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<document id="1DD0AC1F85C740962F6FE5A6F43B49AE" ID-DOI="10.1649/1065.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="5369896" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630268191031" checkinUser="valdenar" docAuthor="Anderson, Robert S." docDate="1807" docId="038387CCFFD3FFDBC2F3FB0A2FCAFBFA" docLanguage="en" docName="1065.1.pdf" docOrigin="The Coleopterists Bulletin 62 (1)" docSource="http://dx.doi.org/10.1649/1065.1" docStyle="DocumentStyle:76939A7658C5F80CF4BB6818E728552A.2:TheColeopBull.2003-2009.journal_article.1cover" docStyleId="76939A7658C5F80CF4BB6818E728552A" docStyleName="TheColeopBull.2003-2009.journal_article.1cover" docStyleVersion="2" docTitle="Cryptorhynchus Illiger 1807" docType="treatment" docVersion="3" lastPageNumber="169" masterDocId="FFBAFFB4FFD2FFD9C34CFFE72C26FFD1" masterDocTitle="A Review of the Genus Cryptorhynchus Illiger 1807 in the United States and Canada (Curculionidae: Cryptorhynchinae)" masterLastPageNumber="180" masterPageNumber="168" pageNumber="168" updateTime="1699205265088" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="B9636F53D81A54D4A24B601093CD6557">A Review of the Genus Cryptorhynchus Illiger 1807 in the United States and Canada (Curculionidae: Cryptorhynchinae)</mods:title>
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<mods:title id="D9493D19B46F614976905CD11262568D">The Coleopterists Bulletin</mods:title>
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<mods:date id="58061959320DBF04D2DCF9706DB628C0">1807</mods:date>
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<subSubSection id="C3306551FFD3FFD8C2F3FB0A2E84FAD2" box="[447,674,1261,1283]" pageId="1" pageNumber="168" type="nomenclature">
<paragraph id="8B9536DAFFD3FFD8C2F3FB0A2E84FAD2" blockId="1.[447,674,1261,1283]" box="[447,674,1261,1283]" pageId="1" pageNumber="168">
<heading id="D0DD81B6FFD3FFD8C2F3FB0A2E84FAD2" box="[447,674,1261,1283]" centered="true" fontSize="9" level="2" pageId="1" pageNumber="168" reason="2">
<taxonomicName id="4C2A4D59FFD3FFD8C2F3FB0A2E84FAD2" ID-CoL="8HF62" authority="Illiger" authorityName="Illiger" authorityYear="1807" box="[447,674,1261,1283]" class="Insecta" family="Curculionidae" genus="Cryptorhynchus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="168" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAC8FFD3FFD8C2F3FB0A2E84FAD2" bold="true" box="[447,674,1261,1283]" pageId="1" pageNumber="168">
<emphasis id="B95EEAC8FFD3FFD8C2F3FB0A2E78FAD2" bold="true" box="[447,606,1261,1283]" italics="true" pageId="1" pageNumber="168">Cryptorhynchus</emphasis>
Illiger
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3306551FFD3FFD8C3DEFAC52F1BFA83" pageId="1" pageNumber="168" type="reference_group">
<paragraph id="8B9536DAFFD3FFD8C3DEFAC52F1BFA83" blockId="1.[146,974,1314,1362]" pageId="1" pageNumber="168">
<taxonomicName id="4C2A4D59FFD3FFD8C3DEFAC52DC0FAE9" ID-CoL="8HF62" authority="Illiger 1807: 330" authorityName="Illiger" authorityPageNumber="330" authorityYear="1807" box="[146,486,1314,1336]" class="Insecta" family="Curculionidae" genus="Cryptorhynchus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="168" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAC8FFD3FFD8C3DEFAC52D17FAE6" box="[146,305,1314,1335]" italics="true" pageId="1" pageNumber="168">Cryptorhynchus</emphasis>
Illiger 1807:330
</taxonomicName>
.
<emphasis id="B95EEAC8FFD3FFD8C2BAFAC52F1BFA83" italics="true" pageId="1" pageNumber="168">(Complete generic synonymy is presented in Alonso-Zarazaga and Lyal (1999) and is not repeated here.)</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3306551FFD3FFD8C3E6FA8A2CD6FA4C" pageId="1" pageNumber="168" type="type_taxon">
<paragraph id="8B9536DAFFD3FFD8C3E6FA8A2CD6FA4C" blockId="1.[146,975,1389,1570]" pageId="1" pageNumber="168">
<materialsCitation id="3B423C87FFD3FFD8C3E6FA8A2CCDFA4C" collectionCode="ICZN" pageId="1" pageNumber="168" specimenCode="ICZN 1967" specimenCount="1" typeStatus="holotype">
<emphasis id="B95EEAC8FFD3FFD8C3E6FA8A2D0BFA52" bold="true" box="[170,301,1389,1411]" pageId="1" pageNumber="168">
<typeStatus id="54918878FFD3FFD8C3E6FA8A2CFAFA52" box="[170,220,1389,1411]" pageId="1" pageNumber="168">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C2A4D59FFD3FFD8C278FA8A2E49FA52" ID-CoL="32HFP" authority="Linnaeus 1758" authorityName="Linnaeus" authorityYear="1758" box="[308,623,1389,1411]" class="Insecta" family="Curculionidae" genus="Curculio" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="168" phylum="Arthropoda" rank="species" species="lapathi">
<emphasis id="B95EEAC8FFD3FFD8C278FA8A2DF3FA53" box="[308,469,1389,1410]" italics="true" pageId="1" pageNumber="168">Curculio lapathi</emphasis>
Linnaeus 1758
</taxonomicName>
by subsequent designation (
<specimenCode id="DB8C9EA1FFD3FFD8C0DCFA8A2CE1FA4C" collectionCode="ICZN" pageId="1" pageNumber="168">ICZN 1967</specimenCode>
:83)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3306551FFD3FFDBC3E6FA452FCAFBFA" lastPageId="2" lastPageNumber="169" pageId="1" pageNumber="168" type="diagnosis">
<paragraph id="8B9536DAFFD3FFD8C3E6FA452FE9F9F3" blockId="1.[146,975,1389,1570]" pageId="1" pageNumber="168">
<emphasis id="B95EEAC8FFD3FFD8C3E6FA452D34FA69" bold="true" box="[170,274,1442,1464]" pageId="1" pageNumber="168">Diagnosis.</emphasis>
Within North America,
<taxonomicName id="4C2A4D59FFD3FFD8C151FA452E9AFA66" authorityName="Illiger" authorityYear="1807" box="[541,700,1442,1463]" class="Insecta" family="Curculionidae" genus="Cryptorhynchus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="168" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAC8FFD3FFD8C151FA452E9AFA66" box="[541,700,1442,1463]" italics="true" pageId="1" pageNumber="168">Cryptorhynchus</emphasis>
</taxonomicName>
can be recognized by the following combination of characters. Body size
<quantity id="4CD29B3FFFD3FFD8C131FA5A2ED2FA02" box="[637,756,1469,1491]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="9.5" metricValueMin="4.5" pageId="1" pageNumber="168" unit="mm" value="7.0" valueMax="9.5" valueMin="4.5">4.59.5 mm</quantity>
in length. Head with eyes separated by slightly less than width of rostrum; eyes partially covered by postocular lobe when head in repose. Rostrum more or less evenly curved, mandibles prominent when closed with visible inner acute tooth. Pronotum
</paragraph>
<paragraph id="8B9536DAFFD3FFD8C16DF9D92E67F980" blockId="1.[545,577,1598,1617]" box="[545,577,1598,1617]" pageId="1" pageNumber="168">168</paragraph>
<paragraph id="8B9536DAFFD0FFDBC3E3FF252DD1FE8F" blockId="2.[175,1005,194,889]" pageId="2" pageNumber="169">slightly produced over head but vertex visible in dorsal view. Elytra with intervals flat or with intervals 3 and 5 very slightly convex and with tufts or patches of erect dark scales giving them a more elevated appearance; vestiture of broad scales only, no hairs. Middle coxae widely separated by distance greater than width of coxa. Legs with femora each with one (may be small) or two teeth; tibiae with outer edge simple, not carinate.</paragraph>
<paragraph id="8B9536DAFFD0FFDBC38BFE832EE1FDCA" blockId="2.[175,1005,194,889]" pageId="2" pageNumber="169">
<emphasis id="B95EEAC8FFD0FFDBC38BFE832D0EFEAB" bold="true" box="[199,296,356,378]" pageId="2" pageNumber="169">Diversity.</emphasis>
<taxonomicName id="4C2A4D59FFD0FFDBC278FE832DF5FEA8" authorityName="Illiger" authorityYear="1807" box="[308,467,356,377]" class="Insecta" family="Curculionidae" genus="Cryptorhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAC8FFD0FFDBC278FE832DF5FEA8" box="[308,467,356,377]" italics="true" pageId="2" pageNumber="169">Cryptorhynchus</emphasis>
</taxonomicName>
is a large, cosmopolitan genus. Only a very few species are temperate and the vast majority are tropical in their distributions. For example, some 120+ species are known from Central America (OBrien and Wibmer 1982).
<bibRefCitation id="EFBB4B2BFFD0FFDBC214FE532E2EFE1B" author="Anderson" box="[344,520,436,458]" pageId="2" pageNumber="169" pagination="722 - 815" refId="ref6623" refString="Anderson, R. S. 2002. Chapter 131. Curculionidae [pp. 722 - 815]. In: American Beetles Volume 2 (R. H. Arnett, Jr, M. C. Thomas, and P. Skelley, editors). CRC Press." type="journal article" year="2002">Anderson (2002)</bibRefCitation>
mistakenly stated that only four species are present in North America (erroneously excluding
<taxonomicName id="4C2A4D59FFD0FFDBC19EFE282FB2FE34" authority="LeConte" authorityName="LeConte" box="[722,916,463,485]" class="Insecta" family="Curculionidae" genus="Cryptorhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="169" phylum="Arthropoda" rank="species" species="tristis">
<emphasis id="B95EEAC8FFD0FFDBC19EFE282F08FE35" box="[722,814,463,484]" italics="true" pageId="2" pageNumber="169">C. tristis</emphasis>
LeConte
</taxonomicName>
and
<taxonomicName id="4C2A4D59FFD0FFDBC099FE282D5CFDD1" authority="(Sleeper))" authorityName="Sleeper. Jens Prena" authorityYear="1955" baseAuthorityName="Sleeper" class="Insecta" family="Curculionidae" genus="Sternochetus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="169" phylum="Arthropoda" rank="species" species="woodruffi">
<emphasis id="B95EEAC8FFD0FFDBC099FE282D28FE2E" italics="true" pageId="2" pageNumber="169">C. woodruffi</emphasis>
(Sleeper))
</taxonomicName>
. With the nomenclatural changes reported herein, five species are now recognized as occurring in North America.
</paragraph>
<paragraph id="8B9536DAFFD0FFDBC38BFDC72EA9FC93" blockId="2.[175,1005,194,889]" pageId="2" pageNumber="169">
<emphasis id="B95EEAC8FFD0FFDBC38BFDC72D90FDE7" bold="true" box="[199,438,544,566]" pageId="2" pageNumber="169">Nomenclatural changes.</emphasis>
<taxonomicName id="4C2A4D59FFD0FFDBC2F1FDC62F85FDE7" authority="LeConte 1876: 254" authorityName="LeConte" authorityPageNumber="254" authorityYear="1876" box="[445,931,544,566]" class="Insecta" family="Curculionidae" genus="Cryptorhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="169" phylum="Arthropoda" rank="species" species="minutissimus">
<emphasis id="B95EEAC8FFD0FFDBC2F1FDC62EC5FDE7" box="[445,739,545,566]" italics="true" pageId="2" pageNumber="169">Cryptorhynchus minutissimus</emphasis>
LeConte 1876:254
</taxonomicName>
is here removed from
<taxonomicName id="4C2A4D59FFD0FFDBC21FFDDC2DD4FD81" authorityName="Illiger" authorityYear="1807" box="[339,498,571,592]" class="Insecta" family="Curculionidae" genus="Cryptorhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAC8FFD0FFDBC21FFDDC2DD4FD81" box="[339,498,571,592]" italics="true" pageId="2" pageNumber="169">Cryptorhynchus</emphasis>
</taxonomicName>
and provisionally placed in the genus
<taxonomicName id="4C2A4D59FFD0FFDBC0E8FDDC2D05FDBD" authority="Schoenherr" authorityName="Schoenherr" class="Insecta" family="Curculionidae" genus="Acalles" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAC8FFD0FFDBC0E8FDDC2FCAFD81" box="[932,1004,571,592]" italics="true" pageId="2" pageNumber="169">Acalles</emphasis>
Schoenherr
</taxonomicName>
as
<taxonomicName id="4C2A4D59FFD0FFDBC21EFDB12EB8FDBD" authority="(LeConte)" authorityName="Anderson" authorityYear="1807" baseAuthorityName="Anderson" baseAuthorityYear="1807" box="[338,670,598,620]" class="Insecta" family="Curculionidae" genus="Acalles" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="169" phylum="Arthropoda" rank="species" species="minutissimus" status="comb. nov.">
<emphasis id="B95EEAC8FFD0FFDBC21EFDB12E01FDBA" box="[338,551,598,619]" italics="true" pageId="2" pageNumber="169">Acalles minutissimus</emphasis>
(LeConte)
</taxonomicName>
,
<taxonomicNameLabel id="A26D57B3FFD0FFDBC1E3FDB12F4FFDBD" box="[687,873,598,620]" pageId="2" pageNumber="169" rank="species">new combination</taxonomicNameLabel>
. It is much smaller in size than other North American
<taxonomicName id="4C2A4D59FFD0FFDBC120FD962F2DFD57" authorityName="Illiger" authorityYear="1807" box="[620,779,625,646]" class="Insecta" family="Curculionidae" genus="Cryptorhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAC8FFD0FFDBC120FD962F2DFD57" box="[620,779,625,646]" italics="true" pageId="2" pageNumber="169">Cryptorhynchus</emphasis>
</taxonomicName>
and is very similar to
<taxonomicName id="4C2A4D59FFD0FFDBC3E3FD6B2E04FD70" authority="Blatchley. Both" authorityName="Blatchley. Both" box="[175,546,651,673]" class="Insecta" family="Curculionidae" genus="Acalles" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="169" phylum="Arthropoda" rank="species" species="sablensis">
<emphasis id="B95EEAC8FFD0FFDBC3E3FD6B2D45FD70" box="[175,355,652,673]" italics="true" pageId="2" pageNumber="169">Acalles sablensis</emphasis>
Blatchley. Both
</taxonomicName>
possess a distinct and relatively long metepisternal suture, similar abdominal structure (length of ventrites and depth of sutures), lack teeth on the femora, and have a very similar dorsal scale pattern. Undoubtedly, the definition of
<taxonomicName id="4C2A4D59FFD0FFDBC14AFD3B2E68FD20" box="[518,590,732,753]" class="Insecta" family="Curculionidae" genus="Acalles" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAC8FFD0FFDBC14AFD3B2E68FD20" box="[518,590,732,753]" italics="true" pageId="2" pageNumber="169">Acalles</emphasis>
</taxonomicName>
will require modification itself as the continued inclusion of both Old World and New World species in the same genus is questionable. The combination
<taxonomicName id="4C2A4D59FFD0FFDBC153FCF52F6BFCF9" authority="Say" authorityName="Say" authorityYear="1831" box="[543,845,786,808]" class="Insecta" family="Curculionidae" genus="Cryptorhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="169" phylum="Arthropoda" rank="species" species="obliquus">
<emphasis id="B95EEAC8FFD0FFDBC153FCF52F3AFCF6" box="[543,796,786,807]" italics="true" pageId="2" pageNumber="169">Cryptorhynchus obliquus</emphasis>
Say
</taxonomicName>
has also been resurrected (see discussion under that species).
</paragraph>
<paragraph id="8B9536DAFFD0FFDBC38BFCAF2F84FCA8" blockId="2.[175,1005,194,889]" pageId="2" pageNumber="169">
Based upon these changes, some modifications to couplet 25 of the key to genera of Cryptorhynchinae in
<bibRefCitation id="EFBB4B2BFFD0FFDBC2B9FC842E87FCA8" author="Anderson" box="[501,673,867,889]" pageId="2" pageNumber="169" pagination="722 - 815" refId="ref6623" refString="Anderson, R. S. 2002. Chapter 131. Curculionidae [pp. 722 - 815]. In: American Beetles Volume 2 (R. H. Arnett, Jr, M. C. Thomas, and P. Skelley, editors). CRC Press." type="journal article" year="2002">Anderson (2002)</bibRefCitation>
can be made as follows.
</paragraph>
<paragraph id="8B9536DAFFD0FFDBC3FEFC4D2FCAFC0B" blockId="2.[178,1004,938,1068]" pageId="2" pageNumber="169">
25(24) Elytra with alternate intervals 3, 5, 7 and 9 elevated and sharply carinate, lacking any tufts of erect scales.....................................
<taxonomicName id="4C2A4D59FFD0FFDBC0D1FC222FCAFC0B" box="[925,1004,965,986]" class="Insecta" family="Curculionidae" genus="Eubulus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAC8FFD0FFDBC0D1FC222FCAFC0B" box="[925,1004,965,986]" italics="true" pageId="2" pageNumber="169">Eubulus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8B9536DAFFD0FFDBC3FEFC072FCAFBFA" blockId="2.[178,1004,938,1068]" pageId="2" pageNumber="169">
- Elytra with intervals 3 and 5 flat to very slightly elevated, not carinate, with tufts or patches of suberect or erect broad, dark scales giving them a more elevated appearance..................................
<taxonomicName id="4C2A4D59FFD0FFDBC001FBF12FCAFBFA" authorityName="Illiger" authorityYear="1807" box="[845,1004,1046,1067]" class="Insecta" family="Curculionidae" genus="Cryptorhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAC8FFD0FFDBC001FBF12FCAFBFA" box="[845,1004,1046,1067]" italics="true" pageId="2" pageNumber="169">Cryptorhynchus</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
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