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<document id="B4489E9A308D6B1377AFD11718E6BBF0" ID-CLB-Dataset="8932" ID-DOI="10.11646/zootaxa.4821.2.3" ID-GBIF-Dataset="02e15568-9697-42ad-af97-54abb128a40f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4440916" ID-ZooBank="F0EBDD5F-5DBE-47A9-B946-881DF3A21BA1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609217280184" checkinUser="plazi" docAuthor="Jiang, Ri-Xin, Bai, Xing-Long, Ren, Guo-Dong, Yang, Hui-Yu &amp; Wang, Shuo" docDate="2020" docId="038382170D4970392FBD46EDFCA7FF6A" docLanguage="en" docName="zootaxa.4821.2.3.pdf" docOrigin="Zootaxa 4821 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hexarhopalus (Hexarhopalus) mangshanicus , Ren &amp; xu 2011" docType="treatment" docVersion="9" lastPageNumber="286" masterDocId="FFBAFA6F0D4170302F2A4056FFBBFFBF" masterDocTitle="A taxonomic revision of the genus Hexarhopalus (Coleoptera, Tenebrionidae Cnodalonini) from China" masterLastPageNumber="304" masterPageNumber="277" pageNumber="285" updateTime="1698848438170" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6BC08CE12E45F8B763A7584513AB196B">A taxonomic revision of the genus Hexarhopalus (Coleoptera, Tenebrionidae Cnodalonini) from China</mods:title>
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<mods:namePart id="CF5D65DCE6483C63210940B89833DF0E">Jiang, Ri-Xin</mods:namePart>
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<mods:namePart id="586E49FD3784B9D984CB41754C0D4771">Bai, Xing-Long</mods:namePart>
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<mods:affiliation id="0E225E7E4A494A0AD21EDA82AAA1680A">College of Life Science, Institute of Life Science and Green Development, Hebei University, Baoding, Hebei, 071002, P. R. China Corresponding author. &amp; bxlngg @ 163. com; https: // orcid. org / 0000 - 0002 - 0448 - 8272 &amp; gdren @ hbu. edu. cn; https: // orcid. org / 0000 - 0001 - 5808 - 9122</mods:affiliation>
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<treatment id="038382170D4970392FBD46EDFCA7FF6A" ID-DOI="http://doi.org/10.5281/zenodo.4441052" ID-GBIF-Taxon="176580101" ID-Zenodo-Dep="4441052" LSID="urn:lsid:plazi:treatment:038382170D4970392FBD46EDFCA7FF6A" httpUri="http://treatment.plazi.org/id/038382170D4970392FBD46EDFCA7FF6A" lastPageId="9" lastPageNumber="286" pageId="8" pageNumber="285">
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<heading id="D0DD846D0D4970382FBD46EDFC2AF96A" bold="true" box="[151,913,1723,1750]" fontSize="11" level="1" pageId="8" pageNumber="285" reason="1">
<taxonomicName id="4C2A48820D4970382FBD46EDFC2AF96A" authority="Ren &amp; Xu, 2011" authorityName=", Ren &amp; xu" authorityYear="2011" box="[151,913,1723,1750]" class="Insecta" family="Tenebrionidae" genus="Hexarhopalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="285" phylum="Arthropoda" rank="species" species="mangshanicus" subGenus="Hexarhopalus">
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<emphasis id="B95EEF130D4970382FBD46EDFD78F969" bold="true" box="[151,707,1723,1750]" italics="true" pageId="8" pageNumber="285">Hexarhopalus (Hexarhopalus) mangshanicus</emphasis>
<bibRefCitation id="EFBB4EF00D4970382DE046EAFC2AF96A" author="Ren, G. - D. &amp; Xu, J. - S." box="[714,913,1723,1750]" pageId="8" pageNumber="285" pagination="23 - 34" refId="ref11188" refString="Ren, G. - D. &amp; Xu, J. - S. (2011) The genus Hexarhopalus Fairmaire, 1891 in China, with description of three new species (Coleoptera, Tenebrionidae, Cnodalonini), Acta Zoologica Academiae Scientiarum Hungaricae, 57, 23 - 34." type="journal article" year="2011">Ren &amp; Xu, 2011</bibRefCitation>
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<paragraph id="8B9533010D4970382FBD4689FE72F946" blockId="8.[151,913,1723,1785]" box="[151,457,1759,1785]" pageId="8" pageNumber="285">
<heading id="D0DD846D0D4970382FBD4689FE72F946" bold="true" box="[151,457,1759,1785]" fontSize="11" level="1" pageId="8" pageNumber="285" reason="1">
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(
<figureCitation id="13112F840D4970382FB54689FEBEF946" box="[159,261,1759,1785]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStartId-0="9.[151,250,1464,1488]" captionStartId-1="10.[151,250,1208,1232]" captionTargetBox-0="[233,1349,243,1435]" captionTargetBox-1="[177,1406,237,1179]" captionTargetId-0="figure-40@9.[225,1363,237,1441]" captionTargetId-1="figure-53@10.[177,1411,237,1184]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURE 8. Habitus of Hexarhopalus (H.) mangshanicus. A. Male; B. Female." captionText-1="FIGURE 9. Diagnostic features of Hexarhopalus (H.) mangshanicus (AG. Male; HI. Female). A. Head and pronotum, in dorsal view; B. Antenna; C. Aedeagus, ventral view; D. Same, lateral view; E. Same, dorsal view; F. Spiculum gastrale; G. Abdominal ventrite VIII of male; H. Ventrite VIII of female and spiculum; I. Ovipositor." figureDoi-0="http://doi.org/10.5281/zenodo.4440936" figureDoi-1="http://doi.org/10.5281/zenodo.4440938" httpUri-0="https://zenodo.org/record/4440936/files/figure.png" httpUri-1="https://zenodo.org/record/4440938/files/figure.png" pageId="8" pageNumber="285">Figs 89</figureCitation>
,
<figureCitation id="13112F840D4970382E3B4689FEF8F946" box="[273,323,1759,1785]" captionStart="FIGURE 20" captionStartId="22.[151,250,1872,1896]" captionTargetBox="[210,1371,351,1840]" captionTargetId="figure-58@22.[201,1386,345,1848]" captionTargetPageId="22" captionText="FIGURE 20. Types of Chinese Hexarhopalus species.A. Hexarhopalus (H.) mangshanicus, Ren &amp; xu, 2011, holotype, male; B. Hexarhopalus (L.) bisinuatus, Ren &amp; xu, 2011, holotype, male; C. Hexarhopalus (L.) xui, Ren &amp; xu, 2011, holotype, female; D. Hexarhopalus (H.) attenuatus (Pic, 1922), paralectotype; E. Same, in lateral view (20AC deposited in MHBU; 20D deposited in HNHM; figs 20DE were taken by Tamás Németh)." figureDoi="http://doi.org/10.5281/zenodo.4440980" httpUri="https://zenodo.org/record/4440980/files/figure.png" pageId="8" pageNumber="285">20A</figureCitation>
,
<figureCitation id="13112F840D4970382E7A4689FE3AF946" box="[336,385,1759,1785]" captionStart="FIGURE 21" captionStartId="23.[151,250,1041,1065]" captionTargetBox="[206,1381,186,1013]" captionTargetId="figure-18@23.[201,1386,181,1018]" captionTargetPageId="23" captionText="FIGURE 21. Lateral groove of pronotum of Chinese Hexarhopalus species (not include Hexarhopalus (L.) xui). A. Hexarhopalus (H.) attenuatus; B. Hexarhopalus (H.) liuyixiaoi sp. nov.; C. H. (H.) sculptilis; D. Hexarhopalus (H.) mangshanicus; E. H. (H.) sculpticollis; F. H. (L.) qiujianyueae sp. nov.; G. Hexarhopalus (L.) bisinuatus; H. Hexarhopalus (L.) difformis; I. Hexarhopalus (L.) qiului sp. nov.; J. Hexarhopalus (L.) xuhaoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4440984" httpUri="https://zenodo.org/record/4440984/files/figure.png" pageId="8" pageNumber="285">21D</figureCitation>
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<figureCitation id="13112F840D4970382EA44689FE7AF946" box="[398,449,1759,1785]" captionStart="FIGURE 22" captionStartId="24.[151,250,1733,1757]" captionTargetBox="[201,1386,309,1710]" captionTargetId="figure-78@24.[201,1386,309,1710]" captionTargetPageId="24" captionText="FIGURE 22. Habitat of Chinese Hexarhopalus species. A. Mr. Yi-Xiao Liu working at night in the type locality of Hexarhopalus (H.) liuyixiaoi sp. nov.: Fushou Mountain (Pingjiang County, Hunan Province); B. General environment of Fushou Mountain; CD. Living specimen of Hexarhopalus (H.) liuyixiaoi sp. nov. climbing on a tree at night; E. Type locality of Hexarhopalus (L.) qiujianyueae sp. nov.: Dawei Mountain (Pingbian Miao Autonomous County, Yunnan Province); F. Living specimen of Hexarhopalus (H.) sculptilis in a rotten wood; G. Microbiotop of Hexarhopalus (H.) sculptilis; H. Living specimen of Hexarhopalus (H.) mangshanicus, at night; I. Microbiotop of Hexarhopalus (L.) difformis; J. Living specimen of Hexarhopalus (H.) difformis; K. Living specimen of Hexarhopalus (L.) qiului sp. nov. (Figs 22EG, 22IJ taken by Dr. Hao Xu; Fig. 22H taken by Mr. Xiang Zhu; Fig. 22K taken by Dr. Lu Qiu)." figureDoi="http://doi.org/10.5281/zenodo.4440989" httpUri="https://zenodo.org/record/4440989/files/figure.png" pageId="8" pageNumber="285">22H</figureCitation>
)
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<taxonomicName id="4C2A48820D4970382FBD4772FCAFF883" authority="Ren &amp; Xu, 2011" authorityName=", Ren &amp; xu" authorityYear="2011" box="[151,788,1828,1852]" class="Insecta" family="Tenebrionidae" genus="Hexarhopalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="285" phylum="Arthropoda" rank="species" species="mangshanicus" subGenus="Hexarhopalus">
<emphasis id="B95EEF130D4970382FBD4772FE9DF883" box="[151,294,1828,1852]" italics="true" pageId="8" pageNumber="285">Hexarhopalus</emphasis>
(
<emphasis id="B95EEF130D4970382E1F4772FE7FF883" box="[309,452,1828,1852]" italics="true" pageId="8" pageNumber="285">Hexarhopalus</emphasis>
)
<emphasis id="B95EEF130D4970382EF94772FDDFF883" box="[467,612,1828,1852]" italics="true" pageId="8" pageNumber="285">mangshanicus</emphasis>
<materialsCitation id="3B42395C0D4970382D404772FCAFF883" ID-GBIF-Occurrence="3015210332" box="[618,788,1828,1852]" country="China" location="Mangshan" pageId="8" pageNumber="285" specimenCount="1" stateProvince="Hunan">
<bibRefCitation id="EFBB4EF00D4970382D404772FCAFF883" author="Ren, G. - D. &amp; Xu, J. - S." box="[618,788,1828,1852]" pageId="8" pageNumber="285" pagination="23 - 34" refId="ref11188" refString="Ren, G. - D. &amp; Xu, J. - S. (2011) The genus Hexarhopalus Fairmaire, 1891 in China, with description of three new species (Coleoptera, Tenebrionidae, Cnodalonini), Acta Zoologica Academiae Scientiarum Hungaricae, 57, 23 - 34." type="journal article" year="2011">Ren &amp; Xu, 2011</bibRefCitation>
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.
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locality:
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,
<collectingRegion id="49EEFDE30D4970382CD24772FB85F884" box="[1016,1086,1828,1851]" country="China" name="Hunan" pageId="8" pageNumber="285">Hunan</collectingRegion>
,
<location id="8EF565DA0D4970382B624772FB0EF883" LSID="urn:lsid:plazi:treatment:038382170D4970392FBD46EDFCA7FF6A:8EF565DA0D4970382B624772FB0EF883" box="[1096,1205,1828,1852]" country="China" name="Mangshan" pageId="8" pageNumber="285" stateProvince="Hunan">Mangshan</location>
.
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Material examined (
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,
<specimenCount id="9D2CF8880D4970382ED0473CFDADF83B" box="[506,534,1898,1924]" pageId="8" pageNumber="285" type="male">19</specimenCount>
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♂♂,
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<emphasis id="B95EEF130D4970382D72473CFDDDF83B" bold="true" box="[600,614,1898,1924]" pageId="8" pageNumber="285">4</emphasis>
♀♀
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<emphasis id="B95EEF130D4970382DB2473CFD13F83B" bold="true" box="[664,680,1898,1924]" pageId="8" pageNumber="285">).</emphasis>
<typeStatus id="54918DA30D4970382D9A473CFCA6F83B" box="[688,797,1898,1924]" pageId="8" pageNumber="285" type="holotype">Holotype</typeStatus>
:
<materialsCitation id="3B42395C0D4970382C03473CFDCCF817" ID-GBIF-Occurrence="3015210309" collectionCode="MHBU" country="China" elevation="800" location="Mangshan" pageId="8" pageNumber="285" specimenCount="1" specimenCount-male="1" stateProvince="Hunan" typeStatus="holotype">
<specimenCount id="9D2CF8880D4970382C03473CFCEDF83A" box="[809,854,1898,1925]" pageId="8" pageNumber="285" type="male">
<emphasis id="B95EEF130D4970382C03473CFC8CF83B" bold="true" box="[809,823,1898,1924]" pageId="8" pageNumber="285">1</emphasis>
</specimenCount>
,
<collectingCountry id="F33D73910D4970382C49473CFC7BF83B" box="[867,960,1898,1924]" name="China" pageId="8" pageNumber="285">CHINA</collectingCountry>
:
<collectingRegion id="49EEFDE30D4970382CE6473DFBA0F83A" box="[972,1051,1899,1925]" country="China" name="Hunan" pageId="8" pageNumber="285">Hunan</collectingRegion>
: labeles:
<location id="8EF565DA0D4970382BA3473CFABEF83B" LSID="urn:lsid:plazi:treatment:038382170D4970392FBD46EDFCA7FF6A:8EF565DA0D4970382BA3473CFABEF83B" box="[1161,1285,1898,1924]" country="China" name="Mangshan" pageId="8" pageNumber="285" stateProvince="Hunan">Mangshan</location>
,
<quantity id="4CD29EE40D4970382A38473CFAE7F83A" box="[1298,1372,1898,1925]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="285" unit="m" value="800.0">
<elevation id="0007D4320D4970382A38473CFAE7F83A" box="[1298,1372,1898,1925]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="285" unit="m" value="800.0">800 m</elevation>
</quantity>
, Sep-tember 1960, collector unknown (
<collectionCode id="ED3BABC40D4970382D3C47D8FDD4F817" box="[534,623,1934,1960]" pageId="8" pageNumber="285">MHBU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B42395C0D4970382DA847D8FAA7F84E" ID-GBIF-Occurrence="3015210318" collectingDate="2019-06-04" collectingDateMax="2019-06-06" collectingDateMin="2019-06-04" collectionCode="QUST, MYNU" country="China" county="Yangshan County" elevation="1600" latitude="24.922966" location="Guangdongdiyifeng" longLatPrecision="1" longitude="112.98771" municipality="Qingyuan City" pageId="8" pageNumber="285" specimenCount="44" specimenCount-female="8" specimenCount-male="36" stateProvince="Guangdong">
Other material examined:
<specimenCount id="9D2CF8880D4970382C8F47D8FC4AF816" box="[933,1009,1934,1961]" pageId="8" pageNumber="285" type="male">
<emphasis id="B95EEF130D4970382C8F47D8FC7BF817" bold="true" box="[933,960,1934,1960]" pageId="8" pageNumber="285">18</emphasis>
♂♂
</specimenCount>
,
<specimenCount id="9D2CF8880D4970382B2B47D8FBFAF816" box="[1025,1089,1934,1961]" pageId="8" pageNumber="285" type="female">
<emphasis id="B95EEF130D4970382B2B47D8FBB4F817" bold="true" box="[1025,1039,1934,1960]" pageId="8" pageNumber="285">4</emphasis>
♀♀
</specimenCount>
,
<collectingCountry id="F33D73910D4970382B6547D8FB11F817" box="[1103,1194,1934,1960]" name="China" pageId="8" pageNumber="285">CHINA</collectingCountry>
:
<collectingRegion id="49EEFDE30D4970382B9C47D8FA81F817" box="[1206,1338,1934,1960]" country="China" name="Guangdong" pageId="8" pageNumber="285">Guangdong</collectingRegion>
: labeled:
<collectingRegion id="49EEFDE30D4970382FB147E4FE98F873" box="[155,291,1970,1996]" country="China" name="Guangdong" pageId="8" pageNumber="285">Guangdong</collectingRegion>
,
<collectingMunicipality id="6BF1A97B0D4970382E0147E4FE70F873" box="[299,459,1970,1996]" pageId="8" pageNumber="285">Qingyuan City</collectingMunicipality>
(
<emphasis id="B95EEF130D4970382EF247E5FD97F875" bold="true" box="[472,556,1971,1994]" pageId="8" pageNumber="285">Ȉẑḿ</emphasis>
),
<collectingCounty id="62F44B8D0D4970382D1547E4FD44F873" box="[575,767,1970,1996]" pageId="8" pageNumber="285">Yangshan County</collectingCounty>
(
<emphasis id="B95EEF130D4970382C2647E5FCE4F875" bold="true" box="[780,863,1971,1994]" pageId="8" pageNumber="285">ṞƜss</emphasis>
),
<location id="8EF565DA0D4970382C5947E4FBF5F873" LSID="urn:lsid:plazi:treatment:038382170D4970392FBD46EDFCA7FF6A:8EF565DA0D4970382C5947E4FBF5F873" box="[883,1102,1970,1996]" country="China" county="Yangshan County" latitude="24.922966" longLatPrecision="1" longitude="112.98771" municipality="Qingyuan City" name="Guangdongdiyifeng" pageId="8" pageNumber="285" stateProvince="Guangdong">Guangdongdiyifeng</location>
(
<emphasis id="B95EEF130D4970382B7647E5FB5DF875" bold="true" box="[1116,1254,1971,1994]" pageId="8" pageNumber="285">ḞṪë̃ḍ</emphasis>
),
<geoCoordinate id="EE1E55C60D4970382BD147E4FF11F84E" degrees="112" direction="east" minutes="59" orientation="longitude" pageId="8" pageNumber="285" precision="1" seconds="15.76" value="112.98771">112°5915.76” E</geoCoordinate>
,
<geoCoordinate id="EE1E55C60D4970382F9F4780FEDCF84E" box="[181,359,2006,2033]" degrees="24" direction="north" minutes="55" orientation="latitude" pageId="8" pageNumber="285" precision="1" seconds="22.68" value="24.922966">24°5522.68” N</geoCoordinate>
, H:
<quantity id="4CD29EE40D4970382EBE4780FE58F84F" box="[404,483,2006,2032]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="285" unit="m" value="1600.0">
<elevation id="0007D4320D4970382EBE4780FE58F84F" box="[404,483,2006,2032]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="285" unit="m" value="1600.0">1600m</elevation>
</quantity>
,
<date id="FF9415C10D4970382EC44780FDC0F84F" box="[494,635,2006,2032]" pageId="8" pageNumber="285" value="2019-06-04" valueMax="2019-06-06" valueMin="2019-06-04">
<collectingDate id="EFD0EC290D4970382EC44780FDC0F84F" box="[494,635,2006,2032]" pageId="8" pageNumber="285" value="2019-06-04" valueMax="2019-06-06" valueMin="2019-06-04">2019.VI.46</collectingDate>
</date>
, Qiu Jian-Yue leg (
<specimenCount id="9D2CF8880D4970382C7C4780FCC4F84E" box="[854,895,2006,2033]" pageId="8" pageNumber="285" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D2CF8880D4970382CA14780FC0EF84E" box="[907,949,2006,2033]" pageId="8" pageNumber="285" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED3BABC40D4970382CE84780FBB7F84F" box="[962,1036,2006,2032]" pageId="8" pageNumber="285">QUST</collectionCode>
;
<specimenCount id="9D2CF8880D4970382B3C4780FBDEF84E" box="[1046,1125,2006,2033]" pageId="8" pageNumber="285" type="male">17 ♂♂</specimenCount>
,
<specimenCount id="9D2CF8880D4970382B454780FB15F84E" box="[1135,1198,2006,2033]" pageId="8" pageNumber="285" type="female">3 ♀♀</specimenCount>
,
<collectionCode id="ED3BABC40D4970382B914781FAAEF84E" box="[1211,1301,2007,2033]" pageId="8" pageNumber="285">MYNU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C330608A0D4870392FED40C1FCA7FF6A" pageId="9" pageNumber="286" type="discussion">
<paragraph id="8B9533010D4870392FED40C1FCA7FF6A" blockId="9.[151,1437,151,213]" pageId="9" pageNumber="286">
<emphasis id="B95EEF130D4870392FED40C1FE86FF0E" bold="true" box="[199,317,151,177]" pageId="9" pageNumber="286">Remarks:</emphasis>
<taxonomicName id="4C2A48820D4870392E6240C1FD72FF0E" authorityName=", Ren &amp; xu" authorityYear="2011" box="[328,713,151,177]" class="Insecta" family="Tenebrionidae" genus="Hexarhopalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="286" phylum="Arthropoda" rank="species" species="mangshanicus" subGenus="Hexarhopalus">
<emphasis id="B95EEF130D4870392E6240C1FE5DFF0E" box="[328,486,151,177]" italics="true" pageId="9" pageNumber="286">Hexarhopalus</emphasis>
(
<emphasis id="B95EEF130D4870392ED040C1FDADFF0F" box="[506,534,151,176]" italics="true" pageId="9" pageNumber="286">H.</emphasis>
)
<emphasis id="B95EEF130D4870392D0340C1FD72FF0E" box="[553,713,151,177]" italics="true" pageId="9" pageNumber="286">mangshanicus</emphasis>
</taxonomicName>
was originally described from
<collectingRegion id="49EEFDE30D4870392B1240C1FB48FF0E" box="[1080,1267,151,177]" country="China" name="Hunan" pageId="9" pageNumber="286">Hunan Province</collectingRegion>
,
<collectingCountry id="F33D73910D4870392A2840C1FAF3FF0E" box="[1282,1352,151,177]" name="China" pageId="9" pageNumber="286">China</collectingCountry>
, and is herein newly recorded from
<collectingRegion id="49EEFDE30D4870392EFC40EDFD7DFF6A" box="[470,710,187,213]" country="China" name="Guangdong" pageId="9" pageNumber="286">Guangdong Province</collectingRegion>
,
<collectingCountry id="F33D73910D4870392DFB40EDFCA3FF6A" box="[721,792,187,213]" name="China" pageId="9" pageNumber="286">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>