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<document id="1A2D8179AAC5628661FD940EBF4767C9" ID-DOI="10.5281/zenodo.191157" ID-GBIF-Dataset="3709de53-c30a-4ec4-9676-30edc0511523" ID-ISSN="1175-5326" ID-Zenodo-Dep="191157" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461124408753" checkinUser="plazi" docAuthor="Kånneby, Tobias, Todaro, Antonio &amp; Jondelius, Ulf" docDate="2009" docId="03834469FFB50318FF02F8DEF46DF839" docLanguage="en" docName="zt02278p046.pdf" docOrigin="Zootaxa 2278" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ichthydium skandicum Kånneby, Todaro &amp; Jondelius, 2009, n. sp." docType="treatment" docVersion="8" lastPageNumber="30" masterDocId="FFBA3C11FFB4031CFF95FFECF10FFF81" masterDocTitle="One new species and records of Ichthydium Ehrenberg, 1830 (Gastrotricha: Chaetonotida) from Sweden with a key to the genus" masterLastPageNumber="46" masterPageNumber="26" pageNumber="27" updateTime="1698596439301" updateUser="plazi">
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<mods:title id="1831B4569EABDCA45FF1B362F8BE7487">One new species and records of Ichthydium Ehrenberg, 1830 (Gastrotricha: Chaetonotida) from Sweden with a key to the genus</mods:title>
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<mods:namePart id="7E03E498FDE96EA45AD7747B403D1A5B">Kånneby, Tobias</mods:namePart>
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<mods:namePart id="697B88388CCACDC9E8CC538CAE3F9543">Todaro, Antonio</mods:namePart>
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<mods:namePart id="4F08E89C39F031251304FA7CD0CEFD23">Jondelius, Ulf</mods:namePart>
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<mods:date id="728B0DF9A49F8FF748E804412A203817">2009</mods:date>
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<mods:identifier id="9345754C79460AE0F87A88E070969293" type="Zenodo-Dep">191157</mods:identifier>
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<treatment id="03834469FFB50318FF02F8DEF46DF839" ID-DOI="http://doi.org/10.5281/zenodo.5627837" ID-GBIF-Taxon="119626088" ID-Zenodo-Dep="5627837" LSID="urn:lsid:plazi:treatment:03834469FFB50318FF02F8DEF46DF839" httpUri="http://treatment.plazi.org/id/03834469FFB50318FF02F8DEF46DF839" lastPageId="4" lastPageNumber="30" pageId="1" pageNumber="27">
<subSubSection id="C330A6F4FFB5031DFF02F8DEF093F8EE" pageId="1" pageNumber="27" type="nomenclature">
<paragraph id="8B95F57FFFB5031DFF02F8DEF0F7F8CD" blockId="1.[151,504,1842,1903]" box="[151,504,1842,1868]" pageId="1" pageNumber="27">
<heading id="D0DD4213FFB5031DFF02F8DEF0F7F8CD" bold="true" box="[151,504,1842,1868]" fontSize="11" level="1" pageId="1" pageNumber="27" reason="1">
<emphasis id="B95E296DFFB5031DFF02F8DEF0F7F8CD" bold="true" box="[151,504,1842,1868]" pageId="1" pageNumber="27">
<taxonomicName id="4C2A8EFCFFB5031DFF02F8DEF0A1F8CD" ID-CoL="6MTQP" box="[151,430,1842,1868]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="1" pageNumber="27" phylum="Gastrotricha" rank="species" species="skandicum" status="sp. nov.">
<emphasis id="B95E296DFFB5031DFF02F8DEF0A1F8CD" bold="true" box="[151,430,1842,1868]" italics="true" pageId="1" pageNumber="27">Ichthydium skandicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D9416FFB5031DFE20F8DEF0F7F8CD" box="[437,504,1842,1868]" pageId="1" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B95F57FFFB5031DFF02F8B9F093F8EE" blockId="1.[151,504,1842,1903]" box="[151,412,1877,1903]" pageId="1" pageNumber="27">
(
<figureCitation id="1311E9FAFFB5031DFF0AF8B9F000F8EE" box="[159,271,1877,1903]" captionStart="FIGURE 1. A C" captionStartId="2.[151,255,1184,1208]" captionTargetBox="[172,1425,199,1148]" captionTargetId="figure@2.[155,1440,174,1192]" captionTargetPageId="2" captionText="FIGURE 1. A C) Different representations of Ichthydium skandicum n. sp. Schematic drawing of A) dorsal view and B) ventral view. C) Detailed drawing of dorsal view. Scale bar: 20 µm." httpUri="https://zenodo.org/record/191158/files/figure.png" pageId="1" pageNumber="27">Figures 1</figureCitation>
,
<figureCitation id="1311E9FAFFB5031DFE88F8B9F021F8EE" box="[285,302,1877,1903]" captionStart="FIGURE 2. A C" captionStartId="3.[151,255,1269,1293]" captionTargetBox="[155,1436,195,1245]" captionTargetId="figure@3.[155,1440,194,1246]" captionTargetPageId="3" captionText="FIGURE 2. A C) Photographs of live specimens of Ichthydium skandicum n. sp. A) Habitus. B) Ventral view showing distribution of ciliary tufts in the trunk region. C) Dorsal view of posterior trunk region showing distribution of scales. Scale bars: A, B 20 µm; C, 10 µm." httpUri="https://zenodo.org/record/191159/files/figure.png" pageId="1" pageNumber="27">2</figureCitation>
;
<tableCitation id="C6A8C0C4FFB5031DFEAEF8B9F09FF8EE" box="[315,400,1877,1903]" captionStart="TABLE 1" captionStartId="4.[151,241,267,291]" captionTargetBox="[151,1436,395,1112]" captionTargetPageId="4" captionText="TABLE 1. Morphometric data (µm) for Ichthydium skandicum n. sp. Specimen H, 1, 2 and 3 were found in Jämtland 2008 while specimens 4, 5 and 6 were found in Bohuslän 2009. L, Length; W, Width; H, Holotype; Na, Not available. Character Specimen" httpUri="http://table.plazi.org/id/DF55A5F7FFB00318FF02FEE7F3A1FEF9" pageId="1" pageNumber="27" tableUuid="DF55A5F7FFB00318FF02FEE7F3A1FEF9">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C330A6F4FFB5031EFF02F84EF0E6F9B0" lastPageId="2" lastPageNumber="28" pageId="1" pageNumber="27" type="materials_examined">
<paragraph id="8B95F57FFFB5031DFF02F84EF431F862" blockId="1.[151,1437,1954,2019]" pageId="1" pageNumber="27">
<emphasis id="B95E296DFFB5031DFF02F84EF05BF83D" bold="true" box="[151,340,1954,1980]" pageId="1" pageNumber="27">
<typeStatus id="54914BDDFFB5031DFF02F84EF1D6F83D" box="[151,217,1954,1980]" pageId="1" pageNumber="27">Type</typeStatus>
material
</emphasis>
:
<typeStatus id="54914BDDFFB5031DFEF3F84EF0D3F83D" box="[358,476,1954,1980]" pageId="1" pageNumber="27" type="holotype">Holotype</typeStatus>
mounted in glycerin with 10 % borax neutralized formalin. Department of Invertebrate Zoology, Swedish Museum of Natural History, Stockholm,
<collectingCountry id="F33DB5EFFFB5031DFC70F825F549F862" box="[997,1094,1993,2019]" name="Sweden" pageId="1" pageNumber="27">Sweden</collectingCountry>
. Accession no. 7753.
</paragraph>
<caption id="DF55A5F7FFB6031EFF02FB4CF267FB5B" httpUri="https://zenodo.org/record/191158/files/figure.png" pageId="2" pageNumber="28" targetBox="[172,1425,199,1148]" targetPageId="2">
<paragraph id="8B95F57FFFB6031EFF02FB4CF267FB5B" blockId="2.[151,1436,1184,1242]" pageId="2" pageNumber="28">
<emphasis id="B95E296DFFB6031EFF02FB4CF053FB39" bold="true" box="[151,348,1184,1208]" pageId="2" pageNumber="28">FIGURE 1. AC)</emphasis>
Different representations of
<taxonomicName id="4C2A8EFCFFB6031EFD18FB4CF272FB36" box="[653,893,1184,1207]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="2" pageNumber="28" phylum="Gastrotricha" rank="species" species="skandicum" status="sp. nov.">
<emphasis id="B95E296DFFB6031EFD18FB4CF272FB36" box="[653,893,1184,1207]" italics="true" pageId="2" pageNumber="28">Ichthydium skandicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D9416FFB6031EFC16FB4CF2B3FB36" box="[899,956,1184,1207]" pageId="2" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
Schematic drawing of A) dorsal view and B) ventral view. C) Detailed drawing of dorsal view. Scale bar: 20 µm.
</paragraph>
</caption>
<paragraph id="8B95F57FFFB6031EFF50FAE4F23EFAC8" blockId="2.[151,1438,1288,1972]" pageId="2" pageNumber="28">
<emphasis id="B95E296DFFB6031EFF50FAE4F085FAA3" bold="true" box="[197,394,1288,1314]" pageId="2" pageNumber="28">Other material:</emphasis>
Six specimens which were measured and photographed but are no longer extant. Several additional specimens in 95% Ethanol for genetic studies.
</paragraph>
<paragraph id="8B95F57FFFB6031EFF50FAB9F36BFA17" blockId="2.[151,1438,1288,1972]" pageId="2" pageNumber="28">
<emphasis id="B95E296DFFB6031EFF50FAB9F06AFAEE" bold="true" box="[197,357,1365,1391]" pageId="2" pageNumber="28">
<typeStatus id="54914BDDFFB6031EFF50FAB9F00DFAEE" box="[197,258,1365,1391]" pageId="2" pageNumber="28">Type</typeStatus>
locality
</emphasis>
: A small pool in a bog close to highway E14 between Åre and Storlien, Jämtland,
<collectingCountry id="F33DB5EFFFB6031EFAADFABAF497FAF1" box="[1336,1432,1366,1392]" name="Sweden" pageId="2" pageNumber="28">Sweden</collectingCountry>
,
<date id="FF94D3BFFFB6031EFF02FA90F005FA17" box="[151,266,1404,1430]" pageId="2" pageNumber="28" value="2008-07">July 2008</date>
(
<geoCoordinate id="EE1E93B8FFB6031EFE8CFA90F0BBFA17" box="[281,436,1404,1430]" direction="north" orientation="latitude" pageId="2" pageNumber="28" precision="1" value="63.354134">63º21.248N</geoCoordinate>
;
<geoCoordinate id="EE1E93B8FFB6031EFE55FA90F357FA17" box="[448,600,1404,1430]" direction="east" orientation="longitude" pageId="2" pageNumber="28" precision="1" value="12.59005">12º35.403E</geoCoordinate>
).
</paragraph>
<paragraph id="8B95F57FFFB6031EFF50FA4EF0E6F9B0" blockId="2.[151,1438,1288,1972]" pageId="2" pageNumber="28">
<emphasis id="B95E296DFFB6031EFF50FA4EF08BFA3D" bold="true" box="[197,388,1442,1468]" pageId="2" pageNumber="28">Other localities</emphasis>
: Rockpool on Skaftö among
<taxonomicName id="4C2A8EFCFFB6031EFD48FA4FF256FA3D" box="[733,857,1443,1468]" class="Sphagnopsida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" order="Sphagnales" pageId="2" pageNumber="28" phylum="Bryophyta" rank="genus">
<emphasis id="B95E296DFFB6031EFD48FA4FF256FA3D" box="[733,857,1443,1468]" italics="true" pageId="2" pageNumber="28">Sphagnum</emphasis>
</taxonomicName>
sp., Bohuslän,
<collectingCountry id="F33DB5EFFFB6031EFB9BFA4FF561FA3C" box="[1038,1134,1443,1469]" name="Sweden" pageId="2" pageNumber="28">Sweden</collectingCountry>
,
<date id="FF94D3BFFFB6031EFBEFFA4FF5E3FA3C" box="[1146,1260,1443,1469]" pageId="2" pageNumber="28" value="2009-07">July 2009</date>
(
<geoCoordinate id="EE1E93B8FFB6031EFB68FA4FF498FA3C" box="[1277,1431,1443,1469]" direction="north" orientation="latitude" pageId="2" pageNumber="28" precision="1" value="58.247284">58º14.837N</geoCoordinate>
;
<geoCoordinate id="EE1E93B8FFB6031EFF02FA26F020FA65" box="[151,303,1482,1508]" direction="east" orientation="longitude" pageId="2" pageNumber="28" precision="1" value="11.437817">11º26.269E</geoCoordinate>
); Artificial pond on Skaftö among
<taxonomicName id="4C2A8EFCFFB6031EFD44FA26F241FA62" box="[721,846,1482,1507]" class="Sphagnopsida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" order="Sphagnales" pageId="2" pageNumber="28" phylum="Bryophyta" rank="genus">
<emphasis id="B95E296DFFB6031EFD44FA26F241FA62" box="[721,846,1482,1507]" italics="true" pageId="2" pageNumber="28">Sphagnum</emphasis>
</taxonomicName>
sp., Bohuslän,
<collectingCountry id="F33DB5EFFFB6031EFB9DFA26F566FA65" box="[1032,1129,1482,1508]" name="Sweden" pageId="2" pageNumber="28">Sweden</collectingCountry>
,
<date id="FF94D3BFFFB6031EFBE0FA26F5E5FA65" box="[1141,1258,1482,1508]" pageId="2" pageNumber="28" value="2009-07">July 2009</date>
(
<geoCoordinate id="EE1E93B8FFB6031EFB6EFA26F498FA65" box="[1275,1431,1482,1508]" direction="north" orientation="latitude" pageId="2" pageNumber="28" precision="1" value="58.2439">58º14.634N</geoCoordinate>
;
<geoCoordinate id="EE1E93B8FFB6031EFF02FA1CF04FF98B" box="[151,320,1520,1546]" direction="east" orientation="longitude" pageId="2" pageNumber="28" precision="1" value="11.453667">11º27.220E</geoCoordinate>
); Rockpool on Blåbergsholmen among
<taxonomicName id="4C2A8EFCFFB6031EFCDEFA1DF2DCF98B" box="[843,979,1521,1546]" class="Sphagnopsida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" order="Sphagnales" pageId="2" pageNumber="28" phylum="Bryophyta" rank="genus">
<emphasis id="B95E296DFFB6031EFCDEFA1DF2DCF98B" box="[843,979,1521,1546]" italics="true" pageId="2" pageNumber="28">Sphagnum</emphasis>
</taxonomicName>
sp., Bohuslän,
<collectingCountry id="F33DB5EFFFB6031EFB36FA1CF402F98B" box="[1187,1293,1520,1546]" name="Sweden" pageId="2" pageNumber="28">Sweden</collectingCountry>
,
<date id="FF94D3BFFFB6031EFA89FA1CF491F98B" box="[1308,1438,1520,1546]" pageId="2" pageNumber="28" value="2009-07">July 2009</date>
(
<geoCoordinate id="EE1E93B8FFB6031EFF0AF9FBF035F9B0" box="[159,314,1559,1585]" direction="north" orientation="latitude" pageId="2" pageNumber="28" precision="1" value="58.2496">58º14.976N</geoCoordinate>
;
<geoCoordinate id="EE1E93B8FFB6031EFED3F9FBF0D2F9B0" box="[326,477,1559,1585]" direction="east" orientation="longitude" pageId="2" pageNumber="28" precision="1" value="11.439716">11º26.383E</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection id="C330A6F4FFB6031EFF50F9D1F041F9FF" pageId="2" pageNumber="28" type="etymology">
<paragraph id="8B95F57FFFB6031EFF50F9D1F041F9FF" blockId="2.[151,1438,1288,1972]" pageId="2" pageNumber="28">
<emphasis id="B95E296DFFB6031EFF50F9D1F043F9D6" bold="true" box="[197,332,1597,1623]" pageId="2" pageNumber="28">Etymology</emphasis>
: The species name refers to the Scandinavian mountain range “Skanderna” where the species was first found.
