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<document id="6B9E0F0CF3280C3A5D6C0A9FA3553543" ID-CLB-Dataset="28431" ID-DOI="10.11646/zootaxa.4564.1.2" ID-GBIF-Dataset="d3a56aac-92aa-48f2-b9fe-9576a4c2a0ec" ID-ISSN="1175-5326" ID-Zenodo-Dep="2588621" ID-ZooBank="27F70B5F-24DD-4C9E-BBA7-2B87476BD49C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552162070752" checkinUser="plazi" docAuthor="Gunter, Nicole L. &amp; Weir, Thomas A." docDate="2019" docId="03832713FF8F8D57A3DAFD0251579082" docLanguage="en" docName="zootaxa.4564.1.2.pdf" docOrigin="Zootaxa 4564 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepanus tozerensis Gunter &amp; Weir 2019, new species" docType="treatment" docUuid="A24FB341-9F8B-469E-9861-BFC1108392C1" docUuidSource="ZooBank" docVersion="9" lastPageNumber="78" masterDocId="FFBA5F6BFFAB8D72A34DFFB65569910A" masterDocTitle="Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): key to species groups and description of 14 new species from the L. pygmaeus species group" masterLastPageNumber="80" masterPageNumber="41" pageNumber="77" updateTime="1698700752634" updateUser="ExternalLinkService">
<mods:mods id="4D8897CF18896F0B467B4E5A56DFB8F8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6545F7E294D2FB7E827AD25A7AFF2B92">
<mods:title id="9FC97B6BBEB5FE6EA132AE94F2A28FBE">Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): key to species groups and description of 14 new species from the L. pygmaeus species group</mods:title>
</mods:titleInfo>
<mods:name id="34FE130D06FE1E344187AFC030210A26" type="personal">
<mods:role id="4612E89652D647B57C40EA2D5BAB9562">
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<mods:namePart id="D2E0C90F5E1429D7A679F0B8568D87B5">Gunter, Nicole L.</mods:namePart>
</mods:name>
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<mods:namePart id="88F09E41ABE2A32359614CA294B1D494">Weir, Thomas A.</mods:namePart>
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<mods:title id="BE9D55BB77850819B0FC9C8568313970">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="403C2D4AB573982964EF20E6BE3D3D6E">
<mods:date id="E0885C386933BA223C2F73D834B50BFF">2019</mods:date>
<mods:detail id="16419037827D565EEBD12294A216EA59" type="pubDate">
<mods:number id="C50DA396D507D0BAB1D4F8B856BDE76A">2019-03-05</mods:number>
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<mods:number id="BE25E2052586703C974896E1C1E9842F">4564</mods:number>
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<mods:classification id="D0D9BB7800E2CCC9C205E8AA8615C4D5">journal article</mods:classification>
<mods:identifier id="FD2C7E54C34C95BBC13DA7FA05269BA3" type="CLB-Dataset">28431</mods:identifier>
<mods:identifier id="5DF1A5502BD6988C7CB30D0D1C3CD7E0" type="DOI">10.11646/zootaxa.4564.1.2</mods:identifier>
<mods:identifier id="9984DC1B1EA45D848E99CE8FBE27DF90" type="GBIF-Dataset">d3a56aac-92aa-48f2-b9fe-9576a4c2a0ec</mods:identifier>
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<mods:identifier id="431B8516BFE126EA451FD4CE723C7D14" type="Zenodo-Dep">2588621</mods:identifier>
<mods:identifier id="090F2F98F1D4B0B0C33067A44707E737" type="ZooBank">27F70B5F-24DD-4C9E-BBA7-2B87476BD49C</mods:identifier>
</mods:mods>
<treatment id="03832713FF8F8D57A3DAFD0251579082" ID-DOI="http://doi.org/10.5281/zenodo.5922836" ID-GBIF-Taxon="154366360" ID-Zenodo-Dep="5922836" LSID="urn:lsid:zoobank.org:act:A24FB341-9F8B-469E-9861-BFC1108392C1" httpUri="http://treatment.plazi.org/id/03832713FF8F8D57A3DAFD0251579082" lastPageId="37" lastPageNumber="78" pageId="36" pageNumber="77">
<subSubSection id="C330C58EFF8F8D56A3DAFD02578993C4" box="[151,736,692,718]" pageId="36" pageNumber="77" type="nomenclature">
<paragraph id="8B959605FF8F8D56A3DAFD02578993C4" blockId="36.[151,736,692,751]" box="[151,736,692,718]" pageId="36" pageNumber="77">
<heading id="D0DD2169FF8F8D56A3DAFD02578993C4" bold="true" box="[151,736,692,718]" fontSize="11" level="1" pageId="36" pageNumber="77" reason="1">
<emphasis id="B95E4A17FF8F8D56A3DAFD02578993C4" bold="true" box="[151,736,692,718]" pageId="36" pageNumber="77">
<taxonomicName id="4C2AED86FF8F8D56A3DAFD02572D93C4" ID-CoL="3T8VR" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[151,580,692,718]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="77" phylum="Arthropoda" rank="species" species="tozerensis" status="sp. nov.">
<emphasis id="B95E4A17FF8F8D56A3DAFD02541493C4" bold="true" box="[151,381,692,718]" italics="true" pageId="36" pageNumber="77">Lepanus tozerensis</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A26DF76CFF8F8D56A11DFD02578993C4" box="[592,736,692,718]" pageId="36" pageNumber="77" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFF8F8D56A3DAFD6054FE93E5" box="[151,407,726,751]" pageId="36" pageNumber="77" type="description">
<paragraph id="8B959605FF8F8D56A3DAFD6054FE93E5" blockId="36.[151,736,692,751]" box="[151,407,726,751]" pageId="36" pageNumber="77">
(
<figureCitation id="13118A80FF8F8D56A3D2FD60546B93E5" box="[159,258,726,751]" captionStart="FIGURE 7" captionStartId="17.[151,250,1974,1996]" captionTargetBox="[497,1058,239,1909]" captionTargetId="figure@17.[403,1184,193,1953]" captionTargetPageId="17" captionText="FIGURE 7. Habitus of species within the Lepanus pygmaeus species group. A, Lepanus pygmaeus; B, Lepanus sauroni; C, Lepanus tozerensis. All specimens in comparative scale. Scale bar 1 mm." httpUri="https://zenodo.org/record/2588635/files/figure.png" pageId="36" pageNumber="77">Figs. 7C</figureCitation>
,
<figureCitation id="13118A80FF8F8D56A240FD61544593E5" box="[269,300,727,751]" captionStart="FIGURE 9" captionStartId="19.[151,250,1201,1223]" captionTargetBox="[205,1310,206,1167]" captionTargetId="figure@19.[194,1392,193,1180]" captionTargetPageId="19" captionText="FIGURE 9. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus pecki; B, Lepanus podocarp; C: Lepanus pungalina; D, Lepanus pygmaeus; E, Lepanus sauroni; F, Lepanus tozerensis. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588639/files/figure.png" pageId="36" pageNumber="77">9F</figureCitation>
,
<figureCitation id="13118A80FF8F8D56A27BFD61540F93E5" box="[310,358,727,751]" captionStart="FIGURE 12" captionStartId="25.[151,250,1905,1927]" captionTargetBox="[231,1341,208,1744]" captionTargetId="figure@25.[230,1357,193,1890]" captionTargetPageId="25" captionText="FIGURE 12. Left, dorsal, and right view of aedeagi of species within the Lepanus pygmaeus species group. A, Lepanus podocarp; B, Lepanus pungalina; C, Lepanus pygmaeus; D, Lepanus sauroni; E: Lepanus tozerensis. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar." httpUri="https://zenodo.org/record/2588645/files/figure.png" pageId="36" pageNumber="77">12E</figureCitation>
,
<figureCitation id="13118A80FF8F8D56A23CFD6154E793E5" box="[369,398,727,751]" captionStart="FIGURE 14" captionStartId="31.[151,250,1220,1242]" captionTargetBox="[151,1436,375,1198]" captionTargetId="figure@31.[151,1436,375,1198]" captionTargetPageId="31" captionText="FIGURE 14. Map of known records for 10 species within the Lepanus pygmaeus species group overlaid with predicted distribution boundary of L. pygmaeus (dashed line) taken from Fig. 13 map." httpUri="https://zenodo.org/record/2588649/files/figure.png" pageId="36" pageNumber="77">14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFF8F8D56A3DAFCA854CA955D" pageId="36" pageNumber="77" type="materials_examined">
<paragraph id="8B959605FF8F8D56A3DAFCA854CA955D" blockId="36.[151,1437,798,2011]" pageId="36" pageNumber="77">
<emphasis id="B95E4A17FF8F8D56A3DAFCA854CD923D" bold="true" box="[151,420,798,823]" pageId="36" pageNumber="77">
Type series.
<materialsCitation id="3B429C58FF8F8D56A263FCA854CD923D" ID-GBIF-Occurrence="2012745317" box="[302,420,798,823]" collectingDate="1986-07-05" collectingDateMax="68-06-03" collectingDateMin="1986-07-05" collectionCode="ANIC" collectorName="T. Weir" determinerName="T. A. Weir" latitude="-12.716666" location="Berlesate" longLatPrecision="1292" longitude="143.28334" pageId="36" pageNumber="77" specimenCode="ANIC 1061, CYP5" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="549128A7FF8F8D56A263FCA854F6923D" box="[302,415,798,823]" pageId="36" pageNumber="77" type="holotype">Holotype</typeStatus>
:
</materialsCitation>
</emphasis>
<specimenCount id="9D2C5D8CFF8F8D56A2E0FCA954AB923C" box="[429,450,799,822]" pageId="36" pageNumber="77" type="male">♂</specimenCount>
“
<geoCoordinate id="EE1EF0C2FF8F8D56A29BFCA95759923D" box="[470,560,799,823]" direction="south" orientation="latitude" pageId="36" pageNumber="77" precision="555" value="-12.43">12.43 S</geoCoordinate>
<geoCoordinate id="EE1EF0C2FF8F8D56A174FCA957C8923C" box="[569,673,799,823]" direction="east" orientation="longitude" pageId="36" pageNumber="77" precision="555" value="143.17">143.17 E</geoCoordinate>
<locationDeviation id="9A278F27FF8F8D56A1E7FCA856FE923C" box="[682,919,798,823]" location="Mt. Tozer" pageId="36" pageNumber="77">
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ENE Mt. Tozer
</locationDeviation>
, QLD,
<date id="FF94B0C5FF8F8D56A0A2FCA851F7923D" box="[1007,1182,798,823]" pageId="36" pageNumber="77" value="1986-07-05" valueMax="1986-07-10" valueMin="1986-07-05">
<collectingDate id="EFD0492DFF8F8D56A0A2FCA851F7923D" box="[1007,1182,798,823]" pageId="36" pageNumber="77" value="1986-07-05" valueMax="1986-07-10" valueMin="1986-07-05">5-10 July 1986</collectingDate>
</date>
,
<collectorName id="26DFF3D3FF8F8D56A7E6FCA9506D923D" box="[1195,1284,798,823]" pageId="36" pageNumber="77">T. Weir</collectorName>
” / “
<location id="8EF5C0DEFF8F8D56A67FFCA850F4923D" LSID="urn:lsid:plazi:treatment:03832713FF8F8D57A3DAFD0251579082:8EF5C0DEFF8F8D56A67FFCA850F4923D" box="[1330,1437,798,823]" latitude="-12.716666" longLatPrecision="1292" longitude="143.28334" name="Berlesate" pageId="36" pageNumber="77">Berlesate</location>
<specimenCode id="DB8C3E7EFF8F8D56A3DAFCF554729251" box="[151,283,834,859]" collectionCode="ANIC" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="36" pageNumber="77">ANIC 1061</specimenCode>
open forest litter” [
<geoCoordinate id="EE1EF0C2FF8F8D56A2B5FCF5573D9251" box="[504,596,835,859]" direction="south" orientation="latitude" pageId="36" pageNumber="77" precision="925" value="-12.716666">12°43'S</geoCoordinate>
,
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] / “ANIC Database No. 25-
<date id="FF94B0C5FF8F8D56A74FFCF451329251" box="[1026,1115,834,859]" pageId="36" pageNumber="77" value="68-06-03">
<collectingDate id="EFD0492DFF8F8D56A74FFCF451329251" box="[1026,1115,834,859]" pageId="36" pageNumber="77" value="68-06-03">060368</collectingDate>
</date>
” / ”
<taxonomicName id="4C2AED86FF8F8D56A7C8FCF5518F9251" authorityName=", Matthews" authorityYear="1974" box="[1157,1254,835,859]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="79" phylum="Arthropoda" rank="genus">Lepanus</taxonomicName>
<specimenCode id="DB8C3E7EFF8F8D56A7A3FCF4505B9251" box="[1262,1330,834,859]" collectionCode="CYP" country="Cyprus" lsid="urn:lsid:biocol.org:col:13586" name="Ministry of Agriculture, Natural Resources and Environment" pageId="36" pageNumber="77" type="Herbarium">CYP5</specimenCode>
det.
<determinerName id="7E497AC1FF8F8D56A626FCF555A49275" pageId="36" pageNumber="77">T.A. Weir</determinerName>
2012” / “ANIC Image” in ANIC.
<materialsCitation id="3B429C58FF8F8D56A117FCD057FA92A9" ID-GBIF-Occurrence="2012745318" collectingDate="1986-07-05" collectingDateMax="1986-07-10" collectingDateMin="1986-07-05" collectionCode="ANIC" collectorName="T. A. Weir" latitude="-12.716666" location="Mount Tozer" longLatPrecision="1292" longitude="143.28334" pageId="36" pageNumber="77" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B95E4A17FF8F8D56A117FCD057B29275" bold="true" box="[602,731,870,895]" pageId="36" pageNumber="77">
<typeStatus id="549128A7FF8F8D56A117FCD057BF9275" box="[602,726,870,895]" pageId="36" pageNumber="77" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<quantity id="4CD23BE0FF8F8D56A1A9FCD156769275" box="[740,799,870,895]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="36" pageNumber="77" unit="km" value="9.0">9 km</quantity>
ENE
<location id="8EF5C0DEFF8F8D56A02AFCD156929274" LSID="urn:lsid:plazi:treatment:03832713FF8F8D57A3DAFD0251579082:8EF5C0DEFF8F8D56A02AFCD156929274" box="[871,1019,871,894]" latitude="-12.716666" longLatPrecision="1292" longitude="143.28334" name="Mount Tozer" pageId="36" pageNumber="77">Mount Tozer</location>
, [
<geoCoordinate id="EE1EF0C2FF8F8D56A75DFCD151059275" box="[1040,1132,871,895]" direction="south" orientation="latitude" pageId="36" pageNumber="77" precision="925" value="-12.716666">12°43'S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FF8F8D56A734FCD1518D9275" box="[1145,1252,871,895]" direction="east" orientation="longitude" pageId="36" pageNumber="77" precision="925" value="143.28334">143°17'E</geoCoordinate>
],
<date id="FF94B0C5FF8F8D56A7B4FCD050F19275" box="[1273,1432,870,895]" pageId="36" pageNumber="77" value="1986-07-05" valueMax="1986-07-10" valueMin="1986-07-05">
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</date>
,
<collectorName id="26DFF3D3FF8F8D56A3DAFC3D546F92A9" box="[151,262,906,931]" pageId="36" pageNumber="77">T.A. Weir</collectorName>
(25-065529 ×
<specimenCount id="9D2C5D8CFF8F8D56A2E6FC3D54BB92A8" box="[427,466,907,931]" pageId="36" pageNumber="77" type="male">1 ♂</specimenCount>
×
<specimenCount id="9D2C5D8CFF8F8D56A2BFFC3D577592A8" box="[498,540,907,931]" pageId="36" pageNumber="77" type="female">1 ♀</specimenCount>
in
<collectionCode id="ED3B0EC0FF8F8D56A10FFC3D57E292A9" box="[578,651,907,931]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="36" pageNumber="77">ANIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FF8F8D56A1D2FC3D507C9505" ID-GBIF-Occurrence="2012745308" collectingDate="1968-05-11" collectingDateMax="69-06-03" collectingDateMin="1968-05-11" collectionCode="ANIC, QM" collectorName="T. A. Weir &amp; G. B. Monteith &amp; Iron Range &amp; Cape &amp; Peninsula" elevation="5" latitude="-12.733334" location="Chilli Beach" longLatPrecision="1292" longitude="143.28334" pageId="36" pageNumber="77" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="York" typeStatus="paratype">
<locationDeviation id="9A278F27FF8F8D56A1D2FC3D56DB92A8" box="[671,946,906,931]" location="Mount Tozer" pageId="36" pageNumber="77">
<quantity id="4CD23BE0FF8F8D56A1D2FC3D57B092A9" box="[671,729,906,931]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="36" pageNumber="77" unit="km" value="7.0">7 km</quantity>
ENE Mount Tozer
</locationDeviation>
, [
<geoCoordinate id="EE1EF0C2FF8F8D56A088FC3D514892A9" box="[965,1057,907,931]" direction="south" orientation="latitude" pageId="36" pageNumber="77" precision="925" value="-12.716666">12°43'S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FF8F8D56A763FC3D51F092A9" box="[1070,1177,907,931]" direction="east" orientation="longitude" pageId="36" pageNumber="77" precision="925" value="143.28334">143°17'E</geoCoordinate>
],
<date id="FF94B0C5FF8F8D56A7E0FC3C504992A9" box="[1197,1312,906,931]" pageId="36" pageNumber="77" value="1986-07-06">
<collectingDate id="EFD0492DFF8F8D56A7E0FC3C504992A9" box="[1197,1312,906,931]" pageId="36" pageNumber="77" value="1986-07-06">6.vii.1986</collectingDate>
</date>
,
<collectorName id="26DFF3D3FF8F8D56A660FC3D50F592A9" box="[1325,1436,906,931]" pageId="36" pageNumber="77">T.A. Weir</collectorName>
(25-
<date id="FF94B0C5FF8F8D56A38CFC18547092CD" box="[193,281,942,967]" pageId="36" pageNumber="77" value="69-06-03">
<collectingDate id="EFD0492DFF8F8D56A38CFC18547092CD" box="[193,281,942,967]" pageId="36" pageNumber="77" value="69-06-03">060369</collectingDate>
</date>
2 ×
<specimenCount id="9D2C5D8CFF8F8D56A217FC19540692CC" box="[346,367,943,966]" pageId="36" pageNumber="77" type="male">♂</specimenCount>
2 ×
<specimenCount id="9D2C5D8CFF8F8D56A2FDFC1954AC92CC" box="[432,453,943,966]" pageId="36" pageNumber="77" type="female">♀</specimenCount>
in
<collectionCode id="ED3B0EC0FF8F8D56A2BEFC19575E92CD" box="[499,567,943,967]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="36" pageNumber="77">ANIC</collectionCode>
and 1 ×
<specimenCount id="9D2C5D8CFF8F8D56A1E1FC1957A892CC" box="[684,705,943,966]" pageId="36" pageNumber="77" type="male">♂</specimenCount>
1 ×
<specimenCount id="9D2C5D8CFF8F8D56A04FFC19567E92CC" box="[770,791,943,966]" pageId="36" pageNumber="77" type="female">♀</specimenCount>
in
<collectionCode id="ED3B0EC0FF8F8D56A009FC19561F92CD" box="[836,886,943,967]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="36" pageNumber="77" type="Museum">QM</collectionCode>
);
<location id="8EF5C0DEFF8F8D56A0C3FC18514B92CD" LSID="urn:lsid:plazi:treatment:03832713FF8F8D57A3DAFD0251579082:8EF5C0DEFF8F8D56A0C3FC18514B92CD" box="[910,1058,942,967]" latitude="-12.733334" longLatPrecision="1292" longitude="143.28334" name="Chilli Beach" pageId="36" pageNumber="77" stateProvince="York">Chilli Beach</location>
,
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,
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,
<quantity id="4CD23BE0FF8F8D56A62BFC1850FE92CC" box="[1382,1431,942,967]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="77" unit="m" value="5.0">
<elevation id="00077136FF8F8D56A62BFC1850FE92CC" box="[1382,1431,942,967]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="77" unit="m" value="5.0">5 m</elevation>
</quantity>
,
<date id="FF94B0C5FF8F8D56A3DAFC64547E92E1" box="[151,279,978,1003]" pageId="36" pageNumber="77" value="2010-12-11">
<collectingDate id="EFD0492DFF8F8D56A3DAFC64547E92E1" box="[151,279,978,1003]" pageId="36" pageNumber="77" value="2010-12-11">11.xii.2010</collectingDate>
</date>
,
<collectorName id="26DFF3D3FF8F8D56A264FC6554A492E1" box="[297,461,978,1003]" pageId="36" pageNumber="77">G.B. Monteith</collectorName>
(25-
<collectingDate id="EFD0492DFF8F8D56A149FC6457EC92E1" box="[516,645,978,1003]" isEnumeration="true" pageId="36" pageNumber="77">
<date id="FF94B0C5FF8F8D56A149FC64573592E1" box="[516,604,978,1003]" pageId="36" pageNumber="77" value="27-06-08">060827</date>
<date id="FF94B0C5FF8F8D56A124FC6557EC92E1" box="[617,645,979,1003]" pageId="36" pageNumber="77">x1</date>
</collectingDate>
♀ in
<collectionCode id="ED3B0EC0FF8F8D56A198FC65567392E1" box="[725,794,979,1003]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="36" pageNumber="77">ANIC</collectionCode>
also as COL1459);
<collectorName id="26DFF3D3FF8F8D56A75DFC6551FC92E1" box="[1040,1173,979,1003]" pageId="36" pageNumber="77">Iron Range</collectorName>
,
<collectorName id="26DFF3D3FF8F8D56A7EBFC6551B692E1" box="[1190,1247,979,1003]" pageId="36" pageNumber="77">Cape</collectorName>
<collectingRegion id="49EE58E7FF8F8D56A7A1FC64504A92E1" box="[1260,1315,978,1003]" country="United Kingdom" name="York" pageId="36" pageNumber="77">York</collectingRegion>
<collectorName id="26DFF3D3FF8F8D56A662FC6450F592E1" box="[1327,1436,978,1003]" pageId="36" pageNumber="77">Peninsula</collectorName>
(general), [
<geoCoordinate id="EE1EF0C2FF8F8D56A25CFC4154049505" box="[273,365,1015,1039]" direction="south" orientation="latitude" pageId="36" pageNumber="77" precision="925" value="-12.733334">12°44'S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FF8F8D56A234FC41548C9505" box="[377,485,1015,1039]" direction="east" orientation="longitude" pageId="36" pageNumber="77" precision="925" value="143.28334">143°17'E</geoCoordinate>
],
<date id="FF94B0C5FF8F8D56A2B5FC4057FB9505" box="[504,658,1014,1039]" pageId="36" pageNumber="77" value="1968-05-11" valueMax="1968-05-17" valueMin="1968-05-11">
<collectingDate id="EFD0492DFF8F8D56A2B5FC4057FB9505" box="[504,658,1014,1039]" pageId="36" pageNumber="77" value="1968-05-11" valueMax="1968-05-17" valueMin="1968-05-11">11–17.v.1968</collectingDate>
</date>
,
<collectorName id="26DFF3D3FF8F8D56A1D0FC4156549505" box="[669,829,1014,1039]" pageId="36" pageNumber="77">G.B. Monteith</collectorName>
(25-058520
<date id="FF94B0C5FF8F8D56A083FC4156839505" box="[974,1002,1015,1039]" pageId="36" pageNumber="77">
<collectingDate id="EFD0492DFF8F8D56A083FC4156839505" box="[974,1002,1015,1039]" pageId="36" pageNumber="77">x1</collectingDate>
</date>
sex undetermined in
<collectionCode id="ED3B0EC0FF8F8D56A796FC4150649505" box="[1243,1293,1015,1039]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="36" pageNumber="77" type="Museum">QM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FF8F8D56A66CFC4154F6955D" ID-GBIF-Occurrence="2012745319" collectingDate="1989-06-27" collectingDateMax="1989-07-12" collectingDateMin="1989-06-27" collectionCode="ANIC" collectorName="T. A. Weir" elevation="520" latitude="-13.733334" location="McIlwraith Range" longLatPrecision="1290" longitude="143.33333" pageId="36" pageNumber="77" specimenCount="1" stateProvince="York" typeStatus="paratype">
<locationDeviation id="9A278F27FF8F8D56A66CFC41544A9539" location="Bald Hill" pageId="36" pageNumber="77">
<quantity id="4CD23BE0FF8F8D56A66CFC41500F9505" box="[1313,1382,1014,1039]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="36" pageNumber="77" unit="km" value="11.0">11 km</quantity>
NW of Bald Hill
</locationDeviation>
,
<location id="8EF5C0DEFF8F8D56A27CFBAC576A9539" LSID="urn:lsid:plazi:treatment:03832713FF8F8D57A3DAFD0251579082:8EF5C0DEFF8F8D56A27CFBAC576A9539" box="[305,515,1050,1075]" latitude="-13.733334" longLatPrecision="1290" longitude="143.33333" name="McIlwraith Range" pageId="36" pageNumber="77" stateProvince="York">McIlwraith Range</location>
, [
<geoCoordinate id="EE1EF0C2FF8F8D56A155FBAD57069539" box="[536,623,1051,1075]" direction="south" orientation="latitude" pageId="36" pageNumber="77" precision="925" value="-13.733334">13°44'S</geoCoordinate>
<geoCoordinate id="EE1EF0C2FF8F8D56A135FBAD578B9539" box="[632,738,1051,1075]" direction="east" orientation="longitude" pageId="36" pageNumber="77" precision="925" value="143.33333">143°20'E</geoCoordinate>
],
<quantity id="4CD23BE0FF8F8D56A1BAFBAC562B9538" box="[759,834,1050,1075]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="36" pageNumber="77" unit="m" value="520.0">
<elevation id="00077136FF8F8D56A1BAFBAC562B9538" box="[759,834,1050,1075]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="36" pageNumber="77" unit="m" value="520.0">520 m</elevation>
</quantity>
,
<date id="FF94B0C5FF8F8D56A01DFBAC51709539" box="[848,1049,1050,1075]" pageId="36" pageNumber="77" value="1989-06-27" valueMax="1989-07-12" valueMin="1989-06-27">
<collectingDate id="EFD0492DFF8F8D56A01DFBAC51709539" box="[848,1049,1050,1075]" pageId="36" pageNumber="77" value="1989-06-27" valueMax="1989-07-12" valueMin="1989-06-27">27.vi–12.vii.1989</collectingDate>
</date>
,
<collectorName id="26DFF3D3FF8F8D56A76AFBAD51FE9539" box="[1063,1175,1050,1075]" pageId="36" pageNumber="77">T.A. Weir</collectorName>
(25-
<collectingDate id="EFD0492DFF8F8D56A787FBAC50209539" box="[1226,1353,1050,1075]" isEnumeration="true" pageId="36" pageNumber="77">
<date id="FF94B0C5FF8F8D56A787FBAC504A9539" box="[1226,1315,1050,1075]" pageId="36" pageNumber="77" value="70-06-03">060370</date>
<date id="FF94B0C5FF8F8D56A661FBAD50209539" box="[1324,1353,1051,1075]" pageId="36" pageNumber="77">x1</date>
</collectingDate>
♀, 25- 0 60371
<date id="FF94B0C5FF8F8D56A3BFFB895467955D" box="[242,270,1087,1111]" pageId="36" pageNumber="77">
<collectingDate id="EFD0492DFF8F8D56A3BFFB895467955D" box="[242,270,1087,1111]" pageId="36" pageNumber="77">x3</collectingDate>
</date>
♀ in
<collectionCode id="ED3B0EC0FF8F8D56A200FB8954FE955D" box="[333,407,1087,1111]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="36" pageNumber="77">ANIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFF8F8D56A38AFBD4541E9665" pageId="36" pageNumber="77" type="description">
<paragraph id="8B959605FF8F8D56A38AFBD454B59595" blockId="36.[151,1437,798,2011]" pageId="36" pageNumber="77">
<emphasis id="B95E4A17FF8F8D56A38AFBD454309571" bold="true" box="[199,345,1122,1147]" pageId="36" pageNumber="77">Description.</emphasis>
Head and pronotum dark brown and elytra either uniformly light brown or light brown with red humeri. Antennal clubs grey.
</paragraph>
<paragraph id="8B959605FF8F8D56A38AFB1C51A895C8" blockId="36.[151,1437,798,2011]" box="[199,1217,1194,1219]" pageId="36" pageNumber="77">
Total length:
<quantity id="4CD23BE0FF8F8D56A211FB1D54DC95C8" box="[348,437,1195,1219]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="36" pageNumber="77" unit="mm" value="2.2">2.2 mm</quantity>
.
<typeStatus id="549128A7FF8F8D56A2F2FB1C574E95C9" box="[447,551,1194,1219]" pageId="36" pageNumber="77" type="holotype">Holotype</typeStatus>
measurements: Total length
<quantity id="4CD23BE0FF8F8D56A026FB1D56AA95C8" box="[875,963,1195,1219]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="36" pageNumber="77" unit="mm" value="2.2">2.2 mm</quantity>
, elytral width
<quantity id="4CD23BE0FF8F8D56A729FB1C51D495C8" box="[1124,1213,1194,1219]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="36" pageNumber="77" unit="mm" value="1.5">1.5 mm</quantity>
.
</paragraph>
<paragraph id="8B959605FF8F8D56A38AFB7857AA9459" blockId="36.[151,1437,798,2011]" pageId="36" pageNumber="77">Male: Head: U-shaped between clypeal teeth, which are short and upturned. Basal carina present but interrupted medially. Margin slightly angulate at clypeogenall suture. Genal angle weakly defined. Dorsal part of eyes narrow in size, separated by interocular space approximately 21 times eye width (32:1.5); eye canthus not dividing eye. Head 1.4 × wider than long (42:30).</paragraph>
<paragraph id="8B959605FF8F8D56A38AFAE857829491" blockId="36.[151,1437,798,2011]" pageId="36" pageNumber="77">Prothorax: Hypomeral surface finely reticulate with light punctation. Hypomeral striae about half the length of hypomeron. Pronotum 1.7 × wider than long (67:40).</paragraph>
<paragraph id="8B959605FF8F8D56A38AFA10512E94E9" blockId="36.[151,1437,798,2011]" pageId="36" pageNumber="77">Elytra: Surface smooth. Striae 1–5 double with very fine punctate edges. All striae on disc not equal width, striae 6 is not as wide, striae 7 appears single. Elytra 1.25 × wider than long (80:65).</paragraph>
<paragraph id="8B959605FF8F8D56A38AFA59559A9745" blockId="36.[151,1437,798,2011]" pageId="36" pageNumber="77">Legs: Front edge of protibiae truncate; apical digit somewhat elongate and sharp; 2 teeth on outer edge. Inner edge of metatibiae almost straight. Metatarsi with 2nd tarsomere subequal to the 5th. Tarsal claws minute and toothed.</paragraph>
<paragraph id="8B959605FF8F8D56A38AF9EC507997B0" blockId="36.[151,1437,798,2011]" pageId="36" pageNumber="77">Abdomen: Pygidium with almost circular depression containing a round tubercle in upper third. Depression occupies slightly more than half of the pygidium. Upper edge of pygidium broadly angulate and the depression extends under this edge. Abdominal surface slightly reticulate right across. Segment 6 finely punctate.</paragraph>
<paragraph id="8B959605FF8F8D56A38AF97051BE962D" blockId="36.[151,1437,798,2011]" pageId="36" pageNumber="77">Pterothorax: Medial lobe of metaventrite finely punctate and broadly margined between mesocoxae. Lateral lobe of metaventrite smooth and punctate with setae. Meso-metaventral suture very slightly arched. Mesoventrite with slight notch anteriorly. Mesoventrite and mesepimeron smooth. Metanepisternum reticulate.</paragraph>
<paragraph id="8B959605FF8F8D56A38AF884541E9665" blockId="36.[151,1437,798,2011]" pageId="36" pageNumber="77">Female: Apical digit absent with apical spur extending straight from front edge. Abdominal ventrite 6 slightly longer than in male.</paragraph>
</subSubSection>
<subSubSection id="C330C58EFF8F8D56A38AF8CC56349698" box="[199,861,1914,1939]" pageId="36" pageNumber="77" type="etymology">
<paragraph id="8B959605FF8F8D56A38AF8CC56349698" blockId="36.[151,1437,798,2011]" box="[199,861,1914,1939]" pageId="36" pageNumber="77">
<emphasis id="B95E4A17FF8F8D56A38AF8CC54279699" bold="true" box="[199,334,1914,1939]" pageId="36" pageNumber="77">Etymology.</emphasis>
Named for the
<typeStatus id="549128A7FF8F8D56A2B0F8CD57449699" box="[509,557,1915,1939]" pageId="36" pageNumber="77">type</typeStatus>
locality near Mount Tozer.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFF8F8D57A38AF828546991DE" lastPageId="37" lastPageNumber="78" pageId="36" pageNumber="77" type="distribution">
<paragraph id="8B959605FF8F8D57A38AF828546991DE" blockId="36.[151,1437,798,2011]" lastBlockId="37.[151,1436,151,392]" lastPageId="37" lastPageNumber="78" pageId="36" pageNumber="77">
<emphasis id="B95E4A17FF8F8D56A38AF828540996BD" bold="true" box="[199,352,1950,1975]" pageId="36" pageNumber="77">Distribution.</emphasis>
Known only from the CYP IBRA region of northern
<collectingRegion id="49EE58E7FF8F8D56A09EF828513296BD" box="[979,1115,1950,1975]" country="Australia" name="Queensland" pageId="36" pageNumber="77">Queensland</collectingRegion>
. Most specimens have been collected from open forests of Kutini-Payamu (Iron Range) National Park to McIlwrath Range in the CYP Interim Biogeographic Regionalisation for
<collectingCountry id="F33DD695FF8E8D57A16AFF2157E491BA" box="[551,653,151,176]" name="Australia" pageId="37" pageNumber="78">Australia</collectingCountry>
region, with one specimen from Chilli Beach collected in rainforest (
<figureCitation id="13118A80FF8E8D57A3D2FF0A559A91DE" box="[159,243,188,213]" captionStart="FIGURE 14" captionStartId="31.[151,250,1220,1242]" captionTargetBox="[151,1436,375,1198]" captionTargetId="figure@31.[151,1436,375,1198]" captionTargetPageId="31" captionText="FIGURE 14. Map of known records for 10 species within the Lepanus pygmaeus species group overlaid with predicted distribution boundary of L. pygmaeus (dashed line) taken from Fig. 13 map." httpUri="https://zenodo.org/record/2588649/files/figure.png" pageId="37" pageNumber="78">Fig. 14</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFF8E8D57A38AFF6951579082" pageId="37" pageNumber="78" type="discussion">
<paragraph id="8B959605FF8E8D57A38AFF6951579082" blockId="37.[151,1436,151,392]" pageId="37" pageNumber="78">
<emphasis id="B95E4A17FF8E8D57A38AFF69542691F2" bold="true" box="[199,335,223,248]" pageId="37" pageNumber="78">Comments.</emphasis>
<taxonomicName id="4C2AED86FF8E8D57A22CFF57572891F2" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[353,577,225,248]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="78" phylum="Arthropoda" rank="species" species="tozerensis">
<emphasis id="B95E4A17FF8E8D57A22CFF57572891F2" box="[353,577,225,248]" italics="true" pageId="37" pageNumber="78">Lepanus tozerensis</emphasis>
</taxonomicName>
was previously referred to as “
<taxonomicName id="4C2AED86FF8E8D57A0ACFF57512991F2" authorityName=", Matthews" authorityYear="1974" box="[993,1088,225,248]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis id="B95E4A17FF8E8D57A0ACFF57512991F2" box="[993,1088,225,248]" italics="true" pageId="37" pageNumber="78">Lepanus</emphasis>
</taxonomicName>
CYP5” under the informal nomenclature sensu
<bibRefCitation id="EFBBEBF4FF8E8D57A2C9FEB357059016" author="Yeates, D. &amp; Gunter, N. L. &amp; Weir, T. &amp; Cameron, S. L." box="[388,620,260,285]" pageId="37" pageNumber="78" pagination="92 - 99" refId="ref30201" refString="Yeates, D. K, Gunter, N. L., Weir, T. &amp; Cameron, S. L. (2011) Developing a benchmark invertebrate group for biodiversity modelling. In: West, J. (Ed.), 21 st Century Taxonomy: accelerating research and discovery of Australia's biodiversity, Taxonomy Research and Identification Network Final Report- March, 2011, pp. 92 - 99. Available from: https: // www. environment. gov. au / system / files / pages / 56 c 08 f 6 f- 7 bf 7 - 4594 - 8175 - b 4 c 143110156 / files / hub-taxonomy. pdf (accessed 12 December 2018)" type="journal article" year="2011">
Yeates
<emphasis id="B95E4A17FF8E8D57A295FEB3577D9017" box="[472,532,260,285]" italics="true" pageId="37" pageNumber="78">et al.</emphasis>
(2011)
</bibRefCitation>
. The specimen from Chilli Beach that was sequenced in the study of
<bibRefCitation id="EFBBEBF4FF8E8D57A3DAFE9E5416904A" author="Gunter, N. L. &amp; Monteith, G. B. &amp; Cameron, S. L. &amp; Weir, T." box="[151,383,295,320]" pageId="37" pageNumber="78" pagination="1 - 27" refId="ref29280" refString="Gunter, N. L., Monteith, G. B., Cameron, S. L. &amp; Weir, T. A (2018) Evidence from Australian mesic zone dung beetles support Gondwanan vicariance and Mesozoic origin of the Scarabaeinae (Coleoptera: Scarabaeidae). Invertebrate Systematics and Evolution, 1 - 27. [Advance online publication] https: // doi. org / 10.1163 / 1876312 X- 00002171" type="book chapter" year="2018">
Gunter
<emphasis id="B95E4A17FF8E8D57A3A2FE9F5443904A" box="[239,298,295,320]" italics="true" pageId="37" pageNumber="78">et al.</emphasis>
(2018)
</bibRefCitation>
is designated as a
<typeStatus id="549128A7FF8E8D57A12DFE9E57A8904A" box="[608,705,296,320]" pageId="37" pageNumber="78" type="paratype">paratype</typeStatus>
here (ANIC 25-
<date id="FF94B0C5FF8E8D57A0CCFE9156B5904A" box="[897,988,295,320]" pageId="37" pageNumber="78" value="27-06-08">060827</date>
). It was recovered as a member of the monsoonal clade of
<taxonomicName id="4C2AED86FF8E8D57A235FEFB54BE906E" authorityName=", Matthews" authorityYear="1974" box="[376,471,333,356]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis id="B95E4A17FF8E8D57A235FEFB54BE906E" box="[376,471,333,356]" italics="true" pageId="37" pageNumber="78">Lepanus</emphasis>
</taxonomicName>
, which contains species in the
<taxonomicName id="4C2AED86FF8E8D57A07DFEFB56D4906E" authorityName="in Matthews" authorityYear="1974" box="[816,957,333,356]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="78" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FF8E8D57A07DFEFB5656906E" box="[816,831,333,356]" italics="true" pageId="37" pageNumber="78">L</emphasis>
.
<emphasis id="B95E4A17FF8E8D57A000FEFB56D4906E" box="[845,957,333,356]" italics="true" pageId="37" pageNumber="78">pygmaeus</emphasis>
</taxonomicName>
species group. This species is known from only
<specimenCount id="9D2C5D8CFF8E8D57A39DFED954019082" box="[208,360,367,392]" pageId="37" pageNumber="78" type="generic">15 specimens</specimenCount>
collected either in dung baited pitfall traps or berlesed leaf litter.
</paragraph>
</subSubSection>
</treatment>
</document>
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