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<document id="16C03A2364CCB44C5F41DD2070340087" ID-CLB-Dataset="28431" ID-DOI="10.11646/zootaxa.4564.1.2" ID-GBIF-Dataset="d3a56aac-92aa-48f2-b9fe-9576a4c2a0ec" ID-ISSN="1175-5326" ID-Zenodo-Dep="2588621" ID-ZooBank="27F70B5F-24DD-4C9E-BBA7-2B87476BD49C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552162070752" checkinUser="plazi" docAuthor="Gunter, Nicole L. &amp; Weir, Thomas A." docDate="2019" docId="03832713FFBE8D64A3DAFE44571F9017" docLanguage="en" docName="zootaxa.4564.1.2.pdf" docOrigin="Zootaxa 4564 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepanus feehani Gunter &amp; Weir 2019, new species" docType="treatment" docUuid="A4AA742E-9A6D-49FD-A952-DCB8C472ACD8" docUuidSource="ZooBank" docVersion="9" lastPageNumber="63" masterDocId="FFBA5F6BFFAB8D72A34DFFB65569910A" masterDocTitle="Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): key to species groups and description of 14 new species from the L. pygmaeus species group" masterLastPageNumber="80" masterPageNumber="41" pageNumber="62" updateTime="1698700752634" updateUser="ExternalLinkService">
<mods:mods id="BC4B12F6CCB17F4FB0BB61CE1B533C8E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="3C5C1D0E2C908D6BB44EE965A6D9BDB6">Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): key to species groups and description of 14 new species from the L. pygmaeus species group</mods:title>
</mods:titleInfo>
<mods:name id="D21DD054F5C3719CD133B883BD0DA3C3" type="personal">
<mods:role id="A402317F8AB0FD91534AC5D5F1C37E42">
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<mods:namePart id="69DABB75936F97DADF87111C545FDF86">Gunter, Nicole L.</mods:namePart>
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<mods:namePart id="523E581AB3E9143B32E66CC629D8A698">Weir, Thomas A.</mods:namePart>
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<mods:title id="D3129F024A25AE4D0AB8149492F6F381">Zootaxa</mods:title>
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<mods:part id="5B6ED762E62064544DB08B88F9CAC164">
<mods:date id="90543E348FCA8A54C6A0AFD2897D3188">2019</mods:date>
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<mods:number id="389F293FA5964000AE176058B6319323">2019-03-05</mods:number>
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<mods:number id="F007F8DF9227C60884D1B574B8082814">4564</mods:number>
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<mods:classification id="296796055145AC9AF2144ABF0357BE16">journal article</mods:classification>
<mods:identifier id="32A0BDE7737C031D56260E0EF5181FE3" type="CLB-Dataset">28431</mods:identifier>
<mods:identifier id="013EC45185E80A5115D6EADE07510987" type="DOI">10.11646/zootaxa.4564.1.2</mods:identifier>
<mods:identifier id="D8328F2D627755F9536A9DB88FB4D163" type="GBIF-Dataset">d3a56aac-92aa-48f2-b9fe-9576a4c2a0ec</mods:identifier>
<mods:identifier id="F746009CFCB6334AC0B6E502AE7C5FD7" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03832713FFBE8D64A3DAFE44571F9017" ID-DOI="http://doi.org/10.5281/zenodo.5922806" ID-GBIF-Taxon="154366347" ID-Zenodo-Dep="5922806" LSID="urn:lsid:zoobank.org:act:A4AA742E-9A6D-49FD-A952-DCB8C472ACD8" httpUri="http://treatment.plazi.org/id/03832713FFBE8D64A3DAFE44571F9017" lastPageId="22" lastPageNumber="63" pageId="21" pageNumber="62">
<subSubSection id="C330C58EFFBE8D67A3DAFE4457B49306" box="[151,733,498,524]" pageId="21" pageNumber="62" type="nomenclature">
<paragraph id="8B959605FFBE8D67A3DAFE4457B49306" blockId="21.[151,733,498,558]" box="[151,733,498,524]" pageId="21" pageNumber="62">
<heading id="D0DD2169FFBE8D67A3DAFE4457B49306" bold="true" box="[151,733,498,524]" fontSize="11" level="1" pageId="21" pageNumber="62" reason="1">
<emphasis id="B95E4A17FFBE8D67A3DAFE4457B49306" bold="true" box="[151,733,498,524]" pageId="21" pageNumber="62">
<taxonomicName id="4C2AED86FFBE8D67A3DAFE4457539306" ID-CoL="6P7VZ" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[151,570,498,524]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="62" phylum="Arthropoda" rank="species" species="feehani" status="sp. nov.">
<emphasis id="B95E4A17FFBE8D67A3DAFE4454039306" bold="true" box="[151,362,498,524]" italics="true" pageId="21" pageNumber="62">Lepanus feehani</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A26DF76CFFBE8D67A10AFE4457B49306" box="[583,733,498,524]" pageId="21" pageNumber="62" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFFBE8D67A3DAFDA354C39324" box="[151,426,533,558]" pageId="21" pageNumber="62" type="description">
<paragraph id="8B959605FFBE8D67A3DAFDA354C39324" blockId="21.[151,733,498,558]" box="[151,426,533,558]" pageId="21" pageNumber="62">
(
<figureCitation id="13118A80FFBE8D67A3D2FDA3546E9324" box="[159,263,533,558]" captionStart="FIGURE 5" captionStartId="15.[151,250,1945,1967]" captionTargetBox="[245,1336,223,1911]" captionTargetId="figure@15.[233,1355,193,1926]" captionTargetPageId="15" captionText="FIGURE 5. Habitus of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil. All specimens in comparative scale. Scale bar 1 mm." httpUri="https://zenodo.org/record/2588631/files/figure.png" pageId="21" pageNumber="62">Figs. 5C</figureCitation>
,
<figureCitation id="13118A80FFBE8D67A25EFDA0545E9324" box="[275,311,534,558]" captionStart="FIGURE 8" captionStartId="18.[151,250,1896,1918]" captionTargetBox="[190,1384,223,1858]" captionTargetId="figure@18.[173,1427,195,1873]" captionTargetPageId="18" captionText="FIGURE 8. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil; G, Lepanus lingziae; H, Lepanus mckenziei; I, Lepanus menendezae. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588637/files/figure.png" pageId="21" pageNumber="62">8C</figureCitation>
,
<figureCitation id="13118A80FFBE8D67A209FDA0541E9324" box="[324,375,534,558]" captionStart="FIGURE 10" captionStartId="23.[151,250,1909,1931]" captionTargetBox="[167,1333,208,1895]" captionTargetId="figure@23.[164,1428,193,1895]" captionTargetPageId="23" captionText="FIGURE 10. Left, dorsal, and right view of aedeagi of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar." httpUri="https://zenodo.org/record/2588641/files/figure.png" pageId="21" pageNumber="62">10C</figureCitation>
,
<figureCitation id="13118A80FFBE8D67A2C9FDA054CB9324" box="[388,418,534,558]" captionStart="FIGURE 14" captionStartId="31.[151,250,1220,1242]" captionTargetBox="[151,1436,375,1198]" captionTargetId="figure@31.[151,1436,375,1198]" captionTargetPageId="31" captionText="FIGURE 14. Map of known records for 10 species within the Lepanus pygmaeus species group overlaid with predicted distribution boundary of L. pygmaeus (dashed line) taken from Fig. 13 map." httpUri="https://zenodo.org/record/2588649/files/figure.png" pageId="21" pageNumber="62">14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFFBE8D67A3DAFDEB57179508" pageId="21" pageNumber="62" type="materials_examined">
<paragraph id="8B959605FFBE8D67A3DAFDEB57179508" blockId="21.[151,1437,605,2034]" pageId="21" pageNumber="62">
<emphasis id="B95E4A17FFBE8D67A3DAFDEB54C4937C" bold="true" box="[151,429,605,630]" pageId="21" pageNumber="62">
Type series.
