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<document id="D491B6A46A45A35B77B49C83BAE4EB1E" ID-CLB-Dataset="5429" ID-DOI="10.11646/zootaxa.4981.3.3" ID-GBIF-Dataset="bd4313a1-f17a-4745-ba2d-655124108edc" ID-ISSN="1175-5326" ID-Zenodo-Dep="4921137" ID-ZooBank="CB36C384-5538-494A-8802-4F821C7E25FE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623280791010" checkinUser="plazi" docAuthor="Chernyshev, Alexei V. &amp; Goedert, James L." docDate="2021" docId="038287AEC147174DFF7F593508CD0D0A" docLanguage="en" docName="zootaxa.4981.3.3.pdf" docOrigin="Zootaxa 4981 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ammonicera benhami Squires &amp; Goedert 1996" docType="treatment" docVersion="8" lastPageNumber="475" masterDocId="FFBBFFD6C142174BFFE85A1D090F0F6B" masterDocTitle="New species of latest Eocene / earliest Oligocene microgastropods (Heterobranchia Orbitestellidae and Omalogyridae) from the Gries Ranch Formation, Lewis County, Washington State, USA" masterLastPageNumber="480" masterPageNumber="469" pageNumber="474" updateTime="1699009098020" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="F43490FC439F96FD35426807F29B01CC">
<mods:title id="C248BB556BAE1C9C2FA058FE49BFDD6F">New species of latest Eocene / earliest Oligocene microgastropods (Heterobranchia Orbitestellidae and Omalogyridae) from the Gries Ranch Formation, Lewis County, Washington State, USA</mods:title>
</mods:titleInfo>
<mods:name id="3FDFF997DA739B0F8A1D62C767CEE2B5" type="personal">
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</mods:role>
<mods:namePart id="8E6ABB11A1B2FB9A71724F2C32A751C8">Chernyshev, Alexei V.</mods:namePart>
<mods:affiliation id="EA88C7D93F81E8FE957318A0DC602E07">A. V. Zhirmunsky National Scientific Center of Marine Biology, Far Eastern Branch, Russian Academy of Sciences, Palchevskogo Street 17, Vladivostok 690041, Russia.</mods:affiliation>
</mods:name>
<mods:name id="8285FFDA54FBDA61FC6E8DAA2868F7FC" type="personal">
<mods:role id="D810E6D0DAED4E9B8336E5370DBBB259">
<mods:roleTerm id="D41007A3ACEBADEFE971ACBB6ADB3041">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9AB1E9AABE71939718461C709502221F">Goedert, James L.</mods:namePart>
<mods:affiliation id="7BA53B8B59DD46FC58629284AA16C9A1">Burke Museum of Natural History and Culture, University of Washington, Seattle, WA 98195, USA.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="99B532F8646DAAA57903A6ABF81EAA0E">text</mods:typeOfResource>
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<mods:titleInfo id="F9B3D76FB12F50FE159AB6943055BB3D">
<mods:title id="DB384F07C5B547242FA3368783E92565">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1544C28B800B931857B5981881DF951B">
<mods:date id="C0AEA164CCCA400AD958384E0E284EA2">2021</mods:date>
<mods:detail id="CB6B90E0DF4439AE4D146B439B8C77E7" type="pubDate">
<mods:number id="E930A64B484E6248E63F6DE0FF0795DC">2021-06-08</mods:number>
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<mods:detail id="78B3D0FF57C6B7FB741E8BFD2B7CA8B1" type="volume">
<mods:number id="E9CCABE0374C61B4DD46688A8FB7B278">4981</mods:number>
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<mods:detail id="AFFF9C10DDB8744B0F6AA7AE55EEAFD6" type="issue">
<mods:number id="F44458D5942E009EB8F694EDB1498EAF">3</mods:number>
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<mods:start id="12CCD84D44F222A03EC5035C1EF4EEB0">469</mods:start>
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<mods:classification id="0FFC0FE8B4AB703211E23895DAF970D5">journal article</mods:classification>
<mods:identifier id="D2B097001946F0ADC3F49869D79D9BF1" type="CLB-Dataset">5429</mods:identifier>
<mods:identifier id="7FF4B0F7135743FD466DE7E6E3172FC9" type="DOI">10.11646/zootaxa.4981.3.3</mods:identifier>
<mods:identifier id="458C78BB15527E963244172E87715F20" type="GBIF-Dataset">bd4313a1-f17a-4745-ba2d-655124108edc</mods:identifier>
<mods:identifier id="9FB935CA790BDAFA7A24B7AF298E8FC2" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D5A872A0FFD36F0A685D3A12A8664E0E" type="Zenodo-Dep">4921137</mods:identifier>
<mods:identifier id="FFC41C2F6970004DB78F714D1FC0FFB8" type="ZooBank">CB36C384-5538-494A-8802-4F821C7E25FE</mods:identifier>
</mods:mods>
<treatment id="038287AEC147174DFF7F593508CD0D0A" ID-DOI="http://doi.org/10.5281/zenodo.5109636" ID-GBIF-Taxon="183548934" ID-Zenodo-Dep="5109636" LSID="urn:lsid:plazi:treatment:038287AEC147174DFF7F593508CD0D0A" httpUri="http://treatment.plazi.org/id/038287AEC147174DFF7F593508CD0D0A" lastPageId="6" lastPageNumber="475" pageId="5" pageNumber="474">
<subSubSection id="C3316533C147174EFF7F59350BED0C29" box="[151,738,807,834]" pageId="5" pageNumber="474" type="nomenclature">
<paragraph id="8B9436B8C147174EFF7F59350BED0C29" blockId="5.[151,738,807,870]" box="[151,738,807,834]" pageId="5" pageNumber="474">
<heading id="D0DC81D4C147174EFF7F59350BED0C29" bold="true" box="[151,738,807,834]" fontSize="11" level="1" pageId="5" pageNumber="474" reason="1">
<taxonomicName id="4C2B4D3BC147174EFF7F59350BED0C29" authority="Squires &amp; Goedert, 1996" authorityName="Squires &amp; Goedert" authorityYear="1996" box="[151,738,807,834]" class="Gastropoda" family="Omalogyridae" genus="Ammonicera" kingdom="Animalia" order="Heterostropha" pageId="5" pageNumber="474" phylum="Mollusca" rank="species" species="benhami">
<emphasis id="B95FEAAAC147174EFF7F59350BED0C29" bold="true" box="[151,738,807,834]" pageId="5" pageNumber="474">
<emphasis id="B95FEAAAC147174EFF7F593508AA0C29" bold="true" box="[151,421,807,834]" italics="true" pageId="5" pageNumber="474">Ammonicera benhami</emphasis>
Squires &amp; Goedert, 1996
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3316533C147174EFF7F595108260C0D" box="[151,297,844,870]" pageId="5" pageNumber="474" type="description">
<paragraph id="8B9436B8C147174EFF7F595108260C0D" blockId="5.[151,738,807,870]" box="[151,297,844,870]" pageId="5" pageNumber="474">
<figureCitation id="13102A3DC147174EFF7F595108260C0D" box="[151,297,844,870]" captionStart="FIGURE 3" captionStartId="5.[151,250,1941,1965]" captionTargetBox="[177,1411,1110,1917]" captionTargetId="figure-221@5.[177,1411,1110,1917]" captionTargetPageId="5" captionText="FIGURE 3. Ammonicera benhami (SEM), specimen MIMB 41136–1, apical view (A, C) and apical view at oblique angle (B, E); specimen MIMB 41136–2, umbilical view (D); general view (A, B); detail of the protoconch (C-E). Scale bars: A, B—100 μm; C-D—20 μm." figureDoi="http://doi.org/10.5281/zenodo.4921147" httpUri="https://zenodo.org/record/4921147/files/figure.png" pageId="5" pageNumber="474">Figure 3A–E</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3316533C147174EFF7F59890A9B0CC5" box="[151,916,916,943]" pageId="5" pageNumber="474" type="reference_group">
<paragraph id="8B9436B8C147174EFF7F59890A9B0CC5" blockId="5.[151,916,916,943]" box="[151,916,916,943]" pageId="5" pageNumber="474">
<taxonomicName id="4C2B4D3BC147174EFF7F59890A0B0CC4" authority="Squires &amp; Goedert, 1996, p. 235" authorityName="Squires &amp; Goedert" authorityPageNumber="235" authorityYear="1996" box="[151,772,916,943]" class="Gastropoda" family="Omalogyridae" genus="Ammonicera" kingdom="Animalia" order="Heterostropha" pageId="5" pageNumber="474" phylum="Mollusca" rank="species" species="benhami">
<emphasis id="B95FEAAAC147174EFF7F598908820CC5" box="[151,397,916,942]" italics="true" pageId="5" pageNumber="474">Ammonicera benhami</emphasis>
Squires &amp; Goedert, 1996, p. 235
</taxonomicName>
, figs. 27–32.
</paragraph>
</subSubSection>
<subSubSection id="C3316533C147174DFF7F59C10A7C0E0F" lastPageId="6" lastPageNumber="475" pageId="5" pageNumber="474" type="materials_examined">
<paragraph id="8B9436B8C147174EFF7F59C10D6A0B54" blockId="5.[151,1436,988,1087]" pageId="5" pageNumber="474">
<emphasis id="B95FEAAAC147174EFF7F59C108890C9D" bold="true" box="[151,390,988,1014]" pageId="5" pageNumber="474">Examined material.</emphasis>
<materialsCitation id="3B433CE5C147174EFE6759C00AF20C9D" ID-GBIF-Occurrence="3323273306" box="[399,1021,988,1015]" collectionCode="MIMB" pageId="5" pageNumber="474" specimenCode="MIMB 41136" specimenCount="4">
<specimenCode id="DB8D9EC3C147174EFE6759C00B230C9D" box="[399,556,988,1015]" collectionCode="MIMB" country="Russia" httpUri="http://grbio.org/cool/063z-3mpp" name="Museum of the Institute of Marine Biology" pageId="5" pageNumber="474">MIMB 41136</specimenCode>
(lot of
<specimenCount id="9D2DFD31C147174EFD6E59C10A190C9C" box="[646,790,988,1015]" pageId="5" pageNumber="474" type="generic">3 specimens</specimenCount>
), and
<specimenCount id="9D2DFD31C147174EFCB659C00AF20C9D" box="[862,1021,988,1015]" pageId="5" pageNumber="474" type="generic">one specimen</specimenCount>
</materialsCitation>
,
<materialsCitation id="3B433CE5C147174EFBE259C10B010B71" ID-GBIF-Occurrence="3323273308" collectionCode="SBMNH" pageId="5" pageNumber="474" specimenCode="SBMNH 467057" specimenCount="1">
<specimenCode id="DB8D9EC3C147174EFBE259C10DC10C9C" box="[1034,1230,988,1015]" collectionCode="SBMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" lsid="urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" pageId="5" pageNumber="474" type="Museum">SBMNH 467057</specimenCode>
, all from the
<typeStatus id="5490881AC147174EFA8459C00C930C9C" box="[1388,1436,989,1015]" pageId="5" pageNumber="474">type</typeStatus>
locality. One additional specimen
</materialsCitation>
,
<materialsCitation id="3B433CE5C147174EFDF05E1D0A170B71" ID-GBIF-Occurrence="3323273302" box="[536,792,1024,1051]" collectionCode="SBMNH" pageId="5" pageNumber="474" specimenCode="SBMNH 467058" specimenCount="1">
<specimenCode id="DB8D9EC3C147174EFDF05E1D0BD70B70" box="[536,728,1024,1051]" collectionCode="SBMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" lsid="urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" pageId="5" pageNumber="474" type="Museum">SBMNH 467058</specimenCode>
, from
</materialsCitation>
<collectionCode id="ED3AAE7DC147174EFCF65E1D0A880B71" box="[798,903,1024,1050]" pageId="5" pageNumber="474">LACMIP</collectionCode>
loc. 16940 [Crescent/McIntosh Formation transition zone of
<bibRefCitation id="EFBA4B49C147174EFEDF5E390B530B55" author="Squires, R. L. &amp; Goedert, J. L." box="[311,604,1059,1086]" pageId="5" pageNumber="474" pagination="254 - 269" refId="ref7043" refString="Squires, R. L. &amp; Goedert, J. L. (1995) New species of middle Eocene gastropods from the northern Doty Hills, southwestern Washington. The Veliger, 38, 254 - 269." type="journal article" year="1995">Squires &amp; Goedert (1995)</bibRefCitation>
] is herein tentatively identified as
<taxonomicName id="4C2B4D3BC147174EFC375E390D510B55" authorityName="Squires &amp; Goedert" authorityYear="1996" box="[991,1118,1060,1086]" class="Gastropoda" family="Omalogyridae" genus="Ammonicera" kingdom="Animalia" order="Heterostropha" pageId="5" pageNumber="474" phylum="Mollusca" rank="species" species="benhami">
<emphasis id="B95FEAAAC147174EFC375E390D510B55" box="[991,1118,1060,1086]" italics="true" pageId="5" pageNumber="474">A. benhami</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF546630C147174EFF7F5D88085C089D" ID-DOI="http://doi.org/10.5281/zenodo.4921147" ID-Zenodo-Dep="4921147" httpUri="https://zenodo.org/record/4921147/files/figure.png" pageId="5" pageNumber="474" startId="5.[151,250,1941,1965]" targetBox="[177,1411,1110,1917]" targetPageId="5">
<paragraph id="8B9436B8C147174EFF7F5D88085C089D" blockId="5.[151,1437,1941,2038]" pageId="5" pageNumber="474">
<emphasis id="B95FEAAAC147174EFF7F5D88081C08C6" bold="true" box="[151,275,1941,1965]" pageId="5" pageNumber="474">FIGURE 3.</emphasis>
<taxonomicName id="4C2B4D3BC147174EFEF25D8B0B4808C5" authority="(SEM)" baseAuthorityName="SEM" box="[282,583,1941,1966]" class="Gastropoda" family="Omalogyridae" genus="Ammonicera" kingdom="Animalia" order="Heterostropha" pageId="5" pageNumber="474" phylum="Mollusca" rank="species" species="benhami">
<emphasis id="B95FEAAAC147174EFEF25D8B08F608C6" box="[282,505,1941,1965]" italics="true" pageId="5" pageNumber="474">Ammonicera benhami</emphasis>
(SEM)
</taxonomicName>
, specimen MIMB 41136–1, apical view (A, C) and apical view at oblique angle (B, E); specimen MIMB 41136–2, umbilical view (D); general view (A, B); detail of the protoconch (C-E). Scale bars: A, B—100 μm; C-D—20 μm.
</paragraph>
</caption>
<paragraph id="8B9436B8C144174DFF2F5A8B0D4E0FDB" blockId="6.[151,1437,150,609]" box="[199,1089,150,177]" pageId="6" pageNumber="475">
<emphasis id="B95FEAAAC144174DFF2F5A8B087A0FDB" bold="true" box="[199,373,150,176]" pageId="6" pageNumber="475">Type material.</emphasis>
<materialsCitation id="3B433CE5C144174DFE945A8B0BA40FDB" ID-GBIF-Occurrence="3323273315" box="[380,683,150,176]" collectionCode="LACMIP" pageId="6" pageNumber="475" specimenCode="LACMIP 11368" specimenCount="1" typeStatus="holotype">
<typeStatus id="5490881AC144174DFE945A8B08E60FDB" box="[380,489,150,176]" pageId="6" pageNumber="475" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DB8D9EC3C144174DFE1B5A8B0BA40FDB" box="[499,683,150,176]" collectionCode="LACMIP" pageId="6" pageNumber="475">LACMIP 11368</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B433CE5C144174DFD5E5A8A0D320FDB" ID-GBIF-Occurrence="3323273320" box="[694,1085,150,177]" collectionCode="LACMIP" pageId="6" pageNumber="475" specimenCode="LACMIP 11369-11371" specimenCount="1" typeStatus="paratype">
<typeStatus id="5490881AC144174DFD5E5A8A0A280FDA" box="[694,807,151,177]" pageId="6" pageNumber="475" type="paratype">Paratypes</typeStatus>
:
<specimenCode id="DB8D9EC3C144174DFCDA5A8B0D320FDB" box="[818,1085,150,176]" collectionCode="LACMIP" pageId="6" pageNumber="475">LACMIP 11369–11371</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9436B8C144174DFF2F5AA7088A0F92" blockId="6.[151,1437,150,609]" pageId="6" pageNumber="475">
<materialsCitation id="3B433CE5C144174DFF2F5AA7088E0F92" ID-GBIF-Occurrence="3323273322" collectionCode="LACMIP" country="United States of America" county="Eocene" location="Larch Mountain" municipality="Crescent Formation" pageId="6" pageNumber="475" specimenCount="1" stateProvince="Washington" typeStatus="holotype">
<emphasis id="B95FEAAAC144174DFF2F5AA7086B0FBF" bold="true" box="[199,356,186,212]" pageId="6" pageNumber="475">
<typeStatus id="5490881AC144174DFF2F5AA7080D0FBF" box="[199,258,186,212]" pageId="6" pageNumber="475">Type</typeStatus>
locality.
</emphasis>
<collectionCode id="ED3AAE7DC144174DFE825AA708DC0FBF" box="[362,467,186,212]" pageId="6" pageNumber="475">LACMIP</collectionCode>
loc. 16655,
<location id="8EF46063C144174DFDB25AA70A1E0FBF" LSID="urn:lsid:plazi:treatment:038287AEC147174DFF7F593508CD0D0A:8EF46063C144174DFDB25AA70A1E0FBF" box="[602,785,186,212]" country="United States of America" county="Eocene" municipality="Crescent Formation" name="Larch Mountain" pageId="6" pageNumber="475" stateProvince="Washington">Larch Mountain</location>
,
<location id="8EF46063C144174DFCF25AA70A950FBF" LSID="urn:lsid:plazi:treatment:038287AEC147174DFF7F593508CD0D0A:8EF46063C144174DFCF25AA70A950FBF" box="[794,922,186,212]" country="United States of America" county="Eocene" municipality="Crescent Formation" name="Black Hills" pageId="6" pageNumber="475" stateProvince="Washington">Black Hills</location>
,
<collectingRegion id="49EFF85AC144174DFC4C5AA70D650FBF" box="[932,1130,186,212]" country="United States of America" name="Washington" pageId="6" pageNumber="475">Washington State</collectingRegion>
,
<collectingCountry id="F33C7628C144174DFB9B5AA70DA00FBF" box="[1139,1199,186,212]" name="United States of America" pageId="6" pageNumber="475">USA</collectingCountry>
;
<collectingMunicipality id="6BF0ACC2C144174DFB515AA70C960FBF" box="[1209,1433,186,212]" pageId="6" pageNumber="475">Crescent Formation</collectingMunicipality>
, middle early
<collectingCounty id="62F54E34C144174DFEC35AC2088E0F92" box="[299,385,223,249]" pageId="6" pageNumber="475">Eocene</collectingCounty>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9436B8C144174DFF2F5B1F0A7C0E0F" blockId="6.[151,1437,150,609]" pageId="6" pageNumber="475">
<emphasis id="B95FEAAAC144174DFF2F5B1F08B90E77" bold="true" box="[199,438,258,284]" pageId="6" pageNumber="475">Emended diagnosis.</emphasis>
<taxonomicName id="4C2B4D3BC144174DFE555B1E0B430E76" authorityName="Vayssiere" authorityYear="1893" box="[445,588,259,285]" class="Gastropoda" family="Omalogyridae" genus="Ammonicera" kingdom="Animalia" order="Heterostropha" pageId="6" pageNumber="475" phylum="Mollusca" rank="genus">
<emphasis id="B95FEAAAC144174DFE555B1E0B430E76" box="[445,588,259,285]" italics="true" pageId="6" pageNumber="475">Ammonicera</emphasis>
</taxonomicName>
with protoconch sculptured by two spiral cords (prominent central and low outer) separated by a spiral groove; teleoconch with one wide subperipheral groove and low subperipheral cord on both apical and basal surfaces, and prominent nodelike axial ribs.
</paragraph>
</subSubSection>
<subSubSection id="C3316533C144174DFF2F5B7308CD0D0A" pageId="6" pageNumber="475" type="discussion">
<paragraph id="8B9436B8C144174DFF2F5B7308CD0D0A" blockId="6.[151,1437,150,609]" pageId="6" pageNumber="475">
<emphasis id="B95FEAAAC144174DFF2F5B7308340EE3" bold="true" box="[199,315,366,392]" pageId="6" pageNumber="475">Remarks.</emphasis>
The size range of
<taxonomicName id="4C2B4D3BC144174DFDFB5B720B9C0EE3" baseAuthorityName="SEM" box="[531,659,366,393]" class="Gastropoda" family="Omalogyridae" genus="Ammonicera" kingdom="Animalia" order="Heterostropha" pageId="6" pageNumber="475" phylum="Mollusca" rank="species" species="benhami">
<emphasis id="B95FEAAAC144174DFDFB5B720B9C0EE3" box="[531,659,366,393]" italics="true" pageId="6" pageNumber="475">A. benhami</emphasis>
</taxonomicName>
is
<quantity id="4CD39B5DC144174DFD505B730A670EE2" box="[696,872,366,393]" metricMagnitude="-4" metricUnit="m" metricValue="4.95" metricValueMax="5.8999999999999995" metricValueMin="4.0" pageId="6" pageNumber="475" unit="mm" value="0.495" valueMax="0.59" valueMin="0.4">0.4 to 0.59 mm</quantity>
in diameter (Squires &amp; Goedert 1996); specimen MIMB 41136–1 (
<figureCitation id="13102A3DC144174DFEB75B8F08B70EC7" box="[351,440,402,428]" captionStart="FIGURE 3" captionStartId="5.[151,250,1941,1965]" captionTargetBox="[177,1411,1110,1917]" captionTargetId="figure-221@5.[177,1411,1110,1917]" captionTargetPageId="5" captionText="FIGURE 3. Ammonicera benhami (SEM), specimen MIMB 41136–1, apical view (A, C) and apical view at oblique angle (B, E); specimen MIMB 41136–2, umbilical view (D); general view (A, B); detail of the protoconch (C-E). Scale bars: A, B—100 μm; C-D—20 μm." figureDoi="http://doi.org/10.5281/zenodo.4921147" httpUri="https://zenodo.org/record/4921147/files/figure.png" pageId="6" pageNumber="475">Fig. 3A</figureCitation>
) has a diameter of
<quantity id="4CD39B5DC144174DFD7C5B8F0BF30EC6" box="[660,764,402,429]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="475" unit="mm" value="0.5">0.50 mm</quantity>
.
<taxonomicName id="4C2B4D3BC144174DFCE05B8E0AF00EC7" baseAuthorityName="SEM" box="[776,1023,402,429]" class="Gastropoda" family="Omalogyridae" genus="Ammonicera" kingdom="Animalia" order="Heterostropha" pageId="6" pageNumber="475" phylum="Mollusca" rank="species" species="benhami">
<emphasis id="B95FEAAAC144174DFCE05B8E0AF00EC7" box="[776,1023,402,429]" italics="true" pageId="6" pageNumber="475">Ammonicera benhami</emphasis>
</taxonomicName>
is only known from three localities, two in the Black Hills southwest of Olympia,
<collectingRegion id="49EFF85AC144174DFD4E5BAB0A210EBB" box="[678,814,438,464]" country="United States of America" name="Washington" pageId="6" pageNumber="475">Washington</collectingRegion>
, and now tentatively from a single locality (weathered and no longer accessible) in the Doty Hills, Lewis County,
<collectingRegion id="49EFF85AC144174DFCD05BC70ACE0E9F" box="[824,961,474,500]" country="United States of America" name="Washington" pageId="6" pageNumber="475">Washington</collectingRegion>
. The two localities in the Black Hills represent deposition between basalt boulders within the tidal zone (
<bibRefCitation id="EFBA4B49C144174DFC835BE30D8D0D73" author="Squires, R. L. &amp; Goedert, J. L." box="[875,1154,510,536]" pageId="6" pageNumber="475" pagination="253 - 266" refId="ref6998" refString="Squires, R. L. &amp; Goedert, J. L. (1994) New species of early Eocene small to minute mollusks from the Crescent Formation, Black Hills, southwestern Washington. The Veliger, 37, 253 - 266." type="journal article" year="1994">Squires &amp; Goedert 1994</bibRefCitation>
, 1996) whereas the Doty Hills localities may represent shallow water taxa transported a short distance into deeper waters off volcanic islands (
<bibRefCitation id="EFBA4B49C144174DFF77585B08BA0D0A" author="Squires, R. L. &amp; Goedert, J. L." box="[159,437,582,609]" pageId="6" pageNumber="475" pagination="254 - 269" refId="ref7043" refString="Squires, R. L. &amp; Goedert, J. L. (1995) New species of middle Eocene gastropods from the northern Doty Hills, southwestern Washington. The Veliger, 38, 254 - 269." type="journal article" year="1995">Squires &amp; Goedert 1995</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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