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<document id="76195F58B5A66CD773305703DD791357" ID-CLB-Dataset="67686" ID-DOI="10.11646/phytotaxa.549.2.1" ID-GBIF-Dataset="01d0463f-e0c0-45e6-b52e-958ee36cdcdd" ID-ISSN="1179-3163" ID-Zenodo-Dep="6622694" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1654674188001" checkinUser="plazi" docAuthor="Swanepoel, Wessel &amp; Manzitto-Tripp, Erin A." docDate="2022" docId="038387B1881FFFCB48F2FCEAFE74D7F8" docLanguage="en" docName="phytotaxa.549.2.1.pdf" docOrigin="Phytotaxa 549 (2)" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Petalidium sesfonteinense Swanepoel &amp; E. Tripp 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="134" masterDocId="FFBAFFC9881EFFCC487AFFDEFFD4D52C" masterDocTitle="Petalidium sesfonteinense (Acanthaceae), a new species from the Kaokoveld, Namibia" masterLastPageNumber="135" masterPageNumber="127" pageNumber="128" updateTime="1699339088775" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="DED726E9BFB542C5AB5D7BC4541D8755" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7A14C7B28D6DC0E969CB45AC092035AB">
<mods:title id="80AC6DADDFE64623A5CFA9792E9AA6BF">Petalidium sesfonteinense (Acanthaceae), a new species from the Kaokoveld, Namibia</mods:title>
</mods:titleInfo>
<mods:name id="16C61EA6955F8299170C6A273C7FDBDF" type="personal">
<mods:role id="B8CA0D4FED3C7B4BAA10391B3C2AD86C">
<mods:roleTerm id="E5A45C5B6AF05B84A549F07094E5E6EE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="82EC21167D38A40E00E81253420FFE9A">Swanepoel, Wessel</mods:namePart>
<mods:affiliation id="C5A445FDA7269B62669F5DB53765F518">H. G. W. J. Schweickerdt Herbarium, Department of Plant and Soil Sciences, University of Pretoria, Pretoria, 0002 South Africa. Postal address: P. O. Box 21168, Windhoek, Namibia.</mods:affiliation>
</mods:name>
<mods:name id="4098A73A48D35D27485255C0DC851795" type="personal">
<mods:role id="179E64254DD58AC47F250ECBF4C35D62">
<mods:roleTerm id="3D34EA14F7B669D7EE8270D4FCD0A5B6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0E4DAE1FB31430CCB8FE64A403E45952">Manzitto-Tripp, Erin A.</mods:namePart>
<mods:affiliation id="9638E4B066305CF7AF420E324A515819">University of Colorado, Department of Ecology &amp; Evolutionary Biology, &amp; Museum of Natural History, UCB 350, Boulder, Colorado, 80309 U. S. A.</mods:affiliation>
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<mods:titleInfo id="FFB4E96F3F9FA5175D180C8175F68496">
<mods:title id="E3DEC35301BA3C623CC8FEEE9C956709">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="F05EDFAADD5D1B272CFB8E8B029575E5">
<mods:date id="E597DC5A4F408E30E3302160C691DDC8">2022</mods:date>
<mods:detail id="2B23A3CC3D60178F46D680EA56D0E5B2" type="pubDate">
<mods:number id="7B430F81949E41BFF2F69D246F769385">2022-06-08</mods:number>
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<mods:detail id="C1BA8657B86DC53FA04F0F48DEEA3B15" type="volume">
<mods:number id="4EAE39E12DEC849ADE839B3E7CE39052">549</mods:number>
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<mods:number id="BBDFBA6959E08CC2406E9479E8616407">2</mods:number>
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<mods:classification id="4231452744700822A1BE89B9BB43B8AF">journal article</mods:classification>
<mods:identifier id="4A1881984275EA2C29E0EC4613F338A6" type="CLB-Dataset">67686</mods:identifier>
<mods:identifier id="06FEF2933C65CF05AF4E32373CF2A736" type="DOI">10.11646/phytotaxa.549.2.1</mods:identifier>
<mods:identifier id="C0DDAAE770AEC99E477DB57D89ED8BA4" type="GBIF-Dataset">01d0463f-e0c0-45e6-b52e-958ee36cdcdd</mods:identifier>
<mods:identifier id="DFFDC0E0663678469BC7C295B0B8DA8C" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="60983BF6FB69F30579AF5DB70A5ECF03" type="Zenodo-Dep">6622694</mods:identifier>
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<treatment id="038387B1881FFFCB48F2FCEAFE74D7F8" ID-DOI="http://doi.org/10.5281/zenodo.6625062" ID-GBIF-Taxon="195810331" ID-Zenodo-Dep="6625062" LSID="urn:lsid:plazi:treatment:038387B1881FFFCB48F2FCEAFE74D7F8" httpUri="http://treatment.plazi.org/id/038387B1881FFFCB48F2FCEAFE74D7F8" lastPageId="7" lastPageNumber="134" pageId="1" pageNumber="128">
<subSubSection id="C330652C881FFFCD48F2FCEAFC40D662" box="[136,916,820,847]" pageId="1" pageNumber="128" type="nomenclature">
<paragraph id="8B9536A7881FFFCD48F2FCEAFC40D662" blockId="1.[136,916,820,847]" box="[136,916,820,847]" pageId="1" pageNumber="128">
<heading id="D0DD81CB881FFFCD48F2FCEAFC40D662" box="[136,916,820,847]" fontSize="11" level="2" pageId="1" pageNumber="128" reason="3">
<taxonomicName id="4C2A4D24881FFFCD48F2FCEAFD67D662" ID-CoL="9BXKS" authority="Swanepoel &amp; E.Tripp" authorityName="Swanepoel &amp; E. Tripp" authorityYear="2022" box="[136,691,820,847]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="128" phylum="Tracheophyta" rank="species" species="sesfonteinense" status="sp. nov.">
<emphasis id="B95EEAB5881FFFCD48F2FCEAFE61D662" bold="true" box="[136,437,820,847]" italics="true" pageId="1" pageNumber="128">Petalidium sesfonteinense</emphasis>
Swanepoel &amp; E.Tripp
</taxonomicName>
,
<emphasis id="B95EEAB5881FFFCD4AC4FCEBFCC7D662" box="[702,787,821,846]" italics="true" pageId="1" pageNumber="128">
<taxonomicNameLabel id="A26D57CE881FFFCD4AC4FCEBFCC7D662" box="[702,787,821,846]" pageId="1" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13112A22881FFFCD4B58FCEAFC5FD662" box="[802,907,820,846]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="4.[136,229,1915,1937]" captionStartId-1="5.[136,229,1963,1985]" captionStartId-2="6.[136,229,1414,1436]" captionTargetBox-0="[151,1435,187,1886]" captionTargetBox-1="[151,1435,389,1938]" captionTargetBox-2="[183,1404,190,1390]" captionTargetId-0="figure-17@4.[151,1435,187,1887]" captionTargetId-1="figure-123@5.[151,1435,389,1938]" captionTargetId-2="figure-17@6.[183,1404,190,1390]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 1. Petalidium sesfonteinense. A. Habit. Plant about 40 cm tall. B. Flower, shoots, and leaves; greyish leaves are still densely covered in trichomes; green leaves almost glabrous. Photographs: W. Swanepoel. Swanepoel 568." captionText-1="FIGURE 2. Petalidium sesfonteinense. Morphology of flowers. A–D. Flowers in front view showing variation in corolla colour and lobe margins, with two yellow nectar guides. E. Corolla in lateral view. F. Corolla in dorsal view. G. Corolla in ventral view. Scale bar = 5 mm. Photographs: W. Swanepoel. Swanepoel 568 (E, F &amp; G)." captionText-2="FIGURE 3. Known distribution of Petalidium sesfonteinense based on herbarium specimens." figureDoi-0="http://doi.org/10.5281/zenodo.6622696" figureDoi-1="http://doi.org/10.5281/zenodo.6622698" figureDoi-2="http://doi.org/10.5281/zenodo.6622700" httpUri-0="https://zenodo.org/record/6622696/files/figure.png" httpUri-1="https://zenodo.org/record/6622698/files/figure.png" httpUri-2="https://zenodo.org/record/6622700/files/figure.png" pageId="1" pageNumber="128">Figs. 1–3</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C330652C881FFFCD48F2FCA1FD24D00E" pageId="1" pageNumber="128" type="diagnosis">
<paragraph id="8B9536A7881FFFCD48F2FCA1FD24D00E" blockId="1.[136,1452,895,1386]" pageId="1" pageNumber="128">
<emphasis id="B95EEAB5881FFFCD48F2FCA1FEDFD6B9" bold="true" box="[136,267,895,917]" pageId="1" pageNumber="128">Diagnosis:—</emphasis>
A woody shrub up to
<quantity id="4CD29B42881FFFCD49A5FC5EFDD2D6B9" box="[479,518,896,917]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="128" unit="m" value="1.0">1 m</quantity>
tall, morphologically most similar to
<taxonomicName id="4C2A4D24881FFFCD4B0DFC5EFB9FD6B9" authorityName="Swanepoel" authorityYear="2020" box="[887,1099,896,917]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="128" phylum="Tracheophyta" rank="species" species="kaokoense">
<emphasis id="B95EEAB5881FFFCD4B0DFC5EFB9FD6B9" box="[887,1099,896,917]" italics="true" pageId="1" pageNumber="128">Petalidium kaokoense</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2A4D24881FFFCD4C06FC5EFB38D6B9" authorityName="Clarke" authorityYear="1899" box="[1148,1260,896,917]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="128" phylum="Tracheophyta" rank="species" species="variabile">
<emphasis id="B95EEAB5881FFFCD4C06FC5EFB38D6B9" box="[1148,1260,896,917]" italics="true" pageId="1" pageNumber="128">P. variabile</emphasis>
</taxonomicName>
; from
<taxonomicName id="4C2A4D24881FFFCD4D55FC5EFA7FD6B9" authorityName="Swanepoel" authorityYear="2020" box="[1327,1451,896,917]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="128" phylum="Tracheophyta" rank="species" species="kaokoense">
<emphasis id="B95EEAB5881FFFCD4D55FC5EFA7FD6B9" box="[1327,1451,896,917]" italics="true" pageId="1" pageNumber="128">P. kaokoense</emphasis>
</taxonomicName>
it differs in having an indumentum on vegetative parts of relatively slender dendritic trichomes (
<emphasis id="B95EEAB5881FFFCD4C1EFC7AFBACD695" box="[1124,1144,932,953]" italics="true" pageId="1" pageNumber="128">vs</emphasis>
. relatively stout stalked-stellate trichomes interspersed with dendritic trichomes), bracts oblanceolate (
<emphasis id="B95EEAB5881FFFCD4BF3FC16FC40D6F1" box="[905,916,968,989]" italics="true" pageId="1" pageNumber="128">v</emphasis>
s. linear-oblanceolate or linear-lanceolate), bracteoles symmetrically elliptic or narrowly ovate with long multicellular glandular trichomes and venation weak or not prominent (
<emphasis id="B95EEAB5881FFFCD4DEBFC32FA71D12D" box="[1425,1445,1004,1025]" italics="true" pageId="1" pageNumber="128">vs</emphasis>
. asymmetrically elliptic-oblong, long multicellular glandular trichomes absent, reticulation prominent), corolla with expanded portion of tube glabrous externally (
<emphasis id="B95EEAB5881FFFCD4A5AFBEAFDE0D165" box="[544,564,1076,1097]" italics="true" pageId="1" pageNumber="128">vs</emphasis>
. strigose), upper lobes connate for 20–30% of their length and obovate (vs. 50%, oblong), lobes variously coloured (white, pink, magenta, apricot, yellow, or cream) and in contrast with maroon throat and mouth (
<emphasis id="B95EEAB5881FFFCD4D28FB86FAB2D141" box="[1362,1382,1112,1133]" italics="true" pageId="1" pageNumber="128">vs</emphasis>
. lobes, throat, and mouth maroon), seeds cordate (
<emphasis id="B95EEAB5881FFFCD4A22FBA2FDB8D1BD" box="[600,620,1148,1169]" italics="true" pageId="1" pageNumber="128">vs</emphasis>
. discoid or ovate); from
<taxonomicName id="4C2A4D24881FFFCD4B20FBA2FC1DD1BD" authorityName="Clarke" authorityYear="1899" box="[858,969,1148,1169]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="128" phylum="Tracheophyta" rank="species" species="variabile">
<emphasis id="B95EEAB5881FFFCD4B20FBA2FC1DD1BD" box="[858,969,1148,1169]" italics="true" pageId="1" pageNumber="128">P. variabile</emphasis>
</taxonomicName>
it differs in indumentum on young stems different from that on leaves (
<emphasis id="B95EEAB5881FFFCD49F9FB7EFE43D199" box="[387,407,1184,1205]" italics="true" pageId="1" pageNumber="128">vs</emphasis>
. similar), indumentum on leaves dendritic (
<emphasis id="B95EEAB5881FFFCD4B41FB7EFC9BD199" box="[827,847,1184,1205]" italics="true" pageId="1" pageNumber="128">vs</emphasis>
. strigose), bracteoles usually with long multicellular trichomes (
<emphasis id="B95EEAB5881FFFCD48BFFB1AFF0DD1F5" box="[197,217,1220,1241]" italics="true" pageId="1" pageNumber="128">vs</emphasis>
. absent), expanded portion of corolla tube glabrous externally (
<emphasis id="B95EEAB5881FFFCD4B32FB1AFC88D1F5" box="[840,860,1220,1241]" italics="true" pageId="1" pageNumber="128">vs</emphasis>
. with short appressed simple trichomes), all corolla lobes of same colour and shade (
<emphasis id="B95EEAB5881FFFCD49D2FB36FE68D1D1" box="[424,444,1256,1277]" italics="true" pageId="1" pageNumber="128">vs</emphasis>
. front corolla lobe differently coloured or shaded than other lobes), capsule flattened ellipsoid or ovoid, sides rugose or smooth (
<emphasis id="B95EEAB5881FFFCD49D2FAD2FE17D00D" box="[424,451,1292,1313]" italics="true" pageId="1" pageNumber="128">vs.</emphasis>
flattened ovoid, sides smooth).
</paragraph>
</subSubSection>
<subSubSection id="C330652C881FFFCE48F2FAEEFF3CD6A4" lastPageId="2" lastPageNumber="129" pageId="1" pageNumber="128" type="materials_examined">
<paragraph id="8B9536A7881FFFCD48F2FAEEFBBAD046" blockId="1.[136,1452,895,1386]" pageId="1" pageNumber="128">
<emphasis id="B95EEAB5881FFFCD48F2FAEEFF16D069" bold="true" box="[136,194,1328,1349]" pageId="1" pageNumber="128">Type:</emphasis>
—
<materialsCitation id="3B423CFA881FFFCD48A1FAEEFBBED046" ID-GBIF-Occurrence="3804421301" collectingDate="2021-06-14" collectionCode="WIND, COLO, PRE, PRU" country="Namibia" elevation="477" location="Ganamub River" municipality="Eastern" pageId="1" pageNumber="128" specimenCount="1" stateProvince="Kunene" typeStatus="holotype">
<collectingCountry id="F33D7637881FFFCD48A1FAEEFE93D069" box="[219,327,1328,1349]" name="Namibia" pageId="1" pageNumber="128">NAMIBIA</collectingCountry>
.
<collectingRegion id="49EEF845881FFFCD492AFAEEFE3ED06A" box="[336,490,1328,1350]" country="Namibia" name="Kunene" pageId="1" pageNumber="128">Kunene Region</collectingRegion>
:
<collectingMunicipality id="6BF1ACDD881FFFCD4989FAEEFDEFD069" box="[499,571,1328,1349]" pageId="1" pageNumber="128">Eastern</collectingMunicipality>
tributary to
<location id="8EF5607C881FFFCD4ACEFAEEFC98D06A" LSID="urn:lsid:plazi:treatment:038387B1881FFFCB48F2FCEAFE74D7F8:8EF5607C881FFFCD4ACEFAEEFC98D06A" box="[692,844,1328,1350]" country="Namibia" municipality="Eastern" name="Ganamub River" pageId="1" pageNumber="128" stateProvince="Kunene">Ganamub River</location>
in mountainous area
<locationDeviation id="9A272F85881FFFCD4C67FAEEFA7CD06A" box="[1053,1448,1328,1350]" firstLocation="Giribesvlakte" pageId="1" pageNumber="128" secondLocation="Hoanib River">between Giribesvlakte and Hoanib River</locationDeviation>
, 1913AB,
<quantity id="4CD29B42881FFFCD4961FA8AFE8ED045" box="[283,346,1364,1385]" metricMagnitude="2" metricUnit="m" metricValue="4.77" pageId="1" pageNumber="128" unit="m" value="477.0">
<elevation id="0007D194881FFFCD4961FA8AFE8ED045" box="[283,346,1364,1385]" metricMagnitude="2" metricUnit="m" metricValue="4.77" pageId="1" pageNumber="128" unit="m" value="477.0">477 m</elevation>
</quantity>
,
<date id="FF941067881FFFCD4919FA8AFE31D045" box="[355,485,1364,1385]" pageId="1" pageNumber="128" value="2021-06-14">
<collectingDate id="EFD0E98F881FFFCD4919FA8AFE31D045" box="[355,485,1364,1385]" pageId="1" pageNumber="128" value="2021-06-14">14 June 2021</collectingDate>
</date>
,
<emphasis id="B95EEAB5881FFFCD4995FA8AFD55D045" box="[495,641,1364,1385]" italics="true" pageId="1" pageNumber="128">Swanepoel 568</emphasis>
(
<typeStatus id="54918805881FFFCD4AF4FA8AFD30D046" box="[654,740,1364,1386]" pageId="1" pageNumber="128" type="holotype">holotype</typeStatus>
<collectionCode id="ED3BAE62881FFFCD4A93FA8AFCE6D045" box="[745,818,1364,1385]" country="Namibia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13705" lsid="urn:lsid:biocol.org:col:13705" name="National Botanical Research Institute" pageId="1" pageNumber="128" type="Herbarium">WIND</collectionCode>
!; isotypes
<collectionCode id="ED3BAE62881FFFCD4BEEFA8AFC09D045" box="[916,989,1364,1385]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15540" lsid="urn:lsid:biocol.org:col:15540" name="University of Colorado Herbarium" pageId="1" pageNumber="128" type="Herbarium">COLO</collectionCode>
!,
<collectionCode id="ED3BAE62881FFFCD4B93FA8AFBC8D045" box="[1001,1052,1364,1385]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="1" pageNumber="128" type="Herbarium">PRE</collectionCode>
!,
<collectionCode id="ED3BAE62881FFFCD4C53FA8AFBB4D046" box="[1065,1120,1364,1386]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14594" lsid="urn:lsid:biocol.org:col:14594" name="University of Pretoria" pageId="1" pageNumber="128" type="Herbarium">PRU</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9536A7881FFFCE48F2FA47FC58D66C" blockId="1.[136,1452,1432,2071]" lastBlockId="2.[136,1452,158,2021]" lastPageId="2" lastPageNumber="129" pageId="1" pageNumber="128">
Erect woody shrub to
<quantity id="4CD29B42881FFFCD49F7FA46FE68D09F" box="[397,444,1432,1459]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="128" unit="m" value="1.0">1 m</quantity>
tall; all relatively young vegetative parts with dense indumentum of relatively slender white dendritic trichomes, glabrescent or nearly so.
<emphasis id="B95EEAB5881FFFCD4A93FA62FCFED0FA" box="[745,810,1468,1494]" italics="true" pageId="1" pageNumber="128">Stems</emphasis>
: main stem up to
<quantity id="4CD29B42881FFFCD4C72FA62FB8FD0FB" box="[1032,1115,1468,1495]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="128" unit="mm" value="80.0">80 mm</quantity>
diam., bark fissured, creamcoloured or greyish white; bark on distal stems peeling in long, thin, narrow strips; young stems quadrangular with very short simple glandular and short to long eglandular trichomes, often multicellular, also bi-furcating and dendritic or long-stalked stellate trichomes, or trichomes absent; pale green with cystoliths linear or appearing circular.
<emphasis id="B95EEAB5881FFFCD4D24F9F7FA78D36E" box="[1374,1452,1577,1602]" italics="true" pageId="1" pageNumber="128">Leaves</emphasis>
opposite and decussate, petiolate; laminae narrowly ovate, ovate or elliptic, rarely oblanceolate, 15–44 ×
<quantity id="4CD29B42881FFFCD4D42F992FA73D34B" box="[1336,1447,1612,1639]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.7" metricValueMin="0.8" pageId="1" pageNumber="128" unit="mm" value="17.5" valueMax="27.0" valueMin="8.0">8–27 mm</quantity>
, with dense indumentum of dendritic trichomes, grey to greyish green, becoming green to yellowish green with age, cystoliths inconspicuous; apices acute, obtuse or rounded, sometimes minutely apiculate, bases attenuate, decurrent, rarely rounded, margins entire; midribs prominent both sides, lateral veins 4 to 6 each side, usually not prominent, petioles up to
<quantity id="4CD29B42881FFFCD495EF902FEA6D3DB" box="[292,370,1756,1783]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="128" unit="mm" value="15.0">15 mm</quantity>
long.
<emphasis id="B95EEAB5881FFFCD49CDF902FDC7D3DA" box="[439,531,1756,1782]" italics="true" pageId="1" pageNumber="128">Flowers</emphasis>
in short dichasia, bracts foliaceous, oblanceolate, up to 15.0 ×
<quantity id="4CD29B42881FFFCD4CB6F902FAF7D3DB" box="[1228,1315,1756,1783]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="128" unit="mm" value="2.2">2.2 mm</quantity>
, apex acute, sessile; pedicels (below bracteoles)
<quantity id="4CD29B42881FFFCD4A62F8DEFD4BD237" box="[536,671,1792,1819]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="1" pageNumber="128" unit="mm" value="0.85" valueMax="1.2" valueMin="0.5">0.5–1.2 mm</quantity>
long; bracteoles symmetrically elliptic or narrowly ovate, coriaceous, forming prominent bulging on bracteole pair from which corolla limb emerges from one side, apex acute, sometimes slightly acuminate or asymmetric, pale green to yellowish green, reticulation slightly or not prominent, midrib straight, venation indistinct, pale green, ca. 10.6–13.3 ×
<quantity id="4CD29B42881FFFCD4ACCF8B2FC96D2AB" box="[694,834,1900,1927]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" metricValueMax="6.8" metricValueMin="5.4" pageId="1" pageNumber="128" unit="mm" value="6.1" valueMax="6.8" valueMin="5.4">5.4–6.8 mm</quantity>
, abaxially with dense glandular trichomes of various lengths, few to scattered long multicellular glandular trichomes, sometimes with very long multicellular sparsely branched eglandular simple or dendritic trichomes in addition, strigose towards base, sericous adaxially, margin lanate towards apex, cystoliths visible.
<taxonomicName id="4C2A4D24881FFFCD4983F806FDEED2DE" box="[505,570,2008,2034]" class="Demospongiae" family="Phloeodictyidae" genus="Calyx" kingdom="Animalia" order="Haplosclerida" pageId="1" pageNumber="128" phylum="Porifera" rank="genus">
<emphasis id="B95EEAB5881FFFCD4983F806FDEED2DE" box="[505,570,2008,2034]" italics="true" pageId="1" pageNumber="128">Calyx</emphasis>
</taxonomicName>
<quantity id="4CD29B42881FFFCD4A38F806FD1DD2DF" box="[578,713,2008,2035]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" metricValueMax="7.2" metricValueMin="6.6" pageId="1" pageNumber="128" unit="mm" value="6.9" valueMax="7.2" valueMin="6.6">6.6–7.2 mm</quantity>
long including basal tube of ca.
<quantity id="4CD29B42881FFFCD4C46F806FBABD2DF" box="[1084,1151,2008,2035]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="128" unit="mm" value="2.0">2 mm</quantity>
, strigose in places on both surfaces; lobes 4, lanceolate, acute, anticous lobe indistinctly bifid, anticous and upper ca. 5.6–7.8 ×
<quantity id="4CD29B42881FFFCD4CA9F822FA8EDD3B" box="[1235,1370,2044,2071]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="1" pageNumber="128" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.5–1.7 mm</quantity>
, folded lengthwise, laterals 5.6–7.0 ×
<quantity id="4CD29B42881CFFCE49ABFF40FD8DD595" box="[465,601,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="2" pageNumber="129" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.9–1.1 mm</quantity>
.
<emphasis id="B95EEAB5881CFFCE4A1BFF40FD63D594" box="[609,695,158,184]" italics="true" pageId="2" pageNumber="129">Corolla</emphasis>
<quantity id="4CD29B42881CFFCE4AC6FF40FCE7D595" box="[700,819,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="2" pageNumber="129" unit="mm" value="22.0" valueMax="24.0" valueMin="20.0">20–24 mm</quantity>
long with lobes straightened, narrow unexpanded portion cylindrical, slightly widening towards throat, flattened, ca.
<quantity id="4CD29B42881CFFCE4B67FF1CFCB8D5F1" box="[797,876,194,221]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="129" unit="mm" value="10.0">10 mm</quantity>
long, ca.
<quantity id="4CD29B42881CFFCE4BA2FF1CFBFAD5F1" box="[984,1070,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="129" unit="mm" value="3.2">3.2 mm</quantity>
diam, expanded portion ca.
<quantity id="4CD29B42881CFFCE4D11FF1CFA78D5F1" box="[1387,1452,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="129" unit="mm" value="6.0">6 mm</quantity>
long, outside glabrous, inside puberulous on area immediately above insertion of filaments up to mouth, otherwise glabrous; inside of anticous portion of throat yellow, terminating in two narrowly triangular separate markings (nectar guides) on proximal portion of lobes; lobes white, pink, magenta, apricot, yellow or cream, expanded portion (throat and mouth) maroon, front lobe including nectar guides maroon dotted proximally, all lobes patent and with long, stiff, patent, white eglandular trichomes, upper lobes obovate, 5.8–8.2 ×
<quantity id="4CD29B42881CFFCE4BF8FEA8FBDFD4BD" box="[898,1035,374,401]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="4.3" metricValueMin="3.5" pageId="2" pageNumber="129" unit="mm" value="3.9" valueMax="4.3" valueMin="3.5">3.5–4.3 mm</quantity>
, connate for 20–30% of their length, overlapping or not, apices rounded, retuse, lateral lobes oblong, 5.2–7.8 ×
<quantity id="4CD29B42881CFFCE4BBFFE44FB9AD499" box="[965,1102,410,437]" metricMagnitude="-3" metricUnit="m" metricValue="3.85" metricValueMax="4.5" metricValueMin="3.2" pageId="2" pageNumber="129" unit="mm" value="3.85" valueMax="4.5" valueMin="3.2">3.2–4.5 mm</quantity>
, apices retuse or truncate, front lobe broadly obovate, 5.5–8.8 ×
<quantity id="4CD29B42881CFFCE4981FE60FD51D4F5" box="[507,645,446,473]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" metricValueMax="7.2" metricValueMin="6.2" pageId="2" pageNumber="129" unit="mm" value="6.7" valueMax="7.2" valueMin="6.2">6.2–7.2 mm</quantity>
, apex rounded, retuse, all lobe margins entire or irregularly denticulate to dentate; palate prominently transversely 3–7-ribbed.
<emphasis id="B95EEAB5881CFFCE4A86FE3CFCB8D4D0" box="[764,876,482,508]" italics="true" pageId="2" pageNumber="129">Filaments</emphasis>
didynamous, inserted dorsally in throat, each long and short filament fused for ca.
<quantity id="4CD29B42881CFFCE4985FDD8FD8CD70D" box="[511,600,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="129" unit="mm" value="1.4">1.4 mm</quantity>
at base, fused part prominent, adnate to tube, free parts tapering towards apex, flattened, with scattered short eglandular trichomes, long filament
<quantity id="4CD29B42881CFFCE4BCEFDF4FBEFD769" box="[948,1083,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.7" metricValueMin="4.3" pageId="2" pageNumber="129" unit="mm" value="5.0" valueMax="5.7" valueMin="4.3">4.3–5.7 mm</quantity>
long, short filament
<quantity id="4CD29B42881CFFCE4D5FFDF4FA78D769" box="[1317,1452,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.3" metricValueMin="2.2" pageId="2" pageNumber="129" unit="mm" value="2.75" valueMax="3.3" valueMin="2.2">2.2–3.3 mm</quantity>
long; filament curtain reduced (
<bibRefCitation id="EFBB4B56881CFFCE4990FD90FD7BD744" author="Manktelow, M." box="[490,687,590,616]" pageId="2" pageNumber="129" pagination="129 - 160" refId="ref5064" refString="Manktelow, M. (2000) The filament curtain: a structure important to systematics and pollination biology in the Acanthaceae. Botanical Journal of the Linnean Society 133: 129 - 160. https: // doi. org / 10.1006 / bojl. 1999.0309" type="journal article" year="2000">Manktelow 2000</bibRefCitation>
); anthers 2-thecous, thecae oblong, subequal,
<quantity id="4CD29B42881CFFCE4CC9FD90FAEED745" box="[1203,1338,590,617]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.9" metricValueMin="2.1" pageId="2" pageNumber="129" unit="mm" value="2.5" valueMax="2.9" valueMin="2.1">2.1–2.9 mm</quantity>
long with scattered short glandular and eglandular trichomes, apex rounded, with minute spurs at base.
<emphasis id="B95EEAB5881CFFCE4CC9FDADFAE7D7A1" box="[1203,1331,627,653]" italics="true" pageId="2" pageNumber="129">Gynoecium</emphasis>
<quantity id="4CD29B42881CFFCE4D41FDACFF60D79D" metricMagnitude="-2" metricUnit="m" metricValue="1.41" metricValueMax="1.56" metricValueMin="1.26" pageId="2" pageNumber="129" unit="mm" value="14.1" valueMax="15.6" valueMin="12.6">12.6–15.6 mm</quantity>
long; ovary ovoid, laterally compressed, 1.5–2.0 × 1.5–1.7 × 1.0–
<quantity id="4CD29B42881CFFCE4BEAFD48FC32D79D" box="[912,998,662,689]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="129" unit="mm" value="1.2">1.2 mm</quantity>
, situated on fleshy disc, glabrous, ovules elliptic, ca.
<quantity id="4CD29B42881CFFCE4974FD64FEB2D7F9" box="[270,358,698,725]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="129" unit="mm" value="0.8">0.8 mm</quantity>
long; style filiform,
<quantity id="4CD29B42881CFFCE4A2CFD64FD39D7F9" box="[598,749,698,725]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" metricValueMax="1.25" metricValueMin="0.99" pageId="2" pageNumber="129" unit="mm" value="11.2" valueMax="12.5" valueMin="9.9">9.9–12.5 mm</quantity>
long, with scattered short eglandular trichomes, stigma lobes linear, slightly flattened, subequal, longer lobe
<quantity id="4CD29B42881CFFCE4ADDFD00FCE5D7D5" box="[679,817,734,761]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="2" pageNumber="129" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0.6–0.8 mm</quantity>
long, shorter lobe
<quantity id="4CD29B42881CFFCE4C74FD00FB4CD7D5" box="[1038,1176,734,761]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="2" pageNumber="129" unit="mm" value="0.6" valueMax="0.7" valueMin="0.5">0.5–0.7 mm</quantity>
long.
<emphasis id="B95EEAB5881CFFCE4C9EFD00FA94D7D4" box="[1252,1344,734,760]" italics="true" pageId="2" pageNumber="129">Capsule</emphasis>
flattened ellipsoid or ovoid, 6.2–6.5 × 3.3–3.6 × 1.4–2.0 mm, chestnut, glossy, sides rugose or smooth, glabrous.
<emphasis id="B95EEAB5881CFFCE4D76FCDCFA98D630" box="[1292,1356,770,796]" italics="true" pageId="2" pageNumber="129">Seeds</emphasis>
cordate, ca. 2.6 ×
<quantity id="4CD29B42881CFFCE4893FCF8FE95D66D" box="[233,321,806,833]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="129" unit="mm" value="1.8">1.8 mm</quantity>
, densely covered with white hygroscopic trichomes.
</paragraph>
<paragraph id="8B9536A7881CFFCE48C7FC94FF3CD6A4" blockId="2.[136,1452,158,2021]" pageId="2" pageNumber="129">
<emphasis id="B95EEAB5881CFFCE48C7FC94FE8BD648" bold="true" box="[189,351,842,868]" pageId="2" pageNumber="129">Phenology:—</emphasis>
Flowers have been recorded from March to June. Fruits have been recorded from April to October.
</paragraph>
</subSubSection>
<subSubSection id="C330652C881CFFCE48C7FC4CFEB7D1DC" pageId="2" pageNumber="129" type="distribution">
<paragraph id="8B9536A7881CFFCE48C7FC4CFA5DD134" blockId="2.[136,1452,158,2021]" pageId="2" pageNumber="129">
<emphasis id="B95EEAB5881CFFCE48C7FC4CFDD1D680" bold="true" box="[189,517,914,940]" pageId="2" pageNumber="129">Distribution and habitat:—</emphasis>
At present,
<taxonomicName id="4C2A4D24881CFFCE4AFCFC4CFC7ED680" authorityName="Swanepoel &amp; E. Tripp" authorityYear="2022" box="[646,938,914,940]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="sesfonteinense">
<emphasis id="B95EEAB5881CFFCE4AFCFC4CFC7ED680" box="[646,938,914,940]" italics="true" pageId="2" pageNumber="129">Petalidium sesfonteinense</emphasis>
</taxonomicName>
is only known from the vicinity of Sesfontein and the surrounding mountainous area up to Tomakas in the west and Warmquelle in the east (
<figureCitation id="13112A22881CFFCE4CBCFC68FAD8D6FC" box="[1222,1292,950,976]" captionStart="FIGURE 3" captionStartId="6.[136,229,1414,1436]" captionTargetBox="[183,1404,190,1390]" captionTargetId="figure-17@6.[183,1404,190,1390]" captionTargetPageId="6" captionText="FIGURE 3. Known distribution of Petalidium sesfonteinense based on herbarium specimens." figureDoi="http://doi.org/10.5281/zenodo.6622700" httpUri="https://zenodo.org/record/6622700/files/figure.png" pageId="2" pageNumber="129">Fig. 3</figureCitation>
). It occurs on hillsides, along seasonally dry riverbeds and at the base of rocky outcrops at elevations of
<quantity id="4CD29B42881CFFCE4CE8FC04FAF5D6D9" box="[1170,1313,986,1013]" metricMagnitude="2" metricUnit="m" metricValue="7.05" metricValueMax="10.7" metricValueMin="3.4" pageId="2" pageNumber="129" unit="m" value="705.0" valueMax="1070.0" valueMin="340.0">340–1070 m</quantity>
a.s.l., about
<quantity id="4CD29B42881CFFCE48F2FC20FED3D134" box="[136,263,1022,1049]" metricMagnitude="5" metricUnit="m" metricValue="1.0" metricValueMax="1.25" metricValueMin="0.75" pageId="2" pageNumber="129" unit="km" value="100.0" valueMax="125.0" valueMin="75.0">75–125 km</quantity>
from the Atlantic Ocean. Average annual rainfall in the area is
<quantity id="4CD29B42881CFFCE4BB1FC20FBB4D135" box="[971,1120,1022,1049]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="2" pageNumber="129" unit="mm" value="125.0" valueMax="150.0" valueMin="100.0">100–150 mm</quantity>
(
<bibRefCitation id="EFBB4B56881CFFCE4C14FC20FAA9D134" author="Mendelsohn, J. &amp; Jarvis, A. &amp; Roberts, C. &amp; Robertson, T." box="[1134,1405,1022,1048]" pageId="2" pageNumber="129" refId="ref5111" refString="Mendelsohn, J., Jarvis, A., Roberts, C. &amp; Robertson, T. (2002) Atlas of Namibia. Philip, Cape Town, 200 pp." type="book" year="2002">
Mendelsohn
<emphasis id="B95EEAB5881CFFCE4D7BFC21FAEDD134" box="[1281,1337,1022,1048]" italics="true" pageId="2" pageNumber="129">et al.</emphasis>
2002
</bibRefCitation>
).
</paragraph>
<paragraph id="8B9536A7881CFFCE48C7FBFCFEB7D1DC" blockId="2.[136,1452,158,2021]" pageId="2" pageNumber="129">
<emphasis id="B95EEAB5881CFFCE48C7FBFCFE04D110" bold="true" box="[189,464,1058,1084]" pageId="2" pageNumber="129">Conservation status:—</emphasis>
<taxonomicName id="4C2A4D24881CFFCE49ABFBFCFD21D110" authorityName="Swanepoel &amp; E. Tripp" authorityYear="2022" box="[465,757,1058,1084]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="sesfonteinense">
<emphasis id="B95EEAB5881CFFCE49ABFBFCFD21D110" box="[465,757,1058,1084]" italics="true" pageId="2" pageNumber="129">Petalidium sesfonteinense</emphasis>
</taxonomicName>
is locally common and probably more widespread in suitable habitats than currently recorded. It is here considered not in immediate conservation danger, because it occurs in sparsely to unpopulated areas and does not seem to be utilised by humans. However, it is browsed by livestock (goats) of the local inhabitants (pers. obs.). The area of occupancy is estimated at &lt;
<quantity id="4CD29B42881CFFCE4BB1FB50FBEFD187" box="[971,1083,1166,1195]" metricMagnitude="7" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="129" unit="km" value="20000.0">20000 km</quantity>
² (
<quantity id="4CD29B42881CFFCE4C2BFB50FB61D187" box="[1105,1205,1166,1195]" metricMagnitude="6" metricUnit="m" metricValue="2.25" pageId="2" pageNumber="129" unit="km" value="2250.0">2250 km</quantity>
²) with less than 10 (8) subpopulations. However, since no decline in population size or numbers is known, it is here ranked as Least Concern (LC) (
<bibRefCitation id="EFBB4B56881CFFCE48B7FB08FE83D1DC" author="IUCN &amp; Cambridge, U. K." box="[205,343,1238,1264]" pageId="2" pageNumber="129" refId="ref5029" refString="IUCN (2012) IUCN red list categories and criteria: Vers. 3.1. 2 nd edn. Gland, Switzerland and Cambridge, U. K., iv + 32 pp." type="book" year="2012">IUCN 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C330652C881CFFCB48C7FB24FE74D7F8" lastPageId="7" lastPageNumber="134" pageId="2" pageNumber="129" type="etymology">
<paragraph id="8B9536A7881CFFCE48C7FB24FA71D038" blockId="2.[136,1452,158,2021]" box="[189,1445,1274,1301]" pageId="2" pageNumber="129">
<emphasis id="B95EEAB5881CFFCE48C7FB24FEB0D038" bold="true" box="[189,356,1274,1300]" pageId="2" pageNumber="129">Etymology:—</emphasis>
The specific epithet refers to Sesfontein, a village in northwestern
<collectingCountry id="F33D7637881CFFCE4C36FB24FB78D038" box="[1100,1196,1274,1300]" name="Namibia" pageId="2" pageNumber="129">Namibia</collectingCountry>
near the
<typeStatus id="54918805881CFFCE4D6EFB25FA90D039" box="[1300,1348,1275,1301]" pageId="2" pageNumber="129">type</typeStatus>
locality.
</paragraph>
<paragraph id="8B9536A7881CFFCE48C7FAC0FDCED070" blockId="2.[136,1452,158,2021]" pageId="2" pageNumber="129">
<emphasis id="B95EEAB5881CFFCE48C7FAC0FEF2D014" bold="true" box="[189,294,1310,1336]" pageId="2" pageNumber="129">Notes:—</emphasis>
Some of the morphological features to distinguish between
<taxonomicName id="4C2A4D24881CFFCE4BC2FAC0FB0ED014" authorityName="Swanepoel &amp; E. Tripp" authorityYear="2022" box="[952,1242,1310,1336]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="sesfonteinense">
<emphasis id="B95EEAB5881CFFCE4BC2FAC0FB0ED014" box="[952,1242,1310,1336]" italics="true" pageId="2" pageNumber="129">Petalidium sesfonteinense</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A4D24881CFFCE4C9DFAC1FAA2D014" authorityName="Swanepoel" authorityYear="2020" box="[1255,1398,1310,1336]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="kaokoense">
<emphasis id="B95EEAB5881CFFCE4C9DFAC1FAA2D014" box="[1255,1398,1310,1336]" italics="true" pageId="2" pageNumber="129">P. kaokoense</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C2A4D24881CFFCE48F2FA9DFEDDD070" authorityName="Clarke" authorityYear="1899" box="[136,265,1346,1372]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="variabile">
<emphasis id="B95EEAB5881CFFCE48F2FA9DFEDDD070" box="[136,265,1346,1372]" italics="true" pageId="2" pageNumber="129">P. variabile</emphasis>
</taxonomicName>
are provided in
<tableCitation id="C6A8031C881CFFCE49BAFA9CFDC7D070" box="[448,531,1346,1372]" captionStart="TABLE 1" captionStartId="3.[136,224,160,184]" captionTargetPageId="3" captionText="TABLE 1. Prominent morphological differences between Petalidium sesfonteinense, P. kaokoense, and P. variabile." httpUri="http://table.plazi.org/id/DF55662F881DFFCF48F2FF7EFA8BD594" pageId="2" pageNumber="129" tableUuid="DF55662F881DFFCF48F2FF7EFA8BD594">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8B9536A7881CFFCE48C7FAB8FC57D21C" blockId="2.[136,1452,158,2021]" pageId="2" pageNumber="129">
Hitherto in herbaria,
<taxonomicName id="4C2A4D24881CFFCE49D2FAB8FD1FD0AC" authorityName="Swanepoel &amp; E. Tripp" authorityYear="2022" box="[424,715,1382,1408]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="sesfonteinense">
<emphasis id="B95EEAB5881CFFCE49D2FAB8FD1FD0AC" box="[424,715,1382,1408]" italics="true" pageId="2" pageNumber="129">Petalidium sesfonteinense</emphasis>
</taxonomicName>
has mostly been confused with
<taxonomicName id="4C2A4D24881CFFCE4C4BFAB9FB65D0AC" authorityName="Clarke" authorityYear="1899" box="[1073,1201,1382,1408]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="variabile">
<emphasis id="B95EEAB5881CFFCE4C4BFAB9FB65D0AC" box="[1073,1201,1382,1408]" italics="true" pageId="2" pageNumber="129">P. variabile</emphasis>
</taxonomicName>
, a species from which it differs in indumentum, leaf, inflorescence, and flower characters (
<tableCitation id="C6A8031C881CFFCE4B04FA54FC07D088" box="[894,979,1418,1444]" captionStart="TABLE 1" captionStartId="3.[136,224,160,184]" captionTargetPageId="3" captionText="TABLE 1. Prominent morphological differences between Petalidium sesfonteinense, P. kaokoense, and P. variabile." httpUri="http://table.plazi.org/id/DF55662F881DFFCF48F2FF7EFA8BD594" pageId="2" pageNumber="129" tableUuid="DF55662F881DFFCF48F2FF7EFA8BD594">Table 1</tableCitation>
&amp;
<figureCitation id="13112A22881CFFCE4B8CFA54FBEED088" box="[1014,1082,1418,1444]" captionStart="FIGURE 4" captionStartId="6.[136,229,1974,1996]" captionTargetBox="[183,1403,1462,1950]" captionTargetId="figure-31@6.[183,1403,1462,1950]" captionTargetPageId="6" captionText="FIGURE 4.Flowers of twospecies related toPetalidium sesfonteinense,for purposesof comparison.A.P.kaokoense.Notethe one-coloured lobes of similar shade with relatively long yellow nectar guides (Hartmann Mountains, Kaokoveld). B. P. variabile. Note the difference in shade between lobes and the relatively short yellow nectar guides (Okahandja, central Namibia). Photographs: W. Swanepoel." figureDoi="http://doi.org/10.5281/zenodo.6622702" httpUri="https://zenodo.org/record/6622702/files/figure.png" pageId="2" pageNumber="129">Fig. 4</figureCitation>
). Distribution of the two species, does not overlap:
<taxonomicName id="4C2A4D24881CFFCE4923FA71FDC9D0E4" authorityName="Swanepoel &amp; E. Tripp" authorityYear="2022" box="[345,541,1454,1481]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="sesfonteinense">
<emphasis id="B95EEAB5881CFFCE4923FA71FEBED0E4" box="[345,362,1455,1480]" italics="true" pageId="2" pageNumber="129">P</emphasis>
.
<emphasis id="B95EEAB5881CFFCE4901FA70FDC9D0E4" box="[379,541,1454,1480]" italics="true" pageId="2" pageNumber="129">sesfonteinense</emphasis>
</taxonomicName>
occurs in northwestern
<collectingCountry id="F33D7637881CFFCE4B4CFA70FC42D0E4" box="[822,918,1454,1480]" name="Namibia" pageId="2" pageNumber="129">Namibia</collectingCountry>
in the vicinity of Sesfontein, and
<taxonomicName id="4C2A4D24881CFFCE4D52FA71FA7FD0E4" authorityName="Clarke" authorityYear="1899" box="[1320,1451,1454,1480]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="variabile">
<emphasis id="B95EEAB5881CFFCE4D52FA71FA7FD0E4" box="[1320,1451,1454,1480]" italics="true" pageId="2" pageNumber="129">P. variabile</emphasis>
</taxonomicName>
from the Bergsig area
<quantity id="4CD29B42881CFFCE49FAFA0CFE01D0C0" box="[384,469,1490,1516]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="129" unit="km" value="100.0">100 km</quantity>
to the south-southeast and southwards to west central
<collectingCountry id="F33D7637881CFFCE4C49FA0CFB47D0C0" box="[1075,1171,1490,1516]" name="Namibia" pageId="2" pageNumber="129">Namibia</collectingCountry>
and in northern
<collectingCountry id="F33D7637881CFFCE4D31FA0CFA7FD0C0" box="[1355,1451,1490,1516]" name="Namibia" pageId="2" pageNumber="129">Namibia</collectingCountry>
near Otavi and Grootfontein. Records of
<taxonomicName id="4C2A4D24881CFFCE4A2FFA29FD02D33C" authorityName="Clarke" authorityYear="1899" box="[597,726,1526,1552]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="variabile">
<emphasis id="B95EEAB5881CFFCE4A2FFA29FD02D33C" box="[597,726,1526,1552]" italics="true" pageId="2" pageNumber="129">P. variabile</emphasis>
</taxonomicName>
in
<collectingCountry id="F33D7637881CFFCE4A83FA28FC9FD33C" box="[761,843,1526,1552]" name="Angola" pageId="2" pageNumber="129">Angola</collectingCountry>
have not yet been vetted by the first author and might represent a different taxon.
<taxonomicName id="4C2A4D24881CFFCE49CDF9C4FD0ED318" authorityName="Swanepoel &amp; E. Tripp" authorityYear="2022" box="[439,730,1562,1588]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="sesfonteinense">
<emphasis id="B95EEAB5881CFFCE49CDF9C4FD0ED318" box="[439,730,1562,1588]" italics="true" pageId="2" pageNumber="129">Petalidium sesfonteinense</emphasis>
</taxonomicName>
can also be confused with
<taxonomicName id="4C2A4D24881CFFCE4C7BF9C5FB75D318" authorityName="Meyer" authorityYear="1973" box="[1025,1185,1562,1588]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="ohopohense">
<emphasis id="B95EEAB5881CFFCE4C7BF9C5FB75D318" box="[1025,1185,1562,1588]" italics="true" pageId="2" pageNumber="129">P. ohopohense</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A4D24881CFFCE4CD7F9C5FA70D318" authorityName="Merxm. &amp; Hainz" authorityYear="1955" box="[1197,1444,1562,1588]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="pilosi-bracteolatum">
<emphasis id="B95EEAB5881CFFCE4CD7F9C5FA70D318" box="[1197,1444,1562,1588]" italics="true" pageId="2" pageNumber="129">P. pilosi-bracteolatum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A4D24881CFFCE48F2F9E1FE87D374" authorityName="Meyer" authorityYear="1961" box="[136,339,1599,1624]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="rossmannianum">
<emphasis id="B95EEAB5881CFFCE48F2F9E1FE87D374" box="[136,339,1599,1624]" italics="true" pageId="2" pageNumber="129">P. rossmannianum</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C2A4D24881CFFCE49F0F9E1FDF5D374" authorityName="Moore" authorityYear="1880" box="[394,545,1598,1624]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="welwitschii">
<emphasis id="B95EEAB5881CFFCE49F0F9E1FDF5D374" box="[394,545,1598,1624]" italics="true" pageId="2" pageNumber="129">P.welwitschii</emphasis>
</taxonomicName>
, especially in herbarium specimens where fresh flowering material is not available. However, distribution of
<taxonomicName id="4C2A4D24881CFFCE49E4F9BDFD8ED350" authorityName="Swanepoel &amp; E. Tripp" authorityYear="2022" box="[414,602,1634,1660]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="sesfonteinense">
<emphasis id="B95EEAB5881CFFCE49E4F9BDFD8ED350" box="[414,602,1634,1660]" italics="true" pageId="2" pageNumber="129">P. sesfonteinense</emphasis>
</taxonomicName>
does not overlap with these species, with
<taxonomicName id="4C2A4D24881CFFCE4C52F9BDFACBD350" authorityName="Merxm. &amp; Hainz" authorityYear="1955" box="[1064,1311,1634,1660]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="pilosi-bracteolatum">
<emphasis id="B95EEAB5881CFFCE4C52F9BDFACBD350" box="[1064,1311,1634,1660]" italics="true" pageId="2" pageNumber="129">P. pilosi-bracteolatum</emphasis>
</taxonomicName>
occurring to the south and the other three species to the east or the north of
<taxonomicName id="4C2A4D24881CFFCE4B41F959FC23D38C" authorityName="Swanepoel &amp; E. Tripp" authorityYear="2022" box="[827,1015,1670,1696]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="sesfonteinense">
<emphasis id="B95EEAB5881CFFCE4B41F959FC23D38C" box="[827,1015,1670,1696]" italics="true" pageId="2" pageNumber="129">P. sesfonteinense</emphasis>
</taxonomicName>
. Other species of
<taxonomicName id="4C2A4D24881CFFCE4CC5F958FAEED38C" authorityName="Nees von Esenbeck" authorityYear="1832" box="[1215,1338,1670,1696]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="genus">
<emphasis id="B95EEAB5881CFFCE4CC5F958FAEED38C" box="[1215,1338,1670,1696]" italics="true" pageId="2" pageNumber="129">Petalidium</emphasis>
</taxonomicName>
occurring within the range of
<taxonomicName id="4C2A4D24881CFFCE490BF975FDE6D3E8" authorityName="Swanepoel &amp; E. Tripp" authorityYear="2022" box="[369,562,1706,1732]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="sesfonteinense">
<emphasis id="B95EEAB5881CFFCE490BF975FDE6D3E8" box="[369,562,1706,1732]" italics="true" pageId="2" pageNumber="129">P. sesfonteinense</emphasis>
</taxonomicName>
are
<taxonomicName id="4C2A4D24881CFFCE4A13F975FBD5D3E9" authority="Meyer (1967: 505)" authorityName="Meyer" authorityPageNumber="505" authorityYear="1967" box="[617,1025,1706,1733]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="angustitibum">
<emphasis id="B95EEAB5881CFFCE4A13F975FCCED3E8" box="[617,794,1706,1732]" italics="true" pageId="2" pageNumber="129">P. angustitibum</emphasis>
<bibRefCitation id="EFBB4B56881CFFCE4B5FF975FBD5D3E9" author="Meyer, P. G." box="[805,1025,1706,1733]" pageId="2" pageNumber="129" pagination="505 - 515" refId="ref5198" refString="Meyer, P. G. (1967) Beitrage zur kenntnis der Acanthaceen Sudwestafrikas. Mitteilungen der Botanischen Staatssammlung Munchen 6: 505 - 515. [https: // www. biodiversitylibrary. org / item / 51962 # page / 537 / mode / 1 up]" type="journal article" year="1967">Meyer (1967: 505)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2A4D24881CFFCE4C6BF975FB73D3E8" authorityName="Moore" authorityYear="1880" box="[1041,1191,1707,1732]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="coccineum">
<emphasis id="B95EEAB5881CFFCE4C6BF975FB73D3E8" box="[1041,1191,1707,1732]" italics="true" pageId="2" pageNumber="129">P. coccineum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A4D24881CFFCE4CC3F975FA87D3E8" authorityName="Moore" authorityYear="1880" box="[1209,1363,1706,1732]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="halimoides">
<emphasis id="B95EEAB5881CFFCE4CC3F975FA87D3E8" box="[1209,1363,1706,1732]" italics="true" pageId="2" pageNumber="129">P. halimoides</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C2A4D24881CFFCE4DEDF975FF30D3C4" authorityName="Clarke" authorityYear="1899" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="lanatum">
<emphasis id="B95EEAB5881CFFCE4DEDF975FF30D3C4" italics="true" pageId="2" pageNumber="129">P. lanatum</emphasis>
</taxonomicName>
. All the mentioned species, including
<taxonomicName id="4C2A4D24881CFFCE4AF4F911FC9FD3C4" authorityName="Swanepoel &amp; E. Tripp" authorityYear="2022" box="[654,843,1742,1768]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="sesfonteinense">
<emphasis id="B95EEAB5881CFFCE4AF4F911FC9FD3C4" box="[654,843,1742,1768]" italics="true" pageId="2" pageNumber="129">P. sesfonteinense</emphasis>
</taxonomicName>
, are from the group composed of plants with irregular, four-parted calyces (
<bibRefCitation id="EFBB4B56881CFFCE490BF92CFDE2D220" author="Obermeijer, A. A." box="[369,566,1778,1804]" pageId="2" pageNumber="129" pagination="151 - 162" refId="ref5433" refString="Obermeijer, A. A. (1936) The South African species of Petalidium. Annals of the Transvaal Museum 18: 151 - 162." type="journal article" year="1936">Obermeijer 1936</bibRefCitation>
,
<bibRefCitation id="EFBB4B56881CFFCE4A39F92CFCDDD220" author="Tripp, E. A. &amp; Tsai, Y. E. &amp; Zhuang, Y. &amp; Dexter, K. G." box="[579,777,1778,1805]" pageId="2" pageNumber="129" pagination="1 - 17" refId="ref5642" refString="Tripp, E. A., Tsai, Y. E., Zhuang, Y. &amp; Dexter, K. G. (2017) RADseq dataset with 90 % missing data fully resolves recent radiation of Petalidium (Acanthaceae) in the ultra-arid deserts of Namibia. Ecology and Evolution 7: 1 - 17. https: // doi. org / 10.1002 / ece 3.3274" type="journal article" year="2017">
Tripp
<emphasis id="B95EEAB5881CFFCE4AF3F92DFD68D220" box="[649,700,1778,1804]" italics="true" pageId="2" pageNumber="129">et al</emphasis>
. 2017
</bibRefCitation>
). A key to
<emphasis id="B95EEAB5881CFFCE4BFDF92DFB3CD220" box="[903,1256,1778,1804]" italics="true" pageId="2" pageNumber="129">
<taxonomicName id="4C2A4D24881CFFCE4BFDF92DFB9DD220" authorityName="Swanepoel &amp; E. Tripp" authorityYear="2022" box="[903,1097,1778,1804]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="sesfonteinense">P. sesfonteinense</taxonomicName>
,
<taxonomicName id="4C2A4D24881CFFCE4C2CF92DFB3CD220" authorityName="Swanepoel" authorityYear="2020" box="[1110,1256,1778,1804]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="kaokoense">P. kaokoense</taxonomicName>
</emphasis>
,
<taxonomicName id="4C2A4D24881CFFCE4C82F92DFAAED220" authorityName="Clarke" authorityYear="1899" box="[1272,1402,1778,1804]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="species" species="variabile">
<emphasis id="B95EEAB5881CFFCE4C82F92DFAAED220" box="[1272,1402,1778,1804]" italics="true" pageId="2" pageNumber="129">P. variabile</emphasis>
</taxonomicName>
and other species of
<taxonomicName id="4C2A4D24881CFFCE4945F8C8FE6ED21C" authorityName="Nees von Esenbeck" authorityYear="1832" box="[319,442,1814,1840]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="129" phylum="Tracheophyta" rank="genus">
<emphasis id="B95EEAB5881CFFCE4945F8C8FE6ED21C" box="[319,442,1814,1840]" italics="true" pageId="2" pageNumber="129">Petalidium</emphasis>
</taxonomicName>
within its distribution range is provided.
</paragraph>
<paragraph id="8B9536A7881CFFCE48C7F8E4FBE5D2C9" blockId="2.[136,1452,158,2021]" pageId="2" pageNumber="129">
Indumentum is very useful in differentiating among the mentioned taxa and consists mainly of
<specimenCount id="9D2CFD2E881CFFCE4D77F8E4FA5ED279" box="[1293,1418,1850,1877]" pageId="2" pageNumber="129" type="generic" typeStatus="types">three types</specimenCount>
of trichomes: simple (unbranched), stellate, and dendritic. The simple trichomes are either single or multi-celled and gland-tipped or not. The stellate trichomes are either stalked or sessile with arms radiating from a common point. The dendritic trichomes are stalked with the arms spirally arranged, opposite and distichous, or opposite and decussate. Each taxon has a unique combination of trichomes (see
<tableCitation id="C6A8031C881CFFCE4A83F814FC98D2C8" box="[761,844,1994,2020]" captionStart="TABLE 1" captionStartId="3.[136,224,160,184]" captionTargetPageId="3" captionText="TABLE 1. Prominent morphological differences between Petalidium sesfonteinense, P. kaokoense, and P. variabile." httpUri="http://table.plazi.org/id/DF55662F881DFFCF48F2FF7EFA8BD594" pageId="2" pageNumber="129" tableUuid="DF55662F881DFFCF48F2FF7EFA8BD594">Table 1</tableCitation>
and key to species).
</paragraph>
<caption id="DF55662F881DFFCF48F2FF7EFA8BD594" ID-Table-UUID="DF55662F881DFFCF48F2FF7EFA8BD594" box="[136,1375,160,184]" httpUri="http://table.plazi.org/id/DF55662F881DFFCF48F2FF7EFA8BD594" pageId="3" pageNumber="130" startId="3.[136,224,160,184]" targetBox="[144,1440,203,2063]" targetIsTable="true" targetPageId="3">
<paragraph id="8B9536A7881DFFCF48F2FF7EFA8BD594" blockId="3.[136,1375,160,184]" box="[136,1375,160,184]" pageId="3" pageNumber="130">
<emphasis id="B95EEAB5881DFFCF48F2FF7EFF2FD594" bold="true" box="[136,251,160,184]" pageId="3" pageNumber="130">TABLE 1.</emphasis>
Prominent morphological differences between
<taxonomicName id="4C2A4D24881DFFCF4A8CFF7EFBD8D594" authorityName="Swanepoel &amp; E. Tripp" authorityYear="2022" box="[758,1036,160,184]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="130" phylum="Tracheophyta" rank="species" species="sesfonteinense">
<emphasis id="B95EEAB5881DFFCF4A8CFF7EFBD8D594" box="[758,1036,160,184]" italics="true" pageId="3" pageNumber="130">Petalidium sesfonteinense</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A4D24881DFFCF4C63FF7FFB76D594" authorityName="Swanepoel" authorityYear="2020" box="[1049,1186,160,184]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="130" phylum="Tracheophyta" rank="species" species="kaokoense">
<emphasis id="B95EEAB5881DFFCF4C63FF7FFB76D594" box="[1049,1186,160,184]" italics="true" pageId="3" pageNumber="130">P. kaokoense</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C2A4D24881DFFCF4CA7FF7FFA8FD594" authorityName="Clarke" authorityYear="1899" box="[1245,1371,160,184]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="130" phylum="Tracheophyta" rank="species" species="variabile">P. variabile</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B9536A7881DFFCF48EAFF15FA43DD2C" pageId="3" pageNumber="130">
<table id="F92AC407881D003348EAFF15FA74DD23" box="[144,1440,203,2063]" gridcols="4" gridrows="20" pageId="3" pageNumber="130">
<tr id="351A34E5881D003348EAFF15FA74D5F2" box="[144,1440,203,222]" gridrow="0" pageId="3" pageNumber="130">
<th id="76CB5D99881D003348EAFF15FE5FD5F2" box="[144,395,203,222]" gridcol="0" gridrow="0" pageId="3" pageNumber="130">
<emphasis id="B95EEAB5881DFFCF48EAFF15FF3AD5F2" bold="true" box="[144,238,203,222]" pageId="3" pageNumber="130">Character</emphasis>
</th>
<th id="76CB5D99881D003349E0FF15FD36D5F2" box="[410,738,203,222]" gridcol="1" gridrow="0" pageId="3" pageNumber="130">
<taxonomicName id="4C2A4D24881DFFCF49E0FF15FDFBD5F2" authorityName="Swanepoel &amp; E. Tripp" authorityYear="2022" box="[410,559,203,222]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="130" phylum="Tracheophyta" rank="species" species="sesfonteinense">
<emphasis id="B95EEAB5881DFFCF49E0FF15FDFBD5F2" bold="true" box="[410,559,203,222]" italics="true" pageId="3" pageNumber="130">P. sesfonteinense</emphasis>
</taxonomicName>
</th>
<th id="76CB5D99881D00334A89FF15FB93D5F2" box="[755,1095,203,222]" gridcol="2" gridrow="0" pageId="3" pageNumber="130">
<taxonomicName id="4C2A4D24881DFFCF4A89FF15FCB0D5F2" authorityName="Swanepoel" authorityYear="2020" box="[755,868,203,222]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="130" phylum="Tracheophyta" rank="species" species="kaokoense">
<emphasis id="B95EEAB5881DFFCF4A89FF15FCB0D5F2" bold="true" box="[755,868,203,222]" italics="true" pageId="3" pageNumber="130">P. kaokoense</emphasis>
</taxonomicName>
</th>
<th id="76CB5D99881D00334C2DFF15FA74D5F2" box="[1111,1440,203,222]" gridcol="3" gridrow="0" pageId="3" pageNumber="130">
<emphasis id="B95EEAB5881DFFCF4C2DFF15FB6DD5F2" bold="true" box="[1111,1209,203,222]" italics="true" pageId="3" pageNumber="130">P. variabile</emphasis>
</th>
</tr>
<tr id="351A34E5881D003348EAFF2BFA74D494" box="[144,1440,245,440]" gridrow="1" pageId="3" pageNumber="130">
<th id="76CB5D99881D003348EAFF2BFE5FD494" box="[144,395,245,440]" gridcol="0" gridrow="1" pageId="3" pageNumber="130">
<emphasis id="B95EEAB5881DFFCF48EAFE93FEDCD44C" bold="true" box="[144,264,333,352]" pageId="3" pageNumber="130">Indumentum</emphasis>
</th>
<td id="76CB5D99881D003349E0FF2BFD36D494" box="[410,738,245,440]" gridcol="1" gridrow="1" pageId="3" pageNumber="130">Very short glandular trichomes, short to long simple trichomes, often multicellular, also bi-forked, dendritic or long-stalked stellate trichomes, some trichome types absent; dendritic trichomes appearing slender, branches relatively long</td>
<td id="76CB5D99881D00334A89FF2BFB93D494" box="[755,1095,245,440]" gridcol="2" gridrow="1" pageId="3" pageNumber="130">Short-stalked stellate and dendritic trichomes, appearing relatively stout; stalk of stellate trichomes and branches relatively short</td>
<td id="76CB5D99881D00334C2DFF2BFA74D494" box="[1111,1440,245,440]" gridcol="3" gridrow="1" pageId="3" pageNumber="130">Very short appressed trichomes (densely strigose, trichomes pointing backwards), and short glandular trichomes</td>
</tr>
<tr id="351A34E5881D003348EAFE11FA74D774" box="[144,1440,463,600]" gridrow="2" pageId="3" pageNumber="130">
<th id="76CB5D99881D003348EAFE11FE5FD774" box="[144,395,463,600]" gridcol="0" gridrow="2" pageId="3" pageNumber="130">
<emphasis id="B95EEAB5881DFFCF48EAFDD4FE5FD730" bold="true" box="[144,395,521,541]" pageId="3" pageNumber="130">Young stems (indumentum)</emphasis>
</th>
<td id="76CB5D99881D003349E0FE11FD36D774" box="[410,738,463,600]" gridcol="1" gridrow="2" pageId="3" pageNumber="130">Very short glandular trichomes, short to long simple trichomes, often multicellular, also bi-forked, dendritic or long-stalked stellate trichomes; some types lacking</td>
<td id="76CB5D99881D00334A89FE11FB93D774" box="[755,1095,463,600]" gridcol="2" gridrow="2" pageId="3" pageNumber="130">Stellate trichomes, interspersed with a few dendritic ones</td>
<td id="76CB5D99881D00334C2DFE11FA74D774" box="[1111,1440,463,600]" gridcol="3" gridrow="2" pageId="3" pageNumber="130">Densely strigose interspersed with short glandular trichomes</td>
</tr>
<tr id="351A34E5881D003348EAFDB0FA74D790" box="[144,1440,622,700]" gridrow="3" pageId="3" pageNumber="130">
<th id="76CB5D99881D003348EAFDB0FE5FD790" box="[144,395,622,700]" gridcol="0" gridrow="3" pageId="3" pageNumber="130">
<emphasis id="B95EEAB5881DFFCF48EAFDA3FEC7D781" bold="true" box="[144,275,637,685]" pageId="3" pageNumber="130">Leaves (indumentum)</emphasis>
</th>
<td id="76CB5D99881D003349E0FDB0FD36D790" box="[410,738,622,700]" gridcol="1" gridrow="3" pageId="3" pageNumber="130">Dendritic trichomes</td>
<td id="76CB5D99881D00334A89FDB0FB93D790" box="[755,1095,622,700]" gridcol="2" gridrow="3" pageId="3" pageNumber="130">Stalked stellate trichomes, interspersed with dendritic and very short glandular trichomes</td>
<td id="76CB5D99881D00334C2DFDB0FA74D790" box="[1111,1440,622,700]" gridcol="3" gridrow="3" pageId="3" pageNumber="130">Densely strigose trichomes interspersed with short glandular trichomes</td>
</tr>
<tr id="351A34E5881D003348EAFD0DFA74D628" box="[144,1440,723,772]" gridrow="4" pageId="3" pageNumber="130">
<th id="76CB5D99881D003348EAFD0DFE5FD628" box="[144,395,723,772]" gridcol="0" gridrow="4" pageId="3" pageNumber="130">
<emphasis id="B95EEAB5881DFFCF48EAFD3FFEC5D7D8" bold="true" box="[144,273,737,756]" pageId="3" pageNumber="130">Inflorescences</emphasis>
</th>
<td id="76CB5D99881D003349E0FD0DFD36D628" box="[410,738,723,772]" gridcol="1" gridrow="4" pageId="3" pageNumber="130">Axis not becoming spiny with age</td>
<td id="76CB5D99881D00334A89FD0DFB93D628" box="[755,1095,723,772]" gridcol="2" gridrow="4" pageId="3" pageNumber="130">Axis sometimes becoming spiny with age</td>
<td id="76CB5D99881D00334C2DFD0DFA74D628" box="[1111,1440,723,772]" gridcol="3" gridrow="4" pageId="3" pageNumber="130">Axis sometimes becoming spiny with age</td>
</tr>
<tr id="351A34E5881D003348EAFCC4FA74D602" box="[144,1440,794,814]" gridrow="5" pageId="3" pageNumber="130">
<th id="76CB5D99881D003348EAFCC4FE5FD602" box="[144,395,794,814]" gridcol="0" gridrow="5" pageId="3" pageNumber="130">
<emphasis id="B95EEAB5881DFFCF48EAFCC4FEC7D601" bold="true" box="[144,275,794,813]" pageId="3" pageNumber="130">Bracts (shape)</emphasis>
</th>
<td id="76CB5D99881D003349E0FCC4FD36D602" box="[410,738,794,814]" gridcol="1" gridrow="5" pageId="3" pageNumber="130">Oblanceolate</td>
<td id="76CB5D99881D00334A89FCC4FB93D602" box="[755,1095,794,814]" gridcol="2" gridrow="5" pageId="3" pageNumber="130">Linear-lanceolate or linear-oblanceolate</td>
<td id="76CB5D99881D00334C2DFCC4FA74D602" box="[1111,1440,794,814]" gridcol="3" gridrow="5" pageId="3" pageNumber="130">Oblanceolate</td>
</tr>
<tr id="351A34E5881D003348EAFC9AFA74D69C" box="[144,1440,836,944]" gridrow="6" pageId="3" pageNumber="130">
<th id="76CB5D99881D003348EAFC9AFE5FD69C" box="[144,395,836,944]" gridcol="0" gridrow="6" pageId="3" pageNumber="130">
<emphasis id="B95EEAB5881DFFCF48EAFCBCFEB6D6BE" bold="true" box="[144,354,866,914]" pageId="3" pageNumber="130">Bracteoles Shape &amp; reticulation</emphasis>
</th>
<td id="76CB5D99881D003349E0FC9AFD36D69C" box="[410,738,836,944]" gridcol="1" gridrow="6" pageId="3" pageNumber="130">Symmetrically elliptic or narrowly ovate; reticulation weak or not prominent; midrib indistinct, straight; venation indistinct, pale green</td>
<td id="76CB5D99881D00334A89FC9AFB93D69C" box="[755,1095,836,944]" gridcol="2" gridrow="6" pageId="3" pageNumber="130">Asymmetrically elliptic-oblong; reticulation prominent; midrib distinct, curved; venation distinct, dark green to maroon</td>
<td id="76CB5D99881D00334C2DFC9AFA74D69C" box="[1111,1440,836,944]" gridcol="3" gridrow="6" pageId="3" pageNumber="130">Asymmetrically elliptic-oblong; reticulation weak or not prominent; midrib distinct, prominent or slightly so</td>
</tr>
<tr id="351A34E5881D003348EAFC18FA74D6DB" box="[144,1440,966,1015]" gridrow="7" pageId="3" pageNumber="130" rowspan-0="1" rowspan-2="1" rowspan-3="1">
<td id="76CB5D99881D003349E0FC18FD36D6DB" box="[410,738,966,1015]" gridcol="1" gridrow="7" pageId="3" pageNumber="130">Glandular trichomes of various lengths and few to scattered long</td>
</tr>
<tr id="351A34E5881D003348EAFBDFFA74D1A5" box="[144,1440,1025,1161]" gridrow="8" pageId="3" pageNumber="130">
<th id="76CB5D99881D003348EAFBDFFE5FD1A5" box="[144,395,1025,1161]" gridcol="0" gridrow="8" pageId="3" pageNumber="130">
<emphasis id="B95EEAB5881DFFCF48DFFBCEFF25D16C" bold="true" pageId="3" pageNumber="130">Indumentum (abaxially)</emphasis>
</th>
<td id="76CB5D99881D003349E0FBDFFD36D1A5" box="[410,738,1025,1161]" gridcol="1" gridrow="8" pageId="3" pageNumber="130">multicellular glandular trichomes, sometimes also with very long multicellular sparsely branched eglandular simple or dendritic trichomes</td>
<td id="76CB5D99881D00334A89FBDFFB93D1A5" box="[755,1095,1025,1161]" gridcol="2" gridrow="8" pageId="3" pageNumber="130">Stalked stellate, dendritic and very short glandular trichomes</td>
<td id="76CB5D99881D00334C2DFBDFFA74D1A5" box="[1111,1440,1025,1161]" gridcol="3" gridrow="8" pageId="3" pageNumber="130">Densely strigose interspersed with short glandular trichomes</td>
</tr>
<tr id="351A34E5881D003348EAFB7EFA74D19F" box="[144,1440,1184,1203]" gridrow="9" pageId="3" pageNumber="130">
<th id="76CB5D99881D003348EAFB7EFA74D19F" box="[144,1440,1184,1203]" colspan="4" colspanRight="3" gridcol="0" gridrow="9" pageId="3" pageNumber="130">
<emphasis id="B95EEAB5881DFFCF48EAFB7EFF00D19F" bold="true" box="[144,212,1184,1203]" pageId="3" pageNumber="130">Corolla</emphasis>
</th>
</tr>
<tr id="351A34E5881D003348EAFB14FA74D1D6" box="[144,1440,1226,1274]" gridrow="10" pageId="3" pageNumber="130">
<th id="76CB5D99881D003348EAFB14FE5FD1D6" box="[144,395,1226,1274]" gridcol="0" gridrow="10" pageId="3" pageNumber="130">
<emphasis id="B95EEAB5881DFFCF48DFFB14FEDED1D6" bold="true" pageId="3" pageNumber="130">Indumentum (expanded part, outside)</emphasis>
</th>
<td id="76CB5D99881D003349E0FB14FD36D1D6" box="[410,738,1226,1274]" gridcol="1" gridrow="10" pageId="3" pageNumber="130">Glabrous</td>
<td id="76CB5D99881D00334A89FB14FB93D1D6" box="[755,1095,1226,1274]" gridcol="2" gridrow="10" pageId="3" pageNumber="130">Densely strigose</td>
<td id="76CB5D99881D00334C2DFB14FA74D1D6" box="[1111,1440,1226,1274]" gridcol="3" gridrow="10" pageId="3" pageNumber="130">Strigose</td>
</tr>
<tr id="351A34E5881D003348EAFACFFA74D06E" box="[144,1440,1297,1346]" gridrow="11" pageId="3" pageNumber="130">
<th id="76CB5D99881D003348EAFACFFE5FD06E" box="[144,395,1297,1346]" gridcol="0" gridrow="11" pageId="3" pageNumber="130">
<emphasis id="B95EEAB5881DFFCF48EAFAFEFF29D01F" bold="true" box="[144,253,1312,1331]" pageId="3" pageNumber="130">Upper lobes</emphasis>
</th>
<td id="76CB5D99881D003349E0FACFFD36D06E" box="[410,738,1297,1346]" gridcol="1" gridrow="11" pageId="3" pageNumber="130">Connate for 20–30% the length; obovate</td>
<td id="76CB5D99881D00334A89FACFFB93D06E" box="[755,1095,1297,1346]" gridcol="2" gridrow="11" pageId="3" pageNumber="130">Connate for 50% the length; oblong</td>
<td id="76CB5D99881D00334C2DFACFFA74D06E" box="[1111,1440,1297,1346]" gridcol="3" gridrow="11" pageId="3" pageNumber="130">Connate for 33–50% the length; narrowly ovate or oblong-elliptic</td>
</tr>
<tr id="351A34E5881D003348EAFA87FA74D0A5" box="[144,1440,1369,1417]" gridrow="12" pageId="3" pageNumber="130">
<th id="76CB5D99881D003348EAFA87FE5FD0A5" box="[144,395,1369,1417]" gridcol="0" gridrow="12" pageId="3" pageNumber="130">
<emphasis id="B95EEAB5881DFFCF48EAFA87FED7D0A5" bold="true" box="[144,259,1369,1417]" pageId="3" pageNumber="130">Lobes (orientation)</emphasis>
</th>
<td id="76CB5D99881D003349E0FA87FD36D0A5" box="[410,738,1369,1417]" gridcol="1" gridrow="12" pageId="3" pageNumber="130">Patent</td>
<td id="76CB5D99881D00334A89FA87FB93D0A5" box="[755,1095,1369,1417]" gridcol="2" gridrow="12" pageId="3" pageNumber="130">Lateral and upper lobes patent, front lobe ca. at 45 degrees</td>
<td id="76CB5D99881D00334C2DFA87FA74D0A5" box="[1111,1440,1369,1417]" gridcol="3" gridrow="12" pageId="3" pageNumber="130">Upper lobes in line with tube to sub-patent, other lobes patent</td>
</tr>
<tr id="351A34E5881D003348EAFA7EFA74D09F" box="[144,1440,1440,1459]" gridrow="13" pageId="3" pageNumber="130">
<th id="76CB5D99881D003348EAFA7EFE5FD09F" box="[144,395,1440,1459]" gridcol="0" gridrow="13" pageId="3" pageNumber="130">
<emphasis id="B95EEAB5881DFFCF48EAFA7EFE9BD09F" bold="true" box="[144,335,1440,1459]" pageId="3" pageNumber="130">Lobes (indumentum)</emphasis>
</th>
<td id="76CB5D99881D003349E0FA7EFD36D09F" box="[410,738,1440,1459]" gridcol="1" gridrow="13" pageId="3" pageNumber="130">Patent trichomes on all lobes</td>
<td id="76CB5D99881D00334A89FA7EFB93D09F" box="[755,1095,1440,1459]" gridcol="2" gridrow="13" pageId="3" pageNumber="130">Patent trichomes only on front lobe</td>
<td id="76CB5D99881D00334C2DFA7EFA74D09F" box="[1111,1440,1440,1459]" gridcol="3" gridrow="13" pageId="3" pageNumber="130">Patent trichomes on all lobes</td>
</tr>
<tr id="351A34E5881D003348EAFA14FA74D3E4" box="[144,1440,1482,1736]" gridrow="14" pageId="3" pageNumber="130">
<th id="76CB5D99881D003348EAFA14FE5FD3E4" box="[144,395,1482,1736]" gridcol="0" gridrow="14" pageId="3" pageNumber="130">
<emphasis id="B95EEAB5881DFFCF48EAF9E1FF04D37E" bold="true" box="[144,208,1599,1618]" pageId="3" pageNumber="130">Colour</emphasis>
</th>
<td id="76CB5D99881D003349E0FA14FD36D3E4" box="[410,738,1482,1736]" gridcol="1" gridrow="14" pageId="3" pageNumber="130">Lobes white, pink, magenta, apricot, yellow, or cream, maroon dotted proximally; throat and mouth maroon, terminating in two conspicuous narrow triangular markings on proximal part of lobes; two yellow nectar guides on front lobe narrowly triangular, relatively long, widely separate</td>
<td id="76CB5D99881D00334A89FA14FB93D3E4" box="[755,1095,1482,1736]" gridcol="2" gridrow="14" pageId="3" pageNumber="130">Lobes, throat, and mouth uniformly maroon; two yellow nectar guides on front lobe oblong or oblong-triangular, relatively long, connate towards base</td>
<td id="76CB5D99881D00334C2DFA14FA74D3E4" box="[1111,1440,1482,1736]" gridcol="3" gridrow="14" pageId="3" pageNumber="130">Front corolla lobe differently coloured or shaded than upper and sometimes also lateral lobes: white, pink, or crimson; upper and lateral lobes: brown, dirty orange, pink, or crimson; throat and mouth maroon; two yellow nectar guides on front lobe narrowly triangular, relatively short, separate</td>
</tr>
<tr id="351A34E5881D003348EAF900FA74D3DE" box="[144,1440,1758,1778]" gridrow="15" pageId="3" pageNumber="130">
<th id="76CB5D99881D003348EAF900FE5FD3DE" box="[144,395,1758,1778]" gridcol="0" gridrow="15" pageId="3" pageNumber="130">
<emphasis id="B95EEAB5881DFFCF48EAF900FEC5D3DD" bold="true" box="[144,273,1758,1777]" pageId="3" pageNumber="130">Stigma (lobes)</emphasis>
</th>
<td id="76CB5D99881D003349E0F900FD36D3DE" box="[410,738,1758,1778]" gridcol="1" gridrow="15" pageId="3" pageNumber="130">Sub-equal</td>
<td id="76CB5D99881D00334A89F900FB93D3DE" box="[755,1095,1758,1778]" gridcol="2" gridrow="15" pageId="3" pageNumber="130">Unequal</td>
<td id="76CB5D99881D00334C2DF900FA74D3DE" box="[1111,1440,1758,1778]" gridcol="3" gridrow="15" pageId="3" pageNumber="130">Sub-equal</td>
</tr>
<tr id="351A34E5881D003348EAF8D7FA74D215" box="[144,1440,1801,1849]" gridrow="16" pageId="3" pageNumber="130">
<th id="76CB5D99881D003348EAF8D7FE5FD215" box="[144,395,1801,1849]" gridcol="0" gridrow="16" pageId="3" pageNumber="130">
<emphasis id="B95EEAB5881DFFCF48EAF8C9FF0DD206" bold="true" box="[144,217,1815,1834]" pageId="3" pageNumber="130">Capsule</emphasis>
</th>
<td id="76CB5D99881D003349E0F8D7FD36D215" box="[410,738,1801,1849]" gridcol="1" gridrow="16" pageId="3" pageNumber="130">Flattened ellipsoid or ovoid, rugose or smooth</td>
<td id="76CB5D99881D00334A89F8D7FB93D215" box="[755,1095,1801,1849]" gridcol="2" gridrow="16" pageId="3" pageNumber="130">Flattened ovoid, smooth</td>
<td id="76CB5D99881D00334C2DF8D7FA74D215" box="[1111,1440,1801,1849]" gridcol="3" gridrow="16" pageId="3" pageNumber="130">Flattened ovoid, smooth</td>
</tr>
<tr id="351A34E5881D003348EAF88EFA74D24F" box="[144,1440,1872,1891]" gridrow="17" pageId="3" pageNumber="130">
<th id="76CB5D99881D003348EAF88EFE5FD24F" box="[144,395,1872,1891]" gridcol="0" gridrow="17" pageId="3" pageNumber="130">
<emphasis id="B95EEAB5881DFFCF48EAF88EFF17D24F" bold="true" box="[144,195,1872,1891]" pageId="3" pageNumber="130">Seeds</emphasis>
</th>
<td id="76CB5D99881D003349E0F88EFD36D24F" box="[410,738,1872,1891]" gridcol="1" gridrow="17" pageId="3" pageNumber="130">Cordate</td>
<td id="76CB5D99881D00334A89F88EFB93D24F" box="[755,1095,1872,1891]" gridcol="2" gridrow="17" pageId="3" pageNumber="130">Discoid or ovate</td>
<td id="76CB5D99881D00334C2DF88EFA74D24F" box="[1111,1440,1872,1891]" gridcol="3" gridrow="17" pageId="3" pageNumber="130">Cordate</td>
</tr>
<tr id="351A34E5881D003348EAF8A4FA74D2A1" box="[144,1440,1914,1933]" gridrow="18" pageId="3" pageNumber="130">
<th id="76CB5D99881D003348EAF8A4FE5FD2A1" box="[144,395,1914,1933]" gridcol="0" gridrow="18" pageId="3" pageNumber="130">
<emphasis id="B95EEAB5881DFFCF48EAF8A4FF3FD2A1" bold="true" box="[144,235,1914,1933]" pageId="3" pageNumber="130">Cystoliths</emphasis>
</th>
<td id="76CB5D99881D003349E0F8A4FD36D2A1" box="[410,738,1914,1933]" gridcol="1" gridrow="18" pageId="3" pageNumber="130">Linear or appearing circular</td>
<td id="76CB5D99881D00334A89F8A4FB93D2A1" box="[755,1095,1914,1933]" gridcol="2" gridrow="18" pageId="3" pageNumber="130">Linear</td>
<td id="76CB5D99881D00334C2DF8A4FA74D2A1" box="[1111,1440,1914,1933]" gridcol="3" gridrow="18" pageId="3" pageNumber="130">Linear or appearing circular</td>
</tr>
<tr id="351A34E5881D003348EAF87AFA74DD23" box="[144,1440,1956,2063]" gridrow="19" pageId="3" pageNumber="130">
<th id="76CB5D99881D003348EAF87AFE5FDD23" box="[144,395,1956,2063]" gridcol="0" gridrow="19" pageId="3" pageNumber="130">
<emphasis id="B95EEAB5881DFFCF48EAF80EFF2BD2CF" bold="true" box="[144,255,2000,2019]" pageId="3" pageNumber="130">Distribution</emphasis>
</th>
<td id="76CB5D99881D003349E0F87AFD36DD23" box="[410,738,1956,2063]" gridcol="1" gridrow="19" pageId="3" pageNumber="130">Sesfontein and the surrounding mountainous area up to Tomakas in the west and Warmquelle in the east</td>
<td id="76CB5D99881D00334A89F87AFB93DD23" box="[755,1095,1956,2063]" gridcol="2" gridrow="19" pageId="3" pageNumber="130">Hartmann Mountains north of Orupembe to 20 km south of the Kunene River and at Omungwindi ca. 40 km east of the Hartmann Mountains</td>
<td id="76CB5D99881D00334C2DF87AFA74DD23" box="[1111,1440,1956,2063]" gridcol="3" gridrow="19" pageId="3" pageNumber="130">From the Bergsig area southwards to west central Namibia and in northern Namibia near Otavi and Grootfontein</td>
</tr>
</table>
</paragraph>
<caption id="DF55662F881AFFC848F2F8A5FBE6D298" ID-DOI="http://doi.org/10.5281/zenodo.6622696" ID-Zenodo-Dep="6622696" httpUri="https://zenodo.org/record/6622696/files/figure.png" pageId="4" pageNumber="131" startId="4.[136,229,1915,1937]" targetBox="[151,1435,187,1886]" targetPageId="4">
<paragraph id="8B9536A7881AFFC848F2F8A5FBE6D298" blockId="4.[136,1451,1915,1973]" pageId="4" pageNumber="131">
<emphasis id="B95EEAB5881AFFC848F2F8A5FF2BD2BC" bold="true" box="[136,255,1915,1937]" pageId="4" pageNumber="131">FIGURE 1.</emphasis>
<taxonomicName id="4C2A4D24881AFFC8497CF8A5FDD6D2BC" authorityName="Swanepoel &amp; E. Tripp" authorityYear="2022" box="[262,514,1915,1936]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="131" phylum="Tracheophyta" rank="species" species="sesfonteinense">
<emphasis id="B95EEAB5881AFFC8497CF8A5FDD6D2BC" box="[262,514,1915,1936]" italics="true" pageId="4" pageNumber="131">Petalidium sesfonteinense</emphasis>
</taxonomicName>
.
<emphasis id="B95EEAB5881AFFC84A75F8A5FDF2D2BC" bold="true" box="[527,550,1915,1936]" pageId="4" pageNumber="131">A.</emphasis>
Habit. Plant about 40 cm tall.
<emphasis id="B95EEAB5881AFFC84B2FF8A5FCBFD2BC" bold="true" box="[853,875,1915,1936]" pageId="4" pageNumber="131">B.</emphasis>
Flower, shoots, and leaves; greyish leaves are still densely covered in trichomes; green leaves almost glabrous. Photographs: W. Swanepoel.
<emphasis id="B95EEAB5881AFFC84BE0F841FBF8D298" box="[922,1068,1951,1972]" italics="true" pageId="4" pageNumber="131">Swanepoel 568</emphasis>
.
</paragraph>
</caption>
<paragraph id="8B9536A7881AFFC948C7F83DFCBFD440" blockId="4.[136,1452,2019,2082]" lastBlockId="5.[136,1452,158,365]" lastPageId="5" lastPageNumber="132" pageId="4" pageNumber="131">
<bibRefCitation id="EFBB4B56881AFFC848C7F83DFE5CD2D1" author="Tripp, E. A. &amp; Tsai, Y. E. &amp; Zhuang, Y. &amp; Dexter, K. G." box="[189,392,2019,2046]" pageId="4" pageNumber="131" pagination="1 - 17" refId="ref5642" refString="Tripp, E. A., Tsai, Y. E., Zhuang, Y. &amp; Dexter, K. G. (2017) RADseq dataset with 90 % missing data fully resolves recent radiation of Petalidium (Acanthaceae) in the ultra-arid deserts of Namibia. Ecology and Evolution 7: 1 - 17. https: // doi. org / 10.1002 / ece 3.3274" type="journal article" year="2017">
Tripp
<emphasis id="B95EEAB5881AFFC8497BF83AFEE5D2D1" box="[257,305,2019,2045]" italics="true" pageId="4" pageNumber="131">et al</emphasis>
. (2017)
</bibRefCitation>
conducted a phylogenomic study of
<taxonomicName id="4C2A4D24881AFFC84B5AF83DFC4FD2D1" authorityName="Nees von Esenbeck" authorityYear="1832" box="[800,923,2019,2045]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="131" phylum="Tracheophyta" rank="genus">
<emphasis id="B95EEAB5881AFFC84B5AF83DFC4FD2D1" box="[800,923,2019,2045]" italics="true" pageId="4" pageNumber="131">Petalidium</emphasis>
</taxonomicName>
, sampling several species including
<taxonomicName id="4C2A4D24881AFFC84D4AF83DFEFEDD0D" authorityName="Swanepoel &amp; E. Tripp" authorityYear="2022" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="131" phylum="Tracheophyta" rank="species" species="sesfonteinense">
<emphasis id="B95EEAB5881AFFC84D4AF83DFEFEDD0D" italics="true" pageId="4" pageNumber="131">Petalidium sesfonteinense</emphasis>
</taxonomicName>
(therein labelled as “
<taxonomicName id="4C2A4D24881AFFC84A62F7D9FD2ADD0D" box="[536,766,2055,2081]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="131" phylum="Tracheophyta" rank="species" species="variabile">
<emphasis id="B95EEAB5881AFFC84A62F7D9FD2ADD0D" box="[536,766,2055,2081]" italics="true" pageId="4" pageNumber="131">Petalidium variabile</emphasis>
</taxonomicName>
16-20”), along with
<taxonomicName id="4C2A4D24881AFFC84B9DF7D6FB60DD0D" authorityName="Meyer" authorityYear="1961" box="[999,1204,2056,2081]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="131" phylum="Tracheophyta" rank="species" species="rossmannianum">
<emphasis id="B95EEAB5881AFFC84B9DF7D6FB60DD0D" box="[999,1204,2056,2081]" italics="true" pageId="4" pageNumber="131">P. rossmannianum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A4D24881AFFC84CBBF7D6FA8EDD0D" authorityName="Moore" authorityYear="1880" box="[1217,1370,2055,2081]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="131" phylum="Tracheophyta" rank="species" species="welwitschii">
<emphasis id="B95EEAB5881AFFC84CBBF7D6FA8EDD0D" box="[1217,1370,2055,2081]" italics="true" pageId="4" pageNumber="131">P. welwitschii</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C2A4D24881AFFC94DEDF7D6FF29D594" authorityName="Swanepoel" authorityYear="2020" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" lastPageId="5" lastPageNumber="132" order="Lamiales" pageId="4" pageNumber="131" phylum="Tracheophyta" rank="species" species="kaokoense">
<emphasis id="B95EEAB5881AFFC94DEDF7D6FF29D594" italics="true" lastPageId="5" lastPageNumber="132" pageId="4" pageNumber="131">P. kaokoense</emphasis>
</taxonomicName>
(therein labelled with an arrow as “
<taxonomicName id="4C2A4D24881BFFC94AE2FF40FC85D594" box="[664,849,158,185]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="132" phylum="Tracheophyta" rank="species" species="undefined-8">
<emphasis id="B95EEAB5881BFFC94AE2FF40FCC7D594" box="[664,787,158,184]" italics="true" pageId="5" pageNumber="132">Petalidium</emphasis>
sp. 8
</taxonomicName>
[
<taxonomicName id="4C2A4D24881BFFC94B1BFF40FC39D594" box="[865,1005,158,184]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" isUncertain="true" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="132" phylum="Tracheophyta" rank="species" species="variabile">
cf.
<emphasis id="B95EEAB5881BFFC94BF2FF40FC39D594" box="[904,1005,158,184]" italics="true" pageId="5" pageNumber="132">variabile</emphasis>
</taxonomicName>
]”). These four species together formed a strongly supported clade (Clades 7, 8, &amp; 9
<bibRefCitation id="EFBB4B56881BFFC94AF8FF1CFC95D5F0" author="Tripp, E. A. &amp; Tsai, Y. E. &amp; Zhuang, Y. &amp; Dexter, K. G." box="[642,833,194,221]" pageId="5" pageNumber="132" pagination="1 - 17" refId="ref5642" refString="Tripp, E. A., Tsai, Y. E., Zhuang, Y. &amp; Dexter, K. G. (2017) RADseq dataset with 90 % missing data fully resolves recent radiation of Petalidium (Acanthaceae) in the ultra-arid deserts of Namibia. Ecology and Evolution 7: 1 - 17. https: // doi. org / 10.1002 / ece 3.3274" type="journal article" year="2017">
Tripp
<emphasis id="B95EEAB5881BFFC94ABDFF1DFD2CD5F0" box="[711,760,194,220]" italics="true" pageId="5" pageNumber="132">et al</emphasis>
. 2017
</bibRefCitation>
) that is sister to Clade 10, which contained
<taxonomicName id="4C2A4D24881BFFC94D4AFF1CFDE8D42D" authority="Meuse ex Meyer (1961: 66)" authorityName="Meyer" authorityPageNumber="66" authorityYear="1961" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="132" phylum="Tracheophyta" rank="species" species="crispum">
<emphasis id="B95EEAB5881BFFC94D4AFF1CFF36D42C" italics="true" pageId="5" pageNumber="132">Petalidium crispum</emphasis>
Meuse ex
<bibRefCitation id="EFBB4B56881BFFC94910FF39FDE8D42D" author="Meyer, P. G." box="[362,572,230,257]" pageId="5" pageNumber="132" pagination="59 - 72" refId="ref5145" refString="Meyer, P. G. (1961) Beitrage zur kenntnis der Acanthaceen Sudwestafrikas (III). Mitteilungen der Botanischen Staatssammlung Munchen 4: 59 - 72. [https: // www. biodiversitylibrary. org / item / 52384 # page / 747 / mode / 1 up]" type="journal article" year="1961">Meyer (1961: 66)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2A4D24881BFFC94A36FF39FD30D42C" authorityName="Moore" authorityYear="1880" box="[588,740,231,256]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="132" phylum="Tracheophyta" rank="species" species="coccineum">
<emphasis id="B95EEAB5881BFFC94A36FF39FD30D42C" box="[588,740,231,256]" italics="true" pageId="5" pageNumber="132">P. coccineum</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C2A4D24881BFFC94B51FF39FB3ED42D" authority="Obermeijer (1936: 161)" authorityName="Obermeijer" authorityPageNumber="161" authorityYear="1936" box="[811,1258,230,257]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="132" phylum="Tracheophyta" rank="species" species="bracteatum">
<emphasis id="B95EEAB5881BFFC94B51FF39FC1FD42C" box="[811,971,230,256]" italics="true" pageId="5" pageNumber="132">P. bracteatum</emphasis>
<bibRefCitation id="EFBB4B56881BFFC94BACFF38FB3ED42D" author="Obermeijer, A. A." box="[982,1258,230,257]" pageId="5" pageNumber="132" pagination="151 - 162" refId="ref5433" refString="Obermeijer, A. A. (1936) The South African species of Petalidium. Annals of the Transvaal Museum 18: 151 - 162." type="journal article" year="1936">Obermeijer (1936: 161)</bibRefCitation>
</taxonomicName>
. More sampling combined with next-generation sequencing is needed to further elucidate phylogenetic relatedness of this new species to other species of
<taxonomicName id="4C2A4D24881BFFC9492CFEF0FE05D464" authorityName="Nees von Esenbeck" authorityYear="1832" box="[342,465,302,328]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="132" phylum="Tracheophyta" rank="genus">
<emphasis id="B95EEAB5881BFFC9492CFEF0FE05D464" box="[342,465,302,328]" italics="true" pageId="5" pageNumber="132">Petalidium</emphasis>
</taxonomicName>
, including the widespread
<taxonomicName id="4C2A4D24881BFFC94A80FEF1FCADD464" box="[762,889,302,328]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="132" phylum="Tracheophyta" rank="species" species="variabile">
<emphasis id="B95EEAB5881BFFC94A80FEF1FCADD464" box="[762,889,302,328]" italics="true" pageId="5" pageNumber="132">P. variabile</emphasis>
</taxonomicName>
(Manzitto-Tripp
<emphasis id="B95EEAB5881BFFC94C40FEF1FBBED464" box="[1082,1130,302,328]" italics="true" pageId="5" pageNumber="132">et al</emphasis>
. in prep.). In turn, this would help to clarify geographical limits of these closely related species.
</paragraph>
<caption id="DF55662F881BFFC948F2F875FD7DDD25" ID-DOI="http://doi.org/10.5281/zenodo.6622698" ID-Zenodo-Dep="6622698" httpUri="https://zenodo.org/record/6622698/files/figure.png" pageId="5" pageNumber="132" startId="5.[136,229,1963,1985]" targetBox="[151,1435,389,1938]" targetPageId="5">
<paragraph id="8B9536A7881BFFC948F2F875FD7DDD25" blockId="5.[136,1452,1963,2057]" pageId="5" pageNumber="132">
<emphasis id="B95EEAB5881BFFC948F2F875FF29D2EC" bold="true" box="[136,253,1963,1985]" pageId="5" pageNumber="132">FIGURE 2.</emphasis>
<taxonomicName id="4C2A4D24881BFFC94979F875FE29D2EC" authorityName="Swanepoel &amp; E. Tripp" authorityYear="2022" box="[259,509,1963,1984]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="132" phylum="Tracheophyta" rank="species" species="sesfonteinense">
<emphasis id="B95EEAB5881BFFC94979F875FE29D2EC" box="[259,509,1963,1984]" italics="true" pageId="5" pageNumber="132">Petalidium sesfonteinense</emphasis>
</taxonomicName>
. Morphology of flowers.
<emphasis id="B95EEAB5881BFFC94A8EF875FCFCD2EC" bold="true" box="[756,808,1963,1984]" pageId="5" pageNumber="132">A–D.</emphasis>
Flowers in front view showing variation in corolla colour and lobe margins, with two yellow nectar guides.
<emphasis id="B95EEAB5881BFFC94A70F811FDF5D2C8" bold="true" box="[522,545,1999,2020]" pageId="5" pageNumber="132">E.</emphasis>
Corolla in lateral view.
<emphasis id="B95EEAB5881BFFC94B7FF811FCC3D2C8" bold="true" box="[773,791,1999,2020]" pageId="5" pageNumber="132">F.</emphasis>
Corolla in dorsal view.
<emphasis id="B95EEAB5881BFFC94B81F811FBC0D2C9" bold="true" box="[1019,1044,1999,2021]" pageId="5" pageNumber="132">G.</emphasis>
Corolla in ventral view. Scale bar = 5 mm. Photographs: W. Swanepoel.
<emphasis id="B95EEAB5881BFFC949D8F82DFDE0DD24" box="[418,564,2035,2056]" italics="true" pageId="5" pageNumber="132">Swanepoel 568</emphasis>
(E, F &amp; G).
</paragraph>
</caption>
<caption id="DF55662F8818FFCA48F2FA58FBD9D0B0" ID-DOI="http://doi.org/10.5281/zenodo.6622700" ID-Zenodo-Dep="6622700" box="[136,1037,1414,1436]" httpUri="https://zenodo.org/record/6622700/files/figure.png" pageId="6" pageNumber="133" startId="6.[136,229,1414,1436]" targetBox="[183,1404,190,1390]" targetPageId="6">
<paragraph id="8B9536A78818FFCA48F2FA58FBD9D0B0" blockId="6.[136,1037,1414,1436]" box="[136,1037,1414,1436]" pageId="6" pageNumber="133">
<emphasis id="B95EEAB58818FFCA48F2FA58FF2AD0B7" bold="true" box="[136,254,1414,1436]" pageId="6" pageNumber="133">FIGURE 3.</emphasis>
Known distribution of
<taxonomicName id="4C2A4D248818FFCA49A5FA58FD0DD0B7" authorityName="Swanepoel &amp; E. Tripp" authorityYear="2022" box="[479,729,1414,1435]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="133" phylum="Tracheophyta" rank="species" species="sesfonteinense">
<emphasis id="B95EEAB58818FFCA49A5FA58FD0DD0B7" box="[479,729,1414,1435]" italics="true" pageId="6" pageNumber="133">Petalidium sesfonteinense</emphasis>
</taxonomicName>
based on herbarium specimens.
</paragraph>
</caption>
<caption id="DF55662F8818FFCA48F2F868FA96DD38" ID-DOI="http://doi.org/10.5281/zenodo.6622702" ID-Zenodo-Dep="6622702" httpUri="https://zenodo.org/record/6622702/files/figure.png" pageId="6" pageNumber="133" startId="6.[136,229,1974,1996]" targetBox="[183,1403,1462,1950]" targetPageId="6">
<paragraph id="8B9536A78818FFCA48F2F868FA96DD38" blockId="6.[136,1452,1974,2068]" pageId="6" pageNumber="133">
<emphasis id="B95EEAB58818FFCA48F2F868FF2FD2E7" bold="true" box="[136,251,1974,1996]" pageId="6" pageNumber="133">FIGURE 4.</emphasis>
Flowers of two species related to
<taxonomicName id="4C2A4D248818FFCA4A48F868FCFED2E7" authorityName="Swanepoel &amp; E. Tripp" authorityYear="2022" box="[562,810,1974,1995]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="133" phylum="Tracheophyta" rank="species" species="sesfonteinense">
<emphasis id="B95EEAB58818FFCA4A48F868FCFED2E7" box="[562,810,1974,1995]" italics="true" pageId="6" pageNumber="133">Petalidium sesfonteinense</emphasis>
</taxonomicName>
, for purposes of comparison.
<emphasis id="B95EEAB58818FFCA4C46F868FB80D2E7" bold="true" box="[1084,1108,1974,1995]" pageId="6" pageNumber="133">A.</emphasis>
<emphasis id="B95EEAB58818FFCA4C2DF869FB03D2E7" box="[1111,1239,1974,1996]" italics="true" pageId="6" pageNumber="133">
<taxonomicName id="4C2A4D248818FFCA4C2DF869FB07D2E7" authorityName="Swanepoel" authorityYear="2020" box="[1111,1235,1974,1996]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="133" phylum="Tracheophyta" rank="species" species="kaokoense">P.kaokoense</taxonomicName>
.
</emphasis>
Note the one-coloured lobes of similar shade with relatively long yellow nectar guides (Hartmann Mountains, Kaokoveld).
<emphasis id="B95EEAB58818FFCA4C3CF804FB89D2C3" bold="true" box="[1094,1117,2010,2031]" pageId="6" pageNumber="133">B.</emphasis>
<taxonomicName id="4C2A4D248818FFCA4C18F805FB04D2C3" box="[1122,1232,2010,2032]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="133" phylum="Tracheophyta" rank="species" species="variabile">
<emphasis id="B95EEAB58818FFCA4C18F805FB04D2C3" box="[1122,1232,2010,2032]" italics="true" pageId="6" pageNumber="133">P. variabile</emphasis>
</taxonomicName>
. Note the difference in shade between lobes and the relatively short yellow nectar guides (Okahandja, central Namibia). Photographs: W. Swanepoel.
</paragraph>
</caption>
<paragraph id="8B9536A78819FFCB48C7FF40FE74D7F8" blockId="7.[136,1452,158,725]" pageId="7" pageNumber="134">
<emphasis id="B95EEAB58819FFCB48C7FF40FD21D594" bold="true" box="[189,757,158,184]" pageId="7" pageNumber="134">
Additional specimens examined (
<typeStatus id="549188058819FFCB4A35FF40FD1FD594" box="[591,715,158,184]" pageId="7" pageNumber="134" type="paratype">paratypes</typeStatus>
):—
</emphasis>
<collectingCountry id="F33D76378819FFCB4A8FFF40FCA7D594" box="[757,883,158,184]" name="Namibia" pageId="7" pageNumber="134">NAMIBIA</collectingCountry>
,
<collectingRegion id="49EEF8458819FFCB4BFBFF41FBEDD594" box="[897,1081,158,185]" country="Namibia" name="Kunene" pageId="7" pageNumber="134">Kunene Region</collectingRegion>
:—1813: North of Sesfontein, (– CD),
<date id="FF9410678819FFCB48B3FF1CFEA9D5F0" box="[201,381,194,220]" pageId="7" pageNumber="134" value="2004-03-31">31 March 2004</date>
,
<emphasis id="B95EEAB58819FFCB49F1FF1CFDC9D5F0" box="[395,541,194,220]" italics="true" pageId="7" pageNumber="134">Burke 04080</emphasis>
(WIND!);
<quantity id="4CD29B428819FFCB4ADBFF1CFD3FD5F0" box="[673,747,194,220]" metricMagnitude="4" metricUnit="m" metricValue="4.2" pageId="7" pageNumber="134" unit="km" value="42.0">42 km</quantity>
before Sesfontein from Puros, (–CD),
<quantity id="4CD29B428819FFCB4CD5FF1CFB2FD5F1" box="[1199,1275,194,221]" metricMagnitude="2" metricUnit="m" metricValue="4.46" pageId="7" pageNumber="134" unit="m" value="446.0">446 m</quantity>
,
<date id="FF9410678819FFCB4D73FF1CFA73D5F0" box="[1289,1447,194,221]" pageId="7" pageNumber="134" value="2014-05-18">18 May 2014</date>
,
<emphasis id="B95EEAB58819FFCB48F2FF38FE85D42C" box="[136,337,230,256]" italics="true" pageId="7" pageNumber="134">Klaassen EK2548</emphasis>
(WIND!); Hills
<quantity id="4CD29B428819FFCB4A72FF38FD94D42C" box="[520,576,230,256]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="134" unit="km" value="2.0">2 km</quantity>
north of Okarondokavi, (–CD),
<quantity id="4CD29B428819FFCB4BDEFF38FC38D42D" box="[932,1004,230,257]" metricMagnitude="2" metricUnit="m" metricValue="7.42" pageId="7" pageNumber="134" unit="m" value="742.0">742 m</quantity>
,
<date id="FF9410678819FFCB4B8CFF38FB58D42C" box="[1014,1164,230,257]" pageId="7" pageNumber="134" value="2022-05-14">14 May 2022</date>
,
<emphasis id="B95EEAB58819FFCB4CECFF38FAEBD42C" box="[1174,1343,230,256]" italics="true" pageId="7" pageNumber="134">Swanepoel 579</emphasis>
(WIND); Hills
<quantity id="4CD29B428819FFCB48BFFED4FEC7D408" box="[197,275,266,293]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="134" unit="km" value="2.5">2.5 km</quantity>
north of Okarondokavi, (–CD),
<quantity id="4CD29B428819FFCB4A04FED4FD13D409" box="[638,711,266,293]" metricMagnitude="2" metricUnit="m" metricValue="7.6" pageId="7" pageNumber="134" unit="m" value="760.0">760 m</quantity>
,
<date id="FF9410678819FFCB4AA8FED4FCBED408" box="[722,874,266,293]" pageId="7" pageNumber="134" value="2022-05-14">14 May 2022</date>
,
<emphasis id="B95EEAB58819FFCB4B0FFED4FBCBD408" box="[885,1055,266,292]" italics="true" pageId="7" pageNumber="134">Swanepoel 580</emphasis>
(WIND);
<quantity id="4CD29B428819FFCB4CEEFED4FB19D408" box="[1172,1229,266,292]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="134" unit="km" value="6.0">6 km</quantity>
south of Okovikuta on road to Ganamub, (–CD),
<quantity id="4CD29B428819FFCB49ABFEF0FDCED465" box="[465,538,302,329]" metricMagnitude="2" metricUnit="m" metricValue="6.94" pageId="7" pageNumber="134" unit="m" value="694.0">694 m</quantity>
,
<date id="FF9410678819FFCB4A5CFEF0FD6AD464" box="[550,702,302,329]" pageId="7" pageNumber="134" value="2022-05-14">14 May 2022</date>
,
<emphasis id="B95EEAB58819FFCB4AB3FEF0FCA7D464" box="[713,883,302,328]" italics="true" pageId="7" pageNumber="134">Swanepoel 581</emphasis>
(WIND!).—1913:
<quantity id="4CD29B428819FFCB4C36FEF0FB47D464" box="[1100,1171,302,328]" metricMagnitude="4" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="134" unit="km" value="33.0">33 km</quantity>
from Sesfontein turn-off to Hoanib River, (–AB),
<quantity id="4CD29B428819FFCB49DFFE8CFE3BD441" box="[421,495,338,365]" metricMagnitude="2" metricUnit="m" metricValue="3.47" pageId="7" pageNumber="134" unit="m" value="347.0">347 m</quantity>
,
<date id="FF9410678819FFCB4987FE8CFD4CD440" box="[509,664,338,365]" pageId="7" pageNumber="134" value="2014-05-17">17 May 2014</date>
,
<emphasis id="B95EEAB58819FFCB4ADEFE8CFCBBD440" box="[676,879,338,364]" italics="true" pageId="7" pageNumber="134">Klaassen EK2542</emphasis>
(WIND!);
<quantity id="4CD29B428819FFCB4B8AFE8CFBEDD440" box="[1008,1081,338,364]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="134" unit="km" value="20.0">20 km</quantity>
Noord van Sesfontein, dolomiet berge, (–BA),
<date id="FF9410678819FFCB4953FEA8FE15D4BC" box="[297,449,374,401]" pageId="7" pageNumber="134" value="1976-05-21">21 May 1976</date>
,
<emphasis id="B95EEAB58819FFCB49B6FEA8FD9FD4BC" box="[460,587,374,400]" italics="true" pageId="7" pageNumber="134">Viljoen 313</emphasis>
(WIND!); ca.
<quantity id="4CD29B428819FFCB4A8AFEA8FCFCD4BC" box="[752,808,374,400]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="134" unit="km" value="3.0">3 km</quantity>
W of Sesfontein, (–BA),
<quantity id="4CD29B428819FFCB4C3DFEA8FB44D4BD" box="[1095,1168,374,401]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="134" unit="m" value="600.0">600 m</quantity>
,
<date id="FF9410678819FFCB4CE1FEA8FAECD4BD" box="[1179,1336,374,401]" pageId="7" pageNumber="134" value="1985-04-11">11 April 1985</date>
,
<emphasis id="B95EEAB58819FFCB4D39FEA8FEF6D498" italics="true" pageId="7" pageNumber="134">Jacobsen &amp; Moss K130</emphasis>
(WIND!); Dry water course in calcrete hills NW of Sesfontein, (–BA),
<date id="FF9410678819FFCB4C46FE44FAC2D499" box="[1084,1302,410,437]" pageId="7" pageNumber="134" value="1995-09-11">11 September 1995</date>
,
<emphasis id="B95EEAB58819FFCB4D5AFE44FA7FD498" box="[1312,1451,410,436]" italics="true" pageId="7" pageNumber="134">Sullivan 350</emphasis>
(WIND!);
<quantity id="4CD29B428819FFCB4885FE60FEECD4F4" box="[255,312,446,472]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="134" unit="km" value="2.0">2 km</quantity>
NW of Sesfontein on D 3707, extremely dry slate slopes hosting on
<taxonomicName id="4C2A4D248819FFCB4C47FE60FB5AD4F4" box="[1085,1166,446,472]" class="Magnoliopsida" family="Acanthaceae" genus="Ruellia" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="134" phylum="Tracheophyta" rank="genus">
<emphasis id="B95EEAB58819FFCB4C47FE60FB5AD4F4" box="[1085,1166,446,472]" italics="true" pageId="7" pageNumber="134">Ruellia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A4D248819FFCB4CE6FE61FB27D4F4" box="[1180,1267,447,472]" class="Magnoliopsida" family="Acanthaceae" genus="Justicia" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="134" phylum="Tracheophyta" rank="genus">
<emphasis id="B95EEAB58819FFCB4CE6FE61FB27D4F4" box="[1180,1267,447,472]" italics="true" pageId="7" pageNumber="134">Justicia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2A4D248819FFCB4D53FE60FA70D4F4" authorityName="Nees von Esenbeck" authorityYear="1832" box="[1321,1444,446,472]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="134" phylum="Tracheophyta" rank="genus">
<emphasis id="B95EEAB58819FFCB4D53FE60FA70D4F4" box="[1321,1444,446,472]" italics="true" pageId="7" pageNumber="134">Petalidium</emphasis>
</taxonomicName>
, (–BA),
<date id="FF9410678819FFCB48A5FE3CFE5BD4D0" box="[223,399,482,508]" pageId="7" pageNumber="134" value="2010-03-26">26 March 2010</date>
,
<emphasis id="B95EEAB58819FFCB49E1FE3DFD60D4D0" box="[411,692,482,509]" italics="true" pageId="7" pageNumber="134">Tripp &amp; Dexter 865–869</emphasis>
(PRE!, WIND!); Mountains
<quantity id="4CD29B428819FFCB4B84FE3CFBECD4D0" box="[1022,1080,482,509]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="134" unit="km" value="5.0">5 km</quantity>
southwest of Sesfontein, (–BA),
<quantity id="4CD29B428819FFCB48F2FDD8FF04D70D" box="[136,208,518,545]" metricMagnitude="2" metricUnit="m" metricValue="4.99" pageId="7" pageNumber="134" unit="m" value="499.0">499 m</quantity>
,
<date id="FF9410678819FFCB48A1FDD8FEA5D70C" box="[219,369,518,545]" pageId="7" pageNumber="134" value="2022-05-12">12 May 2022</date>
,
<emphasis id="B95EEAB58819FFCB4906FDD8FDF1D70C" box="[380,549,518,544]" italics="true" pageId="7" pageNumber="134">Swanepoel 578</emphasis>
(WIND!); Hoanib Poort
<quantity id="4CD29B428819FFCB4B46FDD8FC55D70C" box="[828,897,518,544]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="134" unit="km" value="11.0">11 km</quantity>
east of Sesfontein on road to Warmquelle, (–BA),
<quantity id="4CD29B428819FFCB48F2FDF4FF05D769" box="[136,209,554,581]" metricMagnitude="2" metricUnit="m" metricValue="6.09" pageId="7" pageNumber="134" unit="m" value="609.0">609 m</quantity>
,
<date id="FF9410678819FFCB48A7FDF4FEA1D768" box="[221,373,554,581]" pageId="7" pageNumber="134" value="2022-05-15">15 May 2022</date>
,
<emphasis id="B95EEAB58819FFCB49FAFDF4FDFFD768" box="[384,555,554,580]" italics="true" pageId="7" pageNumber="134">Swanepoel 582</emphasis>
(WIND!);
<quantity id="4CD29B428819FFCB4AD3FDF4FD36D768" box="[681,738,554,580]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="134" unit="km" value="6.0">6 km</quantity>
east of Ganamub on road to Sesfontein, (–BA),
<quantity id="4CD29B428819FFCB4D7AFDF4FA9DD769" box="[1280,1353,554,581]" metricMagnitude="2" metricUnit="m" metricValue="6.31" pageId="7" pageNumber="134" unit="m" value="631.0">631 m</quantity>
,
<date id="FF9410678819FFCB4D2FFDF4FF17D744" pageId="7" pageNumber="134" value="2022-05-25">25 May 2022</date>
,
<emphasis id="B95EEAB58819FFCB48B4FD90FEACD744" box="[206,376,590,616]" italics="true" pageId="7" pageNumber="134">Swanepoel 584</emphasis>
(WIND!);
<quantity id="4CD29B428819FFCB498CFD90FDE4D744" box="[502,560,590,616]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="134" unit="km" value="9.0">9 km</quantity>
east-northeast of Warmquelle in poort on road to Beesvlakte, (–BB),
<quantity id="4CD29B428819FFCB4D40FD90FA57D745" box="[1338,1411,590,617]" metricMagnitude="2" metricUnit="m" metricValue="9.19" pageId="7" pageNumber="134" unit="m" value="919.0">919 m</quantity>
,
<date id="FF9410678819FFCB4DF5FD90FF29D7A0" pageId="7" pageNumber="134" value="2022-05-12">12 May 2022</date>
,
<emphasis id="B95EEAB58819FFCB497DFDACFE64D7A0" box="[263,432,626,652]" italics="true" pageId="7" pageNumber="134">Swanepoel 576</emphasis>
(WIND!);
<quantity id="4CD29B428819FFCB4A56FDACFDA6D7A0" box="[556,626,626,652]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="134" unit="km" value="12.0">12 km</quantity>
east-northeast of Warmquelle in poort on road to Beesvlakte, (–BB),
<quantity id="4CD29B428819FFCB4D09FDACFF74D79D" metricMagnitude="3" metricUnit="m" metricValue="1.068" pageId="7" pageNumber="134" unit="m" value="1068.0">1068 m</quantity>
,
<date id="FF9410678819FFCB48D1FD48FE95D79C" box="[171,321,662,689]" pageId="7" pageNumber="134" value="2022-05-12">12 May 2022</date>
,
<emphasis id="B95EEAB58819FFCB4936FD48FE21D79D" box="[332,501,662,689]" italics="true" pageId="7" pageNumber="134">Swanepoel 577</emphasis>
(WIND!);
<quantity id="4CD29B428819FFCB4A0BFD48FD63D79C" box="[625,695,662,688]" metricMagnitude="4" metricUnit="m" metricValue="3.7" pageId="7" pageNumber="134" unit="km" value="37.0">37 km</quantity>
from Khowarib on road to Palmwag, (–DB),
<quantity id="4CD29B428819FFCB4CCAFD48FAD2D79D" box="[1200,1286,662,689]" metricMagnitude="3" metricUnit="m" metricValue="1.0059999999999998" pageId="7" pageNumber="134" unit="m" value="1006.0">1006 m</quantity>
,
<date id="FF9410678819FFCB4D6BFD48FA73D79C" box="[1297,1447,662,689]" pageId="7" pageNumber="134" value="2022-05-12">12 May 2022</date>
,
<emphasis id="B95EEAB58819FFCB48F2FD64FEE6D7F9" box="[136,306,698,725]" italics="true" pageId="7" pageNumber="134">Swanepoel 575</emphasis>
(WIND).
</paragraph>
</subSubSection>
</treatment>
</document>
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