<materialsCitation id="3B429C58FFBE8D67A27FFDEB54C4937C" ID-GBIF-Occurrence="2012745420" box="[306,429,605,630]" collectingDate="1980-10-05" collectingDateMax="79-06-03" collectingDateMin="1980-10-05" collectionCode="ANIC" collectorName="T. Weir" latitude="-15.17" location="Rounded Hill" longLatPrecision="555" longitude="145.21666" pageId="21" pageNumber="62" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="549128A7FFBE8D67A27FFDEB54C1937C" box="[306,424,605,630]" pageId="21" pageNumber="62" type="holotype">Holotype</typeStatus>
:
</materialsCitation>
</emphasis>
<specimenCount id="9D2C5D8CFFBE8D67A2FBFDE854A5937F" box="[438,460,606,629]" pageId="21" pageNumber="62" type="male">♂</specimenCount>
“
<geoCoordinate id="EE1EF0C2FFBE8D67A292FDEB5746937C" box="[479,559,605,630]" direction="south" orientation="latitude" pageId="21" pageNumber="62" precision="555" value="-15.17">15.17S</geoCoordinate>
<geoCoordinate id="EE1EF0C2FFBE8D67A17AFDEB57FC937C" box="[567,661,605,630]" direction="east" orientation="longitude" pageId="21" pageNumber="62" precision="555" value="145.13">145.13E</geoCoordinate>
<quantity id="4CD23BE0FFBE8D67A1D0FDEB57A6937C" box="[669,719,605,630]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="62" unit="km" value="1.0">1km</quantity>
N of
<location id="8EF5C0DEFFBE8D67A05FFDEB56C2937C" LSID="urn:lsid:plazi:treatment:03832713FFBE8D64A3DAFE44571F9017:8EF5C0DEFFBE8D67A05FFDEB56C2937C" box="[786,939,605,630]" latitude="-15.17" longLatPrecision="555" longitude="145.21666" name="Rounded Hill" pageId="21" pageNumber="62">Rounded Hill</location>
QLD,
<date id="FF94B0C5FFBE8D67A0B7FDEB51FE937C" box="[1018,1175,605,630]" pageId="21" pageNumber="62" value="1980-10-05" valueMax="1980-10-06" valueMin="1980-10-05">
<collectingDate id="EFD0492DFFBE8D67A0B7FDEB51FE937C" box="[1018,1175,605,630]" pageId="21" pageNumber="62" value="1980-10-05" valueMax="1980-10-06" valueMin="1980-10-05">5-6 Oct. 1980</collectingDate>
</date>
<collectorName id="26DFF3D3FFBE8D67A7D3FDE85191937C" box="[1182,1272,605,630]" pageId="21" pageNumber="62">T. Weir</collectorName>
” [15°17’00’S,
<geoCoordinate id="EE1EF0C2FFBE8D67A3DAFD3754439390" box="[151,298,641,666]" direction="east" orientation="longitude" pageId="21" pageNumber="62" precision="15" value="145.21666">145°13’00”E</geoCoordinate>
]/ “
<collectingMethod id="526BEE12FFBE8D67A21EFD3757499393" box="[339,544,641,666]" pageId="21" pageNumber="62">human dung trap</collectingMethod>
open forest” / “more in spirit tube” / “ANIC Database No. 25-
<date id="FF94B0C5FFBE8D67A669FD3750149390" box="[1316,1405,641,666]" pageId="21" pageNumber="62" value="79-06-03">
<collectingDate id="EFD0492DFFBE8D67A669FD3750149390" box="[1316,1405,641,666]" pageId="21" pageNumber="62" value="79-06-03">060379</collectingDate>
</date>
” / “
<taxonomicName id="4C2AED86FFBE8D67A3ECFD10546B93B4" authorityName=", Matthews" authorityYear="1974" box="[161,258,678,702]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="79" phylum="Arthropoda" rank="genus">Lepanus</taxonomicName>
NQ26 det.
