">L. grosseorum</emphasis>
</taxonomicName>
) by the absence of precloacal pores in females of the new species. The new species differs from
<taxonomicName id="4C2A468B5B1C335FFDD043594614FA09" box="[621,770,1404,1429]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="calchaqui">
<emphasis id="B95EE11A5B1C335FFDD043594614FA09" box="[621,770,1404,1429]" italics="true" pageId="2" pageNumber="35">L. calchaqui</emphasis>
</taxonomicName>
by the absence of dorsolateral whitish stripes, invariably present in males of the latter, and by having more middorsal scales, both in male and females (
<number id="3DE22F2F5B1C335FFE1A43E944D3FA7A" box="[423,453,1484,1510]" pageId="2" pageNumber="35" value="68.0">68</number>
vs.
<number id="3DE22F2F5B1C335FFE4443E9470FFA7A" box="[505,537,1484,1510]" pageId="2" pageNumber="35" value="63.0">63</number>
, and
<number id="3DE22F2F5B1C335FFDE743E947B5FA7A" box="[602,675,1484,1510]" pageId="2" pageNumber="35" value="69.0" valueMax="71.0" valueMin="67.0">67–71</number>
vs.
<number id="3DE22F2F5B1C335FFD6B43E94634FA7A" box="[726,802,1484,1510]" pageId="2" pageNumber="35" value="61.0" valueMax="64.0" valueMin="58.0">58–64</number>
, respectively). It differs from
<taxonomicName id="4C2A468B5B1C335FFB3443E94012FA79" box="[1161,1284,1484,1509]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="quilmes">
<emphasis id="B95EE11A5B1C335FFB3443E94012FA79" box="[1161,1284,1484,1509]" italics="true" pageId="2" pageNumber="35">L. quilmes</emphasis>
</taxonomicName>
by the absence of bold, symmetrical dorsal pattern in adult males. The number of midbody scales (
<number id="3DE22F2F5B1C335FFEE3403944BCF9AA" box="[350,426,1564,1590]" pageId="2" pageNumber="35" value="60.0" valueMax="62.0" valueMin="58.0">58–62</number>
) in
<taxonomicName id="4C2A468B5B1C335FFE674039474AF9A9" box="[474,604,1564,1589]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="cinereus">
<emphasis id="B95EE11A5B1C335FFE674039474AF9A9" box="[474,604,1564,1589]" italics="true" pageId="2" pageNumber="35">L. cinereus</emphasis>
</taxonomicName>
is higher than in
<taxonomicName id="4C2A468B5B1C335FFC91403946DBF9A9" box="[812,973,1564,1589]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="chacoensis">
<emphasis id="B95EE11A5B1C335FFC91403946DBF9A9" box="[812,973,1564,1589]" italics="true" pageId="2" pageNumber="35">L. chacoensis</emphasis>
</taxonomicName>
(
<number id="3DE22F2F5B1C335FFC604039413AF9AA" box="[989,1068,1564,1590]" pageId="2" pageNumber="35" value="45.0" valueMax="48.0" valueMin="42.0">42–48</number>
), but lower than in
<taxonomicName id="4C2A468B5B1C335FFAAE40394493F9C1" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="irregularis">
<emphasis id="B95EE11A5B1C335FFAAE40394493F9C1" italics="true" pageId="2" pageNumber="35">L. irregularis</emphasis>
</taxonomicName>
(
<number id="3DE22F2F5B1C335FFE2A406144F5F9C2" box="[407,483,1604,1630]" pageId="2" pageNumber="35" value="67.0" valueMax="74.0" valueMin="60.0">60–74</number>
) and
<taxonomicName id="4C2A468B5B1C335FFD95406147F7F9C1" box="[552,737,1604,1629]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="uspallatensis">
<emphasis id="B95EE11A5B1C335FFD95406147F7F9C1" box="[552,737,1604,1629]" italics="true" pageId="2" pageNumber="35">L. uspallatensis</emphasis>
</taxonomicName>
(
<number id="3DE22F2F5B1C335FFD4F40614657F9C2" box="[754,833,1604,1630]" pageId="2" pageNumber="35" value="68.0" valueMax="76.0" valueMin="60.0">60–76</number>
). From the latter, the new species further differs by having keeled dorsal scales (smooth or faintly keeled in
<taxonomicName id="4C2A468B5B1C335FFB97404941F2F919" box="[1066,1252,1644,1669]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="uspallatensis">
<emphasis id="B95EE11A5B1C335FFB97404941F2F919" box="[1066,1252,1644,1669]" italics="true" pageId="2" pageNumber="35">L. uspallatensis</emphasis>
</taxonomicName>
), less middorsal scales (
<number id="3DE22F2F5B1C335FFE6640B14730F932" box="[475,550,1684,1710]" pageId="2" pageNumber="35" value="69.0" valueMax="71.0" valueMin="67.0">67–71</number>
<emphasis id="B95EE11A5B1C335FFD9040B1476EF931" box="[557,632,1684,1709]" italics="true" pageId="2" pageNumber="35">versus</emphasis>
<number id="3DE22F2F5B1C335FFD3D40B147DBF932" box="[640,717,1684,1710]" pageId="2" pageNumber="35" value="85.5" valueMax="94.0" valueMin="77.0">77–94</number>
), and underside of the tail inmaculate (spotted in
<taxonomicName id="4C2A468B5B1C335FFAAE40B144B7F949" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="uspallatensis">
<emphasis id="B95EE11A5B1C335FFAAE40B144B7F949" italics="true" pageId="2" pageNumber="35">L. uspallatensis</emphasis>
</taxonomicName>
).
<taxonomicName id="4C2A468B5B1C335FFE7F409947BBF949" box="[450,685,1724,1749]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="cinereus">
<emphasis id="B95EE11A5B1C335FFE7F409947BBF949" box="[450,685,1724,1749]" italics="true" pageId="2" pageNumber="35">Liolaemus cinereus</emphasis>
</taxonomicName>
further differs from
<taxonomicName id="4C2A468B5B1C335FFC0240994149F949" box="[959,1119,1724,1749]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="montanezi">
<emphasis id="B95EE11A5B1C335FFC0240994149F949" box="[959,1119,1724,1749]" italics="true" pageId="2" pageNumber="35">L. montanezi</emphasis>
</taxonomicName>
(the only other species of the genus syntopic to it) in having different dorsal pattern and coloration, shorter tail, keel less marked on the dorsal scales, and lacking of antehumeral black pigment.
