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<document id="E9DFB51027175AE2E1E6A97115C6762D" ID-DOI="10.11646/zootaxa.3887.2.6" ID-GBIF-Dataset="856e1d56-1b48-4e94-ae6d-671bfdeac333" ID-ISSN="1175-5326" ID-Zenodo-Dep="229415" ID-ZooBank="D7FDE4F1-B238-4DBF-BCC4-BAC719EF7E0A" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1460977280099" checkinUser="plazi" docAuthor="Görföl, Tamás, Csorba, Gábor, Eger, Judith L., Son, Nguyen Truong &amp; Francis, Charles M." docDate="2014" docId="038187E6FFB8F455FF6CF9E4FC76FB11" docLanguage="en" docName="zt03887p250.pdf" docOrigin="Zootaxa 3887 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hypsugo dolichodon Görföl et al. 2014, n. sp." docType="treatment" docVersion="5" lastPageNumber="247" masterDocId="FFB8FF9EFFBAF45DFFFBFFB6FF89FFBA" masterDocTitle="Canines make the difference: a new species of Hypsugo (Chiroptera: Vespertilionidae) from Laos and Vietnam" masterLastPageNumber="250" masterPageNumber="239" pageNumber="241" updateTime="1731503338885" updateUser="jonas">
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<mods:title id="5351068E55BAC3F58F90623090FEC9D2">Canines make the difference: a new species of Hypsugo (Chiroptera: Vespertilionidae) from Laos and Vietnam</mods:title>
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<mods:namePart id="378E8F952969D7D46BA6CBC1EA259F03">Görföl, Tamás</mods:namePart>
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<mods:namePart id="27B0A30C96A117FF3087CFD145EF29F5">Csorba, Gábor</mods:namePart>
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<mods:namePart id="B6226510485A4F4FCBDC54B651E8A710">Eger, Judith L.</mods:namePart>
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<mods:namePart id="FFFEE3D11AF9C27647FD5B56E67C4131">Son, Nguyen Truong</mods:namePart>
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<treatment id="038187E6FFB8F455FF6CF9E4FC76FB11" LSID="urn:lsid:plazi:treatment:038187E6FFB8F455FF6CF9E4FC76FB11" httpUri="http://treatment.plazi.org/id/038187E6FFB8F455FF6CF9E4FC76FB11" lastPageId="8" lastPageNumber="247" pageId="2" pageNumber="241">
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<heading id="D0DF819CFFB8F45FFF6CF9E4FE53F9D7" bold="true" box="[151,474,1618,1645]" fontSize="11" level="1" pageId="2" pageNumber="241" reason="1">
<emphasis id="B95CEAE2FFB8F45FFF6CF9E4FE53F9D7" bold="true" box="[151,474,1618,1645]" pageId="2" pageNumber="241">
<taxonomicName id="4C284D73FFB8F45FFF6CF9E4FE18F9D6" ID-CoL="8LVDV" authority="Görföl et al., 2014" authorityName="Görföl et al." authorityYear="2014" box="[151,401,1618,1644]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="2" pageNumber="241" phylum="Chordata" rank="species" species="dolichodon" status="sp. nov.">
<emphasis id="B95CEAE2FFB8F45FFF6CF9E4FE18F9D6" bold="true" box="[151,401,1618,1644]" italics="true" pageId="2" pageNumber="241">Hypsugo dolichodon</emphasis>
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<taxonomicNameLabel id="A26F5799FFB8F45FFE63F9E5FE53F9D7" box="[408,474,1619,1645]" pageId="2" pageNumber="241" rank="species">n. sp.</taxonomicNameLabel>
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<figureCitation id="13132A75FFB8F45FFF6CF9C3FE89F934" box="[151,256,1653,1678]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="3.[151,250,1908,1930]" captionStartId-1="5.[151,250,782,804]" captionStartId-2="5.[151,250,1497,1519]" captionStartId-3="6.[151,250,1936,1958]" captionStartId-4="7.[151,250,1146,1168]" captionTargetBox-0="[350,1229,484,1885]" captionTargetBox-1="[427,1159,184,760]" captionTargetBox-2="[171,1408,853,1481]" captionTargetBox-3="[151,1436,193,1914]" captionTargetBox-4="[157,1429,193,1115]" captionTargetId-0="figure@3.[340,1248,481,1886]" captionTargetId-1="figure@5.[427,1160,184,760]" captionTargetId-2="figure@5.[171,1409,853,1481]" captionTargetId-3="figure@6.[151,1436,193,1914]" captionTargetId-4="figure@7.[151,1436,193,1125]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURE 1. Lateral view of skulls of (a) H. pulveratus BM (NH) 94.9. 1.18 from Fujian, China (b) H. dolichodon n. sp. ROM 110459 (holotype) and (c) H. cf. pulveratus IEBR M- 1610 from Vietnam. Scale = 5 mm" captionText-1="FIGURE 2. Portrait of a living specimen of H. dolichodon n. sp. (ROM 110807, from Vietnam)." captionText-2="FIGURE 3. Dorsal (left) and ventral (right) hairs of (a) H. pulveratus BM (NH) 70.7. 18.12 (holotype) and (b) H. dolichodon n. sp. ROM 110459 (holotype)." captionText-3="FIGURE 4. Dorsal, lateral, and ventral views of skulls and occlusal and lateral views of mandibles of H. pulveratus ROM 110653, from Laos (left) and H. dolichodon n. sp. ROM 110462, paratype (right). Scale = 5 mm." captionText-4="FIGURE 5. Occlusal view of left upper (left side) and right lower (right side) dentition of (a) H. pulveratus BM (NH) 70.7. 18.12 (holotype) and (b) H. dolichodon n. sp. ROM 110459 (holotype). Scale = 3 mm" httpUri-0="https://zenodo.org/record/229416/files/figure.png" httpUri-1="https://zenodo.org/record/229417/files/figure.png" httpUri-2="https://zenodo.org/record/229418/files/figure.png" httpUri-3="https://zenodo.org/record/229419/files/figure.png" httpUri-4="https://zenodo.org/record/229420/files/figure.png" pageId="2" pageNumber="241">Figs. 15</figureCitation>
</paragraph>
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<treatmentCitationGroup id="AB3811DEFFB8F45FFF6CF90EFDCCF977" box="[151,581,1719,1741]" pageId="2" pageNumber="241">
<taxonomicName id="4C284D73FFB8F45FFF6CF90EFF78F977" ID-CoL="5348" box="[151,241,1720,1741]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="2" pageNumber="241" phylum="Chordata" rank="genus">
<emphasis id="B95CEAE2FFB8F45FFF6CF90EFF78F977" box="[151,241,1720,1741]" italics="true" pageId="2" pageNumber="241">Hypsugo</emphasis>
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CMF sp.A:
<treatmentCitation id="0A8910E1FFB8F45FFE94F901FDCCF977" author="Francis" box="[367,581,1719,1741]" page="7" pageId="2" pageNumber="241" year="2010">
<bibRefCitation id="EFB94B01FFB8F45FFE94F901FDCCF977" author="Francis" box="[367,581,1719,1741]" etAl="et al." firstAuthor="Francis" page="7" pageId="2" pageNumber="241" pagination="12575" refId="ref5760" refString="Francis, C. M., Borisenko, A. V., Ivanova, N. V., Eger, J. L., Lim, B. K., Guillen-Servent, A., Kruskop, S. V., Mackie, I. &amp; Hebert, P. D. N. (2010) The role of DNA barcodes in understanding and conservation of mammal diversity in Southeast Asia. PLoS ONE, 5, e 12575." type="journal article" year="2010">
Francis
<emphasis id="B95CEAE2FFB8F45FFE3BF90EFE7AF977" box="[448,499,1719,1741]" italics="true" pageId="2" pageNumber="241">et al.</emphasis>
2010: 7
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</paragraph>
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<subSubSection id="C332657BFFB8F45EFF6CF94FFDE9FF6F" lastPageId="3" lastPageNumber="242" pageId="2" pageNumber="241" type="materials_examined">
<paragraph id="8B9736F0FFB8F45FFF6CF94FFCB2F88C" blockId="2.[151,1437,1785,2029]" pageId="2" pageNumber="241">
<materialsCitation id="3B403CADFFB8F45FFF6CF94FFCB2F88C" collectingDate="1997-05-05" collectionCode="ROM" collectorName="Charles M. Francis &amp; Antonio Guillen" country="Laos" elevation="150" latitude="14.95" location="Xe Kaman" longLatPrecision="1286" longitude="107.13333" municipality="Ban Paam" pageId="2" pageNumber="240" specimenCode="ROM 110459" specimenCount="1" specimenCount-female="1" stateProvince="Attapu Province" typeStatus="holotype">
<emphasis id="B95CEAE2FFB8F45FFF6CF94FFE82F8A8" bold="true" box="[151,267,1785,1810]" pageId="2" pageNumber="241">
<typeStatus id="54938852FFB8F45FFF6CF94FFE8FF8A8" box="[151,262,1785,1810]" pageId="2" pageNumber="241" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DB8E9E8BFFB8F45FFEE8F94CFE26F8A8" box="[275,431,1785,1810]" pageId="2" pageNumber="241">ROM 110459</specimenCode>
(collector number # 970505.05), immature
<specimenCount id="9D2EFD79FFB8F45FFC5FF94FFC7AF8A8" box="[932,1011,1785,1810]" count="1" pageId="2" pageNumber="241" type="female">female</specimenCount>
, in alcohol, skull removed. Collected by
<collectorName id="26DD5326FFB8F45FFF41F8ABFE19F88C" box="[186,400,1821,1846]" pageId="2" pageNumber="241">Charles M. Francis</collectorName>
and
<collectorName id="26DD5326FFB8F45FFE3DF8ABFDF4F88C" box="[454,637,1821,1846]" pageId="2" pageNumber="241">Antonio Guillén</collectorName>
on 5 May, 1997.
</materialsCitation>
</paragraph>
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<emphasis id="B95CEAE2FFB8F45FFF3CF8F7FEEDF8E0" bold="true" box="[199,356,1857,1882]" pageId="2" pageNumber="241">Type locality</emphasis>
.
<geoCoordinate id="EE1C5037FFB8F45FFE82F8F7FE55F8E0" box="[377,476,1857,1882]" degrees="14" direction="north" minutes="57" orientation="latitude" pageId="2" pageNumber="241" precision="925" value="14.95">14°57N</geoCoordinate>
,
<geoCoordinate id="EE1C5037FFB8F45FFE15F8F4FDD5F8E0" box="[494,604,1858,1882]" degrees="107" direction="east" minutes="08" orientation="longitude" pageId="2" pageNumber="241" precision="925" value="107.13333">107°08E</geoCoordinate>
,
<collectingMunicipality id="6BF3AC8AFFB8F45FFD95F8F4FD67F8E3" box="[622,750,1858,1881]" pageId="2" pageNumber="241">Xe Kaman</collectingMunicipality>
proposed dam site,
<quantity id="4CD09B15FFB8F45FFC12F8F4FBBEF8E0" box="[1001,1079,1857,1882]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="241" unit="km" value="10.0">
<locationDeviation id="9A252FD2FFB8F45FFC12F8F4FBBEF8E0" box="[1001,1079,1857,1882]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="241" unit="km" value="10.0">10 km</locationDeviation>
</quantity>
east from
<collectingMunicipality id="6BF3AC8AFFB8F45FFB39F8F4FAB7F8E3" box="[1218,1342,1858,1881]" pageId="2" pageNumber="241">Ban Paam</collectingMunicipality>
,
<collectingRegion id="49ECF812FFB8F45FFAABF8F4FF77F8C4" pageId="2" pageNumber="241">Attapu Province</collectingRegion>
,
<collectingCountry id="F33F7660FFB8F45FFEF3F8D0FEBDF8C7" box="[264,308,1894,1917]" name="Laos" pageId="2" pageNumber="241" value="Laos">Lao</collectingCountry>
PDR,
<quantity id="4CD09B15FFB8F45FFE84F8D3FE4FF8C7" box="[383,454,1893,1918]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="241" unit="m" value="150.0">
<elevation id="0005D1C3FFB8F45FFE84F8D3FE4FF8C7" box="[383,454,1893,1918]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="241" unit="m" value="150.0">150 m</elevation>
</quantity>
a.s.l.
</paragraph>
<paragraph id="8B9736F0FFB8F45FFF3CF83FFD76F857" blockId="2.[151,1437,1785,2029]" pageId="2" pageNumber="241">
<materialsCitation id="3B403CADFFB8F45FFF3CF83FFADDF818" box="[199,1364,1929,1954]" collectingDate="1997-05-06" collectionCode="ROM" country="Laos" elevation="150" latitude="14.966666" location="Xe Kaman" longLatPrecision="1295" longitude="107.21667" municipality="Ban Paam" pageId="2" pageNumber="240" specimenCode="ROM 110462" specimenCount="1" specimenCount-male="1" stateProvince="Attapu Province" typeStatus="paratype">
<typeStatus id="54938852FFB8F45FFF3CF83FFEB6F818" box="[199,319,1929,1954]" pageId="2" pageNumber="241" type="paratype">
<emphasis id="B95CEAE2FFB8F45FFF3CF83FFEB6F818" bold="true" box="[199,319,1929,1954]" pageId="2" pageNumber="241">Paratypes</emphasis>
</typeStatus>
.
<specimenCode id="DB8E9E8BFFB8F45FFEB6F83CFE61F818" box="[333,488,1929,1954]" pageId="2" pageNumber="241">
<collectionCode id="ED39AE35FFB8F45FFEB6F83CFE04F818" box="[333,397,1930,1954]" collectionName="Canada Entomology Department, Royal Ontario Museum" pageId="2" pageNumber="241">ROM</collectionCode>
110462
</specimenCode>
(# 970506.12), immature
<specimenCount id="9D2EFD79FFB8F45FFCEBF83FFCC0F818" box="[784,841,1929,1954]" count="1" pageId="2" pageNumber="241" type="male">male</specimenCount>
, collected at the
<typeStatus id="54938852FFB8F45FFBF3F83CFBB1F818" box="[1032,1080,1930,1954]" pageId="2" pageNumber="241">type</typeStatus>
locality on
<date id="FF961030FFB8F45FFB44F83FFA83F818" box="[1215,1290,1929,1954]" pageId="2" pageNumber="241">6 May</date>
, 1997;
</materialsCitation>
<materialsCitation id="3B403CADFFB8F45FFAA6F83CFD72F857" collectingDate="1997-05-06" collectionCode="ROM" country="Laos" elevation="150" latitude="14.96666667" location="Xe Kaman" longLatPrecision="1286" longitude="107.21666667" municipality="Xe Kaman" pageId="2" pageNumber="241" specimenCode="ROM 110464" specimenCount="1" specimenCount-female="1" stateProvince="Attapu Province">
<specimenCode id="DB8E9E8BFFB8F45FFAA6F83CFF63F873" pageId="2" pageNumber="241">ROM 110464</specimenCode>
(# 970508.21), immature
<specimenCount id="9D2EFD79FFB8F45FFDEEF806FDEDF873" box="[533,612,1968,1993]" pageId="2" pageNumber="241" type="female">female</specimenCount>
, collected at
<geoCoordinate id="EE1C5037FFB8F45FFD00F807FCDEF873" box="[763,855,1964,1993]" degrees="14" direction="north" minutes="58" orientation="latitude" pageId="2" pageNumber="241" precision="925" value="14.966666">14°58N</geoCoordinate>
<geoCoordinate id="EE1C5037FFB8F45FFCA4F807FC41F873" box="[863,968,1964,1993]" degrees="107" direction="east" minutes="13" orientation="longitude" pageId="2" pageNumber="241" precision="925" value="107.21667">107°13E</geoCoordinate>
,
<collectingRegion id="49ECF812FFB8F45FFC2EF807FB18F873" box="[981,1169,1968,1993]" pageId="2" pageNumber="241">Attapu Province</collectingRegion>
,
<collectingCountry id="F33F7660FFB8F45FFB66F807FB40F872" box="[1181,1225,1969,1992]" name="Laos" pageId="2" pageNumber="241" value="Laos">Lao</collectingCountry>
PDR,
<quantity id="4CD09B15FFB8F45FFAECF806FAE9F872" box="[1303,1376,1968,1993]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="241" unit="m" value="150.0">
<elevation id="0005D1C3FFB8F45FFAECF806FAE9F872" box="[1303,1376,1968,1993]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="241" unit="m" value="150.0">150 m</elevation>
</quantity>
a.s.l. on
<date id="FF961030FFB8F45FFF41F863FE8DF857" box="[186,260,2005,2029]" pageId="2" pageNumber="241">8 May</date>
, 1997. All bodies in alcohol, skulls removed
</materialsCitation>
.
</paragraph>
<paragraph id="8B9736F0FFB9F45EFF3CFF21FDE9FF6F" blockId="3.[151,1437,151,429]" pageId="3" pageNumber="242">
<materialsCitation id="3B403CADFFB9F45EFF3CFF21FDE9FF6F" collectionCode="ROM" country="Vietnam" elevation="100" latitude="11.416667" location="Cat Tien National Park" longLatPrecision="1295" longitude="107.433334" pageId="3" pageNumber="242" specimenCode="ROM 110807" specimenCount="2" specimenCount-adult="1" specimenCount-female="1" stateProvince="Dong Nai">
<emphasis id="B95CEAE2FFB9F45EFF3CFF21FE28FF0A" bold="true" box="[199,417,151,176]" pageId="3" pageNumber="242">Referred material</emphasis>
.
<collectingCountry id="F33F7660FFB9F45EFE49FF2EFDB1FF0A" box="[434,568,152,176]" name="Vietnam" pageId="3" pageNumber="242" value="Vietnam">VIETNAM</collectingCountry>
:
<specimenCode id="DB8E9E8BFFB9F45EFDBEFF2EFD6CFF0A" box="[581,741,152,176]" pageId="3" pageNumber="242">ROM 110807</specimenCode>
,
<specimenCount id="9D2EFD79FFB9F45EFD0FFF21FCA5FF0A" box="[756,812,151,176]" count="1" pageId="3" pageNumber="242" type="adult">adult</specimenCount>
<specimenCount id="9D2EFD79FFB9F45EFCCDFF21FC0CFF0A" box="[822,901,151,176]" count="1" pageId="3" pageNumber="242" type="female">female</specimenCount>
, dry skin and skull,
<geoCoordinate id="EE1C5037FFB9F45EFB8CFF21FB50FF0A" box="[1143,1241,151,176]" degrees="11" direction="north" minutes="25" orientation="latitude" pageId="3" pageNumber="242" precision="925" value="11.416667">11°25N</geoCoordinate>
,
<geoCoordinate id="EE1C5037FFB9F45EFB13FF21FADAFF0A" box="[1256,1363,151,176]" degrees="107" direction="east" minutes="26" orientation="longitude" pageId="3" pageNumber="242" precision="925" value="107.433334">107°26E</geoCoordinate>
<collectingRegion id="49ECF812FFB9F45EFAA5FF2EFF4BFF6F" country="Vietnam" name="Dong Nai" pageId="3" pageNumber="242">Dong Nai</collectingRegion>
,
<location id="8EF7602BFFB9F45EFF36FF0AFE5BFF6F" LSID="urn:lsid:plazi:treatment:038187E6FFB8F455FF6CF9E4FC76FB11:8EF7602BFFB9F45EFF36FF0AFE5BFF6F" box="[205,466,188,213]" country="Vietnam" latitude="11.416667" longLatPrecision="1295" longitude="107.433334" name="Cat Tien National Park" pageId="3" pageNumber="242" stateProvince="Dong Nai">Cat Tien National Park</location>
,
<quantity id="4CD09B15FFB9F45EFE26FF0AFDADFF6E" box="[477,548,188,212]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="242" unit="m" value="100.0">
<elevation id="0005D1C3FFB9F45EFE26FF0AFDADFF6E" box="[477,548,188,212]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="242" unit="m" value="100.0">100 m</elevation>
</quantity>
a.s.l.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C332657BFFB9F45EFF3CFF69FE39FEFA" pageId="3" pageNumber="242" type="etymology">
<paragraph id="8B9736F0FFB9F45EFF3CFF69FE39FEFA" blockId="3.[151,1437,151,429]" pageId="3" pageNumber="242">
<emphasis id="B95CEAE2FFB9F45EFF3CFF69FEC7FF42" bold="true" box="[199,334,223,248]" pageId="3" pageNumber="242">Etymology.</emphasis>
From the Greek words “dolichos” (= long, enlarged) and “oodontos” (= tooth) referring to the long upper canine of the new species which separates it from SE Asian congeners. The proposed English name is Long-toothed Pipistrelle.
</paragraph>
</subSubSection>
<subSubSection id="C332657BFFB9F45EFF3CFEFAFEBCFE16" pageId="3" pageNumber="242" type="diagnosis">
<paragraph id="8B9736F0FFB9F45EFF3CFEFAFEBCFE16" blockId="3.[151,1437,151,429]" pageId="3" pageNumber="242">
<emphasis id="B95CEAE2FFB9F45EFF3CFEFAFEC8FEDF" bold="true" box="[199,321,332,357]" pageId="3" pageNumber="242">Diagnosis.</emphasis>
A medium-sized representative of
<taxonomicName id="4C284D73FFB9F45EFD19FEFBFCCFFEDE" box="[738,838,333,356]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="3" pageNumber="242" phylum="Chordata" rank="genus">
<emphasis id="B95CEAE2FFB9F45EFD19FEFBFCCFFEDE" box="[738,838,333,356]" italics="true" pageId="3" pageNumber="242">Hypsugo</emphasis>
</taxonomicName>
(FA
<quantity id="4CD09B15FFB9F45EFC73FEFAFBA4FEDE" box="[904,1069,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="3.6799999999999997" metricValueMax="3.84" metricValueMin="3.52" pageId="3" pageNumber="242" unit="mm" value="36.8" valueMax="38.4" valueMin="35.2">35.2-38.4 mm</quantity>
, CCL
<quantity id="4CD09B15FFB9F45EFB84FEFAFAB6FEDE" box="[1151,1343,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="1.2879999999999998" metricValueMax="1.331" metricValueMin="1.2449999999999999" pageId="3" pageNumber="242" unit="mm" value="12.879999999999999" valueMax="13.31" valueMin="12.45">12.45-13.31 mm</quantity>
) with a relatively robust skull, almost flat cranial profile and well-developed upper canines which are almost twice as high as P4 (
<figureCitation id="13132A75FFB9F45EFF19FE22FEAAFE16" box="[226,291,404,429]" captionStart="FIGURE 1" captionStartId="3.[151,250,1908,1930]" captionTargetBox="[350,1229,484,1885]" captionTargetId="figure@3.[340,1248,481,1886]" captionTargetPageId="3" captionText="FIGURE 1. Lateral view of skulls of (a) H. pulveratus BM (NH) 94.9. 1.18 from Fujian, China (b) H. dolichodon n. sp. ROM 110459 (holotype) and (c) H. cf. pulveratus IEBR M- 1610 from Vietnam. Scale = 5 mm" httpUri="https://zenodo.org/record/229416/files/figure.png" pageId="3" pageNumber="242">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF576678FFB9F45EFF6CF8C2FB84F812" httpUri="https://zenodo.org/record/229416/files/figure.png" pageId="3" pageNumber="242" targetBox="[350,1229,484,1885]" targetPageId="3">
<paragraph id="8B9736F0FFB9F45EFF6CF8C2FB84F812" blockId="3.[151,1436,1908,1960]" pageId="3" pageNumber="242">
<emphasis id="B95CEAE2FFB9F45EFF6CF8C2FE84F833" bold="true" box="[151,269,1908,1930]" pageId="3" pageNumber="242">FIGURE 1</emphasis>
. Lateral view of skulls of (a)
<taxonomicName id="4C284D73FFB9F45EFDC4F8C3FD43F830" box="[575,714,1908,1930]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="3" pageNumber="242" phylum="Chordata" rank="species" species="pulveratus">
<emphasis id="B95CEAE2FFB9F45EFDC4F8C3FD43F830" box="[575,714,1908,1930]" italics="true" pageId="3" pageNumber="242">H. pulveratus</emphasis>
</taxonomicName>
BM(NH) 94.9.1.18 from Fujian, China (b)
<taxonomicName id="4C284D73FFB9F45EFB72F8C3FA93F830" box="[1161,1306,1908,1930]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="3" pageNumber="242" phylum="Chordata" rank="species" species="dolichodon" status="sp. nov.">
<emphasis id="B95CEAE2FFB9F45EFB72F8C3FA93F830" box="[1161,1306,1908,1930]" italics="true" pageId="3" pageNumber="242">H. dolichodon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F5799FFB9F45EFADAF8C2FAD2F833" box="[1313,1371,1908,1929]" pageId="3" pageNumber="242" rank="species">
<emphasis id="B95CEAE2FFB9F45EFADAF8C2FAD2F833" bold="true" box="[1313,1371,1908,1929]" pageId="3" pageNumber="242">n. sp.</emphasis>
</taxonomicNameLabel>
ROM 110459 (holotype) and (c)
<taxonomicName id="4C284D73FFB9F45EFE5EF825FDC6F812" box="[421,591,1938,1960]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="3" pageNumber="242" phylum="Chordata" rank="species" species="pulveratus">
<emphasis id="B95CEAE2FFB9F45EFE5EF825FE36F812" box="[421,447,1939,1960]" italics="true" pageId="3" pageNumber="242">H.</emphasis>
cf.
<emphasis id="B95CEAE2FFB9F45EFE1FF824FDC6F812" box="[484,591,1938,1960]" italics="true" pageId="3" pageNumber="242">pulveratus</emphasis>
</taxonomicName>
IEBR M-1610 from Vietnam. Scale= 5 mm
</paragraph>
</caption>
<subSubSection id="C332657BFFBEF458FF3CFF21FDC8F84C" lastPageId="5" lastPageNumber="244" pageId="4" pageNumber="243" type="description">
<paragraph id="8B9736F0FFBEF459FF3CFF21FDC9FEDF" blockId="4.[151,1437,151,357]" pageId="4" pageNumber="243">
<emphasis id="B95CEAE2FFBEF459FF3CFF21FED0FF0A" bold="true" box="[199,345,151,176]" pageId="4" pageNumber="243">Description.</emphasis>
Medium-sized
<taxonomicName id="4C284D73FFBEF459FDF0FF2FFDE6FF0A" box="[523,623,153,176]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="243" phylum="Chordata" rank="genus">
<emphasis id="B95CEAE2FFBEF459FDF0FF2FFDE6FF0A" box="[523,623,153,176]" italics="true" pageId="4" pageNumber="243">Hypsugo</emphasis>
</taxonomicName>
(
<tableCitation id="C6AA034BFFBEF459FD85FF21FD5AFF0A" box="[638,723,151,176]" captionStart="TABLE 1" captionStartId="4.[151,239,404,427]" captionTargetBox="[159,1372,486,1948]" captionTargetPageId="4" captionText="TABLE 1. Selected external and craniodental measurements (in mm) of H. pulveratus and H. dolichodon n. sp. Values are given as mean, ± SD (n ≥ 2), min-max, (n). Acronyms and definitions for measurements are given in the text." httpUri="http://table.plazi.org/id/DF576678FFBEF459FF6CFE22FAC4FE70" pageId="4" pageNumber="243" tableUuid="DF576678FFBEF459FF6CFE22FAC4FE70">Table 1</tableCitation>
). The colour of the wings and other skin parts is nearly black. The ears are relatively short and wide, somewhat rounded with a short, squarish tragus (
<figureCitation id="13132A75FFBEF459FB63FF0AFB68FF6E" box="[1176,1249,188,213]" captionStart="FIGURE 2" captionStartId="5.[151,250,782,804]" captionTargetBox="[427,1159,184,760]" captionTargetId="figure@5.[427,1160,184,760]" captionTargetPageId="5" captionText="FIGURE 2. Portrait of a living specimen of H. dolichodon n. sp. (ROM 110807, from Vietnam)." httpUri="https://zenodo.org/record/229417/files/figure.png" pageId="4" pageNumber="243">Fig. 2</figureCitation>
). The dorsal fur (based on colour photographs and the dry study skin of the
<collectionCode id="ED39AE35FFBEF459FCC3FF56FCF1FF42" LSID="urn:lsid:biocol.org:col:34778" box="[824,888,224,248]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="4" pageNumber="243">ROM</collectionCode>
<date id="FF961030FFBEF459FC84FF56FC5BFF42" box="[895,978,224,248]" pageId="4" pageNumber="243" value="07-11-08">110807</date>
specimen) is blackish brown, with paler tips. On the ventrum the hairs have two bands of colour: the basal half is blackish brown and the distal half dirty white (
<figureCitation id="13132A75FFBEF459FF11FE91FEBCFEFA" box="[234,309,295,320]" captionStart="FIGURE 3" captionStartId="5.[151,250,1497,1519]" captionTargetBox="[171,1408,853,1481]" captionTargetId="figure@5.[171,1409,853,1481]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal (left) and ventral (right) hairs of (a) H. pulveratus BM (NH) 70.7. 18.12 (holotype) and (b) H. dolichodon n. sp. ROM 110459 (holotype)." httpUri="https://zenodo.org/record/229418/files/figure.png" pageId="4" pageNumber="243">Fig. 3</figureCitation>
). The Epiblema is very low and elongated, the calcar extends to half of the free edge of the uropatagium and is poorly developed.
</paragraph>
<caption id="DF576678FFBEF459FF6CFE22FAC4FE70" ID-Table-UUID="DF576678FFBEF459FF6CFE22FAC4FE70" httpUri="http://table.plazi.org/id/DF576678FFBEF459FF6CFE22FAC4FE70" pageId="4" pageNumber="243" startId="4.[151,239,404,427]" targetBox="[159,1372,486,1948]" targetIsTable="true" targetPageId="4">
<paragraph id="8B9736F0FFBEF459FF6CFE22FAC4FE70" blockId="4.[151,1433,403,459]" pageId="4" pageNumber="243">
<emphasis id="B95CEAE2FFBEF459FF6CFE22FE8DFE11" bold="true" box="[151,260,404,427]" pageId="4" pageNumber="243">TABLE 1</emphasis>
. Selected external and craniodental measurements (in mm) of
<taxonomicName id="4C284D73FFBEF459FC53FE22FBB0FE11" box="[936,1081,403,427]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="243" phylum="Chordata" rank="species" species="pulveratus">
<emphasis id="B95CEAE2FFBEF459FC53FE22FBB0FE11" box="[936,1081,403,427]" italics="true" pageId="4" pageNumber="243">H. pulveratus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C284D73FFBEF459FB94FE22FA80FE11" box="[1135,1289,403,427]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="243" phylum="Chordata" rank="species" species="dolichodon" status="sp. nov.">
<emphasis id="B95CEAE2FFBEF459FB94FE22FA80FE11" box="[1135,1289,403,427]" italics="true" pageId="4" pageNumber="243">H. dolichodon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F5799FFBEF459FAEBFE22FAC4FE11" box="[1296,1357,404,427]" pageId="4" pageNumber="243" rank="species">
<emphasis id="B95CEAE2FFBEF459FAEBFE22FAC4FE11" bold="true" box="[1296,1357,404,427]" pageId="4" pageNumber="243">n. sp.</emphasis>
</taxonomicNameLabel>
Values are given as mean, ± SD (n ≥ 2), min-max, (n). Acronyms and definitions for measurements are given in the text.
</paragraph>
</caption>
<paragraph id="8B9736F0FFBEF459FF64FE50FA90F826" pageId="4" pageNumber="243">
<table id="F928C450FFBE0BA2FF64FE50FAD5F826" box="[159,1372,486,1948]" gridcols="5" gridrows="21" pageId="4" pageNumber="243">
<tr id="351834B2FFBE0BA2FF64FE50FAD5FE47" box="[159,1372,486,509]" gridrow="0" pageId="4" pageNumber="243">
<th id="76C95DCEFFBE0BA2FF64FE50FF74FE47" box="[159,253,486,509]" gridcol="0" gridrow="0" pageId="4" pageNumber="243">character</th>
<th id="76C95DCEFFBE0BA2FEABFE50FD89FE47" box="[336,512,486,509]" gridcol="1" gridrow="0" pageId="4" pageNumber="243">
<taxonomicName id="4C284D73FFBEF459FEABFE5EFE53FE46" box="[336,474,486,509]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="243" phylum="Chordata" rank="species" species="pulveratus">
<emphasis id="B95CEAE2FFBEF459FEABFE5EFE53FE46" box="[336,474,486,509]" italics="true" pageId="4" pageNumber="243">H. pulveratus</emphasis>
</taxonomicName>
</th>
<th id="76C95DCEFFBE0BA2FD9FFE50FC9AFE47" box="[612,787,486,509]" gridcol="2" gridrow="0" pageId="4" pageNumber="243">
<taxonomicName id="4C284D73FFBEF459FD9FFE5EFD67FE46" box="[612,750,486,509]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="243" phylum="Chordata" rank="species" species="pulveratus">
<emphasis id="B95CEAE2FFBEF459FD9FFE5EFD67FE46" box="[612,750,486,509]" italics="true" pageId="4" pageNumber="243">H. pulveratus</emphasis>
</taxonomicName>
</th>
<th id="76C95DCEFFBE0BA2FC8CFE50FBAFFE47" box="[887,1062,486,509]" gridcol="3" gridrow="0" pageId="4" pageNumber="243">
<taxonomicName id="4C284D73FFBEF459FC8CFE5EFB88FE46" box="[887,1025,486,509]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="243" phylum="Chordata" rank="species" species="pulveratus">
<emphasis id="B95CEAE2FFBEF459FC8CFE5EFB88FE46" box="[887,1025,486,509]" italics="true" pageId="4" pageNumber="243">H. pulveratus</emphasis>
</taxonomicName>
</th>
<th id="76C95DCEFFBE0BA2FB69FE50FAD5FE47" box="[1170,1372,486,509]" gridcol="4" gridrow="0" pageId="4" pageNumber="243">
<taxonomicName id="4C284D73FFBEF459FB69FE5EFAABFE46" box="[1170,1314,486,509]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="243" phylum="Chordata" rank="species" species="dolichodon" status="sp. nov.">
<emphasis id="B95CEAE2FFBEF459FB69FE5EFAABFE46" box="[1170,1314,486,509]" italics="true" pageId="4" pageNumber="243">H. dolichodon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F5799FFBEF459FAD2FE51FAD5FE46" box="[1321,1372,487,508]" pageId="4" pageNumber="243" rank="species">
<emphasis id="B95CEAE2FFBEF459FAD2FE51FAD5FE46" bold="true" box="[1321,1372,487,508]" pageId="4" pageNumber="243">n.sp.</emphasis>
</taxonomicNameLabel>
</th>
</tr>
<tr id="351834B2FFBE0BA2FF64FDA6FAD5FDFE" box="[159,1372,528,580]" gridrow="1" pageId="4" pageNumber="243">
<th id="76C95DCEFFBE0BA2FF64FDA6FF74FDFE" box="[159,253,528,580]" gridcol="0" gridrow="1" pageId="4" pageNumber="243">FA</th>
<td id="76C95DCEFFBE0BA2FEABFDA6FD89FDFE" box="[336,512,528,580]" gridcol="1" gridrow="1" pageId="4" pageNumber="243">34.6 ± 1.39 32.437.6 (32)</td>
<td id="76C95DCEFFBE0BA2FD9FFDA6FC9AFDFE" box="[612,787,528,580]" gridcol="2" gridrow="1" pageId="4" pageNumber="243">34.2 ± 0.99 32.435.8 (19)</td>
<td id="76C95DCEFFBE0BA2FC8CFDA6FBAFFDFE" box="[887,1062,528,580]" gridcol="3" gridrow="1" pageId="4" pageNumber="243">35.4 ± 1.63 32.437.6 (11)</td>
<td id="76C95DCEFFBE0BA2FB69FDA6FAD5FDFE" box="[1170,1372,528,580]" gridcol="4" gridrow="1" pageId="4" pageNumber="243">37.0 ± 1.30 35.238.4 (4)</td>
</tr>
<tr id="351834B2FFBE0BA2FF64FDEEFAD5FD36" box="[159,1372,600,652]" gridrow="2" pageId="4" pageNumber="243">
<th id="76C95DCEFFBE0BA2FF64FDEEFF74FD36" box="[159,253,600,652]" gridcol="0" gridrow="2" pageId="4" pageNumber="243">TIB</th>
<td id="76C95DCEFFBE0BA2FEABFDEEFD89FD36" box="[336,512,600,652]" gridcol="1" gridrow="2" pageId="4" pageNumber="243">13.9 ± 0.53 12.914.7 (11)</td>
<td id="76C95DCEFFBE0BA2FD9FFDEEFC9AFD36" box="[612,787,600,652]" gridcol="2" gridrow="2" pageId="4" pageNumber="243">13.7 ± 0.42 13.014.1 (5)</td>
<td id="76C95DCEFFBE0BA2FC8CFDEEFBAFFD36" box="[887,1062,600,652]" gridcol="3" gridrow="2" pageId="4" pageNumber="243">14.0 ± 0.62 12.914.7 (6)</td>
<td id="76C95DCEFFBE0BA2FB69FDEEFAD5FD36" box="[1170,1372,600,652]" gridcol="4" gridrow="2" pageId="4" pageNumber="243">14.4 ± 0.20 14.214.5 (2)</td>
</tr>
<tr id="351834B2FFBE0BA2FF64FD16FAD5FD6E" box="[159,1372,672,724]" gridrow="3" pageId="4" pageNumber="243">
<th id="76C95DCEFFBE0BA2FF64FD16FF74FD6E" box="[159,253,672,724]" gridcol="0" gridrow="3" pageId="4" pageNumber="243">HF</th>
<td id="76C95DCEFFBE0BA2FEABFD16FD89FD6E" box="[336,512,672,724]" gridcol="1" gridrow="3" pageId="4" pageNumber="243">6.1 ± 0.79 4.78.2 (24)</td>
<td id="76C95DCEFFBE0BA2FD9FFD16FC9AFD6E" box="[612,787,672,724]" gridcol="2" gridrow="3" pageId="4" pageNumber="243">6.0 ± 0.81 4.78.0 (14)</td>
<td id="76C95DCEFFBE0BA2FC8CFD16FBAFFD6E" box="[887,1062,672,724]" gridcol="3" gridrow="3" pageId="4" pageNumber="243">6.3 ± 0.78 5.48.2 (10)</td>
<td id="76C95DCEFFBE0BA2FB69FD16FAD5FD6E" box="[1170,1372,672,724]" gridcol="4" gridrow="3" pageId="4" pageNumber="243">6.4 ± 0.50 5.97.0 (4)</td>
</tr>
<tr id="351834B2FFBE0BA2FF64FD5EFAD5FCA6" box="[159,1372,744,796]" gridrow="4" pageId="4" pageNumber="243">
<th id="76C95DCEFFBE0BA2FF64FD5EFF74FCA6" box="[159,253,744,796]" gridcol="0" gridrow="4" pageId="4" pageNumber="243">GLS-i</th>
<td id="76C95DCEFFBE0BA2FEABFD5EFD89FCA6" box="[336,512,744,796]" gridcol="1" gridrow="4" pageId="4" pageNumber="243">14.01 ± 0.38 13.2614.64 (26)</td>
<td id="76C95DCEFFBE0BA2FD9FFD5EFC9AFCA6" box="[612,787,744,796]" gridcol="2" gridrow="4" pageId="4" pageNumber="243">13.89 ± 0.38 13.2614.28 (11)</td>
<td id="76C95DCEFFBE0BA2FC8CFD5EFBAFFCA6" box="[887,1062,744,796]" gridcol="3" gridrow="4" pageId="4" pageNumber="243">14.12 ± 0.40 13.3214.64 (13)</td>
<td id="76C95DCEFFBE0BA2FB69FD5EFAD5FCA6" box="[1170,1372,744,796]" gridcol="4" gridrow="4" pageId="4" pageNumber="243">14.25 ± 0.43 13.8314.83 (4)</td>
</tr>
<tr id="351834B2FFBE0BA2FF64FC86FAD5FCDE" box="[159,1372,816,868]" gridrow="5" pageId="4" pageNumber="243">
<th id="76C95DCEFFBE0BA2FF64FC86FF74FCDE" box="[159,253,816,868]" gridcol="0" gridrow="5" pageId="4" pageNumber="243">GLS</th>
<td id="76C95DCEFFBE0BA2FEABFC86FD89FCDE" box="[336,512,816,868]" gridcol="1" gridrow="5" pageId="4" pageNumber="243">13.75 ± 0.25 13.2614.11 (26)</td>
<td id="76C95DCEFFBE0BA2FD9FFC86FC9AFCDE" box="[612,787,816,868]" gridcol="2" gridrow="5" pageId="4" pageNumber="243">13.66 ± 0.22 13.2613.94 (11)</td>
<td id="76C95DCEFFBE0BA2FC8CFC86FBAFFCDE" box="[887,1062,816,868]" gridcol="3" gridrow="5" pageId="4" pageNumber="243">13.81 ± 0.27 13.3214.11 (13)</td>
<td id="76C95DCEFFBE0BA2FB69FC86FAD5FCDE" box="[1170,1372,816,868]" gridcol="4" gridrow="5" pageId="4" pageNumber="243">13.76 ± 0.41 13.3014.29 (4)</td>
</tr>
<tr id="351834B2FFBE0BA2FF64FCCEFAD5FC16" box="[159,1372,888,940]" gridrow="6" pageId="4" pageNumber="243">
<th id="76C95DCEFFBE0BA2FF64FCCEFF74FC16" box="[159,253,888,940]" gridcol="0" gridrow="6" pageId="4" pageNumber="243">CCL</th>
<td id="76C95DCEFFBE0BA2FEABFCCEFD89FC16" box="[336,512,888,940]" gridcol="1" gridrow="6" pageId="4" pageNumber="243">12.63 ± 0.30 12.0013.20 (28)</td>
<td id="76C95DCEFFBE0BA2FD9FFCCEFC9AFC16" box="[612,787,888,940]" gridcol="2" gridrow="6" pageId="4" pageNumber="243">12.53 ± 0.22 12.1212.81 (12)</td>
<td id="76C95DCEFFBE0BA2FC8CFCCEFBAFFC16" box="[887,1062,888,940]" gridcol="3" gridrow="6" pageId="4" pageNumber="243">12.75 ± 0.31 12.0013.20 (13)</td>
<td id="76C95DCEFFBE0BA2FB69FCCEFAD5FC16" box="[1170,1372,888,940]" gridcol="4" gridrow="6" pageId="4" pageNumber="243">12.86 ± 0.36 12.4513.31 (4)</td>
</tr>
<tr id="351834B2FFBE0BA2FF64FC76FAD5FC4E" box="[159,1372,960,1012]" gridrow="7" pageId="4" pageNumber="243">
<th id="76C95DCEFFBE0BA2FF64FC76FF74FC4E" box="[159,253,960,1012]" gridcol="0" gridrow="7" pageId="4" pageNumber="243">C1C1W</th>
<td id="76C95DCEFFBE0BA2FEABFC76FD89FC4E" box="[336,512,960,1012]" gridcol="1" gridrow="7" pageId="4" pageNumber="243">4.22 ± 0.19 3.724.53 (30)</td>
<td id="76C95DCEFFBE0BA2FD9FFC76FC9AFC4E" box="[612,787,960,1012]" gridcol="2" gridrow="7" pageId="4" pageNumber="243">4.27 ± 0.18 4.004.53 (12)</td>
<td id="76C95DCEFFBE0BA2FC8CFC76FBAFFC4E" box="[887,1062,960,1012]" gridcol="3" gridrow="7" pageId="4" pageNumber="243">4.20 ± 0.20 3.724.42 (14)</td>
<td id="76C95DCEFFBE0BA2FB69FC76FAD5FC4E" box="[1170,1372,960,1012]" gridcol="4" gridrow="7" pageId="4" pageNumber="243">4.17 ± 0.19 4.004.40 (4)</td>
</tr>
<tr id="351834B2FFBE0BA2FF64FBBEFAD5FB86" box="[159,1372,1032,1084]" gridrow="8" pageId="4" pageNumber="243">
<th id="76C95DCEFFBE0BA2FF64FBBEFF74FB86" box="[159,253,1032,1084]" gridcol="0" gridrow="8" pageId="4" pageNumber="243">M3M3W</th>
<td id="76C95DCEFFBE0BA2FEABFBBEFD89FB86" box="[336,512,1032,1084]" gridcol="1" gridrow="8" pageId="4" pageNumber="243">5.73 ± 0.21 5.206.11 (29)</td>
<td id="76C95DCEFFBE0BA2FD9FFBBEFC9AFB86" box="[612,787,1032,1084]" gridcol="2" gridrow="8" pageId="4" pageNumber="243">5.69 ± 0.13 5.345.83 (11)</td>
<td id="76C95DCEFFBE0BA2FC8CFBBEFBAFFB86" box="[887,1062,1032,1084]" gridcol="3" gridrow="8" pageId="4" pageNumber="243">5.78 ± 0.26 5.206.11 (14)</td>
<td id="76C95DCEFFBE0BA2FB69FBBEFAD5FB86" box="[1170,1372,1032,1084]" gridcol="4" gridrow="8" pageId="4" pageNumber="243">5.77 ± 0.20 5.526.00 (4)</td>
</tr>
<tr id="351834B2FFBE0BA2FF64FBE6FAD5FB3E" box="[159,1372,1104,1156]" gridrow="9" pageId="4" pageNumber="243">
<th id="76C95DCEFFBE0BA2FF64FBE6FF74FB3E" box="[159,253,1104,1156]" gridcol="0" gridrow="9" pageId="4" pageNumber="243">RW</th>
<td id="76C95DCEFFBE0BA2FEABFBE6FD89FB3E" box="[336,512,1104,1156]" gridcol="1" gridrow="9" pageId="4" pageNumber="243">4.95 ± 0.22 4.535.45 (30)</td>
<td id="76C95DCEFFBE0BA2FD9FFBE6FC9AFB3E" box="[612,787,1104,1156]" gridcol="2" gridrow="9" pageId="4" pageNumber="243">4.95 ± 0.18 4.625.24 (12)</td>
<td id="76C95DCEFFBE0BA2FC8CFBE6FBAFFB3E" box="[887,1062,1104,1156]" gridcol="3" gridrow="9" pageId="4" pageNumber="243">4.95 ± 0.26 4.535.45 (14)</td>
<td id="76C95DCEFFBE0BA2FB69FBE6FAD5FB3E" box="[1170,1372,1104,1156]" gridcol="4" gridrow="9" pageId="4" pageNumber="243">5.20 ± 0.35 4.875.65 (4)</td>
</tr>
<tr id="351834B2FFBE0BA2FF64FB2EFAD5FB76" box="[159,1372,1176,1228]" gridrow="10" pageId="4" pageNumber="243">
<th id="76C95DCEFFBE0BA2FF64FB2EFF74FB76" box="[159,253,1176,1228]" gridcol="0" gridrow="10" pageId="4" pageNumber="243">IOW</th>
<td id="76C95DCEFFBE0BA2FEABFB2EFD89FB76" box="[336,512,1176,1228]" gridcol="1" gridrow="10" pageId="4" pageNumber="243">3.73 ± 0.15 3.333.99 (31)</td>
<td id="76C95DCEFFBE0BA2FD9FFB2EFC9AFB76" box="[612,787,1176,1228]" gridcol="2" gridrow="10" pageId="4" pageNumber="243">3.75 ± 0.14 3.503.97 (12)</td>
<td id="76C95DCEFFBE0BA2FC8CFB2EFBAFFB76" box="[887,1062,1176,1228]" gridcol="3" gridrow="10" pageId="4" pageNumber="243">3.70 ± 0.17 3.333.99 (15)</td>
<td id="76C95DCEFFBE0BA2FB69FB2EFAD5FB76" box="[1170,1372,1176,1228]" gridcol="4" gridrow="10" pageId="4" pageNumber="243">3.86 ± 0.06 3.773.90 (4)</td>
</tr>
<tr id="351834B2FFBE0BA2FF64FB56FAD5FAAE" box="[159,1372,1248,1300]" gridrow="11" pageId="4" pageNumber="243">
<th id="76C95DCEFFBE0BA2FF64FB56FF74FAAE" box="[159,253,1248,1300]" gridcol="0" gridrow="11" pageId="4" pageNumber="243">ZYW</th>
<td id="76C95DCEFFBE0BA2FEABFB56FD89FAAE" box="[336,512,1248,1300]" gridcol="1" gridrow="11" pageId="4" pageNumber="243">8.46 ± 0.18 8.158.80 (21)</td>
<td id="76C95DCEFFBE0BA2FD9FFB56FC9AFAAE" box="[612,787,1248,1300]" gridcol="2" gridrow="11" pageId="4" pageNumber="243">8.39 ± 0.15 8.168.62 (11)</td>
<td id="76C95DCEFFBE0BA2FC8CFB56FBAFFAAE" box="[887,1062,1248,1300]" gridcol="3" gridrow="11" pageId="4" pageNumber="243">8.52 ± 0.20 8.158.80 (8)</td>
<td id="76C95DCEFFBE0BA2FB69FB56FAD5FAAE" box="[1170,1372,1248,1300]" gridcol="4" gridrow="11" pageId="4" pageNumber="243">8.43 ± 0.35 8.188.68 (2)</td>
</tr>
<tr id="351834B2FFBE0BA2FF64FA9EFAD5FAE6" box="[159,1372,1320,1372]" gridrow="12" pageId="4" pageNumber="243">
<th id="76C95DCEFFBE0BA2FF64FA9EFF74FAE6" box="[159,253,1320,1372]" gridcol="0" gridrow="12" pageId="4" pageNumber="243">MAW</th>
<td id="76C95DCEFFBE0BA2FEABFA9EFD89FAE6" box="[336,512,1320,1372]" gridcol="1" gridrow="12" pageId="4" pageNumber="243">7.24 ± 0.13 6.947.56 (27)</td>
<td id="76C95DCEFFBE0BA2FD9FFA9EFC9AFAE6" box="[612,787,1320,1372]" gridcol="2" gridrow="12" pageId="4" pageNumber="243">7.23 ± 0.12 7.007.37 (12)</td>
<td id="76C95DCEFFBE0BA2FC8CFA9EFBAFFAE6" box="[887,1062,1320,1372]" gridcol="3" gridrow="12" pageId="4" pageNumber="243">7.27 ± 0.15 6.947.56 (12)</td>
<td id="76C95DCEFFBE0BA2FB69FA9EFAD5FAE6" box="[1170,1372,1320,1372]" gridcol="4" gridrow="12" pageId="4" pageNumber="243">7.66 ± 0.14 7.477.77 (4)</td>
</tr>
<tr id="351834B2FFBE0BA2FF64FAC6FAD5FA1E" box="[159,1372,1392,1444]" gridrow="13" pageId="4" pageNumber="243">
<th id="76C95DCEFFBE0BA2FF64FAC6FF74FA1E" box="[159,253,1392,1444]" gridcol="0" gridrow="13" pageId="4" pageNumber="243">BCW</th>
<td id="76C95DCEFFBE0BA2FEABFAC6FD89FA1E" box="[336,512,1392,1444]" gridcol="1" gridrow="13" pageId="4" pageNumber="243">6.75 ± 0.16 6.186.97 (27)</td>
<td id="76C95DCEFFBE0BA2FD9FFAC6FC9AFA1E" box="[612,787,1392,1444]" gridcol="2" gridrow="13" pageId="4" pageNumber="243">6.79 ± 0.12 6.586.97 (11)</td>
<td id="76C95DCEFFBE0BA2FC8CFAC6FBAFFA1E" box="[887,1062,1392,1444]" gridcol="3" gridrow="13" pageId="4" pageNumber="243">6.72 ± 0.20 6.186.96 (13)</td>
<td id="76C95DCEFFBE0BA2FB69FAC6FAD5FA1E" box="[1170,1372,1392,1444]" gridcol="4" gridrow="13" pageId="4" pageNumber="243">6.97 ± 0.17 6.797.12 (3)</td>
</tr>
<tr id="351834B2FFBE0BA2FF64FA0EFAD5FA56" box="[159,1372,1464,1516]" gridrow="14" pageId="4" pageNumber="243">
<th id="76C95DCEFFBE0BA2FF64FA0EFF74FA56" box="[159,253,1464,1516]" gridcol="0" gridrow="14" pageId="4" pageNumber="243">BCH</th>
<td id="76C95DCEFFBE0BA2FEABFA0EFD89FA56" box="[336,512,1464,1516]" gridcol="1" gridrow="14" pageId="4" pageNumber="243">4.81 ± 0.13 4.625.10 (25)</td>
<td id="76C95DCEFFBE0BA2FD9FFA0EFC9AFA56" box="[612,787,1464,1516]" gridcol="2" gridrow="14" pageId="4" pageNumber="243">4.78 ± 0.10 4.624.97 (10)</td>
<td id="76C95DCEFFBE0BA2FC8CFA0EFBAFFA56" box="[887,1062,1464,1516]" gridcol="3" gridrow="14" pageId="4" pageNumber="243">4.82 ± 0.15 4.625.10 (12)</td>
<td id="76C95DCEFFBE0BA2FB69FA0EFAD5FA56" box="[1170,1372,1464,1516]" gridcol="4" gridrow="14" pageId="4" pageNumber="243">4.84 ± 0.15 4.675.04 (4)</td>
</tr>
<tr id="351834B2FFBE0BA2FF64F9B6FAD5F98E" box="[159,1372,1536,1588]" gridrow="15" pageId="4" pageNumber="243">
<th id="76C95DCEFFBE0BA2FF64F9B6FF74F98E" box="[159,253,1536,1588]" gridcol="0" gridrow="15" pageId="4" pageNumber="243">AOB</th>
<td id="76C95DCEFFBE0BA2FEABF9B6FD89F98E" box="[336,512,1536,1588]" gridcol="1" gridrow="15" pageId="4" pageNumber="243">0.85 ± 0.19 0.541.17 (22)</td>
<td id="76C95DCEFFBE0BA2FD9FF9B6FC9AF98E" box="[612,787,1536,1588]" gridcol="2" gridrow="15" pageId="4" pageNumber="243">0.87 ± 0.14 0.761.10 (7)</td>
<td id="76C95DCEFFBE0BA2FC8CF9B6FBAFF98E" box="[887,1062,1536,1588]" gridcol="3" gridrow="15" pageId="4" pageNumber="243">0.87 ± 0.21 0.541.17 (13)</td>
<td id="76C95DCEFFBE0BA2FB69F9B6FAD5F98E" box="[1170,1372,1536,1588]" gridcol="4" gridrow="15" pageId="4" pageNumber="243">0.96 ± 0.07 0.921.06 (4)</td>
</tr>
<tr id="351834B2FFBE0BA2FF64F9FEFAD5F9C6" box="[159,1372,1608,1660]" gridrow="16" pageId="4" pageNumber="243">
<th id="76C95DCEFFBE0BA2FF64F9FEFF74F9C6" box="[159,253,1608,1660]" gridcol="0" gridrow="16" pageId="4" pageNumber="243">CM3L</th>
<td id="76C95DCEFFBE0BA2FEABF9FEFD89F9C6" box="[336,512,1608,1660]" gridcol="1" gridrow="16" pageId="4" pageNumber="243">4.88 ± 0.14 4.575.15 (33)</td>
<td id="76C95DCEFFBE0BA2FD9FF9FEFC9AF9C6" box="[612,787,1608,1660]" gridcol="2" gridrow="16" pageId="4" pageNumber="243">4.87 ± 0.14 4.625.11 (14)</td>
<td id="76C95DCEFFBE0BA2FC8CF9FEFBAFF9C6" box="[887,1062,1608,1660]" gridcol="3" gridrow="16" pageId="4" pageNumber="243">4.91 ± 0.14 4.645.15 (15)</td>
<td id="76C95DCEFFBE0BA2FB69F9FEFAD5F9C6" box="[1170,1372,1608,1660]" gridcol="4" gridrow="16" pageId="4" pageNumber="243">5.05 ± 0.15 4.845.19 (4)</td>
</tr>
<tr id="351834B2FFBE0BA2FF64F926FAD5F97E" box="[159,1372,1680,1732]" gridrow="17" pageId="4" pageNumber="243">
<th id="76C95DCEFFBE0BA2FF64F926FF74F97E" box="[159,253,1680,1732]" gridcol="0" gridrow="17" pageId="4" pageNumber="243">CP4L</th>
<td id="76C95DCEFFBE0BA2FEABF926FD89F97E" box="[336,512,1680,1732]" gridcol="1" gridrow="17" pageId="4" pageNumber="243">1.99 ± 0.12 1.662.23 (33)</td>
<td id="76C95DCEFFBE0BA2FD9FF926FC9AF97E" box="[612,787,1680,1732]" gridcol="2" gridrow="17" pageId="4" pageNumber="243">2.01 ± 0.11 1.812.17 (14)</td>
<td id="76C95DCEFFBE0BA2FC8CF926FBAFF97E" box="[887,1062,1680,1732]" gridcol="3" gridrow="17" pageId="4" pageNumber="243">1.97 ± 0.14 1.662.23 (15)</td>
<td id="76C95DCEFFBE0BA2FB69F926FAD5F97E" box="[1170,1372,1680,1732]" gridcol="4" gridrow="17" pageId="4" pageNumber="243">2.09 ± 0.20 1.802.24 (4)</td>
</tr>
<tr id="351834B2FFBE0BA2FF64F96EFAD5F8B6" box="[159,1372,1752,1804]" gridrow="18" pageId="4" pageNumber="243">
<th id="76C95DCEFFBE0BA2FF64F96EFF74F8B6" box="[159,253,1752,1804]" gridcol="0" gridrow="18" pageId="4" pageNumber="243">ML</th>
<td id="76C95DCEFFBE0BA2FEABF96EFD89F8B6" box="[336,512,1752,1804]" gridcol="1" gridrow="18" pageId="4" pageNumber="243">9.65 ± 0.21 9.159.96 (31)</td>
<td id="76C95DCEFFBE0BA2FD9FF96EFC9AF8B6" box="[612,787,1752,1804]" gridcol="2" gridrow="18" pageId="4" pageNumber="243">9.59 ± 0.18 9.299.92 (14)</td>
<td id="76C95DCEFFBE0BA2FC8CF96EFBAFF8B6" box="[887,1062,1752,1804]" gridcol="3" gridrow="18" pageId="4" pageNumber="243">9.75 ± 0.15 9.509.96 (14)</td>
<td id="76C95DCEFFBE0BA2FB69F96EFAD5F8B6" box="[1170,1372,1752,1804]" gridcol="4" gridrow="18" pageId="4" pageNumber="243">10.00 ± 0.28 9.5910.21 (4)</td>
</tr>
<tr id="351834B2FFBE0BA2FF64F896FAD5F8EE" box="[159,1372,1824,1876]" gridrow="19" pageId="4" pageNumber="243">
<th id="76C95DCEFFBE0BA2FF64F896FF74F8EE" box="[159,253,1824,1876]" gridcol="0" gridrow="19" pageId="4" pageNumber="243">cm3L</th>
<td id="76C95DCEFFBE0BA2FEABF896FD89F8EE" box="[336,512,1824,1876]" gridcol="1" gridrow="19" pageId="4" pageNumber="243">5.20 ± 0.14 4.865.47 (33)</td>
<td id="76C95DCEFFBE0BA2FD9FF896FC9AF8EE" box="[612,787,1824,1876]" gridcol="2" gridrow="19" pageId="4" pageNumber="243">5.19 ± 0.14 4.985.47 (14)</td>
<td id="76C95DCEFFBE0BA2FC8CF896FBAFF8EE" box="[887,1062,1824,1876]" gridcol="3" gridrow="19" pageId="4" pageNumber="243">5.24 ± 0.13 5.015.39 (15)</td>
<td id="76C95DCEFFBE0BA2FB69F896FAD5F8EE" box="[1170,1372,1824,1876]" gridcol="4" gridrow="19" pageId="4" pageNumber="243">5.49 ± 0.14 5.345.65 (4)</td>
</tr>
<tr id="351834B2FFBE0BA2FF64F8DEFAD5F826" box="[159,1372,1896,1948]" gridrow="20" pageId="4" pageNumber="243">
<th id="76C95DCEFFBE0BA2FF64F8DEFF74F826" box="[159,253,1896,1948]" gridcol="0" gridrow="20" pageId="4" pageNumber="243">CPH</th>
<td id="76C95DCEFFBE0BA2FEABF8DEFD89F826" box="[336,512,1896,1948]" gridcol="1" gridrow="20" pageId="4" pageNumber="243">2.75 ± 0.11 2.523.02 (33)</td>
<td id="76C95DCEFFBE0BA2FD9FF8DEFC9AF826" box="[612,787,1896,1948]" gridcol="2" gridrow="20" pageId="4" pageNumber="243">2.71 ± 0.09 2.522.85 (14)</td>
<td id="76C95DCEFFBE0BA2FC8CF8DEFBAFF826" box="[887,1062,1896,1948]" gridcol="3" gridrow="20" pageId="4" pageNumber="243">2.79 ± 0.13 2.583.02 (15)</td>
<td id="76C95DCEFFBE0BA2FB69F8DEFAD5F826" box="[1170,1372,1896,1948]" gridcol="4" gridrow="20" pageId="4" pageNumber="243">2.90 ± 0.20 2.723.11 (3)</td>
</tr>
</table>
</paragraph>
<caption id="DF576678FFBFF458FF6CFCB8FBE7FC9E" box="[151,1134,782,804]" httpUri="https://zenodo.org/record/229417/files/figure.png" pageId="5" pageNumber="244" targetBox="[427,1159,184,760]" targetPageId="5">
<paragraph id="8B9736F0FFBFF458FF6CFCB8FBE7FC9E" blockId="5.[151,1134,782,804]" box="[151,1134,782,804]" pageId="5" pageNumber="244">
<emphasis id="B95CEAE2FFBFF458FF6CFCB8FE84FC99" bold="true" box="[151,269,782,804]" pageId="5" pageNumber="244">FIGURE 2</emphasis>
. Portrait of a living specimen of
<taxonomicName id="4C284D73FFBFF458FDA0FCB9FD65FC9E" box="[603,748,782,804]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="5" pageNumber="244" phylum="Chordata" rank="species" species="dolichodon" status="sp. nov.">
<emphasis id="B95CEAE2FFBFF458FDA0FCB9FD65FC9E" box="[603,748,782,804]" italics="true" pageId="5" pageNumber="244">H. dolichodon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F5799FFBFF458FD09FCB8FCA2FC99" box="[754,811,782,803]" pageId="5" pageNumber="244" rank="species">
<emphasis id="B95CEAE2FFBFF458FD09FCB8FCA2FC99" bold="true" box="[754,811,782,803]" pageId="5" pageNumber="244">n. sp.</emphasis>
</taxonomicNameLabel>
(ROM 110807, from Vietnam).
</paragraph>
</caption>
<caption id="DF576678FFBFF458FF6CFA6FFE34F9B4" httpUri="https://zenodo.org/record/229418/files/figure.png" pageId="5" pageNumber="244" targetBox="[171,1408,853,1481]" targetPageId="5">
<paragraph id="8B9736F0FFBFF458FF6CFA6FFE34F9B4" blockId="5.[151,1436,1497,1550]" pageId="5" pageNumber="244">
<emphasis id="B95CEAE2FFBFF458FF6CFA6FFE9BFA54" bold="true" box="[151,274,1497,1519]" pageId="5" pageNumber="244">FIGURE 3.</emphasis>
Dorsal (left) and ventral (right) hairs of (a)
<taxonomicName id="4C284D73FFBFF458FD35FA6CFCD1FA55" box="[718,856,1497,1519]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="5" pageNumber="244" phylum="Chordata" rank="species" species="pulveratus">
<emphasis id="B95CEAE2FFBFF458FD35FA6CFCD1FA55" box="[718,856,1497,1519]" italics="true" pageId="5" pageNumber="244">H. pulveratus</emphasis>
</taxonomicName>
BM(NH) 70.7.18.12 (holotype) and (b)
<taxonomicName id="4C284D73FFBFF458FB0AFA6CFA08FA55" box="[1265,1409,1497,1519]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="5" pageNumber="244" phylum="Chordata" rank="species" species="dolichodon" status="sp. nov.">
<emphasis id="B95CEAE2FFBFF458FB0AFA6CFA08FA55" box="[1265,1409,1497,1519]" italics="true" pageId="5" pageNumber="244">H. dolichodon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F5799FFBFF458FA7CFA6CFF3CF9B7" pageId="5" pageNumber="244" rank="species">
<emphasis id="B95CEAE2FFBFF458FA7CFA6CFF3CF9B7" bold="true" pageId="5" pageNumber="244">n. sp.</emphasis>
</taxonomicNameLabel>
ROM 110459 (holotype).
</paragraph>
</caption>
<paragraph id="8B9736F0FFBFF458FF3CF980FDF3F898" blockId="5.[151,1437,1590,2038]" pageId="5" pageNumber="244">
The skull is rather large with a relatively robust, long and narrow rostrum (
<figureCitation id="13132A75FFBFF458FBF2F980FBC3F9F5" box="[1033,1098,1590,1615]" captionStart="FIGURE 1" captionStartId="3.[151,250,1908,1930]" captionTargetBox="[350,1229,484,1885]" captionTargetId="figure@3.[340,1248,481,1886]" captionTargetPageId="3" captionText="FIGURE 1. Lateral view of skulls of (a) H. pulveratus BM (NH) 94.9. 1.18 from Fujian, China (b) H. dolichodon n. sp. ROM 110459 (holotype) and (c) H. cf. pulveratus IEBR M- 1610 from Vietnam. Scale = 5 mm" httpUri="https://zenodo.org/record/229416/files/figure.png" pageId="5" pageNumber="244">Fig. 1</figureCitation>
). The cranial profile is evenly ascending with no depression between the rostrum and the flattened braincase. The highest point of the skull is the posterior part of the braincase. The supraorbital tubercles are distinct. The zygoma has no dorsal eminences. The skull lacks a sagittal crest; the lambdoid crest is pronounced and reaches the posterior-most part of the lambdoid. The anterior palatal emargination posteriorly extends to the tip of P2; the posterior palatal emargination has a wide and strong palatal spike. The anteorbital foramen is large. The basioccipital pits are shallow, but visible. The auditory bullae are relatively large (
<figureCitation id="13132A75FFBFF458FDDCF8BCFDE4F898" box="[551,621,1802,1827]" captionStart="FIGURE 4" captionStartId="6.[151,250,1936,1958]" captionTargetBox="[151,1436,193,1914]" captionTargetId="figure@6.[151,1436,193,1914]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal, lateral, and ventral views of skulls and occlusal and lateral views of mandibles of H. pulveratus ROM 110653, from Laos (left) and H. dolichodon n. sp. ROM 110462, paratype (right). Scale = 5 mm." httpUri="https://zenodo.org/record/229419/files/figure.png" pageId="5" pageNumber="244">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8B9736F0FFBFF458FF3CF89BFDC8F84C" blockId="5.[151,1437,1590,2038]" pageId="5" pageNumber="244">
The second upper incisor (
<date id="FF961030FFBFF458FE0BF898FD80F8FC" box="[496,521,1838,1862]" pageId="5" pageNumber="244">I2</date>
) is clearly bifid. The cavity of
<date id="FF961030FFBFF458FC9FF898FCF2F8FC" box="[868,891,1838,1862]" pageId="5" pageNumber="244">I3</date>
is directed labially. The
<date id="FF961030FFBFF458FB74F898FB2EF8FC" box="[1167,1191,1838,1862]" pageId="5" pageNumber="244">I3</date>
is shorter than
<date id="FF961030FFBFF458FAA8F898FAE5F8FC" box="[1363,1388,1838,1862]" pageId="5" pageNumber="244">I2</date>
, but its basal dimensions are larger. The
<date id="FF961030FFBFF458FDC8F8E7FDC3F8D3" box="[563,586,1873,1897]" pageId="5" pageNumber="244">I3</date>
and C are not in contact. The upper canine is relatively slender; its height much exceeds that of P4. P2 is about 2/3 of the crown area of the
<date id="FF961030FFBFF458FC7BF8C2FC10F836" box="[896,921,1908,1932]" pageId="5" pageNumber="244">I3</date>
, displaced lingually, and is visible between C and P4 when viewed from the labial side. P4 has a well-developed antero-lingual cingular cusp. In the lower dentition
<date id="FF961030FFBFF458FEFAF80CFEEBF869" box="[257,354,1978,2003]" pageId="5" pageNumber="244">i1 and i2</date>
present three lobes, whereas
<date id="FF961030FFBFF458FD52F80CFD37F869" box="[681,702,1978,2003]" pageId="5" pageNumber="244">i3</date>
possesses an additional fourth lobe on the lingual side. The lower molars are clearly myotodont (
<figureCitation id="13132A75FFBFF458FE14F86BFDBAF84C" box="[495,563,2013,2038]" captionStart="FIGURE 5" captionStartId="7.[151,250,1146,1168]" captionTargetBox="[157,1429,193,1115]" captionTargetId="figure@7.[151,1436,193,1125]" captionTargetPageId="7" captionText="FIGURE 5. Occlusal view of left upper (left side) and right lower (right side) dentition of (a) H. pulveratus BM (NH) 70.7. 18.12 (holotype) and (b) H. dolichodon n. sp. ROM 110459 (holotype). Scale = 3 mm" httpUri="https://zenodo.org/record/229420/files/figure.png" pageId="5" pageNumber="244">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF576678FFBCF45BFF6CF826FBEEF87E" httpUri="https://zenodo.org/record/229419/files/figure.png" pageId="6" pageNumber="245" targetBox="[151,1436,193,1914]" targetPageId="6">
<paragraph id="8B9736F0FFBCF45BFF6CF826FBEEF87E" blockId="6.[151,1436,1936,1989]" pageId="6" pageNumber="245">
<emphasis id="B95CEAE2FFBCF45BFF6CF826FE86F81F" bold="true" box="[151,271,1936,1958]" pageId="6" pageNumber="245">FIGURE 4</emphasis>
. Dorsal, lateral, and ventral views of skulls and occlusal and lateral views of mandibles of
<taxonomicName id="4C284D73FFBCF45BFB36F827FAD3F81C" box="[1229,1370,1936,1958]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="6" pageNumber="245" phylum="Chordata" rank="species" species="pulveratus">
<emphasis id="B95CEAE2FFBCF45BFB36F827FAD3F81C" box="[1229,1370,1936,1958]" italics="true" pageId="6" pageNumber="245">H. pulveratus</emphasis>
</taxonomicName>
ROM 110653, from Laos (left) and
<taxonomicName id="4C284D73FFBCF45BFE3AF806FDDBF87E" box="[449,594,1966,1989]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="6" pageNumber="245" phylum="Chordata" rank="species" species="dolichodon" status="sp. nov.">
<emphasis id="B95CEAE2FFBCF45BFE3AF806FDDBF87E" box="[449,594,1966,1989]" italics="true" pageId="6" pageNumber="245">H. dolichodon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F5799FFBCF45BFDA3F819FD1BF87E" box="[600,658,1967,1988]" pageId="6" pageNumber="245" rank="species">
<emphasis id="B95CEAE2FFBCF45BFDA3F819FD1BF87E" bold="true" box="[600,658,1967,1988]" pageId="6" pageNumber="245">n. sp.</emphasis>
</taxonomicNameLabel>
ROM 110462, paratype (right). Scale= 5 mm.
</paragraph>
</caption>
<caption id="DF576678FFBDF45AFF6CFBCCFBA3FB15" httpUri="https://zenodo.org/record/229420/files/figure.png" pageId="7" pageNumber="246" targetBox="[157,1429,193,1115]" targetPageId="7">
<paragraph id="8B9736F0FFBDF45AFF6CFBCCFBA3FB15" blockId="7.[151,1436,1146,1199]" pageId="7" pageNumber="246">
<emphasis id="B95CEAE2FFBDF45AFF6CFBCCFE98FB35" bold="true" box="[151,273,1146,1168]" pageId="7" pageNumber="246">FIGURE 5</emphasis>
. Occlusal view of left upper (left side) and right lower (right side) dentition of (a)
<taxonomicName id="4C284D73FFBDF45AFB5EFBCDFABDFB2A" box="[1189,1332,1146,1168]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="246" phylum="Chordata" rank="species" species="pulveratus">
<emphasis id="B95CEAE2FFBDF45AFB5EFBCDFABDFB2A" box="[1189,1332,1146,1168]" italics="true" pageId="7" pageNumber="246">H. pulveratus</emphasis>
</taxonomicName>
BM(NH) 70.7.18.12 (holotype) and (b)
<taxonomicName id="4C284D73FFBDF45AFE3CFB2CFDD1FB15" box="[455,600,1177,1199]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="246" phylum="Chordata" rank="species" species="dolichodon" status="sp. nov.">
<emphasis id="B95CEAE2FFBDF45AFE3CFB2CFDD1FB15" box="[455,600,1177,1199]" italics="true" pageId="7" pageNumber="246">H. dolichodon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F5799FFBDF45AFDA5FB2FFD1EFB15" box="[606,663,1177,1199]" pageId="7" pageNumber="246" rank="species">
<emphasis id="B95CEAE2FFBDF45AFDA5FB2FFD18FB14" bold="true" box="[606,657,1177,1198]" pageId="7" pageNumber="246">n. sp</emphasis>
.
</taxonomicNameLabel>
ROM 110459 (holotype). Scale= 3 mm
</paragraph>
</caption>
<subSubSection id="C332657BFFBDF45AFF3CFB60FE90F9B5" pageId="7" pageNumber="246" type="diagnosis">
<paragraph id="8B9736F0FFBDF45AFF3CFB60FCAAFA8C" blockId="7.[151,1437,1238,2019]" pageId="7" pageNumber="246">
<emphasis id="B95CEAE2FFBDF45AFF3CFB60FEE5FB55" bold="true" box="[199,364,1238,1263]" pageId="7" pageNumber="246">Comparisons.</emphasis>
<taxonomicName id="4C284D73FFBDF45AFE8DFB61FD93FB54" box="[374,538,1238,1262]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="246" phylum="Chordata" rank="species" species="dolichodon" status="sp. nov.">
<emphasis id="B95CEAE2FFBDF45AFE8DFB61FD93FB54" box="[374,538,1238,1262]" italics="true" pageId="7" pageNumber="246">H. dolichodon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F5799FFBDF45AFDDFFB60FDEFFB55" box="[548,614,1238,1263]" pageId="7" pageNumber="246" rank="species">
<emphasis id="B95CEAE2FFBDF45AFDDFFB60FDEFFB55" bold="true" box="[548,614,1238,1263]" pageId="7" pageNumber="246">n. sp.</emphasis>
</taxonomicNameLabel>
is distinguishable by the combination of the black or nearly black ear, muzzle and wings, relatively large P2 visible in lateral view, and shallow basioccipital pits from all other known Southeast Asian species of
<taxonomicName id="4C284D73FFBDF45AFE32FAA9FDA4FA8C" box="[457,557,1311,1334]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="246" phylum="Chordata" rank="genus">
<emphasis id="B95CEAE2FFBDF45AFE32FAA9FDA4FA8C" box="[457,557,1311,1334]" italics="true" pageId="7" pageNumber="246">Hypsugo</emphasis>
</taxonomicName>
except
<taxonomicName id="4C284D73FFBDF45AFD7FFAA9FC95FA8C" box="[644,796,1310,1334]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="246" phylum="Chordata" rank="species" species="pulveratus">
<emphasis id="B95CEAE2FFBDF45AFD7FFAA9FC95FA8C" box="[644,796,1310,1334]" italics="true" pageId="7" pageNumber="246">H. pulveratus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9736F0FFBDF45AFF3CFAF4FE90F9B5" blockId="7.[151,1437,1238,2019]" pageId="7" pageNumber="246">
Although with overlapping values,
<taxonomicName id="4C284D73FFBDF45AFDA1FAF5FD75FAE0" box="[602,764,1346,1370]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="246" phylum="Chordata" rank="species" species="dolichodon" status="sp. nov.">
<emphasis id="B95CEAE2FFBDF45AFDA1FAF5FD75FAE0" box="[602,764,1346,1370]" italics="true" pageId="7" pageNumber="246">H. dolichodon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F5799FFBDF45AFCFFFAF4FCB5FAE1" box="[772,828,1346,1371]" pageId="7" pageNumber="246" rank="species">
<emphasis id="B95CEAE2FFBDF45AFCFFFAF4FCB5FAE1" bold="true" box="[772,828,1346,1371]" pageId="7" pageNumber="246">n.sp.</emphasis>
</taxonomicNameLabel>
is significantly (p&lt;0.05) larger than
<taxonomicName id="4C284D73FFBDF45AFB18FAF5FAF4FAE0" box="[1251,1405,1346,1370]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="246" phylum="Chordata" rank="species" species="pulveratus">
<emphasis id="B95CEAE2FFBDF45AFB18FAF5FAF4FAE0" box="[1251,1405,1346,1370]" italics="true" pageId="7" pageNumber="246">H. pulveratus</emphasis>
</taxonomicName>
in the following measurements: FA, MAW, BCW, CM3L, ML and cm
<specimenCount id="9D2EFD79FFBDF45AFC5CFAD1FC43FAC5" box="[935,970,1383,1407]" pageId="7" pageNumber="246" type="generic">3L.</specimenCount>
The epiblema of
<taxonomicName id="4C284D73FFBDF45AFB61FAD1FAB4FAC4" box="[1178,1341,1382,1406]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="246" phylum="Chordata" rank="species" species="dolichodon" status="sp. nov.">
<emphasis id="B95CEAE2FFBDF45AFB61FAD1FAB4FAC4" box="[1178,1341,1382,1406]" italics="true" pageId="7" pageNumber="246">H. dolichodon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F5799FFBDF45AFABCFAD0FAF6FAC4" box="[1351,1407,1382,1407]" pageId="7" pageNumber="246" rank="species">
<emphasis id="B95CEAE2FFBDF45AFABCFAD0FAF0FAC5" bold="true" box="[1351,1401,1382,1407]" pageId="7" pageNumber="246">n.sp</emphasis>
.
</taxonomicNameLabel>
is narrow and elongated (vs. more definite in
<taxonomicName id="4C284D73FFBDF45AFD84FA3DFC91FA18" box="[639,792,1418,1442]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="246" phylum="Chordata" rank="species" species="pulveratus">
<emphasis id="B95CEAE2FFBDF45AFD84FA3DFC91FA18" box="[639,792,1418,1442]" italics="true" pageId="7" pageNumber="246">H. pulveratus</emphasis>
</taxonomicName>
). The rostral profile of the skull is almost flat (vs. with a distinctly elevated frontal region). Both upper and lower canines become narrow beyond the cingular ring and their height exceeds that of the corresponding second premolars (vs. relatively and absolutely shorter in
<taxonomicName id="4C284D73FFBDF45AFB02FA65FA1BFA50" box="[1273,1426,1490,1514]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="246" phylum="Chordata" rank="species" species="pulveratus">
<emphasis id="B95CEAE2FFBDF45AFB02FA65FA1BFA50" box="[1273,1426,1490,1514]" italics="true" pageId="7" pageNumber="246">H. pulveratus</emphasis>
</taxonomicName>
) (
<figureCitation id="13132A75FFBDF45AFF64FA40FF65F9B5" box="[159,236,1526,1551]" captionStart="FIGURE 1" captionStartId="3.[151,250,1908,1930]" captionTargetBox="[350,1229,484,1885]" captionTargetId="figure@3.[340,1248,481,1886]" captionTargetPageId="3" captionText="FIGURE 1. Lateral view of skulls of (a) H. pulveratus BM (NH) 94.9. 1.18 from Fujian, China (b) H. dolichodon n. sp. ROM 110459 (holotype) and (c) H. cf. pulveratus IEBR M- 1610 from Vietnam. Scale = 5 mm" httpUri="https://zenodo.org/record/229416/files/figure.png" pageId="7" pageNumber="246">Figs. 1</figureCitation>
,
<figureCitation id="13132A75FFBDF45AFF01FA41FE82F9B5" box="[250,267,1527,1551]" captionStart="FIGURE 4" captionStartId="6.[151,250,1936,1958]" captionTargetBox="[151,1436,193,1914]" captionTargetId="figure@6.[151,1436,193,1914]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal, lateral, and ventral views of skulls and occlusal and lateral views of mandibles of H. pulveratus ROM 110653, from Laos (left) and H. dolichodon n. sp. ROM 110462, paratype (right). Scale = 5 mm." httpUri="https://zenodo.org/record/229419/files/figure.png" pageId="7" pageNumber="246">4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C332657BFFBDF45AFF3CF9ACFCBCF859" pageId="7" pageNumber="246" type="discussion">
<paragraph id="8B9736F0FFBDF45AFF3CF9ACFB32F979" blockId="7.[151,1437,1238,2019]" pageId="7" pageNumber="246">
<emphasis id="B95CEAE2FFBDF45AFF3CF9ACFDC8F989" bold="true" box="[199,577,1562,1587]" pageId="7" pageNumber="246">Principal Component Analysis</emphasis>
. In addition to the morphological comparisons and univariate statistical analyses, Principal Component Analysis (PCA) was used to compare
<taxonomicName id="4C284D73FFBDF45AFC56F989FBCFF9EC" box="[941,1094,1598,1622]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="246" phylum="Chordata" rank="species" species="pulveratus">
<emphasis id="B95CEAE2FFBDF45AFC56F989FBCFF9EC" box="[941,1094,1598,1622]" italics="true" pageId="7" pageNumber="246">H. pulveratus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C284D73FFBDF45AFB84F989FAA9F9EC" box="[1151,1312,1598,1622]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="246" phylum="Chordata" rank="species" species="dolichodon" status="sp. nov.">
<emphasis id="B95CEAE2FFBDF45AFB84F989FAA9F9EC" box="[1151,1312,1598,1622]" italics="true" pageId="7" pageNumber="246">H. dolichodon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F5799FFBDF45AFAD3F988FAE1F9ED" box="[1320,1384,1598,1623]" pageId="7" pageNumber="246" rank="species">
<emphasis id="B95CEAE2FFBDF45AFAD3F988FAE1F9ED" bold="true" box="[1320,1384,1598,1623]" pageId="7" pageNumber="246">n. sp.</emphasis>
</taxonomicNameLabel>
The PCA results on 14 craniodental measurements showed good separation between the two species (
<figureCitation id="13132A75FFBDF45AFB1EF9D4FAA2F9C1" box="[1253,1323,1634,1659]" captionStart="FIGURE 6" captionStartId="9.[151,250,1359,1381]" captionTargetBox="[167,1420,195,1338]" captionTargetId="figure@9.[167,1420,195,1338]" captionTargetPageId="9" captionText="FIGURE 6. Principal Component Analysis based on 14 craniodental characters of H. pulveratus (black circles) and H. dolichodon n. sp. (empty triangles)." httpUri="https://zenodo.org/record/229421/files/figure.png" pageId="7" pageNumber="246">Fig. 6</figureCitation>
,
<tableCitation id="C6AA034BFFBDF45AFAC3F9D4FA06F9C0" box="[1336,1423,1634,1659]" captionStart="TABLE 2" captionStartId="8.[151,239,152,175]" captionTargetBox="[159,1236,202,886]" captionTargetPageId="8" captionText="TABLE 2. Factor loadings of craniodental characters obtained by the PCA." httpUri="http://table.plazi.org/id/DF576678FFB2F455FF6CFF2EFC35FF15" pageId="7" pageNumber="246" tableUuid="DF576678FFB2F455FF6CFF2EFC35FF15">Table 2</tableCitation>
). PC1 is a size axis and PC2 indicates the wideness of the skull, with high loadings on IOW, BCW and MAW, indicating
<taxonomicName id="4C284D73FFBDF45AFEF5F91DFE27F978" box="[270,430,1706,1730]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="246" phylum="Chordata" rank="species" species="dolichodon" status="sp. nov.">
<emphasis id="B95CEAE2FFBDF45AFEF5F91DFE27F978" box="[270,430,1706,1730]" italics="true" pageId="7" pageNumber="246">H. dolichodon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F5799FFBDF45AFE4EF91CFE7AF979" box="[437,499,1706,1731]" pageId="7" pageNumber="246" rank="species">
<emphasis id="B95CEAE2FFBDF45AFE4EF91CFE7AF979" bold="true" box="[437,499,1706,1731]" pageId="7" pageNumber="246">n. sp.</emphasis>
</taxonomicNameLabel>
has a relatively wider braincase with greater interorbital width.
</paragraph>
<paragraph id="8B9736F0FFBDF45AFF3CF978FCBCF859" blockId="7.[151,1437,1238,2019]" pageId="7" pageNumber="246">
<emphasis id="B95CEAE2FFBDF45AFF3CF978FE4CF95D" bold="true" box="[199,453,1742,1767]" pageId="7" pageNumber="246">Genetic relationships</emphasis>
.
<collectionCode id="ED39AE35FFBDF45AFE2EF979FD9BF95D" LSID="urn:lsid:biocol.org:col:15536" box="[469,530,1743,1767]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="7" pageNumber="246">DNA</collectionCode>
barcodes for
<specimenCount id="9D2EFD79FFBDF45AFD4BF979FCB5F95D" box="[688,828,1742,1767]" pageId="7" pageNumber="246" type="generic">4 specimens</specimenCount>
of
<taxonomicName id="4C284D73FFBDF45AFC9FF979FB8FF95C" box="[868,1030,1742,1766]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="246" phylum="Chordata" rank="species" species="dolichodon" status="sp. nov.">
<emphasis id="B95CEAE2FFBDF45AFC9FF979FB8FF95C" box="[868,1030,1742,1766]" italics="true" pageId="7" pageNumber="246">H. dolichodon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F5799FFBDF45AFBF4F978FBD9F95D" box="[1039,1104,1742,1767]" pageId="7" pageNumber="246" rank="species">
<emphasis id="B95CEAE2FFBDF45AFBF4F978FBD9F95D" bold="true" box="[1039,1104,1742,1767]" pageId="7" pageNumber="246">n. sp.</emphasis>
</taxonomicNameLabel>
from
<collectingCountry id="F33F7660FFBDF45AFB63F979FB4DF95C" box="[1176,1220,1743,1766]" name="Laos" pageId="7" pageNumber="246">Lao</collectingCountry>
PDR and
<collectingCountry id="F33F7660FFBDF45AFAC6F978FA15F95D" box="[1341,1436,1742,1767]" name="Vietnam" pageId="7" pageNumber="246">Vietnam</collectingCountry>
deposited in
<collectionCode id="ED39AE35FFBDF45AFEC8F944FEFFF8B0" LSID="urn:lsid:biocol.org:col:34778" box="[307,374,1778,1802]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="7" pageNumber="246">ROM</collectionCode>
, as well as
<specimenCount id="9D2EFD79FFBDF45AFDF5F944FD17F8B1" box="[526,670,1778,1803]" pageId="7" pageNumber="246" type="generic">3 specimens</specimenCount>
from
<collectingCountry id="F33F7660FFBDF45AFD14F944FCC7F8B1" box="[751,846,1778,1803]" name="Vietnam" pageId="7" pageNumber="246">Vietnam</collectingCountry>
which are deposited in
<collectionCode id="ED39AE35FFBDF45AFB83F945FB46F8B0" box="[1144,1231,1779,1802]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="7" pageNumber="246">ZMMU</collectionCode>
(these additional specimens were not examined as part of this study) were published by
<bibRefCitation id="EFB94B01FFBDF45AFC4DF8A0FB12F895" author="Francis" box="[950,1179,1814,1839]" etAl="et al." firstAuthor="Francis" pageId="7" pageNumber="246" pagination="12575" refId="ref5760" refString="Francis, C. M., Borisenko, A. V., Ivanova, N. V., Eger, J. L., Lim, B. K., Guillen-Servent, A., Kruskop, S. V., Mackie, I. &amp; Hebert, P. D. N. (2010) The role of DNA barcodes in understanding and conservation of mammal diversity in Southeast Asia. PLoS ONE, 5, e 12575." type="journal article" year="2010">
Francis
<emphasis id="B95CEAE2FFBDF45AFBEBF8A1FBC0F894" box="[1040,1097,1814,1838]" italics="true" pageId="7" pageNumber="246">et al.</emphasis>
(2010)
</bibRefCitation>
as “
<taxonomicName id="4C284D73FFBDF45AFB35F8A1FABBF894" box="[1230,1330,1815,1838]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="246" phylum="Chordata" rank="genus">
<emphasis id="B95CEAE2FFBDF45AFB35F8A1FABBF894" box="[1230,1330,1815,1838]" italics="true" pageId="7" pageNumber="246">Hypsugo</emphasis>
</taxonomicName>
CMF sp. A”. No intraspecific variation in the barcode was detected within the series, even from distinct localities, suggesting that all of these specimens represent the same species. The sequences differed by 12.414.0% (mean 13.7%) from the next nearest neighbour for which
<collectionCode id="ED39AE35FFBDF45AFD2CF834FC9DF820" LSID="urn:lsid:biocol.org:col:15536" box="[727,788,1922,1946]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="7" pageNumber="246">DNA</collectionCode>
barcodes were available, which was
<taxonomicName id="4C284D73FFBDF45AFB41F835FADAF820" box="[1210,1363,1922,1946]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="246" phylum="Chordata" rank="species" species="pulveratus">
<emphasis id="B95CEAE2FFBDF45AFB41F835FADAF820" box="[1210,1363,1922,1946]" italics="true" pageId="7" pageNumber="246">H. pulveratus</emphasis>
</taxonomicName>
. They diverged by 15.6 to 18.0% (mean 16.4%) from a series of
<taxonomicName id="4C284D73FFBDF45AFCDDF811FC3AF804" box="[806,947,1958,1982]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="246" phylum="Chordata" rank="species" species="cadornae">
<emphasis id="B95CEAE2FFBDF45AFCDDF811FC3AF804" box="[806,947,1958,1982]" italics="true" pageId="7" pageNumber="246">H. cadornae</emphasis>
</taxonomicName>
. These levels of divergence clearly support the recognition of
<taxonomicName id="4C284D73FFBDF45AFE9EF87DFD8CF858" box="[357,517,1994,2018]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="246" phylum="Chordata" rank="species" species="dolichodon" status="sp. nov.">
<emphasis id="B95CEAE2FFBDF45AFE9EF87DFD8CF858" box="[357,517,1994,2018]" italics="true" pageId="7" pageNumber="246">H. dolichodon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F5799FFBDF45AFDF7F87CFDC2F859" box="[524,587,1994,2019]" pageId="7" pageNumber="246" rank="species">
<emphasis id="B95CEAE2FFBDF45AFDF7F87CFDC2F859" bold="true" box="[524,587,1994,2019]" pageId="7" pageNumber="246">n. sp.</emphasis>
</taxonomicNameLabel>
as a distinct species.
</paragraph>
</subSubSection>
<caption id="DF576678FFB2F455FF6CFF2EFC35FF15" ID-Table-UUID="DF576678FFB2F455FF6CFF2EFC35FF15" box="[151,956,151,175]" httpUri="http://table.plazi.org/id/DF576678FFB2F455FF6CFF2EFC35FF15" pageId="8" pageNumber="247" startId="8.[151,239,152,175]" targetBox="[159,1236,202,886]" targetIsTable="true" targetPageId="8">
<paragraph id="8B9736F0FFB2F455FF6CFF2EFC35FF15" blockId="8.[151,956,151,175]" box="[151,956,151,175]" pageId="8" pageNumber="247">
<emphasis id="B95CEAE2FFB2F455FF6CFF2EFE8AFF15" bold="true" box="[151,259,152,175]" pageId="8" pageNumber="247">TABLE 2</emphasis>
. Factor loadings of craniodental characters obtained by the PCA.
</paragraph>
</caption>
<paragraph id="8B9736F0FFB2F455FF64FF7CFB3BFCCC" pageId="8" pageNumber="247">
<table id="F928C450FFB20BA2FF64FF7CFB5DFCCC" box="[159,1236,202,886]" gridcols="4" gridrows="17" pageId="8" pageNumber="247">
<tr id="351834B2FFB20BA2FF64FF7CFB5DFF5A" box="[159,1236,202,224]" gridrow="0" pageId="8" pageNumber="247">
<th id="76C95DCEFFB20BA2FF64FF7CFE3BFF5A" box="[159,434,202,224]" gridcol="0" gridrow="0" pageId="8" pageNumber="247">Character</th>
<th id="76C95DCEFFB20BA2FD94FF7CFD21FF5A" box="[623,680,202,224]" gridcol="1" gridrow="0" pageId="8" pageNumber="247">PC1</th>
<th id="76C95DCEFFB20BA2FC7AFF7CFC4BFF5A" box="[897,962,202,224]" gridcol="2" gridrow="0" pageId="8" pageNumber="247">PC2</th>
<th id="76C95DCEFFB20BA2FB69FF7CFB5DFF5A" box="[1170,1236,202,224]" gridcol="3" gridrow="0" pageId="8" pageNumber="247">PC3</th>
</tr>
<tr id="351834B2FFB20BA2FF64FF42FB5DFEB0" box="[159,1236,244,266]" gridrow="1" pageId="8" pageNumber="247">
<th id="76C95DCEFFB20BA2FF64FF42FE3BFEB0" box="[159,434,244,266]" gridcol="0" gridrow="1" pageId="8" pageNumber="247">GLS-i</th>
<td id="76C95DCEFFB20BA2FD94FF42FD21FEB0" box="[623,680,244,266]" gridcol="1" gridrow="1" pageId="8" pageNumber="247">0.837</td>
<td id="76C95DCEFFB20BA2FC7AFF42FC4BFEB0" box="[897,962,244,266]" gridcol="2" gridrow="1" pageId="8" pageNumber="247">-0.140</td>
<td id="76C95DCEFFB20BA2FB69FF42FB5DFEB0" box="[1170,1236,244,266]" gridcol="3" gridrow="1" pageId="8" pageNumber="247">0.105</td>
</tr>
<tr id="351834B2FFB20BA2FF64FEABFB5DFE89" box="[159,1236,285,307]" gridrow="2" pageId="8" pageNumber="247">
<th id="76C95DCEFFB20BA2FF64FEABFE3BFE89" box="[159,434,285,307]" gridcol="0" gridrow="2" pageId="8" pageNumber="247">GLS</th>
<td id="76C95DCEFFB20BA2FD94FEABFD21FE89" box="[623,680,285,307]" gridcol="1" gridrow="2" pageId="8" pageNumber="247">0.871</td>
<td id="76C95DCEFFB20BA2FC7AFEABFC4BFE89" box="[897,962,285,307]" gridcol="2" gridrow="2" pageId="8" pageNumber="247">-0.291</td>
<td id="76C95DCEFFB20BA2FB69FEABFB5DFE89" box="[1170,1236,285,307]" gridcol="3" gridrow="2" pageId="8" pageNumber="247">0.054</td>
</tr>
<tr id="351834B2FFB20BA2FF64FEF0FB5DFEE6" box="[159,1236,326,348]" gridrow="3" pageId="8" pageNumber="247">
<th id="76C95DCEFFB20BA2FF64FEF0FE3BFEE6" box="[159,434,326,348]" gridcol="0" gridrow="3" pageId="8" pageNumber="247">CCL</th>
<td id="76C95DCEFFB20BA2FD94FEF0FD21FEE6" box="[623,680,326,348]" gridcol="1" gridrow="3" pageId="8" pageNumber="247">0.932</td>
<td id="76C95DCEFFB20BA2FC7AFEF0FC4BFEE6" box="[897,962,326,348]" gridcol="2" gridrow="3" pageId="8" pageNumber="247">-0.202</td>
<td id="76C95DCEFFB20BA2FB69FEF0FB5DFEE6" box="[1170,1236,326,348]" gridcol="3" gridrow="3" pageId="8" pageNumber="247">-0.032</td>
</tr>
<tr id="351834B2FFB20BA2FF64FEC6FB5DFE3C" box="[159,1236,368,390]" gridrow="4" pageId="8" pageNumber="247">
<th id="76C95DCEFFB20BA2FF64FEC6FE3BFE3C" box="[159,434,368,390]" gridcol="0" gridrow="4" pageId="8" pageNumber="247">C1C1W</th>
<td id="76C95DCEFFB20BA2FD94FEC6FD21FE3C" box="[623,680,368,390]" gridcol="1" gridrow="4" pageId="8" pageNumber="247">0.678</td>
<td id="76C95DCEFFB20BA2FC7AFEC6FC4BFE3C" box="[897,962,368,390]" gridcol="2" gridrow="4" pageId="8" pageNumber="247">-0.167</td>
<td id="76C95DCEFFB20BA2FB69FEC6FB5DFE3C" box="[1170,1236,368,390]" gridcol="3" gridrow="4" pageId="8" pageNumber="247">0.156</td>
</tr>
<tr id="351834B2FFB20BA2FF64FE2FFB5DFE15" box="[159,1236,409,431]" gridrow="5" pageId="8" pageNumber="247">
<th id="76C95DCEFFB20BA2FF64FE2FFE3BFE15" box="[159,434,409,431]" gridcol="0" gridrow="5" pageId="8" pageNumber="247">M3M3W</th>
<td id="76C95DCEFFB20BA2FD94FE2FFD21FE15" box="[623,680,409,431]" gridcol="1" gridrow="5" pageId="8" pageNumber="247">0.862</td>
<td id="76C95DCEFFB20BA2FC7AFE2FFC4BFE15" box="[897,962,409,431]" gridcol="2" gridrow="5" pageId="8" pageNumber="247">-0.142</td>
<td id="76C95DCEFFB20BA2FB69FE2FFB5DFE15" box="[1170,1236,409,431]" gridcol="3" gridrow="5" pageId="8" pageNumber="247">0.057</td>
</tr>
<tr id="351834B2FFB20BA2FF64FE74FB5DFE62" box="[159,1236,450,472]" gridrow="6" pageId="8" pageNumber="247">
<th id="76C95DCEFFB20BA2FF64FE74FE3BFE62" box="[159,434,450,472]" gridcol="0" gridrow="6" pageId="8" pageNumber="247">IOW</th>
<td id="76C95DCEFFB20BA2FD94FE74FD21FE62" box="[623,680,450,472]" gridcol="1" gridrow="6" pageId="8" pageNumber="247">0.418</td>
<td id="76C95DCEFFB20BA2FC7AFE74FC4BFE62" box="[897,962,450,472]" gridcol="2" gridrow="6" pageId="8" pageNumber="247">0.648</td>
<td id="76C95DCEFFB20BA2FB69FE74FB5DFE62" box="[1170,1236,450,472]" gridcol="3" gridrow="6" pageId="8" pageNumber="247">0.355</td>
</tr>
<tr id="351834B2FFB20BA2FF64FE5AFB5DFDB8" box="[159,1236,492,514]" gridrow="7" pageId="8" pageNumber="247">
<th id="76C95DCEFFB20BA2FF64FE5AFE3BFDB8" box="[159,434,492,514]" gridcol="0" gridrow="7" pageId="8" pageNumber="247">MAW</th>
<td id="76C95DCEFFB20BA2FD94FE5AFD21FDB8" box="[623,680,492,514]" gridcol="1" gridrow="7" pageId="8" pageNumber="247">0.724</td>
<td id="76C95DCEFFB20BA2FC7AFE5AFC4BFDB8" box="[897,962,492,514]" gridcol="2" gridrow="7" pageId="8" pageNumber="247">0.483</td>
<td id="76C95DCEFFB20BA2FB69FE5AFB5DFDB8" box="[1170,1236,492,514]" gridcol="3" gridrow="7" pageId="8" pageNumber="247">-0.08</td>
</tr>
<tr id="351834B2FFB20BA2FF64FDA3FB5DFD91" box="[159,1236,533,555]" gridrow="8" pageId="8" pageNumber="247">
<th id="76C95DCEFFB20BA2FF64FDA3FE3BFD91" box="[159,434,533,555]" gridcol="0" gridrow="8" pageId="8" pageNumber="247">BCW</th>
<td id="76C95DCEFFB20BA2FD94FDA3FD21FD91" box="[623,680,533,555]" gridcol="1" gridrow="8" pageId="8" pageNumber="247">0.515</td>
<td id="76C95DCEFFB20BA2FC7AFDA3FC4BFD91" box="[897,962,533,555]" gridcol="2" gridrow="8" pageId="8" pageNumber="247">0.672</td>
<td id="76C95DCEFFB20BA2FB69FDA3FB5DFD91" box="[1170,1236,533,555]" gridcol="3" gridrow="8" pageId="8" pageNumber="247">0.306</td>
</tr>
<tr id="351834B2FFB20BA2FF64FD88FB5DFDEE" box="[159,1236,574,596]" gridrow="9" pageId="8" pageNumber="247">
<th id="76C95DCEFFB20BA2FF64FD88FE3BFDEE" box="[159,434,574,596]" gridcol="0" gridrow="9" pageId="8" pageNumber="247">BCH</th>
<td id="76C95DCEFFB20BA2FD94FD88FD21FDEE" box="[623,680,574,596]" gridcol="1" gridrow="9" pageId="8" pageNumber="247">0.575</td>
<td id="76C95DCEFFB20BA2FC7AFD88FC4BFDEE" box="[897,962,574,596]" gridcol="2" gridrow="9" pageId="8" pageNumber="247">-0.527</td>
<td id="76C95DCEFFB20BA2FB69FD88FB5DFDEE" box="[1170,1236,574,596]" gridcol="3" gridrow="9" pageId="8" pageNumber="247">0.398</td>
</tr>
<tr id="351834B2FFB20BA2FF64FDDEFB5DFDC4" box="[159,1236,616,638]" gridrow="10" pageId="8" pageNumber="247">
<th id="76C95DCEFFB20BA2FF64FDDEFE3BFDC4" box="[159,434,616,638]" gridcol="0" gridrow="10" pageId="8" pageNumber="247">CM3L</th>
<td id="76C95DCEFFB20BA2FD94FDDEFD21FDC4" box="[623,680,616,638]" gridcol="1" gridrow="10" pageId="8" pageNumber="247">0.886</td>
<td id="76C95DCEFFB20BA2FC7AFDDEFC4BFDC4" box="[897,962,616,638]" gridcol="2" gridrow="10" pageId="8" pageNumber="247">0.114</td>
<td id="76C95DCEFFB20BA2FB69FDDEFB5DFDC4" box="[1170,1236,616,638]" gridcol="3" gridrow="10" pageId="8" pageNumber="247">-0.297</td>
</tr>
<tr id="351834B2FFB20BA2FF64FD27FB5DFD1D" box="[159,1236,657,679]" gridrow="11" pageId="8" pageNumber="247">
<th id="76C95DCEFFB20BA2FF64FD27FE3BFD1D" box="[159,434,657,679]" gridcol="0" gridrow="11" pageId="8" pageNumber="247">CP4L</th>
<td id="76C95DCEFFB20BA2FD94FD27FD21FD1D" box="[623,680,657,679]" gridcol="1" gridrow="11" pageId="8" pageNumber="247">0.682</td>
<td id="76C95DCEFFB20BA2FC7AFD27FC4BFD1D" box="[897,962,657,679]" gridcol="2" gridrow="11" pageId="8" pageNumber="247">-0.056</td>
<td id="76C95DCEFFB20BA2FB69FD27FB5DFD1D" box="[1170,1236,657,679]" gridcol="3" gridrow="11" pageId="8" pageNumber="247">-0.600</td>
</tr>
<tr id="351834B2FFB20BA2FF64FD0CFB5DFD6A" box="[159,1236,698,720]" gridrow="12" pageId="8" pageNumber="247">
<th id="76C95DCEFFB20BA2FF64FD0CFE3BFD6A" box="[159,434,698,720]" gridcol="0" gridrow="12" pageId="8" pageNumber="247">ML</th>
<td id="76C95DCEFFB20BA2FD94FD0CFD21FD6A" box="[623,680,698,720]" gridcol="1" gridrow="12" pageId="8" pageNumber="247">0.857</td>
<td id="76C95DCEFFB20BA2FC7AFD0CFC4BFD6A" box="[897,962,698,720]" gridcol="2" gridrow="12" pageId="8" pageNumber="247">-0.039</td>
<td id="76C95DCEFFB20BA2FB69FD0CFB5DFD6A" box="[1170,1236,698,720]" gridcol="3" gridrow="12" pageId="8" pageNumber="247">-0.064</td>
</tr>
<tr id="351834B2FFB20BA2FF64FD52FB5DFD40" box="[159,1236,740,762]" gridrow="13" pageId="8" pageNumber="247">
<th id="76C95DCEFFB20BA2FF64FD52FE3BFD40" box="[159,434,740,762]" gridcol="0" gridrow="13" pageId="8" pageNumber="247">cm3L</th>
<td id="76C95DCEFFB20BA2FD94FD52FD21FD40" box="[623,680,740,762]" gridcol="1" gridrow="13" pageId="8" pageNumber="247">0.840</td>
<td id="76C95DCEFFB20BA2FC7AFD52FC4BFD40" box="[897,962,740,762]" gridcol="2" gridrow="13" pageId="8" pageNumber="247">0.270</td>
<td id="76C95DCEFFB20BA2FB69FD52FB5DFD40" box="[1170,1236,740,762]" gridcol="3" gridrow="13" pageId="8" pageNumber="247">-0.296</td>
</tr>
<tr id="351834B2FFB20BA2FF64FCBBFB5DFC99" box="[159,1236,781,803]" gridrow="14" pageId="8" pageNumber="247">
<th id="76C95DCEFFB20BA2FF64FCBBFE3BFC99" box="[159,434,781,803]" gridcol="0" gridrow="14" pageId="8" pageNumber="247">CPH</th>
<td id="76C95DCEFFB20BA2FD94FCBBFD21FC99" box="[623,680,781,803]" gridcol="1" gridrow="14" pageId="8" pageNumber="247">0.765</td>
<td id="76C95DCEFFB20BA2FC7AFCBBFC4BFC99" box="[897,962,781,803]" gridcol="2" gridrow="14" pageId="8" pageNumber="247">-0.166</td>
<td id="76C95DCEFFB20BA2FB69FCBBFB5DFC99" box="[1170,1236,781,803]" gridcol="3" gridrow="14" pageId="8" pageNumber="247">0.313</td>
</tr>
<tr id="351834B2FFB20BA2FF64FC80FB5DFCF6" box="[159,1236,822,844]" gridrow="15" pageId="8" pageNumber="247">
<th id="76C95DCEFFB20BA2FF64FC80FE3BFCF6" box="[159,434,822,844]" gridcol="0" gridrow="15" pageId="8" pageNumber="247">Eigenvalue</th>
<td id="76C95DCEFFB20BA2FD94FC80FD21FCF6" box="[623,680,822,844]" gridcol="1" gridrow="15" pageId="8" pageNumber="247">8.101</td>
<td id="76C95DCEFFB20BA2FC7AFC80FC4BFCF6" box="[897,962,822,844]" gridcol="2" gridrow="15" pageId="8" pageNumber="247">1.693</td>
<td id="76C95DCEFFB20BA2FB69FC80FB5DFCF6" box="[1170,1236,822,844]" gridcol="3" gridrow="15" pageId="8" pageNumber="247">1.064</td>
</tr>
<tr id="351834B2FFB20BA2FF64FCD6FB5DFCCC" box="[159,1236,864,886]" gridrow="16" pageId="8" pageNumber="247">
<th id="76C95DCEFFB20BA2FF64FCD6FE3BFCCC" box="[159,434,864,886]" gridcol="0" gridrow="16" pageId="8" pageNumber="247">Percentage of variance (%)</th>
<td id="76C95DCEFFB20BA2FD94FCD6FD21FCCC" box="[623,680,864,886]" gridcol="1" gridrow="16" pageId="8" pageNumber="247">57.9</td>
<td id="76C95DCEFFB20BA2FC7AFCD6FC4BFCCC" box="[897,962,864,886]" gridcol="2" gridrow="16" pageId="8" pageNumber="247">12.1</td>
<td id="76C95DCEFFB20BA2FB69FCD6FB5DFCCC" box="[1170,1236,864,886]" gridcol="3" gridrow="16" pageId="8" pageNumber="247">7.6</td>
</tr>
</table>
</paragraph>
<subSubSection id="C332657BFFB2F455FF3CFC0CFC76FB11" pageId="8" pageNumber="247" type="distribution">
<paragraph id="8B9736F0FFB2F455FF3CFC0CFC76FB11" blockId="8.[151,1437,954,1195]" pageId="8" pageNumber="247">
<emphasis id="B95CEAE2FFB2F455FF3CFC0CFDC7FC69" bold="true" box="[199,590,954,979]" pageId="8" pageNumber="247">Distribution and ecological notes</emphasis>
. The known distribution of this species is limited to
<collectingCountry id="F33F7660FFB2F455FB67FC0DFB41FC68" box="[1180,1224,955,978]" name="Laos" pageId="8" pageNumber="247">Lao</collectingCountry>
PDR and southern
<collectingCountry id="F33F7660FFB2F455FF6CFC68FF73FC4D" box="[151,250,990,1015]" name="Vietnam" pageId="8" pageNumber="247">Vietnam</collectingCountry>
. All of the
<collectingCountry id="F33F7660FFB2F455FE8EFC69FE28FC4C" box="[373,417,991,1014]" name="Laos" pageId="8" pageNumber="247">Lao</collectingCountry>
specimens were captured in mist nets near the Xe Kaman, a moderately large, fast-flowing river
<quantity id="4CD09B15FFB2F455FF2FFBB4FECEFBA0" box="[212,327,1026,1051]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="8" pageNumber="247" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">50100 m</quantity>
across, flowing through lowland evergreen rainforest in the Dong Amphan National Biodiversity Area. Two of them were caught in nets set perpendicular to the river, along its bank, and one was set across the mouth of a small tributary. All of the
<collectionCode id="ED39AE35FFB2F455FDC1FBFDFDF3FBD9" LSID="urn:lsid:biocol.org:col:34778" box="[570,634,1099,1123]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="8" pageNumber="247">ROM</collectionCode>
and
<collectionCode id="ED39AE35FFB2F455FD4BFBFDFC8EFBD8" box="[688,775,1099,1122]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="8" pageNumber="247">ZMMU</collectionCode>
Vietnamese specimens came from Cat Tien National Park. The
<collectionCode id="ED39AE35FFB2F455FF37FBD9FE85FB3D" LSID="urn:lsid:biocol.org:col:34778" box="[204,268,1135,1159]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="8" pageNumber="247">ROM</collectionCode>
specimen was collected near the headquarters of the Park, close to the Dong Nai River. This area is generally described as lowland rain forest with dipterocarp trees (
<bibRefCitation id="EFB94B01FFB2F455FC8FFB25FC7BFB11" author="Scott" box="[884,1010,1171,1195]" firstAuthor="Scott" pageId="8" pageNumber="247" refId="ref6476" refString="Scott, D. A. (1989) A Directory of Asian Wetlands. IUCN, Gland, 1181 pp." type="book" year="1989">Scott 1989</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>