276 lines
31 KiB
XML
276 lines
31 KiB
XML
<document id="24164AC0015485D4EB4DAD29AAACB8AD" ID-DOI="10.5281/zenodo.207073" ID-GBIF-Dataset="a3815507-6100-464e-aaf6-60d6820a1eec" ID-ISSN="1175-5326" ID-Zenodo-Dep="207073" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460228776049" checkinUser="plazi" docAuthor="Torres-Carvajal, Omar, Etheridge, Richard & Queiroz, Kevin De" docDate="2011" docId="038287FDFF9B606769838A91D5E9F9FF" docLanguage="en" docName="zt02752p044.pdf" docOrigin="Zootaxa 2752" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Morunasaurus Dunn 1933" docType="treatment" docVersion="9" lastPageNumber="33" masterDocId="FFBBFF85FFBB604769148B6ED174FF93" masterDocTitle="A systematic revision of Neotropical lizards in the clade Hoplocercinae (Squamata: Iguania)" masterLastPageNumber="44" masterPageNumber="1" pageNumber="33" updateTime="1698238838839" updateUser="plazi">
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<mods:titleInfo id="7DE1A36E3A898AFABE76440B6EAD9505">
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<mods:title id="C3995EF260C20BDB31DC7A8A7F8A6EF5">A systematic revision of Neotropical lizards in the clade Hoplocercinae (Squamata: Iguania)</mods:title>
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<mods:namePart id="10BF13C7BD774A04434B8DC9DF134DCD">Torres-Carvajal, Omar</mods:namePart>
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<mods:namePart id="8A663F9ED1AD31C272DF1F996795BB5F">Etheridge, Richard</mods:namePart>
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<mods:namePart id="67FAA5DEF063E9DE6E1273FED89F8A68">Queiroz, Kevin De</mods:namePart>
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<mods:typeOfResource id="EF341B79F984603F4E5F96DD4EAD3DC3">text</mods:typeOfResource>
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<mods:title id="2FD37B2064FA9F3C0E47DF88728037CE">Zootaxa</mods:title>
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<mods:part id="9CA0813C37E2BED92044B7720AE654C7">
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<mods:date id="8B21206E5607EB8BCDE94208EB278C17">2011</mods:date>
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<mods:detail id="641C2E4EABCD70925FBFDB0EDC403EFE" type="volume">
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<mods:number id="E54862485FDB053D0E804381C391CF22">2752</mods:number>
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<mods:identifier id="989274D6A353B4D9CF94114192783824" type="DOI">10.5281/zenodo.207073</mods:identifier>
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<mods:identifier id="D1ACA48C915250EDA6245B0C0DDF961F" type="ISSN">1175-5326</mods:identifier>
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<treatment id="038287FDFF9B606769838A91D5E9F9FF" ID-DOI="http://doi.org/10.5281/zenodo.6184981" ID-GBIF-Taxon="119387052" ID-Zenodo-Dep="6184981" LSID="urn:lsid:plazi:treatment:038287FDFF9B606769838A91D5E9F9FF" httpUri="http://treatment.plazi.org/id/038287FDFF9B606769838A91D5E9F9FF" lastPageNumber="33" pageId="32" pageNumber="33">
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<subSubSection id="C3316560FF9B606769838A91D315FDE8" pageId="32" pageNumber="33" type="nomenclature">
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<paragraph id="8B9436EBFF9B606769838A91D0AEFD8A" blockId="32.[151,474,511,537]" box="[151,474,511,537]" pageId="32" pageNumber="33">
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<heading id="D0DC8187FF9B606769838A91D0AEFD8A" bold="true" box="[151,474,511,537]" fontSize="11" level="1" pageId="32" pageNumber="33" reason="1">
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<taxonomicName id="4C2B4D68FF9B606769838A91D0AEFD8A" authority="Dunn 1933" authorityName="Dunn" authorityYear="1933" box="[151,474,511,537]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="genus">
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<emphasis id="B95FEAF9FF9B606769838A91D0AEFD8A" bold="true" box="[151,474,511,537]" pageId="32" pageNumber="33">
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<emphasis id="B95FEAF9FF9B606769838A91D038FD8A" bold="true" box="[151,332,511,537]" italics="true" pageId="32" pageNumber="33">Morunasaurus</emphasis>
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<bibRefCitation id="EFBA4B1AFF9B606768478A91D0AEFD8A" author="Dunn" box="[339,474,511,537]" pageId="32" pageNumber="39" refString="Dunn, E. R. (1933) Amphibians and reptiles from El Valle de Anton, Panama. Occasional Papers of the Boston Society of Natural History, 8, 65 - 79." type="journal article" year="1933">Dunn 1933</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8B9436EBFF9B606769838928D315FDE8" blockId="32.[151,609,582,635]" pageId="32" pageNumber="33">Proposed standard English name: manticores Proposed standard Spanish name: mantícoras</paragraph>
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</subSubSection>
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<subSubSection id="C3316560FF9B6067698389F3D563FC53" pageId="32" pageNumber="33" type="reference_group">
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<paragraph id="8B9436EBFF9B6067698389F3D042FD63" blockId="32.[151,1436,669,752]" pageId="32" pageNumber="33">
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<taxonomicName id="4C2B4D68FF9B6067698389F3D06CFD20" box="[151,280,669,691]" class="Reptilia" family="Hoplocercidae" genus="Hoplocercus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="genus">
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<emphasis id="B95FEAF9FF9B6067698389F3D06CFD20" box="[151,280,669,691]" italics="true" pageId="32" pageNumber="33">Hoplocercus</emphasis>
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</taxonomicName>
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(part)
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<bibRefCitation id="EFBA4B1AFF9B6067684889F3D34AFD20" author="O'Shaughnessy" box="[348,574,669,691]" pageId="32" pageNumber="40" refString="O'Shaughnessy, A. W. E. (1881) An account of the collection of lizards made by Mr. Buckley in Ecuador, and now in the British Museum, with descriptions of the new species. Proceedings of the Zoological Society of London, 1881, 227 - 245." type="proceedings paper" year="1881">O’Shaughnessy (1881</bibRefCitation>
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:244);
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<bibRefCitation id="EFBA4B1AFF9B60676B6B89F3D217FD20" author="Boulenger" box="[639,867,669,691]" pageId="32" pageNumber="38" refString="Boulenger, G. A. (1885) Catalogue of the Lizards in the British Museum (Natural History) II. Taylor and Francis, London, 497 pp." type="book" year="1885">Boulenger (1885:200)</bibRefCitation>
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;
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<bibRefCitation id="EFBA4B1AFF9B60676A7989F0D520FD20" author="Burt" box="[877,1108,669,691]" pageId="32" pageNumber="38" refString="Burt, C. E. & Burt, M. D. (1933) A preliminary check list of the lizards of South America. Transactions of the Academy of Sciences of St Louis, 28, 1 - 104." type="journal article" year="1933">Burt & Burt (1933:26)</bibRefCitation>
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.
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<taxonomicName id="4C2B4D68FF9B6067698389D3D0A8FD42" authority="Dunn (1933:75)" authorityName="Dunn" authorityPageNumber="75" authorityYear="1933" box="[151,476,699,722]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="genus">
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<emphasis id="B95FEAF9FF9B6067698389D3D058FD41" box="[151,300,701,722]" italics="true" pageId="32" pageNumber="33">Morunasaurus</emphasis>
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<bibRefCitation id="EFBA4B1AFF9B6067682089D3D0A8FD42" author="Dunn" box="[308,476,699,722]" pageId="32" pageNumber="39" refString="Dunn, E. R. (1933) Amphibians and reptiles from El Valle de Anton, Panama. Occasional Papers of the Boston Society of Natural History, 8, 65 - 79." type="journal article" year="1933">Dunn (1933:75)</bibRefCitation>
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</taxonomicName>
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;
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<bibRefCitation id="EFBA4B1AFF9B606768FC89D3D216FD41" author="Peters" box="[488,866,700,722]" pageId="32" pageNumber="40" refString="Peters, J. A. & Donoso-Barros, R. (1970) Catalogue of the neotropical Squamata: Part II. Lizards and amphisbaenians. United States National Museum Bulletin, 297, 1 - 293." type="journal article" year="1970">Peters & Donoso-Barros (1970:204)</bibRefCitation>
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.
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<typeStatus id="54908849FF9B60676A7989D3D2EBFD41" box="[877,927,701,722]" pageId="32" pageNumber="33">Type</typeStatus>
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species (by original designation):
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<taxonomicName id="4C2B4D68FF9B60676C1389D3D042FD63" authority="Dunn." authorityName="Dunn." class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="groi">
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<emphasis id="B95FEAF9FF9B60676C1389D3D184FD63" italics="true" pageId="32" pageNumber="33">Morunasaurus groi</emphasis>
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Dunn.
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</taxonomicName>
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</paragraph>
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<paragraph id="8B9436EBFF9B606769838879D563FC53" blockId="32.[151,1437,791,1644]" pageId="32" pageNumber="33">
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<emphasis id="B95FEAF9FF9B606769838879D062FCA3" bold="true" box="[151,278,791,816]" pageId="32" pageNumber="33">Definition.</emphasis>
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<taxonomicName id="4C2B4D68FF9B6067680B8877D325FCBC" authority="Dunn 1933" authorityName="Dunn" authorityYear="1933" box="[287,593,791,816]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="genus">
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<emphasis id="B95FEAF9FF9B6067680B8877D0B0FCA3" box="[287,452,793,816]" italics="true" pageId="32" pageNumber="33">Morunasaurus</emphasis>
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<bibRefCitation id="EFBA4B1AFF9B606768DA8876D325FCBC" author="Dunn" box="[462,593,791,815]" pageId="32" pageNumber="39" refString="Dunn, E. R. (1933) Amphibians and reptiles from El Valle de Anton, Panama. Occasional Papers of the Boston Society of Natural History, 8, 65 - 79." type="journal article" year="1933">Dunn 1933</bibRefCitation>
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</taxonomicName>
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, converted clade name, is here defined as referring to the most inclusive crown clade containing
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<taxonomicName id="4C2B4D68FF9B606768BE8853D266FCC7" authority="Dunn 1933" authorityName="Dunn" authorityYear="1933" box="[426,786,828,852]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="groi">
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<emphasis id="B95FEAF9FF9B606768BE8853D3F2FCC7" box="[426,646,829,852]" italics="true" pageId="32" pageNumber="33">Morunasaurus groi</emphasis>
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<bibRefCitation id="EFBA4B1AFF9B60676B9B8853D266FCC7" author="Dunn" box="[655,786,828,852]" pageId="32" pageNumber="39" refString="Dunn, E. R. (1933) Amphibians and reptiles from El Valle de Anton, Panama. Occasional Papers of the Boston Society of Natural History, 8, 65 - 79." type="journal article" year="1933">Dunn 1933</bibRefCitation>
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</taxonomicName>
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, but not
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<taxonomicName id="4C2B4D68FF9B60676A6D8852D457FCC7" authority="Fitzinger 1843" authorityName="Fitzinger" authorityYear="1843" box="[889,1315,828,852]" class="Reptilia" family="Hoplocercidae" genus="Hoplocercus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="spinosus">
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<emphasis id="B95FEAF9FF9B60676A6D8852D504FCC7" box="[889,1136,828,852]" italics="true" pageId="32" pageNumber="33">Hoplocercus spinosus</emphasis>
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<bibRefCitation id="EFBA4B1AFF9B60676D6E8852D457FCC7" author="Fitzinger" box="[1146,1315,828,852]" pageId="32" pageNumber="39" refString="Fitizinger, L. (1843) Systema Reptilium, Fasciculus Primus, Amblyglossae. Baumuller und Seidel, Wien, 106 pp." type="book" year="1843">Fitzinger 1843</bibRefCitation>
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</taxonomicName>
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, or
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<taxonomicName id="4C2B4D68FF9B60676C458852D364FCE4" authority="Bocourt 1874" authorityName="Bocourt" authorityYear="1874" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="heterolepis">
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<emphasis id="B95FEAF9FF9B60676C458852D02FFCEB" italics="true" pageId="32" pageNumber="33">Enyalioides heterolepis</emphasis>
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(
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<bibRefCitation id="EFBA4B1AFF9B6067687E8831D37CFCE4" author="Bocourt" box="[362,520,863,887]" pageId="32" pageNumber="38" refString="Bocourt, M. F. (1874) Deux notes sur quelques sauriens de l'Amerique tropicale. Annales des Sciences Naturelles. Zoologie et Biologie Animale, 19, 1 - 5." type="journal article" year="1874">Bocourt 1874</bibRefCitation>
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)
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</taxonomicName>
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, or
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<taxonomicName id="4C2B4D68FF9B60676B2D880ED543FCE4" authority="Boulenger 1885" authorityName="Boulenger" authorityYear="1885" box="[569,1079,863,888]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="oshaughnessyi">
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<emphasis id="B95FEAF9FF9B60676B2D880ED213FCEB" box="[569,871,864,888]" italics="true" pageId="32" pageNumber="33">Enyalioides oshaughnessyi</emphasis>
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(
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<bibRefCitation id="EFBA4B1AFF9B60676A628831D55BFCE4" author="Boulenger" box="[886,1071,863,887]" pageId="32" pageNumber="38" refString="Boulenger, G. A. (1885) Catalogue of the Lizards in the British Museum (Natural History) II. Taylor and Francis, London, 497 pp." type="book" year="1885">Boulenger 1885</bibRefCitation>
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)
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</taxonomicName>
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. Reference phylogeny:
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<figureCitation id="13102A6EFF9B60676C2A8831D4E8FCE4" box="[1342,1436,863,887]" captionStart="FIGURE 3" captionStartId="12.[151,250,1517,1540]" captionTargetBox="[280,1305,321,1491]" captionTargetId="figure@12.[274,1314,306,1505]" captionTargetPageId="12" captionText="FIGURE 3. Caudal segments from the anterior third of the tail of ten species of Hoplocercinae in lateral view. (A) Enyalioides cofanorum, (B) E. heterolepis, (C) E. laticeps, (D) E. microlepis, (E) E. oshaughnessyi, (F) E. palpebralis, (G) E. praestabilis, (H) E. rubrigularis, (I) E. touzeti, and (J) Morunasaurus annularis. Scale bars = 5 mm. Illustration by D. Paucar." httpUri="https://zenodo.org/record/207076/files/figure.png" pageId="32" pageNumber="33">
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Figure
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<quantity id="4CD39B0EFF9B60676C9A8831D4E8FCE4" box="[1422,1436,863,887]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="32" pageNumber="33" unit="in" value="3.0">3</quantity>
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</figureCitation>
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in Torres-Carvajal & de Queiroz (2009), in which it is assumed that
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<taxonomicName id="4C2B4D68FF9B60676A8D88EBD506FC0F" box="[921,1138,901,924]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="groi">
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<emphasis id="B95FEAF9FF9B60676A8D88EBD506FC0F" box="[921,1138,901,924]" italics="true" pageId="32" pageNumber="33">Morunasaurus groi</emphasis>
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</taxonomicName>
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is part of a clade that also includes
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<taxonomicName id="4C2B4D68FF9B606769EF88C7D0F8FC53" box="[251,396,936,960]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="annularis">
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<emphasis id="B95FEAF9FF9B606769EF88C7D0F8FC53" box="[251,396,936,960]" italics="true" pageId="32" pageNumber="33">M. annularis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C2B4D68FF9B606768D688C7D317FC53" box="[450,611,937,960]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="peruvianus">
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<emphasis id="B95FEAF9FF9B606768D688C7D317FC53" box="[450,611,937,960]" italics="true" pageId="32" pageNumber="33">M. peruvianus</emphasis>
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</taxonomicName>
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, but none of the other taxa in that tree.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3316560FF9B606769D188A5D0B5FA97" pageId="32" pageNumber="33" type="diagnosis">
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<paragraph id="8B9436EBFF9B606769D188A5D23EFB2F" blockId="32.[151,1437,791,1644]" pageId="32" pageNumber="33">
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<emphasis id="B95FEAF9FF9B606769D188A5D04BFC77" bold="true" box="[197,319,971,996]" pageId="32" pageNumber="33">Diagnosis.</emphasis>
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<taxonomicName id="4C2B4D68FF9B6067685E88A3D09BFC77" box="[330,495,973,996]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="genus">
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<emphasis id="B95FEAF9FF9B6067685E88A3D09BFC77" box="[330,495,973,996]" italics="true" pageId="32" pageNumber="33">Morunasaurus</emphasis>
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</taxonomicName>
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can be distinguished from
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<taxonomicName id="4C2B4D68FF9B60676A2788A2D2B5FC77" box="[819,961,972,996]" class="Reptilia" family="Hoplocercidae" genus="Hoplocercus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="genus">
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<emphasis id="B95FEAF9FF9B60676A2788A2D2B5FC77" box="[819,961,972,996]" italics="true" pageId="32" pageNumber="33">Hoplocercus</emphasis>
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</taxonomicName>
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by having a tail that is roughly circular (rather than depressed) in cross section and longer than the body (i.e., tail length> SVL). It differs from both
|
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<taxonomicName id="4C2B4D68FF9B60676C76889FD181FBBF" class="Reptilia" family="Hoplocercidae" genus="Hoplocercus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="genus">
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||
<emphasis id="B95FEAF9FF9B60676C76889FD181FBBF" italics="true" pageId="32" pageNumber="33">Hoplocercus</emphasis>
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</taxonomicName>
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and
|
||
<taxonomicName id="4C2B4D68FF9B6067683F8F7AD0C4FBBF" box="[299,432,1044,1068]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="genus">
|
||
<emphasis id="B95FEAF9FF9B6067683F8F7AD0C4FBBF" box="[299,432,1044,1068]" italics="true" pageId="32" pageNumber="33">Enyalioides</emphasis>
|
||
</taxonomicName>
|
||
in having projecting scales (spines) on thigh, shin, and pes (projecting scales also present in
|
||
<taxonomicName id="4C2B4D68FF9B606769A18F57D025FBC3" box="[181,337,1080,1104]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="heterolepis">
|
||
<emphasis id="B95FEAF9FF9B606769A18F57D025FBC3" box="[181,337,1080,1104]" italics="true" pageId="32" pageNumber="33">E. heterolepis</emphasis>
|
||
</taxonomicName>
|
||
, but not as spines); a dentary bone extending posteriorly above the anterior surangular foramen (
|
||
<bibRefCitation id="EFBA4B1AFF9B6067698A8F32D0CBFBE7" author="Wiens" box="[158,447,1116,1140]" pageId="32" pageNumber="41" refString="Wiens, J. J. & Etheridge, R. (2003) Phylogenetic relationships of hoplocercid lizards: Coding and combining meristic, morphometric, and polymorphic data using step matrices. Herpetologica, 59, 375 - 398." type="journal article" year="2003">Wiens & Etheridge 2003</bibRefCitation>
|
||
), and in lacking a parietal eye.
|
||
<taxonomicName id="4C2B4D68FF9B60676A3C8F33D2B9FBE7" box="[808,973,1117,1140]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="genus">
|
||
<emphasis id="B95FEAF9FF9B60676A3C8F33D2B9FBE7" box="[808,973,1117,1140]" italics="true" pageId="32" pageNumber="33">Morunasaurus</emphasis>
|
||
</taxonomicName>
|
||
also differs from
|
||
<taxonomicName id="4C2B4D68FF9B60676D888F32D454FBE7" box="[1180,1312,1116,1140]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="genus">
|
||
<emphasis id="B95FEAF9FF9B60676D888F32D454FBE7" box="[1180,1312,1116,1140]" italics="true" pageId="32" pageNumber="33">Enyalioides</emphasis>
|
||
</taxonomicName>
|
||
in that the posterior whorl of each caudal segment is composed of greatly enlarged, projecting, spinous scales at least two times as large as the scales of the immediately anterior whorl.
|
||
</paragraph>
|
||
<paragraph id="8B9436EBFF9B606769D38FA9D0B5FA97" blockId="32.[151,1437,791,1644]" pageId="32" pageNumber="33">
|
||
<emphasis id="B95FEAF9FF9B606769D38FA9D013FB73" bold="true" box="[199,359,1223,1248]" pageId="32" pageNumber="33">Composition.</emphasis>
|
||
Three species:
|
||
<taxonomicName id="4C2B4D68FF9B60676B348FA7D53DFB73" authority="O'Shaughnessy 1881" authorityName="O'Shaughnessy" authorityYear="1881" box="[544,1097,1224,1248]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="annularis">
|
||
<emphasis id="B95FEAF9FF9B60676B348FA7D24EFB73" box="[544,826,1224,1248]" italics="true" pageId="32" pageNumber="33">Morunasaurus annularis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBA4B1AFF9B60676A598FA6D534FB73" author="O'Shaughnessy" box="[845,1088,1224,1248]" pageId="32" pageNumber="40" refString="O'Shaughnessy, A. W. E. (1881) An account of the collection of lizards made by Mr. Buckley in Ecuador, and now in the British Museum, with descriptions of the new species. Proceedings of the Zoological Society of London, 1881, 227 - 245." type="proceedings paper" year="1881">O'Shaughnessy 1881</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2B4D68FF9B60676D438FA7D448FB73" authority="Dunn 1933" authorityName="Dunn" authorityYear="1933" box="[1111,1340,1224,1248]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="groi">
|
||
<emphasis id="B95FEAF9FF9B60676D438FA7D5DAFB73" box="[1111,1198,1225,1248]" italics="true" pageId="32" pageNumber="33">M. groi</emphasis>
|
||
<bibRefCitation id="EFBA4B1AFF9B60676DAC8FA7D448FB73" author="Dunn" box="[1208,1340,1224,1248]" pageId="32" pageNumber="39" refString="Dunn, E. R. (1933) Amphibians and reptiles from El Valle de Anton, Panama. Occasional Papers of the Boston Society of Natural History, 8, 65 - 79." type="journal article" year="1933">Dunn 1933</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C2B4D68FF9B60676C698FA7D0C9FA97" authority="Kohler 2003" authorityName="Kohler" authorityYear="2003" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="peruvianus">
|
||
<emphasis id="B95FEAF9FF9B60676C698FA7D060FA97" italics="true" pageId="32" pageNumber="33">M. peruvianus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBA4B1AFF9B606768368F82D0C1FA97" author="Kohler" box="[290,437,1260,1284]" pageId="32" pageNumber="40" refString="Kohler, G. (2003) A new species of Morunasaurus from Peru. Senckenbergiana Biologica, 82, 235 - 241." type="journal article" year="2003">Köhler 2003</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3316560FF9B606769D38E7ED5E9F9FF" pageId="32" pageNumber="33" type="discussion">
|
||
<paragraph id="8B9436EBFF9B606769D38E7ED02BF9B7" blockId="32.[151,1437,791,1644]" pageId="32" pageNumber="33">
|
||
<emphasis id="B95FEAF9FF9B606769D38E7ED04FFABB" bold="true" box="[199,315,1296,1320]" pageId="32" pageNumber="33">Remarks.</emphasis>
|
||
The phylogenetic position of
|
||
<taxonomicName id="4C2B4D68FF9B60676B888E7FD235FABB" box="[668,833,1297,1320]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="genus">
|
||
<emphasis id="B95FEAF9FF9B60676B888E7FD235FABB" box="[668,833,1297,1320]" italics="true" pageId="32" pageNumber="33">Morunasaurus</emphasis>
|
||
</taxonomicName>
|
||
relative to other hoplocercine taxa remains unclear. Etheridge & de Queiroz (1988) inferred a phylogeny in which
|
||
<taxonomicName id="4C2B4D68FF9B60676A4A8E5BD577FADF" box="[862,1027,1333,1356]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="genus">
|
||
<emphasis id="B95FEAF9FF9B60676A4A8E5BD577FADF" box="[862,1027,1333,1356]" italics="true" pageId="32" pageNumber="33">Morunasaurus</emphasis>
|
||
</taxonomicName>
|
||
was paraphyletic and deeply nested within
|
||
<taxonomicName id="4C2B4D68FF9B606769F18E36D01DFAE3" box="[229,361,1368,1392]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="genus">
|
||
<emphasis id="B95FEAF9FF9B606769F18E36D01DFAE3" box="[229,361,1368,1392]" italics="true" pageId="32" pageNumber="33">Enyalioides</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFBA4B1AFF9B606768638E36D3EBFAE3" author="Wiens" box="[375,671,1368,1392]" pageId="32" pageNumber="41" refString="Wiens, J. J. & Etheridge, R. (2003) Phylogenetic relationships of hoplocercid lizards: Coding and combining meristic, morphometric, and polymorphic data using step matrices. Herpetologica, 59, 375 - 398." type="journal article" year="2003">Wiens & Etheridge (2003)</bibRefCitation>
|
||
found
|
||
<taxonomicName id="4C2B4D68FF9B60676BF98E37D2E6FAE3" box="[749,914,1369,1392]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="genus">
|
||
<emphasis id="B95FEAF9FF9B60676BF98E37D2E6FAE3" box="[749,914,1369,1392]" italics="true" pageId="32" pageNumber="33">Morunasaurus</emphasis>
|
||
</taxonomicName>
|
||
to be either paraphyletic, or monophyletic and sister to
|
||
<taxonomicName id="4C2B4D68FF9B606769E08E12D09DFA07" box="[244,489,1404,1428]" class="Reptilia" family="Hoplocercidae" genus="Hoplocercus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="spinosus">
|
||
<emphasis id="B95FEAF9FF9B606769E08E12D09DFA07" box="[244,489,1404,1428]" italics="true" pageId="32" pageNumber="33">Hoplocercus spinosus</emphasis>
|
||
</taxonomicName>
|
||
. Torres-Carvajal & de Queiroz (2009) found
|
||
<taxonomicName id="4C2B4D68FF9B60676AF78E13D5FCFA07" box="[995,1160,1405,1428]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="genus">
|
||
<emphasis id="B95FEAF9FF9B60676AF78E13D5FCFA07" box="[995,1160,1405,1428]" italics="true" pageId="32" pageNumber="33">Morunasaurus</emphasis>
|
||
</taxonomicName>
|
||
to be nested within
|
||
<taxonomicName id="4C2B4D68FF9B60676C738E13D184FA2B" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="genus">
|
||
<emphasis id="B95FEAF9FF9B60676C738E13D184FA2B" italics="true" pageId="32" pageNumber="33">Enyalioides</emphasis>
|
||
</taxonomicName>
|
||
, but they could not reject the monophyly of
|
||
<taxonomicName id="4C2B4D68FF9B60676BFE8ECED21AFA2B" box="[746,878,1440,1464]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="genus">
|
||
<emphasis id="B95FEAF9FF9B60676BFE8ECED21AFA2B" box="[746,878,1440,1464]" italics="true" pageId="32" pageNumber="33">Enyalioides</emphasis>
|
||
</taxonomicName>
|
||
statistically; however, their study did not include
|
||
<taxonomicName id="4C2B4D68FF9B606769838EABD19EFA4F" box="[151,234,1477,1500]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="groi">
|
||
<emphasis id="B95FEAF9FF9B606769838EABD19EFA4F" box="[151,234,1477,1500]" italics="true" pageId="32" pageNumber="33">M. groi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54908849FF9B606769ED8EAAD05FFA4F" box="[249,299,1476,1500]" pageId="32" pageNumber="33">type</typeStatus>
|
||
species), or nuclear gene sequences from
|
||
<taxonomicName id="4C2B4D68FF9B60676A118EABD2D2FA4F" box="[773,934,1477,1500]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="peruvianus">
|
||
<emphasis id="B95FEAF9FF9B60676A118EABD2D2FA4F" box="[773,934,1477,1500]" italics="true" pageId="32" pageNumber="33">M. peruvianus</emphasis>
|
||
</taxonomicName>
|
||
. If
|
||
<taxonomicName id="4C2B4D68FF9B60676ADB8EABD555FA4F" box="[975,1057,1477,1500]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="groi">
|
||
<emphasis id="B95FEAF9FF9B60676ADB8EABD555FA4F" box="[975,1057,1477,1500]" italics="true" pageId="32" pageNumber="33">M. groi</emphasis>
|
||
</taxonomicName>
|
||
is not closely related to
|
||
<taxonomicName id="4C2B4D68FF9B60676C218EABD1BFF993" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="annularis">
|
||
<emphasis id="B95FEAF9FF9B60676C218EABD1BFF993" italics="true" pageId="32" pageNumber="33">M. annularis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2B4D68FF9B606769EB8E87D0D4F993" box="[255,416,1513,1536]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="peruvianus">
|
||
<emphasis id="B95FEAF9FF9B606769EB8E87D0D4F993" box="[255,416,1513,1536]" italics="true" pageId="32" pageNumber="33">M. peruvianus</emphasis>
|
||
</taxonomicName>
|
||
, then according to our phylogenetic definition, only
|
||
<taxonomicName id="4C2B4D68FF9B60676AFF8E87D56DF993" box="[1003,1049,1513,1536]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="groi">
|
||
<emphasis id="B95FEAF9FF9B60676AFF8E87D56DF993" box="[1003,1049,1513,1536]" italics="true" pageId="32" pageNumber="33">groi</emphasis>
|
||
</taxonomicName>
|
||
(the
|
||
<typeStatus id="54908849FF9B60676D458E87D5F5F992" box="[1105,1153,1513,1537]" pageId="32" pageNumber="33">type</typeStatus>
|
||
species) will be included in
|
||
<taxonomicName id="4C2B4D68FF9B606769A08D63D02DF9B7" box="[180,345,1549,1572]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="genus">
|
||
<emphasis id="B95FEAF9FF9B606769A08D63D02DF9B7" box="[180,345,1549,1572]" italics="true" pageId="32" pageNumber="33">Morunasaurus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9436EBFF9B606769D38D5ED5E9F9FF" blockId="32.[151,1437,791,1644]" pageId="32" pageNumber="33">
|
||
The vernacular name "manticores" ("mantícoras") refers to animals that inhabit mountains near the Moruna (the basis of the name
|
||
<taxonomicName id="4C2B4D68FF9B606768758D3BD372F9FF" box="[353,518,1621,1644]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="genus">
|
||
<emphasis id="B95FEAF9FF9B606768758D3BD372F9FF" box="[353,518,1621,1644]" italics="true" pageId="32" pageNumber="33">Morunasaurus</emphasis>
|
||
</taxonomicName>
|
||
) in E. R. R. Eddison's fantasy novel
|
||
<emphasis id="B95FEAF9FF9B60676AB78D3AD5E2F9FF" box="[931,1174,1620,1644]" italics="true" pageId="32" pageNumber="33">The Worm Ouroboros</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |