665 lines
77 KiB
XML
665 lines
77 KiB
XML
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<mods:title id="19D5EF1A68CC146D583D878F9EA934F7">Revising the taxonomy of Darevskia valentini (Boettger, 1892) and Darevskia rudis (Bedriaga, 1886) (Squamata, Lacertidae): a Morpho-Phylogenetic integrated study in a complex Anatolian scenario</mods:title>
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<mods:namePart id="BA9ED1CF175E349CCD21E2D8EC007837">Arribas, Oscar</mods:namePart>
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<mods:namePart id="83F0E3B488E0C98046A18A5348BA00DA">Candan, Kamil</mods:namePart>
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<mods:namePart id="161C77B041A42C14853DF45726F9B6CF">Kumlutaş, Yusuf</mods:namePart>
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<mods:namePart id="04F1D7871E5D125BABE1B844D28EC445">Gül, Serkan</mods:namePart>
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<treatment id="038387B0424CFE78FF3B4F561298F9D9" ID-DOI="http://doi.org/10.5281/zenodo.7517913" ID-GBIF-Taxon="208791118" ID-Zenodo-Dep="7517913" LSID="urn:lsid:plazi:treatment:038387B0424CFE78FF3B4F561298F9D9" httpUri="http://treatment.plazi.org/id/038387B0424CFE78FF3B4F561298F9D9" lastPageId="22" lastPageNumber="23" pageId="18" pageNumber="19">
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<taxonomicName id="4C2A4D25424CFE7CFF3B4F5612E7F9E8" authority="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz, 2022" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" box="[151,465,1616,1642]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="josefschmidtleri" status="sp. nov.">
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<emphasis id="B95EEAB4424CFE7CFF3B4F5612E7F9E8" box="[151,465,1616,1642]" italics="true" pageId="18" pageNumber="19">Darevskia josefschmidtleri</emphasis>
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</taxonomicName>
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<emphasis id="B95EEAB4424CFE7CFE754F571100F9E9" bold="true" box="[473,566,1617,1643]" pageId="18" pageNumber="19">
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<taxonomicNameLabel id="A26D57CF424CFE7CFE754F571100F9E9" box="[473,566,1617,1643]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9536A6424CFE7CFF3B4F72129FF90C" blockId="18.[151,566,1616,1679]" box="[151,425,1652,1679]" pageId="18" pageNumber="19">
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<emphasis id="B95EEAB4424CFE7CFF3B4F72129FF90C" bold="true" box="[151,425,1652,1679]" pageId="18" pageNumber="19">
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(Appendix 5;
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<figureCitation id="13112A23424CFE7CFE974F7212ACF90C" box="[315,410,1652,1678]" captionStart="FIGURE 12" captionStartId="22.[152,255,955,979]" captionTargetBox="[151,1413,392,928]" captionTargetId="figure-129@22.[151,1435,381,931]" captionTargetPageId="22" captionText="FIGURE 12. a) Darevskia spitzenbergerae wernermayeri ssp. nov. (Paratype; nº 12, Male; Başeğmez Village, Çaldıran, Turkey); b) Darevskia mirabilis stat. nov. (Paratype; nº 5, Female; Ovit Pass, Kaçkar Mountains, Rize, Turkey); c) Darevskia rudis bolkardaghica (Paratype; nº 1, Male; Karagöl, Ulukışla, Niğde, Central Anatolia, Turkey); d) Darevskia rudis lantzicyreni comb. nov. (nº 23, male; Kümbet Village, Zara, Turkey); e) Darevskia josefschmidtleri sp. nov. (Paratype; nº 20, Male; Yukarınarlıca Village, Çatak, Van, Turkey); f) Darevskia valentini (nº 9, Male; Tepeler Village, Ardahan, Turkey) and temporal area of an Armenian specimen (Karvansaray, Martuni District, Armenia); g) Darevskia spitzenbergerae spitzenbergerae stat. et comb. nov. (nº 1, Male; Cilo Sat Mountains, Hakkari, Turkey)- Also, temporal area of other specimen from the same locality. The new nomenclature proposed in the text is used." figureDoi="http://doi.org/10.5281/zenodo.7517939" httpUri="https://zenodo.org/record/7517939/files/figure.png" pageId="18" pageNumber="19">Fig. 12e</figureCitation>
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).
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C330652D424CFE7CFF3B4FBA167AF8F7" pageId="18" pageNumber="19" type="description">
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<paragraph id="8B9536A6424CFE7CFF3B4FBA1288F954" blockId="18.[151,446,1724,1750]" box="[151,446,1724,1750]" pageId="18" pageNumber="19">
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<emphasis id="B95EEAB4424CFE7CFF3B4FBA1288F954" bold="true" box="[151,446,1724,1750]" pageId="18" pageNumber="19">Synonymy/Chresonymy:</emphasis>
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</paragraph>
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<paragraph id="8B9536A6424CFE7CFF3B4E041235F8BA" blockId="18.[151,1437,1793,1909]" pageId="18" pageNumber="19">
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<taxonomicName id="4C2A4D25424CFE7CFF3B4E04111BF89B" authority="" Zwischenpopulation" authorityName="Zwischenpopulation" box="[151,557,1793,1817]" class="Reptilia" family="Lacertidae" genus="Lacerta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="valentini">
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<emphasis id="B95EEAB4424CFE7CFF3B4E04127FF89B" box="[151,329,1793,1817]" italics="true" pageId="18" pageNumber="19">Lacerta valentini</emphasis>
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“Zwischenpopulation
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</taxonomicName>
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||
”(intermediate population);
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<bibRefCitation id="EFBB4B57424CFE7CFCE34E0717E0F89B" author="Eiselt, J. & Darevsky, I. S. & Schmidtler, J. F." box="[847,1238,1793,1817]" pageId="18" pageNumber="19" pagination="1 - 18" refId="ref25991" refString="Eiselt, J., Darevsky, I. S. & Schmidtler, J. F. (1992) Untersuchungen an Felseneidechsen (Lacerta saxicola komplex) in der ostlichen Turkei, I. Lacerta valentini Boettger. Annalen des Naturhistorischen Museums in Wien, 93 (B), 1 - 18." type="journal article" year="1992">Eiselt, Darevsky & Schmidtler, 1992</bibRefCitation>
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||
. (from “Çaldıran”- sic.!-)
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</paragraph>
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<taxonomicName id="4C2A4D25424CFE7CFF3B4E381268F8D4" baseAuthorityName="Boettger" baseAuthorityYear="1892" box="[151,350,1854,1878]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="valentini">
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<emphasis id="B95EEAB4424CFE7CFF3B4E381268F8D4" box="[151,350,1854,1878]" italics="true" pageId="18" pageNumber="19">Darevskia valentini</emphasis>
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</taxonomicName>
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||
“Clade B”;
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||
<bibRefCitation id="EFBB4B57424CFE7CFE764E38119CF8D4" author="Candan, K. & Kornilios, P. & Ayaz, D. & Kumlutas, Y. & Gul, S. & Yildirim-Caynak, E. & Ilgaz, C." box="[474,682,1854,1878]" pageId="18" pageNumber="19" pagination="665 - 681" refId="ref25168" refString="Candan, K., Kornilios, P., Ayaz, D., Kumlutas, Y., Gul, S., Yildirim-Caynak, E. & Ilgaz, C. (2021) Cryptic genetic structure within Valentin's Lizard, Darevskia valentini (Boettger, 1892) (Squamata, Lacertidae), with implications for systematics and origins of parthenogenesis. Systematics and Biodiversity, 19 (7), 665 - 681. https: // doi. org / 10.1080 / 14772000.2021.1909171" type="journal article" year="2021">
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||
Candan
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||
<emphasis id="B95EEAB4424CFE7CFD824E391157F8D4" box="[558,609,1854,1878]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
|
||
(2021)
|
||
</bibRefCitation>
|
||
(same locality as this study)
|
||
</paragraph>
|
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<paragraph id="8B9536A6424CFE7CFF3B4E5B167AF8F7" blockId="18.[151,1437,1793,1909]" box="[151,1356,1885,1909]" pageId="18" pageNumber="19">
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<emphasis id="B95EEAB4424CFE7CFF3B4E5B115CF8F7" bold="true" box="[151,618,1885,1909]" pageId="18" pageNumber="19">ZooBank registration (http://zoobank.org):</emphasis>
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urn:lsid:zoobank.org:act:
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<uuid id="FF8C0C73424CFE7CFCCA4E5B167EF8F7" box="[870,1352,1885,1909]" pageId="18" pageNumber="19">56CFE08E-164E-485B-8F4C-94EA76293128</uuid>
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</uri>
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||
.
|
||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C330652D424CFE7BFF3B4EA517D1FEE6" lastPageId="21" lastPageNumber="22" pageId="18" pageNumber="19" type="materials_examined">
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<paragraph id="8B9536A6424CFE7CFF3B4EA5108CF863" blockId="18.[151,1436,1955,2018]" pageId="18" pageNumber="19">
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<materialsCitation id="3B423CFB424CFE7CFF3B4EA5108CF863" ID-GBIF-Occurrence="4012805377" collectingDate="2016-06-24" collectionCode="ZDEU" collectorName="Kamil Candan" country="Turkey" location="Basegmez Village" pageId="18" pageNumber="19" specimenCode="ZDEU220" specimenCount="1" stateProvince="Van" typeStatus="holotype">
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<emphasis id="B95EEAB4424CFE7CFF3B4EA5123BF83F" bold="true" box="[151,269,1955,1981]" pageId="18" pageNumber="19">
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<typeStatus id="54918804424CFE7CFF3B4EA5123EF83F" box="[151,264,1955,1981]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
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:
|
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</emphasis>
|
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<specimenCode id="DB8C9EDD424CFE7CFEB94EA512BFF83F" box="[277,393,1955,1981]" collectionCode="ZDEU" country="Turkey" httpUri="http://grbio.org/cool/s077-eqrn" name="Zoology Department, Ege University" pageId="18" pageNumber="19">ZDEU220</specimenCode>
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||
/2016 (n.2). ♁,
|
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<location id="8EF5607D424CFE7CFD944EA21030F83F" LSID="urn:lsid:plazi:treatment:038387B0424CFE78FF3B4F561298F9D9:8EF5607D424CFE7CFD944EA21030F83F" box="[568,774,1955,1982]" country="Turkey" name="Basegmez Village" pageId="18" pageNumber="19" stateProvince="Van">Başeğmez Village</location>
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, Çaldıran,
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<collectingRegion id="49EEF844424CFE7CFC2C4EA21099F83C" box="[896,943,1956,1982]" country="Turkey" name="Van" pageId="18" pageNumber="19">Van</collectingRegion>
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,
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<collectingCountry id="F33D7636424CFE7CFC154EA5173DF83F" box="[953,1035,1955,1981]" name="Turkey" pageId="18" pageNumber="19">Turkey</collectingCountry>
|
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. leg.
|
||
<collectorName id="26DF5370424CFE7CFBEA4EA517DAF83F" box="[1094,1260,1955,1981]" pageId="18" pageNumber="19">Kamil Candan</collectorName>
|
||
, Nurettin Beşer and Mehmet Kürşat Şahin,
|
||
<date id="FF941066424CFE7CFE664EC1117AF863" box="[458,588,1991,2017]" pageId="18" pageNumber="19" value="2016-06-24">
|
||
<collectingDate id="EFD0E98E424CFE7CFE664EC1117AF863" box="[458,588,1991,2017]" pageId="18" pageNumber="19" value="2016-06-24">24.06.2016</collectingDate>
|
||
</date>
|
||
. Conserved in
|
||
<collectionCode id="ED3BAE63424CFE7CFD5E4ECE100BF860" box="[754,829,1992,2018]" country="Turkey" httpUri="http://grbio.org/cool/s077-eqrn" name="Zoology Department, Ege University" pageId="18" pageNumber="19">ZDEU</collectionCode>
|
||
collection.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<caption id="DF55662E424DFE7DFF3B4F54146AF90D" ID-DOI="http://doi.org/10.5281/zenodo.7517933" ID-Zenodo-Dep="7517933" httpUri="https://zenodo.org/record/7517933/files/figure.png" pageId="19" pageNumber="20" startId="19.[151,254,1618,1643]" targetBox="[228,1957,510,1590]" targetPageId="19">
|
||
<paragraph id="8B9536A6424DFE7DFF3B4F54146AF90D" blockId="19.[151,2037,1618,1679]" pageId="19" pageNumber="20">
|
||
<emphasis id="B95EEAB4424DFE7DFF3B4F541212F9E9" bold="true" box="[151,292,1618,1643]" pageId="19" pageNumber="20">FIGURE 10.</emphasis>
|
||
Parsimony networks corresponding to
|
||
<emphasis id="B95EEAB4424DFE7DFD034F5411D1F9E9" box="[687,743,1618,1643]" italics="true" pageId="19" pageNumber="20">Cyt-b</emphasis>
|
||
(A) and
|
||
<emphasis id="B95EEAB4424DFE7DFCEC4F5410B4F9E9" box="[832,898,1618,1643]" italics="true" pageId="19" pageNumber="20">MC1R</emphasis>
|
||
(B) represent reconstruction of the studied group. Numbers within parentheses represent a mutational step, black circles missing haplotypes, and colored circles haplotypes. The circle area is proportional to the number of individuals. The new nomenclature proposed in the text is used.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF55662E424AFE7AFF344FD71791F867" ID-DOI="http://doi.org/10.5281/zenodo.7517937" ID-Zenodo-Dep="7517937" httpUri="https://zenodo.org/record/7517937/files/figure.png" pageId="20" pageNumber="21" startId="20.[152,253,1745,1769]" targetBox="[449,1137,190,1710]" targetPageId="20">
|
||
<paragraph id="8B9536A6424AFE7AFF344FD71791F867" blockId="20.[151,1437,1745,2022]" pageId="20" pageNumber="21">
|
||
<emphasis id="B95EEAB4424AFE7AFF344FD71212F96B" bold="true" box="[152,292,1745,1769]" pageId="20" pageNumber="21">FIGURE 11.</emphasis>
|
||
Escalation of the calf area in the
|
||
<taxonomicName id="4C2A4D25424AFE7AFD2A4FD7118FF96B" baseAuthorityName="Bedriaga" baseAuthorityYear="1886" box="[646,697,1745,1769]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="rudis">
|
||
<emphasis id="B95EEAB4424AFE7AFD2A4FD7118FF96B" box="[646,697,1745,1769]" italics="true" pageId="20" pageNumber="21">rudis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2A4D25424AFE7AFD424FD71073F96B" baseAuthorityName="Boettger" baseAuthorityYear="1892" box="[750,837,1745,1769]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="valentini">
|
||
<emphasis id="B95EEAB4424AFE7AFD424FD71073F96B" box="[750,837,1745,1769]" italics="true" pageId="20" pageNumber="21">valentini</emphasis>
|
||
</taxonomicName>
|
||
complexes. Scale size (great, medium, small) and keeling development (weak keeling, medium keeling, strong keeling). The new nomenclature proposed in the text is used.
|
||
<taxonomicName id="4C2A4D25424AFE7AFB544FF016AAF88F" baseAuthorityName="Darevsky & Eiselt" baseAuthorityYear="1967" box="[1272,1436,1781,1805]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="subSpecies" species="valentini" subSpecies="lantzicyreni">
|
||
<emphasis id="B95EEAB4424AFE7AFB544FF016AAF88F" box="[1272,1436,1781,1805]" italics="true" pageId="20" pageNumber="21">D. r. lantzicyreni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAB4424AFE7AFF3B4E1C1230F8B0" bold="true" box="[151,262,1818,1842]" pageId="20" pageNumber="21">comb. nov.</emphasis>
|
||
(medium scale size, medium keeling);
|
||
<taxonomicName id="4C2A4D25424AFE7AFD2D4E1C100AF8B3" authority="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz, 2022" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" box="[641,828,1817,1841]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="josefschmidtleri" status="sp. nov.">
|
||
<emphasis id="B95EEAB4424AFE7AFD2D4E1C100AF8B3" box="[641,828,1817,1841]" italics="true" pageId="20" pageNumber="21">D. josefschmidtleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAB4424AFE7AFCEC4E1C10BBF8B0" bold="true" box="[832,909,1818,1842]" pageId="20" pageNumber="21">
|
||
<taxonomicNameLabel id="A26D57CF424AFE7AFCEC4E1C10BBF8B0" box="[832,909,1818,1842]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(small scale size, weak keeling);
|
||
<taxonomicName id="4C2A4D25424AFE7AFB624E1C16ABF8B3" baseAuthorityName="Eiselt, Darevsky & Schmidtler" baseAuthorityYear="1992" box="[1230,1437,1817,1841]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="subSpecies" species="valentini" subSpecies="spitzenbergerae">
|
||
<emphasis id="B95EEAB4424AFE7AFB624E1C16ABF8B3" box="[1230,1437,1817,1841]" italics="true" pageId="20" pageNumber="21">D. s. spitzenbergerae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAB4424AFE7AFF3B4E381262F8D4" bold="true" box="[151,340,1854,1878]" pageId="20" pageNumber="21">stat. et comb. nov.</emphasis>
|
||
(small scale size, weak keeling);
|
||
<emphasis id="B95EEAB4424AFE7AFD0D4E381056F8D7" box="[673,864,1854,1877]" italics="true" pageId="20" pageNumber="21">D. s. wernermayeri</emphasis>
|
||
<emphasis id="B95EEAB4424AFE7AFCCA4E381089F8D4" bold="true" box="[870,959,1854,1878]" pageId="20" pageNumber="21">ssp. nov.</emphasis>
|
||
(medium scale size, weak keeling);
|
||
<taxonomicName id="4C2A4D25424AFE7AFA8B4E3816AAF8D7" baseAuthorityName="Boettger" baseAuthorityYear="1892" box="[1319,1436,1853,1877]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="valentini">
|
||
<emphasis id="B95EEAB4424AFE7AFA8B4E3816AAF8D7" box="[1319,1436,1853,1877]" italics="true" pageId="20" pageNumber="21">D. valentini</emphasis>
|
||
</taxonomicName>
|
||
(medium scale size, weak keeling);
|
||
<emphasis id="B95EEAB4424AFE7AFE5A4E6411B1F8FB" box="[502,647,1889,1913]" italics="true" pageId="20" pageNumber="21">
|
||
D. b.
|
||
<taxonomicName id="4C2A4D25424AFE7AFD874E6711B1F8FB" baseAuthorityName="Mehely" baseAuthorityYear="1909" box="[555,647,1889,1913]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="bithynica">bithynica</taxonomicName>
|
||
</emphasis>
|
||
(small scale size, weak keeling);
|
||
<emphasis id="B95EEAB4424AFE7AFC7C4E641708F8FB" box="[976,1086,1889,1913]" italics="true" pageId="20" pageNumber="21">D. b. tristis</emphasis>
|
||
(medium scale size, weak keeling);
|
||
<emphasis id="B95EEAB4424AFE7AFF3B4E801215F81F" box="[151,291,1925,1949]" italics="true" pageId="20" pageNumber="21">D. o. bischoffi</emphasis>
|
||
<emphasis id="B95EEAB4424AFE7AFE8B4E8012AEF81C" bold="true" box="[295,408,1926,1950]" pageId="20" pageNumber="21">
|
||
<taxonomicNameLabel id="A26D57CF424AFE7AFE8B4E8012AEF81C" box="[295,408,1926,1950]" pageId="20" pageNumber="21" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(great scale size, strongly keeling);
|
||
<taxonomicName id="4C2A4D25424AFE7AFD564E801049F81F" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" baseAuthorityName="Lantz & Cyren" baseAuthorityYear="1936" box="[762,895,1925,1949]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="subSpecies" species="obscura" subSpecies="obscura">
|
||
<emphasis id="B95EEAB4424AFE7AFD564E801049F81F" box="[762,895,1925,1949]" italics="true" pageId="20" pageNumber="21">D. o. obscura</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAB4424AFE7AFC284E801776F81C" bold="true" box="[900,1088,1926,1950]" pageId="20" pageNumber="21">stat. et comb. nov.</emphasis>
|
||
(great scale size, strongly keeling);
|
||
<emphasis id="B95EEAB4424AFE7AFF3B4EAC125EF843" box="[151,360,1961,1985]" italics="true" pageId="20" pageNumber="21">D. o. macromaculata</emphasis>
|
||
<emphasis id="B95EEAB4424AFE7AFEC14EAC12E8F840" bold="true" box="[365,478,1962,1986]" pageId="20" pageNumber="21">comb. nov.</emphasis>
|
||
(great scale size, strongly keeling);
|
||
<taxonomicName id="4C2A4D25424AFE7AFCED4EAC1734F843" authorityName="Arribas, Ilgaz, Kumlutas, Durmus, Avci & Uzum" authorityYear="2013" box="[833,1026,1961,1985]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="subSpecies" species="rudis" subSpecies="bolkardaghica">
|
||
<emphasis id="B95EEAB4424AFE7AFCED4EAC1734F843" box="[833,1026,1961,1985]" italics="true" pageId="20" pageNumber="21">D. r. bolkardaghica</emphasis>
|
||
</taxonomicName>
|
||
(medium scale size, medium keeling);
|
||
<taxonomicName id="4C2A4D25424AFE7AFA284EAC13FCF867" baseAuthorityName="Bedriaga" baseAuthorityYear="1886" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="rudis">
|
||
<emphasis id="B95EEAB4424AFE7AFA284EAC13FCF867" italics="true" pageId="20" pageNumber="21">D. rudis</emphasis>
|
||
</taxonomicName>
|
||
(medium scale size, medium keeling);
|
||
<taxonomicName id="4C2A4D25424AFE7AFDFD4EC811FFF867" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" baseAuthorityName="Arribas, Ilgaz, Kumlutas, Durmus, Avci & Uzum" baseAuthorityYear="2013" box="[593,713,1997,2021]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="mirabilis" status="stat. nov.">
|
||
<emphasis id="B95EEAB4424AFE7AFDFD4EC811FFF867" box="[593,713,1997,2021]" italics="true" pageId="20" pageNumber="21">D. mirabilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAB4424AFE7AFD634EC8101BF864" bold="true" box="[719,813,1998,2022]" pageId="20" pageNumber="21">
|
||
<taxonomicNameLabel id="A26D57CF424AFE7AFD634EC8101BF864" box="[719,813,1998,2022]" pageId="20" pageNumber="21">stat. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(medium scale size, medium keeling.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9536A6424BFE7BFF6B499117D1FEE6" blockId="21.[151,1437,151,2036]" pageId="21" pageNumber="22">
|
||
<materialsCitation id="3B423CFB424BFE7BFF6B49911609FF33" ID-GBIF-Occurrence="4012805479" box="[199,1343,151,179]" collectionCode="ZDEU" collectorName="Same" pageId="21" pageNumber="22" specimenCode="ZDEU221" specimenCount="11" specimenCount-female="9" specimenCount-subadult="2" typeStatus="paratype">
|
||
<emphasis id="B95EEAB4424BFE7BFF6B4991127EFF33" bold="true" box="[199,328,151,177]" pageId="21" pageNumber="22">
|
||
<typeStatus id="54918804424BFE7BFF6B49911275FF33" box="[199,323,151,177]" pageId="21" pageNumber="22" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
8 ♁♁,
|
||
<specimenCount id="9D2CFD2F424BFE7BFE0F499112D3FF33" box="[419,485,151,177]" pageId="21" pageNumber="22" type="female">8 ♀♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D2CFD2F424BFE7BFE584991115CFF33" box="[500,618,151,177]" pageId="21" pageNumber="22" type="subadult">2 subadult</specimenCount>
|
||
<specimenCount id="9D2CFD2F424BFE7BFDDF499111A9FF33" box="[627,671,151,177]" pageId="21" pageNumber="22" type="female">♀♀</specimenCount>
|
||
.
|
||
<collectorName id="26DF5370424BFE7BFD02499111DAFF33" box="[686,748,151,177]" pageId="21" pageNumber="22">Same</collectorName>
|
||
locality, date and collectors as holotype.
|
||
<specimenCode id="DB8C9EDD424BFE7BFB6649911609FF33" box="[1226,1343,151,177]" collectionCode="ZDEU" country="Turkey" httpUri="http://grbio.org/cool/s077-eqrn" name="Zoology Department, Ege University" pageId="21" pageNumber="22">ZDEU221</specimenCode>
|
||
</materialsCitation>
|
||
/2016, 6 ♁♁,
|
||
<materialsCitation id="3B423CFB424BFE7BFF7C49BC12DFFF7A" ID-GBIF-Occurrence="4012805332" collectingDate="2016-06-22" collectionCode="ZDEU" collectorName="Kamil Candan & Nurettin Beser & Mehmet Kursat Sahin" country="Turkey" location="Cirisli Village" municipality="Cat" pageId="21" pageNumber="22" specimenCode="ZDEU222" specimenCount="13" specimenCount-female="12" specimenCount-juvenile="1" stateProvince="Erzurum" typeStatus="paratype">
|
||
<specimenCount id="9D2CFD2F424BFE7BFF7C49BC1228FF57" box="[208,286,186,213]" pageId="21" pageNumber="22" type="female">12 ♀♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D2CFD2F424BFE7BFE8049BC12A8FF56" box="[300,414,186,212]" pageId="21" pageNumber="22" type="juvenile">1 juvenile</specimenCount>
|
||
,
|
||
<location id="8EF5607D424BFE7BFE0649BC117DFF56" LSID="urn:lsid:plazi:treatment:038387B0424CFE78FF3B4F561298F9D9:8EF5607D424BFE7BFE0649BC117DFF56" box="[426,587,186,212]" country="Turkey" municipality="Cat" name="Cirisli Village" pageId="21" pageNumber="22" stateProvince="Erzurum">Çirişli Village</location>
|
||
,
|
||
<collectingMunicipality id="6BF1ACDC424BFE7BFDFA49BC11B6FF56" box="[598,640,186,212]" pageId="21" pageNumber="22">Çat</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49EEF844424BFE7BFD2049BD11C7FF57" box="[652,753,187,213]" country="Turkey" name="Erzurum" pageId="21" pageNumber="22">Erzurum</collectingRegion>
|
||
,
|
||
<collectingCountry id="F33D7636424BFE7BFD5049BC1078FF56" box="[764,846,186,212]" name="Turkey" pageId="21" pageNumber="22">Turkey</collectingCountry>
|
||
. leg.
|
||
<collectorName id="26DF5370424BFE7BFC2549BC1719FF56" box="[905,1071,186,212]" pageId="21" pageNumber="22">Kamil Candan</collectorName>
|
||
,
|
||
<collectorName id="26DF5370424BFE7BFB9749BC17D4FF57" box="[1083,1250,186,213]" pageId="21" pageNumber="22">Nurettin Beşer</collectorName>
|
||
,
|
||
<collectorName id="26DF5370424BFE7BFB4149BC13ECFF7A" pageId="21" pageNumber="22">Mehmet Kürşat Şahin</collectorName>
|
||
,
|
||
<date id="FF941066424BFE7BFF4A49D81251FF7A" box="[230,359,222,248]" pageId="21" pageNumber="22" value="2016-06-22">
|
||
<collectingDate id="EFD0E98E424BFE7BFF4A49D81251FF7A" box="[230,359,222,248]" pageId="21" pageNumber="22" value="2016-06-22">22.06.2016</collectingDate>
|
||
</date>
|
||
.
|
||
<specimenCode id="DB8C9EDD424BFE7BFEDF49D812DFFF7A" box="[371,489,222,248]" collectionCode="ZDEU" country="Turkey" httpUri="http://grbio.org/cool/s077-eqrn" name="Zoology Department, Ege University" pageId="21" pageNumber="22">ZDEU222</specimenCode>
|
||
</materialsCitation>
|
||
/2016, 5 ♁♁,
|
||
<materialsCitation id="3B423CFB424BFE7BFD2849D81080FE9F" ID-GBIF-Occurrence="4012805338" collectingDate="2016-01-07" collectionCode="ZDEU" collectorName="Kamil Candan & Nurettin Beser & Mehmet Kursat Sahin" country="Turkey" location="Palandoken Mountain" pageId="21" pageNumber="22" specimenCode="ZDEU119" specimenCount="9" specimenCount-female="8" specimenCount-juvenile="1" stateProvince="Erzurum" typeStatus="paratype">
|
||
<specimenCount id="9D2CFD2F424BFE7BFD2849D811F3FF7B" box="[644,709,222,249]" pageId="21" pageNumber="22" type="female">8 ♀♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D2CFD2F424BFE7BFD7F49D81070FF7A" box="[723,838,222,248]" pageId="21" pageNumber="22" type="juvenile">1 juvenile</specimenCount>
|
||
,
|
||
<location id="8EF5607D424BFE7BFCFE49D8177AFF7A" LSID="urn:lsid:plazi:treatment:038387B0424CFE78FF3B4F561298F9D9:8EF5607D424BFE7BFCFE49D8177AFF7A" box="[850,1100,222,248]" country="Turkey" name="Palandoken Mountain" pageId="21" pageNumber="22" stateProvince="Erzurum">Palandöken Mountain</location>
|
||
,
|
||
<collectingRegion id="49EEF844424BFE7BFBF449D9178BFF7B" box="[1112,1213,223,249]" country="Turkey" name="Erzurum" pageId="21" pageNumber="22">Erzurum</collectingRegion>
|
||
,
|
||
<collectingCountry id="F33D7636424BFE7BFB6449D8162CFF7A" box="[1224,1306,222,248]" name="Turkey" pageId="21" pageNumber="22">Turkey</collectingCountry>
|
||
. leg.
|
||
<collectorName id="26DF5370424BFE7BFAFA49D813D9FE9E" pageId="21" pageNumber="22">Kamil Candan</collectorName>
|
||
,
|
||
<collectorName id="26DF5370424BFE7BFF5748041295FE9E" box="[251,419,258,284]" pageId="21" pageNumber="22">Nurettin Beşer</collectorName>
|
||
,
|
||
<collectorName id="26DF5370424BFE7BFE024804119EFE9E" box="[430,680,258,284]" pageId="21" pageNumber="22">Mehmet Kürşat Şahin</collectorName>
|
||
,
|
||
<date id="FF941066424BFE7BFD1848041003FE9E" box="[692,821,258,284]" pageId="21" pageNumber="22" value="2016-01-07">
|
||
<collectingDate id="EFD0E98E424BFE7BFD1848041003FE9E" box="[692,821,258,284]" pageId="21" pageNumber="22" value="2016-01-07">01.07.2016</collectingDate>
|
||
</date>
|
||
.
|
||
<specimenCode id="DB8C9EDD424BFE7BFCED48041080FE9F" box="[833,950,258,285]" collectionCode="ZDEU" country="Turkey" httpUri="http://grbio.org/cool/s077-eqrn" name="Zoology Department, Ege University" pageId="21" pageNumber="22">ZDEU119</specimenCode>
|
||
</materialsCitation>
|
||
/2015, 1 ♁♁,
|
||
<materialsCitation id="3B423CFB424BFE7BFBFD4804178AFEC2" ID-GBIF-Occurrence="4012805333" collectingDate="2015-07-26" collectionCode="ZDEU" collectorName="Kamil Candan & Elif Yildirim Caynak" country="Turkey" location="Balik Lake" municipality="Taslicay" pageId="21" pageNumber="22" specimenCode="ZDEU121" specimenCount="5" specimenCount-female="5" stateProvince="Agri" typeStatus="paratype">
|
||
<specimenCount id="9D2CFD2F424BFE7BFBFD480417A4FE9E" box="[1105,1170,258,284]" pageId="21" pageNumber="22" type="female">3 ♀♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D2CFD2F424BFE7BFB0C480417E9FE9E" box="[1184,1247,258,284]" pageId="21" pageNumber="22" type="female">2 ♀♀</specimenCount>
|
||
subadults,
|
||
<location id="8EF5607D424BFE7BFACC480413E4FEBD" LSID="urn:lsid:plazi:treatment:038387B0424CFE78FF3B4F561298F9D9:8EF5607D424BFE7BFACC480413E4FEBD" country="Turkey" municipality="Taslicay" name="Balik Lake" pageId="21" pageNumber="22" stateProvince="Agri">Balık Lake</location>
|
||
,
|
||
<collectingMunicipality id="6BF1ACDC424BFE7BFF714823120DFEBD" box="[221,315,293,319]" pageId="21" pageNumber="22">Taşlıçay</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49EEF844424BFE7BFEE84820124CFEC2" box="[324,378,294,320]" country="Turkey" name="Agri" pageId="21" pageNumber="22">Ağrı</collectingRegion>
|
||
,
|
||
<collectingCountry id="F33D7636424BFE7BFE29482312E1FEBD" box="[389,471,293,319]" name="Turkey" pageId="21" pageNumber="22">Turkey</collectingCountry>
|
||
. leg.
|
||
<collectorName id="26DF5370424BFE7BFDBD48231183FEBD" box="[529,693,293,319]" pageId="21" pageNumber="22">Kamil Candan</collectorName>
|
||
,
|
||
<collectorName id="26DF5370424BFE7BFD6D48231086FEBD" box="[705,944,293,319]" pageId="21" pageNumber="22">Elif Yıldırım Caynak</collectorName>
|
||
,
|
||
<date id="FF941066424BFE7BFC174823170BFEC2" box="[955,1085,293,320]" pageId="21" pageNumber="22" value="2015-07-26">
|
||
<collectingDate id="EFD0E98E424BFE7BFC174823170BFEC2" box="[955,1085,293,320]" pageId="21" pageNumber="22" value="2015-07-26">26.07.2015</collectingDate>
|
||
</date>
|
||
.
|
||
<specimenCode id="DB8C9EDD424BFE7BFBEB4823178AFEC2" box="[1095,1212,293,320]" collectionCode="ZDEU" country="Turkey" httpUri="http://grbio.org/cool/s077-eqrn" name="Zoology Department, Ege University" pageId="21" pageNumber="22">ZDEU121</specimenCode>
|
||
</materialsCitation>
|
||
/2015, 2 ♁♁,
|
||
<materialsCitation id="3B423CFB424BFE7BFAFA482017D5FEE6" ID-GBIF-Occurrence="4012805325" collectingDate="2015-07-25" collectorName="Kamil Candan & Elif Yildirim Caynak" country="Turkey" location="Guzeldere Village" municipality="Hinis" pageId="21" pageNumber="22" specimenCount="4" specimenCount-female="4" stateProvince="Erzurum" typeStatus="paratype">
|
||
<specimenCount id="9D2CFD2F424BFE7BFAFA482016A0FEC2" box="[1366,1430,294,320]" pageId="21" pageNumber="22" type="female">4 ♀♀</specimenCount>
|
||
,
|
||
<location id="8EF5607D424BFE7BFF3B484F1253FEE1" LSID="urn:lsid:plazi:treatment:038387B0424CFE78FF3B4F561298F9D9:8EF5607D424BFE7BFF3B484F1253FEE1" box="[151,357,329,355]" country="Turkey" municipality="Hinis" name="Guzeldere Village" pageId="21" pageNumber="22" stateProvince="Erzurum">Güzeldere Village</location>
|
||
,
|
||
<collectingMunicipality id="6BF1ACDC424BFE7BFEC3484C1298FEE6" box="[367,430,330,356]" pageId="21" pageNumber="22">Hınıs</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49EEF844424BFE7BFE16484C1129FEE6" box="[442,543,330,356]" country="Turkey" name="Erzurum" pageId="21" pageNumber="22">Erzurum</collectingRegion>
|
||
,
|
||
<collectingCountry id="F33D7636424BFE7BFD86484F114DFEE1" box="[554,635,329,355]" name="Turkey" pageId="21" pageNumber="22">Turkey</collectingCountry>
|
||
, leg.
|
||
<collectorName id="26DF5370424BFE7BFD1A484F106DFEE1" box="[694,859,329,355]" pageId="21" pageNumber="22">Kamil Candan</collectorName>
|
||
,
|
||
<collectorName id="26DF5370424BFE7BFCCA484F1760FEE1" box="[870,1110,329,355]" pageId="21" pageNumber="22">Elif Yıldırım Caynak</collectorName>
|
||
,
|
||
<date id="FF941066424BFE7BFBCD484F17D5FEE6" box="[1121,1251,329,356]" pageId="21" pageNumber="22" value="2015-07-25">
|
||
<collectingDate id="EFD0E98E424BFE7BFBCD484F17D5FEE6" box="[1121,1251,329,356]" pageId="21" pageNumber="22" value="2015-07-25">25.07.2015</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C330652D424BFE7BFF6B486B175EFC41" pageId="21" pageNumber="22" type="etymology">
|
||
<paragraph id="8B9536A6424BFE7BFF6B486B1093FE29" blockId="21.[151,1437,151,2036]" pageId="21" pageNumber="22">
|
||
<emphasis id="B95EEAB4424BFE7BFF6B486B1291FE05" bold="true" box="[199,423,365,391]" pageId="21" pageNumber="22">Derivatio nominis:</emphasis>
|
||
The specific epithet refers to Josef Friederich Schmidtler (born 1942), for his remarkable work on the knowledge of Turkish herpetofauna and its rich diversity.
|
||
</paragraph>
|
||
<paragraph id="8B9536A6424BFE7BFF6B48B2175EFC41" blockId="21.[151,1437,151,2036]" pageId="21" pageNumber="22">
|
||
<emphasis id="B95EEAB4424BFE7BFF6B48B211A6FE4C" bold="true" box="[199,656,436,462]" pageId="21" pageNumber="22">Comparative diagnosis (Morphology):</emphasis>
|
||
<taxonomicName id="4C2A4D25424BFE7BFD3B48B210F4FE4C" authority="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz, 2022" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" box="[663,962,436,462]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="josefschmidtleri" status="sp. nov.">
|
||
<emphasis id="B95EEAB4424BFE7BFD3B48B210F4FE4C" box="[663,962,436,462]" italics="true" pageId="21" pageNumber="22">Darevskia josefschmidtleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAB4424BFE7BFC6548B31714FE4D" bold="true" box="[969,1058,437,463]" pageId="21" pageNumber="22">
|
||
<taxonomicNameLabel id="A26D57CF424BFE7BFC6548B31714FE4D" box="[969,1058,437,463]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is a medium sized species (adults SVL:
|
||
<quantity id="4CD29B43424BFE7BFF7348DE1290FE70" box="[223,422,472,499]" metricMagnitude="-2" metricUnit="m" metricValue="6.06" metricValueMax="6.795" metricValueMin="5.325" pageId="21" pageNumber="22" unit="mm" value="60.6" valueMax="67.95" valueMin="53.25">53.25–67.95 mm</quantity>
|
||
). It is characterized by medium or small-sized scales with feeble keeling (even barely visible).
|
||
<taxonomicName id="4C2A4D25424BFE7BFF5648FA111EFD94" authority="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz, 2022" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" box="[250,552,508,534]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="josefschmidtleri" status="sp. nov.">
|
||
<emphasis id="B95EEAB4424BFE7BFF5648FA111EFD94" box="[250,552,508,534]" italics="true" pageId="21" pageNumber="22">Darevskia josefschmidtleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAB4424BFE7BFD9C48FA11BDFD94" bold="true" box="[560,651,508,534]" pageId="21" pageNumber="22">
|
||
<taxonomicNameLabel id="A26D57CF424BFE7BFD9C48FA11BDFD94" box="[560,651,508,534]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C2A4D25424BFE7BFC8E48FB109EFD94" baseAuthorityName="Boettger" baseAuthorityYear="1892" box="[802,936,508,534]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="valentini">
|
||
<emphasis id="B95EEAB4424BFE7BFC8E48FB109EFD94" box="[802,936,508,534]" italics="true" pageId="21" pageNumber="22">D. valentini</emphasis>
|
||
</taxonomicName>
|
||
in that there is a higher number of lamellae (46–59 vs. 42–53) and dorsals (47–58 vs. 41–52);
|
||
<taxonomicName id="4C2A4D25424BFE7BFD604B2610CFFDB8" authority="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz, 2022" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" box="[716,1017,544,570]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="josefschmidtleri" status="sp. nov.">
|
||
<emphasis id="B95EEAB4424BFE7BFD604B2610CFFDB8" box="[716,1017,544,570]" italics="true" pageId="21" pageNumber="22">Darevskia josefschmidtleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAB4424BFE7BFBAC4B26176FFDB8" bold="true" box="[1024,1113,544,570]" pageId="21" pageNumber="22">
|
||
<taxonomicNameLabel id="A26D57CF424BFE7BFBAC4B26176FFDB8" box="[1024,1113,544,570]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
males have a higher number of preanal than
|
||
<taxonomicName id="4C2A4D25424BFE7BFEE84B4212F1FDDF" baseAuthorityName="Boettger" baseAuthorityYear="1892" box="[324,455,579,605]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="valentini">
|
||
<emphasis id="B95EEAB4424BFE7BFEE84B4212F1FDDF" box="[324,455,579,605]" italics="true" pageId="21" pageNumber="22">D. valentini</emphasis>
|
||
</taxonomicName>
|
||
(1–3 vs. 1) while females have different collar scores (8–12 vs. 7–10) and gulars (23–31 vs. 21–29). It also differs from
|
||
<taxonomicName id="4C2A4D25424BFE7BFE564B6E11B6FD03" baseAuthorityName="Boettger" baseAuthorityYear="1892" box="[506,640,615,641]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="valentini">
|
||
<emphasis id="B95EEAB4424BFE7BFE564B6E11B6FD03" box="[506,640,615,641]" italics="true" pageId="21" pageNumber="22">D. valentini</emphasis>
|
||
</taxonomicName>
|
||
in having a greater head relative length (0.18–0.23 vs. 0.16.–0.21) for males.
|
||
<taxonomicName id="4C2A4D25424BFE7BFF4A4B8D1127FD27" authority="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz, 2022" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" box="[230,529,651,677]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="josefschmidtleri" status="sp. nov.">
|
||
<emphasis id="B95EEAB4424BFE7BFF4A4B8D1127FD27" box="[230,529,651,677]" italics="true" pageId="21" pageNumber="22">Darevskia josefschmidtleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAB4424BFE7BFDBB4B8D1146FD27" bold="true" box="[535,624,651,677]" pageId="21" pageNumber="22">
|
||
<taxonomicNameLabel id="A26D57CF424BFE7BFDBB4B8D1146FD27" box="[535,624,651,677]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<emphasis id="B95EEAB4424BFE7BFCAE4B8A17A9FD27" box="[770,1183,651,677]" italics="true" pageId="21" pageNumber="22">
|
||
<taxonomicName id="4C2A4D25424BFE7BFCAE4B8A17B8FD27" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" baseAuthorityName="Eiselt, Darevsky & Schmidtler" baseAuthorityYear="1992" box="[770,1166,651,677]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="subSpecies" species="spitzenbergerae" subSpecies="spitzenbergerae">D. spitzenbergerae spitzenbergerae</taxonomicName>
|
||
<emphasis id="B95EEAB4424BFE7BFB384B8D17A9FD27" bold="true" box="[1172,1183,651,677]" pageId="21" pageNumber="22">s</emphasis>
|
||
</emphasis>
|
||
tat. et comb. nov. (see below) in that there is higher number of tibials (16–24 vs. 15–19) and dorsals (48–58 vs. 44–53) for males. Also, it differs from
|
||
<taxonomicName id="4C2A4D25424BFE7BFE894BD51185FD6E" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" baseAuthorityName="Eiselt, Darevsky & Schmidtler" baseAuthorityYear="1992" box="[293,691,722,748]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="subSpecies" species="spitzenbergerae" subSpecies="spitzenbergerae">
|
||
<emphasis id="B95EEAB4424BFE7BFE894BD51185FD6E" box="[293,691,722,748]" italics="true" pageId="21" pageNumber="22">D. spitzenbergerae spitzenbergerae</emphasis>
|
||
</taxonomicName>
|
||
in having a shorter head relative length (0.18–0.23 vs 0.19–0.24).
|
||
<taxonomicName id="4C2A4D25424BFE7BFF3B4BF012F1FC92" authority="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz, 2022" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" box="[151,455,758,784]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="josefschmidtleri" status="sp. nov.">
|
||
<emphasis id="B95EEAB4424BFE7BFF3B4BF012F1FC92" box="[151,455,758,784]" italics="true" pageId="21" pageNumber="22">Darevskia josefschmidtleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAB4424BFE7BFE7E4BF01119FC92" bold="true" box="[466,559,758,784]" pageId="21" pageNumber="22">
|
||
<taxonomicNameLabel id="A26D57CF424BFE7BFE7E4BF01119FC92" box="[466,559,758,784]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from “Clade A” from
|
||
<bibRefCitation id="EFBB4B57424BFE7BFC3F4BF017B3FC92" author="Candan, K. & Kornilios, P. & Ayaz, D. & Kumlutas, Y. & Gul, S. & Yildirim-Caynak, E. & Ilgaz, C." box="[915,1157,758,784]" pageId="21" pageNumber="22" pagination="665 - 681" refId="ref25168" refString="Candan, K., Kornilios, P., Ayaz, D., Kumlutas, Y., Gul, S., Yildirim-Caynak, E. & Ilgaz, C. (2021) Cryptic genetic structure within Valentin's Lizard, Darevskia valentini (Boettger, 1892) (Squamata, Lacertidae), with implications for systematics and origins of parthenogenesis. Systematics and Biodiversity, 19 (7), 665 - 681. https: // doi. org / 10.1080 / 14772000.2021.1909171" type="journal article" year="2021">
|
||
Candan
|
||
<emphasis id="B95EEAB4424BFE7BFC584BF11706FC92" box="[1012,1072,758,784]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
|
||
(2021)
|
||
</bibRefCitation>
|
||
(described below as
|
||
<taxonomicName id="4C2A4D25424BFE7BFA2D4BF112DAFCB6" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="subSpecies" species="spitzenbergerae" subSpecies="wernermayeri">
|
||
<emphasis id="B95EEAB4424BFE7BFA2D4BF112DAFCB6" italics="true" pageId="21" pageNumber="22">D. spitzenbergerae wernermayeri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAB4424BFE7BFE5F4A1C1156FCB1" bold="true" box="[499,608,793,820]" pageId="21" pageNumber="22">ssp. nov.)</emphasis>
|
||
in that there is a lower number of ventrals (26–29 vs. 26–31), temp 2 (2–6 vs. 4–6) in males and preanals (1–3 vs. 2–3) in females.
|
||
<taxonomicName id="4C2A4D25424BFE7BFCAD4A3B171DFCD5" authority="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz, 2022" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" box="[769,1067,829,855]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="josefschmidtleri" status="sp. nov.">
|
||
<emphasis id="B95EEAB4424BFE7BFCAD4A3B171DFCD5" box="[769,1067,829,855]" italics="true" pageId="21" pageNumber="22">Darevskia josefschmidtleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAB4424BFE7BFB9D4A3817BFFCDA" bold="true" box="[1073,1161,830,856]" pageId="21" pageNumber="22">
|
||
<taxonomicNameLabel id="A26D57CF424BFE7BFB9D4A3817BFFCDA" box="[1073,1161,830,856]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C2A4D25424BFE7BFAB54A3816AAFCD5" baseAuthorityName="Boettger" baseAuthorityYear="1892" box="[1305,1436,829,855]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="valentini">
|
||
<emphasis id="B95EEAB4424BFE7BFAB54A3816AAFCD5" box="[1305,1436,829,855]" italics="true" pageId="21" pageNumber="22">D. valentini</emphasis>
|
||
</taxonomicName>
|
||
in that there is a lower number of maxillary (15–18 vs. 16–20) and dentary teeth (20–22 vs. 18–25).
|
||
<taxonomicName id="4C2A4D25424BFE7BFA864A67127CFC1D" authority="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz, 2022" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="josefschmidtleri" status="sp. nov.">
|
||
<emphasis id="B95EEAB4424BFE7BFA864A67127CFC1D" italics="true" pageId="21" pageNumber="22">Darevskia josefschmidtleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAB4424BFE7BFEFD4A83129DFC1D" bold="true" box="[337,427,901,927]" pageId="21" pageNumber="22">
|
||
<taxonomicNameLabel id="A26D57CF424BFE7BFEFD4A83129DFC1D" box="[337,427,901,927]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from Clade A (
|
||
<taxonomicName id="4C2A4D25424BFE7BFD064A801715FC1D" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" box="[682,1059,901,927]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="subSpecies" species="spitzenbergerae" subSpecies="wernermayeri">
|
||
<emphasis id="B95EEAB4424BFE7BFD064A801715FC1D" box="[682,1059,901,927]" italics="true" pageId="21" pageNumber="22">D. spitzenbergerae wernermayeri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAB4424BFE7BFB864A8317B9FC1D" bold="true" box="[1066,1167,901,927]" pageId="21" pageNumber="22">ssp. nov.</emphasis>
|
||
see below) in that there is a lower number of maxillary (15–18 vs. 19–21) and dentary teeth (20–22 vs. 23–24).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C330652D424BFE7BFF6B4ACA11EAFB1B" pageId="21" pageNumber="22" type="diagnosis">
|
||
<paragraph id="8B9536A6424BFE7BFF6B4ACA11EAFB1B" blockId="21.[151,1437,151,2036]" pageId="21" pageNumber="22">
|
||
<emphasis id="B95EEAB4424BFE7BFF6B4ACA12D7FC64" bold="true" box="[199,481,972,998]" pageId="21" pageNumber="22">Diagnosis (Molecular):</emphasis>
|
||
It can be distinguished from other former
|
||
<taxonomicName id="4C2A4D25424BFE7BFC514ACB17B7FC64" baseAuthorityName="Boettger" baseAuthorityYear="1892" box="[1021,1153,972,998]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="valentini">
|
||
<emphasis id="B95EEAB4424BFE7BFC514ACB17B7FC64" box="[1021,1153,972,998]" italics="true" pageId="21" pageNumber="22">D. valentini</emphasis>
|
||
</taxonomicName>
|
||
populations by unique nucleotide combination located on the mitochondrial gene
|
||
<emphasis id="B95EEAB4424BFE7BFC9C4AF61058FB88" box="[816,878,1008,1034]" italics="true" pageId="21" pageNumber="22">Cyt-b</emphasis>
|
||
and nuclear loci
|
||
<emphasis id="B95EEAB4424BFE7BFB9E4AF6174DFB88" box="[1074,1147,1008,1034]" italics="true" pageId="21" pageNumber="22">MC1R</emphasis>
|
||
. The consensus sequence (
|
||
<emphasis id="B95EEAB4424BFE7BFF0D4D1213E9FBAC" box="[161,223,1044,1070]" italics="true" pageId="21" pageNumber="22">Cyt-b</emphasis>
|
||
) for
|
||
<taxonomicName id="4C2A4D25424BFE7BFEB04D12117AFBAC" authority="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz, 2022" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" box="[284,588,1044,1070]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="josefschmidtleri" status="sp. nov.">
|
||
<emphasis id="B95EEAB4424BFE7BFEB04D12117AFBAC" box="[284,588,1044,1070]" italics="true" pageId="21" pageNumber="22">Darevskia josefschmidtleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAB4424BFE7BFDF44D121181FBAC" bold="true" box="[600,695,1044,1070]" pageId="21" pageNumber="22">
|
||
<taxonomicNameLabel id="A26D57CF424BFE7BFDF44D121181FBAC" box="[600,695,1044,1070]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is found in Appendix 6, together with the respective sequence for the
|
||
<taxonomicName id="4C2A4D25424BFE7BFF464D3E1259FBD0" baseAuthorityName="Boettger" baseAuthorityYear="1892" box="[234,367,1080,1106]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" sensu="stricto" species="valentini">
|
||
<emphasis id="B95EEAB4424BFE7BFF464D3E1259FBD0" box="[234,367,1080,1106]" italics="true" pageId="21" pageNumber="22">D. valentini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26D57CF424BFE7BFED44D3E1283FBD0" box="[376,437,1080,1106]" pageId="21" pageNumber="22" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
In this Table, the thirty simple nucleotide diagnostic characters between the consensus sequences are highlighted. Similarly, the molecular diagnostic characters for
|
||
<taxonomicName id="4C2A4D25424BFE7BFBA54D5D1601FBF7" authority="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz, 2022" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" box="[1033,1335,1115,1141]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="josefschmidtleri" status="sp. nov.">
|
||
<emphasis id="B95EEAB4424BFE7BFBA54D5D1601FBF7" box="[1033,1335,1115,1141]" italics="true" pageId="21" pageNumber="22">Darevskia josefschmidtleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAB4424BFE7BFAEC4D5A16AAFBF4" bold="true" box="[1344,1436,1116,1142]" pageId="21" pageNumber="22">
|
||
<taxonomicNameLabel id="A26D57CF424BFE7BFAEC4D5A16AAFBF4" box="[1344,1436,1116,1142]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
regarding the nuclear loci are shown in Appendix 6.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C330652D424BFE78FF6B4DA5122AFEE7" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="description">
|
||
<paragraph id="8B9536A6424BFE7BFF6B4DA513C7FB63" blockId="21.[151,1437,151,2036]" pageId="21" pageNumber="22">
|
||
<emphasis id="B95EEAB4424BFE7BFF6B4DA512D3FB3F" bold="true" box="[199,485,1187,1213]" pageId="21" pageNumber="22">
|
||
Description of
|
||
<typeStatus id="54918804424BFE7BFED94DA512D6FB3F" box="[373,480,1187,1213]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
An adult male. Tail in regeneration process. Fixed with ethanol and conserved in 96% ethanol.
|
||
</paragraph>
|
||
<paragraph id="8B9536A6424BFE7BFF6B4DEC16A4F9C5" blockId="21.[151,1437,151,2036]" pageId="21" pageNumber="22">
|
||
<emphasis id="B95EEAB4424BFE7BFF6B4DEC120CFA86" box="[199,314,1258,1284]" italics="true" pageId="21" pageNumber="22">Scalation:</emphasis>
|
||
Rostral not in contact with the nostril. Single postnasal on each side. Width of frontonasal (internasal) plate subequal to length, not contact with rostral. Sutures between prefrontal plates and frontal plate straight. Parietal plates in contact with postorbital plates on each side. Supraciliar granules 9 on each side, interrupted series on right, not on left side. Supraciliar plates 6 on each side. Supralabial and sublabial plates 4 and 6 on each side, respectively. Plates in supratemporal region 3 on left, 4 on right. The first supratemporal plate large, narrow towards the back, ends bluntly. Masseteric large, in one piece on each side, separated from the first supratemporal plate by a row of small scales. Tympanic obvious, in two pieces on the left and one piece on right, separated from masseteric by three and two rows of small scales on the left and right, respectively. Eight flat and smooth collaria. Gularia 27. Ventralia contains 6 longitudinal and 28 transverse rows of plates. Preanal scale in two pieces, surrounded by 6 rows of plates. Femoral pores 21 on left, 20 on right side. Subdigital lamellae 27 on left, 26 on right. Tibial scale 19. Dorsalia 53.
|
||
</paragraph>
|
||
<paragraph id="8B9536A6424BFE7BFF6B4F5616AFF90C" blockId="21.[151,1437,151,2036]" pageId="21" pageNumber="22">
|
||
<emphasis id="B95EEAB4424BFE7BFF6B4F561203F9EB" box="[199,309,1616,1641]" italics="true" pageId="21" pageNumber="22">Biometry:</emphasis>
|
||
SVL
|
||
<quantity id="4CD29B43424BFE7BFED64F5612C4F9E8" box="[378,498,1616,1642]" metricMagnitude="-2" metricUnit="m" metricValue="6.468" pageId="21" pageNumber="22" unit="mm" value="64.68">64.68 mm</quantity>
|
||
., pileus width
|
||
<quantity id="4CD29B43424BFE7BFD394F5611CDF9E8" box="[661,763,1616,1642]" metricMagnitude="-3" metricUnit="m" metricValue="6.79" pageId="21" pageNumber="22" unit="mm" value="6.79">6.79 mm</quantity>
|
||
, pileus length
|
||
<quantity id="4CD29B43424BFE7BFC0D4F561723F9E8" box="[929,1045,1616,1642]" metricMagnitude="-2" metricUnit="m" metricValue="1.327" pageId="21" pageNumber="22" unit="mm" value="13.27">13.27 mm</quantity>
|
||
, head width
|
||
<quantity id="4CD29B43424BFE7BFB0B4F56163BF9E8" box="[1191,1293,1616,1642]" metricMagnitude="-3" metricUnit="m" metricValue="8.13" pageId="21" pageNumber="22" unit="mm" value="8.13">8.13 mm</quantity>
|
||
, head length
|
||
<quantity id="4CD29B43424BFE7BFF3B4F72123CF90C" box="[151,266,1652,1678]" metricMagnitude="-2" metricUnit="m" metricValue="1.314" pageId="21" pageNumber="22" unit="mm" value="13.14">13.14 mm</quantity>
|
||
. Length of forelimb
|
||
<quantity id="4CD29B43424BFE7BFE404F72116BF90C" box="[492,605,1652,1678]" metricMagnitude="-2" metricUnit="m" metricValue="1.839" pageId="21" pageNumber="22" unit="mm" value="18.39">18.39 mm</quantity>
|
||
, length of hindlimb
|
||
<quantity id="4CD29B43424BFE7BFC904F721099F90C" box="[828,943,1652,1678]" metricMagnitude="-2" metricUnit="m" metricValue="2.79" pageId="21" pageNumber="22" unit="mm" value="27.9">27.90 mm</quantity>
|
||
. Anal plate width
|
||
<quantity id="4CD29B43424BFE7BFBD94F7517EFF90C" box="[1141,1241,1651,1678]" metricMagnitude="-3" metricUnit="m" metricValue="4.69" pageId="21" pageNumber="22" unit="mm" value="4.69">4.69 mm</quantity>
|
||
, length
|
||
<quantity id="4CD29B43424BFE7BFA9C4F7216A3F90C" box="[1328,1429,1652,1678]" metricMagnitude="-3" metricUnit="m" metricValue="1.71" pageId="21" pageNumber="22" unit="mm" value="1.71">1.71 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9536A6424BFE7BFF6B4F9111ABF829" blockId="21.[151,1437,151,2036]" pageId="21" pageNumber="22">
|
||
<emphasis id="B95EEAB4424BFE7BFF6B4F911151F933" box="[199,615,1687,1714]" italics="true" pageId="21" pageNumber="22">Coloration and pattern (in alcohol):</emphasis>
|
||
The ground color of the dorsum is greenish brown. Dorsal tract with a wide vertebral band composed of fairly irregular transverse spots nost covering almost its complete width. Similar dark spotting is present on each side of the body (temporal or lateral band) that appears reticulated. Between these two dark spots, located both in the middle of the dorsum and flanks, a paler double line extends from the nape to the base of the tail (Appendix 5). There are few pale spots inside the reticulate on dark bands on flanks, with more in the forelimb. A few light bluish spots near the forelimb basis. Belly, along with head and neck, whitish (see photos in Appendix 5), with dark and blue spots in the outermost rows of ventrals. Background color of head plates brownish, with few scattered and small black spots on it.
|
||
</paragraph>
|
||
<paragraph id="8B9536A6424BFE78FF6B4EB3122AFEE7" blockId="21.[151,1437,151,2036]" lastBlockId="22.[151,1437,151,357]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
|
||
<emphasis id="B95EEAB4424BFE7BFF6B4EB312D2F852" bold="true" box="[199,484,1973,2000]" pageId="21" pageNumber="22">
|
||
Variations of
|
||
<typeStatus id="54918804424BFE7BFEC84EB012E9F852" box="[356,479,1974,2000]" pageId="21" pageNumber="22" type="paratype">paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Descriptive statistics and variation range of the morphometric and scalation characters are given in Table 7. Frontonasal (internasal) rarely is in contact with rostral. Sutures between the prefrontal and frontal are usually slanted. Parietal is rarely in contact with postorbital. Masseteric is sometimes divided into two pieces. Tight scales are feeble-keeled, smaller than dorsal ones. In
|
||
<specimenCount id="9D2CFD2F4248FE78FC7B49BD1740FF57" box="[983,1142,187,213]" pageId="22" pageNumber="23" type="generic">one specimen</specimenCount>
|
||
, transverse dark spots on ground color combined with spots on each side, do not form two separate rows. In addition, transverse dark spots, longitudinally above ground color, are faint in
|
||
<specimenCount id="9D2CFD2F4248FE78FD3348051076FE9F" box="[671,832,259,285]" pageId="22" pageNumber="23" type="generic">ten specimens</specimenCount>
|
||
. In four samples, pale spots on the forelimb base were not blue (white). Belly, along with head and neck, yellowish in
|
||
<specimenCount id="9D2CFD2F4248FE78FCF64821173FFEC3" box="[858,1033,295,321]" pageId="22" pageNumber="23" type="generic">nine specimens</specimenCount>
|
||
. In fourteen samples, no spotting on head plates.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF55662E4248FE78FF344ABD1197FB4D" ID-DOI="http://doi.org/10.5281/zenodo.7517939" ID-Zenodo-Dep="7517939" httpUri="https://zenodo.org/record/7517939/files/figure.png" pageId="22" pageNumber="23" startId="22.[152,255,955,979]" targetBox="[151,1413,392,928]" targetPageId="22">
|
||
<paragraph id="8B9536A64248FE78FF344ABD1197FB4D" blockId="22.[151,1437,955,1231]" pageId="22" pageNumber="23">
|
||
<emphasis id="B95EEAB44248FE78FF344ABD121FFC51" bold="true" box="[152,297,955,979]" pageId="22" pageNumber="23">FIGURE 12.</emphasis>
|
||
a)
|
||
<taxonomicName id="4C2A4D254248FE78FEFE4ABD11CBFC51" authority="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz, 2022" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" box="[338,765,955,979]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="subSpecies" species="spitzenbergerae" status="subsp. nov." subSpecies="wernermayeri">
|
||
<emphasis id="B95EEAB44248FE78FEFE4ABD11CBFC51" box="[338,765,955,979]" italics="true" pageId="22" pageNumber="23">Darevskia spitzenbergerae wernermayeri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAB44248FE78FCA44ABD1050FC51" bold="true" box="[776,870,955,979]" pageId="22" pageNumber="23">
|
||
<taxonomicNameLabel id="A26D57CF4248FE78FCA44ABD1050FC51" box="[776,870,955,979]" pageId="22" pageNumber="23" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(Paratype; nº 12, Male; Başeğmez Village, Çaldıran, Turkey); b)
|
||
<taxonomicName id="4C2A4D254248FE78FEBE4AD912EAFC75" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" baseAuthorityName="Arribas, Ilgaz, Kumlutas, Durmus, Avci & Uzum" baseAuthorityYear="2013" box="[274,476,991,1015]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="mirabilis" status="stat. nov.">
|
||
<emphasis id="B95EEAB44248FE78FEBE4AD912EAFC75" box="[274,476,991,1015]" italics="true" pageId="22" pageNumber="23">Darevskia mirabilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAB44248FE78FE4F4AD91175FC75" bold="true" box="[483,579,991,1015]" pageId="22" pageNumber="23">
|
||
<taxonomicNameLabel id="A26D57CF4248FE78FE4F4AD91175FC75" box="[483,579,991,1015]" pageId="22" pageNumber="23">stat. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(Paratype; nº 5, Female; Ovit Pass, Kaçkar Mountains, Rize, Turkey); c)
|
||
<taxonomicName id="4C2A4D254248FE78FA994AD91252FB99" authorityName="Arribas, Ilgaz, Kumlutas, Durmus, Avci & Uzum" authorityYear="2013" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="subSpecies" species="rudis" subSpecies="bolkardaghica">
|
||
<emphasis id="B95EEAB44248FE78FA994AD91252FB99" italics="true" pageId="22" pageNumber="23">Darevskia rudis bolkardaghica</emphasis>
|
||
</taxonomicName>
|
||
(Paratype; nº 1, Male; Karagöl, Ulukışla, Niğde, Central Anatolia, Turkey); d)
|
||
<taxonomicName id="4C2A4D254248FE78FBD34D0516ABFB99" baseAuthorityName="Darevsky & Eiselt" baseAuthorityYear="1967" box="[1151,1437,1027,1051]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="subSpecies" species="rudis" subSpecies="lantzicyreni">
|
||
<emphasis id="B95EEAB44248FE78FBD34D0516ABFB99" box="[1151,1437,1027,1051]" italics="true" pageId="22" pageNumber="23">Darevskia rudis lantzicyreni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAB44248FE78FF3B4D211239FBBD" bold="true" box="[151,271,1063,1087]" pageId="22" pageNumber="23">comb. nov.</emphasis>
|
||
(nº 23, male; Kümbet Village, Zara, Turkey); e)
|
||
<taxonomicName id="4C2A4D254248FE78FC8A4D21170CFBBD" authority="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz, 2022" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" box="[806,1082,1063,1087]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="josefschmidtleri" status="sp. nov.">
|
||
<emphasis id="B95EEAB44248FE78FC8A4D21170CFBBD" box="[806,1082,1063,1087]" italics="true" pageId="22" pageNumber="23">Darevskia josefschmidtleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAB44248FE78FBE84D2117AFFBBD" bold="true" box="[1092,1177,1063,1087]" pageId="22" pageNumber="23">
|
||
<taxonomicNameLabel id="A26D57CF4248FE78FBE84D2117AFFBBD" box="[1092,1177,1063,1087]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(Paratype; nº 20, Male; Yukarınarlıca Village, Çatak, Van, Turkey); f)
|
||
<taxonomicName id="4C2A4D254248FE78FDC04D4D1005FBE1" baseAuthorityName="Boettger" baseAuthorityYear="1892" box="[620,819,1099,1123]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="valentini">
|
||
<emphasis id="B95EEAB44248FE78FDC04D4D1005FBE1" box="[620,819,1099,1123]" italics="true" pageId="22" pageNumber="23">Darevskia valentini</emphasis>
|
||
</taxonomicName>
|
||
(nº 9, Male; Tepeler Village, Ardahan, Turkey) and temporal area of an Armenian specimen (Karvansaray, Martuni District, Armenia); g)
|
||
<taxonomicName id="4C2A4D254248FE78FC364D691679FB05" authorityName="Arribas & Candan & Kornilios & Ayaz & Kumlutaş & Gül & Yilmaz & Caynak & Ilgaz" authorityYear="2022" baseAuthorityName="Eiselt, Darevsky & Schmidtler" baseAuthorityYear="1992" box="[922,1359,1135,1159]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="subSpecies" species="spitzenbergerae" subSpecies="spitzenbergerae">
|
||
<emphasis id="B95EEAB44248FE78FC364D691679FB05" box="[922,1359,1135,1159]" italics="true" pageId="22" pageNumber="23">Darevskia spitzenbergerae spitzenbergerae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EEAB44248FE78FAF94D69123AFB29" bold="true" pageId="22" pageNumber="23">stat. et comb. nov.</emphasis>
|
||
(nº 1, Male; Cilo Sat Mountains, Hakkari, Turkey)- Also, temporal area of other specimen from the same locality. The new nomenclature proposed in the text is used.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C330652D4248FE78FF6B4DFB1104FA72" pageId="22" pageNumber="23" type="distribution">
|
||
<paragraph id="8B9536A64248FE78FF6B4DFB10F2FADD" blockId="22.[151,1436,1277,1627]" pageId="22" pageNumber="23">
|
||
<emphasis id="B95EEAB44248FE78FF6B4DFB1255FA95" bold="true" box="[199,355,1277,1303]" pageId="22" pageNumber="23">Distribution:</emphasis>
|
||
Confirmed localities draw an area around Çaldıran (
|
||
<collectingRegion id="49EEF8444248FE78FC034DFB10EBFA95" box="[943,989,1277,1303]" country="Turkey" name="Van" pageId="22" pageNumber="23">Van</collectingRegion>
|
||
) located in the east of
|
||
<collectingRegion id="49EEF8444248FE78FB7B4DF81635FA9A" box="[1239,1283,1278,1304]" country="Turkey" name="Van" pageId="22" pageNumber="23">Van</collectingRegion>
|
||
Lake, around Balık Lake (
|
||
<collectingRegion id="49EEF8444248FE78FE8E4C27126CFAB9" box="[290,346,1313,1339]" country="Turkey" name="Agri" pageId="22" pageNumber="23">Ağrı</collectingRegion>
|
||
), Palandöken Mountain and around Çat and Hınıs (
|
||
<collectingRegion id="49EEF8444248FE78FC364C271736FAB9" box="[922,1024,1313,1339]" country="Turkey" name="Erzurum" pageId="22" pageNumber="23">Erzurum</collectingRegion>
|
||
) which is in the south of this massif, all in
|
||
<collectingCountry id="F33D76364248FE78FF7A4C43121EFADD" box="[214,296,1349,1375]" name="Turkey" pageId="22" pageNumber="23">Turkey</collectingCountry>
|
||
. Probably also in intermediate areas among these localities.
|
||
</paragraph>
|
||
<paragraph id="8B9536A64248FE78FF6B4C6F1104FA72" blockId="22.[151,1436,1277,1627]" pageId="22" pageNumber="23">
|
||
<emphasis id="B95EEAB44248FE78FF6B4C6F121AFA01" bold="true" box="[199,300,1385,1411]" pageId="22" pageNumber="23">Habitat:</emphasis>
|
||
Subalpine-like vegetation from Irano-Turanian Region, in rocky and stony areas near
|
||
<quantity id="4CD29B434248FE78FA8D4C6F164CFA06" box="[1313,1402,1385,1412]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="22" pageNumber="23" unit="m" value="2000.0">2000 m</quantity>
|
||
or higher:
|
||
<quantity id="4CD29B434248FE78FF404C8B1275FA2A" box="[236,323,1421,1448]" metricMagnitude="3" metricUnit="m" metricValue="2.0949999999999998" pageId="22" pageNumber="23" unit="m" value="2095.0">2095 m</quantity>
|
||
. (Çaldıran),
|
||
<quantity id="4CD29B434248FE78FE604C8B1115FA2A" box="[460,547,1421,1448]" metricMagnitude="3" metricUnit="m" metricValue="2.27" pageId="22" pageNumber="23" unit="m" value="2270.0">2270 m</quantity>
|
||
. (Balık Lake),
|
||
<quantity id="4CD29B434248FE78FD6A4C8B102AFA2A" box="[710,796,1421,1448]" metricMagnitude="3" metricUnit="m" metricValue="2.429" pageId="22" pageNumber="23" unit="m" value="2429.0">2429 m</quantity>
|
||
. (Palandöken),
|
||
<quantity id="4CD29B434248FE78FC644C8B1728FA2A" box="[968,1054,1421,1448]" metricMagnitude="3" metricUnit="m" metricValue="1.946" pageId="22" pageNumber="23" unit="m" value="1946.0">1946 m</quantity>
|
||
. (Çat),
|
||
<quantity id="4CD29B434248FE78FBC24C8B17F3FA2A" box="[1134,1221,1421,1448]" metricMagnitude="3" metricUnit="m" metricValue="2.643" pageId="22" pageNumber="23" unit="m" value="2643.0">2643 m</quantity>
|
||
. (Hınıs).
|
||
<taxonomicName id="4C2A4D254248FE78FA864C8B1223FA49" baseAuthorityName="Darevsky" baseAuthorityYear="1966" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="unisexualis">
|
||
<emphasis id="B95EEAB44248FE78FA864C8B1223FA49" italics="true" pageId="22" pageNumber="23">Darevskia unisexualis</emphasis>
|
||
</taxonomicName>
|
||
was found sympatric in the Palandöken Mountain, while no other reptile species were detected in the same and other localities of the area.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C330652D4248FE78FF6B4CFF1298F9D9" pageId="22" pageNumber="23" type="discussion">
|
||
<paragraph id="8B9536A64248FE78FF6B4CFF1298F9D9" blockId="22.[151,1436,1277,1627]" pageId="22" pageNumber="23">
|
||
<emphasis id="B95EEAB44248FE78FF6B4CFF1264F991" bold="true" box="[199,338,1529,1555]" pageId="22" pageNumber="23">Comments:</emphasis>
|
||
This new taxon seems to be the parental species that gave origin by hybridization to the parthenogenetic
|
||
<taxonomicName id="4C2A4D254248FE78FEFA4F1812CCF9B5" baseAuthorityName="Darevsky" baseAuthorityYear="1966" box="[342,506,1565,1591]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="unisexualis">
|
||
<emphasis id="B95EEAB44248FE78FEFA4F1812CCF9B5" box="[342,506,1565,1591]" italics="true" pageId="22" pageNumber="23">D. unisexualis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2A4D254248FE78FDA14F18119BF9B5" baseAuthorityName="Schmidtler, Eiselt & Darevsky" baseAuthorityYear="1994" box="[525,685,1565,1591]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="sapphirina">
|
||
<emphasis id="B95EEAB44248FE78FDA14F18119BF9B5" box="[525,685,1565,1591]" italics="true" pageId="22" pageNumber="23">D. sapphirina</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2A4D254248FE78FD414F1810F4F9B5" baseAuthorityName="Schmidtler, Eiselt & Darevsky" baseAuthorityYear="1994" box="[749,962,1565,1591]" class="Reptilia" family="Lacertidae" genus="Darevskia" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="bendimahiensis">
|
||
<emphasis id="B95EEAB44248FE78FD414F1810F4F9B5" box="[749,962,1565,1591]" italics="true" pageId="22" pageNumber="23">D. bendimahiensis</emphasis>
|
||
</taxonomicName>
|
||
according to Z-chromosome inheritance (
|
||
<bibRefCitation id="EFBB4B574248FE78FF324F471294F9D9" author="Yanchukov, A. & Tarkhnishvili, D. & Erdolu, M. & Sahin, M. K. & Candan, K. & Murtskhvaladze, M. & Gabelaia, M. & Iankoshvili, G. & Barateli, N. & Ilgaz, C. & Kumlutas, Y. & Matur, F. & Colak, F. & Arakelyan, M. & Galoyan, E." box="[158,418,1601,1627]" pageId="22" pageNumber="23" pagination="293 - 305" refId="ref29305" refString="Yanchukov, A., Tarkhnishvili, D., Erdolu, M., Sahin, M. K., Candan, K., Murtskhvaladze, M., Gabelaia, M., Iankoshvili, G., Barateli, N., Ilgaz, C., Kumlutas, Y., Matur, F., Colak, F., Arakelyan, M. & Galoyan, E. (2022) Precise paternal ancestry of hybrid unisexual ZW lizards (genus Darevskia: Lacertidae: Squamata) revealed by Z-linked genomic markers. Biological Journal of the Linnean Society, 136 (2), 293 - 305. https: // doi. org / 10.1093 / biolinnean / blac 023" type="journal article" year="2022">
|
||
Yanchukov
|
||
<emphasis id="B95EEAB44248FE78FE8A4F441268F9D9" box="[294,350,1601,1627]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
|
||
2022
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |