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<document id="11024BDE7DB662DF6282AF00239ADB21" ID-DOI="10.26879/841" ID-ISSN="1094-8074" ID-Zenodo-Dep="11062827" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1714011148597" checkinUser="felipe" docAuthor="Wang, Hongshan &amp; Dilcher, David L." docDate="2018" docId="03838A22FF98AA2FFF1AFC87FAEA31DF" docLanguage="en" docName="PalaeontolElectron.21.3.34A.pdf" docOrigin="Palaeontologia Electronica (34 A) 21 (3)" docSource="http://dx.doi.org/10.26879/841" docStyle="DocumentStyle:322FC57CC99045555090E8A84CFB029A.4:PalaeontolElectron.2018-.journal_article" docStyleId="322FC57CC99045555090E8A84CFB029A" docStyleName="PalaeontolElectron.2018-.journal_article" docStyleVersion="4" docTitle="Pabiania variloba Upchurch and Dilcher 1990" docType="treatment" docVersion="3" lastPageNumber="12" masterDocId="FFBAF25AFF93AA24FFC4FFEEFFCB377D" masterDocTitle="Early Cretaceous angiosperm leaves from the Dakota Formation, Hoisington III locality, Kansas, USA" masterLastPageNumber="49" masterPageNumber="1" pageNumber="12" updateTime="1715622024402" updateUser="carolina" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="B36D3C49E4251EB17F977C217B417EE8">Early Cretaceous angiosperm leaves from the Dakota Formation, Hoisington III locality, Kansas, USA</mods:title>
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<mods:namePart id="C39EB6A55D6EEBACFBAA73CEAEA9C1C0">Wang, Hongshan</mods:namePart>
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<mods:namePart id="5ED327BE4CE1CD29897B14925892899D">Dilcher, David L.</mods:namePart>
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<treatment id="03838A22FF98AA2FFF1AFC87FAEA31DF" LSID="urn:lsid:plazi:treatment:03838A22FF98AA2FFF1AFC87FAEA31DF" httpUri="http://treatment.plazi.org/id/03838A22FF98AA2FFF1AFC87FAEA31DF" lastPageNumber="12" pageId="11" pageNumber="12">
<subSubSection id="C33068BFFF98AA2FFF1AFC87FD3134FF" box="[222,762,873,898]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8B953B34FF98AA2FFF1AFC87FD3134FF" blockId="11.[222,762,873,930]" box="[222,762,873,898]" pageId="11" pageNumber="12">
<heading id="D0DD8C58FF98AA2FFF1AFC87FD3134FF" box="[222,762,873,898]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="12" reason="7">
<taxonomicName id="4C2A40B7FF98AA2FFF1AFC87FD3134FF" authority="Upchurch and Dilcher, 1990" authorityName="Upchurch and Dilcher" authorityYear="1990" box="[222,762,873,898]" class="Magnoliopsida" genus="Pabiania" higherTaxonomySource="GBIF" kingdom="Plantae" order="Laurales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="variloba">
<emphasis id="B95EE726FF98AA2FFF1AFC87FE6134FF" box="[222,426,873,898]" italics="true" pageId="11" pageNumber="12">Pabiania variloba</emphasis>
Upchurch and Dilcher, 1990
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33068BFFF98AA2FFE6DFC67FB61351F" pageId="11" pageNumber="12" type="description">
<paragraph id="8B953B34FF98AA2FFE6DFC67FDE434DF" blockId="11.[222,762,873,930]" box="[425,559,905,930]" pageId="11" pageNumber="12">
<figureCitation id="131127B1FF98AA2FFE6DFC67FDE434DF" box="[425,559,905,930]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStartId-0="12.[197,288,1708,1731]" captionStartId-1="13.[197,288,1698,1721]" captionTargetBox-0="[240,1392,165,1679]" captionTargetBox-1="[199,1434,168,1648]" captionTargetId-0="figure-13@12.[240,1392,165,1679]" captionTargetId-1="figure-19@13.[199,1434,167,1652]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="FIGURE 7. Pabiania variloba Upchurch and Dilcher, 1990. 1-3, UF15706-24423, specimen showing suprabasal actinodromous primary venation (scale bar equals 1 cm) (1), enlargement of a sinus area to show sinus bracing by secondary veins (scale bar equals 2 mm) (2); and enlargement of basal portion of leaf to show two pairs of basal secondary veins (scale bar equals 2 mm) (3). 4-5, UF15706-30154, specimen showing a long petiole and ocrea-like structure at the base (scale bar equals 5 mm) (4) and enlargement of the petiole to show the ocrea-like structure (scale bar equals 2 mm) (5). 6, UF15706-14832, specimen showing suprabasal actinodromous primary venation and acute apices of the lobes. Scale bar equals 1 cm." captionText-1="FIGURE 8. Pabiania variloba Upchurch and Dilcher, 1990. 1, UF15706-14823, leaf showing basal actinodromous primary venation and basal secondary veins. Scale bar equals 1 cm. 2, UF15706-24464, leaf showing rounded lobe apex. Scale bar equals 1 cm. 3, UF15706-24587, specimen showing a small lobe on the left and entire margin on the right of the leaf. Scale bar equals 5 mm. 4, Enlargement of Figure 8.2 to show straight primary vein extending to lobe apex and two series of loops in the excostal region. Scale bar equals 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.11062841" figureDoi-1="http://doi.org/10.5281/zenodo.11062843" httpUri-0="https://zenodo.org/record/11062841/files/figure.png" httpUri-1="https://zenodo.org/record/11062843/files/figure.png" pageId="11" pageNumber="12">Figures 7-8</figureCitation>
</paragraph>
<paragraph id="8B953B34FF98AA2FFF04FC5DFD513495" blockId="11.[192,793,947,1901]" pageId="11" pageNumber="12">
v. 1981a
<taxonomicName id="4C2A40B7FF98AA2FFEF7FC5DFD6A34B7" authority="Lesquereux" authorityName="Lesquereux" authorityYear="1891" box="[307,673,947,970]" class="Magnoliopsida" family="Sapindaceae" genus="Acerites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="multiformis">
<emphasis id="B95EE726FF98AA2FFEF7FC5DFDC434B7" box="[307,527,947,970]" italics="true" pageId="11" pageNumber="12">Acerites multiformis</emphasis>
Lesquereux
</taxonomicName>
; Retallack and Dilcher, 1981a, p. 38, fig. 2.4.
</paragraph>
<paragraph id="8B953B34FF98AA2FFF04FC1DFC7036FF" blockId="11.[192,793,947,1901]" lastBlockId="11.[840,1441,168,1698]" pageId="11" pageNumber="12">
<emphasis id="B95EE726FF98AA2FFF04FC1DFE913371" bold="true" box="[192,346,1011,1036]" pageId="11" pageNumber="12">Description.</emphasis>
Leaf commonly trilobate, rarely bilobate or unlobed; base ranging from acute to obtuse and tending towards cuneate, with the basalmost portion of the lamina decurrent on the petiole; margin entire; apex of lobes acute to rounded; unlobed leaves broad elliptic to obovate, with rounded apex. Primary venation basal to suprabasal actinodromous in 3-lobed leaves, pinnate in unlobed leaves; midvein moderate; lateral primary veins similar to midvein in thickness, at lower angle than superadjacent secondary veins and tending to be recurved, commonly decurrent on midvein, producing one to three festooned brochidodromous external branches. Secondary venation festooned brochidodromous; secondary veins thin or moderate relative to primary veins, three to six pairs along midvein, with one pair of basilaminar secondary veins present in all leaves, alternate, moderate acute, straight to apically curved, tending to be festooned brochidodromous except near sinuses. Tertiary venation reticulate; tertiary veins thin relative to secondary veins, opposite to alternate, closely but somewhat irregularly spaced, originating at acute to right angles. Quaternary venation intergrading with tertiary veins, generally unbranched; quaternary veins opposite to alternate, non-orthogonal, enclosing somewhat elongate, 4-sided regions. Quinternary venation highly irregular, both ramified and reticulate, arising both from thick lateral branches of the quaternary veins and from the sides of lower order veins; quinternary veins weak, curved, often producing branches to form an open venation. Areolation poorly developed. Marginal venation consisting of a thin fimbrial vein.
</paragraph>
<paragraph id="8B953B34FF98AA2FFC8CFE67FADE36DF" blockId="11.[840,1441,168,1698]" box="[840,1301,393,418]" pageId="11" pageNumber="12">
<emphasis id="B95EE726FF98AA2FFC8CFE67FB2336DF" bold="true" box="[840,1256,393,418]" pageId="11" pageNumber="12">Number of specimens examined.</emphasis>
90.
</paragraph>
<paragraph id="8B953B34FF98AA2FFC8CFE46FC11355F" blockId="11.[840,1441,168,1698]" pageId="11" pageNumber="12">
<emphasis id="B95EE726FF98AA2FFC8CFE46FBA036BF" bold="true" box="[840,1131,424,450]" pageId="11" pageNumber="12">Specimens illustrated.</emphasis>
<collectionCode id="ED3BA3F1FF98AA2FFBBFFE47FB5636BF" box="[1147,1181,425,450]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="11" pageNumber="12" type="Museum">UF</collectionCode>
15706-24423 (
<figureCitation id="131127B1FF98AA2FFA96FE47FC42369F" captionStart="FIGURE 7" captionStartId="12.[197,288,1708,1731]" captionTargetBox="[240,1392,165,1679]" captionTargetId="figure-13@12.[240,1392,165,1679]" captionTargetPageId="12" captionText="FIGURE 7. Pabiania variloba Upchurch and Dilcher, 1990. 1-3, UF15706-24423, specimen showing suprabasal actinodromous primary venation (scale bar equals 1 cm) (1), enlargement of a sinus area to show sinus bracing by secondary veins (scale bar equals 2 mm) (2); and enlargement of basal portion of leaf to show two pairs of basal secondary veins (scale bar equals 2 mm) (3). 4-5, UF15706-30154, specimen showing a long petiole and ocrea-like structure at the base (scale bar equals 5 mm) (4) and enlargement of the petiole to show the ocrea-like structure (scale bar equals 2 mm) (5). 6, UF15706-14832, specimen showing suprabasal actinodromous primary venation and acute apices of the lobes. Scale bar equals 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11062841" httpUri="https://zenodo.org/record/11062841/files/figure.png" pageId="11" pageNumber="12">Figure 7.1-3</figureCitation>
); 30154 (
<figureCitation id="131127B1FF98AA2FFC3BFE27FB57369F" box="[1023,1180,457,482]" captionStart="FIGURE 7" captionStartId="12.[197,288,1708,1731]" captionTargetBox="[240,1392,165,1679]" captionTargetId="figure-13@12.[240,1392,165,1679]" captionTargetPageId="12" captionText="FIGURE 7. Pabiania variloba Upchurch and Dilcher, 1990. 1-3, UF15706-24423, specimen showing suprabasal actinodromous primary venation (scale bar equals 1 cm) (1), enlargement of a sinus area to show sinus bracing by secondary veins (scale bar equals 2 mm) (2); and enlargement of basal portion of leaf to show two pairs of basal secondary veins (scale bar equals 2 mm) (3). 4-5, UF15706-30154, specimen showing a long petiole and ocrea-like structure at the base (scale bar equals 5 mm) (4) and enlargement of the petiole to show the ocrea-like structure (scale bar equals 2 mm) (5). 6, UF15706-14832, specimen showing suprabasal actinodromous primary venation and acute apices of the lobes. Scale bar equals 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11062841" httpUri="https://zenodo.org/record/11062841/files/figure.png" pageId="11" pageNumber="12">Figure 7.4-5</figureCitation>
); 14832 (
<figureCitation id="131127B1FF98AA2FFAD4FE27FA5E369F" box="[1296,1429,457,482]" captionStart="FIGURE 7" captionStartId="12.[197,288,1708,1731]" captionTargetBox="[240,1392,165,1679]" captionTargetId="figure-13@12.[240,1392,165,1679]" captionTargetPageId="12" captionText="FIGURE 7. Pabiania variloba Upchurch and Dilcher, 1990. 1-3, UF15706-24423, specimen showing suprabasal actinodromous primary venation (scale bar equals 1 cm) (1), enlargement of a sinus area to show sinus bracing by secondary veins (scale bar equals 2 mm) (2); and enlargement of basal portion of leaf to show two pairs of basal secondary veins (scale bar equals 2 mm) (3). 4-5, UF15706-30154, specimen showing a long petiole and ocrea-like structure at the base (scale bar equals 5 mm) (4) and enlargement of the petiole to show the ocrea-like structure (scale bar equals 2 mm) (5). 6, UF15706-14832, specimen showing suprabasal actinodromous primary venation and acute apices of the lobes. Scale bar equals 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11062841" httpUri="https://zenodo.org/record/11062841/files/figure.png" pageId="11" pageNumber="12">Figure 7.6</figureCitation>
); 14823 (
<figureCitation id="131127B1FF98AA2FFC65FE07FBEB357F" box="[929,1056,489,514]" captionStart="FIGURE 8" captionStartId="13.[197,288,1698,1721]" captionTargetBox="[199,1434,168,1648]" captionTargetId="figure-19@13.[199,1434,167,1652]" captionTargetPageId="13" captionText="FIGURE 8. Pabiania variloba Upchurch and Dilcher, 1990. 1, UF15706-14823, leaf showing basal actinodromous primary venation and basal secondary veins. Scale bar equals 1 cm. 2, UF15706-24464, leaf showing rounded lobe apex. Scale bar equals 1 cm. 3, UF15706-24587, specimen showing a small lobe on the left and entire margin on the right of the leaf. Scale bar equals 5 mm. 4, Enlargement of Figure 8.2 to show straight primary vein extending to lobe apex and two series of loops in the excostal region. Scale bar equals 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11062843" httpUri="https://zenodo.org/record/11062843/files/figure.png" pageId="11" pageNumber="12">Figure 8.1</figureCitation>
); 24464 (
<figureCitation id="131127B1FF98AA2FFB49FE07FA89357F" box="[1165,1346,489,514]" captionStart="FIGURE 8" captionStartId="13.[197,288,1698,1721]" captionTargetBox="[199,1434,168,1648]" captionTargetId="figure-19@13.[199,1434,167,1652]" captionTargetPageId="13" captionText="FIGURE 8. Pabiania variloba Upchurch and Dilcher, 1990. 1, UF15706-14823, leaf showing basal actinodromous primary venation and basal secondary veins. Scale bar equals 1 cm. 2, UF15706-24464, leaf showing rounded lobe apex. Scale bar equals 1 cm. 3, UF15706-24587, specimen showing a small lobe on the left and entire margin on the right of the leaf. Scale bar equals 5 mm. 4, Enlargement of Figure 8.2 to show straight primary vein extending to lobe apex and two series of loops in the excostal region. Scale bar equals 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11062843" httpUri="https://zenodo.org/record/11062843/files/figure.png" pageId="11" pageNumber="12">Figure 8.2, 8.4</figureCitation>
); 24587 (
<figureCitation id="131127B1FF98AA2FFC8AFDE7FC04355F" box="[846,975,521,546]" captionStart="FIGURE 8" captionStartId="13.[197,288,1698,1721]" captionTargetBox="[199,1434,168,1648]" captionTargetId="figure-19@13.[199,1434,167,1652]" captionTargetPageId="13" captionText="FIGURE 8. Pabiania variloba Upchurch and Dilcher, 1990. 1, UF15706-14823, leaf showing basal actinodromous primary venation and basal secondary veins. Scale bar equals 1 cm. 2, UF15706-24464, leaf showing rounded lobe apex. Scale bar equals 1 cm. 3, UF15706-24587, specimen showing a small lobe on the left and entire margin on the right of the leaf. Scale bar equals 5 mm. 4, Enlargement of Figure 8.2 to show straight primary vein extending to lobe apex and two series of loops in the excostal region. Scale bar equals 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11062843" httpUri="https://zenodo.org/record/11062843/files/figure.png" pageId="11" pageNumber="12">Figure 8.3</figureCitation>
).
</paragraph>
<paragraph id="8B953B34FF98AA2FFC8CFDC6FB61351F" blockId="11.[840,1441,168,1698]" pageId="11" pageNumber="12">
<emphasis id="B95EE726FF98AA2FFC8CFDC6FC39353F" bold="true" box="[840,1010,552,578]" pageId="11" pageNumber="12">Occurrences.</emphasis>
<materialsCitation id="3B423169FF98AA2FFC38FDC7FAA2353F" box="[1020,1385,552,578]" location="Rose Creek" pageId="11" pageNumber="12" specimenCount="1" stateProvince="Nebraska">
<location id="8EF56DEFFF98AA2FFC38FDC7FB47353F" LSID="urn:lsid:plazi:treatment:03838A22FF98AA2FFF1AFC87FAEA31DF:8EF56DEFFF98AA2FFC38FDC7FB47353F" box="[1020,1164,552,578]" name="Rose Creek" pageId="11" pageNumber="12" stateProvince="Nebraska">Rose Creek</location>
locality,
<collectingRegion id="49EEF5D6FF98AA2FFB33FDC7FAA2353F" box="[1271,1385,553,578]" country="United States of America" name="Nebraska" pageId="11" pageNumber="12">Nebraska</collectingRegion>
</materialsCitation>
and Hoisington III locality,
<collectingRegion id="49EEF5D6FF98AA2FFB8DFDA7FB6D351F" box="[1097,1190,585,610]" country="United States of America" name="Kansas" pageId="11" pageNumber="12">Kansas</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C33068BFFF98AA2FFC8CFD87FAEA31DF" pageId="11" pageNumber="12" type="discussion">
<paragraph id="8B953B34FF98AA2FFC8CFD87FA8832DF" blockId="11.[840,1441,168,1698]" pageId="11" pageNumber="12">
<emphasis id="B95EE726FF98AA2FFC8CFD87FC0B35FF" bold="true" box="[840,960,617,642]" pageId="11" pageNumber="12">Remarks.</emphasis>
In Upchurch and Dilchers (1990) diagnosis and description of this species, the primary venation was described as palinactinodromous. Observations based on Hoisington III locality specimens indicate that the primary venation of
<taxonomicName id="4C2A40B7FF98AA2FFA8AFD07FCB1345F" authorityName="Upchurch and Dilcher" authorityYear="1990" class="Magnoliopsida" genus="Pabiania" higherTaxonomySource="GBIF" kingdom="Plantae" order="Laurales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="variloba">
<emphasis id="B95EE726FF98AA2FFA8AFD07FCB1345F" italics="true" pageId="11" pageNumber="12">P. variloba</emphasis>
</taxonomicName>
can vary from basal actinodromous (
<figureCitation id="131127B1FF98AA2FFA96FCE7FCBA343F" captionStart="FIGURE 8" captionStartId="13.[197,288,1698,1721]" captionTargetBox="[199,1434,168,1648]" captionTargetId="figure-19@13.[199,1434,167,1652]" captionTargetPageId="13" captionText="FIGURE 8. Pabiania variloba Upchurch and Dilcher, 1990. 1, UF15706-14823, leaf showing basal actinodromous primary venation and basal secondary veins. Scale bar equals 1 cm. 2, UF15706-24464, leaf showing rounded lobe apex. Scale bar equals 1 cm. 3, UF15706-24587, specimen showing a small lobe on the left and entire margin on the right of the leaf. Scale bar equals 5 mm. 4, Enlargement of Figure 8.2 to show straight primary vein extending to lobe apex and two series of loops in the excostal region. Scale bar equals 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11062843" httpUri="https://zenodo.org/record/11062843/files/figure.png" pageId="11" pageNumber="12">Figure 8.1</figureCitation>
), suprabasal actinodromous (
<figureCitation id="131127B1FF98AA2FFB22FCC7FAA4343F" box="[1254,1391,809,834]" captionStart="FIGURE 7" captionStartId="12.[197,288,1708,1731]" captionTargetBox="[240,1392,165,1679]" captionTargetId="figure-13@12.[240,1392,165,1679]" captionTargetPageId="12" captionText="FIGURE 7. Pabiania variloba Upchurch and Dilcher, 1990. 1-3, UF15706-24423, specimen showing suprabasal actinodromous primary venation (scale bar equals 1 cm) (1), enlargement of a sinus area to show sinus bracing by secondary veins (scale bar equals 2 mm) (2); and enlargement of basal portion of leaf to show two pairs of basal secondary veins (scale bar equals 2 mm) (3). 4-5, UF15706-30154, specimen showing a long petiole and ocrea-like structure at the base (scale bar equals 5 mm) (4) and enlargement of the petiole to show the ocrea-like structure (scale bar equals 2 mm) (5). 6, UF15706-14832, specimen showing suprabasal actinodromous primary venation and acute apices of the lobes. Scale bar equals 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11062841" httpUri="https://zenodo.org/record/11062841/files/figure.png" pageId="11" pageNumber="12">Figure 7.6</figureCitation>
), to suprabasal palinactinodromous (
<figureCitation id="131127B1FF98AA2FFB03FCA7FA94341F" box="[1223,1375,841,866]" captionStart="FIGURE 7" captionStartId="12.[197,288,1708,1731]" captionTargetBox="[240,1392,165,1679]" captionTargetId="figure-13@12.[240,1392,165,1679]" captionTargetPageId="12" captionText="FIGURE 7. Pabiania variloba Upchurch and Dilcher, 1990. 1-3, UF15706-24423, specimen showing suprabasal actinodromous primary venation (scale bar equals 1 cm) (1), enlargement of a sinus area to show sinus bracing by secondary veins (scale bar equals 2 mm) (2); and enlargement of basal portion of leaf to show two pairs of basal secondary veins (scale bar equals 2 mm) (3). 4-5, UF15706-30154, specimen showing a long petiole and ocrea-like structure at the base (scale bar equals 5 mm) (4) and enlargement of the petiole to show the ocrea-like structure (scale bar equals 2 mm) (5). 6, UF15706-14832, specimen showing suprabasal actinodromous primary venation and acute apices of the lobes. Scale bar equals 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11062841" httpUri="https://zenodo.org/record/11062841/files/figure.png" pageId="11" pageNumber="12">Figure 7.1-3</figureCitation>
). The leaves from the Rose Creek locality,
<collectingRegion id="49EEF5D6FF98AA2FFAEAFC87FA6B34FF" box="[1326,1440,873,898]" country="United States of America" name="Nebraska" pageId="11" pageNumber="12">Nebraska</collectingRegion>
described by Upchurch and Dilcher (1990) and stored at the Paleobotany collection in the
<collectingRegion id="49EEF5D6FF98AA2FFA8AFC47FA5434BF" box="[1358,1439,937,962]" country="United States of America" name="Florida" pageId="11" pageNumber="12">Florida</collectingRegion>
Museum of Natural History (more than
<specimenCount id="9D2CF0BDFF98AA2FFAE7FC27FC47337F" count="100" pageId="11" pageNumber="12" type="generic">100 specimens</specimenCount>
) are smaller in size compared with the specimens from the Hoisington III locality,
<collectingRegion id="49EEF5D6FF98AA2FFACCFBE7FAAE335F" box="[1288,1381,1033,1058]" country="United States of America" name="Kansas" pageId="11" pageNumber="12">Kansas</collectingRegion>
. The specimens from the Hoisington III locality also show that the petioles are long and they have winged lamina tissue along both sides. The petiole base may be ocreate (
<figureCitation id="131127B1FF98AA2FFB8AFB67FAC933DF" box="[1102,1282,1161,1186]" captionStart="FIGURE 7" captionStartId="12.[197,288,1708,1731]" captionTargetBox="[240,1392,165,1679]" captionTargetId="figure-13@12.[240,1392,165,1679]" captionTargetPageId="12" captionText="FIGURE 7. Pabiania variloba Upchurch and Dilcher, 1990. 1-3, UF15706-24423, specimen showing suprabasal actinodromous primary venation (scale bar equals 1 cm) (1), enlargement of a sinus area to show sinus bracing by secondary veins (scale bar equals 2 mm) (2); and enlargement of basal portion of leaf to show two pairs of basal secondary veins (scale bar equals 2 mm) (3). 4-5, UF15706-30154, specimen showing a long petiole and ocrea-like structure at the base (scale bar equals 5 mm) (4) and enlargement of the petiole to show the ocrea-like structure (scale bar equals 2 mm) (5). 6, UF15706-14832, specimen showing suprabasal actinodromous primary venation and acute apices of the lobes. Scale bar equals 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11062841" httpUri="https://zenodo.org/record/11062841/files/figure.png" pageId="11" pageNumber="12">Figure 7.4, 7.5</figureCitation>
). Other variations of leaf morphological characters include: (1) lamina from unlobed (
<figureCitation id="131127B1FF98AA2FFB97FB27FB18339F" box="[1107,1235,1225,1250]" captionStart="FIGURE 8" captionStartId="13.[197,288,1698,1721]" captionTargetBox="[199,1434,168,1648]" captionTargetId="figure-19@13.[199,1434,167,1652]" captionTargetPageId="13" captionText="FIGURE 8. Pabiania variloba Upchurch and Dilcher, 1990. 1, UF15706-14823, leaf showing basal actinodromous primary venation and basal secondary veins. Scale bar equals 1 cm. 2, UF15706-24464, leaf showing rounded lobe apex. Scale bar equals 1 cm. 3, UF15706-24587, specimen showing a small lobe on the left and entire margin on the right of the leaf. Scale bar equals 5 mm. 4, Enlargement of Figure 8.2 to show straight primary vein extending to lobe apex and two series of loops in the excostal region. Scale bar equals 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11062843" httpUri="https://zenodo.org/record/11062843/files/figure.png" pageId="11" pageNumber="12">Figure 8.3</figureCitation>
) to trilobed (
<figureCitation id="131127B1FF98AA2FFAB4FB27FC2E327F" captionStart="FIGURE 7" captionStartId="12.[197,288,1708,1731]" captionTargetBox="[240,1392,165,1679]" captionTargetId="figure-13@12.[240,1392,165,1679]" captionTargetPageId="12" captionText="FIGURE 7. Pabiania variloba Upchurch and Dilcher, 1990. 1-3, UF15706-24423, specimen showing suprabasal actinodromous primary venation (scale bar equals 1 cm) (1), enlargement of a sinus area to show sinus bracing by secondary veins (scale bar equals 2 mm) (2); and enlargement of basal portion of leaf to show two pairs of basal secondary veins (scale bar equals 2 mm) (3). 4-5, UF15706-30154, specimen showing a long petiole and ocrea-like structure at the base (scale bar equals 5 mm) (4) and enlargement of the petiole to show the ocrea-like structure (scale bar equals 2 mm) (5). 6, UF15706-14832, specimen showing suprabasal actinodromous primary venation and acute apices of the lobes. Scale bar equals 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11062841" httpUri="https://zenodo.org/record/11062841/files/figure.png" pageId="11" pageNumber="12">Figures 7.1, 7.6</figureCitation>
,
<figureCitation id="131127B1FF98AA2FFC36FB07FBF8327F" box="[1010,1075,1257,1282]" captionStart="FIGURE 8" captionStartId="13.[197,288,1698,1721]" captionTargetBox="[199,1434,168,1648]" captionTargetId="figure-19@13.[199,1434,167,1652]" captionTargetPageId="13" captionText="FIGURE 8. Pabiania variloba Upchurch and Dilcher, 1990. 1, UF15706-14823, leaf showing basal actinodromous primary venation and basal secondary veins. Scale bar equals 1 cm. 2, UF15706-24464, leaf showing rounded lobe apex. Scale bar equals 1 cm. 3, UF15706-24587, specimen showing a small lobe on the left and entire margin on the right of the leaf. Scale bar equals 5 mm. 4, Enlargement of Figure 8.2 to show straight primary vein extending to lobe apex and two series of loops in the excostal region. Scale bar equals 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11062843" httpUri="https://zenodo.org/record/11062843/files/figure.png" pageId="11" pageNumber="12">8.1-2</figureCitation>
); leaf base from acute (
<figureCitation id="131127B1FF98AA2FFA96FB07FCA4325F" captionStart="FIGURE 7" captionStartId="12.[197,288,1708,1731]" captionTargetBox="[240,1392,165,1679]" captionTargetId="figure-13@12.[240,1392,165,1679]" captionTargetPageId="12" captionText="FIGURE 7. Pabiania variloba Upchurch and Dilcher, 1990. 1-3, UF15706-24423, specimen showing suprabasal actinodromous primary venation (scale bar equals 1 cm) (1), enlargement of a sinus area to show sinus bracing by secondary veins (scale bar equals 2 mm) (2); and enlargement of basal portion of leaf to show two pairs of basal secondary veins (scale bar equals 2 mm) (3). 4-5, UF15706-30154, specimen showing a long petiole and ocrea-like structure at the base (scale bar equals 5 mm) (4) and enlargement of the petiole to show the ocrea-like structure (scale bar equals 2 mm) (5). 6, UF15706-14832, specimen showing suprabasal actinodromous primary venation and acute apices of the lobes. Scale bar equals 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11062841" httpUri="https://zenodo.org/record/11062841/files/figure.png" pageId="11" pageNumber="12">Figure 7.6</figureCitation>
) to rounded (
<figureCitation id="131127B1FF98AA2FFBD8FAE7FB6B325F" box="[1052,1184,1289,1314]" captionStart="FIGURE 8" captionStartId="13.[197,288,1698,1721]" captionTargetBox="[199,1434,168,1648]" captionTargetId="figure-19@13.[199,1434,167,1652]" captionTargetPageId="13" captionText="FIGURE 8. Pabiania variloba Upchurch and Dilcher, 1990. 1, UF15706-14823, leaf showing basal actinodromous primary venation and basal secondary veins. Scale bar equals 1 cm. 2, UF15706-24464, leaf showing rounded lobe apex. Scale bar equals 1 cm. 3, UF15706-24587, specimen showing a small lobe on the left and entire margin on the right of the leaf. Scale bar equals 5 mm. 4, Enlargement of Figure 8.2 to show straight primary vein extending to lobe apex and two series of loops in the excostal region. Scale bar equals 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11062843" httpUri="https://zenodo.org/record/11062843/files/figure.png" pageId="11" pageNumber="12">Figure 8.1</figureCitation>
); (2) apex of lamina lobes from acute (
<figureCitation id="131127B1FF98AA2FFBF1FAC7FB7C323F" box="[1077,1207,1321,1346]" captionStart="FIGURE 7" captionStartId="12.[197,288,1708,1731]" captionTargetBox="[240,1392,165,1679]" captionTargetId="figure-13@12.[240,1392,165,1679]" captionTargetPageId="12" captionText="FIGURE 7. Pabiania variloba Upchurch and Dilcher, 1990. 1-3, UF15706-24423, specimen showing suprabasal actinodromous primary venation (scale bar equals 1 cm) (1), enlargement of a sinus area to show sinus bracing by secondary veins (scale bar equals 2 mm) (2); and enlargement of basal portion of leaf to show two pairs of basal secondary veins (scale bar equals 2 mm) (3). 4-5, UF15706-30154, specimen showing a long petiole and ocrea-like structure at the base (scale bar equals 5 mm) (4) and enlargement of the petiole to show the ocrea-like structure (scale bar equals 2 mm) (5). 6, UF15706-14832, specimen showing suprabasal actinodromous primary venation and acute apices of the lobes. Scale bar equals 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11062841" httpUri="https://zenodo.org/record/11062841/files/figure.png" pageId="11" pageNumber="12">Figure 7.6</figureCitation>
), rounded (
<figureCitation id="131127B1FF98AA2FFA96FAC7FCBA321F" captionStart="FIGURE 7" captionStartId="12.[197,288,1708,1731]" captionTargetBox="[240,1392,165,1679]" captionTargetId="figure-13@12.[240,1392,165,1679]" captionTargetPageId="12" captionText="FIGURE 7. Pabiania variloba Upchurch and Dilcher, 1990. 1-3, UF15706-24423, specimen showing suprabasal actinodromous primary venation (scale bar equals 1 cm) (1), enlargement of a sinus area to show sinus bracing by secondary veins (scale bar equals 2 mm) (2); and enlargement of basal portion of leaf to show two pairs of basal secondary veins (scale bar equals 2 mm) (3). 4-5, UF15706-30154, specimen showing a long petiole and ocrea-like structure at the base (scale bar equals 5 mm) (4) and enlargement of the petiole to show the ocrea-like structure (scale bar equals 2 mm) (5). 6, UF15706-14832, specimen showing suprabasal actinodromous primary venation and acute apices of the lobes. Scale bar equals 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11062841" httpUri="https://zenodo.org/record/11062841/files/figure.png" pageId="11" pageNumber="12">Figure 7.1</figureCitation>
), to mucronate (
<figureCitation id="131127B1FF98AA2FFBF7FAA7FB7E321F" box="[1075,1205,1353,1378]" captionStart="FIGURE 8" captionStartId="13.[197,288,1698,1721]" captionTargetBox="[199,1434,168,1648]" captionTargetId="figure-19@13.[199,1434,167,1652]" captionTargetPageId="13" captionText="FIGURE 8. Pabiania variloba Upchurch and Dilcher, 1990. 1, UF15706-14823, leaf showing basal actinodromous primary venation and basal secondary veins. Scale bar equals 1 cm. 2, UF15706-24464, leaf showing rounded lobe apex. Scale bar equals 1 cm. 3, UF15706-24587, specimen showing a small lobe on the left and entire margin on the right of the leaf. Scale bar equals 5 mm. 4, Enlargement of Figure 8.2 to show straight primary vein extending to lobe apex and two series of loops in the excostal region. Scale bar equals 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11062843" httpUri="https://zenodo.org/record/11062843/files/figure.png" pageId="11" pageNumber="12">Figure 8.2</figureCitation>
). All other leaf morphological characters are consistent with those described by Upchurch and Dilcher (1990).
</paragraph>
<paragraph id="8B953B34FF98AA2FFC46FA47FAEA31DF" blockId="11.[840,1441,168,1698]" pageId="11" pageNumber="12">
Heer (1883, plate 38, figure 3) and Lesquereux (1892, plate 54, figures 1-3) described a few specimens with similar leaf morphology and assigned them to
<taxonomicName id="4C2A40B7FF98AA2FFBE5F9E7FC63313F" authority="Heer. Unfortunately" authorityName="Heer. Unfortunately" authorityYear="1882" class="Magnoliopsida" family="Araliaceae" genus="Aralia" kingdom="Plantae" order="Apiales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="groenlandica">
<emphasis id="B95EE726FF98AA2FFBE5F9E7FAC8315F" box="[1057,1283,1545,1570]" italics="true" pageId="11" pageNumber="12">Aralia groenlandica</emphasis>
Heer. Unfortunately
</taxonomicName>
, only primary and secondary venation are observed on these leaves, which makes it difficult to compare them with the specimens from the Rose Creek and Hoisington III localities.
</paragraph>
</subSubSection>
</treatment>
</document>