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<document id="1BCF8A7F3EF216439B29BBF059E0678E" ID-CLB-Dataset="269966" ID-DOI="10.5281/zenodo.5337424" ID-GBIF-Dataset="7c26f212-ab38-4e92-b461-cdb7dc073af5" ID-ISSN="0253-116X" ID-Zenodo-Dep="5337424" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1630178635768" checkinUser="carolina" docAuthor="Puthz, V." docDate="2012" docId="038313247C4BFFCFFF01FEDB3C36E5C8" docLanguage="en" docName="LinzerbiolBeitr.2012.44.2.1431-1475.pdf" docOrigin="Linzer biologische Beiträge 44 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Stenus perpinguis Puthz 2012, nov.sp." docType="treatment" docVersion="7" lastPageNumber="1447" masterDocId="FFBA6B5C7C5BFFDFFFB3FFE33D00E112" masterDocTitle="On the Stenus LATREILLE 1797 from Taiwan with non-spotted elytra (Coleoptera, Staphylinidae) (327 Contribution to the Knowledge of Steninae)" masterLastPageNumber="1475" masterPageNumber="1431" pageNumber="1447" updateTime="1699176368675" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="913591DF0CB17537DE2848A4774B2ADC">On the Stenus LATREILLE 1797 from Taiwan with non-spotted elytra (Coleoptera, Staphylinidae) (327 Contribution to the Knowledge of Steninae)</mods:title>
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<mods:namePart id="A17574CCECB6D0C847DE7335CBA9A872">Puthz, V.</mods:namePart>
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<mods:title id="E2F073AC8BCC2B954551D96D321A3337">Linzer biologische Beiträge</mods:title>
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<mods:part id="2233F4F6FE78195DA3B02811D983A7F1">
<mods:date id="0D19741E9524239D74ED09D0EECA11A9">2012</mods:date>
<mods:detail id="085975446281CDF774E08B63507ACD17" type="pubDate">
<mods:number id="415AEAEB0DF921FE21DFB89C341C2BFC">2012-12-28</mods:number>
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<mods:number id="2EE908D67970D392B425B2B60CACEBA9">44</mods:number>
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<mods:classification id="8AEAC0B6E688F1CAA2728B3F3E4CEA72">journal article</mods:classification>
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<mods:identifier id="05C80F4AA5DC716A9694BB92B4D08DB0" type="DOI">10.5281/zenodo.5337424</mods:identifier>
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<treatment id="038313247C4BFFCFFF01FEDB3C36E5C8" ID-DOI="http://doi.org/10.5281/zenodo.8392886" ID-GBIF-Taxon="213626799" ID-Zenodo-Dep="8392886" LSID="urn:lsid:plazi:treatment:038313247C4BFFCFFF01FEDB3C36E5C8" httpUri="http://treatment.plazi.org/id/038313247C4BFFCFFF01FEDB3C36E5C8" lastPageNumber="1447" pageId="16" pageNumber="1447">
<subSubSection id="C330F1B97C4BFFCFFF01FEDB3F63E05C" box="[178,611,312,334]" pageId="16" pageNumber="1447" type="nomenclature">
<paragraph id="8B95A2327C4BFFCFFF01FEDB3F63E05C" blockId="16.[178,611,312,334]" box="[178,611,312,334]" pageId="16" pageNumber="1447">
<heading id="D0DD155E7C4BFFCFFF01FEDB3F63E05C" box="[178,611,312,334]" fontSize="8" level="2" pageId="16" pageNumber="1447" reason="2">
<emphasis id="B95E7E207C4BFFCFFF01FEDB3F63E05C" bold="true" box="[178,611,312,334]" pageId="16" pageNumber="1447">
<taxonomicName id="4C2AD9B17C4BFFCFFF01FEDB3C6EE05F" ID-CoL="52B4M" authority="Puthz, 2012" authorityName="Puthz" authorityYear="2012" box="[178,366,312,333]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1447" phylum="Arthropoda" rank="species" species="perpinguis" status="sp. nov.">
<emphasis id="B95E7E207C4BFFCFFF01FEDB3C6EE05F" bold="true" box="[178,366,312,333]" italics="true" pageId="16" pageNumber="1447">Stenus perpinguis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26DC35B7C4BFFCFFEC7FEDB3CC0E05C" box="[372,448,312,334]" pageId="16" pageNumber="1447" rank="species">nov.sp.</taxonomicNameLabel>
(
<figureCitation id="1311BEB77C4BFFCFFE7EFEDB3F10E05C" box="[461,528,312,334]" captionStart="Figs 1-16" captionStartId="38.[178,218,1324,1344]" captionTargetBox="[178,1082,220,1305]" captionTargetId="figure-8@38.[178,1085,674,834]" captionTargetPageId="38" captionText="Figs 1-16: Sternite 8 of male (1-8, 13, 14) and of female (9, 10), posterior portion of sternite 9 of male (11, 12), anterior portion of aedeagus (15, 16) of Stenus hirtellus SHARP (1), S. piliferus MOTSCHULSKY (2), S. perpastus nov.sp. (3, PT), S. perpinguis nov.sp. (4, PT), S. perpropinquus nov.sp. (5, PT), S. persimplex nov.sp. (6, PT), S. perturbator nov.sp. (7, PT), S. pervilis nov.sp. (8, PT), S. persculptus nov.sp. (9, PT), S. perspicabilis nov.sp. (10, PT), S. sugayai nov.sp. (11, PT), S. perodiosus nov.sp. (12, PT), S. ascendor (13, 15, PT), S. hanami HROMÁDKA (14, 16, HT). – Scale bar = 0.1mm (1= 2-10; 11=12; 13=14; 15=16)." figureDoi="http://doi.org/10.5281/zenodo.5337426" httpUri="https://zenodo.org/record/5337426/files/figure.png" pageId="16" pageNumber="1447">Figs 4</figureCitation>
,
<figureCitation id="1311BEB77C4BFFCFFDAFFEDB3F37E05C" box="[540,567,312,334]" captionStart="Figs 17-29" captionStartId="39.[178,218,1076,1096]" captionTargetBox="[179,1081,220,1056]" captionTargetId="figure-8@39.[178,1085,583,743]" captionTargetPageId="39" captionText="Figs 17-29: Ventral aspect of aedeagus of Stenus perpusillus nov.sp. (17, PT), S. sugayai nov.sp. (18, 19: Anmashan, PTT), S. perpinguis nov.sp. (20, T 28, PT), S. perspicabilis nov.sp. (21, Lalashan, PT), S. perpastus nov.sp. (22, T 92, PT), S. mysterialis PUTHZ (23, near Tadongshan), S. pervenustus nov.sp. (24, T 138, PT), S. persculptus nov.sp. (25, T 95, PT), S. perpropinquus nov.sp. (26, T 52, PT), S. cirriostentans nov.sp. (27, PT), S. perpauper nov.sp. (28, Fenchifu, PT), S. persculpturatus nov.sp. (29, T 101, PT). – Scale bar = 0.1mm (17= 18, 19, 20= 21-29)." figureDoi="http://doi.org/10.5281/zenodo.5337428" httpUri="https://zenodo.org/record/5337428/files/figure.png" pageId="16" pageNumber="1447">20</figureCitation>
,
<figureCitation id="1311BEB77C4BFFCFFDF1FEDB3F5DE05C" box="[578,605,312,334]" captionStart="Figs 55-69" captionStartId="42.[178,218,1301,1321]" captionTargetBox="[178,1085,219,1281]" captionTargetId="figure-149@42.[178,1085,219,1281]" captionTargetPageId="42" captionText="Figs 55-69: Spermathecae of PTT of Stenus pervenustus nov.sp. (55, T 138), S. persculptus nov.sp. (56, T 107), S. persculpturatus nov.sp. (57, T 88), S. pervilis nov.sp. (58, T 120), S. sugayai nov.sp. (59, Anmashan), S. persubtilis nov.sp. (60, T 199), S. perplicatus nov.sp. (61, Meifeng), S. perturbator nov.sp. (62, T 187), S. persimplex nov.sp. (63, T 177), S. perrarus nov.sp. (64, Fushan), S. perspicabilis nov.sp. (65, T 144), S. perpinguis nov.sp. (66, T 29), S. perodiosus nov.sp. (67, T 84), S. perpunctus nov.sp. (68, T 72), S. perversor nov.sp. (69, T 59)." figureDoi="http://doi.org/10.5281/zenodo.5337434" httpUri="https://zenodo.org/record/5337434/files/figure.png" pageId="16" pageNumber="1447">66</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C330F1B97C4BFFCFFF01FE8B3F92E0CE" pageId="16" pageNumber="1447" type="materials_examined">
<paragraph id="8B95A2327C4BFFCFFF01FE8B3F92E0CE" blockId="16.[178,1085,360,476]" pageId="16" pageNumber="1447">
Type material:
<materialsCitation id="3B42A86F7C4BFFCFFE15FE8B3F1FE06E" ID-GBIF-Occurrence="4417845380" box="[422,543,360,380]" collectingDate="1991-05-18" collectionCode="Smetana" collectorName="A. Smetana" country="Taiwan" elevation="2700" location="Cliff" municipality="Yushan National Park" pageId="16" pageNumber="1447" specimenCount="1" stateProvince="Nantou" typeStatus="holotype">
<typeStatus id="54911C907C4BFFCFFE15FE8B3CF7E06E" box="[422,503,360,380]" pageId="16" pageNumber="1447" type="holotype">Holotype</typeStatus>
()
</materialsCitation>
<materialsCitation id="3B42A86F7C4BFFCFFD97FE8B3E58E086" ID-GBIF-Occurrence="4417845518" collectingDate="1991-05-18" collectionCode="Smetana" collectorName="A. Smetana" country="Taiwan" elevation="2700" location="Cliff" municipality="Yushan National Park" pageId="16" pageNumber="1447" specimenCount="22" stateProvince="Nantou" typeStatus="paratype">
and 9, 13 -
<typeStatus id="54911C907C4BFFCFFD75FE8B3E1EE06E" box="[710,798,360,380]" pageId="16" pageNumber="1447" type="paratype">paratypes</typeStatus>
:
<collectingCountry id="F33DE2A27C4BFFCFFC95FE8B3E6BE06E" box="[806,875,360,380]" name="Taiwan" pageId="16" pageNumber="1447">Taiwan</collectingCountry>
:
<collectingRegion id="49EE6CD07C4BFFCFFCC7FE8B3EF1E06E" box="[884,1009,360,380]" country="Taiwan" name="Nantou" pageId="16" pageNumber="1447">Nantou Hsien</collectingRegion>
:
<collectingMunicipality id="6BF138487C4BFFCFFC49FE8B3C5EE086" pageId="16" pageNumber="1447">Yushan National Park</collectingMunicipality>
, Mun-Li
<location id="8EF5F4E97C4BFFCFFE0BFE633CE3E086" LSID="urn:lsid:plazi:treatment:038313247C4BFFCFFF01FEDB3C36E5C8:8EF5F4E97C4BFFCFFE0BFE633CE3E086" box="[440,483,384,404]" country="Taiwan" municipality="Yushan National Park" name="Cliff" pageId="16" pageNumber="1447" stateProvince="Nantou">Cliff</location>
,
<quantity id="4CD20FD77C4BFFCFFE43FE633F2FE086" box="[496,559,384,404]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="16" pageNumber="1447" unit="m" value="2700.0">
<elevation id="000745017C4BFFCFFE43FE633F2FE086" box="[496,559,384,404]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="16" pageNumber="1447" unit="m" value="2700.0">2700m</elevation>
</quantity>
,
<date id="FF9484F27C4BFFCFFD8FFE633F9AE086" box="[572,666,384,404]" pageId="16" pageNumber="1447" value="1991-05-18">
<collectingDate id="EFD07D1A7C4BFFCFFD8FFE633F9AE086" box="[572,666,384,404]" pageId="16" pageNumber="1447" value="1991-05-18">18.V.1991</collectingDate>
</date>
,
<collectorName id="26DFC7E47C4BFFCFFD15FE633E10E086" box="[678,784,384,404]" pageId="16" pageNumber="1447">A. Smetana</collectorName>
(T 86)
</materialsCitation>
.
<materialsCitation id="3B42A86F7C4BFFCFFCD6FE633F04E0BE" ID-GBIF-Occurrence="4417845335" collectingDate="1990-04-27" collectingDateMax="1991-05-13" collectingDateMin="1990-04-27" collectionCode="cN, cP" collectorName="A. Smetana" country="Taiwan" elevation="2700" location="Cliff" municipality="Yushan National Park" pageId="16" pageNumber="1447" specimenCount="8" stateProvince="Chiayi" typeStatus="paratype">
<typeStatus id="54911C907C4BFFCFFCD6FE633EBDE086" box="[869,957,384,404]" pageId="16" pageNumber="1447" type="paratype">Paratypes</typeStatus>
: 2, 6: ibidem,
<date id="FF9484F27C4BFFCFFE9AFE7B3C87E0BE" box="[297,391,408,428]" pageId="16" pageNumber="1447" value="1991-05-13">
<collectingDate id="EFD07D1A7C4BFFCFFE9AFE7B3C87E0BE" box="[297,391,408,428]" pageId="16" pageNumber="1447" value="1991-05-13">13.V.1991</collectingDate>
</date>
, idem (T 79).
</materialsCitation>
<materialsCitation id="3B42A86F7C4BFFCFFDB8FE7B3939E0BE" ID-GBIF-Occurrence="4417845314" box="[523,1081,408,428]" collectingDate="1990-04-27" collectingDateMax="1991-05-13" collectingDateMin="1990-04-27" collectionCode="cN, cP" collectorName="A. Smetana" country="Taiwan" elevation="2700" location="Cliff" municipality="Yushan National Park" pageId="16" pageNumber="1447" specimenCount="15" stateProvince="Chiayi" typeStatus="paratype">
<collectingRegion id="49EE6CD07C4BFFCFFDB8FE7B3F82E0BE" box="[523,642,408,428]" country="Taiwan" name="Chiayi" pageId="16" pageNumber="1447">Chiayi Hsien</collectingRegion>
: 3, 12: ibidem,
<date id="FF9484F27C4BFFCFFCE9FE7B3EBFE0BE" box="[858,959,408,428]" pageId="16" pageNumber="1447" value="1990-04-27">
<collectingDate id="EFD07D1A7C4BFFCFFCE9FE7B3EBFE0BE" box="[858,959,408,428]" pageId="16" pageNumber="1447" value="1990-04-27">27.IV.1990</collectingDate>
</date>
, idem (T 28)
</materialsCitation>
;
<materialsCitation id="3B42A86F7C4BFFCFFF6CFE533F8EE0CE" ID-GBIF-Occurrence="4417845533" collectingDate="1990-04-27" collectingDateMax="1991-05-13" collectingDateMin="1990-04-27" collectionCode="cN, cP" collectorName="Smetana" country="Taiwan" elevation="2750" location="Ghia" municipality="Yushan National Park" pageId="16" pageNumber="1447" specimenCount="4" stateProvince="Chiayi" typeStatus="paratype">
3, 1:
<collectingMunicipality id="6BF138487C4BFFCFFEF7FE533F08E0D6" box="[324,520,432,452]" pageId="16" pageNumber="1447">Yushan National Park</collectingMunicipality>
, Ta-Ta
<location id="8EF5F4E97C4BFFCFFDFFFE533F79E0D6" LSID="urn:lsid:plazi:treatment:038313247C4BFFCFFF01FEDB3C36E5C8:8EF5F4E97C4BFFCFFDFFFE533F79E0D6" box="[588,633,432,452]" municipality="Yushan National Park" name="Ghia" pageId="16" pageNumber="1447">Ghia</location>
,
<quantity id="4CD20FD77C4BFFCFFD31FE533FC2E0D6" box="[642,706,432,452]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="16" pageNumber="1447" unit="m" value="2750.0">
<elevation id="000745017C4BFFCFFD31FE533FC2E0D6" box="[642,706,432,452]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="16" pageNumber="1447" unit="m" value="2750.0">2750m</elevation>
</quantity>
,
<date id="FF9484F27C4BFFCFFD78FE533E30E0D6" box="[715,816,432,452]" pageId="16" pageNumber="1447" value="1990-04-27">
<collectingDate id="EFD07D1A7C4BFFCFFD78FE533E30E0D6" box="[715,816,432,452]" pageId="16" pageNumber="1447" value="1990-04-27">27.IV.1990</collectingDate>
</date>
, idem (T 27).- HT and PTT in coll.
<collectorName id="26DFC7E47C4BFFCFFEB9FE2B3C55E0CE" box="[266,341,456,476]" pageId="16" pageNumber="1447">Smetana</collectorName>
(Ottawa), PTT also in cN and in cP
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C330F1B97C4BFFCFFF01FE083C8FE508" pageId="16" pageNumber="1447" type="description">
<paragraph id="8B95A2327C4BFFCFFF01FE083FDDE312" blockId="16.[178,733,490,512]" box="[178,733,490,512]" pageId="16" pageNumber="1447">
Description: About as in
<taxonomicName id="4C2AD9B17C4BFFCFFDBBFE083F8DE312" authority="Puthz, 2012" authorityName="Puthz" authorityYear="2012" box="[520,653,491,512]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1447" phylum="Arthropoda" rank="species" species="persubtilis" status="sp. nov.">
<emphasis id="B95E7E207C4BFFCFFDBBFE083F8DE312" box="[520,653,491,512]" italics="true" pageId="16" pageNumber="1447">S. persubtilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26DC35B7C4BFFCFFD20FE083FDDE312" box="[659,733,491,512]" pageId="16" pageNumber="1447" rank="species">nov.sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="8B95A2327C4BFFCFFF01FDF33F6CE334" blockId="16.[178,620,528,550]" box="[178,620,528,550]" pageId="16" pageNumber="1447">
Length: 3.0-
<quantity id="4CD20FD77C4BFFCFFE9DFDF33C76E334" box="[302,374,528,550]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="16" pageNumber="1447" unit="mm" value="3.7">3.7mm</quantity>
(forebody:
<quantity id="4CD20FD77C4BFFCFFE5DFDF33F60E334" box="[494,608,528,550]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="16" pageNumber="1447" unit="mm" value="1.7" valueMax="1.8" valueMin="1.6">1.6-1.8mm</quantity>
).
</paragraph>
<paragraph id="8B95A2327C4BFFCFFF01FDD53EA8E359" blockId="16.[178,936,565,587]" box="[178,936,565,587]" pageId="16" pageNumber="1447">PM of the HT: HW: 32; DE: 16; PW: 25; PL: 25; EW: 30; EL: 26; SL: 20.</paragraph>
<paragraph id="8B95A2327C4BFFCFFF01FDB83E1AE22F" blockId="16.[178,1086,602,829]" pageId="16" pageNumber="1447">Male: Metasternum broadly and shallowly impressed, punctation coarse and dense, interstices smaller than half diameter of punctures. Anterior sternites simple, sternite 5 with a very short, broad, shallow, impunctate impression near posterior margin, sternite 6 and 7 each with a broad and deep median impression, which is extremely finely and densely punctate and pubescent, posterior margin very shallowly emarginate. Sternite 8 (fig. 4). Sternite 9 with long, light apicolateral teeth. Aedeagus (fig. 20), median lobe triangularly narrowed, apex acute, expulsion hooks contiguous; parameres much longer than median lobe, with about 10+2 long setae in apical third.</paragraph>
<paragraph id="8B95A2327C4BFFCFFF01FCAE3EBDE292" blockId="16.[178,1085,844,896]" pageId="16" pageNumber="1447">Female: Sternite 8 at posterior margin slightly projecting, rounded. Apicolateral tooth of valvifer long. Spermatheca (fig. 66), infundibulum narrow and long.</paragraph>
<paragraph id="8B95A2327C4BFFCFFF01FC6C3C8FE508" blockId="16.[178,1086,911,1050]" pageId="16" pageNumber="1447">
Distinctly larger than
<taxonomicName id="4C2AD9B17C4BFFCFFE25FC733F1CE2B6" authorityName="Puthz" authorityYear="2012" box="[406,540,911,933]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1447" phylum="Arthropoda" rank="species" species="persubtilis">
<emphasis id="B95E7E207C4BFFCFFE25FC733F1CE2B6" box="[406,540,911,933]" italics="true" pageId="16" pageNumber="1447">S. persubtilis</emphasis>
</taxonomicName>
, median portion of frons comparatively flat, not distinctly exceeding beyond the level of medial eye margins. Pronotum and elytra with very deep impressions, median impression of pronotum broad and broadly impunctate. Abdominal punctation less dense than in
<taxonomicName id="4C2AD9B17C4BFFCFFDEDFC043FE5E2EE" authorityName="Puthz" authorityYear="2012" box="[606,741,999,1020]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1447" phylum="Arthropoda" rank="species" species="persubtilis">
<emphasis id="B95E7E207C4BFFCFFDEDFC043FE5E2EE" box="[606,741,999,1020]" italics="true" pageId="16" pageNumber="1447">S. persubtilis</emphasis>
</taxonomicName>
, interstices on tergite 7 distinctly larger than punctures.
</paragraph>
</subSubSection>
<subSubSection id="C330F1B97C4BFFCFFF01FBCA3E44E585" pageId="16" pageNumber="1447" type="discussion">
<paragraph id="8B95A2327C4BFFCFFF01FBCA3E44E585" blockId="16.[178,1085,1065,1175]" pageId="16" pageNumber="1447">
Comparative notes: This new species is the largest of the Taiwanese species of the
<taxonomicName id="4C2AD9B17C4BFFCFFF44FBA43C7CE54E" box="[247,380,1095,1116]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1447" phylum="Arthropoda" rank="species" species="cephalotes">
<emphasis id="B95E7E207C4BFFCFFF44FBA43C7CE54E" box="[247,380,1095,1116]" italics="true" pageId="16" pageNumber="1447">S. cephalotes</emphasis>
</taxonomicName>
-group. Beside of the sexual characters it may be distinguished from most of the close relatives by the strong impunctate median impression of the pronotum and by the relatively sparse punctation of the posterior abdomen.
</paragraph>
</subSubSection>
<subSubSection id="C330F1B97C4BFFCFFF01FB443C36E5C8" pageId="16" pageNumber="1447" type="etymology">
<paragraph id="8B95A2327C4BFFCFFF01FB443C36E5C8" blockId="16.[178,1085,1190,1242]" pageId="16" pageNumber="1447">
Etymology: Because of its robust habitus I name this species &quot;
<emphasis id="B95E7E207C4BFFCFFC33FB443EECE5AE" box="[896,1004,1191,1212]" italics="true" pageId="16" pageNumber="1447">perpinguis</emphasis>
&quot; (Lat.= very plump).
</paragraph>
</subSubSection>
</treatment>
</document>
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