</paragraph>
</subSubSection>
<subSubSection id="C3306E835B1D335AFE85473E4017FB9A" lastPageId="7" lastPageNumber="40" pageId="3" pageNumber="36" type="description">
<paragraph id="8B953D085B1D335EFE85473E416AF962" blockId="3.[264,1323,283,1870]" pageId="3" pageNumber="36">
<emphasis id="B95EE11A5B1D335EFE85473E478EFEA9" bold="true" box="[312,664,283,309]" pageId="3" pageNumber="36">
Description of the
<typeStatus id="549183AA5B1D335EFD98473E4785FEA9" box="[549,659,283,309]" pageId="3" pageNumber="36">holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED3BA5CD5B1D335EFD1C473947C8FEAA" box="[673,734,284,310]" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="3" pageNumber="36">MLP</collectionCode>
.R
<number id="3DE22F2F5B1D335EFD424739462BFEAA" box="[767,829,284,310]" pageId="3" pageNumber="36" value="5139.0">5139</number>
, adult male (
<figureCitation id="1311218D5B1D335EFC644739417CFEAA" box="[985,1130,284,310]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="4.[264,367,1454,1478]" captionStartId-1="5.[264,367,1335,1359]" captionStartId-2="6.[264,367,1380,1404]" captionTargetBox-0="[267,1319,794,1430]" captionTargetBox-1="[267,1319,630,1327]" captionTargetBox-2="[268,1318,310,1372]" captionTargetId-0="figure@4.[267,1319,794,1430]" captionTargetId-1="figure@5.[267,1319,628,1329]" captionTargetId-2="figure@6.[268,1319,308,1374]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 1. The holotype of Liolaemus cinereus, adult male, in life. The specimen is actually grey (see Figure 2); the brownish shade is an artifact due to lighting used to portray it." captionText-1="FIGURE 2. The holotype of Liolaemus cinereus in life, showing the grey lining into the prehumeral fold (here extended) and the dark prescapular spot." captionText-2="FIGURE 3. Comparative ventral view in life of the female paratype MLP. R 5140 (left) and the male holotype (right) of Liolaemus cinereus." figureDoi-0="http://doi.org/10.5281/zenodo.273634" figureDoi-1="http://doi.org/10.5281/zenodo.273635" figureDoi-2="http://doi.org/10.5281/zenodo.273636" httpUri-0="https://zenodo.org/record/273634/files/figure.png" httpUri-1="https://zenodo.org/record/273635/files/figure.png" httpUri-2="https://zenodo.org/record/273636/files/figure.png" pageId="3" pageNumber="36">
Figures
<number id="3DE22F2F5B1D335EFB804739417CFEAA" box="[1085,1130,284,310]" pageId="3" pageNumber="36" value="2.0" valueMax="3.0" valueMin="1.0">1–3</number>
</figureCitation>
),
<quantity id="4CD290ED5B1D335EFB3E473941CEFEAA" box="[1155,1240,284,310]" metricMagnitude="-2" metricUnit="m" metricValue="5.9" pageId="3" pageNumber="36" unit="mm" value="59.0">59 mm</quantity>
snoutvent length; regenerated tail
<quantity id="4CD290ED5B1D335EFDDD476147A2FEC2" box="[608,692,324,350]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" pageId="3" pageNumber="36" unit="mm" value="77.0">77 mm</quantity>
long. Some scales on head, body and legs partial or completely black, irregularly scared and with texture change (evidence of disease?). Head
<number id="3DE22F2F5B1D335EFEB547B1442DFE32" box="[264,315,404,430]" pageId="3" pageNumber="36" value="1.28">1.28</number>
times longer than wide. Dorsal head scales smooth. Rostral
<number id="3DE22F2F5B1D335EFB9F47B14143FE32" box="[1058,1109,404,430]" pageId="3" pageNumber="36" value="2.75">2.75</number>
times wider than high, wider than mental, bordered behind by four postrostrals. Nasals anterodorsal in position, nearer to tip of snout than to eye; nostrils circular. Nasals medially separated by five irregular internasals and from the rostral by one postrostral each. Two postnasals on each side. Eight polygonal frontonasals. Two irregular prefrontals. All postrostrals, internasals, postnasals, prefrontals and frontonasals with several scattered brown scale organs. Two fused frontal azygous followed behind by three small frontoparietals. Interparietal pentagonal, with a conspicuous pineal eye. Six parietals; the four posterior to interparietal larger than the interparietal. Occipital scales polygonal and irregular, swollen, slightly rugose, subjuxtaposed; each one with
<number id="3DE22F2F5B1D335EFC8E44D94676FC8A" box="[819,864,764,790]" pageId="3" pageNumber="36" value="2.0" valueMax="3.0" valueMin="1.0">1–3</number>
scale organs. Supraorbital semicircles with
<number id="3DE22F2F5B1D335EFEFE45014476FCA2" box="[323,352,804,830]" pageId="3" pageNumber="36" value="12.0">12</number>
/
<number id="3DE22F2F5B1D335EFED745014490FCA2" box="[362,390,804,830]" pageId="3" pageNumber="36" value="13.0">13</number>
polygonal scales. Ten supraoculars on each side, those medialmost transversely expanded, with
<number id="3DE22F2F5B1D335EFE78456944E4FCFA" box="[453,498,844,870]" pageId="3" pageNumber="36" value="2.0" valueMax="3.0" valueMin="1.0">1–3</number>
scale organs each. Supraoculars separated from superciliaries by one to three rows of small polygonal scales. Two canthals on each side, the anterior transversely expanded, the posterior overlapping the first superciliar. The canthus rostralis includes the postnasal scale that contacts to lorilabials. Five elongate, obliquely overlapping superciliaries, followed by two shorter scales overlapping to the fifth in the opposite direction. A single, short preocular followed by an elongate, concave and curved subocular several times longer than high, overlapped near its posterior end by a thin postocular, the three scales keeled all along. Palpebrals small, granular, juxtaposed. Ciliaries quadrangulars, projecting from the eyelid as a short “eyelash”. Loreals
<number id="3DE22F2F5B1D335EFB1242A941A8FB3A" box="[1199,1214,1164,1190]" pageId="3" pageNumber="36" value="6.0">6</number>
/
<number id="3DE22F2F5B1D335EFB7A42A941C1FB3A" box="[1223,1239,1164,1190]" pageId="3" pageNumber="36" value="4.0">4</number>
, in one or two rows, with several scale organs on each. Lorilabials
<number id="3DE22F2F5B1D335EFC75429146C1FB52" box="[968,983,1204,1230]" pageId="3" pageNumber="36" value="6.0">6</number>
/
<number id="3DE22F2F5B1D335EFC5C429146E6FB52" box="[993,1008,1204,1230]" pageId="3" pageNumber="36" value="7.0">7</number>
, longer than high, the first three on each side higher than supralabials, the fourth subequal, and the remaining lower; all with several scattered scale organs. Seven supralabials, the fourth and fifth the longest, all longer than high. Temporals smooth, polygonal, convex, subimbricate; most with one, more rarely two, scale organs near their posterior or posteroventral border. Aperture of external auditory meatus suboval, higher than long, bordered anterior and posteriorly by small higher-than-long swollen scales, bearing one scale organ each; meatus bordered below by small ovoidal scales longer than high. Mental subpentagonal with concave lateral sides,
<number id="3DE22F2F5B1D335EFE1843D144DCF992" box="[421,458,1524,1550]" pageId="3" pageNumber="36" value="1.6">1.6</number>
times wider than long, bordered laterally by the first infralabial on each side and posteriorly by two large postmentals. Six infralabials on each side, longer than high. Six pair of enlarged postmentals, only the first in medial contact, all others aparted from by convex, imbricated gulars. Only the first pair of postmental in slight contact with the first infralabial, second pair separated from infralabials by one row of sublabials, the third and fourth pair by two rows, and the fifth and sixth by three rows of scales. Posterior gulars flat, imbricated, the posteriormost scales notched on the free margin.
