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<document id="56F0C81335A079EC1EF60002FEAFA3AB" ID-CLB-Dataset="9473" ID-DOI="10.5852/ejt.2020.723.1151" ID-GBIF-Dataset="e4262fe3-082e-420c-a52c-423429fa41a1" ID-Zenodo-Dep="4298029" ID-ZooBank="F4EF2D7D-2482-4B50-9704-E5E49938294F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1606735545470" checkinUser="plazi" docAuthor="Naumova, Tatyana V. &amp; Gagarin, Vladimir G." docDate="2020" docId="038287F2FFB9AF466770FBD66B94563B" docLanguage="en" docName="ejt-723_naumova_gagarin.pdf" docOrigin="European Journal of Taxonomy 723" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Paratrilobus aquaticus Naumova &amp; Gagarin 2020, sp.nov." docType="treatment" docVersion="10" lastPageNumber="169" masterDocId="FFBBFF8AFFBFAF4C6512FFB66A085313" masterDocTitle="Two new nematode species of the genus Paratrilobus Micoletzky, 1922 (Nematoda, Triplonchida) from the water area of Lake Baikal (Russia)" masterLastPageNumber="172" masterPageNumber="159" pageNumber="165" updateTime="1698846909256" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="B3B152D1DF3534C25332431A82D52463" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AD67E90AE97690D1DB9176DB2BC3F1E5">
<mods:title id="808D5654C849548F9BEB3224920B584B">Two new nematode species of the genus Paratrilobus Micoletzky, 1922 (Nematoda, Triplonchida) from the water area of Lake Baikal (Russia)</mods:title>
</mods:titleInfo>
<mods:name id="DB2BDC5075F5EB2192B75B9902B4C4EE" type="personal">
<mods:role id="11527ADFAEEB304378EBA123DA44D195">
<mods:roleTerm id="65BE056E1F3465B65BE5299EBB28326A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9332F8222A3777C95C8A9D9EF79D3367">Naumova, Tatyana V.</mods:namePart>
<mods:nameIdentifier id="0A51D5DDAE9116C639FF9E6E94AA962D" type="LSID">835C52CB-795C-48BF-B5A0-6DA2C303E060</mods:nameIdentifier>
<mods:affiliation id="46FF366AC936535DF1A1C4A3C0DD0D72">Limnological Institute Siberian Branch of the Russian Academy of Sciences, Irkutsk, 664033, Russia.</mods:affiliation>
<mods:nameIdentifier id="3792924C24275C5B893649F176BC4788" type="email">tvnaum@lin.irk.ru</mods:nameIdentifier>
</mods:name>
<mods:name id="EF6A47E6CE57B4410B75941848357599" type="personal">
<mods:role id="C5A04C1BF39CB58CDA4FF115FD7C819E">
<mods:roleTerm id="8FF776890CC984BEBDA9F6F2C70FB130">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E1395FF2493E9F7F806C23D93FE447BB">Gagarin, Vladimir G.</mods:namePart>
<mods:nameIdentifier id="11A65798049F106181188D5161F5D6AF" type="LSID">E28EA8D1-CBFE-4518-AA60-7F4388FBAD64</mods:nameIdentifier>
<mods:affiliation id="5F220A7B1472EF265641A7BADC9B4EB0">Institute for Biology of Inland Waters, Russian Academy of Sciences, Borok, Yaroslavl Province, 152742, Russia.</mods:affiliation>
<mods:nameIdentifier id="A24E15D79AD23D838AA5717EF18B90F2" type="email">gagarin@ibiw.ru</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="ECB7E443D48360CD7FBE97841A523431">text</mods:typeOfResource>
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<mods:titleInfo id="D2B855BF64B5565C2D30598528C38D3C">
<mods:title id="3F3CC3F9E959358ABE673F7557425B39">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="3F4A547144B7FD6FA4D5CD2BCF39947B">
<mods:date id="F37F21746589FE746B455612FF0B4D4A">2020</mods:date>
<mods:detail id="4C23FA391A69562017E150FB15AF81B5" type="pubDate">
<mods:number id="B05FDA230A96A0C73EE0985F44581954">2020-11-16</mods:number>
</mods:detail>
<mods:detail id="35F6C7C269EDD3A107FD81F7B588D33C" type="volume">
<mods:number id="D6F0BB61B9B459E0A2EB6796CAAC0BCF">723</mods:number>
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<mods:classification id="3C9D6FC58BC9248ED158C713DFFD9E18">journal article</mods:classification>
<mods:identifier id="2E13FA8F41CCA93FA1B8CD197719357A" type="CLB-Dataset">9473</mods:identifier>
<mods:identifier id="4A974D113A2973FE18D015D4642AC60E" type="DOI">10.5852/ejt.2020.723.1151</mods:identifier>
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<treatment id="038287F2FFB9AF466770FBD66B94563B" ID-DOI="http://doi.org/10.5281/zenodo.4328053" ID-GBIF-Taxon="170599258" ID-Zenodo-Dep="4328053" LSID="urn:lsid:plazi:treatment:038287F2FFB9AF466770FBD66B94563B" httpUri="http://treatment.plazi.org/id/038287F2FFB9AF466770FBD66B94563B" lastPageId="10" lastPageNumber="169" pageId="6" pageNumber="165">
<subSubSection id="C331656FFFB9AF4A6770FBD669D95768" box="[610,977,1120,1147]" pageId="6" pageNumber="165" type="nomenclature">
<paragraph id="8B9436E4FFB9AF4A6770FBD669D95768" blockId="6.[358,1228,1120,1222]" box="[610,977,1120,1147]" pageId="6" pageNumber="165">
<heading id="D0DC8188FFB9AF4A6770FBD669D95768" box="[610,977,1120,1147]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="165" reason="2">
<taxonomicName id="4C2B4D67FFB9AF4A6770FBD6697E5769" authority="Naumova &amp; Gagarin, 2020" authorityName="Naumova &amp; Gagarin" authorityYear="2020" box="[610,886,1120,1147]" class="Enoplea" family="Tobrilidae" genus="Paratrilobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Triplonchida" pageId="6" pageNumber="165" phylum="Nematoda" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B95FEAF6FFB9AF4A6770FBD6697E5769" bold="true" box="[610,886,1120,1147]" italics="true" pageId="6" pageNumber="165">Paratrilobus aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C578DFFB9AF4A666CFBD769D95768" box="[894,977,1121,1147]" pageId="6" pageNumber="165" rank="species">sp.nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C331656FFFB9AF4A6474FB30697757D5" pageId="6" pageNumber="165" type="description">
<paragraph id="8B9436E4FFB9AF4A6474FB306EC457B2" blockId="6.[358,1228,1120,1222]" box="[358,1228,1158,1185]" pageId="6" pageNumber="165">
<uri id="FFBA3AE6FFB9AF4A6474FB306EC457B2" box="[358,1228,1158,1185]" pageId="6" pageNumber="165">
urn:lsid:zoobank.org:act:
<uuid id="FF8D0C31FFB9AF4A6796FB306EC457B2" box="[644,1228,1158,1185]" pageId="6" pageNumber="165">AB5F0913-BB36-403D-A732-AEFBB93CA939</uuid>
</uri>
</paragraph>
<paragraph id="8B9436E4FFB9AF4A67A6FB1A697757D5" blockId="6.[358,1228,1120,1222]" box="[692,895,1196,1222]" pageId="6" pageNumber="165">
<figureCitation id="13102A61FFB9AF4A67A6FB1A691557D5" box="[692,797,1196,1222]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="7.[189,232,1892,1918]" captionStartId-1="8.[189,232,1858,1884]" captionTargetBox-0="[285,1318,282,1828]" captionTargetBox-1="[258,1330,266,1820]" captionTargetId-0="figure-80@7.[261,1326,264,1856]" captionTargetId-1="figure-110@8.[252,1335,262,1823]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Fig. 3. Paratrilobus aquaticus sp. nov. A. Male anterior end. B. Female vulva region. C. Female posterior body end. D. Male precloacal supplement. E. Male posterior body end. A, D–E: holotype (HM RAS 102/64 768-2); B–C: paratype (HM RAS 102/64 768-4). Scale bars: A = 30 µm; B = 70 µm; C = 60 µm; D = 20 µm; E = 50 µm." captionText-1="Fig. 4. Paratrilobus aquaticus sp. nov., light micrographs. A. Entire male. B. Entire female. C. Female pharyngeal region.D. Male anterior end.E. Female anterior end.F. Female vulva region.G. Male cloacal region. H. Male posterior body end. I. Female posterior body end. A, D, G–H: holotype (HM RAS 102/64 768-2); B–C, I: paratype (HM RAS 102/64 768-4); E–F: paratype (LIN-SB 765-3). Scale bars: A–B = 200 µm; C, H = 50 µm; D–E = 10 µm; F–G, I = 20 µm." figureDoi-0="http://doi.org/10.5281/zenodo.4298037" figureDoi-1="http://doi.org/10.5281/zenodo.4298041" httpUri-0="https://zenodo.org/record/4298037/files/figure.png" httpUri-1="https://zenodo.org/record/4298041/files/figure.png" pageId="6" pageNumber="165">Figs 3–4</figureCitation>
,
<tableCitation id="C6A9035FFFB9AF4A663AFB1A697757D5" box="[808,895,1196,1222]" captionStart="Table 2" captionStartId="9.[190,259,265,291]" captionTargetPageId="9" captionText="Table 2. Measurements (in µm) of Paratrilobus aquaticus sp. nov., presented as mean and range." httpUri="http://table.plazi.org/id/DF54666CFFB6AF4565ACFEBF6F3A5230" pageId="6" pageNumber="165" tableUuid="DF54666CFFB6AF4565ACFEBF6F3A5230">Table 2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C331656FFFB9AF4A65AFFB5969D4559E" pageId="6" pageNumber="165" type="diagnosis">
<paragraph id="8B9436E4FFB9AF4A65AFFB596B3E561A" blockId="6.[189,310,1263,1289]" box="[189,310,1263,1289]" pageId="6" pageNumber="165">
<heading id="D0DC8188FFB9AF4A65AFFB596B3E561A" bold="true" box="[189,310,1263,1289]" fontSize="11" level="3" pageId="6" pageNumber="165" reason="3">
<emphasis id="B95FEAF6FFB9AF4A65AFFB596B3E561A" bold="true" box="[189,310,1263,1289]" pageId="6" pageNumber="165">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B9436E4FFB9AF4A65AFFA9669D4559E" blockId="6.[188,1401,1312,1678]" pageId="6" pageNumber="165">
<taxonomicName id="4C2B4D67FFB9AF4A65AFFA966BDB5629" authority="Naumova &amp; Gagarin, 2020" authorityName="Naumova &amp; Gagarin" authorityYear="2020" box="[189,467,1312,1338]" class="Enoplea" family="Tobrilidae" genus="Paratrilobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Triplonchida" pageId="6" pageNumber="165" phylum="Nematoda" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B95FEAF6FFB9AF4A65AFFA966BDB5629" box="[189,467,1312,1338]" italics="true" pageId="6" pageNumber="165">Paratrilobus aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C578DFFB9AF4A64F0FA97684C5628" box="[482,580,1313,1339]" pageId="6" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
is characterized by a
<quantity id="4CD39B01FFB9AF4A6677FA966E285628" box="[869,1056,1312,1339]" metricMagnitude="0" metricUnit="m" metricValue="2.109" metricValueMax="2.203" metricValueMin="2.0149999999999997" pageId="6" pageNumber="165" unit="mm" value="2109.0" valueMax="2203.0" valueMin="2015.0">2015–2203 µm</quantity>
body length; cuticle finely annulated; crystalloids absent; six inner labial sensillae papilliform; six outer labial sensillae in the shape of smooth non-articulated setae
<quantity id="4CD39B01FFB9AF4A6723FADA68A15695" box="[561,681,1388,1414]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="6" pageNumber="165" unit="mm" value="19.0" valueMax="20.0" valueMin="18.0">18–20 µm</quantity>
long (53–58% of labial region width); four cephalic sensillae in shape of thin and smooth setae
<quantity id="4CD39B01FFB9AF4A6749FA2468DE56BF" box="[603,726,1426,1452]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="6" pageNumber="165" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">10–13 µm</quantity>
long; buccal cavity spacious, barrel-shaped; one pocket connected with buccal cavity by wide gleam and contains two small teeth. Spicules comparatively thin,
<quantity id="4CD39B01FFB9AF4A65AFFA6B6B0256E4" box="[189,266,1501,1528]" metricMagnitude="-2" metricUnit="m" metricValue="5.1" pageId="6" pageNumber="165" unit="mm" value="51.0">51 µm</quantity>
long, 1.2 times as long as cloacal body diameter; gubernaculum in shape of ‘gutter’. Precloacal supplements
<quantity id="4CD39B01FFB9AF4A6445F9B46B8C550F" box="[343,388,1538,1564]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="6" pageNumber="165" unit="in" value="6.0">6 in</quantity>
number, about same size, located approximately at identical distance from each other; supplements echinate, not very protruded over body surface; contents of ampulla located at base; cap armed, numerous small thorns and one large central thorn. Tail slender, comparatively long; three caudal glands and spinneret well developed; subterminal seta not observed.
</paragraph>
</subSubSection>
<subSubSection id="C331656FFFB9AF4A65AFF90E69295410" box="[189,801,1720,1795]" pageId="6" pageNumber="165" type="etymology">
<paragraph id="8B9436E4FFB9AF4A65AFF90E6B4C55C1" blockId="6.[189,324,1720,1746]" box="[189,324,1720,1746]" pageId="6" pageNumber="165">
<heading id="D0DC8188FFB9AF4A65AFF90E6B4C55C1" bold="true" box="[189,324,1720,1746]" fontSize="11" level="3" pageId="6" pageNumber="165" reason="3">
<emphasis id="B95FEAF6FFB9AF4A65AFF90E6B4C55C1" bold="true" box="[189,324,1720,1746]" pageId="6" pageNumber="165">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B9436E4FFB9AF4A65AFF95E69295410" blockId="6.[189,801,1768,1795]" box="[189,801,1768,1795]" pageId="6" pageNumber="165">The species epithet comes from the habitat ('water').</paragraph>
</subSubSection>
<subSubSection id="C331656FFFB9AF4565AFF89A6F7F56D6" lastPageId="9" lastPageNumber="168" pageId="6" pageNumber="165" type="materials_examined">
<paragraph id="8B9436E4FFB9AF4A65AFF89A6BA25455" blockId="6.[189,426,1836,1862]" box="[189,426,1836,1862]" pageId="6" pageNumber="165">
<heading id="D0DC8188FFB9AF4A65AFF89A6BA25455" bold="true" box="[189,426,1836,1862]" fontSize="11" level="3" pageId="6" pageNumber="165" reason="3">
<emphasis id="B95FEAF6FFB9AF4A65AFF89A6BA25455" bold="true" box="[189,426,1836,1862]" pageId="6" pageNumber="165">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B433CB9FFB9AF4A65F8F8EB68DB54FA" ID-GBIF-Occurrence="2985876303" collectedFrom="sand and silt" collectingDate="2012-07-05" collectionCode="HM RAS" collectorName="T. V. Naumova" country="Russia" county="Lake Baikal" elevation="51" latitude="52.07994" location="Lake Baikal" longLatPrecision="1" longitude="105.90368" municipality="Posolsk Bank" pageId="6" pageNumber="165" specimenCode="768-2" specimenCount="1" typeStatus="holotype">
<paragraph id="8B9436E4FFB9AF4A65F8F8EB6B545464" blockId="6.[188,1398,1885,2025]" box="[234,348,1885,1911]" pageId="6" pageNumber="165">
<emphasis id="B95FEAF6FFB9AF4A65F8F8EB6B545464" bold="true" box="[234,348,1885,1911]" pageId="6" pageNumber="165">
<typeStatus id="54908846FFB9AF4A65F8F8EB6B545464" box="[234,348,1885,1911]" pageId="6" pageNumber="165" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B9436E4FFB9AF4A65AFF83568DB54FA" blockId="6.[188,1398,1885,2025]" pageId="6" pageNumber="165">
<collectingCountry id="F33C7674FFB9AF4A65AFF8356B2C548E" box="[189,292,1923,1949]" name="Russia" pageId="6" pageNumber="165">RUSSIA</collectingCountry>
•
<specimenCount id="9D2DFD6DFFB9AF4A6429F8326B4754B3" box="[315,335,1924,1952]" pageId="6" pageNumber="165" type="male">♂</specimenCount>
;
<location id="8EF4603FFFB9AF4A6472F8356BFA548E" LSID="urn:lsid:plazi:treatment:038287F2FFB9AF466770FBD66B94563B:8EF4603FFFB9AF4A6472F8356BFA548E" box="[352,498,1923,1949]" country="Russia" county="Lake Baikal" latitude="52.07994" longLatPrecision="1" longitude="105.90368" municipality="Posolsk Bank" name="Lake Baikal" pageId="6" pageNumber="165">Lake Baikal</location>
,
<collectingMunicipality id="6BF0AC9EFFB9AF4A64EEF8356894548E" box="[508,668,1923,1949]" pageId="6" pageNumber="165">Posolsk Bank</collectingMunicipality>
(underwater elevation of the bottom between the southern and central basins of Lake Baikal);
<geoCoordinate id="EE1F5023FFB9AF4A6734F81E68B654D0" box="[550,702,1960,1987]" degrees="52.07994" direction="north" orientation="latitude" pageId="6" pageNumber="165" precision="1" value="52.07994">52.07994° N</geoCoordinate>
,
<geoCoordinate id="EE1F5023FFB9AF4A67DBF81E696554D0" box="[713,877,1960,1987]" degrees="105.90368" direction="east" orientation="longitude" pageId="6" pageNumber="165" precision="1" value="105.90368">105.90368° E</geoCoordinate>
;
<elevation id="0006D1D7FFB9AF4A666AF81E69F554D0" box="[888,1021,1960,1987]" metricMagnitude="1" metricUnit="m" metricValue="5.1" pageId="6" pageNumber="165" unit="m" value="51.0">
<quantity id="4CD39B01FFB9AF4A666AF81E69BA54D0" box="[888,946,1960,1987]" metricMagnitude="1" metricUnit="m" metricValue="5.1" pageId="6" pageNumber="165" unit="m" value="51.0">51 m</quantity>
depth
</elevation>
;
<date id="FF951024FFB9AF4A611AF81E6E8254D0" box="[1032,1162,1960,1987]" pageId="6" pageNumber="165" value="2012-07-05">
<collectingDate id="EFD1E9CCFFB9AF4A611AF81E6E8254D0" box="[1032,1162,1960,1987]" pageId="6" pageNumber="165" value="2012-07-05">5 Jul. 2012</collectingDate>
</date>
;
<collectorName id="26DE5332FFB9AF4A6187F81F6F3554D0" box="[1173,1341,1961,1987]" pageId="6" pageNumber="165">T.V. Naumova</collectorName>
leg.;
<collectedFrom id="38377519FFB9AF4A65AEF8786B5A54FB" box="[188,338,1998,2024]" pageId="6" pageNumber="165">sand and silt</collectedFrom>
;
<collectionCode id="ED3AAE21FFB9AF4A644CF8796BD954FB" box="[350,465,1998,2025]" pageId="6" pageNumber="165">HM RAS</collectionCode>
, slide 102/64 (
<specimenCode id="DB8D9E9FFFB9AF4A676FF87868CE54FA" box="[637,710,1998,2025]" pageId="6" pageNumber="165">768-2</specimenCode>
).
</paragraph>
</materialsCitation>
<caption id="DF54666CFFB8AF4B65AFF8D2686954FB" ID-DOI="http://doi.org/10.5281/zenodo.4298037" ID-Zenodo-Dep="4298037" httpUri="https://zenodo.org/record/4298037/files/figure.png" pageId="7" pageNumber="166" startId="7.[189,232,1892,1918]" targetBox="[285,1318,282,1828]" targetPageId="7">
<paragraph id="8B9436E4FFB8AF4B65AFF8D2686954FB" blockId="7.[189,1399,1892,2024]" pageId="7" pageNumber="166">
<emphasis id="B95FEAF6FFB8AF4B65AFF8D26B18546D" bold="true" box="[189,272,1892,1918]" pageId="7" pageNumber="166">Fig. 3.</emphasis>
<taxonomicName id="4C2B4D67FFB8AF4B640EF8D26838546D" authority="Naumova &amp; Gagarin, 2020" authorityName="Naumova &amp; Gagarin" authorityYear="2020" box="[284,560,1892,1918]" class="Enoplea" family="Tobrilidae" genus="Paratrilobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Triplonchida" pageId="7" pageNumber="166" phylum="Nematoda" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B95FEAF6FFB8AF4B640EF8D26838546D" box="[284,560,1892,1918]" italics="true" pageId="7" pageNumber="166">Paratrilobus aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C578DFFB8AF4B672FF8D36895546C" box="[573,669,1893,1919]" pageId="7" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B95FEAF6FFB8AF4B67BBF8D268B6546D" bold="true" box="[681,702,1892,1918]" pageId="7" pageNumber="166">A</emphasis>
. Male anterior end.
<emphasis id="B95FEAF6FFB8AF4B66D2F8D269DC546D" bold="true" box="[960,980,1892,1918]" pageId="7" pageNumber="166">B</emphasis>
. Female vulva region.
<emphasis id="B95FEAF6FFB8AF4B61E5F8D26F04546D" bold="true" box="[1271,1292,1892,1918]" pageId="7" pageNumber="166">C</emphasis>
. Female posterior body end.
<emphasis id="B95FEAF6FFB8AF4B64A1F8316BC054B2" bold="true" box="[435,456,1927,1953]" pageId="7" pageNumber="166">D</emphasis>
. Male precloacal supplement.
<emphasis id="B95FEAF6FFB8AF4B662CF831695A54B2" bold="true" box="[830,850,1927,1953]" pageId="7" pageNumber="166">E</emphasis>
. Male posterior body end. A, D–E: holotype (HM RAS 102/64 768-2); B–C: paratype (HM RAS 102/64 768-4). Scale bars: A = 30 µm; B = 70 µm; C = 60 µm; D = 20 µm; E = 50 µm.
</paragraph>
</caption>
<caption id="DF54666CFFB7AF4465AFF8F469AA54F9" ID-DOI="http://doi.org/10.5281/zenodo.4298041" ID-Zenodo-Dep="4298041" httpUri="https://zenodo.org/record/4298041/files/figure.png" pageId="8" pageNumber="167" startId="8.[189,232,1858,1884]" targetBox="[258,1330,266,1820]" targetPageId="8">
<paragraph id="8B9436E4FFB7AF4465AFF8F469AA54F9" blockId="8.[187,1399,1858,2026]" pageId="8" pageNumber="167">
<emphasis id="B95FEAF6FFB7AF4465AFF8F46B03544F" bold="true" box="[189,267,1858,1884]" pageId="8" pageNumber="167">Fig. 4.</emphasis>
<taxonomicName id="4C2B4D67FFB7AF446400F8F46829544F" authority="Naumova &amp; Gagarin, 2020" authorityName="Naumova &amp; Gagarin" authorityYear="2020" box="[274,545,1858,1884]" class="Enoplea" family="Tobrilidae" genus="Paratrilobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Triplonchida" pageId="8" pageNumber="167" phylum="Nematoda" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B95FEAF6FFB7AF446400F8F46829544F" box="[274,545,1858,1884]" italics="true" pageId="8" pageNumber="167">Paratrilobus aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C578DFFB7AF44673BF8F5688E544E" box="[553,646,1859,1885]" pageId="8" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
, light micrographs.
<emphasis id="B95FEAF6FFB7AF446663F8F4698E544F" bold="true" box="[881,902,1858,1884]" pageId="8" pageNumber="167">A</emphasis>
. Entire male.
<emphasis id="B95FEAF6FFB7AF44613EF8F56E37544E" bold="true" box="[1068,1087,1859,1885]" pageId="8" pageNumber="167">B</emphasis>
. Entire female.
<emphasis id="B95FEAF6FFB7AF4461EEF8F46F19544F" bold="true" box="[1276,1297,1858,1884]" pageId="8" pageNumber="167">C</emphasis>
. Female pharyngeal region.
<emphasis id="B95FEAF6FFB7AF44648BF8D06BA65493" bold="true" box="[409,430,1894,1920]" pageId="8" pageNumber="167">D</emphasis>
. Male anterior end.
<emphasis id="B95FEAF6FFB7AF446781F8D068AF5493" bold="true" box="[659,679,1894,1920]" pageId="8" pageNumber="167">E</emphasis>
. Female anterior end.
<emphasis id="B95FEAF6FFB7AF4466B7F8D069BF5493" bold="true" box="[933,951,1894,1920]" pageId="8" pageNumber="167">F</emphasis>
. Female vulva region.
<emphasis id="B95FEAF6FFB7AF4461AFF8D36EDC546C" bold="true" box="[1213,1236,1893,1919]" pageId="8" pageNumber="167">G</emphasis>
. Male cloacal region.
<emphasis id="B95FEAF6FFB7AF446408F83F6B3954B0" bold="true" box="[282,305,1929,1955]" pageId="8" pageNumber="167">H</emphasis>
. Male posterior body end.
<emphasis id="B95FEAF6FFB7AF44676CF83F688154B0" bold="true" box="[638,649,1929,1955]" pageId="8" pageNumber="167">I</emphasis>
. Female posterior body end. A, D, G–H: holotype (HM RAS 102/64 768-2); B–C, I: paratype (HM RAS 102/64 768-4); E–F: paratype (LIN-SB 765-3). Scale bars: A–B = 200 µm; C, H = 50 µm; D–E = 10 µm; F–G, I = 20 µm.
</paragraph>
</caption>
<caption id="DF54666CFFB6AF4565ACFEBF6F3A5230" ID-Table-UUID="DF54666CFFB6AF4565ACFEBF6F3A5230" box="[190,1330,264,291]" httpUri="http://table.plazi.org/id/DF54666CFFB6AF4565ACFEBF6F3A5230" pageId="9" pageNumber="168" targetBox="[305,1261,337,1316]" targetIsTable="true" targetPageId="9">
<paragraph id="8B9436E4FFB6AF4565ACFEBF6F3A5230" blockId="9.[190,1330,264,291]" box="[190,1330,264,291]" pageId="9" pageNumber="168">
<emphasis id="B95FEAF6FFB6AF4565ACFEBF6B295231" bold="true" box="[190,289,264,291]" pageId="9" pageNumber="168">Table 2.</emphasis>
Measurements (in µm) of
<taxonomicName id="4C2B4D67FFB6AF456748FEBE69615231" authority="Naumova &amp; Gagarin, 2020" authorityName="Naumova &amp; Gagarin" authorityYear="2020" box="[602,873,264,290]" class="Enoplea" family="Tobrilidae" genus="Paratrilobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Triplonchida" pageId="9" pageNumber="168" phylum="Nematoda" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B95FEAF6FFB6AF456748FEBE69615231" box="[602,873,264,290]" italics="true" pageId="9" pageNumber="168">Paratrilobus aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C578DFFB6AF456662FEBF69C55230" box="[880,973,265,291]" pageId="9" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
, presented as mean and range.
</paragraph>
</caption>
<paragraph id="8B9436E4FFB6AF456423FEE76E805637" pageId="9" pageNumber="168">
<table id="F92BC444FFB650B36423FEE76EE55637" box="[305,1261,337,1316]" gridcols="3" gridrows="22" pageId="9" pageNumber="168">
<tr id="351B34A6FFB650B36423FEE76EE5527D" box="[305,1261,337,366]" gridrow="0" pageId="9" pageNumber="168">
<th id="76CA5DDAFFB650B36423FEE768AD527D" box="[305,677,337,366]" gridcol="0" gridrow="0" pageId="9" pageNumber="168">
<emphasis id="B95FEAF6FFB6AF456423FEE76BBA5278" bold="true" box="[305,434,337,363]" pageId="9" pageNumber="168">Character</emphasis>
</th>
<th id="76CA5DDAFFB650B36639FEE769B3527D" box="[811,955,337,366]" gridcol="1" gridrow="0" pageId="9" pageNumber="168">
<emphasis id="B95FEAF6FFB6AF456639FEE46995527F" bold="true" box="[811,925,338,364]" pageId="9" pageNumber="168">Holotype</emphasis>
♂
</th>
<th id="76CA5DDAFFB650B36106FEE76EE5527D" box="[1044,1261,337,366]" gridcol="2" gridrow="0" pageId="9" pageNumber="168">
<emphasis id="B95FEAF6FFB6AF45613AFEE46EAD527F" bold="true" box="[1064,1189,338,364]" pageId="9" pageNumber="168">Paratypes</emphasis>
♀♀
</th>
</tr>
<tr id="351B34A6FFB650B36423FEC96EE5528A" box="[305,1261,383,409]" gridrow="1" pageId="9" pageNumber="168">
<th id="76CA5DDAFFB650B36423FEC968AD528A" box="[305,677,383,409]" gridcol="0" gridrow="1" pageId="9" pageNumber="168">Number of specimens</th>
<td id="76CA5DDAFFB650B36639FEC969B3528A" box="[811,955,383,409]" gridcol="1" gridrow="1" pageId="9" pageNumber="168">1</td>
<td id="76CA5DDAFFB650B36106FEC96EE5528A" box="[1044,1261,383,409]" gridcol="2" gridrow="1" pageId="9" pageNumber="168">4</td>
</tr>
<tr id="351B34A6FFB650B36423FE1A6EE552D4" box="[305,1261,428,455]" gridrow="2" pageId="9" pageNumber="168">
<th id="76CA5DDAFFB650B36423FE1A68AD52D4" box="[305,677,428,455]" gridcol="0" gridrow="2" pageId="9" pageNumber="168">Body length (L)</th>
<td id="76CA5DDAFFB650B36639FE1A69B352D4" box="[811,955,428,455]" gridcol="1" gridrow="2" pageId="9" pageNumber="168">2097</td>
<td id="76CA5DDAFFB650B36106FE1A6EE552D4" box="[1044,1261,428,455]" gridcol="2" gridrow="2" pageId="9" pageNumber="168">2079 (2015–2203)</td>
</tr>
<tr id="351B34A6FFB650B36423FE6F6EE552E7" box="[305,1261,473,500]" gridrow="3" pageId="9" pageNumber="168">
<th id="76CA5DDAFFB650B36423FE6F68AD52E7" box="[305,677,473,500]" gridcol="0" gridrow="3" pageId="9" pageNumber="168">
<emphasis id="B95FEAF6FFB6AF456423FE6C6B4852E0" box="[305,320,474,499]" italics="true" pageId="9" pageNumber="168">a</emphasis>
</th>
<td id="76CA5DDAFFB650B36639FE6F69B352E7" box="[811,955,473,500]" gridcol="1" gridrow="3" pageId="9" pageNumber="168">28</td>
<td id="76CA5DDAFFB650B36106FE6F6EE552E7" box="[1044,1261,473,500]" gridcol="2" gridrow="3" pageId="9" pageNumber="168">29 (26–30)</td>
</tr>
<tr id="351B34A6FFB650B36423FDB16EE55132" box="[305,1261,519,545]" gridrow="4" pageId="9" pageNumber="168">
<th id="76CA5DDAFFB650B36423FDB168AD5132" box="[305,677,519,545]" gridcol="0" gridrow="4" pageId="9" pageNumber="168">
<emphasis id="B95FEAF6FFB6AF456423FDB16B485132" box="[305,320,519,545]" italics="true" pageId="9" pageNumber="168">b</emphasis>
</th>
<td id="76CA5DDAFFB650B36639FDB169B35132" box="[811,955,519,545]" gridcol="1" gridrow="4" pageId="9" pageNumber="168">3.9</td>
<td id="76CA5DDAFFB650B36106FDB16EE55132" box="[1044,1261,519,545]" gridcol="2" gridrow="4" pageId="9" pageNumber="168">3.4 (3.3–3.7)</td>
</tr>
<tr id="351B34A6FFB650B36423FD826EE5515C" box="[305,1261,564,591]" gridrow="5" pageId="9" pageNumber="168">
<th id="76CA5DDAFFB650B36423FD8268AD515C" box="[305,677,564,591]" gridcol="0" gridrow="5" pageId="9" pageNumber="168">
<emphasis id="B95FEAF6FFB6AF456423FD836B36515D" box="[305,318,565,590]" italics="true" pageId="9" pageNumber="168">c</emphasis>
</th>
<td id="76CA5DDAFFB650B36639FD8269B3515C" box="[811,955,564,591]" gridcol="1" gridrow="5" pageId="9" pageNumber="168">8.9</td>
<td id="76CA5DDAFFB650B36106FD826EE5515C" box="[1044,1261,564,591]" gridcol="2" gridrow="5" pageId="9" pageNumber="168">8.3 (7.5–8.8)</td>
</tr>
<tr id="351B34A6FFB650B36423FDD66EE5516F" box="[305,1261,608,636]" gridrow="6" pageId="9" pageNumber="168">
<th id="76CA5DDAFFB650B36423FDD668AD516F" box="[305,677,608,636]" gridcol="0" gridrow="6" pageId="9" pageNumber="168">
<emphasis id="B95FEAF6FFB6AF456423FDD66B4C516F" box="[305,324,608,636]" italics="true" pageId="9" pageNumber="168">cʹ</emphasis>
</th>
<td id="76CA5DDAFFB650B36639FDD669B3516F" box="[811,955,608,636]" gridcol="1" gridrow="6" pageId="9" pageNumber="168">5.3</td>
<td id="76CA5DDAFFB650B36106FDD66EE5516F" box="[1044,1261,608,636]" gridcol="2" gridrow="6" pageId="9" pageNumber="168">6.3 (6.0–6.8)</td>
</tr>
<tr id="351B34A6FFB650B36423FD396EE551B9" box="[305,1261,655,682]" gridrow="7" pageId="9" pageNumber="168">
<th id="76CA5DDAFFB650B36423FD3968AD51B9" box="[305,677,655,682]" gridcol="0" gridrow="7" pageId="9" pageNumber="168">V,%</th>
<td id="76CA5DDAFFB650B36639FD3969B351B9" box="[811,955,655,682]" gridcol="1" gridrow="7" pageId="9" pageNumber="168">–</td>
<td id="76CA5DDAFFB650B36106FD396EE551B9" box="[1044,1261,655,682]" gridcol="2" gridrow="7" pageId="9" pageNumber="168">52.4 (51.8–53.0)</td>
</tr>
<tr id="351B34A6FFB650B36423FD0A6EE551C4" box="[305,1261,700,727]" gridrow="8" pageId="9" pageNumber="168">
<th id="76CA5DDAFFB650B36423FD0A68AD51C4" box="[305,677,700,727]" gridcol="0" gridrow="8" pageId="9" pageNumber="168">Labial region diameter</th>
<td id="76CA5DDAFFB650B36639FD0A69B351C4" box="[811,955,700,727]" gridcol="1" gridrow="8" pageId="9" pageNumber="168">34</td>
<td id="76CA5DDAFFB650B36106FD0A6EE551C4" box="[1044,1261,700,727]" gridcol="2" gridrow="8" pageId="9" pageNumber="168">33 (31–35)</td>
</tr>
<tr id="351B34A6FFB650B36423FD5F6EE55017" box="[305,1261,745,772]" gridrow="9" pageId="9" pageNumber="168">
<th id="76CA5DDAFFB650B36423FD5F68AD5017" box="[305,677,745,772]" gridcol="0" gridrow="9" pageId="9" pageNumber="168">Body diameter</th>
<td id="76CA5DDAFFB650B36639FD5F69B35017" box="[811,955,745,772]" gridcol="1" gridrow="9" pageId="9" pageNumber="168">75</td>
<td id="76CA5DDAFFB650B36106FD5F6EE55017" box="[1044,1261,745,772]" gridcol="2" gridrow="9" pageId="9" pageNumber="168">72 (68–80)</td>
</tr>
<tr id="351B34A6FFB650B36423FCA16EE55021" box="[305,1261,791,818]" gridrow="10" pageId="9" pageNumber="168">
<th id="76CA5DDAFFB650B36423FCA168AD5021" box="[305,677,791,818]" gridcol="0" gridrow="10" pageId="9" pageNumber="168">Anal or cloacal diameter</th>
<td id="76CA5DDAFFB650B36639FCA169B35021" box="[811,955,791,818]" gridcol="1" gridrow="10" pageId="9" pageNumber="168">43</td>
<td id="76CA5DDAFFB650B36106FCA16EE55021" box="[1044,1261,791,818]" gridcol="2" gridrow="10" pageId="9" pageNumber="168">41 (39–43)</td>
</tr>
<tr id="351B34A6FFB650B36423FCF26EE5504C" box="[305,1261,836,863]" gridrow="11" pageId="9" pageNumber="168">
<th id="76CA5DDAFFB650B36423FCF268AD504C" box="[305,677,836,863]" gridcol="0" gridrow="11" pageId="9" pageNumber="168">Stoma length</th>
<td id="76CA5DDAFFB650B36639FCF269B3504C" box="[811,955,836,863]" gridcol="1" gridrow="11" pageId="9" pageNumber="168">50</td>
<td id="76CA5DDAFFB650B36106FCF26EE5504C" box="[1044,1261,836,863]" gridcol="2" gridrow="11" pageId="9" pageNumber="168">50 (48–52)</td>
</tr>
<tr id="351B34A6FFB650B36423FCC46EE5509F" box="[305,1261,882,908]" gridrow="12" pageId="9" pageNumber="168">
<th id="76CA5DDAFFB650B36423FCC468AD509F" box="[305,677,882,908]" gridcol="0" gridrow="12" pageId="9" pageNumber="168">Outer labial setae length</th>
<td id="76CA5DDAFFB650B36639FCC469B3509F" box="[811,955,882,908]" gridcol="1" gridrow="12" pageId="9" pageNumber="168">18</td>
<td id="76CA5DDAFFB650B36106FCC46EE5509F" box="[1044,1261,882,908]" gridcol="2" gridrow="12" pageId="9" pageNumber="168">19 (18–20)</td>
</tr>
<tr id="351B34A6FFB650B36423FC296EE550A9" box="[305,1261,927,954]" gridrow="13" pageId="9" pageNumber="168">
<th id="76CA5DDAFFB650B36423FC2968AD50A9" box="[305,677,927,954]" gridcol="0" gridrow="13" pageId="9" pageNumber="168">Cephalic setae length</th>
<td id="76CA5DDAFFB650B36639FC2969B350A9" box="[811,955,927,954]" gridcol="1" gridrow="13" pageId="9" pageNumber="168">11</td>
<td id="76CA5DDAFFB650B36106FC296EE550A9" box="[1044,1261,927,954]" gridcol="2" gridrow="13" pageId="9" pageNumber="168">12 (10–13)</td>
</tr>
<tr id="351B34A6FFB650B36423FC7A6EE550F4" box="[305,1261,972,999]" gridrow="14" pageId="9" pageNumber="168">
<th id="76CA5DDAFFB650B36423FC7A68AD50F4" box="[305,677,972,999]" gridcol="0" gridrow="14" pageId="9" pageNumber="168">Pharynx length</th>
<td id="76CA5DDAFFB650B36639FC7A69B350F4" box="[811,955,972,999]" gridcol="1" gridrow="14" pageId="9" pageNumber="168">536</td>
<td id="76CA5DDAFFB650B36106FC7A6EE550F4" box="[1044,1261,972,999]" gridcol="2" gridrow="14" pageId="9" pageNumber="168">609 (589–630)</td>
</tr>
<tr id="351B34A6FFB650B36423FC4C6EE55706" box="[305,1261,1018,1045]" gridrow="15" pageId="9" pageNumber="168">
<th id="76CA5DDAFFB650B36423FC4C68AD5706" box="[305,677,1018,1045]" gridcol="0" gridrow="15" pageId="9" pageNumber="168">Posterior pharynx end to vulva</th>
<td id="76CA5DDAFFB650B36639FC4C69B35706" box="[811,955,1018,1045]" gridcol="1" gridrow="15" pageId="9" pageNumber="168">–</td>
<td id="76CA5DDAFFB650B36106FC4C6EE55706" box="[1044,1261,1018,1045]" gridcol="2" gridrow="15" pageId="9" pageNumber="168">479 (434–578)</td>
</tr>
<tr id="351B34A6FFB650B36423FB916EE55751" box="[305,1261,1063,1090]" gridrow="16" pageId="9" pageNumber="168">
<th id="76CA5DDAFFB650B36423FB9168AD5751" box="[305,677,1063,1090]" gridcol="0" gridrow="16" pageId="9" pageNumber="168">Posterior pharynx end to cloaca</th>
<td id="76CA5DDAFFB650B36639FB9169B35751" box="[811,955,1063,1090]" gridcol="1" gridrow="16" pageId="9" pageNumber="168">1326</td>
<td id="76CA5DDAFFB650B36106FB916EE55751" box="[1044,1261,1063,1090]" gridcol="2" gridrow="16" pageId="9" pageNumber="168">–</td>
</tr>
<tr id="351B34A6FFB650B36423FBE26EE5577C" box="[305,1261,1108,1135]" gridrow="17" pageId="9" pageNumber="168">
<th id="76CA5DDAFFB650B36423FBE268AD577C" box="[305,677,1108,1135]" gridcol="0" gridrow="17" pageId="9" pageNumber="168">Vulva to anus</th>
<td id="76CA5DDAFFB650B36639FBE269B3577C" box="[811,955,1108,1135]" gridcol="1" gridrow="17" pageId="9" pageNumber="168">–</td>
<td id="76CA5DDAFFB650B36106FBE26EE5577C" box="[1044,1261,1108,1135]" gridcol="2" gridrow="17" pageId="9" pageNumber="168">738 (697–787)</td>
</tr>
<tr id="351B34A6FFB650B36423FB346EE5578E" box="[305,1261,1154,1181]" gridrow="18" pageId="9" pageNumber="168">
<th id="76CA5DDAFFB650B36423FB3468AD578E" box="[305,677,1154,1181]" gridcol="0" gridrow="18" pageId="9" pageNumber="168">Tail length</th>
<td id="76CA5DDAFFB650B36639FB3469B3578E" box="[811,955,1154,1181]" gridcol="1" gridrow="18" pageId="9" pageNumber="168">235</td>
<td id="76CA5DDAFFB650B36106FB346EE5578E" box="[1044,1261,1154,1181]" gridcol="2" gridrow="18" pageId="9" pageNumber="168">253 (240–267)</td>
</tr>
<tr id="351B34A6FFB650B36423FB196EE557D9" box="[305,1261,1199,1226]" gridrow="19" pageId="9" pageNumber="168">
<th id="76CA5DDAFFB650B36423FB1968AD57D9" box="[305,677,1199,1226]" gridcol="0" gridrow="19" pageId="9" pageNumber="168">Spicula length</th>
<td id="76CA5DDAFFB650B36639FB1969B357D9" box="[811,955,1199,1226]" gridcol="1" gridrow="19" pageId="9" pageNumber="168">51</td>
<td id="76CA5DDAFFB650B36106FB196EE557D9" box="[1044,1261,1199,1226]" gridcol="2" gridrow="19" pageId="9" pageNumber="168">–</td>
</tr>
<tr id="351B34A6FFB650B36423FB6A6EE557E4" box="[305,1261,1244,1271]" gridrow="20" pageId="9" pageNumber="168">
<th id="76CA5DDAFFB650B36423FB6A68AD57E4" box="[305,677,1244,1271]" gridcol="0" gridrow="20" pageId="9" pageNumber="168">Gubernaculum length</th>
<td id="76CA5DDAFFB650B36639FB6A69B357E4" box="[811,955,1244,1271]" gridcol="1" gridrow="20" pageId="9" pageNumber="168">20</td>
<td id="76CA5DDAFFB650B36106FB6A6EE557E4" box="[1044,1261,1244,1271]" gridcol="2" gridrow="20" pageId="9" pageNumber="168">–</td>
</tr>
<tr id="351B34A6FFB650B36423FABF6EE55637" box="[305,1261,1289,1316]" gridrow="21" pageId="9" pageNumber="168">
<th id="76CA5DDAFFB650B36423FABF68AD5637" box="[305,677,1289,1316]" gridcol="0" gridrow="21" pageId="9" pageNumber="168">Number of supplements</th>
<td id="76CA5DDAFFB650B36639FABF69B35637" box="[811,955,1289,1316]" gridcol="1" gridrow="21" pageId="9" pageNumber="168">6</td>
<td id="76CA5DDAFFB650B36106FABF6EE55637" box="[1044,1261,1289,1316]" gridcol="2" gridrow="21" pageId="9" pageNumber="168">–</td>
</tr>
</table>
</paragraph>
<paragraph id="8B9436E4FFB6AF4565F8FAD76B6F5668" blockId="9.[189,1399,1377,1477]" box="[234,359,1377,1403]" pageId="9" pageNumber="168">
<emphasis id="B95FEAF6FFB6AF4565F8FAD76B6F5668" bold="true" box="[234,359,1377,1403]" pageId="9" pageNumber="168">
<typeStatus id="54908846FFB6AF4565F8FAD76B6F5668" box="[234,359,1377,1403]" pageId="9" pageNumber="168" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B9436E4FFB6AF4565AFFA306F7F56D6" blockId="9.[189,1399,1377,1477]" pageId="9" pageNumber="168">
<materialsCitation id="3B433CB9FFB6AF4565AFFA306B0456D6" ID-GBIF-Occurrence="2985876301" collectedFrom="sand and silt" collectingDate="2012-07-05" collectionCode="LIN-SB" collectorName="T. V. Naumova" country="Russia" county="Lake Baikal" elevation="51" latitude="52.07994" location="RUSSIA" longLatPrecision="1" longitude="105.90368" municipality="Posolsk Bank" pageId="9" pageNumber="168" specimenCode="765-3, 765-5, 767-5" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<collectingCountry id="F33C7674FFB6AF4565AFFA306B2C56B3" box="[189,292,1414,1440]" name="Russia" pageId="9" pageNumber="168">RUSSIA</collectingCountry>
•
<specimenCount id="9D2DFD6DFFB6AF45642CFA306B8956B3" box="[318,385,1414,1440]" pageId="9" pageNumber="168" type="female">3 ♀♀</specimenCount>
; same collection data as for the holotype;
<collectorName id="26DE5332FFB6AF456667FA306E1756B3" box="[885,1055,1414,1440]" pageId="9" pageNumber="168">T.V. Naumova</collectorName>
leg.;
<collectionCode id="ED3AAE21FFB6AF456170FA306ECD56B3" box="[1122,1221,1414,1440]" country="Russia" httpUri="http://grbio.org/cool/s5tp-ib34" name="Limnological Institute, Siberian Branch, Russian Academy of Sciences" pageId="9" pageNumber="168">LIN–SB</collectionCode>
(
<specimenCode id="DB8D9E9FFFB6AF4561C4FA306F1656B2" box="[1238,1310,1414,1441]" pageId="9" pageNumber="168">765-3</specimenCode>
,
<specimenCode id="DB8D9E9FFFB6AF456039FA306F7A56B2" box="[1323,1394,1414,1441]" pageId="9" pageNumber="168">765-5</specimenCode>
,
<specimenCode id="DB8D9E9FFFB6AF4565AFFA1C6B0B56D6" box="[189,259,1450,1477]" pageId="9" pageNumber="168">767-5</specimenCode>
)
</materialsCitation>
•
<materialsCitation id="3B433CB9FFB6AF456433FA1D6F7B56D6" ID-GBIF-Occurrence="2985876304" box="[289,1395,1450,1477]" collectedFrom="sand and silt" collectingDate="2012-07-05" collectionCode="HM RAS" collectorName="T. V. Naumova" country="Russia" county="Lake Baikal" elevation="51" latitude="52.07994" location="Lake Baikal" longLatPrecision="1" longitude="105.90368" municipality="Posolsk Bank" pageId="9" pageNumber="168" specimenCode="768-4" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D2DFD6DFFB6AF456433FA1D6B4456D6" box="[289,332,1451,1477]" pageId="9" pageNumber="168" type="female">1 ♀</specimenCount>
same collection data as for the holotype;
<collectorName id="26DE5332FFB6AF45663AFA1D69D856D6" box="[808,976,1451,1477]" pageId="9" pageNumber="168">T.V. Naumova</collectorName>
leg.;
<collectionCode id="ED3AAE21FFB6AF45611DFA1D6E7556D6" box="[1039,1149,1451,1477]" pageId="9" pageNumber="168">HM RAS</collectionCode>
slide 102/64 (
<specimenCode id="DB8D9E9FFFB6AF456032FA1C6F6256D6" box="[1312,1386,1450,1477]" pageId="9" pageNumber="168">768-4</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C331656FFFB6AF4665AFFA5B694D5021" lastPageId="10" lastPageNumber="169" pageId="9" pageNumber="168" type="description">
<paragraph id="8B9436E4FFB6AF4565AFFA5B6B465514" blockId="9.[189,334,1517,1543]" box="[189,334,1517,1543]" pageId="9" pageNumber="168">
<heading id="D0DC8188FFB6AF4565AFFA5B6B465514" bold="true" box="[189,334,1517,1543]" fontSize="11" level="3" pageId="9" pageNumber="168" reason="3">
<emphasis id="B95FEAF6FFB6AF4565AFFA5B6B465514" bold="true" box="[189,334,1517,1543]" pageId="9" pageNumber="168">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B9436E4FFB6AF4565F8F9AB6B225524" blockId="9.[189,1400,1565,1921]" box="[234,298,1565,1591]" pageId="9" pageNumber="168">
<emphasis id="B95FEAF6FFB6AF4565F8F9AB6B225524" bold="true" box="[234,298,1565,1591]" pageId="9" pageNumber="168">Male</emphasis>
</paragraph>
<paragraph id="8B9436E4FFB6AF4565AFF9F468055492" blockId="9.[189,1400,1565,1921]" pageId="9" pageNumber="168">
Body comparatively short. Cuticle finely annulated,
<quantity id="4CD39B01FFB6AF456620F9F46981554F" box="[818,905,1602,1629]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="168" unit="mm" value="1.5">1.5 µm</quantity>
thick. Crystalloids absent. Somatic setae sparse, short. Labial region comparatively high, slightly offset from adjacent body, lips well developed. Six inner labial sensillae papilliform. Six outer labial sensillae in shape of smooth setae long 53% of labial region width. Four cephalic sensillae in shape of thin setae. Cheilostom of average size. Buccal cavity spacious, barrel-shaped, with thick walls. One pocket connected with buccal cavity by wide gleam, containing two small teeth. Stoma 1.5 times as long as labial region width. Amphidial fovea cupshaped, opening at level of buccal cavity. Pharynx muscular, comparatively long, expanding gradually along length. Cardiac glands large, rounded,
<quantity id="4CD39B01FFB6AF4567F8F8F46933544E" box="[746,827,1858,1885]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="168" unit="mm" value="30.0">30 µm</quantity>
in diameter. Ventral gland, its canal, ampulla, excretory pore not observed.
</paragraph>
<paragraph id="8B9436E4FFB6AF4665AFF81C6B6052C6" blockId="9.[189,1399,1962,2025]" lastBlockId="10.[187,1399,265,469]" lastPageId="10" lastPageNumber="169" pageId="9" pageNumber="168">
Testes paired, situated to left of intestine; anterior testis outstretched, posterior testis short, reflexed.
<collectingRegion id="49EFF806FFB6AF45605EF81C6F7E54D7" box="[1356,1398,1962,1988]" country="Hungary" name="Vas" pageId="9" pageNumber="168">Vas</collectingRegion>
deferens well developed. Spicules slender, ventral curved, 1.2 times as long as cloacal body diameter. Gubernaculum in shape of gutter, 39% of spicule length. Precloacal supplement
<quantity id="4CD39B01FFB5AF46617BFEBF6E9E5230" box="[1129,1174,265,291]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="10" pageNumber="169" unit="in" value="6.0">6 in</quantity>
number, about one size, located approximately at equal distances from each other. Supplements echinate, not very protruded over body surface. Ampulla contents concentrated in ampullae base. Shoulder absent. Cap armed, numerous small thorns, one large central thorn. Precloacal supplement located anteriorly to spicules level. Tail slender, comparatively long. Three caudal glands, spinneret well developed. Subterminal setae not seen.
</paragraph>
<paragraph id="8B9436E4FFB5AF4665F8FE4A6B4D5105" blockId="10.[187,1401,508,818]" box="[234,325,508,534]" pageId="10" pageNumber="169">
<emphasis id="B95FEAF6FFB5AF4665F8FE4A6B4D5105" bold="true" box="[234,325,508,534]" pageId="10" pageNumber="169">Female</emphasis>
</paragraph>
<paragraph id="8B9436E4FFB5AF4665AFFDA9694D5021" blockId="10.[187,1401,508,818]" pageId="10" pageNumber="169">General morphology similar to that of males in structure of cuticle, anterior body end. Six outer labial sensillae in shape of smooth non-articulated setae long 53–58% of labial region width. Cardia surrounded by three round glands. Rectum length equal to or slightly greater than anal body diameter. Reproductive system didelphic, amphidelphic. Ovaries situated to left of intestine, reflexed. Oocytes numerous. Vulva transverse slit, situated slightly posterior to mid-body. Vulval lips not sclerotized, not protruded outside body contour. Cuticular wrinkles around vulva, vulva glands not observed. Vagina straight line, with well-expressed spherical vaginal chamber, thick walls. Generated eggs in uterus not observed. Tail slender, comparatively long. Subterminal seta not seen.</paragraph>
</subSubSection>
<subSubSection id="C331656FFFB5AF4665AFFCEF6B94563B" pageId="10" pageNumber="169" type="discussion">
<paragraph id="8B9436E4FFB5AF4665AFFCEF6B275060" blockId="10.[189,303,857,883]" box="[189,303,857,883]" pageId="10" pageNumber="169">
<heading id="D0DC8188FFB5AF4665AFFCEF6B275060" bold="true" box="[189,303,857,883]" fontSize="11" level="3" pageId="10" pageNumber="169" reason="3">
<emphasis id="B95FEAF6FFB5AF4665AFFCEF6B275060" bold="true" box="[189,303,857,883]" pageId="10" pageNumber="169">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B9436E4FFB5AF4665AFFC3E6B94563B" blockId="10.[188,1400,904,1320]" pageId="10" pageNumber="169">
<taxonomicName id="4C2B4D67FFB5AF4665AFFC3E6BC650B1" authority="Naumova &amp; Gagarin, 2020" authorityName="Naumova &amp; Gagarin" authorityYear="2020" box="[189,462,904,930]" class="Enoplea" family="Tobrilidae" genus="Paratrilobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Triplonchida" pageId="10" pageNumber="169" phylum="Nematoda" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B95FEAF6FFB5AF4665AFFC3E6BC650B1" box="[189,462,904,930]" italics="true" pageId="10" pageNumber="169">Paratrilobus aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C578DFFB5AF4664CAFC3E683C50B1" box="[472,564,904,930]" pageId="10" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
is similar to
<taxonomicName id="4C2B4D67FFB5AF4667C5FC3F6ECD50B1" authority="Gagarin &amp; Naumova, 2011" authorityName="Gagarin &amp; Naumova" authorityYear="2011" box="[727,1221,904,930]" class="Enoplea" family="Tobrilidae" genus="Paratrilobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Triplonchida" pageId="10" pageNumber="169" phylum="Nematoda" rank="species" species="granulosus">
<emphasis id="B95FEAF6FFB5AF4667C5FC3F697050B1" box="[727,888,904,930]" italics="true" pageId="10" pageNumber="169">P. granulosus</emphasis>
<bibRefCitation id="EFBA4B15FFB5AF466692FC3E6ECD50B1" author="Gagarin V. G. &amp; Naumova T. V." box="[896,1221,904,930]" pageId="10" pageNumber="169" pagination="85 - 95" refId="ref5344" refString="Gagarin V. G. &amp; Naumova T. V. 2011. Paratrilobus granulosus sp. n., Tobrilus longisetosus sp. n. and Domorganus acutus (Tsalolikhin, 1977) from Lake Baikal, Russia. International Journal of Nematology 21: 85 - 95." type="journal article" year="2011">Gagarin &amp; Naumova, 2011</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C2B4D67FFB5AF466011FC3F6B9450D5" authority="(Tsalolichin, 1977)" baseAuthorityName="Tsalolichin" baseAuthorityYear="1977" class="Enoplea" family="Tobrilidae" genus="Paratrilobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Triplonchida" pageId="10" pageNumber="169" phylum="Nematoda" rank="species" species="ultimus">
<emphasis id="B95FEAF6FFB5AF466011FC3F6F7050B1" box="[1283,1400,904,930]" italics="true" pageId="10" pageNumber="169">P. ultimus</emphasis>
(Tsalolichin, 1977)
</taxonomicName>
in the structure of the precloacal supplements. From the former species, it differs in the absence of crystalloids, a comparatively longer pharynx (
<emphasis id="B95FEAF6FFB5AF46666BFC79698050FA" box="[889,904,975,1001]" italics="true" pageId="10" pageNumber="169">b</emphasis>
= 3.3– 3.9 vs
<emphasis id="B95FEAF6FFB5AF466131FC796E3A50FA" box="[1059,1074,975,1001]" italics="true" pageId="10" pageNumber="169">b</emphasis>
=
<quantity id="4CD39B01FFB5AF46615DFC796ECB50FA" box="[1103,1219,975,1002]" metricMagnitude="-1" metricUnit="m" metricValue="1.3208" metricValueMax="1.4986" metricValueMin="1.143" pageId="10" pageNumber="169" unit="in" value="5.2" valueMax="5.9" valueMin="4.5">4.5–5.9 in</quantity>
<taxonomicName id="4C2B4D67FFB5AF4661D4FC666F6B50FA" authorityName="Gagarin &amp; Naumova" authorityYear="2011" box="[1222,1379,975,1001]" class="Enoplea" family="Tobrilidae" genus="Paratrilobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Triplonchida" pageId="10" pageNumber="169" phylum="Nematoda" rank="species" species="granulosus">
<emphasis id="B95FEAF6FFB5AF4661D4FC666F6B50FA" box="[1222,1379,975,1001]" italics="true" pageId="10" pageNumber="169">P. granulosus</emphasis>
</taxonomicName>
), longer outer labial setae (18–20 μm long, 53–58% of labial region width vs 6.5–8.5 μm long, 30– 35% of labial region width in
<taxonomicName id="4C2B4D67FFB5AF46673EFBA168C55723" authorityName="Gagarin &amp; Naumova" authorityYear="2011" box="[556,717,1046,1072]" class="Enoplea" family="Tobrilidae" genus="Paratrilobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Triplonchida" pageId="10" pageNumber="169" phylum="Nematoda" rank="species" species="granulosus">
<emphasis id="B95FEAF6FFB5AF46673EFBA168C55723" box="[556,717,1046,1072]" italics="true" pageId="10" pageNumber="169">P. granulosus</emphasis>
</taxonomicName>
), the vulva position (V = 51.8–53.0% vs 39.1–46.1% in
<taxonomicName id="4C2B4D67FFB5AF4665AFFB8C6B525740" authorityName="Gagarin &amp; Naumova" authorityYear="2011" box="[189,346,1081,1107]" class="Enoplea" family="Tobrilidae" genus="Paratrilobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Triplonchida" pageId="10" pageNumber="169" phylum="Nematoda" rank="species" species="granulosus">
<emphasis id="B95FEAF6FFB5AF4665AFFB8C6B525740" box="[189,346,1081,1107]" italics="true" pageId="10" pageNumber="169">P. granulosus</emphasis>
</taxonomicName>
), the longer stoma (
<quantity id="4CD39B01FFB5AF466752FB8F68B15747" box="[576,697,1081,1108]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="5.2" metricValueMin="4.8" pageId="10" pageNumber="169" unit="mm" value="50.0" valueMax="52.0" valueMin="48.0">48–52 µm</quantity>
long vs
<quantity id="4CD39B01FFB5AF46660BFB8F69985747" box="[793,912,1081,1108]" metricMagnitude="-2" metricUnit="m" metricValue="3.05" metricValueMax="3.3" metricValueMin="2.8" pageId="10" pageNumber="169" unit="mm" value="30.5" valueMax="33.0" valueMin="28.0">28–33 µm</quantity>
long in
<taxonomicName id="4C2B4D67FFB5AF4666F9FB8C6E805740" authorityName="Gagarin &amp; Naumova" authorityYear="2011" box="[1003,1160,1081,1107]" class="Enoplea" family="Tobrilidae" genus="Paratrilobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Triplonchida" pageId="10" pageNumber="169" phylum="Nematoda" rank="species" species="granulosus">
<emphasis id="B95FEAF6FFB5AF4666F9FB8C6E805740" box="[1003,1160,1081,1107]" italics="true" pageId="10" pageNumber="169">P. granulosus</emphasis>
</taxonomicName>
), and the absence of a subterminal seta (
<bibRefCitation id="EFBA4B15FFB5AF4664B2FBEB68D65764" author="Gagarin V. G. &amp; Naumova T. V." box="[416,734,1117,1143]" pageId="10" pageNumber="169" pagination="85 - 95" refId="ref5344" refString="Gagarin V. G. &amp; Naumova T. V. 2011. Paratrilobus granulosus sp. n., Tobrilus longisetosus sp. n. and Domorganus acutus (Tsalolikhin, 1977) from Lake Baikal, Russia. International Journal of Nematology 21: 85 - 95." type="journal article" year="2011">Gagarin &amp; Naumova 2011</bibRefCitation>
). From the latter species, it differs in the longer pharynx (
<emphasis id="B95FEAF6FFB5AF4665D5FB366ADE5789" box="[199,214,1152,1178]" italics="true" pageId="10" pageNumber="169">b</emphasis>
= 3.3–3.9 vs
<emphasis id="B95FEAF6FFB5AF46646EFB366B835789" box="[380,395,1152,1178]" italics="true" pageId="10" pageNumber="169">b</emphasis>
=
<quantity id="4CD39B01FFB5AF4664B9FB36682B5789" box="[427,547,1152,1179]" metricMagnitude="-1" metricUnit="m" metricValue="1.3462" metricValueMax="1.4986" metricValueMin="1.1938" pageId="10" pageNumber="169" unit="in" value="5.3" valueMax="5.9" valueMin="4.7">4.7–5.9 in</quantity>
<taxonomicName id="4C2B4D67FFB5AF46673EFB3768A85789" baseAuthorityName="Tsalolichin" baseAuthorityYear="1977" box="[556,672,1152,1178]" class="Enoplea" family="Tobrilidae" genus="Paratrilobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Triplonchida" pageId="10" pageNumber="169" phylum="Nematoda" rank="species" species="ultimus">
<emphasis id="B95FEAF6FFB5AF46673EFB3768A85789" box="[556,672,1152,1178]" italics="true" pageId="10" pageNumber="169">P. ultimus</emphasis>
</taxonomicName>
), a longer and more slender tail (
<emphasis id="B95FEAF6FFB5AF466122FB376E355789" box="[1072,1085,1153,1178]" italics="true" pageId="10" pageNumber="169">c</emphasis>
= 7.5–8.9,
<emphasis id="B95FEAF6FFB5AF4661DAFBC86ED35789" box="[1224,1243,1150,1178]" italics="true" pageId="10" pageNumber="169">cʹ</emphasis>
= 5.5–6.8 vs
<emphasis id="B95FEAF6FFB5AF4665ACFB126AC357AE" box="[190,203,1188,1213]" italics="true" pageId="10" pageNumber="169">c</emphasis>
= 10.9–16.4, cʹ =
<quantity id="4CD39B01FFB5AF4664BBFB15682A57AE" box="[425,546,1187,1214]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.143" metricValueMin="0.889" pageId="10" pageNumber="169" unit="in" value="4.0" valueMax="4.5" valueMin="3.5">3.5–4.5 in</quantity>
<taxonomicName id="4C2B4D67FFB5AF466739FB12689757AE" baseAuthorityName="Tsalolichin" baseAuthorityYear="1977" box="[555,671,1187,1213]" class="Enoplea" family="Tobrilidae" genus="Paratrilobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Triplonchida" pageId="10" pageNumber="169" phylum="Nematoda" rank="species" species="ultimus">
<emphasis id="B95FEAF6FFB5AF466739FB12689757AE" box="[555,671,1187,1213]" italics="true" pageId="10" pageNumber="169">P. ultimus</emphasis>
</taxonomicName>
), the longer stoma (
<quantity id="4CD39B01FFB5AF466681FB156E0757AD" box="[915,1039,1187,1214]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="5.2" metricValueMin="4.8" pageId="10" pageNumber="169" unit="mm" value="50.0" valueMax="52.0" valueMin="48.0">48–52 µm</quantity>
long vs
<quantity id="4CD39B01FFB5AF46616BFB126ECF57AD" box="[1145,1223,1188,1214]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="10" pageNumber="169" unit="mm" value="21.0">21 µm</quantity>
in
<taxonomicName id="4C2B4D67FFB5AF4661E0FB126F6D57AE" baseAuthorityName="Tsalolichin" baseAuthorityYear="1977" box="[1266,1381,1187,1213]" class="Enoplea" family="Tobrilidae" genus="Paratrilobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Triplonchida" pageId="10" pageNumber="169" phylum="Nematoda" rank="species" species="ultimus">
<emphasis id="B95FEAF6FFB5AF4661E0FB126F6D57AE" box="[1266,1381,1187,1213]" italics="true" pageId="10" pageNumber="169">P. ultimus</emphasis>
</taxonomicName>
), longer outer labial setae (
<quantity id="4CD39B01FFB5AF4664F9FB71686F57F1" box="[491,615,1223,1250]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="10" pageNumber="169" unit="mm" value="19.0" valueMax="20.0" valueMin="18.0">18–20 µm</quantity>
long, 53–58% of labial region width
<emphasis id="B95FEAF6FFB5AF466138FB7E6E4A57F2" box="[1066,1090,1224,1249]" italics="true" pageId="10" pageNumber="169">vs</emphasis>
<quantity id="4CD39B01FFB5AF466158FB716EAF57F1" box="[1098,1191,1223,1250]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="10" pageNumber="169" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 µm</quantity>
long, 25–30% of labial region width in
<taxonomicName id="4C2B4D67FFB5AF4664AFFB5D68275617" baseAuthorityName="Tsalolichin" baseAuthorityYear="1977" box="[445,559,1258,1284]" class="Enoplea" family="Tobrilidae" genus="Paratrilobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Triplonchida" pageId="10" pageNumber="169" phylum="Nematoda" rank="species" species="ultimus">
<emphasis id="B95FEAF6FFB5AF4664AFFB5D68275617" box="[445,559,1258,1284]" italics="true" pageId="10" pageNumber="169">P. ultimus</emphasis>
</taxonomicName>
), and the vulva position (V = 51.8–53.0% vs V = 39–43% in
<taxonomicName id="4C2B4D67FFB5AF4661EBFB5D6F645617" baseAuthorityName="Tsalolichin" baseAuthorityYear="1977" box="[1273,1388,1258,1284]" class="Enoplea" family="Tobrilidae" genus="Paratrilobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Triplonchida" pageId="10" pageNumber="169" phylum="Nematoda" rank="species" species="ultimus">
<emphasis id="B95FEAF6FFB5AF4661EBFB5D6F645617" box="[1273,1388,1258,1284]" italics="true" pageId="10" pageNumber="169">P. ultimus</emphasis>
</taxonomicName>
) (Tsalolichin 1977).
</paragraph>
</subSubSection>
</treatment>
</document>
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