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<document id="2D02D4DB5CEB9B67696CF66973FE4FD3" ID-CLB-Dataset="266155" ID-DOI="10.5281/zenodo.6533434" ID-GBIF-Dataset="0f129a32-9d98-4cbf-81fa-981c04fe43d5" ID-ISSN="1942-1354" ID-Zenodo-Dep="6533434" ID-ZooBank="AE18CA26-20E8-48D3-ABD0-22A0D9891065" IM.bibliography_approvedBy="admin" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="admin" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="admin" checkinTime="1652124454819" checkinUser="felipe" docAuthor="Brown, John W., Razowski, Józef, Aarvik, Leif, Timm, Alicia E. &amp; Copeland, Robert S." docDate="2022" docId="038387ADFFBEDF28B3B17F33FC34FCC2" docLanguage="en" docName="InsectaMundi.2022.0927.1-27.pdf" docOrigin="Insecta Mundi 2022 (927)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Eucosmocydia nigeriana Brown and Razowski 2022, new species" docType="treatment" docVersion="7" lastPageNumber="19" masterDocId="FFBAFFD5FFACDF3CB3717B54FFD2FFEB" masterDocTitle="New species and new combinations in Afrotropical Eucosmocydia Diakonoff, 1988 (Lepidoptera: Tortricidae: Olethreutinae)" masterLastPageNumber="27" masterPageNumber="1" pageNumber="17" updateTime="1699327336214" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="1D8E0A7E479B07F85F11F55DA5313716">New species and new combinations in Afrotropical Eucosmocydia Diakonoff, 1988 (Lepidoptera: Tortricidae: Olethreutinae)</mods:title>
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<mods:namePart id="AB363BAA87C147E9D4AE02DE72EFB82B">Brown, John W.</mods:namePart>
<mods:affiliation id="06BE1A0E6A9888CA12A827538D2CC960">Department of Entomology, National Museum of Natural History Washington, DC 20013 - 7012, USA</mods:affiliation>
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<mods:namePart id="D0F2523549C74E8468D482F207B212B8">Razowski, Józef</mods:namePart>
<mods:affiliation id="E0B2819FDA02C14C2455A7559F9DC661">Polish Academy of Sciences, Institute of Systematic Zoology Sławkowska 17, Krakow, Poland</mods:affiliation>
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<mods:namePart id="1849ACAFC62B96D88B5FD1E4FACF1735">Aarvik, Leif</mods:namePart>
<mods:affiliation id="93E38D551EDECE44453C6DA7B7A9EEB9">Natural History Museum, University of Oslo P. O. Box 1172, Blindern, NO- 0318 Oslo, Norway</mods:affiliation>
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<mods:namePart id="738653F4EC148A4275D8A1E26A0118B1">Timm, Alicia E.</mods:namePart>
<mods:affiliation id="5AE635BABFA6CB30E646CAA2CAB23F69">Department of Agricultural Biology, Colorado State University Fort Collins, CO 80521 - 1177, USA</mods:affiliation>
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<mods:namePart id="DF10CA9C826B360D87D1ACA840A64879">Copeland, Robert S.</mods:namePart>
<mods:affiliation id="6FF5C56AB4B4DC99E24CAE66611FB9DD">International Centre of Insect Physiology and Ecology P. O. Box 30772, Nairobi, Kenya</mods:affiliation>
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<treatment id="038387ADFFBEDF28B3B17F33FC34FCC2" ID-DOI="http://doi.org/10.5281/zenodo.6533460" ID-GBIF-Taxon="212971220" ID-Zenodo-Dep="6533460" LSID="urn:lsid:plazi:treatment:038387ADFFBEDF28B3B17F33FC34FCC2" httpUri="http://treatment.plazi.org/id/038387ADFFBEDF28B3B17F33FC34FCC2" lastPageId="20" lastPageNumber="19" pageId="18" pageNumber="17">
<subSubSection id="C3306530FFBEDF2EB3B17F33FC7EFB6F" box="[192,940,1127,1156]" pageId="18" pageNumber="17" type="nomenclature">
<paragraph id="8B9536BBFFBEDF2EB3B17F33FC7EFB6F" blockId="18.[192,940,1127,1189]" box="[192,940,1127,1156]" pageId="18" pageNumber="17">
<heading id="D0DD81D7FFBEDF2EB3B17F33FC7EFB6F" box="[192,940,1127,1156]" fontSize="12" level="2" pageId="18" pageNumber="17" reason="2">
<emphasis id="B95EEAA9FFBEDF2EB3B17F33FC7EFB6F" bold="true" box="[192,940,1127,1156]" pageId="18" pageNumber="17">
<taxonomicName id="4C2A4D38FFBEDF2EB3B17F33FCD9FB6F" authority="Brown and Razowski" authorityName="Brown and Razowski" authorityYear="2022" box="[192,779,1127,1156]" class="Insecta" family="Tortricidae" genus="Eucosmocydia" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="nigeriana" status="sp. nov.">
<emphasis id="B95EEAA9FFBEDF2EB3B17F33FE26FB68" bold="true" box="[192,500,1127,1156]" italics="true" pageId="18" pageNumber="17">Eucosmocydia nigeriana</emphasis>
Brown and Razowski
</taxonomicName>
, 
<taxonomicNameLabel id="A26D57D2FFBEDF2EB0667F33FC7EFB6F" box="[791,940,1127,1156]" pageId="18" pageNumber="17" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3306530FFBEDF2EB3B17FDEFE5DFB4E" box="[192,399,1162,1189]" pageId="18" pageNumber="17" type="description">
<paragraph id="8B9536BBFFBEDF2EB3B17FDEFE5DFB4E" blockId="18.[192,940,1127,1189]" box="[192,399,1162,1189]" pageId="18" pageNumber="17">
<figureCitation id="13112A3EFFBEDF2EB3B17FDEFECBFB4E" box="[192,281,1162,1189]" captionStart="Figures 2–9" captionStartId="10.[192,279,1537,1563]" captionTargetBox="[192,1439,213,1507]" captionTargetId="figure-158@10.[192,1439,213,1510]" captionTargetPageId="10" captionText="Figures 2–9. Adults of Eucosmocydia. 2) E. pappeana, male holotype. 3) E. pappeana, female paratype. 4) E. deinbolliana, male paratype. 5) E. mixographa, female. 6) E. lecaniodiscana, male holotype. 7) E. lecaniodiscana, female paratype. 8) E. nigeriana, male holotype. 9) E. nigeriana, paratype female." figureDoi="http://doi.org/10.5281/zenodo.6533438" httpUri="https://zenodo.org/record/6533438/files/figure.png" pageId="18" pageNumber="17">Fig. 8, 9</figureCitation>
, 
<figureCitation id="13112A3EFFBEDF2EB2547FDFFE90FB4E" box="[293,322,1163,1189]" captionStart="Figures 20–27" captionStartId="15.[192,279,1799,1825]" captionTargetBox="[207,1425,213,1770]" captionTargetId="figure-12@15.[207,1425,213,1773]" captionTargetPageId="15" captionText="Figures 20–27. Male genitalia of Eucosmocydia. 20) E. pappeana, USNM slide 153,769. 21) E. deinbolliana, USNM slide 153,700. 22) E. ugandensis, slide NHMO 3762. 23) E. lecaniodiscana, USNM slide 124,465. 24) Male genitalia of E. nigeriana, USNM slide 153,740.25) Male genitalia of E. pancoviana, USNM slide 153,695.26) Male genitalia of E. chlorobathra, USNM slide 153,691. 27) Male genitalia of E. kirimiriana, USNM slide 143,441." figureDoi="http://doi.org/10.5281/zenodo.6533442" httpUri="https://zenodo.org/record/6533442/files/figure.png" pageId="18" pageNumber="17">24</figureCitation>
, 
<figureCitation id="13112A3EFFBEDF2EB23D7FDFFEBBFB4E" box="[332,361,1163,1189]" captionStart="Figures 28–35" captionStartId="17.[192,279,1799,1825]" captionTargetBox="[284,1348,216,1770]" captionTargetId="figure-82@17.[284,1351,216,1770]" captionTargetPageId="17" captionText="Figures 28–35. Abdominal segment 7 of males of Eucosmocydia. 28) E. pappeana, USNM slide 143,426. 29) E. ugandensis, NHMO slide 3762. 30) E. deinbolliana, USNM slide 153,700. 31) E. lecaniodiscana, USNM slide 153,704. 32) E. nigeriana, USNM slide 153,740. 33) E. pancoviana, USNM slide 153,695. 34) E. chlorobathra, USNM slide 153,691. 35) E. kirimiriana, USNM slide 143,441." figureDoi="http://doi.org/10.5281/zenodo.6533444" httpUri="https://zenodo.org/record/6533444/files/figure.png" pageId="18" pageNumber="17">32</figureCitation>
, 
<figureCitation id="13112A3EFFBEDF2EB2057FDFFE5DFB4E" box="[372,399,1163,1189]" captionStart="Figure 36–41" captionStartId="19.[192,268,1673,1699]" captionTargetBox="[192,1438,213,1644]" captionTargetId="figure-12@19.[192,1438,213,1647]" captionTargetPageId="19" captionText="Figure 36–41. Female genitalia of Eucosmocydia. 36) E. pappeana from Kenya, USNM slide 153,686. 37) E. pappeana from South Africa, USNM slide 144,514. 38) E. mixographa, USNM slide 143,434. 39) E. nigeriana, USNM slide 153,741. 40) E. chlorobathra from Kenya, USNM slide 143,442. 41) E. chlorobathra from the Seychelles, USNM slide 153,749." figureDoi="http://doi.org/10.5281/zenodo.6533446" httpUri="https://zenodo.org/record/6533446/files/figure.png" pageId="18" pageNumber="17">39</figureCitation>
</paragraph>
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<subSubSection id="C3306530FFBEDF2EB3B17FE0FEC1FA9C" pageId="18" pageNumber="17" type="diagnosis">
<paragraph id="8B9536BBFFBEDF2EB3B17FE0FEC1FA9C" blockId="18.[192,1440,1204,1399]" pageId="18" pageNumber="17">
<emphasis id="B95EEAA9FFBEDF2EB3B17FE0FEEEFB25" bold="true" box="[192,316,1204,1230]" pageId="18" pageNumber="17">Diagnosis.</emphasis>
<taxonomicName id="4C2A4D38FFBEDF2EB2327FE0FD9CFB25" authorityName="Brown and Razowski" authorityYear="2022" box="[323,590,1204,1230]" class="Insecta" family="Tortricidae" genus="Eucosmocydia" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="nigeriana">
<emphasis id="B95EEAA9FFBEDF2EB2327FE0FD9CFB25" box="[323,590,1204,1230]" italics="true" pageId="18" pageNumber="17">Eucosmocydia nigeriana</emphasis>
</taxonomicName>
is nearly indistinguishable from 
<taxonomicName id="4C2A4D38FFBEDF2EB0B27FE1FBADFB25" box="[963,1151,1204,1231]" class="Insecta" family="Tortricidae" genus="Eucosmocydia" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="lecaniodiscana">
<emphasis id="B95EEAA9FFBEDF2EB0B27FE1FC00FB25" box="[963,978,1205,1230]" italics="true" pageId="18" pageNumber="17">E</emphasis>
. 
<emphasis id="B95EEAA9FFBEDF2EB0AE7FE0FBADFB25" box="[991,1151,1204,1230]" italics="true" pageId="18" pageNumber="17">lecaniodiscana</emphasis>
</taxonomicName>
in facies, male secondary features, and genitalia. However, in the female genitalia of 
<taxonomicName id="4C2A4D38FFBEDF2EB02E7F83FC34FB1B" authorityName="Brown and Razowski" authorityYear="2022" box="[863,998,1239,1264]" class="Insecta" family="Tortricidae" genus="Eucosmocydia" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="nigeriana">
<emphasis id="B95EEAA9FFBEDF2EB02E7F83FC34FB1B" box="[863,998,1239,1264]" italics="true" pageId="18" pageNumber="17">E. nigeriana</emphasis>
</taxonomicName>
, the somewhat parallel-sided anteriormost portion of the pocket of the sterigma is slightly shorter than that of 
<taxonomicName id="4C2A4D38FFBEDF2EB08F7FACFB68FAFA" box="[1022,1210,1271,1298]" class="Insecta" family="Tortricidae" genus="Eucosmocydia" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="lecaniodiscana">
<emphasis id="B95EEAA9FFBEDF2EB08F7FACFBDFFAFA" box="[1022,1037,1272,1297]" italics="true" pageId="18" pageNumber="17">E</emphasis>
. 
<emphasis id="B95EEAA9FFBEDF2EB76B7FA3FB68FAFA" box="[1050,1210,1271,1297]" italics="true" pageId="18" pageNumber="17">lecaniodiscana</emphasis>
</taxonomicName>
. Also, the sclerite of abdominal segment 7 of the male (
<figureCitation id="13112A3EFFBEDF2EB13C7E4DFD4CFAD8" box="[589,670,1305,1331]" captionStart="Figures 28–35" captionStartId="17.[192,279,1799,1825]" captionTargetBox="[284,1348,216,1770]" captionTargetId="figure-82@17.[284,1351,216,1770]" captionTargetPageId="17" captionText="Figures 28–35. Abdominal segment 7 of males of Eucosmocydia. 28) E. pappeana, USNM slide 143,426. 29) E. ugandensis, NHMO slide 3762. 30) E. deinbolliana, USNM slide 153,700. 31) E. lecaniodiscana, USNM slide 153,704. 32) E. nigeriana, USNM slide 153,740. 33) E. pancoviana, USNM slide 153,695. 34) E. chlorobathra, USNM slide 153,691. 35) E. kirimiriana, USNM slide 143,441." figureDoi="http://doi.org/10.5281/zenodo.6533444" httpUri="https://zenodo.org/record/6533444/files/figure.png" pageId="18" pageNumber="17">Fig. 32</figureCitation>
) has a small, median, triangular point, rather than rounded as in 
<taxonomicName id="4C2A4D38FFBEDF2EB6FD7E4EFEB2FABF" class="Insecta" family="Tortricidae" genus="Eucosmocydia" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="lecaniodiscana">
<emphasis id="B95EEAA9FFBEDF2EB6FD7E4EFA48FAD8" box="[1420,1434,1306,1331]" italics="true" pageId="18" pageNumber="17">E</emphasis>
. 
<emphasis id="B95EEAA9FFBEDF2EB3B17E6EFEB2FABF" box="[192,352,1338,1364]" italics="true" pageId="18" pageNumber="17">lecaniodiscana</emphasis>
</taxonomicName>
(
<figureCitation id="13112A3EFFBEDF2EB21E7E6EFE6DFABE" box="[367,447,1338,1365]" captionStart="Figures 28–35" captionStartId="17.[192,279,1799,1825]" captionTargetBox="[284,1348,216,1770]" captionTargetId="figure-82@17.[284,1351,216,1770]" captionTargetPageId="17" captionText="Figures 28–35. Abdominal segment 7 of males of Eucosmocydia. 28) E. pappeana, USNM slide 143,426. 29) E. ugandensis, NHMO slide 3762. 30) E. deinbolliana, USNM slide 153,700. 31) E. lecaniodiscana, USNM slide 153,704. 32) E. nigeriana, USNM slide 153,740. 33) E. pancoviana, USNM slide 153,695. 34) E. chlorobathra, USNM slide 153,691. 35) E. kirimiriana, USNM slide 143,441." figureDoi="http://doi.org/10.5281/zenodo.6533444" httpUri="https://zenodo.org/record/6533444/files/figure.png" pageId="18" pageNumber="17">Fig. 31</figureCitation>
). These slight differences in morphology are corroborated by differences in barcodes (see below).
</paragraph>
</subSubSection>
<subSubSection id="C3306530FFBEDF28B3B17ED1FBB9FD5E" lastPageId="20" lastPageNumber="19" pageId="18" pageNumber="17" type="description">
<paragraph id="8B9536BBFFBEDF28B3B17ED1FCFFFEFA" blockId="18.[191,1442,1413,1911]" lastBlockId="20.[192,1440,214,475]" lastPageId="20" lastPageNumber="19" pageId="18" pageNumber="17">
<emphasis id="B95EEAA9FFBEDF2EB3B17ED1FE80FA74" bold="true" box="[192,338,1413,1439]" pageId="18" pageNumber="17">Description.</emphasis>
<emphasis id="B95EEAA9FFBEDF2EB2287ED2FE0AFA4B" box="[345,472,1414,1440]" italics="true" pageId="18" pageNumber="17">Male. Head</emphasis>
. Scales of vertex and frons pale gray mixed with cream; labial palpus weakly upturned, length approximately 1.2 times diameter of compound eye, third segment exposed, scales of labial palpus and basal flagellomeres of antenna concolorous with vertex. 
<emphasis id="B95EEAA9FFBEDF2EB03E7E9DFC4FFA08" box="[847,925,1481,1507]" italics="true" pageId="18" pageNumber="17">Thorax</emphasis>
. Nota and tegula pale brown speckled with cream. Forewing (
<figureCitation id="13112A3EFFBEDF2EB2E07EBEFE06F9EE" box="[401,468,1514,1541]" captionStart="Figures 2–9" captionStartId="10.[192,279,1537,1563]" captionTargetBox="[192,1439,213,1507]" captionTargetId="figure-158@10.[192,1439,213,1510]" captionTargetPageId="10" captionText="Figures 2–9. Adults of Eucosmocydia. 2) E. pappeana, male holotype. 3) E. pappeana, female paratype. 4) E. deinbolliana, male paratype. 5) E. mixographa, female. 6) E. lecaniodiscana, male holotype. 7) E. lecaniodiscana, female paratype. 8) E. nigeriana, male holotype. 9) E. nigeriana, paratype female." figureDoi="http://doi.org/10.5281/zenodo.6533438" httpUri="https://zenodo.org/record/6533438/files/figure.png" pageId="18" pageNumber="17">Fig. 8</figureCitation>
) length 5.0– 
<quantity id="4CD29B5EFFBEDF2EB1197EBEFD13F9EE" box="[616,705,1514,1541]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="18" pageNumber="17" unit="mm" value="5.5">5.5 mm</quantity>
(n = 2); forewing with costa gently arched throughout, termen weakly concave beneath apex, broadly convex in remainder; pattern slightly two-toned, with slender, ill-defined, brown, median line dividing basal 0.5 from distal 0.5, terminating before costa; upperside ground color in basal 0.5 orange yellow, speckled with greyish brown; distal 0.5 with pattern more complex including small irregular patches of pale orange and brown, a larger leaden gray patch in tornus; costal strigulae cream, slender, divisions brownish; speculum near mid-termen with slender, pinkish cream crescent-shaped mark and two black dots. Forewing underside tan with alternating rows of black and cream scales in basal 0.75, except in costal region, hind margin cream. Fringe orange-yellow with brown basal line. Hindwing brown, paler in basal region, with scattered black scales in male. Fringe cream with brown basal line. 
<emphasis id="B95EEAA9FFBEDF2EB0417DA3FC48F8FA" box="[816,922,1783,1809]" italics="true" pageId="18" pageNumber="17">Abdomen</emphasis>
. Segment 7 (
<figureCitation id="13112A3EFFBEDF2EB7567DA3FBA7F8FA" box="[1063,1141,1783,1809]" captionStart="Figures 28–35" captionStartId="17.[192,279,1799,1825]" captionTargetBox="[284,1348,216,1770]" captionTargetId="figure-82@17.[284,1351,216,1770]" captionTargetPageId="17" captionText="Figures 28–35. Abdominal segment 7 of males of Eucosmocydia. 28) E. pappeana, USNM slide 143,426. 29) E. ugandensis, NHMO slide 3762. 30) E. deinbolliana, USNM slide 153,700. 31) E. lecaniodiscana, USNM slide 153,704. 32) E. nigeriana, USNM slide 153,740. 33) E. pancoviana, USNM slide 153,695. 34) E. chlorobathra, USNM slide 153,691. 35) E. kirimiriana, USNM slide 143,441." figureDoi="http://doi.org/10.5281/zenodo.6533444" httpUri="https://zenodo.org/record/6533444/files/figure.png" pageId="18" pageNumber="17">Fig. 32</figureCitation>
) with rounded-trapezoidal sclerite at mid-venter of anterior margin, with a small, median, triangular point on posterior margin. Genitalia (
<figureCitation id="13112A3EFFBEDF2EB3B87C6EFECBF8BE" box="[201,281,1850,1877]" captionStart="Figures 20–27" captionStartId="15.[192,279,1799,1825]" captionTargetBox="[207,1425,213,1770]" captionTargetId="figure-12@15.[207,1425,213,1773]" captionTargetPageId="15" captionText="Figures 20–27. Male genitalia of Eucosmocydia. 20) E. pappeana, USNM slide 153,769. 21) E. deinbolliana, USNM slide 153,700. 22) E. ugandensis, slide NHMO 3762. 23) E. lecaniodiscana, USNM slide 124,465. 24) Male genitalia of E. nigeriana, USNM slide 153,740.25) Male genitalia of E. pancoviana, USNM slide 153,695.26) Male genitalia of E. chlorobathra, USNM slide 153,691. 27) Male genitalia of E. kirimiriana, USNM slide 143,441." figureDoi="http://doi.org/10.5281/zenodo.6533442" httpUri="https://zenodo.org/record/6533442/files/figure.png" pageId="18" pageNumber="17">Fig. 24</figureCitation>
) with tegumen broad, ovoid, with sparse fine setae in dorso-posterior 0.5 representing fused socii valva narrowest in basal 0.5, ventral margin abruptly broadened and rounded at base of cucullus, costa and ventral margins of cucullus somewhat parallel-sided, rounded; caulis long, rodlike; phallus somewhat evenly curved throughout, distinctly expanded at junction with caulis.
</paragraph>
<caption id="DF556633FFBFDF2FB3B17DDDFE7CF8E2" ID-DOI="http://doi.org/10.5281/zenodo.6533446" ID-Zenodo-Dep="6533446" httpUri="https://zenodo.org/record/6533446/files/figure.png" pageId="19" pageNumber="18" startId="19.[192,268,1673,1699]" targetBox="[192,1438,213,1644]" targetPageId="19">
<paragraph id="8B9536BBFFBFDF2FB3B17DDDFE7CF8E2" blockId="19.[191,1440,1673,1801]" pageId="19" pageNumber="18">
<emphasis id="B95EEAA9FFBFDF2FB3B17DDDFEB2F94F" bold="true" box="[192,352,1673,1700]" pageId="19" pageNumber="18">Figure 36–41.</emphasis>
Female genitalia of 
<taxonomicName id="4C2A4D38FFBFDF2FB14E7DDEFD09F94F" authorityName="Diakonoff" authorityYear="1988" box="[575,731,1674,1700]" class="Insecta" family="Tortricidae" genus="Eucosmocydia" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAA9FFBFDF2FB14E7DDEFD09F94F" box="[575,731,1674,1700]" italics="true" pageId="19" pageNumber="18">Eucosmocydia</emphasis>
</taxonomicName>
. 
<emphasis id="B95EEAA9FFBFDF2FB1997DDDFCDCF948" bold="true" box="[744,782,1673,1699]" pageId="19" pageNumber="18">36)</emphasis>
<taxonomicName id="4C2A4D38FFBFDF2FB0657DDFFC49F94F" box="[788,923,1674,1700]" class="Insecta" family="Tortricidae" genus="Eucosmocydia" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="pappeana">
<emphasis id="B95EEAA9FFBFDF2FB0657DDFFCF6F94F" box="[788,804,1675,1700]" italics="true" pageId="19" pageNumber="18">E</emphasis>
. 
<emphasis id="B95EEAA9FFBFDF2FB0407DDFFC49F94F" box="[817,923,1675,1700]" italics="true" pageId="19" pageNumber="18">pappeana</emphasis>
</taxonomicName>
from 
<collectingCountry id="F33D762BFFBFDF2FB0AE7DDEFBF5F94F" box="[991,1063,1674,1700]" name="Kenya" pageId="19" pageNumber="18">Kenya</collectingCountry>
, USNM slide 153,686. 
<emphasis id="B95EEAA9FFBFDF2FB6547DDDFA99F948" bold="true" box="[1317,1355,1673,1699]" pageId="19" pageNumber="18">37)</emphasis>
<taxonomicName id="4C2A4D38FFBFDF2FB6237DDFFED0F92E" class="Insecta" family="Tortricidae" genus="Eucosmocydia" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="pappeana">
<emphasis id="B95EEAA9FFBFDF2FB6237DDFFAB0F94F" box="[1362,1378,1675,1700]" italics="true" pageId="19" pageNumber="18">E</emphasis>
. 
<emphasis id="B95EEAA9FFBFDF2FB61F7DDFFED0F92E" italics="true" pageId="19" pageNumber="18">pappeana</emphasis>
</taxonomicName>
from 
<collectingCountry id="F33D762BFFBFDF2FB2347DFFFE07F92E" box="[325,469,1707,1733]" name="South Africa" pageId="19" pageNumber="18">South Africa</collectingCountry>
, USNM slide 144,514. 
<emphasis id="B95EEAA9FFBFDF2FB1BF7DFFFD21F92E" bold="true" box="[718,755,1707,1733]" pageId="19" pageNumber="18">38)</emphasis>
<taxonomicName id="4C2A4D38FFBFDF2FB18B7DF8FC26F92E" authority=", USNM" authorityName="USNM" box="[762,1012,1707,1734]" class="Insecta" family="Tortricidae" genus="Eucosmocydia" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="mixographa">
<emphasis id="B95EEAA9FFBFDF2FB18B7DF8FCD8F92E" box="[762,778,1708,1733]" italics="true" pageId="19" pageNumber="18">E</emphasis>
. 
<emphasis id="B95EEAA9FFBFDF2FB0647DFFFC4BF92E" box="[789,921,1707,1733]" italics="true" pageId="19" pageNumber="18">mixographa</emphasis>
, USNM
</taxonomicName>
slide 143,434. 
<emphasis id="B95EEAA9FFBFDF2FB7E47DFFFB68F92E" bold="true" box="[1173,1210,1707,1733]" pageId="19" pageNumber="18">39)</emphasis>
<taxonomicName id="4C2A4D38FFBFDF2FB7B07DF8FA72F92E" authority=", USNM" authorityName="USNM" box="[1217,1440,1707,1734]" class="Insecta" family="Tortricidae" genus="Eucosmocydia" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="nigeriana">
<emphasis id="B95EEAA9FFBFDF2FB7B07DF8FB03F92E" box="[1217,1233,1708,1733]" italics="true" pageId="19" pageNumber="18">E</emphasis>
. 
<emphasis id="B95EEAA9FFBFDF2FB7AD7DF8FA96F92E" box="[1244,1348,1708,1733]" italics="true" pageId="19" pageNumber="18">nigeriana</emphasis>
, USNM
</taxonomicName>
slide 153,741. 
<emphasis id="B95EEAA9FFBFDF2FB2127D99FE5BF90C" bold="true" box="[355,393,1741,1767]" pageId="19" pageNumber="18">40)</emphasis>
<taxonomicName id="4C2A4D38FFBFDF2FB2E27D9AFDEFF90C" box="[403,573,1741,1767]" class="Insecta" family="Tortricidae" genus="Eucosmocydia" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="chlorobathra">
<emphasis id="B95EEAA9FFBFDF2FB2E27D9AFE71F90C" box="[403,419,1742,1767]" italics="true" pageId="19" pageNumber="18">E</emphasis>
. 
<emphasis id="B95EEAA9FFBFDF2FB2C37D99FDEFF90C" box="[434,573,1741,1767]" italics="true" pageId="19" pageNumber="18">chlorobathra</emphasis>
</taxonomicName>
from 
<collectingCountry id="F33D762BFFBFDF2FB1F67D99FD1DF90C" box="[647,719,1741,1767]" name="Kenya" pageId="19" pageNumber="18">Kenya</collectingCountry>
, USNM slide 143,442. 
<emphasis id="B95EEAA9FFBFDF2FB0A97D99FC2CF90C" bold="true" box="[984,1022,1741,1767]" pageId="19" pageNumber="18">41)</emphasis>
<taxonomicName id="4C2A4D38FFBFDF2FB7767D9AFB60F90C" box="[1031,1202,1741,1767]" class="Insecta" family="Tortricidae" genus="Eucosmocydia" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="chlorobathra">
<emphasis id="B95EEAA9FFBFDF2FB7767D9AFBC5F90C" box="[1031,1047,1742,1767]" italics="true" pageId="19" pageNumber="18">E</emphasis>
. 
<emphasis id="B95EEAA9FFBFDF2FB7567D99FB60F90C" box="[1063,1202,1741,1767]" italics="true" pageId="19" pageNumber="18">chlorobathra</emphasis>
</taxonomicName>
from the 
<collectingCountry id="F33D762BFFBFDF2FB6597D99FA4EF90C" box="[1320,1436,1741,1767]" name="Seychelles" pageId="19" pageNumber="18">Seychelles</collectingCountry>
, USNM slide 153,749.
</paragraph>
</caption>
<paragraph id="8B9536BBFFB8DF28B2717A4DFE70FE30" blockId="20.[192,1440,214,475]" pageId="20" pageNumber="19">
<emphasis id="B95EEAA9FFB8DF28B2717A4DFE89FED8" bold="true" box="[256,347,281,307]" pageId="20" pageNumber="19">Female.</emphasis>
<emphasis id="B95EEAA9FFB8DF28B2137A4DFDCCFED8" box="[354,542,281,307]" italics="true" pageId="20" pageNumber="19">Head and thorax</emphasis>
. Essentially as described for male, but hindwing uniformly dark brown (
<figureCitation id="13112A3EFFB8DF28B6217A4DFA40FED8" box="[1360,1426,281,307]" captionStart="Figures 2–9" captionStartId="10.[192,279,1537,1563]" captionTargetBox="[192,1439,213,1507]" captionTargetId="figure-158@10.[192,1439,213,1510]" captionTargetPageId="10" captionText="Figures 2–9. Adults of Eucosmocydia. 2) E. pappeana, male holotype. 3) E. pappeana, female paratype. 4) E. deinbolliana, male paratype. 5) E. mixographa, female. 6) E. lecaniodiscana, male holotype. 7) E. lecaniodiscana, female paratype. 8) E. nigeriana, male holotype. 9) E. nigeriana, paratype female." figureDoi="http://doi.org/10.5281/zenodo.6533438" httpUri="https://zenodo.org/record/6533438/files/figure.png" pageId="20" pageNumber="19">Fig. 9</figureCitation>
). 
<emphasis id="B95EEAA9FFB8DF28B3B07A6EFEF8FEBF" box="[193,298,314,340]" italics="true" pageId="20" pageNumber="19">Abdomen</emphasis>
. Venter of segment 7 with broad, subsquare area of faint sclerotization. Female genitalia (
<figureCitation id="13112A3EFFB8DF28B6617A6EFAB2FEBE" box="[1296,1376,314,341]" captionStart="Figure 36–41" captionStartId="19.[192,268,1673,1699]" captionTargetBox="[192,1438,213,1644]" captionTargetId="figure-12@19.[192,1438,213,1647]" captionTargetPageId="19" captionText="Figure 36–41. Female genitalia of Eucosmocydia. 36) E. pappeana from Kenya, USNM slide 153,686. 37) E. pappeana from South Africa, USNM slide 144,514. 38) E. mixographa, USNM slide 143,434. 39) E. nigeriana, USNM slide 153,741. 40) E. chlorobathra from Kenya, USNM slide 143,442. 41) E. chlorobathra from the Seychelles, USNM slide 153,749." figureDoi="http://doi.org/10.5281/zenodo.6533446" httpUri="https://zenodo.org/record/6533446/files/figure.png" pageId="20" pageNumber="19">Fig. 39</figureCitation>
) with pair of ill-defined, bristly pads at posterior margin of segment 7; pocket of ostium triangular with irregularly rounded, slight expansion at anterior end, giving rise to frail part of ductus bursae; ductus bursae slender in posterior 0.5 broader in anterior 0.5 starting at origin of ductus seminalis; corpus round with two thorn-shaped signa of similar size.
</paragraph>
<paragraph id="8B9536BBFFB8DF28B3B17ABEFB85FDCC" blockId="20.[192,1440,490,551]" pageId="20" pageNumber="19">
<emphasis id="B95EEAA9FFB8DF28B3B17ABEFEA0FDEE" bold="true" box="[192,370,490,517]" pageId="20" pageNumber="19">DNA barcodes.</emphasis>
There are three sequences of this species in BOLD (BIN: 
<barcodeIndexNumber id="2ECE05CEFFB8DF28B09F7ABFFBB1FDEE" box="[1006,1123,491,517]" pageId="20" pageNumber="19">AAU2218</barcodeIndexNumber>
), with an average distance of 0.1% among them, and a 4.01% distance to the nearest neighbor, 
<taxonomicName id="4C2A4D38FFB8DF28B0E57959FB83FDCD" box="[916,1105,524,551]" class="Insecta" family="Tortricidae" genus="Eucosmocydia" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="lecaniodiscana">
<emphasis id="B95EEAA9FFB8DF28B0E57959FC76FDCD" box="[916,932,525,550]" italics="true" pageId="20" pageNumber="19">E</emphasis>
. 
<emphasis id="B95EEAA9FFB8DF28B0C07958FB83FDCD" box="[945,1105,524,550]" italics="true" pageId="20" pageNumber="19">lecaniodiscana</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9536BBFFB8DF28B3B17962FBB9FD5E" blockId="20.[192,1440,566,693]" pageId="20" pageNumber="19">
<emphasis id="B95EEAA9FFB8DF28B3B17962FED9FDBB" bold="true" box="[192,267,566,592]" pageId="20" pageNumber="19">Types.</emphasis>
<materialsCitation id="3B423CE6FFB8DF28B2667962FC0EFD9A" ID-GBIF-Occurrence="4406082320" collectingDate="2006-06-19" collectionCode="USNM" collectorName="G. M. Miller &amp; T. M. Kuklenski" county="Nigeria" elevation="240" latitude="7.5008" location="International Institute" longLatPrecision="7" longitude="3.9065" municipality="Ibadan" pageId="20" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Oyo" typeStatus="holotype">
<typeStatus id="54918819FFB8DF28B2667962FEADFDBB" box="[279,383,566,592]" pageId="20" pageNumber="19" type="holotype">Holotype</typeStatus>
<specimenCount id="9D2CFD32FFB8DF28B2FD796DFE73FDA4" box="[396,417,569,591]" pageId="20" pageNumber="19" type="male">♂</specimenCount>
, 
<collectingCounty id="62F44E37FFB8DF28B2C57962FDDBFDBB" box="[436,521,566,592]" pageId="20" pageNumber="19">Nigeria</collectingCounty>
, 
<collectingRegion id="49EEF859FFB8DF28B16B7962FD5CFDBB" box="[538,654,566,592]" country="Nigeria" name="Oyo" pageId="20" pageNumber="19">Oyo State</collectingRegion>
, 
<collectingMunicipality id="6BF1ACC1FFB8DF28B1EE7962FD22FDBB" box="[671,752,566,592]" pageId="20" pageNumber="19">Ibadan</collectingMunicipality>
, 
<location id="8EF56060FFB8DF28B0717962FC2EFDBB" LSID="urn:lsid:plazi:treatment:038387ADFFBEDF28B3B17F33FC34FCC2:8EF56060FFB8DF28B0717962FC2EFDBB" box="[768,1020,566,592]" county="Nigeria" latitude="7.5008" longLatPrecision="7" longitude="3.9065" municipality="Ibadan" name="International Institute" pageId="20" pageNumber="19" stateProvince="Oyo">International Institute</location>
for 
<location id="8EF56060FFB8DF28B7447962FAF3FDBB" LSID="urn:lsid:plazi:treatment:038387ADFFBEDF28B3B17F33FC34FCC2:8EF56060FFB8DF28B7447962FAF3FDBB" box="[1077,1313,566,592]" county="Nigeria" latitude="7.5008" longLatPrecision="7" longitude="3.9065" municipality="Ibadan" name="Tropical Agriculture" pageId="20" pageNumber="19" stateProvince="Oyo">Tropical Agriculture</location>
, 
<geoCoordinate id="EE1E507CFFB8DF28B6437962FA4EFDBB" box="[1330,1436,566,593]" degrees="7.5008" direction="north" orientation="latitude" pageId="20" pageNumber="19" precision="5" value="7.5008">7.5008°N</geoCoordinate>
, 
<geoCoordinate id="EE1E507CFFB8DF28B3B17903FEF7FD99" box="[192,293,599,626]" degrees="3.9065" direction="east" orientation="longitude" pageId="20" pageNumber="19" precision="5" value="3.9065">3.9065°E</geoCoordinate>
, 
<quantity id="4CD29B5EFFB8DF28B2417903FEA5FD99" box="[304,375,599,626]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="20" pageNumber="19" unit="m" value="240.0">
<elevation id="0007D188FFB8DF28B2417903FEA5FD99" box="[304,375,599,626]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="20" pageNumber="19" unit="m" value="240.0">240 m</elevation>
</quantity>
, 
<date id="FF94107BFFB8DF28B2F37903FDDAFD9A" box="[386,520,599,626]" pageId="20" pageNumber="19" value="2006-06-19">
<collectingDate id="EFD0E993FFB8DF28B2F37903FDDAFD9A" box="[386,520,599,626]" pageId="20" pageNumber="19" value="2006-06-19">19 Jun 2006</collectingDate>
</date>
, 
<collectorName id="26DF536DFFB8DF28B1627903FD4CFD9A" box="[531,670,599,626]" pageId="20" pageNumber="19">G. M. Miller</collectorName>
&amp; 
<collectorName id="26DF536DFFB8DF28B1CF790CFCBDFD9A" box="[702,879,599,626]" pageId="20" pageNumber="19">T. M. Kuklenski</collectorName>
(
<collectionCode id="ED3BAE7EFFB8DF28B00F7903FC06FD9A" box="[894,980,599,625]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="19" type="Museum">USNM</collectionCode>
)
</materialsCitation>
. 
<typeStatus id="54918819FFB8DF28B094790CFB83FD99" box="[997,1105,600,626]" pageId="20" pageNumber="19" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D2CFD32FFB8DF28B7137903FB51FD9A" box="[1122,1155,599,625]" pageId="20" pageNumber="19" type="male">1♂</specimenCount>
, 
<specimenCount id="9D2CFD32FFB8DF28B7E17903FB61FD9A" box="[1168,1203,599,625]" pageId="20" pageNumber="19" type="female">1♀</specimenCount>
). 
<emphasis id="B95EEAA9FFB8DF28B7B87903FAF5FD9A" bold="true" box="[1225,1319,599,625]" pageId="20" pageNumber="19">
<collectingCountry id="F33D762BFFB8DF28B7B87903FAF0FD9A" box="[1225,1314,599,625]" name="Nigeria" pageId="20" pageNumber="19">Nigeria</collectingCountry>
:
</emphasis>
<collectingRegion id="49EEF859FFB8DF28B65F7903FA4EFD9A" box="[1326,1436,599,625]" country="Nigeria" name="Oyo" pageId="20" pageNumber="19">Oyo State</collectingRegion>
, Ibadan, International Institute for Tropical Agriculture, 
<geoCoordinate id="EE1E507CFFB8DF28B042792DFC4FFD78" box="[819,925,633,660]" degrees="7.5008" direction="north" orientation="latitude" pageId="20" pageNumber="19" precision="5" value="7.5008">7.5008°N</geoCoordinate>
, 
<geoCoordinate id="EE1E507CFFB8DF28B0D8792DFBDCFD78" box="[937,1038,633,660]" degrees="3.9065" direction="east" orientation="longitude" pageId="20" pageNumber="19" precision="5" value="3.9065">3.9065°E</geoCoordinate>
, 
<quantity id="4CD29B5EFFB8DF28B768792DFBB0FD78" box="[1049,1122,633,659]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="20" pageNumber="19" unit="m" value="240.0">240 m</quantity>
, 
<date id="FF94107BFFB8DF28B71C792DFAF7FD78" box="[1133,1317,633,659]" pageId="20" pageNumber="19" value="2006-03-14" valueMax="2006-03-31" valueMin="2006-03-14">14–31 Mar 2006</date>
(
<specimenCount id="9D2CFD32FFB8DF28B646792DFA8AFD79" box="[1335,1368,633,659]" pageId="20" pageNumber="19" type="female">1♀</specimenCount>
), S. E. Miller &amp; T. M. Kuklenski, 
<date id="FF94107BFFB8DF28B29579CFFDBAFD5F" box="[484,616,666,693]" pageId="20" pageNumber="19" value="2006-06-18">18 Jun 2006</date>
(
<specimenCount id="9D2CFD32FFB8DF28B10879CFFD48FD5F" box="[633,666,667,693]" pageId="20" pageNumber="19" type="male">1♂</specimenCount>
), T. M. Kuklenski &amp; T. Olorode (USNM).
</paragraph>
</subSubSection>
<subSubSection id="C3306530FFB8DF28B3B17990FCC6FCEB" pageId="20" pageNumber="19" type="distribution">
<paragraph id="8B9536BBFFB8DF28B3B17990FCC6FCEB" blockId="20.[192,1439,708,768]" pageId="20" pageNumber="19">
<emphasis id="B95EEAA9FFB8DF28B3B17990FE3DFD35" bold="true" box="[192,495,708,734]" pageId="20" pageNumber="19">Distribution and biology.</emphasis>
<taxonomicName id="4C2A4D38FFB8DF28B2897990FCD4FD35" box="[504,774,708,734]" class="Insecta" family="Tortricidae" genus="Eucosmocydia" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="nigeriana">
<emphasis id="B95EEAA9FFB8DF28B2897990FCD4FD35" box="[504,774,708,734]" italics="true" pageId="20" pageNumber="19">Eucosmocydia nigeriana</emphasis>
</taxonomicName>
is known only from 
<collectingCountry id="F33D762BFFB8DF28B08F7990FB81FD35" box="[1022,1107,708,734]" name="Nigeria" pageId="20" pageNumber="19">Nigeria</collectingCountry>
, where it was collected from ultraviolet light. Nothing is known of the early stages.
</paragraph>
</subSubSection>
<subSubSection id="C3306530FFB8DF28B3B1785BFC34FCC2" box="[192,998,783,810]" pageId="20" pageNumber="19" type="etymology">
<paragraph id="8B9536BBFFB8DF28B3B1785BFC34FCC2" blockId="20.[192,998,783,810]" box="[192,998,783,810]" pageId="20" pageNumber="19">
<emphasis id="B95EEAA9FFB8DF28B3B1785BFE96FCC2" bold="true" box="[192,324,783,809]" pageId="20" pageNumber="19">Etymology.</emphasis>
The specific epithet refers to the country of the 
<typeStatus id="54918819FFB8DF28B0287844FC5BFCC1" box="[857,905,784,810]" pageId="20" pageNumber="19">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>