260 lines
29 KiB
XML
260 lines
29 KiB
XML
<document id="97079444B3AD3054FF91D266B2EE0D49" ID-ISSN="2107-7207" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1683043797380" checkinUser="felipe" docAuthor="PešićK, Vladimir, SmitK, Harry & GurungK, Mer Man" docDate="2023" docId="0382879642172E4EFE60FC72FDD9FDE4" docLanguage="en" docName="Acarologia.63.1.148-168.pdf" docOrigin="Acarologia 63 (1)" docStyle="DocumentStyle:0387F17C4AF9F8D6336E46CC5E5664D0.4:Acarologia.2018-.journal_article" docStyleId="0387F17C4AF9F8D6336E46CC5E5664D0" docStyleName="Acarologia.2018-.journal_article" docStyleVersion="4" docTitle="Atractides (Atractides) bhutanicus PešićK & SmitK & GurungK 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="161" masterDocId="FFBBFFEE421C2E40FF8BFFC4FFA4FF9D" masterDocTitle="Five new species of water mite genus Atractides Koch, 1837 from Bhutan with new records (Acari: Hydrachnidia: Hygrobatidae)" masterLastPageNumber="168" masterPageNumber="148" pageNumber="158" updateTime="1684344273785" updateUser="carolina" zenodo-license-document="CC-BY-4.0">
|
||
<mods:mods id="64DD8DBEDB87D983FF53AD156674E535" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="9BE24AA442C661B0E30DF0F574F0B1FE">
|
||
<mods:title id="54337C8B33B5D4B83E4BBA0038D49114">Five new species of water mite genus Atractides Koch, 1837 from Bhutan with new records (Acari: Hydrachnidia: Hygrobatidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="4F066E8B6ED604539C1FB83E678B9A4B" type="personal">
|
||
<mods:role id="E4CCA7F48EBCEA83DCEB30F6C5D60CE8">
|
||
<mods:roleTerm id="9D2A5E485F13CC6364F2837F5BDD5C4C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="18F573266AB67653CD8CC76A9F58C358">PešićK, Vladimir</mods:namePart>
|
||
<mods:affiliation id="03C5DF2CF3BB48C76C164238AD8507E3">University of Montenegro, Cetinjski put bb, 81000 Podgorica, Montenegro.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="37BFD6E51FE2B3338AFD62D279C721F3" type="personal">
|
||
<mods:role id="1B9FAB5762D528E5A27AA964C9218C8D">
|
||
<mods:roleTerm id="883C63BF171991BD37D26268EDF4204D">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="2FF7528DC5A6B70F4B4763F57AB17A17">SmitK, Harry</mods:namePart>
|
||
<mods:affiliation id="97D573BFD380333565FE0CA38D551A90">Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, the Netherlands.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="2F41370E2BD7F8886A859B95E8F89D3C" type="personal">
|
||
<mods:role id="F8AA74AB046B9AC643CA997CCDBAA064">
|
||
<mods:roleTerm id="8277AF6F095624336F9D72A862CD3924">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A51273BD146F65DE2B1C9272917D1944">GurungK, Mer Man</mods:namePart>
|
||
<mods:affiliation id="97FA967C13E060E6B8A24899635FDEC3">Department of Forest Science, College of Natural Resources, Royal University of Bhutan, Lobesa, & University of Montenegro, Cetinjski put bb, 81000 Podgorica, Montenegro.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="752253E6EAB09CB3D00E8159511C5F2D">text</mods:typeOfResource>
|
||
<mods:relatedItem id="2F0AD21AE54CFC62CB7DE162726E2324" type="host">
|
||
<mods:titleInfo id="53565797D360EBF08BDF26E52B51B0BE">
|
||
<mods:title id="B0D69A2FF77E8BE761E2C962C57AB595">Acarologia</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="1D41E704C46CD87B8137B1FD60969BB2">
|
||
<mods:date id="24148891CF15A778A51E8E6CC6E496B1">2023</mods:date>
|
||
<mods:detail id="5D59BBF7736ABE6696BCB0D3FF23C0BF" type="pubDate">
|
||
<mods:number id="883FBAEF69CC30BA36B6A722E8007157">2023-01-30</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="4AD6DC20F1ADAC0784FEB4BEC45C91DE" type="volume">
|
||
<mods:number id="A5538FC8458F567DE2E0219363265680">63</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="BCA20AE9C3956A2D74B4E1D778B16BF3" type="issue">
|
||
<mods:number id="9D35C7659540C5E219C67217D2A55CFB">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="FC609BD1A814427CEA52E35D4AFEACDB" unit="page">
|
||
<mods:start id="CB9A11D17839BA2A8CAAFB7BC1BFE60B">148</mods:start>
|
||
<mods:end id="A684CE4C856054ECFBECFDDDF04CA6DC">168</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="C18D52D84974DC2D8BCF29619C8487E9">journal article</mods:classification>
|
||
<mods:identifier id="04841C1D2FE18D730E7338198D179279" type="ISSN">2107-7207</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="0382879642172E4EFE60FC72FDD9FDE4" LSID="urn:lsid:plazi:treatment:0382879642172E4EFE60FC72FDD9FDE4" httpUri="http://treatment.plazi.org/id/0382879642172E4EFE60FC72FDD9FDE4" lastPageId="14" lastPageNumber="161" pageId="11" pageNumber="158">
|
||
<subSubSection id="C331650B42172E4BFE60FC72FC4FFB9C" pageId="11" pageNumber="158" type="nomenclature">
|
||
<paragraph id="8B94368042172E4BFE60FC72FBAFFC4D" blockId="11.[491,1035,950,976]" box="[491,1035,950,976]" pageId="11" pageNumber="158">
|
||
<heading id="D0DC81EC42172E4BFE60FC72FBAFFC4D" bold="true" box="[491,1035,950,976]" fontSize="11" level="2" pageId="11" pageNumber="158" reason="3">
|
||
<emphasis id="B95FEA9242172E4BFE60FC72FBAFFC4D" bold="true" box="[491,1035,950,976]" pageId="11" pageNumber="158">
|
||
<taxonomicName id="4C2B4D0342172E4BFE60FC72FC06FC4D" authority="PešićK & SmitK & GurungK, 2023" authorityName="PešićK & SmitK & GurungK" authorityYear="2023" box="[491,930,950,976]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="158" phylum="Arthropoda" rank="species" species="bhutanicus" status="sp. nov." subGenus="Atractides">
|
||
<emphasis id="B95FEA9242172E4BFE60FC72FDCBFC4D" bold="true" box="[491,623,950,976]" italics="true" pageId="11" pageNumber="158">Atractides</emphasis>
|
||
(
|
||
<emphasis id="B95FEA9242172E4BFD0BFC72FCA0FC4D" bold="true" box="[640,772,950,976]" italics="true" pageId="11" pageNumber="158">Atractides</emphasis>
|
||
)
|
||
<emphasis id="B95FEA9242172E4BFC9FFC72FC06FC4D" bold="true" box="[788,930,950,976]" italics="true" pageId="11" pageNumber="158">bhutanicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26C57E942172E4BFC22FC72FBAFFC4D" box="[937,1035,950,976]" pageId="11" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B94368042172E4BFE60FC2AFC4FFB9C" blockId="11.[491,1536,1006,1994]" box="[491,1003,1006,1025]" pageId="11" pageNumber="158">
|
||
Zoobank:
|
||
<uuid id="FF8D0C5542172E4BFDCFFC2AFC4FFB9C" box="[580,1003,1006,1025]" pageId="11" pageNumber="158">F09D45C0-7937-441A-AA3D-CA3D46DE9537</uuid>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C331650B42172E4BFD98FBC9FD2CFBB8" box="[531,648,1037,1061]" pageId="11" pageNumber="158" type="description">
|
||
<paragraph id="8B94368042172E4BFD98FBC9FD2CFBB8" blockId="11.[491,1536,1006,1994]" box="[531,648,1037,1061]" pageId="11" pageNumber="158">
|
||
<figureCitation id="13102A0542172E4BFD98FBC9FD2CFBB8" box="[531,648,1037,1061]" captionStart-0="Figure 7" captionStart-1="Figure 8" captionStartId-0="12.[119,186,2025,2046]" captionStartId-1="13.[515,582,1555,1576]" captionTargetBox-0="[156,1441,221,1969]" captionTargetBox-1="[529,1482,234,1494]" captionTargetId-0="figure-2@12.[156,1441,221,1969]" captionTargetId-1="figure-2@13.[503,1486,221,1500]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="Figure 7 Atractides bhutanicussp. nov., ♂ holotype, Chendebji Chhu, Bhutan. A – idiosoma, ventral view; B – palp, medial view; C – I-L-5 and -6. Scale bars = 100 µm." captionText-1="Figure 8 Atractides bhutanicussp. nov., ♀ paratype, Chendebji Chhu, Bhutan. A – genital field; B – palp, medial view; C – I-L-5 and -6. Scale bar = 100 µm" pageId="11" pageNumber="158">Figures 7-8</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C331650B42172E4BFD98FBF4FD6DFB4F" pageId="11" pageNumber="158" type="materials_examined">
