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<mods:title id="152B7D400B8740D79816047F959964D8">Notes on Oriental and East Palaearctic Manota Williston (Diptera, Mycetophilidae), with the description of seven new species</mods:title>
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<subSubSection id="C3326512FFE4521504FAFC61BF144BF5" pageId="2" pageNumber="379" type="nomenclature">
<paragraph id="8B973699FFE4521504FAFC61BF454B94" blockId="2.[151,368,908,967]" box="[151,368,908,934]" pageId="2" pageNumber="379">
<heading id="D0DF81F5FFE4521504FAFC61BF454B94" bold="true" box="[151,368,908,934]" fontSize="11" level="1" pageId="2" pageNumber="379" reason="1">
<emphasis id="B95CEA8BFFE4521504FAFC61BF454B94" bold="true" box="[151,368,908,934]" pageId="2" pageNumber="379">
<taxonomicName id="4C284D1AFFE4521504FAFC61BF704B94" ID-CoL="8CPXW" authorityName="Hippa &amp; Saigusa" authorityYear="2016" box="[151,325,908,934]" class="Insecta" family="Mycetophilidae" genus="Manota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="incilis">
<emphasis id="B95CEA8BFFE4521504FAFC61BF704B94" bold="true" box="[151,325,908,934]" italics="true" pageId="2" pageNumber="379">Manota incilis</emphasis>
</taxonomicName>
sp.
</emphasis>
</heading>
</paragraph>
<paragraph id="8B973699FFE4521504FAFC43BF144BF5" blockId="2.[151,368,908,967]" box="[151,289,942,967]" pageId="2" pageNumber="379">
<figureCitation id="13132A1CFFE4521504FAFC43BF144BF5" box="[151,289,942,967]" captionStart="FIGURE 2" captionStartId="3.[151,250,1396,1418]" captionTargetBox="[206,1384,534,1364]" captionTargetId="figure@3.[197,1389,522,1374]" captionTargetPageId="3" captionText="FIGURE 2. Manota incilis sp. n. (holotype). A. Antennal flagellomere 4, lateral view. B. Hypopygium, ventral view. C. Hypopygium, dorsal view. D. Gonostylus, dorsal view. Scale 0.1 mm. 1 = juxtagonostylar megaseta, 2 = lobe at dorsomedial margin of gonocoxa." httpUri="https://zenodo.org/record/1052175/files/figure.png" pageId="2" pageNumber="379">Figs. 2 A–D</figureCitation>
</paragraph>
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<subSubSection id="C3326512FFE4521504FAFC1BBFA54FAB" pageId="2" pageNumber="379" type="description">
<paragraph id="8B973699FFE4521504FAFC1BBB6F4F47" blockId="2.[151,1437,1014,2017]" pageId="2" pageNumber="379">
Male. 
<emphasis id="B95CEA8BFFE4521504B2FC1BBF044C3D" bold="true" box="[223,305,1014,1039]" pageId="2" pageNumber="379">Colour</emphasis>
. Head pale brown, vertex darker brown. Antenna brown. Mouthparts pale yellowish. Thorax brown. Legs pale yellowish brown, a patch at base of hind coxa, apices of mid- and hind coxae, mid- and hind trochanters, apical third of hind femur and an indistinct patch baso-ventrally on hind femur infuscated. Wing unicolorous brownish; halter pale brown with darker brown knob. Abdomen brown. All setosity pale, yellowish or brownish. 
<emphasis id="B95CEA8BFFE4521504FAFB6BBEE24CAD" bold="true" box="[151,215,1158,1183]" pageId="2" pageNumber="379">Head</emphasis>
. Antennal flagellomere 4, 
<figureCitation id="13132A1CFFE45215066FFB6BBC544CAD" box="[514,609,1158,1183]" captionStart="FIGURE 2" captionStartId="3.[151,250,1396,1418]" captionTargetBox="[206,1384,534,1364]" captionTargetId="figure@3.[197,1389,522,1374]" captionTargetPageId="3" captionText="FIGURE 2. Manota incilis sp. n. (holotype). A. Antennal flagellomere 4, lateral view. B. Hypopygium, ventral view. C. Hypopygium, dorsal view. D. Gonostylus, dorsal view. Scale 0.1 mm. 1 = juxtagonostylar megaseta, 2 = lobe at dorsomedial margin of gonocoxa." httpUri="https://zenodo.org/record/1052175/files/figure.png" pageId="2" pageNumber="379">Fig. 2 A</figureCitation>
. Palpomere 3 of maxillary palpus with apicomedial thumb-like extension, with 4 apically curved sensilla; palpomere 4 with parasegment; palpomere 5 as long as palpomere 4. Number of strong postocular setae 11. 
<emphasis id="B95CEA8BFFE4521505BCFB23BC1F4CD5" bold="true" box="[465,554,1230,1255]" pageId="2" pageNumber="379">Thorax</emphasis>
. Anepisternum setose, with 41 setae; anterior basalare setose, with 16 setae; preepisternum 2 setose, with 17 setae; laterotergite nonsetose, episternum 3 setose, with 14 setae. 
<emphasis id="B95CEA8BFFE452150081FB1FBB114D39" bold="true" box="[1260,1316,1266,1291]" pageId="2" pageNumber="379">Legs</emphasis>
. Midtibial organ absent. Hind tibial organ present. 
<emphasis id="B95CEA8BFFE45215061FFAFBBC864D1D" bold="true" box="[626,691,1302,1327]" pageId="2" pageNumber="379">Wing</emphasis>
. R1 meeting C well on the basal half of the costal margin; the sclerotized part of M2 not quite extending to the level of the tip of R1; wing length 
<quantity id="4CD09B7CFFE452150052FAD3BAAC4D67" box="[1087,1177,1342,1366]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="379" unit="mm" value="2.3">2.3 mm</quantity>
. 
<emphasis id="B95CEA8BFFE4521500C9FAD0BB734D64" bold="true" box="[1188,1350,1341,1366]" pageId="2" pageNumber="379">Hypopygium,</emphasis>
<figureCitation id="13132A1CFFE452150122FAD0BEE54D4E" captionStart="FIGURE 2" captionStartId="3.[151,250,1396,1418]" captionTargetBox="[206,1384,534,1364]" captionTargetId="figure@3.[197,1389,522,1374]" captionTargetPageId="3" captionText="FIGURE 2. Manota incilis sp. n. (holotype). A. Antennal flagellomere 4, lateral view. B. Hypopygium, ventral view. C. Hypopygium, dorsal view. D. Gonostylus, dorsal view. Scale 0.1 mm. 1 = juxtagonostylar megaseta, 2 = lobe at dorsomedial margin of gonocoxa." httpUri="https://zenodo.org/record/1052175/files/figure.png" pageId="2" pageNumber="379">Figs. 2 B–D</figureCitation>
: Sternite 9 less than one-half of the ventral length of gonocoxa, lateral margins sharply delimited, convex, posterior margin with deep U-shaped medial indentation, anterior margin with similar indentation, the setae similar to the ventral setae of gonocoxa. Ventromedial margin of gonocoxa simple, shallowly sigmoid. Parastylar lobe subtriangular, with several setae at posterior margin. Paraapodemal lobe simple, in ventral view covered by the parastylar lobe. Both sternite 9 and the ventral side of the gonocoxa give a densely setose impression. Dorsomedial margin of gonocoxa simple, slightly sigmoid, forming a rounded lobe posteriorly with the posterior margin of the gonocoxa and bearing a number of short setae. Gonocoxa without any kind of posterolateral lobe dorsally. Gonocoxa with a plate-like lobe posteriorly, situated ventrally from the dorsomedial margin, bearing ca. 20 setae on its ventral surface, among these setae there are several rather large ones which form a curved row. Two juxtagonostylar setae present, both rather weakly modified megasetae, one with fine hair-like apex, the other with more abruptly acute apex, both arising from a common small basal body, in dorsal or ventral view the two setae are conspicuously divergent. Gonostylus simple, semicircular, the setae unmodified. Aedeagus elongate subtriangular, without lateral shoulders, the apex curved ventrally. Hypoproct extending posteriorly near midlength of gonostylus, with transverse ventral lobe (sternite 10) on each side, the latter bearing ca. 6 setae, one of which is stronger than the others, these setae indistinctly arranged in a row. Cerci medially separate, simple. Tergite 9 membranous.
</paragraph>
<paragraph id="8B973699FFE4521504AAF86DBFA54FAB" blockId="2.[151,1437,1014,2017]" box="[199,400,1920,1945]" pageId="2" pageNumber="379">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C3326512FFE4521404AAF849BC0F490D" lastPageId="3" lastPageNumber="380" pageId="2" pageNumber="379" type="discussion">
<paragraph id="8B973699FFE4521504AAF849BBA94FD2" blockId="2.[151,1437,1014,2017]" pageId="2" pageNumber="379">
<emphasis id="B95CEA8BFFE4521504AAF849BF704F8F" bold="true" box="[199,325,1956,1981]" pageId="2" pageNumber="379">Discussion</emphasis>
. In the key to Oriental and Palaearctic 
<taxonomicName id="4C284D1AFFE452150697F848BD644F8E" baseAuthorityName="Hippa" baseAuthorityYear="2011" box="[762,849,1957,1980]" class="Insecta" family="Mycetophilidae" genus="Manota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="379" phylum="Arthropoda" rank="genus">
<emphasis id="B95CEA8BFFE452150697F848BD644F8E" box="[762,849,1957,1980]" italics="true" pageId="2" pageNumber="379">Manota</emphasis>
</taxonomicName>
given by 
<bibRefCitation id="EFB94B68FFE4521507D2F849BA604F8F" author="Hippa" box="[959,1109,1956,1981]" pageId="2" pageNumber="379" refString="Hippa, H., Kjaerandsen, J. &amp; Saigusa, T. (2011) The genus Manota Williston (Diptera, Mycetophilidae) in Japan, with a key to the Palaearctic species of the genus. Zootaxa, 2892, 33 - 46." type="journal article" year="2011">Hippa (2011)</bibRefCitation>
<taxonomicName id="4C284D1AFFE452150036F848BAF04F8E" authorityName="Hippa &amp; Saigusa" authorityYear="2016" box="[1115,1221,1956,1980]" class="Insecta" family="Mycetophilidae" genus="Manota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="incilis">
<emphasis id="B95CEA8BFFE452150036F848BAF04F8E" box="[1115,1221,1956,1980]" italics="true" pageId="2" pageNumber="379">M. incilis</emphasis>
</taxonomicName>
runs to couplet 50, leading to the eastern Palaearctic 
<taxonomicName id="4C284D1AFFE45215064FF824BD934FD3" authority="Hippa &amp; Kjaerandsen" authorityName="Hippa &amp; Kjaerandsen" authorityYear="2010" box="[546,934,1992,2017]" class="Insecta" family="Mycetophilidae" genus="Manota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="indahae">
<emphasis id="B95CEA8BFFE45215064FF824BC934FD2" box="[546,678,1992,2016]" italics="true" pageId="2" pageNumber="379">M. indahae</emphasis>
Hippa &amp; Kjaerandsen
</taxonomicName>
and the Oriental 
<taxonomicName id="4C284D1AFFE452150014F824BB0A4FD3" authority="Hippa" authorityName="Hippa" authorityYear="2007" box="[1145,1343,1992,2017]" class="Insecta" family="Mycetophilidae" genus="Manota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="collina">
<emphasis id="B95CEA8BFFE452150014F824BAC54FD2" box="[1145,1264,1992,2016]" italics="true" pageId="2" pageNumber="379">M. collina</emphasis>
Hippa
</taxonomicName>
and 
<emphasis id="B95CEA8BFFE452150110F824BBA94FD2" box="[1405,1436,1993,2016]" italics="true" pageId="2" pageNumber="379">M.</emphasis>
</paragraph>
<paragraph id="8B973699FFE5521404FAFF75BC0F490D" blockId="3.[151,1436,151,467]" pageId="3" pageNumber="380">
<taxonomicName id="4C284D1AFFE5521404FAFF75BF42489D" authority="Hippa. It" authorityName="Hippa. It" authorityYear="2074" box="[151,375,151,176]" class="Insecta" family="Mycetophilidae" genus="Manota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="380" phylum="Arthropoda" rank="species" species="subcollina">
<emphasis id="B95CEA8BFFE5521404FAFF75BF394882" box="[151,268,152,176]" italics="true" pageId="3" pageNumber="380">subcollina</emphasis>
Hippa. It
</taxonomicName>
differs from all in its broad, almost semicircular gonostylus which is ca.1.3 times longer than broad (versus elongate, over twice as long as broad), from 
<taxonomicName id="4C284D1AFFE552140735FF50BDE948E6" authorityName="Hippa &amp; Kjaerandsen" authorityYear="2010" box="[856,988,188,212]" class="Insecta" family="Mycetophilidae" genus="Manota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="380" phylum="Arthropoda" rank="species" species="indahae">
<emphasis id="B95CEA8BFFE552140735FF50BDE948E6" box="[856,988,188,212]" italics="true" pageId="3" pageNumber="380">M. indahae</emphasis>
</taxonomicName>
by having the apex of aedeagus bent ventrally (versus straight) and from 
<taxonomicName id="4C284D1AFFE55214065EFF0CBC9D48CA" authorityName="Hippa" authorityYear="2007" box="[563,680,224,248]" class="Insecta" family="Mycetophilidae" genus="Manota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="380" phylum="Arthropoda" rank="species" species="collina">
<emphasis id="B95CEA8BFFE55214065EFF0CBC9D48CA" box="[563,680,224,248]" italics="true" pageId="3" pageNumber="380">M. collina</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C284D1AFFE55214068FFF0CBD4348CA" authorityName="Hippa. It" authorityYear="2074" box="[738,886,224,248]" class="Insecta" family="Mycetophilidae" genus="Manota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="380" phylum="Arthropoda" rank="species" species="subcollina">
<emphasis id="B95CEA8BFFE55214068FFF0CBD4348CA" box="[738,886,224,248]" italics="true" pageId="3" pageNumber="380">M subcollina</emphasis>
</taxonomicName>
by the unicolorous wing (versus pale basal part and dark apical part). The broad gonostylus distinguishes 
<taxonomicName id="4C284D1AFFE55214072FFEE8BD85492E" authorityName="Hippa &amp; Saigusa" authorityYear="2016" box="[834,944,260,284]" class="Insecta" family="Mycetophilidae" genus="Manota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="380" phylum="Arthropoda" rank="species" species="incilis">
<emphasis id="B95CEA8BFFE55214072FFEE8BD85492E" box="[834,944,260,284]" italics="true" pageId="3" pageNumber="380">M. incilis</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C284D1AFFE552140793FEE8BAAB492E" authorityName="Hippa &amp; Saigusa" authorityYear="2016" box="[1022,1182,260,284]" class="Insecta" family="Mycetophilidae" genus="Manota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="380" phylum="Arthropoda" rank="species" species="nepalensis">
<emphasis id="B95CEA8BFFE552140793FEE8BAAB492E" box="[1022,1182,260,284]" italics="true" pageId="3" pageNumber="380">M. nepalensis</emphasis>
</taxonomicName>
described below. For further discussion, see 
<taxonomicName id="4C284D1AFFE5521405FAFEC4BC064972" authorityName="Hippa &amp; Saigusa" authorityYear="2016" box="[407,563,296,320]" class="Insecta" family="Mycetophilidae" genus="Manota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="380" phylum="Arthropoda" rank="species" species="nepalensis">
<emphasis id="B95CEA8BFFE5521405FAFEC4BC064972" box="[407,563,296,320]" italics="true" pageId="3" pageNumber="380">M. nepalensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3326512FFE5521404AAFEA1BF8B49E1" pageId="3" pageNumber="380" type="etymology">
<paragraph id="8B973699FFE5521404AAFEA1BF3049BA" blockId="3.[151,1436,151,467]" pageId="3" pageNumber="380">
<emphasis id="B95CEA8BFFE5521404AAFEA1BF7B4957" bold="true" box="[199,334,332,357]" pageId="3" pageNumber="380">Etymology.</emphasis>
The name is Latin, 
<emphasis id="B95CEA8BFFE55214065DFEA1BC494956" box="[560,636,332,356]" italics="true" pageId="3" pageNumber="380">
<taxonomicName id="4C284D1AFFE55214065DFEA1BC424956" authorityName="Hippa &amp; Saigusa" authorityYear="2016" box="[560,631,332,356]" class="Insecta" family="Mycetophilidae" genus="Manota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="380" phylum="Arthropoda" rank="species" species="incilis">incilis</taxonomicName>
,
</emphasis>
cut in, referring to the deeply incised anterior and posterior margins of sternite 9.
</paragraph>
<paragraph id="8B973699FFE5521404AAFE7BBF8B49E1" blockId="3.[151,1436,151,467]" pageId="3" pageNumber="380">
<materialsCitation id="3B403CC4FFE5521404AAFE7BBF8B49E1" ID-GBIF-Occurrence="1701200706" collectingDate="1971-09-21" collectingDateMax="1971-09-22" collectingDateMin="1971-09-21" collectionCode="KMNH" collectorName="A. Nakanishi" country="Nepal" elevation="1950" latitude="28.583334" location="Male" longLatPrecision="1231" longitude="83.38333" municipality="Bogara" pageId="3" pageNumber="380" specimenCount="2" typeStatus="holotype">
<emphasis id="B95CEA8BFFE5521404AAFE7BBF26499D" bold="true" box="[199,275,406,431]" pageId="3" pageNumber="380">
<typeStatus id="5493883BFFE5521404AAFE7BBF3A499D" box="[199,271,406,431]" pageId="3" pageNumber="380">Types</typeStatus>
.
</emphasis>
<typeStatus id="5493883BFFE552140576FE7ABFB7499D" box="[283,386,407,431]" pageId="3" pageNumber="380" type="holotype">
<emphasis id="B95CEA8BFFE552140576FE7ABFB7499D" box="[283,386,407,431]" italics="true" pageId="3" pageNumber="380">Holotype</emphasis>
</typeStatus>
. 
<location id="8EF76042FFE5521405FDFE7BBFF9499D" LSID="urn:lsid:plazi:treatment:0381878FFFE4521404FAFC61BF8B49E1:8EF76042FFE5521405FDFE7BBFF9499D" box="[400,460,406,431]" country="Nepal" latitude="28.583334" longLatPrecision="1231" longitude="83.38333" municipality="Bogara" name="Male" pageId="3" pageNumber="380">Male</location>
, 
<collectingCountry id="F33F7609FFE5521405B4FE7ABC01499D" box="[473,564,407,431]" name="Nepal" pageId="3" pageNumber="380">NEPAL</collectingCountry>
, 
<collectingMunicipality id="6BF3ACE3FFE552140652FE75BCA64982" box="[575,659,408,432]" pageId="3" pageNumber="380">Bogara</collectingMunicipality>
, 
<elevation id="0005D1AAFFE5521406F2FE7ABD09499D" box="[671,828,407,431]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.2" metricValueMin="1.7" pageId="3" pageNumber="380" unit="m" value="1950.0" valueMax="2200.0" valueMin="1700.0">
<quantity id="4CD09B7CFFE5521406F2FE7ABD09499D" box="[671,828,407,431]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.2" metricValueMin="1.7" pageId="3" pageNumber="380" unit="m" value="1950.0" valueMax="2200.0" valueMin="1700.0">1700–2200 m</quantity>
</elevation>
, 
<geoCoordinate id="EE1C505EFFE552140724FE7ABD98499D" box="[841,941,403,431]" direction="east" orientation="longitude" pageId="3" pageNumber="380" precision="925" value="83.38333">83o23’ E</geoCoordinate>
, 
<geoCoordinate id="EE1C505EFFE5521407D4FE7ABA14499D" box="[953,1057,403,431]" direction="north" orientation="latitude" pageId="3" pageNumber="380" precision="925" value="28.583334">28o35’ N</geoCoordinate>
, 
<collectingMethod id="52694E8EFFE552140040FE7BBA89499D" box="[1069,1212,406,431]" pageId="3" pageNumber="380">Malaise trap</collectingMethod>
, 
<collectingDate id="EFD2E9B1FFE5521400A5FE7ABBA2499D" box="[1224,1431,407,431]" pageId="3" pageNumber="380" value="1971-09-21" valueMax="1971-09-22" valueMin="1971-09-21">
<date id="FF961059FFE5521400A5FE7ABBA2499D" box="[1224,1431,407,431]" pageId="3" pageNumber="380" value="1971-09-21" valueMax="1971-09-22" valueMin="1971-09-21">Sept. 21–22. 1971</date>
</collectingDate>
, 
<collectorName id="26DD534FFFE5521404FAFE56BF1E49E1" box="[151,299,442,467]" pageId="3" pageNumber="380">A. Nakanishi</collectorName>
(in 
<collectionCode id="ED39AE5CFFE552140535FE56BF8649E1" box="[344,435,443,467]" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13440" name="Kitakyushu Museum and Institute of Natural History" pageId="3" pageNumber="380">KMNH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>