500 lines
62 KiB
XML
500 lines
62 KiB
XML
<document id="58C53B66A3EBE6CA78900B9B1D2BFBEF" ID-DOI="10.11646/zootaxa.3925.3.9" ID-GBIF-Dataset="5ca114a0-2d44-4298-8d84-c9dde1f79cb9" ID-ISSN="1175-5326" ID-Zenodo-Dep="242277" ID-ZooBank="09962D19-6BDF-4E06-B9B1-2760DDD3236E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461004113901" checkinUser="plazi" docAuthor="Collado, Gonzalo A." docDate="2015" docId="038387E8BC01FFB36DB97CFFFE7C9B6D" docLanguage="en" docName="zt03925p449.pdf" docOrigin="Zootaxa 3925 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heleobia deserticola Collado, 2015, sp. nov." docType="treatment" docVersion="7" lastPageNumber="446" masterDocId="FFBAFF90BC01FFB16D2E7A62FFEA9A7A" masterDocTitle="A new freshwater snail (Caenogastropoda: Cochliopidae) from the Atacama Desert, northern Chile" masterLastPageNumber="449" masterPageNumber="445" pageNumber="445" updateTime="1698565191597" updateUser="plazi">
|
||
<mods:mods id="541B48D1054ACBB727CF4F7B484A1672" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="66A542E7630BD9D3054B87FE0ECF4F67">
|
||
<mods:title id="861D6BC0974A88B8BBD89F7CEE1F613F">A new freshwater snail (Caenogastropoda: Cochliopidae) from the Atacama Desert, northern Chile</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="C7BB3E5E831A6BC2ABF9991F169B4B45" type="personal">
|
||
<mods:role id="094E234557DBDAC11E25488C51F6EBFC">
|
||
<mods:roleTerm id="8AE813B42E25504ED85D37AAF8AF66DF">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1629ABA20632EED3E7A9F80288A83944">Collado, Gonzalo A.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="F79407A76545B3247E96ACE28DE5C25B">text</mods:typeOfResource>
|
||
<mods:relatedItem id="71A18E28F8BE07E320EC41E4D7D6B2BA" type="host">
|
||
<mods:titleInfo id="DC658D7944DAE12F5439EF7826C44FF0">
|
||
<mods:title id="580A6ABD6A5D4432B3F16591962FB8F5">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="D37DF7C77B9C229E972C32A40A4E6099">
|
||
<mods:date id="2BF0683FAE9BBE386D55C384CFF6C32D">2015</mods:date>
|
||
<mods:detail id="553048C2F74A5CE99645A6DB6F8FF506" type="volume">
|
||
<mods:number id="CDE3CCD33A06F20CC1A495406CE907AD">3925</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="1E28C2CA53EA85DED0911D314F116BC8" type="issue">
|
||
<mods:number id="C23B7F13AB9B9051CBDCB866D479660D">3</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="AE051A72DA9A30ADC05B7E58A91865D5" unit="page">
|
||
<mods:start id="89CE711E444127A93A2A9DE14D508C28">445</mods:start>
|
||
<mods:end id="E3686AE21CFBBE02A6D9464A1DBBFA3C">449</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="D2943695795E4DAA1DD5AE4595368EDD">journal article</mods:classification>
|
||
<mods:identifier id="FA754DA37A2DF818ED98758FF3C89229" type="DOI">10.11646/zootaxa.3925.3.9</mods:identifier>
|
||
<mods:identifier id="654B46708F793057D365B4456E3D6F93" type="GBIF-Dataset">5ca114a0-2d44-4298-8d84-c9dde1f79cb9</mods:identifier>
|
||
<mods:identifier id="72D2B6CA1A5A23D5F29BA92BEB86062A" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="F504A9C169F8B91C77F51E5A1E4E477D" type="Zenodo-Dep">242277</mods:identifier>
|
||
<mods:identifier id="A20A768B2DE1CFEA54264A337A43D5B5" type="ZooBank">09962D19-6BDF-4E06-B9B1-2760DDD3236E</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="038387E8BC01FFB36DB97CFFFE7C9B6D" ID-DOI="http://doi.org/10.5281/zenodo.6094942" ID-GBIF-Taxon="119598809" ID-Zenodo-Dep="6094942" LSID="urn:lsid:plazi:treatment:038387E8BC01FFB36DB97CFFFE7C9B6D" httpUri="http://treatment.plazi.org/id/038387E8BC01FFB36DB97CFFFE7C9B6D" lastPageId="2" lastPageNumber="446" pageId="0" pageNumber="445">
|
||
<subSubSection id="C3306575BC01FFB16DB97CFFFDBF9CCD" box="[151,597,1693,1719]" pageId="0" pageNumber="445" type="nomenclature">
|
||
<paragraph id="8B9536FEBC01FFB16DB97CFFFDBF9CCD" blockId="0.[151,597,1693,1719]" box="[151,597,1693,1719]" pageId="0" pageNumber="445">
|
||
<heading id="D0DD8192BC01FFB16DB97CFFFDBF9CCD" bold="true" box="[151,597,1693,1719]" fontSize="11" level="1" pageId="0" pageNumber="445" reason="1">
|
||
<emphasis id="B95EEAECBC01FFB16DB97CFFFDBF9CCD" bold="true" box="[151,597,1693,1719]" pageId="0" pageNumber="445">
|
||
Species
|
||
<taxonomicName id="4C2A4D7DBC01FFB16DD77CFFFE1A9CCD" ID-CoL="3K4QL" box="[249,496,1693,1719]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="0" pageNumber="445" phylum="Mollusca" rank="species" species="deserticola" status="sp. nov.">
|
||
<emphasis id="B95EEAECBC01FFB16DD77CFFFE1A9CCD" bold="true" box="[249,496,1693,1719]" italics="true" pageId="0" pageNumber="445">Heleobia deserticola</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26D5797BC01FFB16CD97CFFFDBF9CCD" box="[503,597,1693,1719]" pageId="0" pageNumber="445" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3306575BC01FFB16DB97C80FD579DDD" pageId="0" pageNumber="445" type="materials_examined">
|
||
<paragraph id="8B9536FEBC01FFB16DB97C80FBEB9D66" blockId="0.[151,1436,1762,2028]" pageId="0" pageNumber="445">
|
||
<emphasis id="B95EEAECBC01FFB16DB97C80FEEF9C83" bold="true" box="[151,261,1762,1785]" pageId="0" pageNumber="445">
|
||
<typeStatus id="5491885CBC01FFB16DB97C80FEEB9C83" box="[151,257,1762,1785]" pageId="0" pageNumber="445" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
(
|
||
<collectionCode id="ED3BAE3BBC01FFB16C3D7C80FE889C83" box="[275,354,1762,1785]" httpUri="http://grbio.org/cool/70mh-t5r1" name="Universita di Cagliari" pageId="0" pageNumber="445">MZUC</collectionCode>
|
||
43067,
|
||
<figureCitation id="13112A7BBC01FFB16C947C80FDE49C83" box="[442,526,1762,1785]" captionStart="FIGURE 1. A – Q" captionStartId="2.[151,255,1751,1774]" captionTargetBox="[151,1436,333,1726]" captionTargetId="figure@2.[151,1436,333,1728]" captionTargetPageId="2" captionText="FIGURE 1. A – Q. Heleobia deserticola sp. nov. A. Shell of the holotype (MZUC 43067). B – K. Shells of the paratypes (MZUC 43068 – 43077). L. Operculum (1.70 mm total length). M – O. Penes of three individuals (0.64, 0.70 and 0.90 mm total length, respectively). Operculum and penes were photographed from both sides. P, Q. Egg capsules of the new species. P. Egg capsules attached to the adult shells. Q. Pre – hatching capsule isolated with a shelled juvenile inside. R. Shell of an adult snail from Carrera Pinto (SL 3.51 mm, SW 1.55 mm, AL 1.35 mm, AW 0.97 mm. S. Penis of this individual (0.75 mm total length)." httpUri="https://zenodo.org/record/242278/files/figure.png" pageId="0" pageNumber="445">Fig. 1A</figureCitation>
|
||
). Collected by G.A. Collado from Aguada de Chorrillos,
|
||
<collectingCountry id="F33D766EBC01FFB1695A7C80FB449C83" box="[1140,1198,1762,1785]" name="Chile" pageId="0" pageNumber="445">Chile</collectingCountry>
|
||
(
|
||
<date id="FF94103EBC01FFB169937C80FA7B9C83" box="[1213,1425,1762,1785]" pageId="0" pageNumber="445" value="2011-11-28">28 November 2011</date>
|
||
).
