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<document id="2361C18423DBFBA02235DF296DA28E47" ID-DOI="10.5281/zenodo.177146" ID-GBIF-Dataset="dd5cde2b-ee92-42f9-8214-f088aa60d7d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="177146" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459850893375" checkinUser="plazi" docAuthor="Fibiger, Michael, Zahiri, Reza &amp; Kononenko, Vladimir S." docDate="2007" docId="03838794FFBBFFF6FF191DA9FE532F3F" docLanguage="en" docName="zt01503p019.pdf" docOrigin="Zootaxa 1503" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Margelana brunnea Fibiger, Zahiri &amp; Kononenko, 2007, sp. n." docType="treatment" docVersion="9" lastPageNumber="18" masterDocId="FFBAFFECFFBAFFF3FF8E1D7EFFAE2B23" masterDocTitle="Description of a new species of the Central Palaearctic genus Margelana Staudinger (Lepidoptera: Noctuidae) from northeastern Iran" masterLastPageNumber="19" masterPageNumber="13" pageNumber="14" updateTime="1698225264547" updateUser="plazi">
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<mods:title id="DB7D7F38FC13C001DCC6ECCF6D99AC18">Description of a new species of the Central Palaearctic genus Margelana Staudinger (Lepidoptera: Noctuidae) from northeastern Iran</mods:title>
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<mods:name id="68D5D0FA9FB8C4C4DE4A59321BCF3BD1" type="personal">
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<mods:namePart id="020EC9E16246A0363E7A62AF98EAC222">Fibiger, Michael</mods:namePart>
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<mods:namePart id="5AE0E515B7ADC82FE1B2FCF609DFF661">Zahiri, Reza</mods:namePart>
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<mods:namePart id="DC0A577D7EEBF5FFB54DC9BB97D09701">Kononenko, Vladimir S.</mods:namePart>
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<mods:part id="5CCE392425F7B6AD3294C833AFAD584F">
<mods:date id="39753913969C33EBEDAD6767C1FCB042">2007</mods:date>
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<treatment id="03838794FFBBFFF6FF191DA9FE532F3F" ID-DOI="http://doi.org/10.5281/zenodo.6243243" ID-GBIF-Taxon="119362458" ID-Zenodo-Dep="6243243" LSID="urn:lsid:plazi:treatment:03838794FFBBFFF6FF191DA9FE532F3F" httpUri="http://treatment.plazi.org/id/03838794FFBBFFF6FF191DA9FE532F3F" lastPageId="5" lastPageNumber="18" pageId="1" pageNumber="14">
<subSubSection id="C3306509FFBBFFF2FF191DA9FEFD2A37" pageId="1" pageNumber="14" type="nomenclature">
<paragraph id="8B953682FFBBFFF2FF191DA9FE722BD2" blockId="1.[151,476,215,276]" box="[151,476,215,241]" pageId="1" pageNumber="14">
<heading id="D0DD81EEFFBBFFF2FF191DA9FE722BD2" bold="true" box="[151,476,215,241]" fontSize="11" level="1" pageId="1" pageNumber="14" reason="1">
<emphasis id="B95EEA90FFBBFFF2FF191DA9FE722BD2" bold="true" box="[151,476,215,241]" pageId="1" pageNumber="14">
<taxonomicName id="4C2A4D01FFBBFFF2FF191DA9FE252BD2" box="[151,395,215,241]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="14" phylum="Arthropoda" rank="species" species="brunnea" status="sp. nov.">
<emphasis id="B95EEA90FFBBFFF2FF191DA9FE252BD2" bold="true" box="[151,395,215,241]" italics="true" pageId="1" pageNumber="14">Margelana brunnea</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26D57EBFFBBFFF2FE141DA9FE722BD2" box="[410,476,215,241]" pageId="1" pageNumber="14" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B953682FFBBFFF2FF191D84FEFD2A37" blockId="1.[151,476,215,276]" box="[151,339,250,276]" pageId="1" pageNumber="14">
(
<figureCitation id="13112A07FFBBFFF2FF2E1D84FEBF2A37" box="[160,273,250,276]" captionStart="FIGURES 1 4" captionStartId="1.[151,270,1455,1479]" captionTargetBox="[151,1435,713,1408]" captionTargetId="figure@1.[151,1436,695,1448]" captionTargetPageId="1" captionText="FIGURES 1 4. Adults of Margelana. 1, Holotype male of Margelana brunnea; 2, paratype female of Margelana brunnea; 3, male of M. flavidior; 4, female of M. versicolor." httpUri="https://zenodo.org/record/177147/files/figure.png" pageId="1" pageNumber="14">Figs. 1, 2</figureCitation>
,
<figureCitation id="13112A07FFBBFFF2FE931D84FE832A37" box="[285,301,250,276]" captionStart="FIGURES 5 7" captionStartId="2.[151,269,1963,1987]" captionTargetBox="[257,1436,201,1914]" captionTargetId="figure@2.[151,1436,194,1939]" captionTargetPageId="2" captionText="FIGURES 5 7. Male genitalia of Margelana. 5, M. brunnea; 6, M. flavidior; 7, M. versicolor." httpUri="https://zenodo.org/record/177148/files/figure.png" pageId="1" pageNumber="14">5</figureCitation>
,
<figureCitation id="13112A07FFBBFFF2FEB41D84FEE72A37" box="[314,329,250,276]" captionStart="FIGURES 8 10" captionStartId="3.[151,270,1641,1665]" captionTargetBox="[151,1433,628,1612]" captionTargetId="figure@3.[151,1436,628,1617]" captionTargetPageId="3" captionText="FIGURES 8 10. Female genitalia of Margelana. 8, M. brunnea; 9, M. flavidior; 10, M. versicolor." httpUri="https://zenodo.org/record/177149/files/figure.png" pageId="1" pageNumber="14">8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3306509FFBBFFF6FF191C37FAD629AF" lastPageId="5" lastPageNumber="18" pageId="1" pageNumber="14" type="description">
<paragraph id="8B953682FFBBFFF2FF191C37FC08295F" blockId="1.[151,1437,329,636]" pageId="1" pageNumber="14">
<emphasis id="B95EEA90FFBBFFF2FF191C37FF4B2A40" bold="true" box="[151,229,329,355]" pageId="1" pageNumber="14">
<typeStatus id="54918820FFBBFFF2FF191C37FF702A40" box="[151,222,329,355]" pageId="1" pageNumber="14">Types</typeStatus>
.
</emphasis>
<typeStatus id="54918820FFBBFFF2FF7D1C34FE202A47" box="[243,398,330,356]" pageId="1" pageNumber="14" type="holotype">HOLOTYPE</typeStatus>
ɗ (
<figureCitation id="13112A07FFBBFFF2FE4E1C34FDA62A47" box="[448,520,330,356]" captionStart="FIGURES 1 4" captionStartId="1.[151,270,1455,1479]" captionTargetBox="[151,1435,713,1408]" captionTargetId="figure@1.[151,1436,695,1448]" captionTargetPageId="1" captionText="FIGURES 1 4. Adults of Margelana. 1, Holotype male of Margelana brunnea; 2, paratype female of Margelana brunnea; 3, male of M. flavidior; 4, female of M. versicolor." httpUri="https://zenodo.org/record/177147/files/figure.png" pageId="1" pageNumber="14">Fig. 1</figureCitation>
),
<collectingCountry id="F33D7612FFBBFFF2FDA81C34FDF62A47" box="[550,600,330,356]" name="Iran" pageId="1" pageNumber="14">Iran</collectingCountry>
, Prov. Golestan, Golestan National Park, Almeh,
<quantity id="4CD29B67FFBBFFF2FB361C34FAB62A47" box="[1208,1304,330,356]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="1" pageNumber="14" unit="m" value="1650.0">1650 m</quantity>
,
<date id="FF941042FFBBFFF2FAA91C34FEA32AAF" pageId="1" pageNumber="14" value="1988-10-02" valueMax="1988-10-12" valueMin="1988-10-02">2 through 12.x.1988</date>
(A. Pazuki); deposited at HMIM.
<typeStatus id="54918820FFBBFFF2FD101C0CFCEE2AAF" box="[670,832,370,396]" pageId="1" pageNumber="14" type="paratype">PARATYPES</typeStatus>
(6 ɗ and 47 Ψ): 6 ɗ and 37 Ψ (
<figureCitation id="13112A07FFBBFFF2FB221C0CFB5A2AAF" box="[1196,1268,370,396]" captionStart="FIGURES 1 4" captionStartId="1.[151,270,1455,1479]" captionTargetBox="[151,1435,713,1408]" captionTargetId="figure@1.[151,1436,695,1448]" captionTargetPageId="1" captionText="FIGURES 1 4. Adults of Margelana. 1, Holotype male of Margelana brunnea; 2, paratype female of Margelana brunnea; 3, male of M. flavidior; 4, female of M. versicolor." httpUri="https://zenodo.org/record/177147/files/figure.png" pageId="1" pageNumber="14">Fig. 2</figureCitation>
), same data as
<typeStatus id="54918820FFBBFFF2FF191CE4FEAF2A97" box="[151,257,410,436]" pageId="1" pageNumber="14" type="holotype">holotype</typeStatus>
, but genitalic preparations 5576, 5559 and 5519 M. Fibiger; 2 Ψ, same data as
<typeStatus id="54918820FFBBFFF2FB1B1CE4FB502A97" box="[1173,1278,410,436]" pageId="1" pageNumber="14" type="holotype">holotype</typeStatus>
, but
<date id="FF941042FFBBFFF2FAB81CE4FA322A97" box="[1334,1436,410,436]" pageId="1" pageNumber="14" value="1994-10-03">3.x.1994</date>
(H. Mirzayans, M. Badii &amp; E. Ebrahimi); 4 Ψ, same data as
<typeStatus id="54918820FFBBFFF2FCD41CBCFC6D2AFF" box="[858,963,450,476]" pageId="1" pageNumber="14" type="holotype">holotype</typeStatus>
, but
<quantity id="4CD29B67FFBBFFF2FC721CBCFBF72AFF" box="[1020,1113,450,476]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="14" unit="m" value="1600.0">1600 m</quantity>
,
<date id="FF941042FFBBFFF2FBEB1CBCFB4C2AFF" box="[1125,1250,450,476]" pageId="1" pageNumber="14" value="2000-09-30">30.ix.2000</date>
(R. Ghayourfar &amp; E. Gilasian); 3 Ψ, same data as
<typeStatus id="54918820FFBBFFF2FDA31C94FD392927" box="[557,663,490,516]" pageId="1" pageNumber="14" type="holotype">holotype</typeStatus>
, but
<quantity id="4CD29B67FFBBFFF2FD5F1C94FC802927" box="[721,814,490,516]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="1" pageNumber="14" unit="m" value="1750.0">1750 m</quantity>
,
<date id="FF941042FFBBFFF2FCB41C94FC012927" box="[826,943,490,516]" pageId="1" pageNumber="14" value="2003-10-26">26.x.2003</date>
(G. Stangelmaier &amp; C. Wieser), 1 ɗ,
<collectingCountry id="F33D7612FFBBFFF2FAE31C94FA322927" box="[1389,1436,490,516]" name="Iran" pageId="1" pageNumber="14">Iran</collectingCountry>
NE, Kandavan,
<date id="FF941042FFBBFFF2FED81F6CFE60290F" box="[342,462,530,556]" pageId="1" pageNumber="14" value="1971-10-16">16.x.1971</date>
, genitalic preparation 5577 M. Fibiger, leg. Naim;
<typeStatus id="54918820FFBBFFF2FBBE1F6CFB0E290F" box="[1072,1184,530,556]" pageId="1" pageNumber="14" type="paratype">paratypes</typeStatus>
deposited in the collections at HMIM; M. Fibiger; LNK, Karlsruhe; NHM, London; G. Ronkay; P. Gyulai; TMB, Budapest; G. Stangelmaier, Villach,
<collectingCountry id="F33D7612FFBBFFF2FE2C1F1CFE52295F" box="[418,508,610,636]" name="Austria" pageId="1" pageNumber="14">Austria</collectingCountry>
; and C. Wieser, Klagenfurt,
<collectingCountry id="F33D7612FFBBFFF2FCC71F1CFC0D295F" box="[841,931,610,636]" name="Austria" pageId="1" pageNumber="14">Austria</collectingCountry>
.
</paragraph>
<caption id="DF55660AFFBBFFF2FF1918D1FD482ECA" httpUri="https://zenodo.org/record/177147/files/figure.png" pageId="1" pageNumber="14" targetBox="[151,1435,713,1408]" targetPageId="1">
<paragraph id="8B953682FFBBFFF2FF1918D1FD482ECA" blockId="1.[151,1436,1455,1513]" pageId="1" pageNumber="14">
<emphasis id="B95EEA90FFBBFFF2FF1918D1FEED2EE4" bold="true" box="[151,323,1455,1479]" pageId="1" pageNumber="14">FIGURES 14.</emphasis>
Adults of
<taxonomicName id="4C2A4D01FFBBFFF2FE3818D1FD852EE5" box="[438,555,1455,1478]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEA90FFBBFFF2FE3818D1FD852EE5" box="[438,555,1455,1478]" italics="true" pageId="1" pageNumber="14">Margelana</emphasis>
</taxonomicName>
. 1, Holotype male of
<taxonomicName id="4C2A4D01FFBBFFF2FC9818D1FC422EE5" box="[790,1004,1455,1478]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="14" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="B95EEA90FFBBFFF2FC9818D1FC422EE5" box="[790,1004,1455,1478]" italics="true" pageId="1" pageNumber="14">Margelana brunnea</emphasis>
</taxonomicName>
; 2, paratype female of
<taxonomicName id="4C2A4D01FFBBFFF2FB6B18D1FF132ECA" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="14" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="B95EEA90FFBBFFF2FB6B18D1FF132ECA" italics="true" pageId="1" pageNumber="14">Margelana brunnea</emphasis>
</taxonomicName>
; 3, male of
<taxonomicName id="4C2A4D01FFBBFFF2FEB018ACFE102ECA" box="[318,446,1490,1513]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="14" phylum="Arthropoda" rank="species" species="flavidior">
<emphasis id="B95EEA90FFBBFFF2FEB018ACFE102ECA" box="[318,446,1490,1513]" italics="true" pageId="1" pageNumber="14">M. flavidior</emphasis>
</taxonomicName>
; 4, female of
<taxonomicName id="4C2A4D01FFBBFFF2FDDC18ACFD4C2ECA" box="[594,738,1490,1513]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="14" phylum="Arthropoda" rank="species" species="versicolor">M. versicolor</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B953682FFBBFFF2FF491B62FEF52CCC" blockId="1.[151,1437,1564,2031]" pageId="1" pageNumber="14">
<emphasis id="B95EEA90FFBBFFF2FF491B62FEF02D15" bold="true" box="[199,350,1564,1590]" pageId="1" pageNumber="14">Description.</emphasis>
Wingspan: male,
<quantity id="4CD29B67FFBBFFF2FDC91B63FD7E2D14" box="[583,720,1565,1591]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" metricValueMax="4.0" metricValueMin="3.8" pageId="1" pageNumber="14" unit="mm" value="39.0" valueMax="40.0" valueMin="38.0">3840 mm</quantity>
; female,
<quantity id="4CD29B67FFBBFFF2FCC91B63FC7F2D14" box="[839,977,1565,1591]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" metricValueMax="4.6" metricValueMin="4.2" pageId="1" pageNumber="14" unit="mm" value="44.0" valueMax="46.0" valueMin="42.0">4246 mm</quantity>
. Labial palps small, second segment heavily tufted ventrally with long hair-like setae, third segment bent downwards, 0.33 times of second. Proboscis absent. Antennae of male bipectinate, female filiform. Frons rounded. Head, thorax, and ground colour of forewing brown; wings relatively narrow, forewing sharply pointed at apex; all transverse lines present, except a basal line, white and contrasting; antemedian line absent at costa, but longitudinal along Sc and R1 veinsand united with orbicular stigma, which from R1 forms a sharply angled V, pointing toward postmedial line, the tip of which is close to, sometimes even touching, postmedian line; postmedian line smooth, slightly bent; subterminal line less prominent; orbicular and reniform stigmata marked by pure white streaks that unite, forming a large, broad V; claviform stigma absent; fringes white. Hindwing of both sexes greyishbrown, males slightly lighter. Abdomen dark brown with a light brown anal tuft. Underside of both wings unicolorous grey, with beige fringes; forewing pattern represented by weakly marked beige postmedian line and reniform stigma.
</paragraph>
<caption id="DF55660AFFB8FFF1FF191AD5FB2B2CE1" box="[151,1157,1963,1987]" httpUri="https://zenodo.org/record/177148/files/figure.png" pageId="2" pageNumber="15" targetBox="[257,1436,201,1914]" targetPageId="2">
<paragraph id="8B953682FFB8FFF1FF191AD5FB2B2CE1" blockId="2.[151,1157,1963,1987]" box="[151,1157,1963,1987]" pageId="2" pageNumber="15">
<emphasis id="B95EEA90FFB8FFF1FF191AD5FEED2CE0" bold="true" box="[151,323,1963,1987]" pageId="2" pageNumber="15">FIGURES 57.</emphasis>
Male genitalia of
<taxonomicName id="4C2A4D01FFB8FFF1FD881AD5FDD52CE1" box="[518,635,1963,1986]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEA90FFB8FFF1FD881AD5FDD52CE1" box="[518,635,1963,1986]" italics="true" pageId="2" pageNumber="15">Margelana</emphasis>
</taxonomicName>
. 5,
<taxonomicName id="4C2A4D01FFB8FFF1FD2A1AD5FC8E2CE1" box="[676,800,1963,1986]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="B95EEA90FFB8FFF1FD2A1AD5FC8E2CE1" box="[676,800,1963,1986]" italics="true" pageId="2" pageNumber="15">M. brunnea</emphasis>
</taxonomicName>
; 6,
<taxonomicName id="4C2A4D01FFB8FFF1FCC71AD5FC662CE1" box="[841,968,1963,1986]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="species" species="flavidior">
<emphasis id="B95EEA90FFB8FFF1FCC71AD5FC662CE1" box="[841,968,1963,1986]" italics="true" pageId="2" pageNumber="15">M. flavidior</emphasis>
</taxonomicName>
; 7,
<taxonomicName id="4C2A4D01FFB8FFF1FC7F1AD5FB2E2CE1" box="[1009,1152,1963,1986]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="species" species="versicolor">M. versicolor</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B953682FFB9FFF0FF491DECFC66291F" blockId="3.[151,1437,146,572]" pageId="3" pageNumber="16">
<emphasis id="B95EEA90FFB9FFF0FF491DECFED82B88" box="[199,374,146,171]" italics="true" pageId="3" pageNumber="16">Male genitalia</emphasis>
(
<figureCitation id="13112A07FFB9FFF0FE041DECFE782B8F" box="[394,470,146,172]" captionStart="FIGURES 5 7" captionStartId="2.[151,269,1963,1987]" captionTargetBox="[257,1436,201,1914]" captionTargetId="figure@2.[151,1436,194,1939]" captionTargetPageId="2" captionText="FIGURES 5 7. Male genitalia of Margelana. 5, M. brunnea; 6, M. flavidior; 7, M. versicolor." httpUri="https://zenodo.org/record/177148/files/figure.png" pageId="3" pageNumber="16">Fig. 5</figureCitation>
). Uncus very broad, 1.5 times longer than wide, without apical hook; tegumen broad, longer than vinculum; pleural sclerite (pleurite) large, dorsal part branch-like, curling, ventral part plate-like, broad and bent; saccus large, stout, pointed; valve broadest at base; with a prominent sacculus that reaches costa; costa with a free process rounded at tip; cucullus half the width of the subbasal valve; corona absent; juxta shield-like, pointed ventrally, dorsally scobinate; transtilla extremely broad, membraneous mesially; digitus thorn-like, positioned close to costal process; clasper oblique, with a tiny ampulla, which is most easily recognised by presence of one or two small setae. Aedeagus and everted vesica (
<figureCitation id="13112A07FFB9FFF0FB421CFCFABC2ABF" box="[1228,1298,386,412]" captionStart="FIGURES 8 10" captionStartId="3.[151,270,1641,1665]" captionTargetBox="[151,1433,628,1612]" captionTargetId="figure@3.[151,1436,628,1617]" captionTargetPageId="3" captionText="FIGURES 8 10. Female genitalia of Margelana. 8, M. brunnea; 9, M. flavidior; 10, M. versicolor." httpUri="https://zenodo.org/record/177149/files/figure.png" pageId="3" pageNumber="16">Fig. 8</figureCitation>
). Aedeagus bent; coecum half as wide as rest of aedeagus; aedeagus with a dorsal membranous ridge from ductus ejaculatorius to carina; carina on left side with a large thorn-like spine; vesica projecting laterally to left, no wider than aedeagus, with a broad, short, subbasal diverticulum; apex of vesica with a similar diverticulum, with a small flat cornutus, from which two very small and short spines arise.
</paragraph>
<caption id="DF55660AFFB9FFF0FF191B17FB162DA2" box="[151,1208,1641,1665]" httpUri="https://zenodo.org/record/177149/files/figure.png" pageId="3" pageNumber="16" targetBox="[151,1433,628,1612]" targetPageId="3">
<paragraph id="8B953682FFB9FFF0FF191B17FB162DA2" blockId="3.[151,1208,1641,1665]" box="[151,1208,1641,1665]" pageId="3" pageNumber="16">
<emphasis id="B95EEA90FFB9FFF0FF191B17FEFF2DA2" bold="true" box="[151,337,1641,1665]" pageId="3" pageNumber="16">FIGURES 810.</emphasis>
Female genitalia of
<emphasis id="B95EEA90FFB9FFF0FDA21B14FD072DA2" box="[556,681,1642,1665]" italics="true" pageId="3" pageNumber="16">
<taxonomicName id="4C2A4D01FFB9FFF0FDA21B14FD0B2DA2" box="[556,677,1642,1665]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="16" phylum="Arthropoda" rank="genus">Margelana</taxonomicName>
.
</emphasis>
8,
<taxonomicName id="4C2A4D01FFB9FFF0FD471B14FCEB2DA2" box="[713,837,1642,1665]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="16" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="B95EEA90FFB9FFF0FD471B14FCEB2DA2" box="[713,837,1642,1665]" italics="true" pageId="3" pageNumber="16">M. brunnea</emphasis>
</taxonomicName>
; 9,
<taxonomicName id="4C2A4D01FFB9FFF0FCE01B14FC402DA2" box="[878,1006,1642,1665]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="16" phylum="Arthropoda" rank="species" species="flavidior">
<emphasis id="B95EEA90FFB9FFF0FCE01B14FC402DA2" box="[878,1006,1642,1665]" italics="true" pageId="3" pageNumber="16">M. flavidior</emphasis>
</taxonomicName>
; 10,
<taxonomicName id="4C2A4D01FFB9FFF0FBAA1B14FB1A2DA2" box="[1060,1204,1642,1665]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="16" phylum="Arthropoda" rank="species" species="versicolor">M. versicolor</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B953682FFB9FFF0FF491BCAFC562CC5" blockId="3.[151,1438,1716,2022]" pageId="3" pageNumber="16">
<emphasis id="B95EEA90FFB9FFF0FF491BCAFE3F2DEE" box="[199,401,1716,1741]" italics="true" pageId="3" pageNumber="16">Female genitalia</emphasis>
(
<figureCitation id="13112A07FFB9FFF0FE2B1BCAFE5F2DED" box="[421,497,1716,1742]" captionStart="FIGURES 8 10" captionStartId="3.[151,270,1641,1665]" captionTargetBox="[151,1433,628,1612]" captionTargetId="figure@3.[151,1436,628,1617]" captionTargetPageId="3" captionText="FIGURES 8 10. Female genitalia of Margelana. 8, M. brunnea; 9, M. flavidior; 10, M. versicolor." httpUri="https://zenodo.org/record/177149/files/figure.png" pageId="3" pageNumber="16">Fig. 8</figureCitation>
). Ovipositor typically apameine-like: heavily sclerotised, dorso-ventrally flattened, clothed with small and long setae, sharply pointed at tip, and with a rounded shoulder i.e., bubble-like pouches laterally at base; two typical apameine sclerotised bars between ovipositor lobes are weekly sclerotised; 8th abdominal segment twice as long ventrally as dorsally; ante-vaginal plate 5 times broader than long; anterior and posterior apophyses stout, apically spatulate, posterior apophyses a little longer; anterior apophyses approximately as long as 8th abdominal segment; ductus bursae relatively short, twice as wide as long; corpus bursae very long, anterior half cylindrical, posterior half ovoid; appendix bursae arising basally on the dorsal side of corpus bursae, ductus seminalis at apex of appendix bursae.
</paragraph>
<caption id="DF55660AFFBEFFF7FF19181EFC872E54" box="[151,809,1376,1400]" httpUri="https://zenodo.org/record/177150/files/figure.png" pageId="4" pageNumber="17" targetBox="[150,1436,194,1350]" targetPageId="4">
<paragraph id="8B953682FFBEFFF7FF19181EFC872E54" blockId="4.[151,809,1376,1400]" box="[151,809,1376,1400]" pageId="4" pageNumber="17">
<emphasis id="B95EEA90FFBEFFF7FF19181EFE8A2E5B" bold="true" box="[151,292,1376,1400]" pageId="4" pageNumber="17">FIGURE 11.</emphasis>
Geographic distribution of
<taxonomicName id="4C2A4D01FFBEFFF7FDC3181FFC8C2E5B" box="[589,802,1377,1400]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="B95EEA90FFBEFFF7FDC3181FFC8C2E5B" box="[589,802,1377,1400]" italics="true" pageId="4" pageNumber="17">Margelana brunnea</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B953682FFBEFFF7FF4818D5FE5A2C5E" blockId="4.[151,1436,1451,2037]" pageId="4" pageNumber="17">
<emphasis id="B95EEA90FFBEFFF7FF4818D5FE772EE6" bold="true" box="[198,473,1451,1477]" pageId="4" pageNumber="17">Differential diagnosis.</emphasis>
<taxonomicName id="4C2A4D01FFBEFFF7FE6E18D5FD642EE7" box="[480,714,1451,1476]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="B95EEA90FFBEFFF7FE6E18D5FD642EE7" box="[480,714,1451,1476]" italics="true" pageId="4" pageNumber="17">Margelana brunnea</emphasis>
</taxonomicName>
is the largest of the three
<taxonomicName id="4C2A4D01FFBEFFF7FC7618D5FBD72EE7" box="[1016,1145,1451,1476]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEA90FFBEFFF7FC7618D5FBD72EE7" box="[1016,1145,1451,1476]" italics="true" pageId="4" pageNumber="17">Margelana</emphasis>
</taxonomicName>
species (
<quantity id="4CD29B67FFBEFFF7FB6A18D5FAC62EE6" box="[1252,1384,1451,1477]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" metricValueMax="4.6" metricValueMin="3.8" pageId="4" pageNumber="17" unit="mm" value="42.0" valueMax="46.0" valueMin="38.0">3846 mm</quantity>
),
<taxonomicName id="4C2A4D01FFBEFFF7FAF218D5FF522ECF" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="flavidior">
<emphasis id="B95EEA90FFBEFFF7FAF218D5FF522ECF" italics="true" pageId="4" pageNumber="17">M. flavidior</emphasis>
</taxonomicName>
(
<figureCitation id="13112A07FFBEFFF7FE8518ADFEFE2ECE" box="[267,336,1491,1517]" captionStart="FIGURES 1 4" captionStartId="1.[151,270,1455,1479]" captionTargetBox="[151,1435,713,1408]" captionTargetId="figure@1.[151,1436,695,1448]" captionTargetPageId="1" captionText="FIGURES 1 4. Adults of Margelana. 1, Holotype male of Margelana brunnea; 2, paratype female of Margelana brunnea; 3, male of M. flavidior; 4, female of M. versicolor." httpUri="https://zenodo.org/record/177147/files/figure.png" pageId="4" pageNumber="17">Fig. 3</figureCitation>
) is middle sized (
<quantity id="4CD29B67FFBEFFF7FD9018ADFD0C2ECE" box="[542,674,1491,1517]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" metricValueMax="3.8" metricValueMin="3.5" pageId="4" pageNumber="17" unit="mm" value="36.5" valueMax="38.0" valueMin="35.0">3538 mm</quantity>
), and
<taxonomicName id="4C2A4D01FFBEFFF7FD6918ADFC2D2ECF" box="[743,899,1491,1516]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="versicolor">
<emphasis id="B95EEA90FFBEFFF7FD6918ADFC2D2ECF" box="[743,899,1491,1516]" italics="true" pageId="4" pageNumber="17">M. versicolor</emphasis>
</taxonomicName>
(
<figureCitation id="13112A07FFBEFFF7FC1A18ADFC742ECE" box="[916,986,1491,1517]" captionStart="FIGURES 1 4" captionStartId="1.[151,270,1455,1479]" captionTargetBox="[151,1435,713,1408]" captionTargetId="figure@1.[151,1436,695,1448]" captionTargetPageId="1" captionText="FIGURES 1 4. Adults of Margelana. 1, Holotype male of Margelana brunnea; 2, paratype female of Margelana brunnea; 3, male of M. flavidior; 4, female of M. versicolor." httpUri="https://zenodo.org/record/177147/files/figure.png" pageId="4" pageNumber="17">Fig. 4</figureCitation>
) is the smallest (
<quantity id="4CD29B67FFBEFFF7FB1318ADFA8C2ECE" box="[1181,1314,1491,1517]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" metricValueMax="3.6" metricValueMin="3.2" pageId="4" pageNumber="17" unit="mm" value="34.0" valueMax="36.0" valueMin="32.0">3236 mm</quantity>
).
<taxonomicName id="4C2A4D01FFBEFFF7FABA18ADFF6C2D37" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="B95EEA90FFBEFFF7FABA18ADFF6C2D37" italics="true" pageId="4" pageNumber="17">M. brunnea</emphasis>
</taxonomicName>
has a brown forewing ground colour with a white pattern; the antemedian line is sharply angled submedially; the postmedian line is slightly bent and smooth; most often the orbicular and reniform stigmata are partially fused; the outer and upper part of the prothoracic collar is white, as are the margins of the tegulae.
<taxonomicName id="4C2A4D01FFBEFFF7FAF21B35FF522DAF" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="flavidior">
<emphasis id="B95EEA90FFBEFFF7FAF21B35FF522DAF" italics="true" pageId="4" pageNumber="17">M. flavidior</emphasis>
</taxonomicName>
has a yellowish ground coloureven subsp.
<emphasis id="B95EEA90FFBEFFF7FCAF1B0DFCDF2DAF" box="[801,881,1651,1676]" italics="true" pageId="4" pageNumber="17">ochrea</emphasis>
is less brownish; the antemedian line is also sharply angled but is usually farther away from the postmedian line; the latter line is almost straight and smooth; the orbicular and reniform stigmata are rarely united; the collar and tegulae are concolorous with the head and thorax. The white-grey hindwing of
<taxonomicName id="4C2A4D01FFBEFFF7FD331B95FC0D2C27" box="[701,931,1771,1796]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="flavidior" subSpecies="ochrea">
<emphasis id="B95EEA90FFBEFFF7FD331B95FC0D2C27" box="[701,931,1771,1796]" italics="true" pageId="4" pageNumber="17">M. flavidior ochrea</emphasis>
</taxonomicName>
separates this from
<taxonomicName id="4C2A4D01FFBEFFF7FB181B95FA8E2C27" box="[1174,1312,1771,1796]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="B95EEA90FFBEFFF7FB181B95FA8E2C27" box="[1174,1312,1771,1796]" italics="true" pageId="4" pageNumber="17">M. brunnea</emphasis>
</taxonomicName>
.
<taxonomicName id="4C2A4D01FFBEFFF7FABE1B95FF7B2C0F" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="versicolor">
<emphasis id="B95EEA90FFBEFFF7FABE1B95FF7B2C0F" italics="true" pageId="4" pageNumber="17">M. versicolor</emphasis>
</taxonomicName>
also has a brown ground colour, but white veins; less sharply angeled antemedian line; slightly bent and jagged postmedian line; separated orbicular and reniform stigmata; and yellowish outer and upper part of the collar and margins of tegulae.
</paragraph>
<paragraph id="8B953682FFBEFFF6FF481AF5FECC2A57" blockId="4.[151,1436,1451,2037]" lastBlockId="5.[151,1437,146,1052]" lastPageId="5" lastPageNumber="18" pageId="4" pageNumber="17">
The male genital armature of all three species is rather similar, but in
<taxonomicName id="4C2A4D01FFBEFFF7FB8B1AF5FB202C87" box="[1029,1166,1931,1956]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="B95EEA90FFBEFFF7FB8B1AF5FB202C87" box="[1029,1166,1931,1956]" italics="true" pageId="4" pageNumber="17">M. brunnea</emphasis>
</taxonomicName>
(
<figureCitation id="13112A07FFBEFFF7FB2E1AF5FB492C86" box="[1184,1255,1931,1957]" captionStart="FIGURES 5 7" captionStartId="2.[151,269,1963,1987]" captionTargetBox="[257,1436,201,1914]" captionTargetId="figure@2.[151,1436,194,1939]" captionTargetPageId="2" captionText="FIGURES 5 7. Male genitalia of Margelana. 5, M. brunnea; 6, M. flavidior; 7, M. versicolor." httpUri="https://zenodo.org/record/177148/files/figure.png" pageId="4" pageNumber="17">Fig. 5</figureCitation>
) the uncus is a little longer and broader than that of
<taxonomicName id="4C2A4D01FFBEFFF7FDCD1ACDFD4F2CEF" box="[579,737,1971,1996]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="versicolor">
<emphasis id="B95EEA90FFBEFFF7FDCD1ACDFD4F2CEF" box="[579,737,1971,1996]" italics="true" pageId="4" pageNumber="17">M. versicolor</emphasis>
</taxonomicName>
(
<figureCitation id="13112A07FFBEFFF7FD7E1ACDFC982CEE" box="[752,822,1971,1997]" captionStart="FIGURES 5 7" captionStartId="2.[151,269,1963,1987]" captionTargetBox="[257,1436,201,1914]" captionTargetId="figure@2.[151,1436,194,1939]" captionTargetPageId="2" captionText="FIGURES 5 7. Male genitalia of Margelana. 5, M. brunnea; 6, M. flavidior; 7, M. versicolor." httpUri="https://zenodo.org/record/177148/files/figure.png" pageId="4" pageNumber="17">Fig. 7</figureCitation>
) and the uncus of
<taxonomicName id="4C2A4D01FFBEFFF7FB831ACDFB362CEF" box="[1037,1176,1971,1996]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="flavidior">
<emphasis id="B95EEA90FFBEFFF7FB831ACDFB362CEF" box="[1037,1176,1971,1996]" italics="true" pageId="4" pageNumber="17">M. flavidior</emphasis>
</taxonomicName>
(
<figureCitation id="13112A07FFBEFFF7FB271ACDFB402CEE" box="[1193,1262,1971,1997]" captionStart="FIGURES 5 7" captionStartId="2.[151,269,1963,1987]" captionTargetBox="[257,1436,201,1914]" captionTargetId="figure@2.[151,1436,194,1939]" captionTargetPageId="2" captionText="FIGURES 5 7. Male genitalia of Margelana. 5, M. brunnea; 6, M. flavidior; 7, M. versicolor." httpUri="https://zenodo.org/record/177148/files/figure.png" pageId="4" pageNumber="17">Fig. 6</figureCitation>
) is shorter and much broader than those of the other two species; the tegumen of
<taxonomicName id="4C2A4D01FFBEFFF7FC2A1AA5FBEC2CD7" box="[932,1090,2011,2036]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="versicolor">
<emphasis id="B95EEA90FFBEFFF7FC2A1AA5FBEC2CD7" box="[932,1090,2011,2036]" italics="true" pageId="4" pageNumber="17">M. versicolor</emphasis>
</taxonomicName>
is narrower than those of the two others; the cucullus of
<taxonomicName id="4C2A4D01FFBFFFF6FE521DECFDD22B88" box="[476,636,146,171]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="versicolor">
<emphasis id="B95EEA90FFBFFFF6FE521DECFDD22B88" box="[476,636,146,171]" italics="true" pageId="5" pageNumber="18">M. versicolor</emphasis>
</taxonomicName>
is the narrowest (
<taxonomicName id="4C2A4D01FFBFFFF6FCDD1DECFC722B88" box="[851,988,146,171]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="B95EEA90FFBFFFF6FCDD1DECFC722B88" box="[851,988,146,171]" italics="true" pageId="5" pageNumber="18">M. brunnea</emphasis>
</taxonomicName>
1.5 times broader;
<taxonomicName id="4C2A4D01FFBFFFF6FB4B1DECFAFD2B88" box="[1221,1363,146,171]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="flavidior">
<emphasis id="B95EEA90FFBFFFF6FB4B1DECFAFD2B88" box="[1221,1363,146,171]" italics="true" pageId="5" pageNumber="18">M. flavidior</emphasis>
</taxonomicName>
is 2.5 times broader); the costal process of
<taxonomicName id="4C2A4D01FFBFFFF6FDC31DC4FD772BF0" box="[589,729,186,211]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="flavidior">
<emphasis id="B95EEA90FFBFFFF6FDC31DC4FD772BF0" box="[589,729,186,211]" italics="true" pageId="5" pageNumber="18">M. flavidior</emphasis>
</taxonomicName>
is more prominent; and the juxta of
<taxonomicName id="4C2A4D01FFBFFFF6FB001DC4FAB62BF0" box="[1166,1304,186,211]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="B95EEA90FFBFFFF6FB001DC4FAB62BF0" box="[1166,1304,186,211]" italics="true" pageId="5" pageNumber="18">M. brunnea</emphasis>
</taxonomicName>
is twice as long as that of
<taxonomicName id="4C2A4D01FFBFFFF6FEC41D9CFE782BD8" box="[330,470,226,251]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="flavidior">
<emphasis id="B95EEA90FFBFFFF6FEC41D9CFE782BD8" box="[330,470,226,251]" italics="true" pageId="5" pageNumber="18">M. flavidior</emphasis>
</taxonomicName>
, and 1.5 times longer than that of
<taxonomicName id="4C2A4D01FFBFFFF6FCE11D9CFBA32BD8" box="[879,1037,226,251]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="versicolor">
<emphasis id="B95EEA90FFBFFFF6FCE11D9CFBA32BD8" box="[879,1037,226,251]" italics="true" pageId="5" pageNumber="18">M. versicolor</emphasis>
</taxonomicName>
. The aedeagus carina spine of
<taxonomicName id="4C2A4D01FFBFFFF6FAF31D9CFF572A00" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="B95EEA90FFBFFFF6FAF31D9CFF572A00" italics="true" pageId="5" pageNumber="18">M. brunnea</emphasis>
</taxonomicName>
(
<figureCitation id="13112A07FFBFFFF6FE871C74FEE12A07" box="[265,335,266,292]" captionStart="FIGURES 5 7" captionStartId="2.[151,269,1963,1987]" captionTargetBox="[257,1436,201,1914]" captionTargetId="figure@2.[151,1436,194,1939]" captionTargetPageId="2" captionText="FIGURES 5 7. Male genitalia of Margelana. 5, M. brunnea; 6, M. flavidior; 7, M. versicolor." httpUri="https://zenodo.org/record/177148/files/figure.png" pageId="5" pageNumber="18">Fig. 5</figureCitation>
) is the largest of the three species, and that of
<taxonomicName id="4C2A4D01FFBFFFF6FCFF1C74FBA12A00" box="[881,1039,266,291]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="versicolor">
<emphasis id="B95EEA90FFBFFFF6FCFF1C74FBA12A00" box="[881,1039,266,291]" italics="true" pageId="5" pageNumber="18">M. versicolor</emphasis>
</taxonomicName>
(
<figureCitation id="13112A07FFBFFFF6FB911C74FBCB2A07" box="[1055,1125,266,292]" captionStart="FIGURES 5 7" captionStartId="2.[151,269,1963,1987]" captionTargetBox="[257,1436,201,1914]" captionTargetId="figure@2.[151,1436,194,1939]" captionTargetPageId="2" captionText="FIGURES 5 7. Male genitalia of Margelana. 5, M. brunnea; 6, M. flavidior; 7, M. versicolor." httpUri="https://zenodo.org/record/177148/files/figure.png" pageId="5" pageNumber="18">Fig. 7</figureCitation>
) is the smallest; the cortunus of
<taxonomicName id="4C2A4D01FFBFFFF6FF691C4CFE2B2A68" box="[231,389,306,331]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="versicolor">
<emphasis id="B95EEA90FFBFFFF6FF691C4CFE2B2A68" box="[231,389,306,331]" italics="true" pageId="5" pageNumber="18">M. versicolor</emphasis>
</taxonomicName>
is the largest, and that of
<taxonomicName id="4C2A4D01FFBFFFF6FD3D1C4CFC952A68" box="[691,827,306,331]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="B95EEA90FFBFFFF6FD3D1C4CFC952A68" box="[691,827,306,331]" italics="true" pageId="5" pageNumber="18">M. brunnea</emphasis>
</taxonomicName>
is the smallest, only slightly smaller than that of
<taxonomicName id="4C2A4D01FFBFFFF6FAF31C4CFF522A50" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="flavidior">
<emphasis id="B95EEA90FFBFFFF6FAF31C4CFF522A50" italics="true" pageId="5" pageNumber="18">M. flavidior</emphasis>
</taxonomicName>
(
<figureCitation id="13112A07FFBFFFF6FE831C24FEFA2A57" box="[269,340,346,372]" captionStart="FIGURES 5 7" captionStartId="2.[151,269,1963,1987]" captionTargetBox="[257,1436,201,1914]" captionTargetId="figure@2.[151,1436,194,1939]" captionTargetPageId="2" captionText="FIGURES 5 7. Male genitalia of Margelana. 5, M. brunnea; 6, M. flavidior; 7, M. versicolor." httpUri="https://zenodo.org/record/177148/files/figure.png" pageId="5" pageNumber="18">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="8B953682FFBFFFF6FF491CFCFAD629AF" blockId="5.[151,1437,146,1052]" pageId="5" pageNumber="18">
The female genitalia differ as follows: the lateral shoulder by the base of the ovipositor is slighty larger in
<taxonomicName id="4C2A4D01FFBFFFF6FF191CD4FE8A2AE0" box="[151,292,426,451]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="flavidior">
<emphasis id="B95EEA90FFBFFFF6FF191CD4FE8A2AE0" box="[151,292,426,451]" italics="true" pageId="5" pageNumber="18">M. flavidior</emphasis>
</taxonomicName>
(
<figureCitation id="13112A07FFBFFFF6FEBB1CD4FED02AE7" box="[309,382,426,452]" captionStart="FIGURES 8 10" captionStartId="3.[151,270,1641,1665]" captionTargetBox="[151,1433,628,1612]" captionTargetId="figure@3.[151,1436,628,1617]" captionTargetPageId="3" captionText="FIGURES 8 10. Female genitalia of Margelana. 8, M. brunnea; 9, M. flavidior; 10, M. versicolor." httpUri="https://zenodo.org/record/177149/files/figure.png" pageId="5" pageNumber="18">Fig. 9</figureCitation>
); the anterior apophysis of
<taxonomicName id="4C2A4D01FFBFFFF6FD4D1CD4FCE22AE0" box="[707,844,426,451]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="B95EEA90FFBFFFF6FD4D1CD4FCE22AE0" box="[707,844,426,451]" italics="true" pageId="5" pageNumber="18">M. brunnea</emphasis>
</taxonomicName>
(
<figureCitation id="13112A07FFBFFFF6FCD31CD4FC0A2AE7" box="[861,932,426,452]" captionStart="FIGURES 8 10" captionStartId="3.[151,270,1641,1665]" captionTargetBox="[151,1433,628,1612]" captionTargetId="figure@3.[151,1436,628,1617]" captionTargetPageId="3" captionText="FIGURES 8 10. Female genitalia of Margelana. 8, M. brunnea; 9, M. flavidior; 10, M. versicolor." httpUri="https://zenodo.org/record/177149/files/figure.png" pageId="5" pageNumber="18">Fig. 8</figureCitation>
) is shorter and apically broader than those of
<taxonomicName id="4C2A4D01FFBFFFF6FF361CACFEEA2AC8" box="[184,324,466,491]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="flavidior">
<emphasis id="B95EEA90FFBFFFF6FF361CACFEEA2AC8" box="[184,324,466,491]" italics="true" pageId="5" pageNumber="18">M. flavidior</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2A4D01FFBFFFF6FEF11CACFDB02AC8" box="[383,542,466,491]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="versicolor">
<emphasis id="B95EEA90FFBFFFF6FEF11CACFDB02AC8" box="[383,542,466,491]" italics="true" pageId="5" pageNumber="18">M. versicolor</emphasis>
</taxonomicName>
(
<figureCitation id="13112A07FFBFFFF6FDA01CACFD282ACF" box="[558,646,466,492]" captionStart="FIGURES 8 10" captionStartId="3.[151,270,1641,1665]" captionTargetBox="[151,1433,628,1612]" captionTargetId="figure@3.[151,1436,628,1617]" captionTargetPageId="3" captionText="FIGURES 8 10. Female genitalia of Margelana. 8, M. brunnea; 9, M. flavidior; 10, M. versicolor." httpUri="https://zenodo.org/record/177149/files/figure.png" pageId="5" pageNumber="18">Fig. 10</figureCitation>
); the 8th abdominal segment of
<taxonomicName id="4C2A4D01FFBFFFF6FB8A1CACFB202AC8" box="[1028,1166,466,491]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="B95EEA90FFBFFFF6FB8A1CACFB202AC8" box="[1028,1166,466,491]" italics="true" pageId="5" pageNumber="18">M. brunnea</emphasis>
</taxonomicName>
is longer than those of the two other species; the ventral ostium plate is broadest in
<taxonomicName id="4C2A4D01FFBFFFF6FCEA1C84FC5F2930" box="[868,1009,506,531]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="flavidior">
<emphasis id="B95EEA90FFBFFFF6FCEA1C84FC5F2930" box="[868,1009,506,531]" italics="true" pageId="5" pageNumber="18">M. flavidior</emphasis>
</taxonomicName>
; the length of the sclerotised ductus bursae of
<taxonomicName id="4C2A4D01FFBFFFF6FE821F5CFE3B2918" box="[268,405,546,571]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="B95EEA90FFBFFFF6FE821F5CFE3B2918" box="[268,405,546,571]" italics="true" pageId="5" pageNumber="18">M. brunnea</emphasis>
</taxonomicName>
is only 50% of that of
<taxonomicName id="4C2A4D01FFBFFFF6FD281F5CFC9C2918" box="[678,818,546,571]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="flavidior">
<emphasis id="B95EEA90FFBFFFF6FD281F5CFC9C2918" box="[678,818,546,571]" italics="true" pageId="5" pageNumber="18">M. flavidior</emphasis>
</taxonomicName>
, and that of
<taxonomicName id="4C2A4D01FFBFFFF6FC491F5CFBC82918" box="[967,1126,546,571]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="versicolor">
<emphasis id="B95EEA90FFBFFFF6FC491F5CFBC82918" box="[967,1126,546,571]" italics="true" pageId="5" pageNumber="18">M. versicolor</emphasis>
</taxonomicName>
is 60%; the corpus bursae of
<taxonomicName id="4C2A4D01FFBFFFF6FF391F34FEEE2940" box="[183,320,586,611]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="B95EEA90FFBFFFF6FF391F34FEEE2940" box="[183,320,586,611]" italics="true" pageId="5" pageNumber="18">M. brunnea</emphasis>
</taxonomicName>
is twice as long as that of
<taxonomicName id="4C2A4D01FFBFFFF6FDF81F34FCAC2940" box="[630,770,586,611]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="flavidior">
<emphasis id="B95EEA90FFBFFFF6FDF81F34FCAC2940" box="[630,770,586,611]" italics="true" pageId="5" pageNumber="18">M. flavidior</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2A4D01FFBFFFF6FCB51F34FC772940" box="[827,985,586,611]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="versicolor">
<emphasis id="B95EEA90FFBFFFF6FCB51F34FC772940" box="[827,985,586,611]" italics="true" pageId="5" pageNumber="18">M. versicolor</emphasis>
</taxonomicName>
; and the distal part of corpus bursae is ovoid in
<taxonomicName id="4C2A4D01FFBFFFF6FE8E1F0CFE2629A8" box="[256,392,626,651]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="B95EEA90FFBFFFF6FE8E1F0CFE2629A8" box="[256,392,626,651]" italics="true" pageId="5" pageNumber="18">M. brunnea</emphasis>
</taxonomicName>
, whereas that of
<taxonomicName id="4C2A4D01FFBFFFF6FDDF1F0CFD7329A8" box="[593,733,626,651]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="flavidior">
<emphasis id="B95EEA90FFBFFFF6FDDF1F0CFD7329A8" box="[593,733,626,651]" italics="true" pageId="5" pageNumber="18">M. flavidior</emphasis>
</taxonomicName>
is globular, and that of
<taxonomicName id="4C2A4D01FFBFFFF6FC7D1F0CFB3F29A8" box="[1011,1169,626,651]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="versicolor">
<emphasis id="B95EEA90FFBFFFF6FC7D1F0CFB3F29A8" box="[1011,1169,626,651]" italics="true" pageId="5" pageNumber="18">M. versicolor</emphasis>
</taxonomicName>
is teardrop-shaped.
</paragraph>
</subSubSection>
<subSubSection id="C3306509FFBFFFF6FF491FE7FEA228EF" pageId="5" pageNumber="18" type="distribution">
<paragraph id="8B953682FFBFFFF6FF491FE7FEA228EF" blockId="5.[151,1437,146,1052]" pageId="5" pageNumber="18">
<emphasis id="B95EEA90FFBFFFF6FF491FE7FDA82990" bold="true" box="[199,518,665,691]" pageId="5" pageNumber="18">Distribution and habitat.</emphasis>
All specimens of
<taxonomicName id="4C2A4D01FFBFFFF6FD621FE4FCD62990" box="[748,888,666,691]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="B95EEA90FFBFFFF6FD621FE4FCD62990" box="[748,888,666,691]" italics="true" pageId="5" pageNumber="18">M. brunnea</emphasis>
</taxonomicName>
are from Golestan National Park, Almeh, in northeastern
<collectingCountry id="F33D7612FFBFFFF6FEBE1FBCFECC29FF" box="[304,354,706,732]" name="Iran" pageId="5" pageNumber="18">Iran</collectingCountry>
, between
<quantity id="4CD29B67FFBFFFF6FE591FBCFDD729FF" box="[471,633,706,732]" metricMagnitude="3" metricUnit="m" metricValue="1.675" metricValueMax="1.75" metricValueMin="1.6" pageId="5" pageNumber="18" unit="m" value="1675.0" valueMax="1750.0" valueMin="1600.0">16001750 m</quantity>
(
<figureCitation id="13112A07FFBFFFF6FD041FBCFD4E29FF" box="[650,736,706,732]" captionStart="FIGURE 11" captionStartId="4.[151,255,1376,1400]" captionTargetBox="[150,1436,194,1350]" captionTargetId="figure@4.[147,1439,194,1352]" captionTargetPageId="4" captionText="FIGURE 11. Geographic distribution of Margelana brunnea." httpUri="https://zenodo.org/record/177150/files/figure.png" pageId="5" pageNumber="18">Fig. 11</figureCitation>
). Golestan National Park was the first national park in
<collectingCountry id="F33D7612FFBFFFF6FAE81FBCFA3629FF" box="[1382,1432,706,732]" name="Iran" pageId="5" pageNumber="18">Iran</collectingCountry>
, covering about 91,000 hectares. The terrain is montainous with altitudes varying from
<quantity id="4CD29B67FFBFFFF6FB301F94FACE2827" box="[1214,1376,746,772]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="2.82" metricValueMin="0.38" pageId="5" pageNumber="18" unit="m" value="1600.0" valueMax="2820.0" valueMin="380.0">3802,820 m</quantity>
. The Almeh Region is located in the southeastern part of the park and consists of two mountain ranges separated by a valley. The habitat is dominated by open areas with oak (
<taxonomicName id="4C2A4D01FFBFFFF6FCC51E44FC032870" box="[843,941,826,851]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B95EEA90FFBFFFF6FCC51E44FC032870" box="[843,941,826,851]" italics="true" pageId="5" pageNumber="18">Quercus</emphasis>
</taxonomicName>
),
<taxonomicName id="4C2A4D01FFBFFFF6FC481E44FB162870" box="[966,1208,826,851]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="18" phylum="Tracheophyta" rank="species" species="communis">
<emphasis id="B95EEA90FFBFFFF6FC481E44FB162870" box="[966,1208,826,851]" italics="true" pageId="5" pageNumber="18">Juniperus communis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2A4D01FFBFFFF6FB7F1E44FACD2870" box="[1265,1379,826,851]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="18" phylum="Tracheophyta" rank="species" species="excelsa">
<emphasis id="B95EEA90FFBFFFF6FB7F1E44FACD2870" box="[1265,1379,826,851]" italics="true" pageId="5" pageNumber="18">J. excelsa</emphasis>
</taxonomicName>
, and with various herbaceous plants and shrubs, including
<taxonomicName id="4C2A4D01FFBFFFF6FC941E1CFCD12858" box="[794,895,866,891]" class="Magnoliopsida" family="Berberidaceae" genus="Berberis" kingdom="Plantae" order="Ranunculales" pageId="5" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B95EEA90FFBFFFF6FC941E1CFCD12858" box="[794,895,866,891]" italics="true" pageId="5" pageNumber="18">Berberis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A4D01FFBFFFF6FC1E1E1CFBA42858" box="[912,1034,866,891]" class="Magnoliopsida" family="Rosaceae" genus="Crataegus" kingdom="Plantae" order="Rosales" pageId="5" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B95EEA90FFBFFFF6FC1E1E1CFBA42858" box="[912,1034,866,891]" italics="true" pageId="5" pageNumber="18">Crataegus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C2A4D01FFBFFFF6FBC01E1CFB662858" box="[1102,1224,866,891]" class="Magnoliopsida" family="Amaranthaceae" genus="Haloxylon" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B95EEA90FFBFFFF6FBC01E1CFB662858" box="[1102,1224,866,891]" italics="true" pageId="5" pageNumber="18">Haloxylon</emphasis>
</taxonomicName>
. All of the specimens of
<taxonomicName id="4C2A4D01FFBFFFF6FE891EF4FE3A2880" box="[263,404,906,931]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="B95EEA90FFBFFFF6FE891EF4FE3A2880" box="[263,404,906,931]" italics="true" pageId="5" pageNumber="18">M. brunnea</emphasis>
</taxonomicName>
were collected at light from late September to early October. The early stages are unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3306509FFBFFFF6FF491EA7FE532F3F" pageId="5" pageNumber="18" type="etymology">
<paragraph id="8B953682FFBFFFF6FF491EA7FE532F3F" blockId="5.[151,1437,146,1052]" pageId="5" pageNumber="18">
<emphasis id="B95EEA90FFBFFFF6FF491EA7FEE228D0" bold="true" box="[199,332,985,1011]" pageId="5" pageNumber="18">Etymology</emphasis>
. The name of the new species refers to the brown forewing ground colour, and is derived from the Latin for brown (
<taxonomicName id="4C2A4D01FFBFFFF6FE02197CFE5F2F3F" box="[396,497,1026,1052]" class="Insecta" family="Noctuidae" genus="Margelana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="brunnea">brunnea</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>