<bibRefCitation id="EFBBEBF4FFBE8D67A2C3FD13575793B4" author="Gunter, N. L. &amp; Monteith, G. B. &amp; Cameron, S. L. &amp; Weir, T." box="[398,574,677,702]" pageId="21" pageNumber="62" pagination="1 - 27" refId="ref29280" refString="Gunter, N. L., Monteith, G. B., Cameron, S. L. &amp; Weir, T. A (2018) Evidence from Australian mesic zone dung beetles support Gondwanan vicariance and Mesozoic origin of the Scarabaeinae (Coleoptera: Scarabaeidae). Invertebrate Systematics and Evolution, 1 - 27. [Advance online publication] https: // doi. org / 10.1163 / 1876312 X- 00002171" type="book chapter" year="2018">T.A.Weir 2018</bibRefCitation>
” in ANIC.
<materialsCitation id="3B429C58FFBE8D67A184FD13560693E8" ID-GBIF-Occurrence="2012745407" collectingDate="1980-10-05" collectingDateMax="1980-10-06" collectingDateMin="1980-10-05" collectionCode="ANIC" collectorName="T. A. Weir" latitude="-15.283334" location="1 km N of Rounded Hill" longLatPrecision="1285" longitude="145.21666" pageId="21" pageNumber="62" specimenCount="2" specimenCount-male="2" stateProvince="Queensland" typeStatus="paratype">
<emphasis id="B95E4A17FFBE8D67A184FD13569E93B4" bold="true" box="[713,1015,677,702]" pageId="21" pageNumber="62">
<typeStatus id="549128A7FFBE8D67A184FD13562393B4" box="[713,842,677,702]" pageId="21" pageNumber="62" type="paratype">Paratypes</typeStatus>
:
<collectingRegion id="49EE58E7FFBE8D67A017FD13569B93B4" box="[858,1010,677,702]" country="Australia" name="Queensland" pageId="21" pageNumber="62">Queensland</collectingRegion>
:
</emphasis>
<location id="8EF5C0DEFFBE8D67A74FFD10505D93B4" LSID="urn:lsid:plazi:treatment:03832713FFBE8D64A3DAFE44571F9017:8EF5C0DEFFBE8D67A74FFD10505D93B4" box="[1026,1332,677,702]" latitude="-15.283334" longLatPrecision="1285" longitude="145.21666" name="1 km N of Rounded Hill" pageId="21" pageNumber="62" stateProvince="Queensland">
<quantity id="4CD23BE0FFBE8D67A74FFD10512893B4" box="[1026,1089,677,702]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="62" unit="km" value="1.0">1 km</quantity>
N of Rounded Hill
</location>
,
<geoCoordinate id="EE1EF0C2FFBE8D67A60EFD1350F593B4" box="[1347,1436,677,702]" direction="south" orientation="latitude" pageId="21" pageNumber="62" precision="925" value="-15.283334">15°17'S</geoCoordinate>
<geoCoordinate id="EE1EF0C2FFBE8D67A3DAFD7F546B93E8" box="[151,258,713,738]" direction="east" orientation="longitude" pageId="21" pageNumber="62" precision="925" value="145.21666">145°13'E</geoCoordinate>
,
<date id="FF94B0C5FFBE8D67A242FD7F54FC93E8" box="[271,405,713,738]" pageId="21" pageNumber="62" value="1980-10-05" valueMax="1980-10-06" valueMin="1980-10-05">
<collectingDate id="EFD0492DFFBE8D67A242FD7F54FC93E8" box="[271,405,713,738]" pageId="21" pageNumber="62" value="1980-10-05" valueMax="1980-10-06" valueMin="1980-10-05">5–6.x.1980</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFBE8D67A2ECFD7C577D93E8" box="[417,532,713,738]" pageId="21" pageNumber="62">T.A. Weir</collectorName>
(25-065530 ×
<specimenCount id="9D2C5D8CFFBE8D67A18EFD7C578793EB" box="[707,750,714,738]" pageId="21" pageNumber="62" type="male">2 ♂</specimenCount>
in
<collectionCode id="ED3B0EC0FFBE8D67A056FD7C560E93E8" box="[795,871,714,738]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="21" pageNumber="62">ANIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFBE8D67A036FD7C5608920C" ID-GBIF-Occurrence="2012745418" collectingDate="1981-05-05" collectingDateMax="1981-05-07" collectingDateMin="1981-05-05" collectionCode="ANIC" collectorName="A. Calder &amp; Feehan" latitude="-15.283334" location="1 km N of Rounded Hill" longLatPrecision="1285" longitude="145.21666" pageId="21" pageNumber="62" specimenCount="1" specimenCount-female="1" stateProvince="Queensland" typeStatus="paratype">
<location id="8EF5C0DEFFBE8D67A036FD7C51F393E8" LSID="urn:lsid:plazi:treatment:03832713FFBE8D64A3DAFE44571F9017:8EF5C0DEFFBE8D67A036FD7C51F393E8" box="[891,1178,713,738]" latitude="-15.283334" longLatPrecision="1285" longitude="145.21666" name="1 km N of Rounded Hill" pageId="21" pageNumber="62" stateProvince="Queensland">
<quantity id="4CD23BE0FFBE8D67A036FD7C56DF93E8" box="[891,950,713,738]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="62" unit="km" value="1.0">1 km</quantity>
N of Rounded Hill
</location>
,
<geoCoordinate id="EE1EF0C2FFBE8D67A7EBFD7F519693E8" box="[1190,1279,713,738]" direction="south" orientation="latitude" pageId="21" pageNumber="62" precision="925" value="-15.283334">15°17'S</geoCoordinate>
<geoCoordinate id="EE1EF0C2FFBE8D67A64AFD7F501B93E8" box="[1287,1394,713,738]" direction="east" orientation="longitude" pageId="21" pageNumber="62" precision="925" value="145.21666">145°13'E</geoCoordinate>
,
<date id="FF94B0C5FFBE8D67A632FD7F5597920C" pageId="21" pageNumber="62" value="1981-05-05" valueMax="1981-05-07" valueMin="1981-05-05">
<collectingDate id="EFD0492DFFBE8D67A632FD7F5597920C" pageId="21" pageNumber="62" value="1981-05-05" valueMax="1981-05-07" valueMin="1981-05-05">5– 7.v.1981</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFBE8D67A246FD585414920C" box="[267,381,749,774]" pageId="21" pageNumber="62">A. Calder</collectorName>
&amp; J.