</paragraph>
</subSubSection>
<subSubSection id="C330A6F4FFB6031EFF50F966F37CF835" pageId="2" pageNumber="28" type="diagnosis">
<paragraph id="8B95F57FFFB6031EFF50F966F37CF835" blockId="2.[151,1438,1288,1972]" pageId="2" pageNumber="28">
<emphasis id="B95E296DFFB6031EFF50F966F031F925" bold="true" box="[197,318,1674,1700]" pageId="2" pageNumber="28">Diagnosis</emphasis>
: Mid-sized
<taxonomicName id="4C2A8EFCFFB6031EFE58F967F35EF925" box="[461,593,1675,1700]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="2" pageNumber="28" phylum="Gastrotricha" rank="genus">
<emphasis id="B95E296DFFB6031EFE58F967F35EF925" box="[461,593,1675,1700]" italics="true" pageId="2" pageNumber="28">Ichthydium</emphasis>
</taxonomicName>
species. Head with three lobes and one pair of ciliary tufts. Furca long and tong-shaped, each branch with very short base. Two pairs of thick sensory bristles, the posterior pair anchored to bi-keeled subtriangular scales. Dorsal body surface naked except for one pair of subtriangular bikeeled scales anchoring the posterior sensory bristles and four pairs of keeled subrectangular scales at the base of the furca. Pharynx large, 3035 µm in length with posterior swelling; pharyngeal-intestinal junction (PhIJ) at U28. Locomotory cilia arranged in two rows in the anterior body region divided into four pairs of tufts in the trunk region. Ventral interciliary area naked except for two subrectangular terminal scales. Specimens observed were in parthenogenetic phase.
</paragraph>
</subSubSection>
<caption id="DF55A5F7FFB7031FFF02FB19F305FAD3" httpUri="https://zenodo.org/record/191159/files/figure.png" pageId="3" pageNumber="29" targetBox="[155,1436,195,1245]" targetPageId="3">
<paragraph id="8B95F57FFFB7031FFF02FB19F305FAD3" blockId="3.[151,1436,1269,1362]" pageId="3" pageNumber="29">
<emphasis id="B95E296DFFB7031FFF02FB19F015FA8C" bold="true" box="[151,282,1269,1293]" pageId="3" pageNumber="29">FIGURE 2.</emphasis>
A
<emphasis id="B95E296DFFB7031FFEA1FB19F04EFA8C" bold="true" box="[308,321,1269,1293]" pageId="3" pageNumber="29"></emphasis>
C) Photographs of live specimens of
<taxonomicName id="4C2A8EFCFFB7031FFD47FB1AF2CCFA8C" box="[722,963,1270,1293]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="3" pageNumber="29" phylum="Gastrotricha" rank="species" species="skandicum" status="sp. nov.">
<emphasis id="B95E296DFFB7031FFD47FB1AF2CCFA8C" box="[722,963,1270,1293]" italics="true" pageId="3" pageNumber="29">Ichthydium skandicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D9416FFB7031FFC5FFB1AF50BFA8C" box="[970,1028,1270,1293]" pageId="3" pageNumber="29" rank="species">n. sp.</taxonomicNameLabel>
A) Habitus. B) Ventral view showing distribution of ciliary tufts in the trunk region. C) Dorsal view of posterior trunk region showing distribution of scales. Scale bars: A, B 20 µm; C, 10 µm.
</paragraph>
</caption>
<subSubSection id="C330A6F4FFB70318FF50FA6CF1D4FA1D" lastPageId="4" lastPageNumber="30" pageId="3" pageNumber="29" type="description">
<paragraph id="8B95F57FFFB7031FFF50FA6CF5AEF977" blockId="3.[151,1437,1408,2014]" pageId="3" pageNumber="29">
<emphasis id="B95E296DFFB7031FFF50FA6CF06FFA1B" bold="true" box="[197,352,1408,1434]" pageId="3" pageNumber="29">Description:</emphasis>
Mid-sized
<taxonomicName id="4C2A8EFCFFB7031FFE72FA6DF364FA1B" box="[487,619,1409,1434]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="3" pageNumber="29" phylum="Gastrotricha" rank="genus">
<emphasis id="B95E296DFFB7031FFE72FA6DF364FA1B" box="[487,619,1409,1434]" italics="true" pageId="3" pageNumber="29">Ichthydium</emphasis>
</taxonomicName>
species, 132142 µm in total body length. Three-lobed head bearing one pair of cephalic ciliary tufts at U6. The longest cilium in each tuft is 20 µm in length. No cephalic plates were detected. Width of head/neck/trunk/furcal base are as follows: 2332/2028/2842/1214 µm at U13/ U28/ U60/ U82. Furca tong-shaped, 2730 µm in length of which the adhesive tubes contribute 2025 µm. A pair of bi-keeled subtriangular scales anchoring the posterior sensory bristles is present at U75. Another pair of sensory bristles arises from the neck region at U20. The sensory bristles are thick, the anterior pair (ASB) measures 1012 µm and the posterior pair (PSB) 20 µm in length. Moreover, four pairs of dorsal subelliptical, keeled scales are present at the posterior end, two pairs situated dorsolaterally at U76 (
<date id="FF94D3BFFFB7031FFB1DF963F5A2F928" box="[1160,1197,1679,1705]" pageId="3" pageNumber="29">6 x</date>
1.5 µm) and U80 (
<date id="FF94D3BFFFB7031FFA18F963F1A8F951" pageId="3" pageNumber="29">6 x</date>
1.5 µm) respectively and two pairs dorsally at U78 (4.5
<date id="FF94D3BFFFB7031FFC02F95AF2A9F951" box="[919,934,1718,1744]" pageId="3" pageNumber="29">6</date>
x 12 µm) and U81 (4.5
<date id="FF94D3BFFFB7031FFB6BF95AF403F951" box="[1278,1292,1718,1744]" pageId="3" pageNumber="29">5</date>
x 12 µm) respectively. Apart from these five pairs of scales the dorsal body cuticle appears naked.
</paragraph>
<paragraph id="8B95F57FFFB7031FFF50F8EFF493F85F" blockId="3.[151,1437,1408,2014]" pageId="3" pageNumber="29">Ventral ciliation consists of locomotor cilia, 2025 µm in length, arranged in two rows. In the trunk region the two rows are divided into four pairs of tufts. The anterior-most pair of tufts is situated in the anterior trunk region. The posterior-most pair of tufts is separated by a greater distance from the others of the same row. Interciliary area naked except for two rectangular keeled terminal scales, 89 µm in length and 44.5 µm in width. Mouth subterminal, 67 µm in diameter. Pharynx, 3035 µm in length, widening towards the pharyngeal intestinal junction at U28. Intestine straight, tapering towards posterior end, anus ventral at U74.</paragraph>
<paragraph id="8B95F57FFFB00318FF02FF74F3EEFF59" blockId="4.[151,1435,152,216]" pageId="4" pageNumber="30">The observed specimens were in parthenogenetic phase. More or less mature eggs, situated on either side or dorsal to the intestine, were seen in all specimens.</paragraph>
<caption id="DF55A5F7FFB00318FF02FEE7F3A1FEF9" ID-Table-UUID="DF55A5F7FFB00318FF02FEE7F3A1FEF9" httpUri="http://table.plazi.org/id/DF55A5F7FFB00318FF02FEE7F3A1FEF9" pageId="4" pageNumber="30" targetBox="[151,1436,395,1112]" targetIsTable="true" targetPageId="4">
<paragraph id="8B95F57FFFB00318FF02FEE7F3A1FEF9" blockId="4.[151,1436,267,376]" pageId="4" pageNumber="30">
<emphasis id="B95E296DFFB00318FF02FEE7F003FEA2" bold="true" box="[151,268,267,291]" pageId="4" pageNumber="30">TABLE 1.</emphasis>
Morphometric data (µm) for
<taxonomicName id="4C2A8EFCFFB00318FDC7FEE0F247FEA2" box="[594,840,268,291]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="skandicum" status="sp. nov.">
<emphasis id="B95E296DFFB00318FDC7FEE0F247FEA2" box="[594,840,268,291]" italics="true" pageId="4" pageNumber="30">Ichthydium skandicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D9416FFB00318FCDAFEE0F285FEA2" box="[847,906,268,291]" pageId="4" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
Specimen H, 1, 2 and 3 were found in Jämtland 2008 while specimens 4, 5 and 6 were found in Bohuslän 2009. L, Length; W, Width; H, Holotype; Na, Not available. Character Specimen
</paragraph>
</caption>
<paragraph id="8B95F57FFFB00318FF0AFE59F305FBD9" pageId="4" pageNumber="30">
<table id="F92A07DFFFB0FCE3FF02FE67F493FBD9" box="[151,1436,395,1112]" gridcols="8" gridrows="7" pageId="4" pageNumber="30">
<tr id="351AF73DFFB0FCE3FF02FE67F493FE4D" box="[151,1436,395,460]" gridrow="0" pageId="4" pageNumber="30">
<th id="76CB9E41FFB0FCE3FF02FE67F305FE4D" box="[151,522,395,460]" gridcol="0" gridrow="0" pageId="4" pageNumber="30">Total body L</th>
<th id="76CB9E41FFB0FCE3FDD0FE67F375FE4D" box="[581,634,395,460]" gridcol="1" gridrow="0" pageId="4" pageNumber="30">H 135</th>
<th id="76CB9E41FFB0FCE3FD25FE67F3EAFE4D" box="[688,741,395,460]" gridcol="2" gridrow="0" pageId="4" pageNumber="30">1 137</th>
<th id="76CB9E41FFB0FCE3FC89FE67F249FE4D" box="[796,838,395,460]" gridcol="3" gridrow="0" pageId="4" pageNumber="30">2 132</th>
<th id="76CB9E41FFB0FCE3FC1CFE67F2B1FE4D" box="[905,958,395,460]" gridcol="4" gridrow="0" pageId="4" pageNumber="30">3 135</th>
<th id="76CB9E41FFB0FCE3FB85FE67F556FE4D" box="[1040,1113,395,460]" gridcol="5" gridrow="0" pageId="4" pageNumber="30">4 140</th>
<th id="76CB9E41FFB0FCE3FB02FE67F5EFFE4D" box="[1175,1248,395,460]" gridcol="6" gridrow="0" pageId="4" pageNumber="30">5 140</th>
<th id="76CB9E41FFB0FCE3FA8BFE67F493FE4D" box="[1310,1436,395,460]" gridcol="7" gridrow="0" pageId="4" pageNumber="30">6 142</th>
</tr>
<tr id="351AF73DFFB0FCE3FF02FE0CF493FDCA" box="[151,1436,480,587]" gridrow="1" pageId="4" pageNumber="30">
<th id="76CB9E41FFB0FCE3FF02FE0CF305FDCA" box="[151,522,480,587]" gridcol="0" gridrow="1" pageId="4" pageNumber="30">Head W Neck W Body W</th>
<td id="76CB9E41FFB0FCE3FDD0FE0CF375FDCA" box="[581,634,480,587]" gridcol="1" gridrow="1" pageId="4" pageNumber="30">30 25 35</td>
<td id="76CB9E41FFB0FCE3FD25FE0CF3EAFDCA" box="[688,741,480,587]" gridcol="2" gridrow="1" pageId="4" pageNumber="30">25 20 33</td>
<td id="76CB9E41FFB0FCE3FC89FE0CF249FDCA" box="[796,838,480,587]" gridcol="3" gridrow="1" pageId="4" pageNumber="30">25 22 32</td>
<td id="76CB9E41FFB0FCE3FC1CFE0CF2B1FDCA" box="[905,958,480,587]" gridcol="4" gridrow="1" pageId="4" pageNumber="30">23 13 28</td>
<td id="76CB9E41FFB0FCE3FB85FE0CF556FDCA" box="[1040,1113,480,587]" gridcol="5" gridrow="1" pageId="4" pageNumber="30">28 22 33</td>
<td id="76CB9E41FFB0FCE3FB02FE0CF5EFFDCA" box="[1175,1248,480,587]" gridcol="6" gridrow="1" pageId="4" pageNumber="30">32 22 30</td>
<td id="76CB9E41FFB0FCE3FA8BFE0CF493FDCA" box="[1310,1436,480,587]" gridcol="7" gridrow="1" pageId="4" pageNumber="30">32 28 42*</td>
</tr>
<tr id="351AF73DFFB0FCE3FF02FDB3F493FD21" box="[151,1436,607,672]" gridrow="2" pageId="4" pageNumber="30">
<th id="76CB9E41FFB0FCE3FF02FDB3F305FD21" box="[151,522,607,672]" gridcol="0" gridrow="2" pageId="4" pageNumber="30">Furca Base W Furca L</th>
<td id="76CB9E41FFB0FCE3FDD0FDB3F375FD21" box="[581,634,607,672]" gridcol="1" gridrow="2" pageId="4" pageNumber="30">13 30</td>
<td id="76CB9E41FFB0FCE3FD25FDB3F3EAFD21" box="[688,741,607,672]" gridcol="2" gridrow="2" pageId="4" pageNumber="30">12 30</td>
<td id="76CB9E41FFB0FCE3FC89FDB3F249FD21" box="[796,838,607,672]" gridcol="3" gridrow="2" pageId="4" pageNumber="30">12 28</td>
<td id="76CB9E41FFB0FCE3FC1CFDB3F2B1FD21" box="[905,958,607,672]" gridcol="4" gridrow="2" pageId="4" pageNumber="30">12 28</td>
<td id="76CB9E41FFB0FCE3FB85FDB3F556FD21" box="[1040,1113,607,672]" gridcol="5" gridrow="2" pageId="4" pageNumber="30">13 27</td>
<td id="76CB9E41FFB0FCE3FB02FDB3F5EFFD21" box="[1175,1248,607,672]" gridcol="6" gridrow="2" pageId="4" pageNumber="30">13 28</td>
<td id="76CB9E41FFB0FCE3FA8BFDB3F493FD21" box="[1310,1436,607,672]" gridcol="7" gridrow="2" pageId="4" pageNumber="30">14 28</td>
</tr>
<tr id="351AF73DFFB0FCE3FF02FD58F493FC9E" box="[151,1436,692,799]" gridrow="3" pageId="4" pageNumber="30">
<th id="76CB9E41FFB0FCE3FF02FD58F305FC9E" box="[151,522,692,799]" gridcol="0" gridrow="3" pageId="4" pageNumber="30">Pharynx L Mouth ø Sensory Bristle Scale L x W</th>
<td id="76CB9E41FFB0FCE3FDD0FD58F375FC9E" box="[581,634,692,799]" gridcol="1" gridrow="3" pageId="4" pageNumber="30">34 7 4 x 4</td>
<td id="76CB9E41FFB0FCE3FD25FD58F3EAFC9E" box="[688,741,692,799]" gridcol="2" gridrow="3" pageId="4" pageNumber="30">31 6 6 x 5</td>
<td id="76CB9E41FFB0FCE3FC89FD58F249FC9E" box="[796,838,692,799]" gridcol="3" gridrow="3" pageId="4" pageNumber="30">30 6 Na</td>
<td id="76CB9E41FFB0FCE3FC1CFD58F2B1FC9E" box="[905,958,692,799]" gridcol="4" gridrow="3" pageId="4" pageNumber="30">30 6 5 x 5</td>
<td id="76CB9E41FFB0FCE3FB85FD58F556FC9E" box="[1040,1113,692,799]" gridcol="5" gridrow="3" pageId="4" pageNumber="30">32 6 4 x 4</td>
<td id="76CB9E41FFB0FCE3FB02FD58F5EFFC9E" box="[1175,1248,692,799]" gridcol="6" gridrow="3" pageId="4" pageNumber="30">33 6 4 x 4</td>
<td id="76CB9E41FFB0FCE3FA8BFD58F493FC9E" box="[1310,1436,692,799]" gridcol="7" gridrow="3" pageId="4" pageNumber="30">35 6 4 x 4</td>
</tr>
<tr id="351AF73DFFB0FCE3FF02FCDFF493FC1F" box="[151,1436,819,926]" gridrow="4" pageId="4" pageNumber="30">
<th id="76CB9E41FFB0FCE3FF02FCDFF305FC1F" box="[151,522,819,926]" gridcol="0" gridrow="4" pageId="4" pageNumber="30">Anterior Sensory Bristle L Posterior Sensory Bristle L Furca Scales Dorsolateral L x W</th>
<td id="76CB9E41FFB0FCE3FDD0FCDFF375FC1F" box="[581,634,819,926]" gridcol="1" gridrow="4" pageId="4" pageNumber="30">~10 20 6 x 2</td>
<td id="76CB9E41FFB0FCE3FD25FCDFF3EAFC1F" box="[688,741,819,926]" gridcol="2" gridrow="4" pageId="4" pageNumber="30">Na 20 Na</td>
<td id="76CB9E41FFB0FCE3FC89FCDFF249FC1F" box="[796,838,819,926]" gridcol="3" gridrow="4" pageId="4" pageNumber="30">Na ~20 Na</td>
<td id="76CB9E41FFB0FCE3FC1CFCDFF2B1FC1F" box="[905,958,819,926]" gridcol="4" gridrow="4" pageId="4" pageNumber="30">~12 ~20 Na</td>
<td id="76CB9E41FFB0FCE3FB85FCDFF556FC1F" box="[1040,1113,819,926]" gridcol="5" gridrow="4" pageId="4" pageNumber="30">13 20 Na</td>
<td id="76CB9E41FFB0FCE3FB02FCDFF5EFFC1F" box="[1175,1248,819,926]" gridcol="6" gridrow="4" pageId="4" pageNumber="30">9 26 5 x 1</td>
<td id="76CB9E41FFB0FCE3FA8BFCDFF493FC1F" box="[1310,1436,819,926]" gridcol="7" gridrow="4" pageId="4" pageNumber="30">10 Na 6 x 1.5</td>
</tr>
<tr id="351AF73DFFB0FCE3FF02FC5EF493FB9C" box="[151,1436,946,1053]" gridrow="5" pageId="4" pageNumber="30">
<th id="76CB9E41FFB0FCE3FF02FC5EF305FB9C" box="[151,522,946,1053]" gridcol="0" gridrow="5" pageId="4" pageNumber="30">Furca Scales Dorsal L x W Ventral Cilia L Ventral Terminal Scales L x W</th>
<td id="76CB9E41FFB0FCE3FDD0FC5EF375FB9C" box="[581,634,946,1053]" gridcol="1" gridrow="5" pageId="4" pageNumber="30">6 x 2 Na 9 x 4</td>
<td id="76CB9E41FFB0FCE3FD25FC5EF3EAFB9C" box="[688,741,946,1053]" gridcol="2" gridrow="5" pageId="4" pageNumber="30">Na ~24 Na</td>
<td id="76CB9E41FFB0FCE3FC89FC5EF249FB9C" box="[796,838,946,1053]" gridcol="3" gridrow="5" pageId="4" pageNumber="30">Na ~25 Na</td>
<td id="76CB9E41FFB0FCE3FC1CFC5EF2B1FB9C" box="[905,958,946,1053]" gridcol="4" gridrow="5" pageId="4" pageNumber="30">Na Na Na</td>
<td id="76CB9E41FFB0FCE3FB85FC5EF556FB9C" box="[1040,1113,946,1053]" gridcol="5" gridrow="5" pageId="4" pageNumber="30">4.5 x 1 20-25 9 x 4.5</td>
<td id="76CB9E41FFB0FCE3FB02FC5EF5EFFB9C" box="[1175,1248,946,1053]" gridcol="6" gridrow="5" pageId="4" pageNumber="30">4.5 x 1 24 8 x 4</td>
<td id="76CB9E41FFB0FCE3FA8BFC5EF493FB9C" box="[1310,1436,946,1053]" gridcol="7" gridrow="5" pageId="4" pageNumber="30">5 x 1 20-25 Na</td>
</tr>
<tr id="351AF73DFFB0FCE3FF02FBADF493FBD9" box="[151,1436,1089,1112]" gridrow="6" pageId="4" pageNumber="30" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76CB9E41FFB0FCE3FF02FBADF305FBD9" box="[151,522,1089,1112]" gridcol="0" gridrow="6" pageId="4" pageNumber="30">* Specimen with large mature egg.</th>
</tr>
</table>
</paragraph>
<paragraph id="8B95F57FFFB00318FF50FB6AF1D4FA1D" blockId="4.[151,1437,1158,1976]" pageId="4" pageNumber="30">
<emphasis id="B95E296DFFB00318FF50FB6AF0C7FB21" bold="true" box="[197,456,1158,1184]" pageId="4" pageNumber="30">Taxonomic remarks:</emphasis>
<taxonomicName id="4C2A8EFCFFB00318FE45FB6AF35BFB1E" box="[464,596,1158,1183]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="genus">
<emphasis id="B95E296DFFB00318FE45FB6AF35BFB1E" box="[464,596,1158,1183]" italics="true" pageId="4" pageNumber="30">Ichthydium</emphasis>
</taxonomicName>
is currently divided into three subgenera (but see
<bibRefCitation id="EFBB888EFFB00318FB0AFB6AF480FB21" author="Balsamo" box="[1183,1423,1158,1184]" pageId="4" pageNumber="42" refString="Balsamo, M., d'Hondt, J. L., Pierboni, L., Grilli, P. (2009) Taxonomic and nomenclatural notes on freshwater Gastrotricha. Zootaxa, 2158, 1 - 19." type="journal article" year="2009">
Balsamo
<emphasis id="B95E296DFFB00318FA98FB6AF44EFB1E" box="[1293,1345,1158,1183]" italics="true" pageId="4" pageNumber="30">et al</emphasis>
. 2009
</bibRefCitation>
):
<taxonomicName id="4C2A8EFCFFB00318FF02FB46F0F1FB45" authority="Schwank, 1990" authorityName="Schwank" authorityYear="1990" box="[151,510,1194,1220]" family="Chaetonotidae" genus="Euichthydium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="genus">
<emphasis id="B95E296DFFB00318FF02FB46F032FB42" box="[151,317,1194,1219]" italics="true" pageId="4" pageNumber="30">Euichthydium</emphasis>
<bibRefCitation id="EFBB888EFFB00318FED0FB46F0F1FB45" author="Schwank" box="[325,510,1194,1220]" pageId="4" pageNumber="43" refString="Schwank, P. (1990) Gastrotricha. In: Schwoerbel, J. &amp; Zwick, P. (eds), Susswasserfauna von Mitteleuropa, Band 3. Gastrotricha und Nemertini. Gustav Fischer Verlag, Stuttgart, Jena, New York, 252 pp." type="book" year="1990">Schwank, 1990</bibRefCitation>
</taxonomicName>
predominantly species with a naked cuticle,
<taxonomicName id="4C2A8EFCFFB00318FB88FB46F497FB45" authority="Schwank, 1990" authorityName="Schwank" authorityYear="1990" box="[1053,1432,1194,1220]" family="Chaetonotidae" genus="Forficulichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="genus">
<emphasis id="B95E296DFFB00318FB88FB46F5DBFB42" box="[1053,1236,1194,1219]" italics="true" pageId="4" pageNumber="30">Forficulichthys</emphasis>
<bibRefCitation id="EFBB888EFFB00318FB49FB46F497FB45" author="Schwank" box="[1244,1432,1194,1220]" pageId="4" pageNumber="43" refString="Schwank, P. (1990) Gastrotricha. In: Schwoerbel, J. &amp; Zwick, P. (eds), Susswasserfauna von Mitteleuropa, Band 3. Gastrotricha und Nemertini. Gustav Fischer Verlag, Stuttgart, Jena, New York, 252 pp." type="book" year="1990">Schwank, 1990</bibRefCitation>
</taxonomicName>
, characterized by species bearing tong-shaped furca with tapering tips and the monotypic
<taxonomicName id="4C2A8EFCFFB00318FB5BFB22F044FA8D" authority="Rudescu, 1967" authorityName="Rudescu" authorityYear="1967" family="Chaetonotidae" genus="Pseudichthydium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="genus">
<emphasis id="B95E296DFFB00318FB5BFB22F493FB66" box="[1230,1436,1230,1255]" italics="true" pageId="4" pageNumber="30">Pseudichthydium</emphasis>
<bibRefCitation id="EFBB888EFFB00318FF02FB1EF044FA8D" author="Rudescu" box="[151,331,1266,1292]" pageId="4" pageNumber="43" refString="Rudescu, L. (1967) Gastrotricha. In: Fauna Republicii Socialiste Romania, Trochelminthes, 2 (3). Academia Republicii Socialiste Romania, Bucharest, 292 pp." type="book" year="1967">Rudescu, 1967</bibRefCitation>
</taxonomicName>
which contains
<taxonomicName id="4C2A8EFCFFB00318FD86FB1EF26DFA8D" authority="Varga, 1950" authorityName="Varga" authorityYear="1950" box="[531,866,1266,1292]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="balatonicum">
<emphasis id="B95E296DFFB00318FD86FB1EF3CBFA8A" box="[531,708,1266,1291]" italics="true" pageId="4" pageNumber="30">I. balatonicum</emphasis>
<bibRefCitation id="EFBB888EFFB00318FD58FB1EF26DFA8D" author="Varga" box="[717,866,1266,1292]" pageId="4" pageNumber="44" refString="Varga, L. (1950) Gastrotrichen aus dem Balaton-See. Annales Instituti Biologiae Pervestigandae Hungarici, 19, 1 - 14 (1949)." type="journal article" year="1950">Varga, 1950</bibRefCitation>
</taxonomicName>
, characterized by a large comb-like keel on its dorsal body surface. It should be emphasized that many species are still not affiliated to any of the three above subgenera (see Appendix 1). It should also be noted that the subgenus
<taxonomicName id="4C2A8EFCFFB00318FC66FAD6F595FAD2" box="[1011,1178,1338,1363]" family="Chaetonotidae" genus="Euichthydium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="genus">
<emphasis id="B95E296DFFB00318FC66FAD6F595FAD2" box="[1011,1178,1338,1363]" italics="true" pageId="4" pageNumber="30">Euichthydium</emphasis>
</taxonomicName>
is invalid; because it contains the
<typeStatus id="54914BDDFFB00318FEB8FAB2F06FFAF9" box="[301,352,1374,1400]" pageId="4" pageNumber="30">type</typeStatus>
of the genus
<taxonomicName id="4C2A8EFCFFB00318FD94FAB2F38AFAF6" box="[513,645,1374,1399]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="genus">
<emphasis id="B95E296DFFB00318FD94FAB2F38AFAF6" box="[513,645,1374,1399]" italics="true" pageId="4" pageNumber="30">Ichthydium</emphasis>
</taxonomicName>
, hence it must bear the same name as the genus (see ICZN Article 44.1).