|
||
<typeStatus id="5491885CBC01FFB16DB97D67FF109D66" box="[151,250,1797,1820]" pageId="0" pageNumber="445" type="holotype">Holotype</typeStatus>
|
||
measurements: SL
|
||
<quantity id="4CD29B1BBC01FFB16CE37D67FDC49D66" box="[461,558,1797,1820]" metricMagnitude="-3" metricUnit="m" metricValue="4.41" pageId="0" pageNumber="445" unit="mm" value="4.41">4.41 mm</quantity>
|
||
, SW
|
||
<quantity id="4CD29B1BBC01FFB16F467D67FD239D66" box="[616,713,1797,1820]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="0" pageNumber="445" unit="mm" value="2.1">2.10 mm</quantity>
|
||
, AL 2.00 mm, AW
|
||
<quantity id="4CD29B1BBC01FFB16EB57D67FC179D66" box="[923,1021,1797,1820]" metricMagnitude="-3" metricUnit="m" metricValue="1.21" pageId="0" pageNumber="445" unit="mm" value="1.21">1.21 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9536FEBC01FFB16DE97D4AFD579DDD" blockId="0.[151,1436,1762,2028]" pageId="0" pageNumber="445">
|
||
<emphasis id="B95EEAECBC01FFB16DE97D4AFEAA9D45" bold="true" box="[199,320,1832,1855]" pageId="0" pageNumber="445">
|
||
<typeStatus id="5491885CBC01FFB16DE97D4AFED19D45" box="[199,315,1832,1855]" pageId="0" pageNumber="445" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
(
|
||
<collectionCode id="ED3BAE3BBC01FFB16C7A7D4AFE499D45" box="[340,419,1832,1855]" httpUri="http://grbio.org/cool/70mh-t5r1" name="Universita di Cagliari" pageId="0" pageNumber="445">MZUC</collectionCode>
|
||
43068–43077,
|
||
<figureCitation id="13112A7BBC01FFB16F797D4AFD389D45" box="[599,722,1832,1855]" captionStart="FIGURE 1. A – Q" captionStartId="2.[151,255,1751,1774]" captionTargetBox="[151,1436,333,1726]" captionTargetId="figure@2.[151,1436,333,1728]" captionTargetPageId="2" captionText="FIGURE 1. A – Q. Heleobia deserticola sp. nov. A. Shell of the holotype (MZUC 43067). B – K. Shells of the paratypes (MZUC 43068 – 43077). L. Operculum (1.70 mm total length). M – O. Penes of three individuals (0.64, 0.70 and 0.90 mm total length, respectively). Operculum and penes were photographed from both sides. P, Q. Egg capsules of the new species. P. Egg capsules attached to the adult shells. Q. Pre – hatching capsule isolated with a shelled juvenile inside. R. Shell of an adult snail from Carrera Pinto (SL 3.51 mm, SW 1.55 mm, AL 1.35 mm, AW 0.97 mm. S. Penis of this individual (0.75 mm total length)." httpUri="https://zenodo.org/record/242278/files/figure.png" pageId="0" pageNumber="445">
|
||
Fig. 1B
|
||
<emphasis id="B95EEAECBC01FFB16F807D4AFD519D45" bold="true" box="[686,699,1832,1855]" pageId="0" pageNumber="445">–</emphasis>
|
||
K
|
||
</figureCitation>
|
||
). Snails from Aguada de Chorrillos,
|
||
<collectingCountry id="F33D766EBC01FFB169AD7D4AFB559D45" box="[1155,1215,1832,1855]" name="Chile" pageId="0" pageNumber="445">Chile</collectingCountry>
|
||
, collected with the
|
||
<typeStatus id="5491885CBC01FFB16DB97D28FF1E9D1B" box="[151,244,1866,1889]" pageId="0" pageNumber="445" type="holotype">holotype</typeStatus>
|
||
by G.A. Collado. Morphometric data of the
|
||
<typeStatus id="5491885CBC01FFB16FF87D28FCD69D1B" box="[726,828,1866,1889]" pageId="0" pageNumber="445" type="paratype">paratypes</typeStatus>
|
||
(n = 10): SL: 4.01 ± 0.31 (3.40
|
||
<emphasis id="B95EEAECBC01FFB169B87D28FB499D1B" bold="true" box="[1174,1187,1866,1889]" pageId="0" pageNumber="445">–</emphasis>
|
||
4.40); SW: 1.99 ± 0.13 (1.80
|
||
<emphasis id="B95EEAECBC01FFB16DE07D0FFF319DFE" bold="true" box="[206,219,1901,1924]" pageId="0" pageNumber="445">–</emphasis>
|
||
2.20); AL: 1.76 ± 0.19 (1.40
|
||
<emphasis id="B95EEAECBC01FFB16F277D0FFDFC9DFE" bold="true" box="[521,534,1901,1924]" pageId="0" pageNumber="445">–</emphasis>
|
||
2.00); AW: 1.22 ± 0.15 (1.00
|
||
<emphasis id="B95EEAECBC01FFB16E677D0FFCBC9DFE" bold="true" box="[841,854,1901,1924]" pageId="0" pageNumber="445">–</emphasis>
|
||
1.40); SW/SL: 0.50 ± 0.02 (0.46
|
||
<emphasis id="B95EEAECBC01FFB169807D0FFB519DFE" bold="true" box="[1198,1211,1901,1924]" pageId="0" pageNumber="445">–</emphasis>
|
||
0.53); AL/SL: 0.44 ± 0.04 (0.35
|
||
<emphasis id="B95EEAECBC01FFB16C2A7DF2FEFB9DDD" bold="true" box="[260,273,1936,1959]" pageId="0" pageNumber="445">–</emphasis>
|
||
0.47); AW/AL: 0.07 ± 0.08 (0.61
|
||
<emphasis id="B95EEAECBC01FFB16F5F7DF2FD949DDD" bold="true" box="[625,638,1936,1959]" pageId="0" pageNumber="445">–</emphasis>
|
||
0.86).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3306575BC01FFB06DE97DD0FDEF9B05" lastPageId="1" lastPageNumber="446" pageId="0" pageNumber="445" type="description">
|
||
<paragraph id="8B9536FEBC01FFB06DE97DD0FEA69B6D" blockId="0.[151,1436,1762,2028]" lastBlockId="1.[151,1437,152,2012]" lastPageId="1" lastPageNumber="446" pageId="0" pageNumber="445">
|
||
<emphasis id="B95EEAECBC01FFB16DE97DD0FEB89DB3" bold="true" box="[199,338,1970,1993]" pageId="0" pageNumber="445">Description.</emphasis>
|
||
Shell small, elongate
|
||
<emphasis id="B95EEAECBC01FFB16F107DD0FDA19DB3" bold="true" box="[574,587,1970,1993]" pageId="0" pageNumber="445">–</emphasis>
|
||
conic, thin, transparent, smooth, suture depth, closed umbilicus (imperforate). Adults with 6 to 7 convex whorls. Aperture ovate, outer lip thin. Operculum corneous, ovate, flat, thin, light brown, paucispiral (
|
||
<figureCitation id="13112A7BBC00FFB06C357AFAFE879AD5" box="[283,365,152,175]" captionStart="FIGURE 1. A – Q" captionStartId="2.[151,255,1751,1774]" captionTargetBox="[151,1436,333,1726]" captionTargetId="figure@2.[151,1436,333,1728]" captionTargetPageId="2" captionText="FIGURE 1. A – Q. Heleobia deserticola sp. nov. A. Shell of the holotype (MZUC 43067). B – K. Shells of the paratypes (MZUC 43068 – 43077). L. Operculum (1.70 mm total length). M – O. Penes of three individuals (0.64, 0.70 and 0.90 mm total length, respectively). Operculum and penes were photographed from both sides. P, Q. Egg capsules of the new species. P. Egg capsules attached to the adult shells. Q. Pre – hatching capsule isolated with a shelled juvenile inside. R. Shell of an adult snail from Carrera Pinto (SL 3.51 mm, SW 1.55 mm, AL 1.35 mm, AW 0.97 mm. S. Penis of this individual (0.75 mm total length)." httpUri="https://zenodo.org/record/242278/files/figure.png" pageId="1" pageNumber="446">Fig. 1L</figureCitation>
|
||
). Foot, mantle and tentacles gray, head black, snout white, visceral sac yellow to light brown. Penis elongated, white
|
||
<emphasis id="B95EEAECBC00FFB06C667AD8FEBF9AAB" bold="true" box="[328,341,186,209]" pageId="1" pageNumber="446">–</emphasis>
|
||
grayish; proximal portion with a wide base, with 5
|
||
<emphasis id="B95EEAECBC00FFB06E567AD8FC6F9AAB" bold="true" box="[888,901,186,209]" pageId="1" pageNumber="446">–</emphasis>
|
||
6 cup
|
||
<emphasis id="B95EEAECBC00FFB06EEF7AD8FC249AAB" bold="true" box="[961,974,186,209]" pageId="1" pageNumber="446">–</emphasis>
|
||
shaped apocrine glands on the border (
|
||
<figureCitation id="13112A7BBC00FFB0685E7AD8FF359A8E" captionStart="FIGURE 1. A – Q" captionStartId="2.[151,255,1751,1774]" captionTargetBox="[151,1436,333,1726]" captionTargetId="figure@2.[151,1436,333,1728]" captionTargetPageId="2" captionText="FIGURE 1. A – Q. Heleobia deserticola sp. nov. A. Shell of the holotype (MZUC 43067). B – K. Shells of the paratypes (MZUC 43068 – 43077). L. Operculum (1.70 mm total length). M – O. Penes of three individuals (0.64, 0.70 and 0.90 mm total length, respectively). Operculum and penes were photographed from both sides. P, Q. Egg capsules of the new species. P. Egg capsules attached to the adult shells. Q. Pre – hatching capsule isolated with a shelled juvenile inside. R. Shell of an adult snail from Carrera Pinto (SL 3.51 mm, SW 1.55 mm, AL 1.35 mm, AW 0.97 mm. S. Penis of this individual (0.75 mm total length)." httpUri="https://zenodo.org/record/242278/files/figure.png" pageId="1" pageNumber="446">
|
||
Fig. 1M
|
||
<emphasis id="B95EEAECBC00FFB06D927ABFFF239A8E" bold="true" box="[188,201,221,244]" pageId="1" pageNumber="446">–</emphasis>
|
||
<collectingCountry id="F33D766EBC00FFB06DE77ABFFF359A8E" box="[201,223,221,244]" name="Iceland" pageId="1" pageNumber="446">O</collectingCountry>
|
||
</figureCitation>
|
||
); distal portion tapers to an elongated conical tip. One or two additional aprocrine glands are located in the convex side of the penis.