</paragraph>
<paragraph id="8B953D085B1D3359FE854129464BFD72" blockId="3.[264,1323,283,1870]" lastBlockId="4.[264,1322,284,750]" lastPageId="4" lastPageNumber="37" pageId="3" pageNumber="36">
Body slender. A skin fold out extended along both flanks, between fore and hindlimbs. Behind the tympanum a “
<collectingCountry id="F33D7D985B1D335EFDF94111474AF8D2" box="[580,604,1844,1870]" name="Italy" pageId="3" pageNumber="36">Y</collectingCountry>
” fold; one branch forward reaches posterodorsally the ear, other extends anteroventrally to the inferior border of the ear, and a third, the longest, runs longitudinally on the neck to fuse with the antehumeral fold. Scales of lateral neck region granular, juxtaposed, most of them with one scale pit each. Antehumeral fold distinct, lined by tiny granular scales. Dorsal scales on neck imbricated, smooth, a few blunt keeled. Scales on dorsolateral neck and body subimbricate, with one scale pit. Dorsal scales on body rhomboidal, imbricate, becoming subimbricate to dorsolateral; blunt keeled. Scales around midbody
<number id="3DE22F2F5B1A3359FD1F442947D5FDBA" box="[674,707,524,550]" pageId="4" pageNumber="37" value="62.0">62</number>
; middorsal scales
<number id="3DE22F2F5B1A3359FC09442946C3FDBA" box="[948,981,524,550]" pageId="4" pageNumber="37" value="68.0">68</number>
. Dorsal body scales grade gradually to the sides into smaller scales juxtaposed, some faintly keeled but mostly smooth. Granular scales dorsally and posteriorly to forelimb insertion. Ventral body scales smooth, rhomboidal, larger than dorsals, imbricate. Ventral scales between postmentals and vent
<number id="3DE22F2F5B1A3359FEC54489448FFD5A" box="[376,409,684,710]" pageId="4" pageNumber="37" value="89.0">89</number>
. Scales on precloacal region, between hindlimb insertions and vent, slightly larger than ventrals. Eight orange precloacal pores.
</paragraph>
<caption id="DF556D805B1A3359FEB5438B417EFA79" ID-DOI="http://doi.org/10.5281/zenodo.273634" httpUri="https://zenodo.org/record/273634/files/figure.png" pageId="4" pageNumber="37" targetBox="[267,1319,794,1430]" targetPageId="4">
<paragraph id="8B953D085B1A3359FEB5438B417EFA79" blockId="4.[264,1322,1454,1509]" pageId="4" pageNumber="37">
<emphasis id="B95EE11A5B1A3359FEB5438B449CFA5A" bold="true" box="[264,394,1454,1478]" pageId="4" pageNumber="37">FIGURE 1.</emphasis>
The holotype of
<taxonomicName id="4C2A468B5B1A3359FDF9438B4605FA59" box="[580,787,1454,1477]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="cinereus">
<emphasis id="B95EE11A5B1A3359FDF9438B4605FA59" box="[580,787,1454,1477]" italics="true" pageId="4" pageNumber="37">
Liolaemus
<taxonomicName id="4C2A468B5B1A3359FD07438B4605FA59" box="[698,787,1454,1477]" pageId="4" pageNumber="37" rank="species" species="cinereus">cinereus</taxonomicName>
</emphasis>
</taxonomicName>
, adult male, in life. The specimen is actually grey (see Figure 2); the brownish shade is an artifact due to lighting used to portray it.
</paragraph>
</caption>
<paragraph id="8B953D085B1A3358FE8540334797FE32" blockId="4.[264,1323,1558,1904]" lastBlockId="5.[264,1323,284,590]" lastPageId="5" lastPageNumber="38" pageId="4" pageNumber="37">
Dorsal scales on forelimbs faintly keeled, imbricate. Infrabrachial scales small, conical and juxtaposed. Infra-antibrachial scales larger, smooth, imbricate; progressively notched to become tridentate on wrist. Dorsal scales on manus large, rounded and notched, smooth, imbricate. Supradigitals smooth, wider than long. Palmar scales strongly keeled, mucronate, imbricate. Tricarinate, mucronate lamellae under digits;
<number id="3DE22F2F5B1A3359FB814093414FF94C" box="[1084,1113,1718,1744]" pageId="4" pageNumber="37" value="22.0">22</number>
under fourth left finger. Dorsal thigh scales faintly keeled and imbricate, become gradually smaller, smooth and juxtaposed to posterior. Patch distinct, formed by
<number id="3DE22F2F5B1A3359FC7E4123411AF8BC" box="[963,1036,1798,1824]" pageId="4" pageNumber="37" value="39.0" valueMax="40.0" valueMin="38.0">38–40</number>
scales larger than the neighbours, faintly keeled but erect and mucronate, subjuxtaposed. Thigh underside and infra-tibiofibular scales large, rounded and smooth, imbricate. Scales on dorsum of pes large, imbricate, smooth. Supradigitals smooth, longer than wide. Scales on sole of pes keeled, mucronate, imbricate. Tricarinate, slightly mucronate lamellae under digits;
<number id="3DE22F2F5B1B3358FAB04761403CFEC2" box="[1293,1322,324,350]" pageId="5" pageNumber="38" value="30.0">30</number>
under fourth left toe. Manus and pes with sharp, slender claws, equal in length to the span of
<number id="3DE22F2F5B1B3358FE9547B14442FE32" box="[296,340,404,430]" pageId="5" pageNumber="38" value="4.5" valueMax="5.0" valueMin="4.0">4–5</number>
distal supradigital scales.
</paragraph>
<paragraph id="8B953D085B1B3358FE854799472FFDD2" blockId="5.[264,1323,284,590]" pageId="5" pageNumber="38">Tail slender, subcircular in section. Caudal scales in discernable annuli throughout all its length, less evident only on suprahemipenial portion. Dorsal and lateral caudal scales keeled. Ventral caudal scales smooth and imbricate proximally, gradually keeled and longer than wide to distal.</paragraph>
<caption id="DF556D805B1B3358FEB5431246B0FAF2" ID-DOI="http://doi.org/10.5281/zenodo.273635" httpUri="https://zenodo.org/record/273635/files/figure.png" pageId="5" pageNumber="38" targetBox="[267,1319,630,1327]" targetPageId="5">
<paragraph id="8B953D085B1B3358FEB5431246B0FAF2" blockId="5.[264,1323,1335,1390]" pageId="5" pageNumber="38">
<emphasis id="B95EE11A5B1B3358FEB543124484FAD3" bold="true" box="[264,402,1335,1359]" pageId="5" pageNumber="38">FIGURE 2.</emphasis>
The holotype of
<taxonomicName id="4C2A468B5B1B3358FDD643124655FAD2" box="[619,835,1335,1358]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="38" phylum="Chordata" rank="species" species="cinereus">
<emphasis id="B95EE11A5B1B3358FDD643124655FAD2" box="[619,835,1335,1358]" italics="true" pageId="5" pageNumber="38">
Liolaemus
<taxonomicName id="4C2A468B5B1B3358FD5443124655FAD2" box="[745,835,1335,1358]" pageId="5" pageNumber="38" rank="species" species="cinereus">cinereus</taxonomicName>
</emphasis>
</taxonomicName>
in life, showing the grey lining into the prehumeral fold (here extended) and the dark prescapular spot.
</paragraph>
</caption>
<paragraph id="8B953D085B1B3358FE8543BA4168F8ED" blockId="5.[264,1323,1439,1905]" pageId="5" pageNumber="38">
<emphasis id="B95EE11A5B1B3358FE8543BA44D0FA25" bold="true" box="[312,454,1439,1465]" pageId="5" pageNumber="38">Coloration.</emphasis>
In life, ground color of head, body and limbs ash-grey. Dorsal surfaces of body, limbs, and tail base with abundant white scales, single or in groups of
<number id="3DE22F2F5B1B3358FB1A43E241C5FA7D" box="[1191,1235,1479,1505]" pageId="5" pageNumber="38" value="3.0" valueMax="4.0" valueMin="2.0">2–4</number>
scales, forming rows more-or-less ordered. Grey nearly uniform in preservative. Brown paravertebral transverse bars, thin and broken, little evident. Middorsal and dorsolateral stripes absent as such or as lighter zones. Lateral surfaces of head with subsymmetrical pattern of dark brown lines, the foremost one runs longitudinally from the canthals to the anterior angle of eye; a second bar vertically crosses the middle of the eye from the superciliaries to the upper border of mouth; a third line runs longitudinally more-or-less continued through the temporals and neck from behind the eye, nearly reaching the prescapular spot. Supraciliaries, infraoculars, lorilabials and supralabials, whitish. Mental, infralabials, sublabials and postmentals whitish with occasional darker borders; definitely white in preservative. Extense dark brown pigment on the throat (
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Figure
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).
</paragraph>
<caption id="DF556D805B18335BFEB5434147F2FA07" ID-DOI="http://doi.org/10.5281/zenodo.273636" httpUri="https://zenodo.org/record/273636/files/figure.png" pageId="6" pageNumber="39" targetBox="[268,1318,310,1372]" targetPageId="6">
<paragraph id="8B953D085B18335BFEB5434147F2FA07" blockId="6.[264,1323,1380,1435]" pageId="6" pageNumber="39">
<emphasis id="B95EE11A5B18335BFEB54341449AFAE0" bold="true" box="[264,396,1380,1404]" pageId="6" pageNumber="39">FIGURE 3.</emphasis>
Comparative ventral view in life of the female paratype MLP. R 5140 (left) and the male holotype (right) of
<taxonomicName id="4C2A468B5B18335BFDB343A147CBFA07" box="[526,733,1412,1435]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="39" phylum="Chordata" rank="species" species="cinereus">
<emphasis id="B95EE11A5B18335BFDB343A147CBFA07" box="[526,733,1412,1435]" italics="true" pageId="6" pageNumber="39">Liolaemus cinereus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B953D085B18335BFE8543E94626F962" blockId="6.[264,1323,1484,1870]" pageId="6" pageNumber="39">
“Pocket” formed by antehumeral fold lined in grey. Absence of bars related to the forelimb insertion as well as postscapular spots. The only evident spot is a rounded dark macula just dorsal to the antehumeral fold (
<figureCitation id="1311218D5B18335BFCBC40394671F9AA" box="[769,871,1564,1590]" captionStart="FIGURE 2" captionStartId="5.[264,367,1335,1359]" captionTargetBox="[267,1319,630,1327]" captionTargetId="figure@5.[267,1319,628,1329]" captionTargetPageId="5" captionText="FIGURE 2. The holotype of Liolaemus cinereus in life, showing the grey lining into the prehumeral fold (here extended) and the dark prescapular spot." figureDoi="http://doi.org/10.5281/zenodo.273635" httpUri="https://zenodo.org/record/273635/files/figure.png" pageId="6" pageNumber="39">
Figure
<number id="3DE22F2F5B18335BFCEB40394671F9AA" box="[854,871,1564,1590]" pageId="6" pageNumber="39" value="2.0">2</number>
</figureCitation>
), constant also in females (see below). On body flanks the ground color extends to the longitudinal fold, gradually fading to white from there. Irregular ink-black areas on head and body, probably reflecting an epithelial disease, obscure the pattern in this specimen.Ventral surface of throat, neck and anterior chest dark brown; gradually fading to immaculate white in venter, underside of forelimbs, hindlimbs and tail. Femoral patch scales white.
</paragraph>
<paragraph id="8B953D085B18335AFE8541294712FD02" blockId="6.[264,1323,1484,1870]" lastBlockId="7.[264,1323,284,1870]" lastPageId="7" lastPageNumber="40" pageId="6" pageNumber="39">
<emphasis id="B95EE11A5B18335BFE85412944A7F8BA" bold="true" box="[312,433,1804,1830]" pageId="6" pageNumber="39">Variation.</emphasis>
Based on the three female
<typeStatus id="549183AA5B18335BFCBF41294663F8BA" box="[770,885,1804,1830]" pageId="6" pageNumber="39">paratypes</typeStatus>
, the only other known specimens of
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<emphasis id="B95EE11A5B18335BFEB5411144FDF8D1" box="[264,491,1844,1869]" italics="true" pageId="6" pageNumber="39">Liolaemus cinereus</emphasis>
</taxonomicName>
. Snout-vent length
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(
<collectionCode id="ED3BA5CD5B18335BFC8D41114678F8D2" box="[816,878,1844,1870]" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="6" pageNumber="39">MLP</collectionCode>
.R
<number id="3DE22F2F5B18335BFC31411146DDF8D2" box="[908,971,1844,1870]" pageId="6" pageNumber="39" value="5140.0">5140</number>
),
<quantity id="4CD290ED5B18335BFC6341114126F8D2" box="[990,1072,1844,1870]" metricMagnitude="-2" metricUnit="m" metricValue="6.3" pageId="6" pageNumber="39" unit="mm" value="63.0">63 mm</quantity>
(
<collectionCode id="ED3BA5CD5B18335BFB834111418EF8D2" box="[1086,1176,1844,1870]" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="6" pageNumber="39">MACN</collectionCode>
<number id="3DE22F2F5B18335BFB22411141FAF8D2" box="[1183,1260,1844,1870]" pageId="6" pageNumber="39" value="38798.0">38798</number>
), and
<quantity id="4CD290ED5B19335AFEB54739444AFEAA" box="[264,348,284,310]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" pageId="7" pageNumber="40" unit="mm" value="62.0">62 mm</quantity>
(
<collectionCode id="ED3BA5CD5B19335AFED0473944D1FEAA" box="[365,455,284,310]" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="7" pageNumber="40">MACN</collectionCode>
<number id="3DE22F2F5B19335AFE6D47394708FEAA" box="[464,542,284,310]" pageId="7" pageNumber="40" value="38799.0">38799</number>
); tail length (complete only in the
<collectionCode id="ED3BA5CD5B19335AFC0047394103FEAA" box="[957,1045,284,310]" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="7" pageNumber="40">MACN</collectionCode>
<number id="3DE22F2F5B19335AFBA24739417CFEAA" box="[1055,1130,284,310]" pageId="7" pageNumber="40" value="38799.0">38799</number>
)
<quantity id="4CD290ED5B19335AFBC6473941C4FEAA" box="[1147,1234,284,310]" metricMagnitude="-2" metricUnit="m" metricValue="8.3" pageId="7" pageNumber="40" unit="mm" value="83.0">83 mm</quantity>
. Scales around midbody
<number id="3DE22F2F5B19335AFE6D4761470FFEC2" box="[464,537,324,350]" pageId="7" pageNumber="40" value="59.5" valueMax="61.0" valueMin="58.0">58–61</number>
(mean
<number id="3DE22F2F5B19335AFDCD476147B2FEC2" box="[624,676,324,350]" pageId="7" pageNumber="40" value="59.3">59.3</number>
±
<number id="3DE22F2F5B19335AFD7F476147E3FEC2" box="[706,757,324,350]" pageId="7" pageNumber="40" value="1.53">1.53</number>
SD); middorsal scales
<number id="3DE22F2F5B19335AFBB847614158FEC2" box="[1029,1102,324,350]" pageId="7" pageNumber="40" value="69.0" valueMax="71.0" valueMin="67.0">67–71</number>
(mean
<number id="3DE22F2F5B19335AFB18476141CFFEC2" box="[1189,1241,324,350]" pageId="7" pageNumber="40" value="69.7">69.7</number>
±
<number id="3DE22F2F5B19335AFB4A4761403CFEC2" box="[1271,1322,324,350]" pageId="7" pageNumber="40" value="2.31">2.31</number>
SD); ventral scales between postmentals and vent
<number id="3DE22F2F5B19335AFCDE474946ADFE1A" box="[867,955,364,390]" pageId="7" pageNumber="40" value="99.0" valueMax="100.0" valueMin="98.0">98–100</number>
(mean
<number id="3DE22F2F5B19335AFBA547494123FE1A" box="[1048,1077,364,390]" pageId="7" pageNumber="40" value="99.0">99</number>
±
<number id="3DE22F2F5B19335AFBE547494171FE1A" box="[1112,1127,364,390]" pageId="7" pageNumber="40" value="1.0">1</number>
SD). Precloacal pores absent. Supralabials
<number id="3DE22F2F5B19335AFDF147B1474DFE32" box="[588,603,404,430]" pageId="7" pageNumber="40" value="7.0">7</number>
(
<number id="3DE22F2F5B19335AFDD247B14768FE32" box="[623,638,404,430]" pageId="7" pageNumber="40" value="6.0">6</number>
/
<quantity id="4CD290ED5B19335AFD3A47B147AEFE32" box="[647,696,404,430]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="7" pageNumber="40" unit="in" value="7.0">7 in</quantity>
<collectionCode id="ED3BA5CD5B19335AFD7E47B14616FE32" box="[707,768,404,430]" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="7" pageNumber="40">MLP</collectionCode>
.R
<number id="3DE22F2F5B19335AFC9F47B14674FE32" box="[802,866,404,430]" pageId="7" pageNumber="40" value="5140.0">5140</number>
); infralabials
<number id="3DE22F2F5B19335AFBB447B14101FE32" box="[1033,1047,404,430]" pageId="7" pageNumber="40" value="6.0">6</number>
(
<number id="3DE22F2F5B19335AFB9647B1412FFE32" box="[1067,1081,404,430]" pageId="7" pageNumber="40" value="6.0">6</number>
/
<quantity id="4CD290ED5B19335AFBF947B14162FE32" box="[1092,1140,404,430]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="7" pageNumber="40" unit="in" value="7.0">7 in</quantity>
<collectionCode id="ED3BA5CD5B19335AFBC247B141AAFE32" box="[1151,1212,404,430]" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="7" pageNumber="40">MLP</collectionCode>
.R
<number id="3DE22F2F5B19335AFB6247B14009FE32" box="[1247,1311,404,430]" pageId="7" pageNumber="40" value="5140.0">5140</number>
). Infradigital lamellae under fourth left toe
<number id="3DE22F2F5B19335AFCB547994647FE4A" box="[776,849,444,470]" pageId="7" pageNumber="40" value="30.5" valueMax="31.0" valueMin="30.0">30–31</number>
(mean
<number id="3DE22F2F5B19335AFC0C479946F2FE4A" box="[945,996,444,470]" pageId="7" pageNumber="40" value="30.7">30.7</number>
±
<number id="3DE22F2F5B19335AFBB647994128FE4A" box="[1035,1086,444,470]" pageId="7" pageNumber="40" value="0.58">0.58</number>
SD). Dorsal scales moderately keeled, but more evident than in male. Femoral patch less marked and less spinose than in male. Other lepidosis features (number of azygous frontals, form and arrangement of body scales, distribution of scale organs) as in the male, except for scales on precloacal region, which in females are rounded, nearly granular, juxtaposed and smaller than ventrals.
</paragraph>
<paragraph id="8B953D085B19335AFE8544894017FB9A" blockId="7.[264,1323,284,1870]" pageId="7" pageNumber="40">
Coloration: Dorsal ground grey, with light brown shade. White scales less evident than in male to totally absent. Dorsal brown serial markings present as thin, discontinuous transverse bars, or as small squarish spots arranged in pairs at both sides of midline. On the tail dorsum these marks fuse on midline. Dark head lines as evident as in male, that one running from behind the eye easily visible from above. This line runs dorsolaterally on neck more-or-less continuously to end in a dark spot just dorsal to the antehumeral fold. Postscapular spot absent. Ill-defined brown stripes on the limbs, all scales finely dotted in brown. Black and white stripes on the posterodorsal surface of thighs. Femoral patch whitish. Throat white speckled with grey. Belly and other underparts, white (
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Figure
<number id="3DE22F2F5B19335AFB5C45C941E7FB9A" box="[1249,1265,1004,1030]" pageId="7" pageNumber="40" value="3.0">3</number>
</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3306E835B19335AFE85423147BBFAF2" pageId="7" pageNumber="40" type="etymology">
<paragraph id="8B953D085B19335AFE85423146D9FBCA" blockId="7.[264,1323,284,1870]" pageId="7" pageNumber="40">
<emphasis id="B95EE11A5B19335AFE85423144D2FBB2" bold="true" box="[312,452,1044,1070]" pageId="7" pageNumber="40">Etymology.</emphasis>
The specific epithet is a Latin adjective meaning “ash-grey”, in reference to the overall background color characteristic in this species.
</paragraph>
<paragraph id="8B953D085B19335AFE85424147BBFAF2" blockId="7.[264,1323,284,1870]" pageId="7" pageNumber="40">
<emphasis id="B95EE11A5B19335AFE8542414766FBE2" bold="true" box="[312,624,1124,1150]" pageId="7" pageNumber="40">Geographic distribution.</emphasis>
<taxonomicName id="4C2A468B5B19335AFDC642414677FBE1" box="[635,865,1124,1149]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="40" phylum="Chordata" rank="species" species="cinereus">
<emphasis id="B95EE11A5B19335AFDC642414677FBE1" box="[635,865,1124,1149]" italics="true" pageId="7" pageNumber="40">Liolaemus cinereus</emphasis>
</taxonomicName>
is known only from the
<typeStatus id="549183AA5B19335AFB2B424141DFFBE2" box="[1174,1225,1124,1150]" pageId="7" pageNumber="40">type</typeStatus>
locality and the immediate vicinity of it. All specimens were found on gravel and sandy soil in a landscape of Monte vegetation associated to the higher portions of the banks of the Blanco River. The dominant vegetation are semi-arid scrubs of
<taxonomicName id="4C2A468B5B19335AFC6542F941B8FB69" box="[984,1198,1244,1269]" class="Magnoliopsida" family="Zygophyllaceae" genus="Larrea" kingdom="Plantae" order="Zygophyllales" pageId="7" pageNumber="40" phylum="Tracheophyta" rank="species" species="divaricata">
<emphasis id="B95EE11A5B19335AFC6542F941B8FB69" box="[984,1198,1244,1269]" italics="true" pageId="7" pageNumber="40">Larrea divaricata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A468B5B19335AFB7942F94478FA81" class="Magnoliopsida" family="Fabaceae" genus="Prosopis" kingdom="Animalia" order="Fabales" pageId="7" pageNumber="40" phylum="Tracheophyta" rank="species" species="alpataco">
<emphasis id="B95EE11A5B19335AFB7942F94478FA81" italics="true" pageId="7" pageNumber="40">Prosopis alpataco</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2A468B5B19335AFE174321477DFA81" box="[426,619,1284,1309]" class="Magnoliopsida" family="Zygophyllaceae" genus="Bulnesia" kingdom="Plantae" order="Zygophyllales" pageId="7" pageNumber="40" phylum="Tracheophyta" rank="species" species="retama">
<emphasis id="B95EE11A5B19335AFE174321477DFA81" box="[426,619,1284,1309]" italics="true" pageId="7" pageNumber="40">Bulnesia retama</emphasis>
</taxonomicName>
, with high percentage of bare soil (
<figureCitation id="1311218D5B19335AFBAF4321416DFA82" box="[1042,1147,1284,1310]" captionStart="FIGURE 4" captionStartId="8.[264,367,1316,1340]" captionTargetBox="[267,1319,594,1292]" captionTargetId="figure@8.[267,1319,594,1292]" captionTargetPageId="8" captionText="FIGURE 4. General view of the habitat of Liolaemus cinereus showing in the foreground bushes of Monte vegetation." figureDoi="http://doi.org/10.5281/zenodo.273637" httpUri="https://zenodo.org/record/273637/files/figure.png" pageId="7" pageNumber="40">
Figure
<number id="3DE22F2F5B19335AFBD44321416DFA82" box="[1129,1147,1284,1310]" pageId="7" pageNumber="40" value="4.0">4</number>
</figureCitation>
). Other lizards syntopic with
<taxonomicName id="4C2A468B5B19335AFE0C43094722FAD9" box="[433,564,1324,1349]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="40" phylum="Chordata" rank="species" species="cinereus">
<emphasis id="B95EE11A5B19335AFE0C43094722FAD9" box="[433,564,1324,1349]" italics="true" pageId="7" pageNumber="40">L. cinereus</emphasis>
</taxonomicName>
are
<taxonomicName id="4C2A468B5B19335AFDD743094615FAD9" box="[618,771,1324,1349]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="40" phylum="Chordata" rank="species" species="montanezi">
<emphasis id="B95EE11A5B19335AFDD743094615FAD9" box="[618,771,1324,1349]" italics="true" pageId="7" pageNumber="40">L. montanezi</emphasis>
</taxonomicName>
(species of the same complex) and the Monte gekkonid lizard
<taxonomicName id="4C2A468B5B19335AFE7B437147B0FAF1" box="[454,678,1364,1389]" class="Reptilia" family="Phyllodactylidae" genus="Homonota" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="40" phylum="Chordata" rank="species" species="fasciata">
<emphasis id="B95EE11A5B19335AFE7B437147B0FAF1" box="[454,678,1364,1389]" italics="true" pageId="7" pageNumber="40">Homonota fasciata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3306E835B193355FE8543594482FDBA" lastPageId="8" lastPageNumber="41" pageId="7" pageNumber="40" type="description">
<paragraph id="8B953D085B19335AFE8543594792F992" blockId="7.[264,1323,284,1870]" pageId="7" pageNumber="40">
<emphasis id="B95EE11A5B19335AFE8543594609FA0A" bold="true" box="[312,799,1404,1430]" pageId="7" pageNumber="40">Natural history notes and Conclusions.</emphasis>
The posterior marginal teeth (verified in the specimen
<collectionCode id="ED3BA5CD5B19335AFE37438144F4FA22" box="[394,482,1444,1470]" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="7" pageNumber="40">MACN</collectionCode>
<number id="3DE22F2F5B19335AFE4B43814757FA22" box="[502,577,1444,1470]" pageId="7" pageNumber="40" value="38799.0">38799</number>
) bear moderately cusped, slightly expanded crowns. An inspection into the large intestine of the same individual reveals undigested hard parts of insects and helminthic parasites.
</paragraph>
<paragraph id="8B953D085B193355FE8540394482FDBA" blockId="7.[264,1323,284,1870]" lastBlockId="8.[264,1323,284,550]" lastPageId="8" lastPageNumber="41" pageId="7" pageNumber="40">
The San Guillermo National Park (
<figureCitation id="1311218D5B19335AFD6140394654F9AA" box="[732,834,1564,1590]" captionStart="FIGURE 5" captionStartId="9.[264,367,1218,1242]" captionTargetBox="[280,1306,322,1183]" captionTargetId="figure@9.[267,1319,306,1194]" captionTargetPageId="9" captionText="FIGURE 5. Map of the San Guillermo National Park (Landsat ETM image captured on 02 / 01 / 2002; Gauss-Kruger coordinates; Transverse Mercator Projection). A yellow circle marks the type locality of Liolaemus cinereus." figureDoi="http://doi.org/10.5281/zenodo.273638" httpUri="https://zenodo.org/record/273638/files/figure.png" pageId="7" pageNumber="40">
Figure
<number id="3DE22F2F5B19335AFC8E40394654F9AA" box="[819,834,1564,1590]" pageId="7" pageNumber="40" value="5.0">5</number>
</figureCitation>
) is a protected natural area of
<collectingCountry id="F33D7D985B19335AFB0B4039403DF9AA" box="[1206,1323,1564,1590]" name="Argentina" pageId="7" pageNumber="40">Argentina</collectingCountry>
covering
<number id="3DE22F2F5B19335AFECB406144D8F9C2" box="[374,462,1604,1630]" pageId="7" pageNumber="40" value="170000.0">170000</number>
hectares of High-Andean Puna landscape (
<bibRefCitation id="EFBB40F95B19335AFC74406141BAF9C2" author="Chebez" box="[969,1196,1604,1630]" pageId="7" pageNumber="43" refString="Chebez, J. C., Rey, N. R. &amp; Williams, J. D. (2005) Reptiles de los Parques Nacionales de la Argentina. LOLA, Buenos Aires, Monogr. 19, 75 pp." type="book" year="2005">
Chébez
<emphasis id="B95EE11A5B19335AFB9740614173F9C1" box="[1066,1125,1604,1629]" italics="true" pageId="7" pageNumber="40">et al.</emphasis>
<number id="3DE22F2F5B19335AFBD1406141BAF9C2" box="[1132,1196,1604,1630]" pageId="7" pageNumber="40" value="2005.0">2005</number>
</bibRefCitation>
). The Park is the core section of the Biosphere Reserve of San Guillermo which, with its surface of
<number id="3DE22F2F5B19335AFEB540B14476F932" box="[264,352,1684,1710]" pageId="7" pageNumber="40" value="981000.0">981000</number>
hectares harbors the larger Argentinean populations of wild camelids (
<emphasis id="B95EE11A5B19335AFB7640B14473F949" italics="true" pageId="7" pageNumber="40">Vicugna vicugna</emphasis>
and
<taxonomicName id="4C2A468B5B19335AFE114099477FF949" box="[428,617,1724,1749]" class="Mammalia" family="Camelidae" genus="Lama" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="40" phylum="Chordata" rank="species" species="guanicoe">
<emphasis id="B95EE11A5B19335AFE114099477FF949" box="[428,617,1724,1749]" italics="true" pageId="7" pageNumber="40">Lama guanicoe</emphasis>
</taxonomicName>
), and gives protection to representative ecosystems and threatened fauna,
<emphasis id="B95EE11A5B19335AFE6040C14707F961" box="[477,529,1764,1789]" italics="true" pageId="7" pageNumber="40">e. g.</emphasis>
the birds
<taxonomicName id="4C2A468B5B19335AFD3140C1462BF961" box="[652,829,1764,1789]" class="Aves" family="Rallidae" genus="Fulica" kingdom="Animalia" order="Gruiformes" pageId="7" pageNumber="40" phylum="Chordata" rank="species" species="cornuta">
<emphasis id="B95EE11A5B19335AFD3140C1462BF961" box="[652,829,1764,1789]" italics="true" pageId="7" pageNumber="40">Fulica cornuta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2A468B5B19335AFCC740C1446CF8BA" authority="Haene et al. 2000" authorityName="Haene et al." authorityYear="2000" class="Aves" family="Rheidae" genus="Pterocnemia" kingdom="Animalia" order="Struthioniformes" pageId="7" pageNumber="40" phylum="Chordata" rank="species" species="tarapacensis">
<emphasis id="B95EE11A5B19335AFCC740C141B8F961" box="[890,1198,1764,1789]" italics="true" pageId="7" pageNumber="40">Pterocnemia tarapacensis</emphasis>
(
<bibRefCitation id="EFBB40F95B19335AFB0240C14464F8BA" author="Haene" pageId="7" pageNumber="43" refString="Haene, E., Montanez, A., Carrizo, A., Bodrati, G., Bono, J., Krauss, G., Merida, E., Nardini, C., Rodriguez, R., Jones, J. &amp; Perez, A. (2000) Primer Inventario de los Animales Vertebrados del Parque Nacional San Guillermo (Provincia de San Juan, Republica Argentina). Aves Argentinas / AOP - P. N. San Guillermo; Bs. As. y Rodeo, 37 pp." type="book" year="2000">
Haene
<emphasis id="B95EE11A5B19335AFAA840C14430F8B9" italics="true" pageId="7" pageNumber="40">et al.</emphasis>
<number id="3DE22F2F5B19335AFE8F41294464F8BA" box="[306,370,1804,1830]" pageId="7" pageNumber="40" value="2000.0">2000</number>
</bibRefCitation>
)
</taxonomicName>
. Large plains between
<number id="3DE22F2F5B19335AFD24412947C2F8BA" box="[665,724,1804,1830]" pageId="7" pageNumber="40" value="3000.0">3000</number>
and
<number id="3DE22F2F5B19335AFCA841294646F8BA" box="[789,848,1804,1830]" pageId="7" pageNumber="40" value="4200.0">4200</number>
meters of altitude surrounded by high cordilleras above the
<number id="3DE22F2F5B19335AFDBE41114728F8D2" box="[515,574,1844,1870]" pageId="7" pageNumber="40" value="4200.0">4200</number>
meters and deep ravines (“quebradas”) between
<number id="3DE22F2F5B19335AFBC1411141A1F8D2" box="[1148,1207,1844,1870]" pageId="7" pageNumber="40" value="2100.0">2100</number>
and
<number id="3DE22F2F5B19335AFB524111403CF8D2" box="[1263,1322,1844,1870]" pageId="7" pageNumber="40" value="3000.0">3000</number>
meters determine isolated spots that seem to have favoured speciation events. Most of the speciation processes in the
<taxonomicName id="4C2A468B5B163355FDED476147A5FEC1" box="[592,691,324,349]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="41" phylum="Chordata" rank="species" species="darwinii">
<emphasis id="B95EE11A5B163355FDED476147A5FEC1" box="[592,691,324,349]" italics="true" pageId="8" pageNumber="41">darwinii</emphasis>
</taxonomicName>
complex may have happened during the Pleistocene and could be related to climatic changes during this period (
<bibRefCitation id="EFBB40F95B163355FC6C474941DAFE1A" author="Morando" box="[977,1228,364,390]" pageId="8" pageNumber="43" refString="Morando, M., Avila, L. J., Baker, J. &amp; Sites Jr., J. W. (2004) Phylogeny and phylogeography of the Liolaemus darwinii complex (Squamata: Liolaemidae): Evidence for introgression and incomplete lineage sorting. Evolution, 58 (4), 842 - 861." type="journal article" year="2004">
Morando
<emphasis id="B95EE11A5B163355FBFA47494195FE19" box="[1095,1155,364,389]" italics="true" pageId="8" pageNumber="41">et al.</emphasis>
<number id="3DE22F2F5B163355FB31474941DAFE1A" box="[1164,1228,364,390]" pageId="8" pageNumber="41" value="2004.0">2004</number>
</bibRefCitation>
). Hence the Reserve is, not unexpectedly, rich in iguanid endemisms:
<taxonomicName id="4C2A468B5B163355FB8147B14032FE31" box="[1084,1316,404,429]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="41" phylum="Chordata" rank="species" species="punae">
<emphasis id="B95EE11A5B163355FB8147B14032FE31" box="[1084,1316,404,429]" italics="true" pageId="8" pageNumber="41">Phymaturus punae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A468B5B163355FEB5479947DAFE4A" authority="Cei et al. 1983" authorityName="Cei et al." authorityYear="1983" box="[264,716,444,470]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="41" phylum="Chordata" rank="species" species="eleodori">
<emphasis id="B95EE11A5B163355FEB5479944E6FE49" box="[264,496,444,469]" italics="true" pageId="8" pageNumber="41">Liolaemus eleodori</emphasis>
(
<bibRefCitation id="EFBB40F95B163355FDB9479947D4FE4A" author="Cei" box="[516,706,444,470]" pageId="8" pageNumber="43" refString="Cei, J. M., Etheridge, R. &amp; Videla, F. (1983) Especies nuevas de Iguanidos del noroeste de la provincia de San Juan (Reserva provincial San Guillermo), Argentina. Deserta, (7), 316 - 323." type="journal article" year="1983">
Cei
<emphasis id="B95EE11A5B163355FD874799476FFE49" box="[570,633,444,469]" italics="true" pageId="8" pageNumber="41">et al.</emphasis>
<number id="3DE22F2F5B163355FD39479947D4FE4A" box="[644,706,444,470]" pageId="8" pageNumber="41" value="1983.0">1983</number>
</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C2A468B5B163355FD6147994014FE4A" authority="Cabrera and Monguillot 2006" authorityName="Cabrera and Monguillot" authorityYear="2006" box="[732,1282,444,470]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="41" phylum="Chordata" rank="species" species="montanezi">
<emphasis id="B95EE11A5B163355FD6147994661FE49" box="[732,887,444,469]" italics="true" pageId="8" pageNumber="41">L. montanezi</emphasis>
(
<bibRefCitation id="EFBB40F95B163355FC36479941ECFE4A" author="Cabrera" box="[907,1274,444,470]" pageId="8" pageNumber="42" refString="Cabrera, M. R. &amp; Monguillot, J. C. (2006) A new Andean species of Liolaemus of the darwinii complex (Reptilia: Iguanidae). Zootaxa, 1106, 35 - 43." type="journal article" year="2006">
Cabrera and Monguillot
<number id="3DE22F2F5B163355FB06479941ECFE4A" box="[1211,1274,444,470]" pageId="8" pageNumber="41" value="2006.0">2006</number>
</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C2A468B5B163355FAAE4799447CFE61" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="41" phylum="Chordata" rank="species" species="cinereus" status="sp. nov.">
<emphasis id="B95EE11A5B163355FAAE4799447CFE61" italics="true" pageId="8" pageNumber="41">L. cinereus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D5C615B163355FEC747C144C9FE62" box="[378,479,484,510]" pageId="8" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
, and several species under study and description by us and other researchers.
</paragraph>
</subSubSection>
</treatment>
</document>
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