<collectorName id="26DFF3D3FFBE8D67A2F2FD5B577A920C" box="[447,531,749,774]" pageId="21" pageNumber="62">Feehan</collectorName>
(25-
<date id="FF94B0C5FFBE8D67A105FD5B57CD920C" box="[584,676,749,774]" pageId="21" pageNumber="62" value="73-06-03">060373</date>
<date id="FF94B0C5FFBE8D67A1E6FD5857A1920C" box="[683,712,750,774]" pageId="21" pageNumber="62">
x
<specimenCount id="9D2C5D8CFFBE8D67A1F0FD5857A1920C" box="[701,712,750,774]" pageId="21" pageNumber="62" type="female">1</specimenCount>
</date>
♀ in
<collectionCode id="ED3B0EC0FFBE8D67A040FD585630920C" box="[781,857,750,774]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="21" pageNumber="62">ANIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFBE8D67A023FD5856089220" ID-GBIF-Occurrence="2012745410" collectingDate="1980-10-05" collectingDateMax="1980-10-06" collectingDateMin="1980-10-05" collectionCode="ANIC" latitude="-15.283334" location="Cardale" longLatPrecision="1285" longitude="145.21666" pageId="21" pageNumber="62" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Queensland" typeStatus="paratype">
<locationDeviation id="9A278F27FFBE8D67A023FD5851F8920C" box="[878,1169,749,774]" location="Rounded Hill" pageId="21" pageNumber="62">
<quantity id="4CD23BE0FFBE8D67A023FD5856C0920C" box="[878,937,749,774]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="62" unit="km" value="1.0">1 km</quantity>
N of Rounded Hill
</locationDeviation>
,
<geoCoordinate id="EE1EF0C2FFBE8D67A7D0FD5B519F920C" box="[1181,1270,749,774]" direction="south" orientation="latitude" pageId="21" pageNumber="62" precision="925" value="-15.283334">15°17'S</geoCoordinate>
<geoCoordinate id="EE1EF0C2FFBE8D67A7B3FD5B5001920C" box="[1278,1384,749,774]" direction="east" orientation="longitude" pageId="21" pageNumber="62" precision="925" value="145.21666">145°13'E</geoCoordinate>
,
<date id="FF94B0C5FFBE8D67A632FD5B54699220" pageId="21" pageNumber="62" value="1980-10-05" valueMax="1980-10-06" valueMin="1980-10-05">
<collectingDate id="EFD0492DFFBE8D67A632FD5B54699220" pageId="21" pageNumber="62" value="1980-10-05" valueMax="1980-10-06" valueMin="1980-10-05">5– 6.x.1980</collectingDate>
</date>
, J.C.
<location id="8EF5C0DEFFBE8D67A204FCA754CD9220" LSID="urn:lsid:plazi:treatment:03832713FFBE8D64A3DAFE44571F9017:8EF5C0DEFFBE8D67A204FCA754CD9220" box="[329,420,785,810]" latitude="-15.283334" longLatPrecision="1285" longitude="145.21666" name="Cardale" pageId="21" pageNumber="62" stateProvince="Queensland">Cardale</location>
(25-
<collectingDate id="EFD0492DFFBE8D67A291FCA757099220" box="[476,608,785,810]" isEnumeration="true" pageId="21" pageNumber="62">
<date id="FF94B0C5FFBE8D67A291FCA757519220" box="[476,568,785,810]" pageId="21" pageNumber="62" value="78-06-03">060378</date>
<date id="FF94B0C5FFBE8D67A10EFCA457099220" box="[579,608,786,810]" pageId="21" pageNumber="62">
x
<specimenCount id="9D2C5D8CFFBE8D67A11FFCA457099220" box="[594,608,786,810]" pageId="21" pageNumber="62" type="male">2</specimenCount>
</date>
</collectingDate>
♂, ×
<specimenCount id="9D2C5D8CFFBE8D67A1E2FCA457B69223" box="[687,735,786,810]" pageId="21" pageNumber="62" type="female">1 ♀</specimenCount>
in
<collectionCode id="ED3B0EC0FFBE8D67A040FCA456309220" box="[781,857,786,810]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="21" pageNumber="62">ANIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFBE8D67A03CFCA457B49244" ID-GBIF-Occurrence="2012745411" collectingDate="1980-10-01" collectingDateMax="76-06-03" collectingDateMin="1980-10-01" collectionCode="ANIC" latitude="-15.05" location="Weir" longLatPrecision="1286" longitude="145.15" pageId="21" pageNumber="62" specimenCount="1" specimenCount-female="1" stateProvince="Queensland" typeStatus="paratype">
<locationDeviation id="9A278F27FFBE8D67A03CFCA451E29220" box="[881,1163,785,810]" location="Mount Webb" pageId="21" pageNumber="62">
<quantity id="4CD23BE0FFBE8D67A03CFCA456D99220" box="[881,944,785,810]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="21" pageNumber="62" unit="km" value="3.0">3 km</quantity>
NE Mount Webb
</locationDeviation>
, [
<geoCoordinate id="EE1EF0C2FFBE8D67A7EEFCA7519D9220" box="[1187,1268,785,810]" direction="south" orientation="latitude" pageId="21" pageNumber="62" precision="925" value="-15.05">15°3'S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFBE8D67A648FCA7500E9220" box="[1285,1383,785,810]" direction="east" orientation="longitude" pageId="21" pageNumber="62" precision="925" value="145.15">145°9'E</geoCoordinate>
],
<collectingDate id="EFD0492DFFBE8D67A632FCA454699244" pageId="21" pageNumber="62" value="1980-10-01" valueMax="1980-10-03" valueMin="1980-10-01">
<date id="FF94B0C5FFBE8D67A632FCA454699244" pageId="21" pageNumber="62" value="1980-10-01" valueMax="1980-10-03" valueMin="1980-10-01">1– 3.x.1980</date>
</collectingDate>
, T.A.
<location id="8EF5C0DEFFBE8D67A207FC8354EB9244" LSID="urn:lsid:plazi:treatment:03832713FFBE8D64A3DAFE44571F9017:8EF5C0DEFFBE8D67A207FC8354EB9244" box="[330,386,821,846]" latitude="-15.05" longLatPrecision="1286" longitude="145.15" name="Weir" pageId="21" pageNumber="62" stateProvince="Queensland">Weir</location>
(25-
<date id="FF94B0C5FFBE8D67A2FBFC83577B9244" box="[438,530,821,846]" pageId="21" pageNumber="62" value="76-06-03">
<collectingDate id="EFD0492DFFBE8D67A2FBFC83577B9244" box="[438,530,821,846]" pageId="21" pageNumber="62" value="76-06-03">060376</collectingDate>
</date>
×
<specimenCount id="9D2C5D8CFFBE8D67A179FC8057099247" box="[564,608,822,846]" pageId="21" pageNumber="62" type="female">1 ♀</specimenCount>
in
<collectionCode id="ED3B0EC0FFBE8D67A1C4FC8057BC9244" box="[649,725,822,846]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="21" pageNumber="62">ANIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFBE8D67A1A7FC8057969278" ID-GBIF-Occurrence="2012745419" collectingDate="1981-04-30" collectingDateMax="72-06-03" collectingDateMin="1981-04-30" collectionCode="ANIC" collectorName="A. Calder" latitude="-15.05" location="Feehan" longLatPrecision="1286" longitude="145.15" pageId="21" pageNumber="62" specimenCount="1" specimenCount-female="1" stateProvince="Queensland" typeStatus="paratype">
<locationDeviation id="9A278F27FFBE8D67A1A7FC8056939244" box="[746,1018,821,846]" location="Mount Webb" pageId="21" pageNumber="62">
<quantity id="4CD23BE0FFBE8D67A1A7FC80564F9244" box="[746,806,821,846]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="21" pageNumber="62" unit="km" value="3.0">3 km</quantity>
NE Mount Webb
</locationDeviation>
, [
<geoCoordinate id="EE1EF0C2FFBE8D67A75DFC8351089244" box="[1040,1121,821,846]" direction="south" orientation="latitude" pageId="21" pageNumber="62" precision="925" value="-15.05">15°3'S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFBE8D67A723FC8351B99244" box="[1134,1232,821,846]" direction="east" orientation="longitude" pageId="21" pageNumber="62" precision="925" value="145.15">145°9'E</geoCoordinate>
],
<date id="FF94B0C5FFBE8D67A7ABFC8350FE9244" box="[1254,1431,821,846]" pageId="21" pageNumber="62" value="1981-04-30" valueMax="1981-05-03" valueMin="1981-04-30">
<collectingDate id="EFD0492DFFBE8D67A7ABFC8350FE9244" box="[1254,1431,821,846]" pageId="21" pageNumber="62" value="1981-04-30" valueMax="1981-05-03" valueMin="1981-04-30">30.iv–3.v.1981</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFBE8D67A3DAFCEC54639278" box="[151,266,857,882]" pageId="21" pageNumber="62">A. Calder</collectorName>
&amp; J.
<location id="8EF5C0DEFFBE8D67A203FCEF54CB9278" LSID="urn:lsid:plazi:treatment:03832713FFBE8D64A3DAFE44571F9017:8EF5C0DEFFBE8D67A203FCEF54CB9278" box="[334,418,857,882]" latitude="-15.05" longLatPrecision="1286" longitude="145.15" name="Feehan" pageId="21" pageNumber="62" stateProvince="Queensland">Feehan</location>
(25-
<date id="FF94B0C5FFBE8D67A29AFCEF575A9278" box="[471,563,857,882]" pageId="21" pageNumber="62" value="72-06-03">
<collectingDate id="EFD0492DFFBE8D67A29AFCEF575A9278" box="[471,563,857,882]" pageId="21" pageNumber="62" value="72-06-03">060372</collectingDate>
</date>
×
<specimenCount id="9D2C5D8CFFBE8D67A118FCEC57EB927B" box="[597,642,858,882]" pageId="21" pageNumber="62" type="female">1 ♀</specimenCount>
in
<collectionCode id="ED3B0EC0FFBE8D67A1E7FCEC579F9278" box="[682,758,858,882]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="21" pageNumber="62">ANIC</collectionCode>
)
</materialsCitation>
; Cape Flattery Heath,
<materialsCitation id="3B429C58FFBE8D67A746FCEF576B92B0" ID-GBIF-Occurrence="2012745415" collectingDate="1976-07-13" collectingDateMax="1976-07-14" collectingDateMin="1976-07-13" collectionCode="QM" collectorName="G. B. &amp; S. R. Monteith" elevation="20" latitude="-15.083333" location="Cooktown" longLatPrecision="21" longitude="145.26666" pageId="21" pageNumber="62" specimenCount="6" specimenCount-female="1" specimenCount-male="5" stateProvince="Queensland" typeStatus="paratype">
<quantity id="4CD23BE0FFBE8D67A746FCEF513F9278" box="[1035,1110,857,882]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="21" pageNumber="62" unit="km" value="45.0">45 km</quantity>
N
<location id="8EF5C0DEFFBE8D67A731FCEF51919278" LSID="urn:lsid:plazi:treatment:03832713FFBE8D64A3DAFE44571F9017:8EF5C0DEFFBE8D67A731FCEF51919278" box="[1148,1272,857,882]" latitude="-15.083333" longLatPrecision="21" longitude="145.26666" name="Cooktown" pageId="21" pageNumber="62" stateProvince="Queensland">Cooktown</location>
, [
<geoCoordinate id="EE1EF0C2FFBE8D67A643FCEF50FE9278" box="[1294,1431,857,882]" direction="south" orientation="latitude" pageId="21" pageNumber="62" precision="15" value="-15.083333">15°05'00&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFBE8D67A3DAFCCB5458929C" box="[151,305,893,918]" direction="east" orientation="longitude" pageId="21" pageNumber="62" precision="15" value="145.26666">145°16'00&quot;E</geoCoordinate>
],
<quantity id="4CD23BE0FFBE8D67A208FCC854EB929F" box="[325,386,894,918]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="21" pageNumber="62" unit="m" value="20.0">
<elevation id="00077136FFBE8D67A208FCC854EB929F" box="[325,386,894,918]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="21" pageNumber="62" unit="m" value="20.0">20 m</elevation>
</quantity>
,
<date id="FF94B0C5FFBE8D67A2C2FCCB572A929C" box="[399,579,893,918]" pageId="21" pageNumber="62" value="1976-07-13" valueMax="1976-07-14" valueMin="1976-07-13">
<collectingDate id="EFD0492DFFBE8D67A2C2FCCB572A929C" box="[399,579,893,918]" pageId="21" pageNumber="62" value="1976-07-13" valueMax="1976-07-14" valueMin="1976-07-13">13–14.vii.1976</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFBE8D67A102FCC85624929C" box="[591,845,893,918]" pageId="21" pageNumber="62">G.B. &amp; S.R. Monteith</collectorName>
(25-015583
<date id="FF94B0C5FFBE8D67A0A8FCCB516B929C" box="[997,1026,893,918]" pageId="21" pageNumber="62">
x
<specimenCount id="9D2C5D8CFFBE8D67A0B9FCCB516B929C" box="[1012,1026,893,918]" pageId="21" pageNumber="62" type="male">5</specimenCount>
</date>
♂, 4 ×
<specimenCount id="9D2C5D8CFFBE8D67A711FCC8511B929F" box="[1116,1138,894,917]" pageId="21" pageNumber="62" type="female">♀</specimenCount>
, 3 × sex undetermined in
<collectionCode id="ED3B0EC0FFBE8D67A3DAFC1455AC92B0" box="[151,197,930,954]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="21" pageNumber="62" type="Museum">QM</collectionCode>
also as T50127–T50138)
</materialsCitation>
;
<materialsCitation id="3B429C58FFBE8D67A15FFC1454DF92D4" ID-GBIF-Occurrence="2012745390" collectingDate="1980-10-04" collectionCode="ANIC" collectorName="Weir" latitude="-15.233334" location="7 km N Hope Vale Mission" longLatPrecision="1285" longitude="145.11667" pageId="21" pageNumber="62" specimenCount="1" specimenCount-female="1" stateProvince="Queensland" typeStatus="paratype">
<location id="8EF5C0DEFFBE8D67A15FFC14560392B0" LSID="urn:lsid:plazi:treatment:03832713FFBE8D64A3DAFE44571F9017:8EF5C0DEFFBE8D67A15FFC14560392B0" box="[530,874,929,954]" latitude="-15.233334" longLatPrecision="1285" longitude="145.11667" name="7 km N Hope Vale Mission" pageId="21" pageNumber="62" stateProvince="Queensland">
<quantity id="4CD23BE0FFBE8D67A15FFC14573B92B0" box="[530,594,929,954]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="21" pageNumber="62" unit="km" value="7.0">7 km</quantity>
N Hope Vale Mission
</location>
, [
<geoCoordinate id="EE1EF0C2FFBE8D67A0CFFC17568B92B0" box="[898,994,929,954]" direction="south" orientation="latitude" pageId="21" pageNumber="62" precision="925" value="-15.233334">15°14'S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFBE8D67A0BEFC17513C92B0" box="[1011,1109,929,954]" direction="east" orientation="longitude" pageId="21" pageNumber="62" precision="925" value="145.11667">145°7'E</geoCoordinate>
],
<date id="FF94B0C5FFBE8D67A723FC1451BE92B0" box="[1134,1239,930,954]" pageId="21" pageNumber="62" value="1980-10-04">
<collectingDate id="EFD0492DFFBE8D67A723FC1451BE92B0" box="[1134,1239,930,954]" pageId="21" pageNumber="62" value="1980-10-04">4.x.1980</collectingDate>
</date>
, T.A.
<collectorName id="26DFF3D3FFBE8D67A66AFC17503692B0" box="[1319,1375,929,954]" pageId="21" pageNumber="62">Weir</collectorName>
(25- 0 60377
<date id="FF94B0C5FFBE8D67A3B4FC70547F92D4" box="[249,278,966,990]" pageId="21" pageNumber="62">
x
<specimenCount id="9D2C5D8CFFBE8D67A246FC70547F92D4" box="[267,278,966,990]" pageId="21" pageNumber="62" type="female">1</specimenCount>
</date>
♀ in
<collectionCode id="ED3B0EC0FFBE8D67A22CFC7054C492D4" box="[353,429,966,990]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="21" pageNumber="62">ANIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFBE8D67A288FC7057139508" ID-GBIF-Occurrence="2012745417" collectingDate="1981-05-07" collectingDateMax="1981-05-10" collectingDateMin="1981-05-07" collectionCode="ANIC" collectorName="Feehan" latitude="-15.266666" location="14 km NW Hope Vale Mission" longLatPrecision="1285" longitude="144.98334" pageId="21" pageNumber="62" specimenCount="2" specimenCount-female="2" stateProvince="Queensland" typeStatus="paratype">
<location id="8EF5C0DEFFBE8D67A288FC70565692D4" LSID="urn:lsid:plazi:treatment:03832713FFBE8D64A3DAFE44571F9017:8EF5C0DEFFBE8D67A288FC70565692D4" box="[453,831,965,990]" latitude="-15.266666" longLatPrecision="1285" longitude="144.98334" name="14 km NW Hope Vale Mission" pageId="21" pageNumber="62" stateProvince="Queensland">
<quantity id="4CD23BE0FFBE8D67A288FC70577892D4" box="[453,529,965,990]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="21" pageNumber="62" unit="km" value="14.0">14 km</quantity>
NW Hope Vale Mission
</location>
, [
<geoCoordinate id="EE1EF0C2FFBE8D67A01BFC7356DC92D4" box="[854,949,965,990]" direction="south" orientation="latitude" pageId="21" pageNumber="62" precision="925" value="-15.266666">15°16'S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFBE8D67A088FC73515F92D4" box="[965,1078,965,990]" direction="east" orientation="longitude" pageId="21" pageNumber="62" precision="925" value="144.98334">144°59'E</geoCoordinate>
],
<date id="FF94B0C5FFBE8D67A700FC70518992D4" box="[1101,1248,966,990]" pageId="21" pageNumber="62" value="1981-05-07" valueMax="1981-05-10" valueMin="1981-05-07">
<collectingDate id="EFD0492DFFBE8D67A700FC70518992D4" box="[1101,1248,966,990]" pageId="21" pageNumber="62" value="1981-05-07" valueMax="1981-05-10" valueMin="1981-05-07">7–10.v.1981</collectingDate>
</date>
, J.
<collectorName id="26DFF3D3FFBE8D67A640FC73500892D4" box="[1293,1377,965,990]" pageId="21" pageNumber="62">Feehan</collectorName>
(25- 0 60374
<date id="FF94B0C5FFBE8D67A3BBFC5C547A9508" box="[246,275,1002,1026]" pageId="21" pageNumber="62">
x
<specimenCount id="9D2C5D8CFFBE8D67A245FC5C547A9508" box="[264,275,1002,1026]" pageId="21" pageNumber="62" type="female">1</specimenCount>
</date>
♀, 25-
<collectingDate id="EFD0492DFFBE8D67A22EFC5F548B9508" box="[355,482,1001,1026]" isEnumeration="true" pageId="21" pageNumber="62">
<date id="FF94B0C5FFBE8D67A22EFC5F54D79508" box="[355,446,1001,1026]" pageId="21" pageNumber="62" value="75-06-03">060375</date>
<date id="FF94B0C5FFBE8D67A288FC5C548B9508" box="[453,482,1002,1026]" pageId="21" pageNumber="62">
x
<specimenCount id="9D2C5D8CFFBE8D67A29AFC5C548B9508" box="[471,482,1002,1026]" pageId="21" pageNumber="62" type="female">1</specimenCount>
</date>
</collectingDate>
♀ in
<collectionCode id="ED3B0EC0FFBE8D67A168FC5C571B9508" box="[549,626,1002,1026]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="21" pageNumber="62">ANIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFFBE8D67A38AFBBB57999634" pageId="21" pageNumber="62" type="description">
<paragraph id="8B959605FFBE8D67A38AFBBB5180952C" blockId="21.[151,1437,605,2034]" box="[199,1257,1037,1062]" pageId="21" pageNumber="62">
<emphasis id="B95E4A17FFBE8D67A38AFBBB5436952C" bold="true" box="[199,351,1037,1062]" pageId="21" pageNumber="62">Description.</emphasis>
Head and pronotum dark brown and elytra light brown. Antennal clubs grey.
</paragraph>
<paragraph id="8B959605FFBE8D67A38AFB87504C9543" blockId="21.[151,1437,605,2034]" box="[199,1317,1073,1098]" pageId="21" pageNumber="62">
Total length: 2.0–
<quantity id="4CD23BE0FFBE8D67A2D5FB84549D9543" box="[408,500,1074,1098]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="21" pageNumber="62" unit="mm" value="2.3">2.3 mm</quantity>
.
<typeStatus id="549128A7FFBE8D67A2B2FB8757059540" box="[511,620,1073,1098]" pageId="21" pageNumber="62" type="holotype">Holotype</typeStatus>
measurements: Total length
<quantity id="4CD23BE0FFBE8D67A0F3FB8451729543" box="[958,1051,1074,1098]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="21" pageNumber="62" unit="mm" value="2.2">2.2 mm</quantity>
, elytral width
<quantity id="4CD23BE0FFBE8D67A789FB8450489543" box="[1220,1313,1074,1098]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="21" pageNumber="62" unit="mm" value="1.4">1.4 mm</quantity>
.
</paragraph>
<paragraph id="8B959605FFBE8D67A38AFBE357B795D0" blockId="21.[151,1437,605,2034]" pageId="21" pageNumber="62">Male: Head: U-shaped between clypeal teeth, which are short and upturned. Margin slightly angulate at clypeogenal suture. Genal angle rounded. Basal carina present but interrupted medially. Dorsal part of eyes narrow in size, separated by interocular space approximately 20 times eye width (30: 1.5); eye canthus not dividing eye. Head 1.2 × wider than long (38:32).</paragraph>
<paragraph id="8B959605FFBE8D67A38AFB5356119428" blockId="21.[151,1437,605,2034]" pageId="21" pageNumber="62">Prothorax: Hypomeral surface finely reticulate with light punctation. Hypomeral striae about half the length of hypomeron. Pronotum 1.7 × wider than long (62:36).</paragraph>
<paragraph id="8B959605FFBE8D67A38AFA9B564B9484" blockId="21.[151,1437,605,2034]" pageId="21" pageNumber="62">Elytra: Surface smooth. Striae 1–4 very narrow and double but almost appearing single, striae 5–7 single, striae 6–7 with discernible punctures in the basal half. All striae on disc equal width. Elytra 1.1 × wider than long (74:68). Elytral flange and groove relatively wide.</paragraph>
<paragraph id="8B959605FFBE8D67A38AFA2C54A094F0" blockId="21.[151,1437,605,2034]" pageId="21" pageNumber="62">Legs: Front edge of protibiae deeply angulate; apical digit elongate and sharp; 2 teeth on outer edge. Inner edge of metatibiae almost straight. Metatarsi with 2nd tarsomere subequal to the 5th. Tarsal claws minute, appearing simple.</paragraph>
<paragraph id="8B959605FFBE8D67A38AF9B3546B9783" blockId="21.[151,1437,605,2034]" pageId="21" pageNumber="62">Abdomen: Pygidium with almost circular depression containing a tubercle appearing like an inverted V in upper third; tip of tubercle darker. Depression occupies more than half of the pygidial disc and extends under the slightly angulate upper edge of pygidium. Abdominal surface slightly reticulate right across. Segment 6 punctate.</paragraph>
<paragraph id="8B959605FFBE8D67A38AF92354649610" blockId="21.[151,1437,605,2034]" pageId="21" pageNumber="62">Pterothorax: Medial lobe of metaventrite finely punctate and narrowly margined between mesocoxae. Lateral lobe of metaventrite smooth and punctate with setae. Meso-metaventral suture very slightly arched. Mesoventrite with slight notch anteriorly. Mesoventrite and mesepimeron smooth. Metanepisternum reticulate.</paragraph>
<paragraph id="8B959605FFBE8D67A38AF89357999634" blockId="21.[151,1437,605,2034]" box="[199,752,1829,1854]" pageId="21" pageNumber="62">
Female: As per the
<taxonomicName id="4C2AED86FFBE8D67A2E1F89057289637" authorityName=": Matthews" authorityYear="1974" box="[428,577,1830,1853]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="62" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFBE8D67A2E1F89054D29637" box="[428,443,1830,1853]" italics="true" pageId="21" pageNumber="62">L</emphasis>
.
<emphasis id="B95E4A17FFBE8D67A281F89057289637" box="[460,577,1830,1853]" italics="true" pageId="21" pageNumber="62">pygmaeus</emphasis>
</taxonomicName>
species group.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFFBE8D67A38AF8FF56FA968C" pageId="21" pageNumber="62" type="etymology">
<paragraph id="8B959605FFBE8D67A38AF8FF56FA968C" blockId="21.[151,1437,605,2034]" pageId="21" pageNumber="62">
<emphasis id="B95E4A17FFBE8D67A38AF8FF543A9668" bold="true" box="[199,339,1865,1890]" pageId="21" pageNumber="62">Etymology.</emphasis>
Named in honour of John Feehan, who has collected many of the specimens in the
<typeStatus id="549128A7FFBE8D67A67AF8FC50F59668" box="[1335,1436,1866,1890]" pageId="21" pageNumber="62" type="paratype">paratype</typeStatus>
series and for his contribution to Australian Dung Beetle project.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFFBE8D67A38AF827566696F8" pageId="21" pageNumber="62" type="distribution">
<paragraph id="8B959605FFBE8D67A38AF827566696F8" blockId="21.[151,1437,605,2034]" pageId="21" pageNumber="62">
<emphasis id="B95E4A17FFBE8D67A38AF827540E96A0" bold="true" box="[199,359,1937,1962]" pageId="21" pageNumber="62">Distribution.</emphasis>
Known only from
<specimenCount id="9D2C5D8CFFBE8D67A10AF824578C96A0" box="[583,741,1937,1962]" pageId="21" pageNumber="62" type="generic">23 specimens</specimenCount>
collected from low elevation open and closed forests north of Cooktown in the WET bioregion (
<figureCitation id="13118A80FFBE8D67A124F80357AE96C4" box="[617,711,1973,1998]" captionStart="FIGURE 14" captionStartId="31.[151,250,1220,1242]" captionTargetBox="[151,1436,375,1198]" captionTargetId="figure@31.[151,1436,375,1198]" captionTargetPageId="31" captionText="FIGURE 14. Map of known records for 10 species within the Lepanus pygmaeus species group overlaid with predicted distribution boundary of L. pygmaeus (dashed line) taken from Fig. 13 map." httpUri="https://zenodo.org/record/2588649/files/figure.png" pageId="21" pageNumber="62">Fig. 14</figureCitation>
). This species may represent a short-range endemic as all specimens have been collected within a
<quantity id="4CD23BE0FFBE8D67A123F86F57D196F8" box="[622,696,2009,2034]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="21" pageNumber="62" unit="km" value="25.0">25 km</quantity>
radius.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFFBD8D64A38AFF21571F9017" pageId="22" pageNumber="63" type="discussion">
<paragraph id="8B959605FFBD8D64A38AFF21571F9017" blockId="22.[151,1436,151,285]" pageId="22" pageNumber="63">
<emphasis id="B95E4A17FFBD8D64A38AFF21543D91BA" bold="true" box="[199,340,151,176]" pageId="22" pageNumber="63">Comments.</emphasis>
<taxonomicName id="4C2AED86FFBD8D64A22DFF2F574D91BA" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[352,548,151,176]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="63" phylum="Arthropoda" rank="species" species="feehani">
<emphasis id="B95E4A17FFBD8D64A22DFF2F574D91BA" box="[352,548,151,176]" italics="true" pageId="22" pageNumber="63">Lepanus feehani</emphasis>
</taxonomicName>
has previously been referred to as “
<taxonomicName id="4C2AED86FFBD8D64A0A6FF2F512791BA" authorityName=", Matthews" authorityYear="1974" box="[1003,1102,153,176]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="B95E4A17FFBD8D64A0A6FF2F512791BA" box="[1003,1102,153,176]" italics="true" pageId="22" pageNumber="63">Lepanus</emphasis>
</taxonomicName>
NQ26” under the informal nomenclature sensu
<bibRefCitation id="EFBBEBF4FFBD8D64A2C8FF0B570491DE" author="Yeates, D. &amp; Gunter, N. L. &amp; Weir, T. &amp; Cameron, S. L." box="[389,621,188,213]" pageId="22" pageNumber="63" pagination="92 - 99" refId="ref30201" refString="Yeates, D. K, Gunter, N. L., Weir, T. &amp; Cameron, S. L. (2011) Developing a benchmark invertebrate group for biodiversity modelling. In: West, J. (Ed.), 21 st Century Taxonomy: accelerating research and discovery of Australia's biodiversity, Taxonomy Research and Identification Network Final Report- March, 2011, pp. 92 - 99. Available from: https: // www. environment. gov. au / system / files / pages / 56 c 08 f 6 f- 7 bf 7 - 4594 - 8175 - b 4 c 143110156 / files / hub-taxonomy. pdf (accessed 12 December 2018)" type="journal article" year="2011">
Yeates
<emphasis id="B95E4A17FFBD8D64A297FF0B577C91DF" box="[474,533,188,213]" italics="true" pageId="22" pageNumber="63">et al.</emphasis>
(2011)
</bibRefCitation>
. The majority of specimens have been collected in dung baited pitfall traps, however specimens have also been collected in berlesed leaf-litter and by yellow-pan traps.
<taxonomicName id="4C2AED86FFBD8D64A668FF5750F291F2" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[1317,1435,223,248]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="63" phylum="Arthropoda" rank="species" species="feehani">
<emphasis id="B95E4A17FFBD8D64A668FF5750F291F2" box="[1317,1435,223,248]" italics="true" pageId="22" pageNumber="63">L. feehani</emphasis>
</taxonomicName>
has been collected both day and at night.
</paragraph>
</subSubSection>
</treatment>
</document>
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