|
||
</paragraph>
|
||
<paragraph id="8B9536FEBC00FFB06DE97B40FDEF9B05" blockId="1.[151,1437,152,2012]" pageId="1" pageNumber="446">
|
||
<emphasis id="B95EEAECBC00FFB06DE97B40FE559B43" bold="true" box="[199,447,290,313]" pageId="1" pageNumber="446">Reproductive biology.</emphasis>
|
||
This species is gonochoric, with direct development. Several individuals sampled in Aguada de Chorrillos bore single egg capsules attached to the shells with an embryo or a juvenile snail inside, depending on the stage of development (
|
||
<figureCitation id="13112A7BBC00FFB06CA77B0AFE1D9B05" box="[393,503,360,383]" captionStart="FIGURE 1. A – Q" captionStartId="2.[151,255,1751,1774]" captionTargetBox="[151,1436,333,1726]" captionTargetId="figure@2.[151,1436,333,1728]" captionTargetPageId="2" captionText="FIGURE 1. A – Q. Heleobia deserticola sp. nov. A. Shell of the holotype (MZUC 43067). B – K. Shells of the paratypes (MZUC 43068 – 43077). L. Operculum (1.70 mm total length). M – O. Penes of three individuals (0.64, 0.70 and 0.90 mm total length, respectively). Operculum and penes were photographed from both sides. P, Q. Egg capsules of the new species. P. Egg capsules attached to the adult shells. Q. Pre – hatching capsule isolated with a shelled juvenile inside. R. Shell of an adult snail from Carrera Pinto (SL 3.51 mm, SW 1.55 mm, AL 1.35 mm, AW 0.97 mm. S. Penis of this individual (0.75 mm total length)." httpUri="https://zenodo.org/record/242278/files/figure.png" pageId="1" pageNumber="446">Fig. 1P, Q</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3306575BC00FFB06DE97BE8FBD09873" pageId="1" pageNumber="446" type="materials_examined">
|
||
<paragraph id="8B9536FEBC00FFB06DE97BE8FBD09873" blockId="1.[151,1437,152,2012]" pageId="1" pageNumber="446">
|
||
<emphasis id="B95EEAECBC00FFB06DE97BE8FE8B9BDB" bold="true" box="[199,353,394,417]" pageId="1" pageNumber="446">
|
||
<typeStatus id="5491885CBC00FFB06DE97BE8FF159BDB" box="[199,255,394,417]" pageId="1" pageNumber="446">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
Aguada de Chorrillos, a small spring located at the Pacific coast in the Atacama Desert, northern
|
||
<collectingCountry id="F33D766EBC00FFB06DB97BCFFF3B9BBE" box="[151,209,429,452]" name="Chile" pageId="1" pageNumber="446">Chile</collectingCountry>
|
||
(
|
||
<figureCitation id="13112A7BBC00FFB06DCE7BCFFECA9BBE" box="[224,288,429,452]" captionStart="FIGURE 2" captionStartId="3.[151,255,664,687]" captionTargetBox="[155,1431,198,635]" captionTargetId="figure@3.[151,1436,194,640]" captionTargetPageId="3" captionText="FIGURE 2. Aguada de Chorrillos, Atacama Desert, northern Chile, the type locality of Heleobia deserticola sp. nov. A. Panoramic view of the system, desert, beach and Pacific Ocean. B. Spring where the snails live." httpUri="https://zenodo.org/record/242279/files/figure.png" pageId="1" pageNumber="446">Fig. 2</figureCitation>
|
||
A, B),
|
||
<quantity id="4CD29B1BBC00FFB06C447BCFFE799BBE" box="[362,403,429,452]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="446" unit="m" value="5.0">5 m</quantity>
|
||
above sea level. The site is located several kilometers north of the mouth of the Copiapó River, which is separated from it by an arid desert. The water of the spring arises from the walls of a canyon about
|
||
<quantity id="4CD29B1BBC00FFB068037BB2FA8E9B9D" box="[1325,1380,464,487]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="446" unit="m" value="10.0">10 m</quantity>
|
||
high that form pools (
|
||
<figureCitation id="13112A7BBC00FFB06C667B90FE629873" box="[328,392,498,521]" captionStart="FIGURE 2" captionStartId="3.[151,255,664,687]" captionTargetBox="[155,1431,198,635]" captionTargetId="figure@3.[151,1436,194,640]" captionTargetPageId="3" captionText="FIGURE 2. Aguada de Chorrillos, Atacama Desert, northern Chile, the type locality of Heleobia deserticola sp. nov. A. Panoramic view of the system, desert, beach and Pacific Ocean. B. Spring where the snails live." httpUri="https://zenodo.org/record/242279/files/figure.png" pageId="1" pageNumber="446">Fig. 2</figureCitation>
|
||
B) at the bottom of the ravine which eventually drains to the sea.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3306575BC00FFB06DE97877FF1D980B" pageId="1" pageNumber="446" type="distribution">
|
||
<paragraph id="8B9536FEBC00FFB06DE97877FF1D980B" blockId="1.[151,1437,152,2012]" pageId="1" pageNumber="446">
|
||
<emphasis id="B95EEAECBC00FFB06DE97877FE019856" bold="true" box="[199,491,533,556]" pageId="1" pageNumber="446">Distribution and habitat.</emphasis>
|
||
The snails were collected from the small puddles of water (c.
|
||
<quantity id="4CD29B1BBC00FFB0699E7877FB1C9856" box="[1200,1270,533,556]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="446" unit="cm" value="10.0">10 cm</quantity>
|
||
depth) on soft substrate in the spring Aguada de Chorrillos. In
|
||
<date id="FF94103EBC00FFB06F87785AFCBD9835" box="[681,855,568,591]" pageId="1" pageNumber="446" value="2011-11">November 2011</date>
|
||
individuals of all size ranges from this locality were sampled.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3306575BC00FFB06DE9781FFF0198CD" pageId="1" pageNumber="446" type="etymology">
|
||
<paragraph id="8B9536FEBC00FFB06DE9781FFF0198CD" blockId="1.[151,1437,152,2012]" pageId="1" pageNumber="446">
|
||
<emphasis id="B95EEAECBC00FFB06DE9781FFEAD98EE" bold="true" box="[199,327,637,660]" pageId="1" pageNumber="446">Etymology.</emphasis>
|
||
The name is a compound noun meaning desert dweller; it is derived from the area of origin of the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3306575BC00FFB06DE978A0FDBA9D55" pageId="1" pageNumber="446" type="discussion">
|
||
<paragraph id="8B9536FEBC00FFB06DE978A0FB339EE6" blockId="1.[151,1437,152,2012]" pageId="1" pageNumber="446">
|
||
<emphasis id="B95EEAECBC00FFB06DE978A0FEDF98A3" bold="true" box="[199,309,706,729]" pageId="1" pageNumber="446">Remarks.</emphasis>
|
||
<taxonomicName id="4C2A4D7DBC00FFB06C1378A0FDF798A3" box="[317,541,706,729]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="deserticola" status="sp. nov.">
|
||
<emphasis id="B95EEAECBC00FFB06C1378A0FDF798A3" box="[317,541,706,729]" italics="true" pageId="1" pageNumber="446">Heleobia deserticola</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAECBC00FFB06F0878A0FD9798A3" bold="true" box="[550,637,706,729]" pageId="1" pageNumber="446">
|
||
<taxonomicNameLabel id="A26D5797BC00FFB06F0878A0FD9798A3" box="[550,637,706,729]" pageId="1" pageNumber="446" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from the other species of the genus by its particular penis morphology (
|
||
<figureCitation id="13112A7BBC00FFB06C427887FE009886" box="[364,490,741,764]" captionStart="FIGURE 1. A – Q" captionStartId="2.[151,255,1751,1774]" captionTargetBox="[151,1436,333,1726]" captionTargetId="figure@2.[151,1436,333,1728]" captionTargetPageId="2" captionText="FIGURE 1. A – Q. Heleobia deserticola sp. nov. A. Shell of the holotype (MZUC 43067). B – K. Shells of the paratypes (MZUC 43068 – 43077). L. Operculum (1.70 mm total length). M – O. Penes of three individuals (0.64, 0.70 and 0.90 mm total length, respectively). Operculum and penes were photographed from both sides. P, Q. Egg capsules of the new species. P. Egg capsules attached to the adult shells. Q. Pre – hatching capsule isolated with a shelled juvenile inside. R. Shell of an adult snail from Carrera Pinto (SL 3.51 mm, SW 1.55 mm, AL 1.35 mm, AW 0.97 mm. S. Penis of this individual (0.75 mm total length)." httpUri="https://zenodo.org/record/242278/files/figure.png" pageId="1" pageNumber="446">
|
||
Fig. 1M
|
||
<emphasis id="B95EEAECBC00FFB06CE87887FE399886" bold="true" box="[454,467,741,764]" pageId="1" pageNumber="446">–</emphasis>
|
||
<collectingCountry id="F33D766EBC00FFB06CFA7887FE009886" box="[468,490,741,764]" name="Iceland" pageId="1" pageNumber="446">O</collectingCountry>
|
||
</figureCitation>
|
||
), including the wide portion of the base of the organ, several apocrine glands located around of the complete margin of the base and the additional aprocrine glands located in the convex side of the central portion of the organ. All these characters as a whole differentiate this species from other species of the group whose penis morphology is known (
|
||
<emphasis id="B95EEAECBC00FFB06CA1792FFE5F991E" box="[399,437,845,868]" italics="true" pageId="1" pageNumber="446">e.g.</emphasis>
|
||
<bibRefCitation id="EFBB4B0FBC00FFB06C95792FFD84991E" author="Hubendick" box="[443,622,845,868]" pageId="1" pageNumber="446" refString="Hubendick, B. (1955) XVIII. The anatomy of the Gastropoda. In: The Percy Sladen Trust Expedition to Lake Titicaca in 1937. Transactions of the Linnean Society of London, 1 (3), pp. 309 - 327." type="journal article" year="1955">Hubendick 1955</bibRefCitation>
|
||
; Gaillard & de
|
||
<bibRefCitation id="EFBB4B0FBC00FFB06E3F792FFC21991E" author="Castellanos" box="[785,971,845,868]" pageId="1" pageNumber="446" refString="Gaillard, M. C. & de Castellanos, Z. A. (1976) Mollusca, Gasteropoda, Hydrobiidae. In: Ringuelet, R. A. (Eds.), Fauna de agua dulce de la Republica Argentina, 15 (2), pp. 1 - 39. [Fundacion para la Educacion, la Ciencia y la Cultura (FECIC), Buenos Aires]" type="journal article" year="1976">Castellanos 1976</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBB4B0FBC00FFB06EFB792FFAE8991E" author="Hershler" box="[981,1282,845,868]" pageId="1" pageNumber="446" refString="Hershler, R. & Thompson, F. G. (1992) A review of the aquatic gastropod subfamily Cochliopinae (Prosobranchia: Hydrobiidae). Malacological Review (Supplement), 5, 1 - 140." type="journal article" year="1992">Hershler & Thompson 1992</bibRefCitation>
|
||
; Collado
|
||
<emphasis id="B95EEAECBC00FFB06849792FFA7F991E" box="[1383,1429,845,868]" italics="true" pageId="1" pageNumber="446">et al</emphasis>
|
||
. 2011;
|
||
<bibRefCitation id="EFBB4B0FBC00FFB06DF17912FDD799FD" author="Ovando" box="[223,573,880,903]" pageId="1" pageNumber="446" refString="Ovando, X. M. C. & De Francesco, C. G. (2011) El genero Heleobia en el noroeste argentino. In: Cazzaniga, N. J. (Eds.), El genero Heleobia (Caenogastropoda, Cochliopidae) en America del Sur. Amici Molluscarum, (Numero Especial), pp. 22 - 25" type="book chapter" year="2011">
|
||
Ovando &
|
||
<collectingCountry id="F33D766EBC00FFB06C4F7912FE6A99FD" box="[353,384,880,903]" name="South Georgia and the South Sandwich Islands" pageId="1" pageNumber="446">De</collectingCountry>
|
||
Francesco 2011
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBB4B0FBC00FFB06F637912FD0C99FD" author="Collado" box="[589,742,880,903]" pageId="1" pageNumber="446" refString="Collado, G. A. (2012) Nuevo registro de distribucion geografica y antecedentes de historia natural de Heleobia chimbaensis (Biese, 1944) (Caenogastropoda: Cochliopidae) en la costa del desierto de Atacama: implicancias para su conservacion. Amici Molluscarum, 20 (2), 13 - 18." type="journal article" year="2012">Collado 2012</bibRefCitation>
|
||
; Collado
|
||
<emphasis id="B95EEAECBC00FFB06E767912FC6699FD" box="[856,908,880,903]" italics="true" pageId="1" pageNumber="446">et al</emphasis>
|
||
. 2013). The sister group of this species is an undescribed population from Carrera Pinto, northern
|
||
<collectingCountry id="F33D766EBC00FFB06FF179F0FCF399D3" box="[735,793,914,937]" name="Chile" pageId="1" pageNumber="446">Chile</collectingCountry>
|
||
(Collado
|
||
<emphasis id="B95EEAECBC00FFB06EA479F0FC5099D3" box="[906,954,914,937]" italics="true" pageId="1" pageNumber="446">et al</emphasis>
|
||
. 2013), a spring separated from Aguada de Chorrillos by over
|
||
<quantity id="4CD29B1BBC00FFB06C4A79D7FE5F99B6" box="[356,437,949,972]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="446" unit="km" value="100.0">100 km</quantity>
|
||
of arid desert. I did not treat these populations as conspecific because of differences in shell and penis morphology (
|
||
<figureCitation id="13112A7BBC00FFB06CB279BAFE319995" box="[412,475,984,1007]" captionStart="FIGURE 1. A – Q" captionStartId="2.[151,255,1751,1774]" captionTargetBox="[151,1436,333,1726]" captionTargetId="figure@2.[151,1436,333,1728]" captionTargetPageId="2" captionText="FIGURE 1. A – Q. Heleobia deserticola sp. nov. A. Shell of the holotype (MZUC 43067). B – K. Shells of the paratypes (MZUC 43068 – 43077). L. Operculum (1.70 mm total length). M – O. Penes of three individuals (0.64, 0.70 and 0.90 mm total length, respectively). Operculum and penes were photographed from both sides. P, Q. Egg capsules of the new species. P. Egg capsules attached to the adult shells. Q. Pre – hatching capsule isolated with a shelled juvenile inside. R. Shell of an adult snail from Carrera Pinto (SL 3.51 mm, SW 1.55 mm, AL 1.35 mm, AW 0.97 mm. S. Penis of this individual (0.75 mm total length)." httpUri="https://zenodo.org/record/242278/files/figure.png" pageId="1" pageNumber="446">Fig. 1</figureCitation>
|
||
R, S). The shell of the snails from Carrera Pinto has a band in the growing whorls not present in
|
||
<taxonomicName id="4C2A4D7DBC00FFB06C277998FE009E6B" box="[265,490,1018,1041]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="deserticola" status="sp. nov.">
|
||
<emphasis id="B95EEAECBC00FFB06C277998FE009E6B" box="[265,490,1018,1041]" italics="true" pageId="1" pageNumber="446">Heleobia deserticola</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26D5797BC00FFB06CDF7998FDAC9E6B" box="[497,582,1018,1041]" pageId="1" pageNumber="446" rank="species">sp. nov.</taxonomicNameLabel>
|
||
while the penis has aprocrine glands only on the convex side of the base. The snails from Aguada de Chorrillos and Carrera Pinto belong to the clade including
|
||
<taxonomicName id="4C2A4D7DBC00FFB06ED77E7FFA819E4E" authority="Biese, 1947" authorityName="Biese" authorityYear="1947" box="[1017,1387,1053,1076]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="transitoria">
|
||
<emphasis id="B95EEAECBC00FFB06ED77E7FFB399E4E" box="[1017,1235,1053,1076]" italics="true" pageId="1" pageNumber="446">Heleobia transitoria</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBB4B0FBC00FFB069CF7E7FFA899E4E" author="Biese" box="[1249,1379,1053,1076]" pageId="1" pageNumber="446" refString="Biese, W. A. (1947) Revision de los moluscos terrestres y de agua dulce provistos de concha de Chile. Parte II, Familia Amnicolidae (continuacion). Boletin del Museo Nacional de Historia Natural, 23, 63 - 77." type="journal article" year="1947">Biese, 1947</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, and a population from Quebrada El León in the Atacama Desert. These snails are not very close to
|
||
<taxonomicName id="4C2A4D7DBC00FFB069947E22FEC69E03" authority="Biese, 1947" authorityName="Biese" authorityYear="1947" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="opachensis">
|
||
<emphasis id="B95EEAECBC00FFB069947E22FA719E2D" box="[1210,1435,1088,1111]" italics="true" pageId="1" pageNumber="446">Heleobia opachensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBB4B0FBC00FFB06DB07E00FECE9E03" author="Biese" box="[158,292,1122,1145]" pageId="1" pageNumber="446" refString="Biese, W. A. (1947) Revision de los moluscos terrestres y de agua dulce provistos de concha de Chile. Parte II, Familia Amnicolidae (continuacion). Boletin del Museo Nacional de Historia Natural, 23, 63 - 77." type="journal article" year="1947">Biese, 1947</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2A4D7DBC00FFB06C147E00FD769E03" authority="Biese, 1947" authorityName="Biese" authorityYear="1947" box="[314,668,1122,1145]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="loaensis">
|
||
<emphasis id="B95EEAECBC00FFB06C147E00FE169E03" box="[314,508,1122,1145]" italics="true" pageId="1" pageNumber="446">Heleobia loaensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBB4B0FBC00FFB06F207E00FD7E9E03" author="Biese" box="[526,660,1122,1145]" pageId="1" pageNumber="446" refString="Biese, W. A. (1947) Revision de los moluscos terrestres y de agua dulce provistos de concha de Chile. Parte II, Familia Amnicolidae (continuacion). Boletin del Museo Nacional de Historia Natural, 23, 63 - 77." type="journal article" year="1947">Biese, 1947</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2A4D7DBC00FFB06F847E00FBD29E03" authority="Biese, 1944" authorityName="Biese" authorityYear="1944" box="[682,1080,1122,1145]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="chimbaensis">
|
||
<emphasis id="B95EEAECBC00FFB06F847E00FC739E03" box="[682,921,1122,1145]" italics="true" pageId="1" pageNumber="446">Heleobia chimbaensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBB4B0FBC00FFB06E857E00FBDB9E03" author="Biese" box="[939,1073,1122,1145]" pageId="1" pageNumber="446" refString="Biese, W. A. (1944) Revision de los moluscos terrestres y de agua dulce provistos de concha de Chile. Parte I, Familia Amnicolidae. Boletin del Museo Nacional de Historia Natural, 22, 169 - 190." type="journal article" year="1944">Biese, 1944</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2A4D7DBC00FFB069687E00FF3F9EE6" authority="Courty, 1907" authorityName="Courty" authorityYear="1907" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="ascotanensis">
|
||
<emphasis id="B95EEAECBC00FFB069687E00FAD09E03" box="[1094,1338,1122,1145]" italics="true" pageId="1" pageNumber="446">Heleobia ascotanensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBB4B0FBC00FFB068657E00FF249EE6" author="Courty" pageId="1" pageNumber="446" refString="Courty, G. (1907) Explorations geologiques dans l'Amerique du Sud. In: Mission Scientifique de G. de Crequi Montfort et E. Senechal de la Grange, Vol. 14: 1 - 208. Paris: Imprimerie Nationale." type="journal article" year="1907">Courty, 1907</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2A4D7DBC00FFB06C277EE7FC639EE6" authority="(Philippi, 1860) (Collado et al. 2013)" authorityName="(Philippi, 1860) (Collado et al." authorityYear="2013" box="[265,905,1157,1180]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="atacamensis">
|
||
<emphasis id="B95EEAECBC00FFB06C277EE7FE1E9EE6" box="[265,500,1157,1180]" italics="true" pageId="1" pageNumber="446">Heleobia atacamensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBB4B0FBC00FFB06F2D7EE7FD779EE6" author="Philippi" box="[515,669,1157,1180]" pageId="1" pageNumber="446" refString="Philippi, R. A. (1860) Reise durch die Wueste Atacama auf Befehl der chilenischen Regierung im Sommer 1853 - 54. E Anton, Halle, 62 pp." type="book chapter" year="1860">Philippi, 1860</bibRefCitation>
|
||
) (Collado
|
||
<emphasis id="B95EEAECBC00FFB06E217EE7FCD49EE6" box="[783,830,1157,1180]" italics="true" pageId="1" pageNumber="446">et al</emphasis>
|
||
. 2013)
|
||
</taxonomicName>
|
||
described from northern
|
||
<collectingCountry id="F33D766EBC00FFB069B67EE7FB3F9EE6" box="[1176,1237,1157,1180]" name="Chile" pageId="1" pageNumber="446">Chile</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9536FEBC00FFB06DE97ECAFDBA9D55" blockId="1.[151,1437,152,2012]" pageId="1" pageNumber="446">
|
||
<emphasis id="B95EEAECBC00FFB06DE97ECAFD069EC5" bold="true" box="[199,748,1191,1215]" pageId="1" pageNumber="446">
|
||
Comparison with the other
|
||
<taxonomicName id="4C2A4D7DBC00FFB06F0A7EC5FD6D9EC5" box="[548,647,1191,1215]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B95EEAECBC00FFB06F0A7EC5FD6D9EC5" bold="true" box="[548,647,1191,1215]" italics="true" pageId="1" pageNumber="446">Heleobia</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
</emphasis>
|
||
<taxonomicName id="4C2A4D7DBC00FFB06FD27ECAFC099EC5" box="[764,995,1192,1215]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="deserticola" status="sp. nov.">
|
||
<emphasis id="B95EEAECBC00FFB06FD27ECAFC099EC5" box="[764,995,1192,1215]" italics="true" pageId="1" pageNumber="446">Heleobia deserticola</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAECBC00FFB06EDD7ECAFBB89EC5" bold="true" box="[1011,1106,1192,1215]" pageId="1" pageNumber="446">
|
||
<taxonomicNameLabel id="A26D5797BC00FFB06EDD7ECAFBB89EC5" box="[1011,1106,1192,1215]" pageId="1" pageNumber="446" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
fits well the morphological characteristics of the shell and penis of the genus as defined by
|
||
<bibRefCitation id="EFBB4B0FBC00FFB06E6D7EA8FB6F9E9B" author="Hershler" box="[835,1157,1226,1249]" pageId="1" pageNumber="446" refString="Hershler, R. & Thompson, F. G. (1992) A review of the aquatic gastropod subfamily Cochliopinae (Prosobranchia: Hydrobiidae). Malacological Review (Supplement), 5, 1 - 140." type="journal article" year="1992">Hershler & Thompson (1992)</bibRefCitation>
|
||
, with the male copulatory organ possessing aprocrine glands. Additionally, the new species was also placed in the genus
|
||
<taxonomicName id="4C2A4D7DBC00FFB069967E8FFAF09F7E" box="[1208,1306,1261,1284]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B95EEAECBC00FFB069967E8FFAF09F7E" box="[1208,1306,1261,1284]" italics="true" pageId="1" pageNumber="446">Heleobia</emphasis>
|
||
</taxonomicName>
|
||
based on a molecular phylogenetic analysis (Collado
|
||
<emphasis id="B95EEAECBC00FFB06F787F72FD6F9F5D" box="[598,645,1296,1319]" italics="true" pageId="1" pageNumber="446">et al</emphasis>
|
||
. 2013). Of the nominal species of
|
||
<taxonomicName id="4C2A4D7DBC00FFB06EDC7F72FBBE9F5D" box="[1010,1108,1296,1319]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B95EEAECBC00FFB06EDC7F72FBBE9F5D" box="[1010,1108,1296,1319]" italics="true" pageId="1" pageNumber="446">Heleobia</emphasis>
|
||
</taxonomicName>
|
||
from northern
|
||
<collectingCountry id="F33D766EBC00FFB069DD7F72FAC59F5D" box="[1267,1327,1296,1319]" name="Chile" pageId="1" pageNumber="446">Chile</collectingCountry>
|
||
,
|
||
<taxonomicName id="4C2A4D7DBC00FFB068147F72FEE49F33" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="deserticola" status="sp. nov.">
|
||
<emphasis id="B95EEAECBC00FFB068147F72FEE49F33" italics="true" pageId="1" pageNumber="446">Heleobia deserticola</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAECBC00FFB06C3A7F50FE819F33" bold="true" box="[276,363,1330,1353]" pageId="1" pageNumber="446">
|
||
<taxonomicNameLabel id="A26D5797BC00FFB06C3A7F50FE819F33" box="[276,363,1330,1353]" pageId="1" pageNumber="446" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is the third representative described from the coast of the Atacama Desert after
|
||
<taxonomicName id="4C2A4D7DBC00FFB069E77F50FA879F33" box="[1225,1389,1330,1353]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="chimbaensis">
|
||
<emphasis id="B95EEAECBC00FFB069E77F50FA879F33" box="[1225,1389,1330,1353]" italics="true" pageId="1" pageNumber="446">H. chimbaensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2A4D7DBC00FFB06DB97F37FEC19F16" box="[151,299,1365,1388]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="transitoria">
|
||
<emphasis id="B95EEAECBC00FFB06DB97F37FEC19F16" box="[151,299,1365,1388]" italics="true" pageId="1" pageNumber="446">H. transitoria</emphasis>
|
||
</taxonomicName>
|
||
. The shell of the new species is little differentiated morphologically from other regional congeners [see Collado
|
||
<emphasis id="B95EEAECBC00FFB06DDC7F1AFECB9FF5" box="[242,289,1400,1423]" italics="true" pageId="1" pageNumber="446">et al</emphasis>
|
||
. (2011),
|
||
<bibRefCitation id="EFBB4B0FBC00FFB06CAC7F1AFDC29FF5" author="Collado" box="[386,552,1400,1423]" pageId="1" pageNumber="446" refString="Collado, G. A. (2012) Nuevo registro de distribucion geografica y antecedentes de historia natural de Heleobia chimbaensis (Biese, 1944) (Caenogastropoda: Cochliopidae) en la costa del desierto de Atacama: implicancias para su conservacion. Amici Molluscarum, 20 (2), 13 - 18." type="journal article" year="2012">Collado (2012)</bibRefCitation>
|
||
, Collado
|
||
<emphasis id="B95EEAECBC00FFB06FA07F1AFD579FF5" box="[654,701,1400,1423]" italics="true" pageId="1" pageNumber="446">et al</emphasis>
|
||
. (2013) for comparative purposes]. The closed umbilicus of the new species resembles that of
|
||
<taxonomicName id="4C2A4D7DBC00FFB06C837FF8FDB89FCB" box="[429,594,1434,1457]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="atacamensis">
|
||
<emphasis id="B95EEAECBC00FFB06C837FF8FDB89FCB" box="[429,594,1434,1457]" italics="true" pageId="1" pageNumber="446">H. atacamensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2A4D7DBC00FFB06F4C7FF8FD319FCB" box="[610,731,1434,1457]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="loaensis">
|
||
<emphasis id="B95EEAECBC00FFB06F4C7FF8FD319FCB" box="[610,731,1434,1457]" italics="true" pageId="1" pageNumber="446">H. loaensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2A4D7DBC00FFB06FC47FF8FC689FCB" box="[746,898,1434,1457]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="opachensis">
|
||
<emphasis id="B95EEAECBC00FFB06FC47FF8FC689FCB" box="[746,898,1434,1457]" italics="true" pageId="1" pageNumber="446">H. opachensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2A4D7DBC00FFB06E947FF8FBA49FCB" box="[954,1102,1434,1457]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="transitoria">
|
||
<emphasis id="B95EEAECBC00FFB06E947FF8FC279FCB" box="[954,973,1434,1457]" italics="true" pageId="1" pageNumber="446">H</emphasis>
|
||
.
|
||
<emphasis id="B95EEAECBC00FFB06EF27FF8FBA49FCB" box="[988,1102,1434,1457]" italics="true" pageId="1" pageNumber="446">transitoria</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C2A4D7DBC00FFB069727FF8FAD79FCB" box="[1116,1341,1434,1457]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="deserticola" status="sp. nov.">
|
||
<emphasis id="B95EEAECBC00FFB069727FF8FAD79FCB" box="[1116,1341,1434,1457]" italics="true" pageId="1" pageNumber="446">Heleobia deserticola</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAECBC00FFB0686B7FF8FA769FCB" bold="true" box="[1349,1436,1434,1457]" pageId="1" pageNumber="446">
|
||
<taxonomicNameLabel id="A26D5797BC00FFB0686B7FF8FA769FCB" box="[1349,1436,1434,1457]" pageId="1" pageNumber="446" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has direct development, similar to the reproductive strategy employed by
|
||
<taxonomicName id="4C2A4D7DBC00FFB06ED87FDFFB499FAE" box="[1014,1187,1469,1492]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="chimbaensis">
|
||
<emphasis id="B95EEAECBC00FFB06ED87FDFFBE39FAE" box="[1014,1033,1469,1492]" italics="true" pageId="1" pageNumber="446">H</emphasis>
|
||
.
|
||
<emphasis id="B95EEAECBC00FFB069317FDFFB499FAE" box="[1055,1187,1469,1492]" italics="true" pageId="1" pageNumber="446">chimbaensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2A4D7DBC00FFB069967FDFFEB79F8D" authority="d'Orbigny, 1835" authorityName="d'Orbigny" authorityYear="1835" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="parchappii">
|
||
<emphasis id="B95EEAECBC00FFB069967FDFFA769FAE" box="[1208,1436,1469,1492]" italics="true" pageId="1" pageNumber="446">Heleobia parchappii</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBB4B0FBC00FFB06DB07F82FEBF9F8D" author="d'Orbigny" box="[158,341,1504,1527]" pageId="1" pageNumber="446" refString="d'Orbigny, A. D. (1835) Synopsis terrestrium et fluviatilium molluscorum in suo per Americam Meridionalem itinere collectorum. Magasin de zoologie, d'anatomie comparee et de palaeontologie, 5 (61 - 62), 1 - 44." type="journal article" year="1835">d’Orbigny, 1835</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2A4D7DBC00FFB06C447F82FCA29F8D" authority="Marcus & Marcus, 1965" authorityName="Marcus & Marcus" authorityYear="1965" box="[362,840,1504,1527]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="miaulis">
|
||
<emphasis id="B95EEAECBC00FFB06C447F82FDC89F8D" box="[362,546,1504,1527]" italics="true" pageId="1" pageNumber="446">Heleobia miaulis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBB4B0FBC00FFB06F1C7F82FCAA9F8D" author="Marcus" box="[562,832,1504,1527]" pageId="1" pageNumber="446" refString="Marcus E. & Marcus E. (1965) On Brazilian supratidal and estuarine snails. Boletim da Faculdade de Filosofia, Ciencias e Letras da Universidade de Sao Paulo, Zoologia, 25, 18 - 103." type="journal article" year="1965">Marcus & Marcus, 1965</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2A4D7DBC00FFB06EAE7F82FAF79F8D" authority="Doering, 1885" authorityName="Doering" authorityYear="1885" box="[896,1309,1504,1527]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="guaranitica">
|
||
<emphasis id="B95EEAECBC00FFB06EAE7F82FB8C9F8D" box="[896,1126,1504,1527]" italics="true" pageId="1" pageNumber="446">Heleobia guaranitica</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBB4B0FBC00FFB069587F82FAFF9F8D" author="Doering" box="[1142,1301,1504,1527]" pageId="1" pageNumber="446" refString="Doering, A. (1885) Apuntes sobre la fauna de moluscos de la Republica Argentina. 5 ª entrega. Boletin de la Academia Nacional de Ciencias en Cordoba, 7 (4), 457 - 474." type="journal article" year="1885">Doering, 1885</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFBB4B0FBC00FFB068037F82FEC19C63" author="Marcus" pageId="1" pageNumber="446" refString="Marcus E. & Marcus E. (1965) On Brazilian supratidal and estuarine snails. Boletim da Faculdade de Filosofia, Ciencias e Letras da Universidade de Sao Paulo, Zoologia, 25, 18 - 103." type="journal article" year="1965">Marcus & Marcus 1965</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBB4B0FBC00FFB06C157C60FE3D9C63" author="Gaillard" box="[315,471,1538,1561]" pageId="1" pageNumber="446" refString="Gaillard, M. C. (1973) Contribucion al conocimiento del genero Littoridina Souyelet 1852 en Argentina. Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, 119 pp. [Argentina]" type="book" year="1973">Gaillard 1973</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBB4B0FBC00FFB06CC67C60FD719C63" author="Cazzaniga" box="[488,667,1538,1561]" pageId="1" pageNumber="446" refString="Cazzaniga, N. J. (1982) Notas sobre los hidrobidos argentinos. 5. Conquiliometria de Littoridina parchappii (d'Orbigny, 1835) (Gastropoda Rissoidea) referida a su ciclo de vida en poblaciones australes. Iheringia, Serie Zoologia, 61, 97 - 118." type="journal article" year="1982">Cazzaniga 1982</bibRefCitation>
|
||
; Collado & Méndez 2011); other species of
|
||
<taxonomicName id="4C2A4D7DBC00FFB069B07C60FAEA9C63" box="[1182,1280,1538,1561]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B95EEAECBC00FFB069B07C60FAEA9C63" box="[1182,1280,1538,1561]" italics="true" pageId="1" pageNumber="446">Heleobia</emphasis>
|
||
</taxonomicName>
|
||
have indirect development (see Collado & Méndez 2011). The black pigmentation of the head of
|
||
<taxonomicName id="4C2A4D7DBC00FFB069317C47FB5D9C46" box="[1055,1207,1573,1596]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="deserticola" status="sp. nov.">
|
||
<emphasis id="B95EEAECBC00FFB069317C47FBD89C46" box="[1055,1074,1573,1596]" italics="true" pageId="1" pageNumber="446">H</emphasis>
|
||
.
|
||
<emphasis id="B95EEAECBC00FFB0696E7C47FB5D9C46" box="[1088,1207,1573,1596]" italics="true" pageId="1" pageNumber="446">deserticola</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAECBC00FFB069907C47FAFE9C46" bold="true" box="[1214,1300,1573,1596]" pageId="1" pageNumber="446">
|
||
<taxonomicNameLabel id="A26D5797BC00FFB069907C47FAFE9C46" box="[1214,1300,1573,1596]" pageId="1" pageNumber="446" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from that of
|
||
<taxonomicName id="4C2A4D7DBC00FFB06DCF7C2AFDC19C25" authority="Bavay, 1904" authorityName="Bavay" authorityYear="1904" box="[225,555,1608,1631]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="neveui">
|
||
<emphasis id="B95EEAECBC00FFB06DCF7C2AFE659C25" box="[225,399,1608,1631]" italics="true" pageId="1" pageNumber="446">Heleobia neveui</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBB4B0FBC00FFB06CB27C2AFDC99C25" author="Bavay" box="[412,547,1608,1631]" pageId="1" pageNumber="446" refString="Bavay, A. (1904) Mission de Crequi - Montfort et Senechal de la Grange en Amerique du Sud. Mollusques terrestres et fluviatiles recoltes par le Dr Neveu - Lemaire. Bulletin de la Societe Zoologique de France, 29, 152 - 156." type="journal article" year="1904">Bavay, 1904</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
because the latter species in general is less pigmented (
|
||
<bibRefCitation id="EFBB4B0FBC00FFB0695A7C2AFAC09C25" author="Hubendick" box="[1140,1322,1608,1631]" pageId="1" pageNumber="446" refString="Hubendick, B. (1955) XVIII. The anatomy of the Gastropoda. In: The Percy Sladen Trust Expedition to Lake Titicaca in 1937. Transactions of the Linnean Society of London, 1 (3), pp. 309 - 327." type="journal article" year="1955">Hubendick 1955</bibRefCitation>
|
||
). The gray pigmentation of the tentacles of
|
||
<taxonomicName id="4C2A4D7DBC00FFB06CDA7C08FD669CFB" box="[500,652,1642,1665]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="deserticola" status="sp. nov.">
|
||
<emphasis id="B95EEAECBC00FFB06CDA7C08FDED9CFB" box="[500,519,1642,1665]" italics="true" pageId="1" pageNumber="446">H</emphasis>
|
||
.
|
||
<emphasis id="B95EEAECBC00FFB06F387C08FD669CFB" box="[534,652,1642,1665]" italics="true" pageId="1" pageNumber="446">deserticola</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAECBC00FFB06FBA7C08FD019CFB" bold="true" box="[660,747,1642,1665]" pageId="1" pageNumber="446">
|
||
<taxonomicNameLabel id="A26D5797BC00FFB06FBA7C08FD019CFB" box="[660,747,1642,1665]" pageId="1" pageNumber="446" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from that of
|
||
<taxonomicName id="4C2A4D7DBC00FFB06EE67C08FAC49CFB" authority="Haas, 1955" authorityName="Haas" authorityYear="1955" box="[968,1326,1642,1665]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="compacta">
|
||
<emphasis id="B95EEAECBC00FFB06EE67C08FB709CFB" box="[968,1178,1642,1665]" italics="true" pageId="1" pageNumber="446">Heleobia compacta</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBB4B0FBC00FFB069877C08FACC9CFB" author="Haas" box="[1193,1318,1642,1665]" pageId="1" pageNumber="446" refString="Haas, F. (1955) XVII. Mollusca: Gastropoda. In: The Percy Sladen Trust Expedition to Lake Titicaca in 1937. Transactions of the Linnean Society of London, 1 (3), 275 - 308." type="journal article" year="1955">Haas, 1955</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2A4D7DBC00FFB068147C08FE579CDE" authority="d'Orbigny, 1835" authorityName="d'Orbigny" authorityYear="1835" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="cumingii">
|
||
<emphasis id="B95EEAECBC00FFB068147C08FF1E9CDE" italics="true" pageId="1" pageNumber="446">Heleobia cumingii</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBB4B0FBC00FFB06C2C7CEFFE5F9CDE" author="d'Orbigny" box="[258,437,1677,1700]" pageId="1" pageNumber="446" refString="d'Orbigny, A. D. (1835) Synopsis terrestrium et fluviatilium molluscorum in suo per Americam Meridionalem itinere collectorum. Magasin de zoologie, d'anatomie comparee et de palaeontologie, 5 (61 - 62), 1 - 44." type="journal article" year="1835">d’Orbigny, 1835</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2A4D7DBC00FFB06CDF7CEFFCDA9CDE" authority="Haas, 1955" authorityName="Haas" authorityYear="1955" box="[497,816,1677,1700]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="aperta">
|
||
<emphasis id="B95EEAECBC00FFB06CDF7CEFFD759CDE" box="[497,671,1677,1700]" italics="true" pageId="1" pageNumber="446">Heleobia aperta</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBB4B0FBC00FFB06F837CEFFCC29CDE" author="Haas" box="[685,808,1677,1700]" pageId="1" pageNumber="446" refString="Haas, F. (1955) XVII. Mollusca: Gastropoda. In: The Percy Sladen Trust Expedition to Lake Titicaca in 1937. Transactions of the Linnean Society of London, 1 (3), 275 - 308." type="journal article" year="1955">Haas, 1955</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from Lake Titicaca, which are almost black over all these structures (
|
||
<bibRefCitation id="EFBB4B0FBC00FFB06C227CD2FE2E9CBD" author="Hubendick" box="[268,452,1712,1735]" pageId="1" pageNumber="446" refString="Hubendick, B. (1955) XVIII. The anatomy of the Gastropoda. In: The Percy Sladen Trust Expedition to Lake Titicaca in 1937. Transactions of the Linnean Society of London, 1 (3), pp. 309 - 327." type="journal article" year="1955">Hubendick 1955</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C2A4D7DBC00FFB06CFB7CD2FDDD9CBD" box="[469,567,1712,1735]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B95EEAECBC00FFB06CFB7CD2FDDD9CBD" box="[469,567,1712,1735]" italics="true" pageId="1" pageNumber="446">Heleobia</emphasis>
|
||
</taxonomicName>
|
||
is divided into two groups on the basis of the size (
|
||
<bibRefCitation id="EFBB4B0FBC00FFB069727CD2FB3D9CBD" author="Biese" box="[1116,1239,1712,1735]" pageId="1" pageNumber="446" refString="Biese, W. A. (1944) Revision de los moluscos terrestres y de agua dulce provistos de concha de Chile. Parte I, Familia Amnicolidae. Boletin del Museo Nacional de Historia Natural, 22, 169 - 190." type="journal article" year="1944">Biese 1944</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFBB4B0FBC00FFB069CC7CD2FAF19CBD" author="Biese" box="[1250,1307,1712,1735]" pageId="1" pageNumber="446" refString="Biese, W. A. (1947) Revision de los moluscos terrestres y de agua dulce provistos de concha de Chile. Parte II, Familia Amnicolidae (continuacion). Boletin del Museo Nacional de Historia Natural, 23, 63 - 77." type="journal article" year="1947">1947</bibRefCitation>
|
||
). The group of
|
||
<taxonomicName id="4C2A4D7DBC00FFB06D947CB0FDDC9C93" authority="Pilsbry, 1911" authorityName="Pilsbry" authorityYear="1911" box="[186,566,1746,1769]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="hatcheri">
|
||
<emphasis id="B95EEAECBC00FFB06D947CB0FE689C93" box="[186,386,1746,1769]" italics="true" pageId="1" pageNumber="446">Heleobia hatcheri</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBB4B0FBC00FFB06CB97CB0FDC79C93" author="Pilsbry" box="[407,557,1746,1769]" pageId="1" pageNumber="446" refString="Pilsbry, H. A. (1911) Non - marine Mollusca of Patagonia. In: Scott, W. B. (Eds.), Reports of the Princeton University Expedition to Patagonia 1896 - 1899. Part 5. Zoology. The University of Princeton, Princeton, NJ, pp. 513 - 633." type="book chapter" year="1911">Pilsbry, 1911</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, which contains small species, with 4–5 shell whorls, and the group of
|
||
<taxonomicName id="4C2A4D7DBC00FFB068AC7CB0FEE19D76" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="parchappii">
|
||
<emphasis id="B95EEAECBC00FFB068AC7CB0FEE19D76" italics="true" pageId="1" pageNumber="446">H. parchappii</emphasis>
|
||
</taxonomicName>
|
||
, which contains larger forms, with 5–8 whorls. The new species belongs to the second group; adult individuals can reach up to seven whorls.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3306575BC00FFB36DE97D58FE7C9B6D" lastPageId="2" lastPageNumber="447" pageId="1" pageNumber="446" type="reference_group">
|
||
<paragraph id="8B9536FEBC00FFB36DE97D58FE7C9B6D" blockId="1.[151,1437,152,2012]" lastBlockId="2.[151,1436,152,279]" lastPageId="2" lastPageNumber="447" pageId="1" pageNumber="446">
|
||
<taxonomicName id="4C2A4D7DBC00FFB06DE97D58FE4D9D2B" ID-CoL="3K4QL" box="[199,423,1850,1873]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="deserticola" status="sp. nov.">
|
||
<emphasis id="B95EEAECBC00FFB06DE97D58FE4D9D2B" box="[199,423,1850,1873]" italics="true" pageId="1" pageNumber="446">Heleobia deserticola</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAECBC00FFB06C807D58FDEE9D2B" bold="true" box="[430,516,1850,1873]" pageId="1" pageNumber="446">
|
||
<taxonomicNameLabel id="A26D5797BC00FFB06C807D58FDEE9D2B" box="[430,516,1850,1873]" pageId="1" pageNumber="446" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is only known from the spring of Aguada de Chorrillos (Collado
|
||
<emphasis id="B95EEAECBC00FFB069E27D58FB169D2B" box="[1228,1276,1850,1873]" italics="true" pageId="1" pageNumber="446">et al</emphasis>
|
||
. 2013; present study). This oasis is an extremely small restricted habitat, surrounded by an arid desert and facing the sea. In northern
|
||
<collectingCountry id="F33D766EBC00FFB06DB97DE2FF3E9DED" box="[151,212,1920,1943]" name="Chile" pageId="1" pageNumber="446">Chile</collectingCountry>
|
||
, several localities with populations of
|
||
<taxonomicName id="4C2A4D7DBC00FFB06FA27DE2FD049DED" ID-CoL="7P4WQ" box="[652,750,1920,1943]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B95EEAECBC00FFB06FA27DE2FD049DED" box="[652,750,1920,1943]" italics="true" pageId="1" pageNumber="446">Heleobia</emphasis>
|
||
</taxonomicName>
|
||
are subjected to anthropogenic pressures, pollution or water scarcity (
|
||
<bibRefCitation id="EFBB4B0FBC00FFB06DD77DC0FE7B9DC3" author="Collado" box="[249,401,1954,1977]" pageId="1" pageNumber="446" refString="Collado, G. A. (2012) Nuevo registro de distribucion geografica y antecedentes de historia natural de Heleobia chimbaensis (Biese, 1944) (Caenogastropoda: Cochliopidae) en la costa del desierto de Atacama: implicancias para su conservacion. Amici Molluscarum, 20 (2), 13 - 18." type="journal article" year="2012">Collado 2012</bibRefCitation>
|
||
). Recently, two species of
|
||
<taxonomicName id="4C2A4D7DBC00FFB06F9C7DC0FCFE9DC3" ID-CoL="7P4WQ" box="[690,788,1954,1977]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B95EEAECBC00FFB06F9C7DC0FCFE9DC3" box="[690,788,1954,1977]" italics="true" pageId="1" pageNumber="446">Heleobia</emphasis>
|
||
</taxonomicName>
|
||
in the Atacama Desert,
|
||
<taxonomicName id="4C2A4D7DBC00FFB069357DC0FB559DC3" ID-CoL="3K4Q9" box="[1051,1215,1954,1977]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="atacamensis">
|
||
<emphasis id="B95EEAECBC00FFB069357DC0FB559DC3" box="[1051,1215,1954,1977]" italics="true" pageId="1" pageNumber="446">H. atacamensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2A4D7DBC00FFB069D87DC0FA769DC3" ID-CoL="3K4QG" box="[1270,1436,1954,1977]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="446" phylum="Mollusca" rank="species" species="chimbaensis">
|
||
<emphasis id="B95EEAECBC00FFB069D87DC0FA769DC3" box="[1270,1436,1954,1977]" italics="true" pageId="1" pageNumber="446">H. chimbaensis</emphasis>
|
||
</taxonomicName>
|
||
were declared Critically Endangered and Vulnerable, respectively (
|
||
<bibRefCitation id="EFBB4B0FBC00FFB06EA17DA7FBD29DA6" author="Collado" box="[911,1080,1989,2012]" pageId="1" pageNumber="446" refString="Collado, G. A. (2013 a) Heleobia atacamensis (Philippi, 1860). In: 10 º Proceso de Clasificacion de Especies. Ministerio del Medio Ambiente, Chile. Available from: http: // www. mma. gob. cl (accessed 7 July 2014)" type="journal article" year="2013" yearSuffix="a">Collado 2013a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFBB4B0FBC00FFB069127DA7FBA09DA6" author="Collado" box="[1084,1098,1989,2012]" pageId="1" pageNumber="446" refString="Collado, G. A. (2013 b) Heleobia chimbaensis (Biese, 1944). In: 10 º Proceso de Clasificacion de Especies. Ministerio del Medio Ambiente, Chile. Available from: http: // www. mma. gob. cl (accessed 7 July 2014)" type="journal article" year="2013" yearSuffix="b">b</bibRefCitation>
|
||
), according to the list of the threatened species of the Ministerio del Medio Ambiente,
|
||
<collectingCountry id="F33D766EBC03FFB36E077AFAFBE89AD5" box="[809,1026,152,175]" name="Chile" pageId="2" pageNumber="447">República de Chile</collectingCountry>
|
||
. It is clear that some biological and population parameters need to be estimated by monitoring
|
||
<taxonomicName id="4C2A4D7DBC03FFB36E357AD8FC169AAB" ID-CoL="3K4QL" box="[795,1020,186,209]" class="Gastropoda" family="Cochliopidae" genus="Heleobia" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="447" phylum="Mollusca" rank="species" species="deserticola" status="sp. nov.">
|
||
<emphasis id="B95EEAECBC03FFB36E357AD8FC169AAB" box="[795,1020,186,209]" italics="true" pageId="2" pageNumber="447">Heleobia deserticola</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAECBC03FFB3692A7AD8FBB69AAB" bold="true" box="[1028,1116,186,209]" pageId="2" pageNumber="447">
|
||
<taxonomicNameLabel id="A26D5797BC03FFB3692A7AD8FBB69AAB" box="[1028,1116,186,209]" pageId="2" pageNumber="447" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
because any alteration to the biotope could affect the population of these snails in Aguada de Chorrillos. There are currently no conservation measures for this particular place.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |