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<mods:title id="807A2062FF3C6956322DB35EE1A566D6">First description of winged stages of Thraulobaetodes Elouard &amp; Hideux 1991 and reclassification of Rhithrocloeoninae (Ephemeroptera, Baetidae)</mods:title>
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<mods:namePart id="4953CBDCE89B7A1F3F387E5B6E9D881F">Kluge, Nikita J.</mods:namePart>
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<subSubSection id="C3306540D93AFFC8ECDD1FEFFEE5BCD6" pageId="3" pageNumber="494" type="nomenclature">
<paragraph id="8B9536CBD93AFFC8ECDD1FEFFD11BCF7" blockId="3.[151,748,896,955]" box="[151,748,896,922]" pageId="3" pageNumber="494">
<heading id="D0DD81A7D93AFFC8ECDD1FEFFD11BCF7" bold="true" box="[151,748,896,922]" fontSize="11" level="1" pageId="3" pageNumber="494" reason="1">
<emphasis id="B95EEAD9D93AFFC8ECDD1FEFFD11BCF7" bold="true" box="[151,748,896,922]" pageId="3" pageNumber="494">
Genus
<taxonomicName id="4C2A4D48D93AFFC8ECA41FEFFD11BCF7" ID-CoL="648BV" authority="Elouard &amp; Hideux 1991" authorityName="Elouard &amp; Hideux" authorityYear="1991" box="[238,748,896,922]" class="Insecta" family="Baetidae" genus="Thraulobaetodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="494" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAD9D93AFFC8ECA41FEFFE47BCF7" bold="true" box="[238,442,896,922]" italics="true" pageId="3" pageNumber="494">Thraulobaetodes</emphasis>
<bibRefCitation id="EFBB4B3AD93AFFC8ED8B1FEFFD11BCF7" author="Elouard" box="[449,748,896,922]" pageId="3" pageNumber="494" refString="Elouard, J. - M. &amp; Hideux, P. (1991) Mayflies of West Africa. Thraulobaetodes, an atypic new genus of crawling Baetidae. In: Alba-Tercedor J. &amp; Sanchez-Ortega A. (Eds). Overview and strategies of Ephemeroptera and Plecoptera. Sandhill Crane Press, Gainesville, pp. 169 - 174." type="book chapter" year="1991">Elouard &amp; Hideux 1991</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9536CBD93AFFC8ECDD1FCDFEE5BCD6" blockId="3.[151,748,896,955]" box="[151,280,930,955]" pageId="3" pageNumber="494">
(
<figureCitation id="13112A4ED93AFFC8ECD51FCDFEEDBCD6" box="[159,272,930,955]" captionStart-0="FIGURES 1 6" captionStart-1="FIGURES 7 10" captionStart-2="FIGURES 11 17" captionStart-3="FIGURES 18 24" captionStart-4="FIGURES 25 30" captionStart-5="FIGURES 31 37" captionStart-6="FIGURES 38 44" captionStart-7="FIGURES 45 48" captionStartId-0="5.[151,264,1861,1884]" captionStartId-1="6.[151,264,1913,1936]" captionStartId-2="7.[151,264,1574,1597]" captionStartId-3="8.[151,264,1772,1795]" captionStartId-4="9.[151,264,1952,1975]" captionStartId-5="10.[151,264,1598,1621]" captionStartId-6="11.[151,264,1844,1867]" captionStartId-7="12.[151,264,1653,1676]" captionTargetBox-0="[151,1435,207,1823]" captionTargetBox-1="[162,1435,193,1880]" captionTargetBox-2="[151,1425,218,1541]" captionTargetBox-3="[151,1436,194,1750]" captionTargetBox-4="[156,1409,214,1931]" captionTargetBox-5="[157,1431,206,1566]" captionTargetBox-6="[170,1415,193,1810]" captionTargetBox-7="[170,1416,193,1631]" captionTargetId-0="figure@5.[151,1436,195,1838]" captionTargetId-1="figure@6.[151,1435,193,1892]" captionTargetId-2="figure@7.[151,1436,193,1554]" captionTargetId-3="figure@8.[151,1436,193,1751]" captionTargetId-4="figure@9.[154,1432,193,1932]" captionTargetId-5="figure@10.[151,1436,193,1577]" captionTargetId-6="figure@11.[170,1416,193,1823]" captionTargetId-7="figure@12.[170,1416,193,1632]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="11" captionTargetPageId-7="12" captionText-0="FIGURES 1 6. Thraulobaetodes cumminsorum, larval mouthparts (on the same scale). 1, labrum, dorsal view; 2, 3, apices of left and right mandibles; 4, labium (at left half, dorsal view; at right half, ventral view with muscles shown by interrupted lines); 5, maxilla, dorsal view; 6, maxillary palp of another specimen." captionText-1="FIGURES 7 10. Thraulobaetodes cumminsorum, larval mouthparts (on the same scale). 7, superlinguae; 8, hypopharynx and maxilla (arrow shows 3 rd segment of maxillary palp); 9, labium; 10, right mandible." captionText-2="FIGURES 11 17. Thraulobaetodes cumminsorum, larva. 11, neck articulation and prothorax, lateral view; 12, lateral view of last instar female larva; 13, 14, 15 cross sections of ridges on pronotum, mesonotum and metanotum; 16, outline of hind protopteron with developing subimaginal hind wing inside; 17, abdominal segment IV with left tergalius, posterior view." captionText-3="FIGURES 18 24. Thraulobaetodes cumminsorum, exuviae of last instar larva spread on slide (Figs 18, 19, 21 and 22 on the same scale). 18, abdominal terga I IX; 19, left half of pronotum and mesonotum; 20, part of pronotum, enlarged; 21, hind leg; 22, head of male larva; 23, abdominal tergum IV; 24, paraprocts." captionText-4="FIGURES 25 30. Thraulobaetodes cumminsorum, legs. 25, larval middle leg, front view; 26, claw, median view; 27 29, fragment of subimaginal leg, front view: 27, fore leg of male; 28, fore leg of female; 29, hind leg of female; 30, tibia and tarsus of subimaginal middle leg, front view (same individual on same scale as Fig 25). Arrow shows patella-tibial suture." captionText-5="FIGURES 31 37. Thraulobaetodes cumminsorum. 31, fore and hind wing on the same scale; 32, hind wing, enlarged; 33, apex of fore wing of another individual; 34, head of female imago; 35, outline of abdominal segment IX of mature male larva with developing subimaginal genitals inside (in left half muscles shown by interrupted lines); 36, exuviae of left gonostylus of male subimago; 37, genitals of male imago, the same individual (muscles shown by interrupted lines; in left half gonostylar muscle not shown, internal sclerotized part of penis shown by interrupted line and dotted; in right half cuticular pigmentation shown by dots). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; m. gs, gonostylar muscle; m. gv 1, m. gv 2, muscles of gonovectis; m. s, sterno-styligeral muscle; us, unistyliger." captionText-6="FIGURES 38 44. Thraulobaetodes cumminsorum. 38, apex of abdomen of male imago, lateral view; 39, genitals of male subimago, ventral view; 40, subimaginal exuviae of genitals; 41, genitals of male imago of the same individual, ventral view; 42, female imago; 43, male imago (arrows show protuberances—remnants of larval spines on abdominal terga); 44, mesonotum, lateral-dorsal view." captionText-7="FIGURES 45 48. Thraulobaetodes cumminsorum. 45, 46, subimaginal exuviae spread on slide (on the same scale): 45, left half of mesonotum; 46, part of mesopleuron and metapleuron, with fragments of fore and middle legs. 47, 48, egg and its enlarged fragment (extracted from female imago). Abbreviation: psa, postsubalar sclerite of mesothorax." httpUri-0="https://zenodo.org/record/242406/files/figure.png" httpUri-1="https://zenodo.org/record/242407/files/figure.png" httpUri-2="https://zenodo.org/record/242408/files/figure.png" httpUri-3="https://zenodo.org/record/242409/files/figure.png" httpUri-4="https://zenodo.org/record/242410/files/figure.png" httpUri-5="https://zenodo.org/record/242411/files/figure.png" httpUri-6="https://zenodo.org/record/242412/files/figure.png" httpUri-7="https://zenodo.org/record/242413/files/figure.png" pageId="3" pageNumber="494">Figs 148</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3306540D93AFFC8ECDD1F84FD53BB72" pageId="3" pageNumber="494" type="materials_examined">
<paragraph id="8B9536CBD93AFFC8ECDD1F84FD53BB72" blockId="3.[151,872,1002,1055]" pageId="3" pageNumber="494">
<typeStatus id="54918869D93AFFC8ECDD1F84FF34BB6D" box="[151,201,1003,1024]" pageId="3" pageNumber="494">Type</typeStatus>
species:
<taxonomicName id="4C2A4D48D93AFFC8ED6C1F85FC98BB6D" authority="Elouard &amp; Hideux 1991" authorityName="Elouard &amp; Hideux" authorityYear="1991" box="[294,869,1002,1024]" class="Insecta" family="Baetidae" genus="Thraulobaetodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="494" phylum="Arthropoda" rank="species" species="cumminsorum">
<emphasis id="B95EEAD9D93AFFC8ED6C1F85FD9ABB6D" box="[294,615,1002,1024]" italics="true" pageId="3" pageNumber="494">Thraulobaetodes cumminsorum</emphasis>
<bibRefCitation id="EFBB4B3AD93AFFC8EE271F85FC98BB6D" author="Elouard" box="[621,869,1002,1024]" pageId="3" pageNumber="494" refString="Elouard, J. - M. &amp; Hideux, P. (1991) Mayflies of West Africa. Thraulobaetodes, an atypic new genus of crawling Baetidae. In: Alba-Tercedor J. &amp; Sanchez-Ortega A. (Eds). Overview and strategies of Ephemeroptera and Plecoptera. Sandhill Crane Press, Gainesville, pp. 169 - 174." type="book chapter" year="1991">Elouard &amp; Hideux 1991</bibRefCitation>
</taxonomicName>
. Rank-free hierarchical name:
<taxonomicName id="4C2A4D48D93AFFC8ED8E1866FD8BBB72" box="[452,630,1033,1055]" class="Insecta" family="Baetidae" genus="Thraulobaetodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="494" phylum="Arthropoda" rank="genus">Thraulobaetodes</taxonomicName>
/fg(1).
</paragraph>
</subSubSection>
<subSubSection id="C3306540D93AFFC8ECDD1823FB79BBE5" pageId="3" pageNumber="494" type="diagnosis">
<paragraph id="8B9536CBD93AFFC8ECDD1823FC39BB08" blockId="3.[151,1156,1100,1160]" box="[151,964,1100,1125]" pageId="3" pageNumber="494">
<emphasis id="B95EEAD9D93AFFC8ECDD1823FEECBB08" bold="true" box="[151,273,1100,1125]" pageId="3" pageNumber="494">Diagnosis.</emphasis>
Coincides with description of
<taxonomicName id="4C2A4D48D93AFFC8EE231823FCCEBB09" box="[617,819,1100,1125]" class="Insecta" family="Baetidae" genus="Thraulobaetodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="494" phylum="Arthropoda" rank="species" species="cumminsorum">
<emphasis id="B95EEAD9D93AFFC8EE231823FCCEBB09" box="[617,819,1100,1125]" italics="true" pageId="3" pageNumber="494">Th. cumminsorum</emphasis>
</taxonomicName>
(see below).
</paragraph>
<paragraph id="8B9536CBD93AFFC8EC8D1800FB79BBE5" blockId="3.[151,1156,1100,1160]" box="[199,1156,1135,1160]" pageId="3" pageNumber="494">
<emphasis id="B95EEAD9D93AFFC8EC8D1800FE9ABBE5" bold="true" box="[199,359,1135,1160]" pageId="3" pageNumber="494">Composition.</emphasis>
Includes the single species
<taxonomicName id="4C2A4D48D93AFFC8EED41800FB7DBBE5" authority="Elouard &amp; Hideux 1991" authorityName="Elouard &amp; Hideux" authorityYear="1991" box="[670,1152,1135,1160]" class="Insecta" family="Baetidae" genus="Thraulobaetodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="494" phylum="Arthropoda" rank="species" species="cumminsorum">
<emphasis id="B95EEAD9D93AFFC8EED41800FC95BBE5" box="[670,872,1135,1160]" italics="true" pageId="3" pageNumber="494">Th. cumminsorum</emphasis>
<bibRefCitation id="EFBB4B3AD93AFFC8EF251800FB7DBBE5" author="Elouard" box="[879,1152,1135,1160]" pageId="3" pageNumber="494" refString="Elouard, J. - M. &amp; Hideux, P. (1991) Mayflies of West Africa. Thraulobaetodes, an atypic new genus of crawling Baetidae. In: Alba-Tercedor J. &amp; Sanchez-Ortega A. (Eds). Overview and strategies of Ephemeroptera and Plecoptera. Sandhill Crane Press, Gainesville, pp. 169 - 174." type="book chapter" year="1991">Elouard &amp; Hideux 1991</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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