</paragraph>
</subSubSection>
<subSubSection id="C330A6F4FFB00318FF50FA4AF507F812" pageId="4" pageNumber="30" type="reference_group">
<paragraph id="8B95F57FFFB00318FF50FA4AF283F9F5" blockId="4.[151,1437,1158,1976]" pageId="4" pageNumber="30">
<taxonomicName id="4C2A8EFCFFB00318FF50FA4AF0C1FA3E" ID-CoL="6MTQP" box="[197,462,1446,1471]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="skandicum" status="sp. nov.">
<emphasis id="B95E296DFFB00318FF50FA4AF0C1FA3E" box="[197,462,1446,1471]" italics="true" pageId="4" pageNumber="30">Ichthydium skandicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D9416FFB00318FE41FA4AF31BFA41" box="[468,532,1446,1472]" pageId="4" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
, by virtue of its tong-shaped furca, most closely resembles species within the subgenus
<taxonomicName id="4C2A8EFCFFB00318FE8FFA26F0EBFA62" box="[282,484,1482,1507]" family="Chaetonotidae" genus="Forficulichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="genus">
<emphasis id="B95E296DFFB00318FE8FFA26F0EBFA62" box="[282,484,1482,1507]" italics="true" pageId="4" pageNumber="30">Forficulichthys</emphasis>
</taxonomicName>
. According to
<bibRefCitation id="EFBB888EFFB00318FD39FA26F28CFA65" author="Schwank" box="[684,899,1482,1508]" pageId="4" pageNumber="43" refString="Schwank, P. (1990) Gastrotricha. In: Schwoerbel, J. &amp; Zwick, P. (eds), Susswasserfauna von Mitteleuropa, Band 3. Gastrotricha und Nemertini. Gustav Fischer Verlag, Stuttgart, Jena, New York, 252 pp." type="book" year="1990">Schwank (1990)</bibRefCitation>
the latter taxon contains four species,
<taxonomicName id="4C2A8EFCFFB00318FA1CFA26F0FCF989" ID-CoL="6N62L" authority="Brunson, 1949" authorityName="Brunson" authorityYear="1949" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="cephalobares">
<emphasis id="B95E296DFFB00318FA1CFA26F038F986" italics="true" pageId="4" pageNumber="30">I. cephalobares</emphasis>
<bibRefCitation id="EFBB888EFFB00318FEAAFA02F0FCF989" author="Brunson" box="[319,499,1518,1544]" pageId="4" pageNumber="42" refString="Brunson, R. B. (1949) Gastrotricha from North America II: Four new species of Ichthydium from Michigan. Papers of the Michigan Academy of Science, Arts and Letters, 33, 59 - 62 (1947)." type="journal article" year="1949">Brunson, 1949</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2A8EFCFFB00318FD95FA02F235F989" ID-CoL="6N62Q" authority="Remane, 1927" authorityName="Remane" authorityYear="1927" box="[512,826,1518,1544]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="forficula">
<emphasis id="B95E296DFFB00318FD95FA02F38EF986" box="[512,641,1518,1543]" italics="true" pageId="4" pageNumber="30">I. forficula</emphasis>
<bibRefCitation id="EFBB888EFFB00318FD1FFA02F235F989" author="Remane" box="[650,826,1518,1544]" pageId="4" pageNumber="43" refString="Remane, A. (1927) Beitrage zur Systematik der Susswassergastrotrichen. Zoologische Jahrbucher Abteilung fur Systematik Oekologie und Geographie der Tiere, 53, 269 - 320." type="journal article" year="1927">Remane, 1927</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2A8EFCFFB00318FCD2FA02F577F989" ID-CoL="6N5R2" authority="Voigt, 1901" authorityName="Voigt" authorityYear="1901" box="[839,1144,1518,1544]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="forcipatum">
<emphasis id="B95E296DFFB00318FCD2FA02F2ECF986" box="[839,995,1518,1543]" italics="true" pageId="4" pageNumber="30">I. forcipatum</emphasis>
<bibRefCitation id="EFBB888EFFB00318FC79FA02F577F989" author="Voigt" box="[1004,1144,1518,1544]" pageId="4" pageNumber="44" refString="Voigt, M. (1901) Diagnosen bisher unbeschriebener Organismen aus Ploner Gewassern. Zoologischer Anzeiger, 25, 35 - 39." type="journal article" year="1901">Voigt, 1901</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C2A8EFCFFB00318FB26FA02F451F986" ID-CoL="6N5QP" box="[1203,1374,1518,1543]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="tanytrichum">
<emphasis id="B95E296DFFB00318FB26FA02F451F986" box="[1203,1374,1518,1543]" italics="true" pageId="4" pageNumber="30">I. tanytrichum</emphasis>
</taxonomicName>
. The only species within this subgenus with both a tong-shaped furca and dorsal scales in the posterior trunk region is
<taxonomicName id="4C2A8EFCFFB00318FF26F9DAF055F9CE" ID-CoL="6N5QP" box="[179,346,1590,1615]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="tanytrichum">
<emphasis id="B95E296DFFB00318FF26F9DAF055F9CE" box="[179,346,1590,1615]" italics="true" pageId="4" pageNumber="30">I. tanytrichum</emphasis>
</taxonomicName>
, with bi-keeled scales that carry the posterior sensory bristles in contrast to
<taxonomicName id="4C2A8EFCFFB00318FB73F9DAF472F9CE" ID-CoL="6MTQP" box="[1254,1405,1590,1615]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="skandicum" status="sp. nov.">
<emphasis id="B95E296DFFB00318FB73F9DAF472F9CE" box="[1254,1405,1590,1615]" italics="true" pageId="4" pageNumber="30">I. skandicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D9416FFB00318FA13F9DAF1B7F9F5" pageId="4" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
which carries an additional four pairs of keeled dorsal scales.
</paragraph>
<paragraph id="8B95F57FFFB00318FF50F992F457F8F1" blockId="4.[151,1437,1158,1976]" pageId="4" pageNumber="30">
<taxonomicName id="4C2A8EFCFFB00318FF50F992F045F916" ID-CoL="7P68Y" box="[197,330,1662,1687]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="genus">
<emphasis id="B95E296DFFB00318FF50F992F045F916" box="[197,330,1662,1687]" italics="true" pageId="4" pageNumber="30">Ichthydium</emphasis>
</taxonomicName>
contains an additional four species that carry scales or spines in the posterior trunk region
<taxonomicName id="4C2A8EFCFFB00318FA1EF992F1F6F93A" ID-CoL="6N5QN" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="crassum">
<emphasis id="B95E296DFFB00318FA1EF992F1F6F93A" italics="true" pageId="4" pageNumber="30">I. crassum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A8EFCFFB00318FE9DF94EF0A3F93A" ID-CoL="6MTQH" box="[264,428,1698,1723]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="diacanthum">
<emphasis id="B95E296DFFB00318FE9DF94EF0A3F93A" box="[264,428,1698,1723]" italics="true" pageId="4" pageNumber="30">I. diacanthum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A8EFCFFB00318FE29F94EF322F93A" ID-CoL="6N5QS" box="[444,557,1698,1723]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="dubium">
<emphasis id="B95E296DFFB00318FE29F94EF322F93A" box="[444,557,1698,1723]" italics="true" pageId="4" pageNumber="30">I. dubium</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2A8EFCFFB00318FDFDF94EF22EF93A" ID-CoL="6MTQG" box="[616,801,1698,1723]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="squamigerum">
<emphasis id="B95E296DFFB00318FDFDF94EF22EF93A" box="[616,801,1698,1723]" italics="true" pageId="4" pageNumber="30">I. squamigerum</emphasis>
</taxonomicName>
.
<taxonomicName id="4C2A8EFCFFB00318FCA5F94EF2A5F93A" ID-CoL="6N5QN" box="[816,938,1698,1723]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="crassum">
<emphasis id="B95E296DFFB00318FCA5F94EF2A5F93A" box="[816,938,1698,1723]" italics="true" pageId="4" pageNumber="30">I. crassum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A8EFCFFB00318FC2EF94EF550F93A" ID-CoL="6MTQH" box="[955,1119,1698,1723]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="diacanthum">
<emphasis id="B95E296DFFB00318FC2EF94EF550F93A" box="[955,1119,1698,1723]" italics="true" pageId="4" pageNumber="30">I. diacanthum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2A8EFCFFB00318FB0CF94EF404F93A" ID-CoL="6N5QS" box="[1177,1291,1698,1723]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="dubium">
<emphasis id="B95E296DFFB00318FB0CF94EF404F93A" box="[1177,1291,1698,1723]" italics="true" pageId="4" pageNumber="30">I. dubium</emphasis>
</taxonomicName>
have spines but apparently lack scales which together with the different shape of the furca, differentiate them from
<taxonomicName id="4C2A8EFCFFB00318FA1FF92AF01AF882" ID-CoL="6MTQP" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="skandicum" status="sp. nov.">
<emphasis id="B95E296DFFB00318FA1FF92AF49BF95E" box="[1418,1428,1734,1759]" italics="true" pageId="4" pageNumber="30">I</emphasis>
.
<emphasis id="B95E296DFFB00318FF02F906F01AF882" box="[151,277,1770,1795]" italics="true" pageId="4" pageNumber="30">skandicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D9416FFB00318FE89F906F053F885" box="[284,348,1770,1796]" pageId="4" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
(
<bibRefCitation id="EFBB888EFFB00318FEF9F906F309F885" author="Daday" box="[364,518,1770,1796]" pageId="4" pageNumber="42" refString="Daday, E. V. (1905) Untersuchungen uber die Susswasser Mikrofauna Paraguays. Zoologica, 44, 1 - 374." type="journal article" year="1905">Daday, 1905</bibRefCitation>
;
<bibRefCitation id="EFBB888EFFB00318FD86F906F3C3F885" author="Schwank" box="[531,716,1770,1796]" pageId="4" pageNumber="43" refString="Schwank, P. (1990) Gastrotricha. In: Schwoerbel, J. &amp; Zwick, P. (eds), Susswasserfauna von Mitteleuropa, Band 3. Gastrotricha und Nemertini. Gustav Fischer Verlag, Stuttgart, Jena, New York, 252 pp." type="book" year="1990">Schwank, 1990</bibRefCitation>
;
<bibRefCitation id="EFBB888EFFB00318FD4CF906F507F885" author="Balsamo" box="[729,1032,1770,1796]" pageId="4" pageNumber="42" refString="Balsamo, M. &amp; Todaro, M. A. (1995) Gastrotrichi del Trentino: le Viotte del Monte Bondone. Studi Trentini di Scienze Naturali - Acta Biologica, 70, 9 - 22." type="journal article" year="1995">Balsamo &amp; Todaro, 1995</bibRefCitation>
).
<taxonomicName id="4C2A8EFCFFB00318FB88F906F5D8F882" ID-CoL="6MTQG" box="[1053,1239,1770,1795]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="squamigerum">
<emphasis id="B95E296DFFB00318FB88F906F5D8F882" box="[1053,1239,1770,1795]" italics="true" pageId="4" pageNumber="30">I. squamigerum</emphasis>
</taxonomicName>
has two pairs of keeled scales at the base of the furca compared to four pairs in
<taxonomicName id="4C2A8EFCFFB00318FC17F8E2F517F8A6" ID-CoL="6MTQP" box="[898,1048,1806,1831]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="skandicum" status="sp. nov.">
<emphasis id="B95E296DFFB00318FC17F8E2F517F8A6" box="[898,1048,1806,1831]" italics="true" pageId="4" pageNumber="30">I. skandicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D9416FFB00318FBB5F8E2F550F8A9" box="[1056,1119,1806,1832]" pageId="4" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
Moreover, at least another noticeable trait distinguishes these two species, that is the furca which is long (2830 µm) and tong-shaped in
<taxonomicName id="4C2A8EFCFFB00318FF02F8BAF023F8EE" ID-CoL="6MTQP" box="[151,300,1878,1903]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="skandicum" status="sp. nov.">
<emphasis id="B95E296DFFB00318FF02F8BAF023F8EE" box="[151,300,1878,1903]" italics="true" pageId="4" pageNumber="30">I. skandicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D9416FFB00318FEA1F8BAF07CF8F1" box="[308,371,1878,1904]" pageId="4" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
and short (1318 µm) and straight in
<taxonomicName id="4C2A8EFCFFB00318FCBAF8BAF2E7F8EE" ID-CoL="6MTQG" box="[815,1000,1878,1903]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="squamigerum">
<emphasis id="B95E296DFFB00318FCBAF8BAF2E7F8EE" box="[815,1000,1878,1903]" italics="true" pageId="4" pageNumber="30">I. squamigerum</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFBB888EFFB00318FBB0F8BAF444F8F1" author="Balsamo" box="[1061,1355,1878,1904]" pageId="4" pageNumber="42" refString="Balsamo, M. &amp; Fregni, E. (1995) Gastrotrichs from interstitial fresh water, with a description of four new species. Hydrobiologia, 302, 163 - 175." type="journal article" year="1995">Balsamo &amp; Fregni, 1995</bibRefCitation>
).
</paragraph>
<paragraph id="8B95F57FFFB00318FF50F896F507F812" blockId="4.[151,1437,1158,1976]" box="[197,1032,1914,1940]" pageId="4" pageNumber="30">
Hence,
<taxonomicName id="4C2A8EFCFFB00318FE8BF896F0BCF812" ID-CoL="6MTQP" box="[286,435,1914,1939]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="skandicum" status="sp. nov.">
<emphasis id="B95E296DFFB00318FE8BF896F0BCF812" box="[286,435,1914,1939]" italics="true" pageId="4" pageNumber="30">I. skandicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D9416FFB00318FE2EF896F0F5F815" box="[443,506,1914,1940]" pageId="4" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
is morphologically closest to
<emphasis id="B95E296DFFB00318FCCFF896F507F812" box="[858,1032,1914,1939]" italics="true" pageId="4" pageNumber="30">
<taxonomicName id="4C2A8EFCFFB00318FCCFF896F50BF812" ID-CoL="6N5QP" box="[858,1028,1914,1939]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="tanytrichum">I. tanytrichum</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C330A6F4FFB00318FF50F872F46DF839" box="[197,1378,1950,1976]" pageId="4" pageNumber="30" type="description">
<paragraph id="8B95F57FFFB00318FF50F872F46DF839" blockId="4.[151,1437,1158,1976]" box="[197,1378,1950,1976]" pageId="4" pageNumber="30">
<emphasis id="B95E296DFFB00318FF50F872F33BF839" bold="true" box="[197,564,1950,1976]" pageId="4" pageNumber="30">Associated gastrotrich fauna:</emphasis>
<taxonomicName id="4C2A8EFCFFB00318FDAEF872F503F839" authority="Kisielewski, 1981" authorityName="Kisielewski" authorityYear="1981" box="[571,1036,1950,1976]" family="Chaetonotidae" genus="Chaetonotus" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="oculifer">
<emphasis id="B95E296DFFB00318FDAEF872F23EF836" box="[571,817,1950,1975]" italics="true" pageId="4" pageNumber="30">Chaetonotus oculifer</emphasis>
<bibRefCitation id="EFBB888EFFB00318FCACF872F503F839" author="Kisielewski" box="[825,1036,1950,1976]" pageId="4" pageNumber="43" refString="Kisielewski, J. (1981) Gastrotricha from raised and transitional peat bogs in Poland. Monografie Fauny Polski, 11. Polska Akademia Nauk, Warszawa, 143 pp." type="book" year="1981">Kisielewski, 1981</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C2A8EFCFFB00318FBD3F872F454F836" box="[1094,1371,1950,1975]" family="Chaetonotidae" genus="Ichthydium" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="30" phylum="Gastrotricha" rank="species" species="diacanthum">
<emphasis id="B95E296DFFB00318FBD3F872F454F836" box="[1094,1371,1950,1975]" italics="true" pageId="4" pageNumber="30">Ichthydium diacanthum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>