|
||
<paragraph id="8B94368042172E4BFD98FBF4FB74FB10" blockId="11.[491,1536,1006,1994]" pageId="11" pageNumber="158">
|
||
<emphasis id="B95FEA9242172E4BFD98FBF4FD10FBD5" bold="true" box="[531,692,1072,1096]" pageId="11" pageNumber="158">Type material</emphasis>
|
||
—
|
||
<materialsCitation id="3B433CDD42172E4BFD69FBF4FB36FBF7" collectingDate="2021-11-08" collectionCode="NBC" country="Bhutan" elevation="2483" latitude="27.47692" location="Chendebji Chhu" longLatPrecision="1" longitude="90.35265" pageId="11" pageNumber="158" specimenCode="MG16" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<typeStatus id="5490882242172E4BFD69FBF4FCE7FBD5" box="[738,835,1072,1096]" pageId="11" pageNumber="158">Holotype</typeStatus>
|
||
<specimenCount id="9D2DFD0942172E4BFCC6FBF4FCC7FBDA" box="[845,867,1072,1095]" pageId="11" pageNumber="158" type="male">♂</specimenCount>
|
||
, dissected and slide mounted (
|
||
<collectionCode id="ED3AAE4542172E4BFB5FFBF4FAABFBD5" box="[1236,1295,1072,1096]" pageId="11" pageNumber="158">NBC</collectionCode>
|
||
),
|
||
<collectingCountry id="F33C761042172E4BFAAFFBF4FAD2FBD5" box="[1316,1398,1072,1096]" name="Bhutan" pageId="11" pageNumber="158">Bhutan</collectingCountry>
|
||
,
|
||
<specimenCode id="DB8D9EFB42172E4BFA0EFBF4FA68FBD5" box="[1413,1484,1072,1096]" collectionCode="MG" country="China" name="Museum of Zoology" pageId="11" pageNumber="158">MG16</specimenCode>
|
||
<location id="8EF4605B42172E4BFE60FB96FD09FBF7" LSID="urn:lsid:plazi:treatment:0382879642172E4EFE60FC72FDD9FDE4:8EF4605B42172E4BFE60FB96FD09FBF7" box="[491,685,1106,1130]" country="Bhutan" latitude="27.47692" longLatPrecision="1" longitude="90.35265" name="Chendebji Chhu" pageId="11" pageNumber="158">Chendebji Chhu</location>
|
||
,
|
||
<geoCoordinate id="EE1F504742172E4BFD3CFB96FC85FBF7" box="[695,801,1106,1130]" degrees="27.47692" direction="north" orientation="latitude" pageId="11" pageNumber="158" precision="1" value="27.47692">27.47692°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1F504742172E4BFCA0FB96FC3AFBF7" box="[811,926,1106,1130]" degrees="90.35265" direction="east" orientation="longitude" pageId="11" pageNumber="158" precision="1" value="90.35265">90.35265°E</geoCoordinate>
|
||
,
|
||
<elevation id="0006D1B342172E4BFC23FB96FBB5FBF7" box="[936,1041,1106,1130]" metricMagnitude="3" metricUnit="m" metricValue="2.483" pageId="11" pageNumber="158" unit="m" value="2483.0">
|
||
<quantity id="4CD39B6542172E4BFC23FB96FC4FFBF7" box="[936,1003,1106,1130]" metricMagnitude="3" metricUnit="m" metricValue="2.483" pageId="11" pageNumber="158" unit="m" value="2483.0">2483 m</quantity>
|
||
asl
|
||
</elevation>
|
||
,
|
||
<date id="FF95104042172E4BFB97FB96FB36FBF7" box="[1052,1170,1106,1130]" pageId="11" pageNumber="158" value="2021-11-08">
|
||
<collectingDate id="EFD1E9A842172E4BFB97FB96FB36FBF7" box="[1052,1170,1106,1130]" pageId="11" pageNumber="158" value="2021-11-08">8 Nov. 2021</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<typeStatus id="5490882242172E4BFB15FB97FAACFBF6" box="[1182,1288,1107,1131]" pageId="11" pageNumber="158">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="9D2DFD0942172E4BFA9FFB96FA96FBF7" box="[1300,1330,1106,1130]" pageId="11" pageNumber="158" type="female">2♀</specimenCount>
|
||
, same site and data as the
|
||
<typeStatus id="5490882242172E4BFD15FBB1FD5BFB10" box="[670,767,1141,1165]" pageId="11" pageNumber="158">holotype</typeStatus>
|
||
,
|
||
<specimenCount id="9D2DFD0942172E4BFC81FBB1FC8FFB10" box="[778,811,1141,1165]" pageId="11" pageNumber="158" type="female">1♀</specimenCount>
|
||
dissected and slide mounted (
|
||
<collectionCode id="ED3AAE4542172E4BFB03FBB1FB60FB10" box="[1160,1220,1141,1165]" pageId="11" pageNumber="158">NBC</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B94368042172E4BFD98FB5CFA01FB32" blockId="11.[491,1536,1006,1994]" box="[531,1445,1175,1200]" pageId="11" pageNumber="158">
|
||
<emphasis id="B95FEA9242172E4BFD98FB5CFD1AFB2D" bold="true" box="[531,702,1176,1200]" pageId="11" pageNumber="158">Other material</emphasis>
|
||
—
|
||
<collectingCountry id="F33C761042172E4BFD6CFB53FC9DFB32" box="[743,825,1175,1199]" name="Bhutan" pageId="11" pageNumber="158">Bhutan</collectingCountry>
|
||
,
|
||
<collectionCode id="ED3AAE4542172E4BFCCFFB53FCCBFB32" box="[836,879,1175,1199]" country="China" name="Museum of Zoology" pageId="11" pageNumber="158">MG</collectionCode>
|
||
13 Rukhubji Chhu,
|
||
<geoCoordinate id="EE1F504742172E4BFBC1FB53FB18FB32" box="[1098,1212,1175,1199]" degrees="27.51174" direction="north" orientation="latitude" pageId="11" pageNumber="158" precision="1" value="27.51174">27.51174°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1F504742172E4BFB4DFB53FA8BFB32" box="[1222,1327,1175,1199]" degrees="90.29711" direction="east" orientation="longitude" pageId="11" pageNumber="158" precision="1" value="90.29711">90.29711°E</geoCoordinate>
|
||
,
|
||
<quantity id="4CD39B6542172E4BFAB2FB53FADFFB32" box="[1337,1403,1175,1199]" metricMagnitude="3" metricUnit="m" metricValue="2.5869999999999997" pageId="11" pageNumber="158" unit="m" value="2587.0">2587 m</quantity>
|
||
asl,
|
||
</paragraph>
|
||
<paragraph id="8B94368042172E4BFE60FB7EFD6DFB4F" blockId="11.[491,1536,1006,1994]" box="[491,713,1210,1234]" pageId="11" pageNumber="158">
|
||
<date id="FF95104042172E4BFE60FB7EFDC2FB4F" box="[491,614,1210,1234]" pageId="11" pageNumber="158" value="2021-11-05">5 Nov. 2021</date>
|
||
,
|
||
<specimenCount id="9D2DFD0942172E4BFDFAFB7EFD30FB4F" box="[625,660,1210,1234]" pageId="11" pageNumber="158" type="male">1♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D2DFD0942172E4BFD14FB7EFD60FB4F" box="[671,708,1210,1234]" pageId="11" pageNumber="158" type="female">6♀</specimenCount>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C331650B42172E4BFD98FB19FB60FAA4" pageId="11" pageNumber="158" type="diagnosis">
|
||
<paragraph id="8B94368042172E4BFD98FB19FB60FAA4" blockId="11.[491,1536,1006,1994]" pageId="11" pageNumber="158">
|
||
<emphasis id="B95FEA9242172E4BFD98FB19FD22FB68" bold="true" box="[531,646,1245,1269]" pageId="11" pageNumber="158">Diagnosis</emphasis>
|
||
— Integument finely striated, muscle insertions unsclerotized; anterior margin of male genital field slightly concave, but without an indentation; Vgl-1 separate from Vgl-2; P-2 with a distoventral projection, ventral margin of P-4 projecting.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C331650B42172E4DFD98FA80FB0CF87C" lastPageId="13" lastPageNumber="160" pageId="11" pageNumber="158" type="description">
|
||
<paragraph id="8B94368042172E4BFD98FA80FA26F994" blockId="11.[491,1536,1006,1994]" pageId="11" pageNumber="158">
|
||
<emphasis id="B95FEA9242172E4BFD98FA80FD32FAC1" bold="true" box="[531,662,1348,1372]" pageId="11" pageNumber="158">Description</emphasis>
|
||
—
|
||
<emphasis id="B95FEA9242172E4BFD4AFA80FCD8FAC1" box="[705,892,1348,1372]" italics="true" pageId="11" pageNumber="158">General features</emphasis>
|
||
– Integument striated, muscle insertions unsclerotized; mediocaudal margin Cx-I convex, apodemes of Cx-II in an acute angle with the median line. Genital field with large Ac in a triangular arrangement. Excretory pore smooth; Vgl-1 not fused to Vgl-2. Palp with sexual dimorphism, P-4 sword seta slender, between ventral setae, nearer to distoventral seta. I-L-5 with seta S-1 slender and bluntly pointed, S-2 shorter and pointed, proximally enlarged; I-L-6 slender, curved, basally slightly thickened (
|
||
<figureCitation id="13102A0542172E4BFA8DFA35FAD2F994" box="[1286,1398,1521,1545]" captionStart="Figure 7" captionStartId="12.[119,186,2025,2046]" captionTargetBox="[156,1441,221,1969]" captionTargetId="figure-2@12.[156,1441,221,1969]" captionTargetPageId="12" captionText="Figure 7 Atractides bhutanicussp. nov., ♂ holotype, Chendebji Chhu, Bhutan. A – idiosoma, ventral view; B – palp, medial view; C – I-L-5 and -6. Scale bars = 100 µm." pageId="11" pageNumber="158">Figure 7C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B94368042172E4BFD98F9D7FB59F9ED" blockId="11.[491,1536,1006,1994]" pageId="11" pageNumber="158">
|
||
<emphasis id="B95FEA9242172E4BFD98F9D7FDEAF9B6" box="[531,590,1555,1579]" italics="true" pageId="11" pageNumber="158">Male</emphasis>
|
||
– Genital plate anterior margin slightly concave, posterior margin deeply indented (
|
||
<figureCitation id="13102A0542172E4BFE79F9F2FDC1F9D3" box="[498,613,1590,1614]" captionStart="Figure 7" captionStartId="12.[119,186,2025,2046]" captionTargetBox="[156,1441,221,1969]" captionTargetId="figure-2@12.[156,1441,221,1969]" captionTargetPageId="12" captionText="Figure 7 Atractides bhutanicussp. nov., ♂ holotype, Chendebji Chhu, Bhutan. A – idiosoma, ventral view; B – palp, medial view; C – I-L-5 and -6. Scale bars = 100 µm." pageId="11" pageNumber="158">Figure 7A</figureCitation>
|
||
); P-2 with a finger-like distoventral protrusion, P-3 ventral margin concave, P-4 thickened, maximum height at level of proximoventral hair (
|
||
<figureCitation id="13102A0542172E4BFB09F99CFB56F9ED" box="[1154,1266,1624,1648]" captionStart="Figure 7" captionStartId="12.[119,186,2025,2046]" captionTargetBox="[156,1441,221,1969]" captionTargetId="figure-2@12.[156,1441,221,1969]" captionTargetPageId="12" captionText="Figure 7 Atractides bhutanicussp. nov., ♂ holotype, Chendebji Chhu, Bhutan. A – idiosoma, ventral view; B – palp, medial view; C – I-L-5 and -6. Scale bars = 100 µm." pageId="11" pageNumber="158">Figure 7B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B94368042172E4BFD98F9BFFBDBF90E" blockId="11.[491,1536,1006,1994]" box="[531,1151,1659,1683]" pageId="11" pageNumber="158">
|
||
<emphasis id="B95FEA9242172E4BFD98F9BFFDC3F90E" box="[531,615,1659,1683]" italics="true" pageId="11" pageNumber="158">Female</emphasis>
|
||
– P-2 ventral margin convex distally (
|
||
<figureCitation id="13102A0542172E4BFB8AF9BFFBD7F90E" box="[1025,1139,1659,1683]" captionStart="Figure 8" captionStartId="13.[515,582,1555,1576]" captionTargetBox="[529,1482,234,1494]" captionTargetId="figure-2@13.[503,1486,221,1500]" captionTargetPageId="13" captionText="Figure 8 Atractides bhutanicussp. nov., ♀ paratype, Chendebji Chhu, Bhutan. A – genital field; B – palp, medial view; C – I-L-5 and -6. Scale bar = 100 µm" pageId="11" pageNumber="158">Figure 8B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B94368042172E4BFD98F95AFBCDF967" blockId="11.[491,1536,1006,1994]" pageId="11" pageNumber="158">
|
||
<emphasis id="B95FEA9242172E4BFD98F95AFD1DF92B" bold="true" box="[531,697,1694,1718]" pageId="11" pageNumber="158">Measurements</emphasis>
|
||
—
|
||
<emphasis id="B95FEA9242172E4BFD68F959FCBAF928" box="[739,798,1693,1717]" italics="true" pageId="11" pageNumber="158">Male</emphasis>
|
||
– Idiosoma L 558, W 428; maximum diameter Dgl-4 27. Coxal shield L 344; Cx-III W 375; Cx-I+II mL 131, Cx-I+II lL 250. Genital field L/W 117/136, ratio 0.86, L Ac-1-3: 52-58, 52, 44-52. Ejaculatory complex L 105.
|
||
</paragraph>
|
||
<paragraph id="8B94368042172E4BFD98F8C1FA3FF880" blockId="11.[491,1536,1006,1994]" box="[531,1435,1797,1821]" pageId="11" pageNumber="158">Palp – Total L 324; dL/H, dL/H ratio: P-1, 33/31, 1.08; P-2, 72/63, 1.15; P-3, 77/48, 1.6;</paragraph>
|
||
<paragraph id="8B94368042172E4BFE60F8E3FA67F8A2" blockId="11.[491,1536,1006,1994]" box="[491,1475,1831,1855]" pageId="11" pageNumber="158">P-4, 106/44, 2.41; P-5, 36/17, 2.16; L ratio P-2/P-4, 0.68. Gnathosoma vL 150, chelicera total</paragraph>
|
||
<paragraph id="8B94368042172E4BFE60F88FFD83F8FF" blockId="11.[491,1536,1006,1994]" box="[491,551,1866,1890]" pageId="11" pageNumber="158">L 217.</paragraph>
|
||
<paragraph id="8B94368042172E4BFD98F8A9FA6CF818" blockId="11.[491,1536,1006,1994]" box="[531,1480,1901,1925]" pageId="11" pageNumber="158">Legs – I-L-5 dL 209, vL 137, dL/vL ratio 1.53, maximum H 63, dL/maximum H 3.31, S-1</paragraph>
|
||
<paragraph id="8B94368042172E4BFE60F854FA1EF83A" blockId="11.[491,1536,1006,1994]" box="[491,1466,1935,1959]" pageId="11" pageNumber="158">L 102, L/W ratio 10.8, S-2 L 80, L/W ratio 5.65, distance S-1-2, 27, dL ratio S-1/2, 1.28; I-L-6</paragraph>
|
||
<paragraph id="8B94368042172E4BFE60F876FB38F857" blockId="11.[491,1536,1006,1994]" box="[491,1180,1970,1994]" pageId="11" pageNumber="158">dL 140, central H 22, dL/central H ratio 6.39; L I-L-5/6 ratio 1.5.</paragraph>
|
||
<caption id="DF54660842102E4CFFFCF82DFE33F788" pageId="12" pageNumber="159" startId="12.[119,186,2025,2046]" targetBox="[156,1441,221,1969]" targetPageId="12" targetType="figure">
|
||
<paragraph id="8B94368042102E4CFFFCF82DFE33F788" blockId="12.[119,1507,2025,2069]" pageId="12" pageNumber="159">
|
||
<emphasis id="B95FEA9242102E4CFFFCF82DFF68F863" bold="true" box="[119,204,2025,2046]" pageId="12" pageNumber="159">Figure 7</emphasis>
|
||
<taxonomicName id="4C2B4D0342102E4CFF53F82DFE0BF862" authority="PešićK & SmitK & GurungK, 2023" authorityName="PešićK & SmitK & GurungK" authorityYear="2023" box="[216,431,2025,2047]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="12" pageNumber="159" phylum="Arthropoda" rank="species" species="bhutanicus" status="sp. nov.">
|
||
<emphasis id="B95FEA9242102E4CFF53F82DFE0BF862" box="[216,431,2025,2047]" italics="true" pageId="12" pageNumber="159">Atractides bhutanicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95FEA9242102E4CFE24F82DFE59F863" bold="true" box="[431,509,2025,2046]" pageId="12" pageNumber="159">
|
||
<taxonomicNameLabel id="A26C57E942102E4CFE24F82DFE59F863" box="[431,509,2025,2046]" pageId="12" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, ♂ holotype, Chendebji Chhu, Bhutan. A – idiosoma, ventral view; B – palp, medial view; C – I-L-5 and -6. Scale bars = 100 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF54660842112E4DFD88F9D7FB9BF9A2" pageId="13" pageNumber="160" startId="13.[515,582,1555,1576]" targetBox="[529,1482,234,1494]" targetPageId="13" targetType="figure">
|
||
<paragraph id="8B94368042112E4DFD88F9D7FB9BF9A2" blockId="13.[513,1501,1555,1599]" pageId="13" pageNumber="160">
|
||
<emphasis id="B95FEA9242112E4DFD88F9D7FDFDF9B5" bold="true" box="[515,601,1555,1576]" pageId="13" pageNumber="160">Figure 8</emphasis>
|
||
<taxonomicName id="4C2B4D0342112E4DFDEEF9D7FC98F9B4" authority="PešićK & SmitK & GurungK, 2023" authorityName="PešićK & SmitK & GurungK" authorityYear="2023" box="[613,828,1555,1577]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="160" phylum="Arthropoda" rank="species" species="bhutanicus" status="sp. nov.">
|
||
<emphasis id="B95FEA9242112E4DFDEEF9D7FC98F9B4" box="[613,828,1555,1577]" italics="true" pageId="13" pageNumber="160">Atractides bhutanicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95FEA9242112E4DFCB7F9D7FC2EF9B5" bold="true" box="[828,906,1555,1576]" pageId="13" pageNumber="160">
|
||
<taxonomicNameLabel id="A26C57E942112E4DFCB7F9D7FC2EF9B5" box="[828,906,1555,1576]" pageId="13" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, ♀ paratype, Chendebji Chhu, Bhutan. A – genital field; B – palp, medial view; C – I-L-5 and -6. Scale bar = 100 µm
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B94368042112E4DFD98F971FA81F88F" blockId="13.[491,1497,1717,2017]" pageId="13" pageNumber="160">
|
||
<emphasis id="B95FEA9242112E4DFD98F971FDC3F950" box="[531,615,1717,1741]" italics="true" pageId="13" pageNumber="160">Female</emphasis>
|
||
– Idiosoma L 713, W 550; maximum diameter Dgl-4, 22. Coxal shield L 378; Cx-III W 447; Cx-I+II mL 112, Cx-I+II lL 253. Genital field L/W 152/167, genital plates L 125-128, pregenital sclerite 49, gonopore L 118, L Ac-1-3: 55-56, 53-55, 46-47.
|
||
</paragraph>
|
||
<paragraph id="8B94368042112E4DFD98F8D9FDB4F8E7" blockId="13.[491,1497,1717,2017]" pageId="13" pageNumber="160">Palp – Total L 412; dL/H, dL/H ratio: P-1, 37/34, 1.07; P-2, 84/57, 1.48; P-3, 116/45, 2.55; P-4, 134/34, 3.91; P-5, 41/16, 2.6; L ratio P-2/P-4, 0.63. Gnathosoma vL 141, chelicera total L 247.</paragraph>
|
||
<paragraph id="8B94368042112E4DFD98F841FB0CF87C" blockId="13.[491,1497,1717,2017]" pageId="13" pageNumber="160">Legs – I-L-5 dL 281, vL 180, dL/vL ratio 1.57, maximum H 66, dL/maximum H 4.25, S-1 L 139, L/W ratio 11.1, S-2 L 106, L/W ratio 6.2, distance S-1-2, 44, dL ratio S-1/2, 1.31; I-L-6 dL 205, central H 20, dL/central H ratio 10.1; L I-L-5/6 ratio 1.37.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C331650B42122E4EFD98FF22FDFDFEBD" pageId="14" pageNumber="161" type="etymology">
|
||
<paragraph id="8B94368042122E4EFD98FF22FDFDFEBD" blockId="14.[491,1519,229,633]" pageId="14" pageNumber="161">
|
||
<emphasis id="B95FEA9242122E4EFD98FF22FD37FF63" bold="true" box="[531,659,230,254]" pageId="14" pageNumber="161">Etymology</emphasis>
|
||
— The new species is named after the country where the new species was collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C331650B42122E4EFD98FEEFFC9CFDA8" pageId="14" pageNumber="161" type="discussion">
|
||
<paragraph id="8B94368042122E4EFD98FEEFFC9CFDA8" blockId="14.[491,1519,229,633]" pageId="14" pageNumber="161">
|
||
<emphasis id="B95FEA9242122E4EFD98FEEFFD2FFEDE" bold="true" box="[531,651,299,323]" pageId="14" pageNumber="161">Discussion</emphasis>
|
||
— In regard to the striated integument and the shape of the palp in the male (P-2 with distoventral projection, ventral margin of P-4 projecting), the new species resembles species of the
|
||
<taxonomicName id="4C2B4D0342122E4EFD0DFEB4FCB9FE15" authorityName="Thor" authorityYear="1899" box="[646,797,368,392]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="14" pageNumber="161" phylum="Arthropoda" rank="species" species="nodipalpis">
|
||
<emphasis id="B95FEA9242122E4EFD0DFEB4FCB9FE15" box="[646,797,368,392]" italics="true" pageId="14" pageNumber="161">A. nodipalpis</emphasis>
|
||
</taxonomicName>
|
||
complex (see above, under discussion of
|
||
<taxonomicName id="4C2B4D0342122E4EFB52FEB4FAFAFE15" authority="PešićK & SmitK & GurungK, 2023" authorityName="PešićK & SmitK & GurungK" authorityYear="2023" box="[1241,1374,368,392]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="14" pageNumber="161" phylum="Arthropoda" rank="species" species="conflatus" status="sp. nov.">
|
||
<emphasis id="B95FEA9242122E4EFB52FEB4FAFAFE15" box="[1241,1374,368,392]" italics="true" pageId="14" pageNumber="161">A. conflatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95FEA9242122E4EFAEAFEB4FA18FE15" bold="true" box="[1377,1468,368,392]" pageId="14" pageNumber="161">
|
||
<taxonomicNameLabel id="A26C57E942122E4EFAEAFEB4FA18FE15" box="[1377,1468,368,392]" pageId="14" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
). However, males in the latter species complex differ from the new species by the shape of genital field with anterior and posterior indentations (anterior margin slightly concave, but without medial identation in
|
||
<taxonomicName id="4C2B4D0342122E4EFD4CFE13FCFAFE72" authority="PešićK & SmitK & GurungK, 2023" authorityName="PešićK & SmitK & GurungK" authorityYear="2023" box="[711,862,471,495]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="14" pageNumber="161" phylum="Arthropoda" rank="species" species="bhutanicus" status="sp. nov.">
|
||
<emphasis id="B95FEA9242122E4EFD4CFE13FCFAFE72" box="[711,862,471,495]" italics="true" pageId="14" pageNumber="161">A. bhutanicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95FEA9242122E4EFCEBFE1CFC1CFE6D" bold="true" box="[864,952,472,496]" pageId="14" pageNumber="161">
|
||
<taxonomicNameLabel id="A26C57E942122E4EFCEBFE1CFC1CFE6D" box="[864,952,472,496]" pageId="14" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
). As
|
||
<bibRefCitation id="EFBA4B7142122E4EFC7FFE13FB36FE72" author="Gerecke R." box="[1012,1170,471,495]" pageId="14" pageNumber="161" refId="ref7779" refString="Gerecke R. 2003. Water mites of the genus Atractides Koch, 1837 (Acari: Parasitengona: Hygrobatidae)" type="book" year="2003">Gerecke (2003)</bibRefCitation>
|
||
pointed out, the development of a ventrodistal protrusion of P-
|
||
<quantity id="4CD39B6542122E4EFCCCFE3EFCCAFD8F" box="[839,878,506,530]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="14" pageNumber="161" unit="in" value="2.0">2 in</quantity>
|
||
males probably has evolved independently in various, not closely related species groups.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C331650B42122E4EFD98FDFBFDD9FDE4" pageId="14" pageNumber="161" type="distribution">
|
||
<paragraph id="8B94368042122E4EFD98FDFBFDD9FDE4" blockId="14.[491,1519,229,633]" pageId="14" pageNumber="161">
|
||
<emphasis id="B95FEA9242122E4EFD98FDFBFD3AFDCA" bold="true" box="[531,670,575,599]" pageId="14" pageNumber="161">Distribution</emphasis>
|
||
—
|
||
<collectingCountry id="F33C761042122E4EFD41FDFBFCB9FDCA" box="[714,797,575,599]" name="Bhutan" pageId="14" pageNumber="161">Bhutan</collectingCountry>
|
||
; know from high-order streams at an elevation above
|
||
<quantity id="4CD39B6542122E4EFAF6FDFBFA6FFDCA" box="[1405,1483,575,599]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="14" pageNumber="161" unit="m" value="2400.0">2400 m</quantity>
|
||
(
|
||
<figureCitation id="13102A0542122E4EFE79FDA5FDD6FDE4" box="[498,626,609,633]" captionStart="Figure 12" captionStartId="19.[515,582,1953,1974]" captionTargetBox="[503,1486,221,1899]" captionTargetId="figure-2@19.[503,1486,221,1899]" captionTargetPageId="19" captionText="Figure 12 Photographs of selected sampling sites. A – Bipgang Chhu (locus typicusAof. indentatus sp. nov. and A.himalayicussp. nov., and sampling site ofA. garhwali); B – Chendebji Chhu (locus typicus of A. bhutanicussp. nov.); C – Dakpay Chhu (locus typicus ofA. mangdensissp. nov.)." pageId="14" pageNumber="161">Figure 12B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |