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<document id="13A7D32E98B4FCE315F1BA46DE9DC9B5" ID-CLB-Dataset="46860" ID-DOI="10.11646/zootaxa.4072.2.9" ID-GBIF-Dataset="f248e718-4700-4062-8387-dc13e722a3fe" ID-ISSN="1175-5326" ID-Zenodo-Dep="264448" ID-ZooBank="1C21A78B-C978-4C2B-BD83-66F54478980E" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460358036549" checkinUser="plazi" docAuthor="Wang, Cheng-Bin &amp; Zhou, Hong-Zhang" docDate="2016" docId="038387C8A85F0408FF22FE0B0DC5FE9E" docLanguage="en" docName="zootaxa.4072.2.9.pdf" docOrigin="Zootaxa 4072 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anemadus perreaui Wang &amp; Zhou, 2016, sp. nov." docType="treatment" docVersion="9" lastPageNumber="286" masterDocId="FFBAFFB0A85C040DFFB5FFEF0F24FF82" masterDocTitle="Two new species of the Anemadus taiwanus species-group (Coleoptera: Leiodidae: Cholevinae: Anemadini) from China" masterLastPageNumber="290" masterPageNumber="282" pageNumber="285" updateTime="1698294126784" updateUser="ExternalLinkService">
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<mods:title id="0F54B43B2A78367C45BC1D5A8B720B02">Two new species of the Anemadus taiwanus species-group (Coleoptera: Leiodidae: Cholevinae: Anemadini) from China</mods:title>
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<mods:namePart id="80125697B809419D39854E1670805B6E">Wang, Cheng-Bin</mods:namePart>
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<mods:namePart id="916E34C71078F2F4C2A301FC5BAECA48">Zhou, Hong-Zhang</mods:namePart>
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<mods:date id="2793C0B23F8B11FA5A4D086AD84ED69E">2016</mods:date>
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<treatment id="038387C8A85F0408FF22FE0B0DC5FE9E" ID-DOI="http://doi.org/10.5281/zenodo.6086546" ID-GBIF-Taxon="119409092" ID-Zenodo-Dep="6086546" LSID="urn:lsid:plazi:treatment:038387C8A85F0408FF22FE0B0DC5FE9E" httpUri="http://treatment.plazi.org/id/038387C8A85F0408FF22FE0B0DC5FE9E" lastPageId="5" lastPageNumber="286" pageId="3" pageNumber="285">
<subSubSection id="C3306555A85F040EFF22FE0B0EA7FD9D" pageId="3" pageNumber="285" type="nomenclature">
<paragraph id="8B9536DEA85F040EFF22FE0B0ECEFE7C" blockId="3.[151,490,484,543]" box="[151,490,484,510]" pageId="3" pageNumber="285">
<heading id="D0DD81B2A85F040EFF22FE0B0ECEFE7C" bold="true" box="[151,490,484,510]" fontSize="11" level="1" pageId="3" pageNumber="285" reason="1">
<emphasis id="B95EEACCA85F040EFF22FE0B0ECEFE7C" bold="true" box="[151,490,484,510]" pageId="3" pageNumber="285">
<taxonomicName id="4C2A4D5DA85F040EFF22FE0B0EA1FE7C" ID-CoL="DWSD" box="[151,389,484,510]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="285" phylum="Arthropoda" rank="species" species="perreaui" status="sp. nov.">
<emphasis id="B95EEACCA85F040EFF22FE0B0EA1FE7C" bold="true" box="[151,389,484,510]" italics="true" pageId="3" pageNumber="285">Anemadus perreaui</emphasis>
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<taxonomicNameLabel id="A26D57B7A85F040EFE39FE0B0ECEFE7C" box="[396,490,484,510]" pageId="3" pageNumber="285" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9536DEA85F040EFF2BFDE90EA7FD9D" blockId="3.[151,490,484,543]" box="[158,387,518,543]" pageId="3" pageNumber="285">
(
<figureCitation id="13112A5BA85F040EFF13FDE90FD7FD9D" box="[166,243,518,543]" captionStart="FIGURE 2" captionStartId="4.[151,250,1646,1668]" captionTargetBox="[171,1415,216,1604]" captionTargetId="figure@4.[151,1436,193,1625]" captionTargetPageId="4" captionText="FIGURE 2. Anemadus perreaui sp. nov.: A, habitus ♂ (dorsal view; Baoxing, Ruobigou); B, habitus ♀ (dorsal view; Baoxing, Ruobigou); C, habitus ♀ (dorsal view; Baoxing, Ganyanggou); D, right elytron ♂ (dorsal view; in glycerine). Anemadus sichuanus sp. nov. ♂: E, habitus ♂ (dorsal view; Baoxing, Pujigou); F, habitus ♀ (dorsal view; Baoxing, Pujigou); G, right elytron ♂ (dorsal view; in glycerine). Scales: 1 mm." httpUri="https://zenodo.org/record/264450/files/figure.png" pageId="3" pageNumber="285">Figs. 2</figureCitation>
AD; 3AL)
</paragraph>
</subSubSection>
<subSubSection id="C3306555A85F040EFF22FDA10A26FD2D" pageId="3" pageNumber="285" type="materials_examined">
<paragraph id="8B9536DEA85F040EFF22FDA10A26FD2D" blockId="3.[151,1437,590,2025]" pageId="3" pageNumber="285">
<emphasis id="B95EEACCA85F040EFF22FDA10DB8FDE5" bold="true" box="[151,668,590,615]" pageId="3" pageNumber="285">
<typeStatus id="5491887CA85F040EFF22FDA10FF6FDE5" box="[151,210,590,615]" pageId="3" pageNumber="285">Type</typeStatus>
material.
<typeStatus id="5491887CA85F040EFEFEFDA10E98FDE5" box="[331,444,590,615]" pageId="3" pageNumber="285" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F33D764EA85F040EFE7DFDA10D03FDE5" box="[456,551,590,615]" name="China" pageId="3" pageNumber="285">CHINA</collectingCountry>
, Sichuan:
</emphasis>
♂, Baoxing, Ruobigou,
<quantity id="4CD29B3BA85F040EFC1BFDA10B21FDE5" box="[942,1029,590,615]" metricMagnitude="3" metricUnit="m" metricValue="1.63" pageId="3" pageNumber="285" unit="m" value="1630.0">1630 m</quantity>
,
<taxonomicName id="4C2A4D5DA85F040EFBA5FDA00BEFFDE5" box="[1040,1227,591,615]" class="Magnoliopsida" family="Fagaceae" genus="Cyclobalanopsis" higherTaxonomySource="CoL" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="285" phylum="Tracheophyta" rank="genus">
<emphasis id="B95EEACCA85F040EFBA5FDA00BEFFDE5" box="[1040,1227,591,615]" italics="true" pageId="3" pageNumber="285">Cyclobalanopsis</emphasis>
</taxonomicName>
forest, from fallen wood,
<date id="FF94101EA85F040EFF5CFD9C0E5AFD09" box="[233,382,627,651]" pageId="3" pageNumber="285" value="2003-08-10">10.VIII.2003</date>
, Jie Wu leg. (IZ-CAS).
<emphasis id="B95EEACCA85F040EFD10FD9D0C02FD09" bold="true" box="[677,806,626,651]" pageId="3" pageNumber="285">
<typeStatus id="5491887CA85F040EFD10FD9D0C05FD09" box="[677,801,626,651]" pageId="3" pageNumber="285" type="paratype">Paratypes</typeStatus>
:
</emphasis>
1♀, same data as
<typeStatus id="5491887CA85F040EFBBAFD9C0B55FD09" box="[1039,1137,627,651]" pageId="3" pageNumber="285" type="holotype">holotype</typeStatus>
(IZ-CAS); 1♀, Baoxing, Ganyanggou,
<quantity id="4CD29B3BA85F040EFE81FD780EAFFD2D" box="[308,395,663,687]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="285" unit="m" value="2000.0">2000 m</quantity>
,
<taxonomicName id="4C2A4D5DA85F040EFE22FD780D76FD2D" box="[407,594,663,687]" class="Magnoliopsida" family="Fagaceae" genus="Cyclobalanopsis" higherTaxonomySource="CoL" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="285" phylum="Tracheophyta" rank="genus">
<emphasis id="B95EEACCA85F040EFE22FD780D76FD2D" box="[407,594,663,687]" italics="true" pageId="3" pageNumber="285">Cyclobalanopsis</emphasis>
</taxonomicName>
forest, from fallen wood,
<date id="FF94101EA85F040EFCC3FD780CD9FD2D" box="[886,1021,663,687]" pageId="3" pageNumber="285" value="2003-08-09">9.VIII.2003</date>
, Jie Wu leg. (IZ-CAS).
</paragraph>
</subSubSection>
<subSubSection id="C3306555A85F040EFF72FD550DB3F86B" pageId="3" pageNumber="285" type="description">
<paragraph id="8B9536DEA85F040EFF72FD550E26FCBD" blockId="3.[151,1437,590,2025]" pageId="3" pageNumber="285">
<emphasis id="B95EEACCA85F040EFF72FD550E7DFD51" bold="true" box="[199,345,698,723]" pageId="3" pageNumber="285">Description.</emphasis>
<emphasis id="B95EEACCA85F040EFED5FD540EBEFD51" box="[352,410,699,723]" italics="true" pageId="3" pageNumber="285">Male</emphasis>
. EBL: 3.00 mm. Length of different body parts: HL: AL: PL: ELL = 0.56: 1.14: 0.68:
<quantity id="4CD29B3BA85F040EFF22FD310FD9FD75" box="[151,253,734,759]" metricMagnitude="-3" metricUnit="m" metricValue="1.61" pageId="3" pageNumber="285" unit="mm" value="1.61">1.61 mm</quantity>
; width: HW: EW: PW: ELW = 0.70: 0.12: 1.16:
<quantity id="4CD29B3BA85F040EFCF5FD300C82FD75" box="[832,934,735,759]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" pageId="3" pageNumber="285" unit="mm" value="1.42">1.42 mm</quantity>
. Proportion of antennomeres from base to tip in µm (length × width): 111 × 62, 147 × 50, 119 × 43, 74 × 42, 79 × 47, 64 × 51, 94 × 71, 41 × 68, 85 × 81, 83 × 89, 155 × 88.
</paragraph>
<paragraph id="8B9536DEA85F040EFF72FCA50A3FFC4D" blockId="3.[151,1437,590,2025]" pageId="3" pageNumber="285">
Habitus (
<figureCitation id="13112A5BA85F040EFE9AFCA50E57FCE1" box="[303,371,842,867]" captionStart="FIGURE 2" captionStartId="4.[151,250,1646,1668]" captionTargetBox="[171,1415,216,1604]" captionTargetId="figure@4.[151,1436,193,1625]" captionTargetPageId="4" captionText="FIGURE 2. Anemadus perreaui sp. nov.: A, habitus ♂ (dorsal view; Baoxing, Ruobigou); B, habitus ♀ (dorsal view; Baoxing, Ruobigou); C, habitus ♀ (dorsal view; Baoxing, Ganyanggou); D, right elytron ♂ (dorsal view; in glycerine). Anemadus sichuanus sp. nov. ♂: E, habitus ♂ (dorsal view; Baoxing, Pujigou); F, habitus ♀ (dorsal view; Baoxing, Pujigou); G, right elytron ♂ (dorsal view; in glycerine). Scales: 1 mm." httpUri="https://zenodo.org/record/264450/files/figure.png" pageId="3" pageNumber="285">Fig. 2</figureCitation>
A) elongated oval, regularly convex and sublustrous. Moderately pigmented: mostly chestnut brown; mouthparts, basal four or five antennomeres and apical half of ultimate antennomere, areas near hind corners of pronotum, elytral apices, and tarsi more or less yellowish. Dorsum continually covered with fine, recumbent, sallow pubescence. Insertions of pubescence on elytra aligned along transverse striolations.
</paragraph>
<paragraph id="8B9536DEA85F040EFF72FC350EC0FB0B" blockId="3.[151,1437,590,2025]" pageId="3" pageNumber="285">
Head retractile, HW/HL = 1.24. Surface covered with strong round punctures, separated about 2.04.0 times of their diameter; interspaces smooth, without visable microsculpture. Clypeofrontal suture distinct. Clypeus trapezoidal, anterior margin almost straight. Compound eyes well developed, EW/HW = 0.12. Antennae (
<figureCitation id="13112A5BA85F040EFA89FBCD0A5AFBB9" box="[1340,1406,1058,1083]" captionStart="FIGURE 3" captionStartId="5.[151,250,1882,1904]" captionTargetBox="[181,1408,345,1845]" captionTargetId="figure@5.[151,1436,337,1861]" captionTargetPageId="5" captionText="FIGURE 3. Anemadus perreaui sp. nov.: A, antenna ♂ (dorsal view); B, proleg ♂ (dorsal view); C, protarsus ♀ (dorsal view); D, pronotum ♂ (dorsal view); E, elytral apex ♂ (dorsoapical view); F, ventrite VIII ♀ (ventral view); G, genital segment ♂ (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view); K, apex of right paramere (dorsal view); L, apex of right paramere (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/264451/files/figure.png" pageId="3" pageNumber="285">Fig. 3</figureCitation>
A) long and slender, AL/HW = 1.63; length of 2th/3th = 1.24; 8th asymmetrical, wider than long; 10th slightly wider than long; 11th elongated pear-like.
</paragraph>
<paragraph id="8B9536DEA85F040EFF72FB7A0A31FA9B" blockId="3.[151,1437,590,2025]" pageId="3" pageNumber="285">
Pronotum (
<figureCitation id="13112A5BA85F040EFEFEFB7B0EB7FB2F" box="[331,403,1172,1197]" captionStart="FIGURE 3" captionStartId="5.[151,250,1882,1904]" captionTargetBox="[181,1408,345,1845]" captionTargetId="figure@5.[151,1436,337,1861]" captionTargetPageId="5" captionText="FIGURE 3. Anemadus perreaui sp. nov.: A, antenna ♂ (dorsal view); B, proleg ♂ (dorsal view); C, protarsus ♀ (dorsal view); D, pronotum ♂ (dorsal view); E, elytral apex ♂ (dorsoapical view); F, ventrite VIII ♀ (ventral view); G, genital segment ♂ (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view); K, apex of right paramere (dorsal view); L, apex of right paramere (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/264451/files/figure.png" pageId="3" pageNumber="285">Fig. 3</figureCitation>
D) transverse, widest at about basal 2/5, PW/PL = 1.69. Sides fairly curved, strongly narrowing forward but moderately and almost rectilinearly narrowing backward from widest; hind corners obtuse. Posterior edge slightly bisinuate. Surface covered with large and rough punctures, separated about 1.02.0 times of their diameter; interspaces smooth, without visable microsculpture. Postero-lateral impressions absent.
</paragraph>
<paragraph id="8B9536DEA85F040EFF72FACB0C9BFA73" blockId="3.[151,1437,590,2025]" pageId="3" pageNumber="285">
Elytra oval, widest at about basal 3/8, ELL/EW = 1.13. Sides regularly curved, gradually narrowing from widest to apices; apices rounded (
<figureCitation id="13112A5BA85F040EFD97FAA70D4CFAE3" box="[546,616,1352,1377]" captionStart="FIGURE 3" captionStartId="5.[151,250,1882,1904]" captionTargetBox="[181,1408,345,1845]" captionTargetId="figure@5.[151,1436,337,1861]" captionTargetPageId="5" captionText="FIGURE 3. Anemadus perreaui sp. nov.: A, antenna ♂ (dorsal view); B, proleg ♂ (dorsal view); C, protarsus ♀ (dorsal view); D, pronotum ♂ (dorsal view); E, elytral apex ♂ (dorsoapical view); F, ventrite VIII ♀ (ventral view); G, genital segment ♂ (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view); K, apex of right paramere (dorsal view); L, apex of right paramere (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/264451/files/figure.png" pageId="3" pageNumber="285">Fig. 3</figureCitation>
E). Sculpture of
<typeStatus id="5491887CA85F040EFC9EFAA60C7FFAE3" box="[811,859,1353,1377]" pageId="3" pageNumber="285">type</typeStatus>
«a» (sensu
<bibRefCitation id="EFBB4B2FA85F040EFC5EFAA70A30FAE3" author="Giachino" box="[1003,1300,1352,1377]" pageId="3" pageNumber="285" refString="Giachino, P. M. &amp; Vailati, D. (1993) Revisione degli Anemadinae Hatch, 1928 (Coleoptera Cholevidae). Monografie di Natura Bresciana, 18, 1 - 314." type="journal article" year="1993">Giachino &amp; Vailati, 1993</bibRefCitation>
): transverse striolations well-marked and irregular, longitudinal striae invisible. When immersed in glycerine, large primitive punctures visible within interior of cuticule, arranging into 8 irregular longitudinal rows (imperceptible in dry specimens) (
<figureCitation id="13112A5BA85F040EFE9FFA5B0E57FA4F" box="[298,371,1460,1485]" captionStart="FIGURE 2" captionStartId="4.[151,250,1646,1668]" captionTargetBox="[171,1415,216,1604]" captionTargetId="figure@4.[151,1436,193,1625]" captionTargetPageId="4" captionText="FIGURE 2. Anemadus perreaui sp. nov.: A, habitus ♂ (dorsal view; Baoxing, Ruobigou); B, habitus ♀ (dorsal view; Baoxing, Ruobigou); C, habitus ♀ (dorsal view; Baoxing, Ganyanggou); D, right elytron ♂ (dorsal view; in glycerine). Anemadus sichuanus sp. nov. ♂: E, habitus ♂ (dorsal view; Baoxing, Pujigou); F, habitus ♀ (dorsal view; Baoxing, Pujigou); G, right elytron ♂ (dorsal view; in glycerine). Scales: 1 mm." httpUri="https://zenodo.org/record/264450/files/figure.png" pageId="3" pageNumber="285">Fig. 2</figureCitation>
D). Sutural striae present but inconspicuous. Interspaces of striolations with microtrichiae, aligning into ca. 5 transverse lines. Metathoracic wings fully developed.
</paragraph>
<paragraph id="8B9536DEA85F040EFF72FA130A58F9DF" blockId="3.[151,1437,590,2025]" pageId="3" pageNumber="285">
Prolegs slender, with basal three protarsomeres weakly expanded (
<figureCitation id="13112A5BA85F040EFC0FFA130CDAF997" box="[954,1022,1532,1557]" captionStart="FIGURE 3" captionStartId="5.[151,250,1882,1904]" captionTargetBox="[181,1408,345,1845]" captionTargetId="figure@5.[151,1436,337,1861]" captionTargetPageId="5" captionText="FIGURE 3. Anemadus perreaui sp. nov.: A, antenna ♂ (dorsal view); B, proleg ♂ (dorsal view); C, protarsus ♀ (dorsal view); D, pronotum ♂ (dorsal view); E, elytral apex ♂ (dorsoapical view); F, ventrite VIII ♀ (ventral view); G, genital segment ♂ (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view); K, apex of right paramere (dorsal view); L, apex of right paramere (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/264451/files/figure.png" pageId="3" pageNumber="285">Fig. 3</figureCitation>
B): TW/BTW = 1.49. Protibiae with outer margin almost straight, while inner margin slightly expanded around middle. Profemora wider than protibiae. Mesotibiae distinctly curved, expansion of basal two mesotarsomeres imperceptible. Metatibiae almost straight.
</paragraph>
<paragraph id="8B9536DEA85F040EFF72F9870E99F927" blockId="3.[151,1437,590,2025]" pageId="3" pageNumber="285">
Genital segment (
<figureCitation id="13112A5BA85F040EFE25F9870EF7F903" box="[400,467,1640,1665]" captionStart="FIGURE 3" captionStartId="5.[151,250,1882,1904]" captionTargetBox="[181,1408,345,1845]" captionTargetId="figure@5.[151,1436,337,1861]" captionTargetPageId="5" captionText="FIGURE 3. Anemadus perreaui sp. nov.: A, antenna ♂ (dorsal view); B, proleg ♂ (dorsal view); C, protarsus ♀ (dorsal view); D, pronotum ♂ (dorsal view); E, elytral apex ♂ (dorsoapical view); F, ventrite VIII ♀ (ventral view); G, genital segment ♂ (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view); K, apex of right paramere (dorsal view); L, apex of right paramere (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/264451/files/figure.png" pageId="3" pageNumber="285">Fig. 3</figureCitation>
G) as typical for the genus, tergite IX with a pair of setae long and strong, as well as several fine setae apically.
</paragraph>
<paragraph id="8B9536DEA85F040EFF7BF95F0E51F8FF" blockId="3.[151,1437,590,2025]" pageId="3" pageNumber="285">
Aedeagus with median lobe moderately wide, weakly and almost rectilinearly narrowed towards a bifid apex in dorsal view, apical indentation wide, deep and subtriangular, and both branches rounded at apex (
<figureCitation id="13112A5BA85F040EFA84F93B0A5CF96F" box="[1329,1400,1748,1773]" captionStart="FIGURE 3" captionStartId="5.[151,250,1882,1904]" captionTargetBox="[181,1408,345,1845]" captionTargetId="figure@5.[151,1436,337,1861]" captionTargetPageId="5" captionText="FIGURE 3. Anemadus perreaui sp. nov.: A, antenna ♂ (dorsal view); B, proleg ♂ (dorsal view); C, protarsus ♀ (dorsal view); D, pronotum ♂ (dorsal view); E, elytral apex ♂ (dorsoapical view); F, ventrite VIII ♀ (ventral view); G, genital segment ♂ (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view); K, apex of right paramere (dorsal view); L, apex of right paramere (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/264451/files/figure.png" pageId="3" pageNumber="285">Fig. 3</figureCitation>
H). Paramere constricted subapically, apex dentiform, pointing outwards and bearing 5 setae, their arrangement as shown in
<figureCitation id="13112A5BA85F040EFEB0F8F30E75F8B7" box="[261,337,1820,1845]" captionStart="FIGURE 3" captionStartId="5.[151,250,1882,1904]" captionTargetBox="[181,1408,345,1845]" captionTargetId="figure@5.[151,1436,337,1861]" captionTargetPageId="5" captionText="FIGURE 3. Anemadus perreaui sp. nov.: A, antenna ♂ (dorsal view); B, proleg ♂ (dorsal view); C, protarsus ♀ (dorsal view); D, pronotum ♂ (dorsal view); E, elytral apex ♂ (dorsoapical view); F, ventrite VIII ♀ (ventral view); G, genital segment ♂ (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view); K, apex of right paramere (dorsal view); L, apex of right paramere (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/264451/files/figure.png" pageId="3" pageNumber="285">Figs. 3</figureCitation>
K&amp;L. In lateral view, median lobe almost straight, parameres wide and rounded at apex (
<figureCitation id="13112A5BA85F040EFA8BF8F30AA5F8B7" box="[1342,1409,1820,1845]" captionStart="FIGURE 3" captionStartId="5.[151,250,1882,1904]" captionTargetBox="[181,1408,345,1845]" captionTargetId="figure@5.[151,1436,337,1861]" captionTargetPageId="5" captionText="FIGURE 3. Anemadus perreaui sp. nov.: A, antenna ♂ (dorsal view); B, proleg ♂ (dorsal view); C, protarsus ♀ (dorsal view); D, pronotum ♂ (dorsal view); E, elytral apex ♂ (dorsoapical view); F, ventrite VIII ♀ (ventral view); G, genital segment ♂ (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view); K, apex of right paramere (dorsal view); L, apex of right paramere (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/264451/files/figure.png" pageId="3" pageNumber="285">Fig. 3</figureCitation>
J). Endophallus of
<typeStatus id="5491887CA85F040EFEF2F8AF0E8DF8DB" box="[327,425,1856,1881]" pageId="3" pageNumber="285" type="holotype">holotype</typeStatus>
everted, with a pair of spine rows on dorsal wall and a pair of large teeth pointing upwards at end (
<figureCitation id="13112A5BA85F040EFF5FF88B0E12F8FF" box="[234,310,1892,1917]" captionStart="FIGURE 3" captionStartId="5.[151,250,1882,1904]" captionTargetBox="[181,1408,345,1845]" captionTargetId="figure@5.[151,1436,337,1861]" captionTargetPageId="5" captionText="FIGURE 3. Anemadus perreaui sp. nov.: A, antenna ♂ (dorsal view); B, proleg ♂ (dorsal view); C, protarsus ♀ (dorsal view); D, pronotum ♂ (dorsal view); E, elytral apex ♂ (dorsoapical view); F, ventrite VIII ♀ (ventral view); G, genital segment ♂ (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view); K, apex of right paramere (dorsal view); L, apex of right paramere (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/264451/files/figure.png" pageId="3" pageNumber="285">Figs. 3</figureCitation>
HJ).
</paragraph>
<paragraph id="8B9536DEA85F040EFF72F8670DB3F86B" blockId="3.[151,1437,590,2025]" pageId="3" pageNumber="285">
<emphasis id="B95EEACCA85F040EFF72F8670E3FF822" box="[199,283,1928,1952]" italics="true" pageId="3" pageNumber="285">Female</emphasis>
. Similar to male in general appearance (
<figureCitation id="13112A5BA85F040EFCBBF8670C46F823" box="[782,866,1928,1953]" captionStart="FIGURE 2" captionStartId="4.[151,250,1646,1668]" captionTargetBox="[171,1415,216,1604]" captionTargetId="figure@4.[151,1436,193,1625]" captionTargetPageId="4" captionText="FIGURE 2. Anemadus perreaui sp. nov.: A, habitus ♂ (dorsal view; Baoxing, Ruobigou); B, habitus ♀ (dorsal view; Baoxing, Ruobigou); C, habitus ♀ (dorsal view; Baoxing, Ganyanggou); D, right elytron ♂ (dorsal view; in glycerine). Anemadus sichuanus sp. nov. ♂: E, habitus ♂ (dorsal view; Baoxing, Pujigou); F, habitus ♀ (dorsal view; Baoxing, Pujigou); G, right elytron ♂ (dorsal view; in glycerine). Scales: 1 mm." httpUri="https://zenodo.org/record/264450/files/figure.png" pageId="3" pageNumber="285">Figs. 2</figureCitation>
B&amp;C), but can be distinguished from following characteristics: pro- and mesotarsi simply linear (
<figureCitation id="13112A5BA85F040EFD5BF8430C1CF847" box="[750,824,1964,1989]" captionStart="FIGURE 3" captionStartId="5.[151,250,1882,1904]" captionTargetBox="[181,1408,345,1845]" captionTargetId="figure@5.[151,1436,337,1861]" captionTargetPageId="5" captionText="FIGURE 3. Anemadus perreaui sp. nov.: A, antenna ♂ (dorsal view); B, proleg ♂ (dorsal view); C, protarsus ♀ (dorsal view); D, pronotum ♂ (dorsal view); E, elytral apex ♂ (dorsoapical view); F, ventrite VIII ♀ (ventral view); G, genital segment ♂ (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view); K, apex of right paramere (dorsal view); L, apex of right paramere (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/264451/files/figure.png" pageId="3" pageNumber="285">Fig. 3</figureCitation>
C); ventrite VIII with spiculum ventrale short and subtriangularly narrowed anteriorly (
<figureCitation id="13112A5BA85F040EFD81F83F0D53F86B" box="[564,631,2000,2025]" captionStart="FIGURE 3" captionStartId="5.[151,250,1882,1904]" captionTargetBox="[181,1408,345,1845]" captionTargetId="figure@5.[151,1436,337,1861]" captionTargetPageId="5" captionText="FIGURE 3. Anemadus perreaui sp. nov.: A, antenna ♂ (dorsal view); B, proleg ♂ (dorsal view); C, protarsus ♀ (dorsal view); D, pronotum ♂ (dorsal view); E, elytral apex ♂ (dorsoapical view); F, ventrite VIII ♀ (ventral view); G, genital segment ♂ (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view); K, apex of right paramere (dorsal view); L, apex of right paramere (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/264451/files/figure.png" pageId="3" pageNumber="285">Fig. 3</figureCitation>
F).
</paragraph>
</subSubSection>
<caption id="DF556656A8580409FF22F9810DDEF962" httpUri="https://zenodo.org/record/264450/files/figure.png" pageId="4" pageNumber="286" targetBox="[171,1415,216,1604]" targetPageId="4">
<paragraph id="8B9536DEA8580409FF22F9810DDEF962" blockId="4.[151,1437,1646,1760]" pageId="4" pageNumber="286">
<emphasis id="B95EEACCA8580409FF22F9810D59F906" bold="true" box="[151,637,1646,1668]" pageId="4" pageNumber="286">
FIGURE 2.
<taxonomicName id="4C2A4D5DA8580409FE96F9810ED1F906" box="[291,501,1646,1668]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="286" phylum="Arthropoda" rank="species" species="perreaui" status="sp. nov.">
<emphasis id="B95EEACCA8580409FE96F9810ED1F906" bold="true" box="[291,501,1646,1668]" italics="true" pageId="4" pageNumber="286">Anemadus perreaui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D57B7A8580409FDB4F9810D7FF906" box="[513,603,1646,1668]" pageId="4" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
: A
</emphasis>
, habitus
<emphasis id="B95EEACCA8580409FD56F9810DD2F906" bold="true" box="[739,758,1646,1668]" pageId="4" pageNumber="286"></emphasis>
(dorsal view; Baoxing, Ruobigou);
<emphasis id="B95EEACCA8580409FBC8F9810BAAF906" bold="true" box="[1149,1166,1646,1668]" pageId="4" pageNumber="286">B</emphasis>
, habitus ♀ (dorsal view; Baoxing, Ruobigou);
<emphasis id="B95EEACCA8580409FEC9F9620EAAF921" bold="true" box="[380,398,1677,1699]" pageId="4" pageNumber="286">C</emphasis>
, habitus ♀ (dorsal view; Baoxing, Ganyanggou);
<emphasis id="B95EEACCA8580409FC1EF9620C99F921" bold="true" box="[939,957,1677,1699]" pageId="4" pageNumber="286">D</emphasis>
, right elytron
<emphasis id="B95EEACCA8580409FBE8F9620B54F921" bold="true" box="[1117,1136,1677,1699]" pageId="4" pageNumber="286"></emphasis>
(dorsal view; in glycerine).
<emphasis id="B95EEACCA8580409FF22F9440D2DF943" bold="true" box="[151,521,1707,1729]" pageId="4" pageNumber="286">
<taxonomicName id="4C2A4D5DA8580409FF22F9440E53F943" box="[151,375,1707,1729]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="286" phylum="Arthropoda" rank="species" species="sichuanus" status="sp. nov.">
<emphasis id="B95EEACCA8580409FF22F9440E53F943" bold="true" box="[151,375,1707,1729]" italics="true" pageId="4" pageNumber="286">Anemadus sichuanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D57B7A8580409FECBF9440EEBF943" box="[382,463,1707,1729]" pageId="4" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
♂: E
</emphasis>
, habitus
<emphasis id="B95EEACCA8580409FDD0F9440D5CF943" bold="true" box="[613,632,1707,1729]" pageId="4" pageNumber="286"></emphasis>
(dorsal view; Baoxing, Pujigou);
<emphasis id="B95EEACCA8580409FC67F9440CFBF943" bold="true" box="[978,991,1707,1729]" pageId="4" pageNumber="286">F</emphasis>
, habitus ♀(dorsal view; Baoxing, Pujigou);
<emphasis id="B95EEACCA8580409FF22F9250F8CF962" bold="true" box="[151,168,1738,1760]" pageId="4" pageNumber="286">G</emphasis>
, right elytron
<emphasis id="B95EEACCA8580409FE8DF9250E6FF962" bold="true" box="[312,331,1738,1760]" pageId="4" pageNumber="286"></emphasis>
(dorsal view; in glycerine). Scales: 1 mm.
</paragraph>
</caption>
<subSubSection id="C3306555A8580409FF72F8E80D01F8A2" box="[199,549,1799,1824]" pageId="4" pageNumber="286" type="distribution">
<paragraph id="8B9536DEA8580409FF72F8E80D01F8A2" blockId="4.[151,1437,1799,2007]" box="[199,549,1799,1824]" pageId="4" pageNumber="286">
<emphasis id="B95EEACCA8580409FF72F8E80E44F8A2" bold="true" box="[199,352,1799,1824]" pageId="4" pageNumber="286">Distribution.</emphasis>
<collectingCountry id="F33D764EA8580409FED2F8E80E8EF8A2" box="[359,426,1799,1824]" name="China" pageId="4" pageNumber="286">China</collectingCountry>
(Sichuan).
</paragraph>
</subSubSection>
<subSubSection id="C3306555A8580408FF72F8C40DC5FE9E" lastPageId="5" lastPageNumber="287" pageId="4" pageNumber="286" type="etymology">
<paragraph id="8B9536DEA8580408FF72F8C40DC5FE9E" blockId="4.[151,1437,1799,2007]" lastBlockId="5.[151,1436,151,285]" lastPageId="5" lastPageNumber="287" pageId="4" pageNumber="286">
<emphasis id="B95EEACCA8580409FF72F8C40E6AF8C6" bold="true" box="[199,334,1835,1860]" pageId="4" pageNumber="286">Etymology.</emphasis>
The specific epithet is dedicated to Dr. Michel Perreau (Université Paris 7, Paris,
<collectingCountry id="F33D764EA8580409FA92F8C30A5CF8C6" box="[1319,1400,1836,1860]" name="France" pageId="4" pageNumber="286">France</collectingCountry>
), a famous taxonomist on
<taxonomicName id="4C2A4D5DA8580409FE22F8A00D2EF8EA" box="[407,522,1871,1896]" class="Insecta" family="Leiodidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="286" phylum="Arthropoda" rank="family">Leiodidae</taxonomicName>
, for his excellent work on Chinese fauna and generous help to our study.
<emphasis id="B95EEACCA8580409FF72F89C0E1FF80E" bold="true" box="[199,315,1907,1932]" pageId="4" pageNumber="286">Remarks.</emphasis>
This new species well resembles
<taxonomicName id="4C2A4D5DA8580409FD69F89B0C9AF809" authority="Perreau" authorityName="Perreau" box="[732,958,1908,1932]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="286" phylum="Arthropoda" rank="species" species="ruzickai">
<emphasis id="B95EEACCA8580409FD69F89B0DC9F809" box="[732,749,1908,1931]" italics="true" pageId="4" pageNumber="286">A</emphasis>
.
<emphasis id="B95EEACCA8580409FCB7F89B0C78F80E" box="[770,860,1908,1932]" italics="true" pageId="4" pageNumber="286">ruzickai</emphasis>
Perreau
</taxonomicName>
and
<taxonomicName id="4C2A4D5DA8580409FBB7F89B0A3DF809" authority="Perreau" authorityName="Perreau" box="[1026,1305,1908,1932]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="286" phylum="Arthropoda" rank="species" species="wolongianus">
<emphasis id="B95EEACCA8580409FBB7F89B0B37F809" box="[1026,1043,1908,1931]" italics="true" pageId="4" pageNumber="286">A</emphasis>
.
<emphasis id="B95EEACCA8580409FB92F89B0B92F80E" box="[1063,1206,1908,1932]" italics="true" pageId="4" pageNumber="286">wolongianus</emphasis>
Perreau
</taxonomicName>
in general appearance and their
<typeStatus id="5491887CA8580409FE30F8770E91F832" box="[389,437,1944,1968]" pageId="4" pageNumber="286">type</typeStatus>
localities also with geographical proximity, but it is easily to distinguish it from the latter two by the combination of the following characteristics: antennomere length of 2th/3th = 1.24; aedeagus with median lobe moderately wide, weakly and almost rectilinearly narrowed towards apex when viewed dorsally, apical indentation wide, deep and subtriangular, and both branches rounded at apex; paramere constricted subapically, apex dentiform, pointing outwards when viewed dorsally, and wide and rounded at apex when viewed laterally; each paramere with 5 setae on inner margin of apex.
</paragraph>
</subSubSection>
</treatment>
</document>