168 lines
20 KiB
XML
168 lines
20 KiB
XML
<document id="7500FDFEA5DBF06EBC4F6DEC8D303330" ID-DOI="10.26879/841" ID-ISSN="1094-8074" ID-Zenodo-Dep="11062827" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1714011148597" checkinUser="felipe" docAuthor="Wang, Hongshan & Dilcher, David L." docDate="2018" docId="03838A22FF8EAA39FEECFBFDFBCD315C" docLanguage="en" docName="PalaeontolElectron.21.3.34A.pdf" docOrigin="Palaeontologia Electronica (34 A) 21 (3)" docSource="http://dx.doi.org/10.26879/841" docStyle="DocumentStyle:322FC57CC99045555090E8A84CFB029A.4:PalaeontolElectron.2018-.journal_article" docStyleId="322FC57CC99045555090E8A84CFB029A" docStyleName="PalaeontolElectron.2018-.journal_article" docStyleVersion="4" docTitle="Anisodromum upchurchii Wang & Dilcher 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="30" masterDocId="FFBAF25AFF93AA24FFC4FFEEFFCB377D" masterDocTitle="Early Cretaceous angiosperm leaves from the Dakota Formation, Hoisington III locality, Kansas, USA" masterLastPageNumber="49" masterPageNumber="1" pageNumber="30" updateTime="1715622024402" updateUser="carolina" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="DCB1CF85495B6FA4567C9AF9AB724477">Early Cretaceous angiosperm leaves from the Dakota Formation, Hoisington III locality, Kansas, USA</mods:title>
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<mods:name id="DDB21CF8DB94FC83F624F3F2A117F690" type="personal">
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<mods:namePart id="3764ADE544ECEEB0AC2DBF852E1A3B2B">Wang, Hongshan</mods:namePart>
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<mods:name id="FF448870BA24FE68F3596137BA826236" type="personal">
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<mods:namePart id="E401BF8A1211857972FFB84996110C77">Dilcher, David L.</mods:namePart>
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<mods:typeOfResource id="5981014A7F710D8856B11E0A88CF31F3">text</mods:typeOfResource>
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<mods:titleInfo id="03C7087CC46B62EBE43887E2CCDE8BE3">
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<mods:title id="3FF675B1E9504A4F144A8425818BFD28">Palaeontologia Electronica</mods:title>
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<mods:part id="B644C02119982B29C1359CA169EEBA18">
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<mods:date id="11373885C6DC3BE0052187E17B26E3E6">2018</mods:date>
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<mods:detail id="3EC77427E202E072961A2F93BCB51813" type="series">
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<mods:title id="BC3211168382F4603C514A7EC75AA02B">34 A</mods:title>
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<mods:detail id="D35DA31F555A4193FFB4520CD56776C0" type="pubDate">
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<mods:number id="CC9DDC4D6E3110602354016F60AE6DEF">2018-09-30</mods:number>
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<mods:number id="22BDA089A9F7F9989F7F70EF9FBDB087">21</mods:number>
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<mods:number id="CB3E9945EF9C06346234C545FCFB5FB8">3</mods:number>
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<mods:start id="B154B07C23A74587E4A271FDFA221465">1</mods:start>
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<mods:url id="F447D5AFF4E8EF2BE3F5ACE4AEF3B041">http://dx.doi.org/10.26879/841</mods:url>
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<mods:classification id="89012B9C7F5509403420F4C77FF98912">journal article</mods:classification>
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<mods:identifier id="1C78241F57FB50F18A9D8078B51FD19F" type="DOI">10.26879/841</mods:identifier>
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<mods:identifier id="61AE455553974BBDDA9BB7CCFF4261B6" type="ISSN">1094-8074</mods:identifier>
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<mods:identifier id="E81A093F6C35EB84C598E020E6F54163" type="Zenodo-Dep">11062827</mods:identifier>
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</mods:mods>
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<treatment id="03838A22FF8EAA39FEECFBFDFBCD315C" LSID="urn:lsid:plazi:treatment:03838A22FF8EAA39FEECFBFDFBCD315C" httpUri="http://treatment.plazi.org/id/03838A22FF8EAA39FEECFBFDFBCD315C" lastPageNumber="30" pageId="29" pageNumber="30">
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<subSubSection id="C33068BFFF8EAA39FEECFBFDFD7A3351" box="[296,689,1043,1068]" pageId="29" pageNumber="30" type="nomenclature">
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<paragraph id="8B953B34FF8EAA39FEECFBFDFD7A3351" blockId="29.[296,689,1043,1100]" box="[296,689,1043,1068]" pageId="29" pageNumber="30">
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<heading id="D0DD8C58FF8EAA39FEECFBFDFD7A3351" box="[296,689,1043,1068]" centered="true" fontSize="10" level="2" pageId="29" pageNumber="30" reason="7">
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<taxonomicName id="4C2A40B7FF8EAA39FEECFBFDFD863351" authority="Wang & Dilcher, 2018" authorityName="Wang & Dilcher" authorityYear="2018" box="[296,589,1043,1068]" class="Magnoliopsida" family="Rosidae" genus="Anisodromum" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="upchurchii" status="sp. nov.">
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<emphasis id="B95EE726FF8EAA39FEECFBFDFD863351" box="[296,589,1043,1068]" italics="true" pageId="29" pageNumber="30">Anisodromum upchurchii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26D5A5DFF8EAA39FD91FBFDFD7A3351" box="[597,689,1043,1068]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33068BFFF8EAA39FE59FBDDFDF03331" box="[413,571,1075,1100]" pageId="29" pageNumber="30" type="description">
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<paragraph id="8B953B34FF8EAA39FE59FBDDFDF03331" blockId="29.[296,689,1043,1100]" box="[413,571,1075,1100]" pageId="29" pageNumber="30">
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<figureCitation id="131127B1FF8EAA39FE59FBDDFDF03331" box="[413,571,1075,1100]" captionStart="FIGURE 19. 1-2" captionStartId="30.[192,283,1458,1481]" captionTargetBox="[312,1317,162,1425]" captionTargetId="figure-105@30.[312,1317,162,1425]" captionTargetPageId="30" captionText="FIGURE 19. 1-2, Anisodromum upchurchii sp. nov., UF15706-24576, middle portion of lamina showing strong primary and secondary veins (scale bar equals 1 cm) (1) and enlargement of an area in Figure 19.1 (indicated by arrow) to show percurrent tertiary veins (scale bar equals 2 mm) (2). 3-4, Anisodromum schimperi (Lesquereux) comb. nov., UF15706-24635, leaf showing uneven spacing of secondary veins and reticulate tertiary veins (scale bar equals 5 mm) (3) and; UF15706-24633, leaf showing secondary venation and reticulate tertiary veins (scale bar equals 5 mm) (4)." figureDoi="http://doi.org/10.5281/zenodo.11062865" httpUri="https://zenodo.org/record/11062865/files/figure.png" pageId="29" pageNumber="30">Figure 19.1-2</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33068BFFF8EAA39FF04FBB0FD1432AA" pageId="29" pageNumber="30" type="diagnosis">
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<paragraph id="8B953B34FF8EAA39FF04FBB0FD1432AA" blockId="29.[192,793,1118,1911]" pageId="29" pageNumber="30">
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<emphasis id="B95EE726FF8EAA39FF04FBB0FE73330A" bold="true" box="[192,440,1118,1144]" pageId="29" pageNumber="30">Specific diagnosis.</emphasis>
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Lamina base acute, decurrent. Margin entire. Primary venation pinnate; primary vein stout, multi-stranded, straight. Secondary venation brochidodromous, secondary veins moderate relative to primary vein, opposite, subopposite or alternate; angle of divergence wide acute, uniformly curved apically; secondary vein spacing distant. Intersecondary veins common, usually opposite to a secondary vein on the other side of lamina. Tertiary veins moderate relative to secondary veins, percurrent, course straight, oriented almost at right angles with primary vein.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33068BFFF8EAA39FF04FA30FACD357F" pageId="29" pageNumber="30" type="description">
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<paragraph id="8B953B34FF8EAA39FF04FA30FB23369F" blockId="29.[192,793,1118,1911]" lastBlockId="29.[840,1441,168,1569]" pageId="29" pageNumber="30">
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<emphasis id="B95EE726FF8EAA39FF04FA30FE91328A" bold="true" box="[192,346,1502,1527]" pageId="29" pageNumber="30">Description.</emphasis>
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Incomplete leaves observed. Apex missing. Base acute, decurrent. Margin entire. Primary venation pinnate; primary vein stout, multi-stranded, straight. Secondary venation brochidodromous, secondary veins moderate relative to primary vein, ca. six pairs per lamina, opposite, subopposite or alternate; angle of divergence wide acute, with lowest pair more acute than pairs above, uniformly curved apically; secondary vein spacing distant (distance between veins
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<quantity id="4CD296D1FF8EAA39FD71F910FF23304A" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="29" pageNumber="30" unit="cm" value="1.25" valueMax="1.5" valueMin="1.0">1 to 1.5 cm</quantity>
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. Intersecondary veins common, one or two per intercostal region; extending less than one half distance of half lamina, usually opposite to a secondary vein on the other side of lamina. Tertiary veins moderate relative to secondary veins, percurrent, course straight; angle of origin
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<collectionCode id="ED3BA3F1FF8EAA39FB77FF06FB12367F" box="[1203,1241,232,258]" country="Italy" lsid="urn:lsid:biocol.org:col:15824" name="Museo Regionale di Scienze Naturali della Valle d'Aosta" pageId="29" pageNumber="30" type="Herbarium">AO</collectionCode>
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(acute on exmedial side and obtuse on admedial side of secondary veins or intersecondary vein) or
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<collectionCode id="ED3BA3F1FF8EAA39FB12FEC6FB35363F" box="[1238,1278,296,322]" country="Italy" lsid="urn:lsid:biocol.org:col:15061" name="Università degli Studi di Roma La Sapienza" pageId="29" pageNumber="30" type="Herbarium">RO</collectionCode>
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(right on primary vein and obtuse on admedial side of secondary or intersecondary veins), primary vein-tertiary vein angle almost right (90º); tertiary vein interval close (interval between veins less than
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<quantity id="4CD296D1FF8EAA39FAFEFE47FA5E36BF" box="[1338,1429,425,450]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="29" pageNumber="30" unit="cm" value="0.5">0.5 cm</quantity>
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). Veins of higher order not observed.
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</paragraph>
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<paragraph id="8B953B34FF8EAA39FC8CFE07FACD357F" blockId="29.[840,1441,168,1569]" box="[840,1286,489,514]" pageId="29" pageNumber="30">
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<emphasis id="B95EE726FF8EAA39FC8CFE07FB23357F" bold="true" box="[840,1256,489,514]" pageId="29" pageNumber="30">Number of specimens examined.</emphasis>
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4.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33068BFFF8EAA39FC8CFDE7FA8735FF" pageId="29" pageNumber="30" type="materials_examined">
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<paragraph id="8B953B34FF8EAA39FC8CFDE7FA82355F" blockId="29.[840,1441,168,1569]" box="[840,1353,521,546]" pageId="29" pageNumber="30">
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<materialsCitation id="3B423169FF8EAA39FC8CFDE7FA82355F" box="[840,1353,521,546]" collectionCode="UF" pageId="29" pageNumber="30" specimenCode="UF15706-24576" specimenCount="1" typeStatus="holotype">
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<emphasis id="B95EE726FF8EAA39FC8CFDE7FC0A355F" bold="true" box="[840,961,521,546]" pageId="29" pageNumber="30">
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<typeStatus id="54918596FF8EAA39FC8CFDE7FC76355F" box="[840,957,521,546]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCode id="DB8C934FFF8EAA39FC0DFDE7FB41355F" box="[969,1162,521,546]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="29" pageNumber="30" type="Museum">UF15706-24576</specimenCode>
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(
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<figureCitation id="131127B1FF8EAA39FB53FDE7FAF5355F" box="[1175,1342,521,546]" captionStart="FIGURE 19. 1-2" captionStartId="30.[192,283,1458,1481]" captionTargetBox="[312,1317,162,1425]" captionTargetId="figure-105@30.[312,1317,162,1425]" captionTargetPageId="30" captionText="FIGURE 19. 1-2, Anisodromum upchurchii sp. nov., UF15706-24576, middle portion of lamina showing strong primary and secondary veins (scale bar equals 1 cm) (1) and enlargement of an area in Figure 19.1 (indicated by arrow) to show percurrent tertiary veins (scale bar equals 2 mm) (2). 3-4, Anisodromum schimperi (Lesquereux) comb. nov., UF15706-24635, leaf showing uneven spacing of secondary veins and reticulate tertiary veins (scale bar equals 5 mm) (3) and; UF15706-24633, leaf showing secondary venation and reticulate tertiary veins (scale bar equals 5 mm) (4)." figureDoi="http://doi.org/10.5281/zenodo.11062865" httpUri="https://zenodo.org/record/11062865/files/figure.png" pageId="29" pageNumber="30">Figure 19.1-2</figureCitation>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8B953B34FF8EAA39FC8CFDC7FA8735FF" blockId="29.[840,1441,168,1569]" pageId="29" pageNumber="30">
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<emphasis id="B95EE726FF8EAA39FC8CFDC7FBA8353F" bold="true" box="[840,1123,553,578]" pageId="29" pageNumber="30">Derivation of epithet.</emphasis>
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In honor of Garland
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<collectionCode id="ED3BA3F1FF8EAA39FA41FDC7FA57353F" box="[1413,1436,553,578]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="29" pageNumber="30">R</collectionCode>
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. Upchurch in recognition of his contributions to Dakota Formation angiosperm paleobotany.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33068BFFF8EAA39FC8CFD67FBCD315C" pageId="29" pageNumber="30" type="discussion">
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<paragraph id="8B953B34FF8EAA39FC8CFD67FB9B337F" blockId="29.[840,1441,168,1569]" pageId="29" pageNumber="30">
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<emphasis id="B95EE726FF8EAA39FC8CFD67FC0B35DF" bold="true" box="[840,960,649,674]" pageId="29" pageNumber="30">Remarks.</emphasis>
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<taxonomicName id="4C2A40B7FF8EAA39FC15FD67FACB35DF" authorityName="Wang & Dilcher" authorityYear="2018" box="[977,1280,649,674]" class="Magnoliopsida" genus="Anisodromum" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="upchurchii">
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<emphasis id="B95EE726FF8EAA39FC15FD67FACB35DF" box="[977,1280,649,674]" italics="true" pageId="29" pageNumber="30">Anisodromum upchurchii</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName id="4C2A40B7FF8EAA39FC8CFD47FBF035BC" authorityName="Upchurch and Dilcher" authorityYear="1990" box="[840,1083,680,706]" class="Magnoliopsida" genus="Anisodromum" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="wolfei">
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<emphasis id="B95EE726FF8EAA39FC8CFD47FBF035BC" box="[840,1083,680,706]" italics="true" pageId="29" pageNumber="30">Anisodromum wolfei</emphasis>
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</taxonomicName>
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in that it has large angles of divergence for secondary veins, large distance between secondary veins, fewer secondary veins, and intersecondary veins arranged opposite to secondary vein on the opposite side of the lamina.
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<taxonomicName id="4C2A40B7FF8EAA39FC8CFCA7FBA7341F" authorityName="Wang & Dilcher" authorityYear="2018" box="[840,1132,841,866]" class="Magnoliopsida" genus="Anisodromum" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="upchurchii">
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<emphasis id="B95EE726FF8EAA39FC8CFCA7FBA7341F" box="[840,1132,841,866]" italics="true" pageId="29" pageNumber="30">Anisodromum upchurchii</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C2A40B7FF8EAA39FB63FCA7FAF2341F" authorityName="Wang & Dilcher" authorityYear="2018" baseAuthorityName="Lesquereux" box="[1191,1337,841,866]" class="Magnoliopsida" genus="Anisodromum" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="schimperi">
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<emphasis id="B95EE726FF8EAA39FB63FCA7FAF2341F" box="[1191,1337,841,866]" italics="true" pageId="29" pageNumber="30">A. schimperi</emphasis>
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</taxonomicName>
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are similar in that they both have irregularly spaced secondary veins. They differ in that the secondary venation of
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<taxonomicName id="4C2A40B7FF8EAA39FC10FC47FBA334BF" authorityName="Wang & Dilcher" authorityYear="2018" baseAuthorityName="Lesquereux" box="[980,1128,937,962]" class="Magnoliopsida" genus="Anisodromum" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="schimperi">
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<emphasis id="B95EE726FF8EAA39FC10FC47FBA334BF" box="[980,1128,937,962]" italics="true" pageId="29" pageNumber="30">A. schimperi</emphasis>
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</taxonomicName>
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is festooned brochidodromous and its tertiary veins are orthogonally reticulate near primary vein.
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</paragraph>
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<paragraph id="8B953B34FF8EAA39FC46FBE7FADF323F" blockId="29.[840,1441,168,1569]" pageId="29" pageNumber="30">
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The wide divergence angle of the secondary veins and the primary-tertiary vein angle of
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<taxonomicName id="4C2A40B7FF8EAA39FAB5FBC7FB85331F" authorityName="Wang & Dilcher" authorityYear="2018" class="Magnoliopsida" genus="Anisodromum" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="upchurchii">
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<emphasis id="B95EE726FF8EAA39FAB5FBC7FB85331F" italics="true" pageId="29" pageNumber="30">Anisodromum upchurchii</emphasis>
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</taxonomicName>
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are similar to Morphotype
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<collectionCode id="ED3BA3F1FF8EAA39FC8CFB86FCBA33FF" box="[840,881,1128,1154]" country="China" name="Chongqing Museum" pageId="29" pageNumber="30">CM</collectionCode>
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05 (
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<taxonomicName id="4C2A40B7FF8EAA39FC65FB87FB6D33FF" box="[929,1190,1129,1154]" class="Magnoliopsida" family="Magnoliaceae" genus="Magnoliaephyllum" kingdom="Plantae" order="Magnoliales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="undetermined">
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<emphasis id="B95EE726FF8EAA39FC65FB87FBB333FF" box="[929,1144,1129,1154]" italics="true" pageId="29" pageNumber="30">Magnoliaephyllum</emphasis>
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sp.
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</taxonomicName>
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) from the mid-Cretaceous Cedar Mountain Formation of
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<collectingRegion id="49EEF5D6FF8EAA39FADFFB67FA9C33DF" box="[1307,1367,1161,1186]" country="United States of America" name="Utah" pageId="29" pageNumber="30">Utah</collectingRegion>
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,
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<collectingCountry id="F33D7BA4FF8EAA39FAADFB66FA6B33DF" box="[1385,1440,1160,1186]" name="United States of America" pageId="29" pageNumber="30">USA</collectingCountry>
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(Harris and Arens, 2016, p. 646-648, figure 4.1-3). They differ in that the divergence angle of secondary veins is larger and the percurrent tertiary veins is less dense in
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<taxonomicName id="4C2A40B7FF8EAA39FBE3FAE6FA9F325C" authorityName="Wang & Dilcher" authorityYear="2018" box="[1063,1364,1288,1313]" class="Magnoliopsida" genus="Anisodromum" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="upchurchii">
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<emphasis id="B95EE726FF8EAA39FBE3FAE6FA9F325C" box="[1063,1364,1288,1313]" italics="true" pageId="29" pageNumber="30">Anisodromum upchurchii</emphasis>
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</taxonomicName>
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compared with those of Morphotype
|
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<collectionCode id="ED3BA3F1FF8EAA39FB01FAC6FB3A323F" box="[1221,1265,1320,1346]" country="China" name="Chongqing Museum" pageId="29" pageNumber="30">CM</collectionCode>
|
|
05.
|
|
</paragraph>
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|
<paragraph id="8B953B34FF8EAA39FC46FAA6FBCD315C" blockId="29.[840,1441,168,1569]" pageId="29" pageNumber="30">
|
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It is unknown if the specimens from the Hoisington III locality represent simple or compound leaves. Therefore, we tentatively assign these specimens to the genus
|
|
<taxonomicName id="4C2A40B7FF8EAA39FB30FA46FA5332BC" authorityName="Upchurch and Dilcher" authorityYear="1990" box="[1268,1432,1448,1473]" class="Magnoliopsida" genus="Anisodromum" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="B95EE726FF8EAA39FB30FA46FA5332BC" box="[1268,1432,1448,1473]" italics="true" pageId="29" pageNumber="30">Anisodromum</emphasis>
|
|
</taxonomicName>
|
|
. Future collection with better preserved specimens may confirm this assignment or help separate them from this genus.
|
|
</paragraph>
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|
</subSubSection>
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</treatment>
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</document> |