616 lines
74 KiB
XML
616 lines
74 KiB
XML
<document id="547DCA7258A51724E78C16199002E200" ID-CLB-Dataset="22642" ID-DOI="10.11646/zootaxa.4514.3.3" ID-GBIF-Dataset="54115d80-05ed-4ee7-b035-1ac714f9c31d" ID-ISSN="1175-5326" ID-Zenodo-Dep="3770231" ID-ZooBank="8F3F3680-5950-4FAB-B8DC-455FAC27535A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588001511746" checkinUser="plazi" docAuthor="Abdel-Dayem, Mahmoud S., Rasool, Iftekhar, Elgharbawy, Ali A., Nagel, Peter & Aldhafer, Hathal M." docDate="2018" docId="03811B61FF87FF9CA5BCFDA0FE29FF27" docLanguage="en" docName="zootaxa.4514.3.3.pdf" docOrigin="Zootaxa 4514 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paussus (Hylotorus) abditus Abdel-Dayem & Rasool & Elgharbawy & Nagel & Aldhafer 2018, sp. n." docType="treatment" docVersion="7" lastPageNumber="358" masterDocId="FFB86319FF89FF8EA52BFB78FFF4FFD0" masterDocTitle="Faunistic inventory and zoogeographical analysis of the ground beetles (Coleoptera, Carabidae) of Garf Raydah Nature Reserve, Southwestern of Saudi Arabia, and description of a new species of Paussinae" masterLastPageNumber="371" masterPageNumber="341" pageNumber="355" updateTime="1698837116288" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="0A69EFF573469D9311401104464B3675" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2606F54A8D5FBD095BA7C3007768891F">
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<mods:title id="89A84FDA362427CF1B7F447CE4F8B597">Faunistic inventory and zoogeographical analysis of the ground beetles (Coleoptera, Carabidae) of Garf Raydah Nature Reserve, Southwestern of Saudi Arabia, and description of a new species of Paussinae</mods:title>
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<mods:name id="FE1B447F962B2D2E3441F6BF76F9D0A1" type="personal">
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<mods:role id="797275C6D891A7AEAFDFFFB4654482DF">
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<mods:roleTerm id="A9941FFF2A01908C8D753831230945D0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="963477891BC5BC2DFA157871532F232B">Abdel-Dayem, Mahmoud S.</mods:namePart>
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<mods:affiliation id="46FDDE53144B1DCB943B32AC5B86D4D8">Entomology Department, Faculty of Science, Cairo University, Giza 12613, Egypt.</mods:affiliation>
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</mods:name>
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<mods:name id="5C19AD7F79C7D2B42F9D98082436E9F5" type="personal">
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<mods:role id="706420926EDA1FB5CC4918492FD1E3C8">
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<mods:roleTerm id="EDEA87BD4C757DA31957E9037FED1383">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B683470B7F49CC16E89C652EA948602E">Rasool, Iftekhar</mods:namePart>
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<mods:affiliation id="A3B7508A128BC3BF6E2B825711BA0235">King Saud University Museum of Arthropods (KSMA), Plant Protection Department, College of Food and Agriculture Sciences, King Saud University, P. O. Box 2460 Riyadh 11451, Saudi Arabia.</mods:affiliation>
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</mods:name>
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<mods:name id="6A2EB7C4FCDDCBFC03864CE19189B056" type="personal">
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<mods:role id="DE59B515379769856BCC590A98DCAFFB">
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<mods:roleTerm id="F124EFB1C3D1965D43585A988968B935">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FCC92D7103E075615A86A1742B90A4EC">Elgharbawy, Ali A.</mods:namePart>
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<mods:affiliation id="232C97B13B1EFE1F389E0D93059069EF">Zoology Department, Faculty of Science, Al Azhar University, Nasr City, Cairo, Egypt.</mods:affiliation>
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</mods:name>
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<mods:name id="90459CD899456FCDE92328EFE2750400" type="personal">
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<mods:roleTerm id="2C4DC32D221120488DEAD11D61267EF6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B2B5576137FEC10B894E3AA69C9D672F">Nagel, Peter</mods:namePart>
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<mods:affiliation id="6F7758C696B9C84159A895A2F1024250">Biogeography Research Group, Department of Environmental Sciences, Faculty of Science, University of Basel, St. Johanns-Vorstadt 10, 4056 Basel, Switzerland</mods:affiliation>
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</mods:role>
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<mods:namePart id="87A2E3E957B3809366C358882EB1D33A">Aldhafer, Hathal M.</mods:namePart>
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<mods:affiliation id="5DAB305012D5117B12C6E3F3AEFAE717">King Saud University Museum of Arthropods (KSMA), Plant Protection Department, College of Food and Agriculture Sciences, King Saud University, P. O. Box 2460 Riyadh 11451, Saudi Arabia.</mods:affiliation>
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<mods:typeOfResource id="4022222275243DDDBAF58B56800A51D2">text</mods:typeOfResource>
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<mods:title id="4F8D327A631F6F821E4ECE838758D1BF">Zootaxa</mods:title>
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<mods:part id="5709BF5F26492416E1302A034D865AFE">
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<mods:date id="57427C21A80BCC9D9D4F7E04F709F081">2018</mods:date>
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<mods:number id="F5A6FD6AE6FFABB5FFB480C615431904">2018-11-08</mods:number>
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<mods:number id="3F9D889761E38FF9455DEBA4F3238EEB">4514</mods:number>
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<mods:identifier id="81C81843C059FB472D25D9758E901D8B" type="DOI">10.11646/zootaxa.4514.3.3</mods:identifier>
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<treatment id="03811B61FF87FF9CA5BCFDA0FE29FF27" ID-DOI="http://doi.org/10.5281/zenodo.3799022" ID-GBIF-Taxon="163754613" ID-Zenodo-Dep="3799022" LSID="urn:lsid:plazi:treatment:03811B61FF87FF9CA5BCFDA0FE29FF27" httpUri="http://treatment.plazi.org/id/03811B61FF87FF9CA5BCFDA0FE29FF27" lastPageId="18" lastPageNumber="358" pageId="14" pageNumber="355">
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<subSubSection id="C332F9FCFF87FF80A5BCFDA0FD78F922" box="[151,652,1752,1778]" pageId="14" pageNumber="355" type="nomenclature">
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<paragraph id="8B97AA77FF87FF80A5BCFDA0FD78F922" blockId="14.[151,652,1752,1811]" box="[151,652,1752,1778]" pageId="14" pageNumber="355">
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<heading id="D0DF1D1BFF87FF80A5BCFDA0FD78F922" bold="true" box="[151,652,1752,1778]" fontSize="11" level="1" pageId="14" pageNumber="355" reason="1">
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<emphasis id="B95C7665FF87FF80A5BCFDA0FD78F922" bold="true" box="[151,652,1752,1778]" pageId="14" pageNumber="355">
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<taxonomicName id="4C28D1F4FF87FF80A5BCFDA0FDC9F922" ID-CoL="8MLD4" authority="Nagel" authorityName="Abdel-Dayem & Rasool & Elgharbawy & Nagel & Aldhafer" authorityYear="2018" box="[151,573,1752,1778]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="355" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Hylotorus">
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||
<emphasis id="B95C7665FF87FF80A5BCFDA0FF0FF922" bold="true" box="[151,251,1752,1778]" italics="true" pageId="14" pageNumber="355">Paussus</emphasis>
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||
(
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||
<emphasis id="B95C7665FF87FF80A420FDA0FE77F922" bold="true" box="[267,387,1752,1778]" italics="true" pageId="14" pageNumber="355">Hylotorus</emphasis>
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||
)
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<emphasis id="B95C7665FF87FF80A4BEFDA0FE19F922" bold="true" box="[405,493,1752,1778]" italics="true" pageId="14" pageNumber="355">abditus</emphasis>
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Nagel
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26FCB1EFF87FF80A761FDA0FD78F922" box="[586,652,1752,1778]" pageId="14" pageNumber="355" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C332F9FCFF87FF80A5BCFD82FEF3F8C3" box="[151,263,1786,1811]" pageId="14" pageNumber="355" type="description">
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<paragraph id="8B97AA77FF87FF80A5BCFD82FEF3F8C3" blockId="14.[151,652,1752,1811]" box="[151,263,1786,1811]" pageId="14" pageNumber="355">
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(
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<figureCitation id="1313B6F2FF87FF80A5B5FD82FF0BF8C3" box="[158,255,1786,1811]" captionStart="FIGURE 2" captionStartId="15.[151,250,1898,1920]" captionTargetBox="[403,1206,208,1867]" captionTargetId="figure@15.[364,1228,193,1884]" captionTargetPageId="15" captionText="FIGURE 2. Paussus (Hylotorus) abditus sp. n. holotype specimen from the Hejaz mountains. The illustration shows the specimen with the appendages of the right side at their broadest view, while the left antennae and legs are shown twisted through 90 degrees, i.e. at their narrowest view (cf. Nagel 2006). The inset shows the “collar” of the anterior pronotum at frontal view. Scale bar: 1 mm. Illustration: E. Weber, Basel, Switzerland. Copyright: P. Nagel." figureDoi="http://doi.org/10.5281/zenodo.3770235" httpUri="https://zenodo.org/record/3770235/files/figure.png" pageId="14" pageNumber="355">Figure 2</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C332F9FCFF87FF9EA5BCFC39FE75FD78" lastPageId="16" lastPageNumber="357" pageId="14" pageNumber="355" type="materials_examined">
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<paragraph id="8B97AA77FF87FF80A5BCFC39FC9DF8AE" blockId="14.[151,1436,1857,2026]" pageId="14" pageNumber="355">
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<emphasis id="B95C7665FF87FF80A5BCFC39FE0AF88A" bold="true" box="[151,510,1857,1882]" pageId="14" pageNumber="355">
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||
<typeStatus id="549314D5FF87FF80A5BCFC39FEF0F88A" box="[151,260,1857,1882]" pageId="14" pageNumber="355" type="holotype">Holotype</typeStatus>
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(hereby designated):
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</emphasis>
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Male; dry-mounted, glued on rectangular, pinned card; antennal clubs, left foreleg, left hind leg detached, labium dissected and glued on same card.
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</paragraph>
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||
<paragraph id="8B97AA77FF87FF80A5ECFCF1FAB1F83A" blockId="14.[151,1436,1857,2026]" pageId="14" pageNumber="355">
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||
<emphasis id="B95C7665FF87FF80A5ECFCF1FE8CF872" bold="true" box="[199,376,1929,1954]" pageId="14" pageNumber="355">Original labels</emphasis>
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||
(rectangular) (
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||
<emphasis id="B95C7665FF87FF80A70FFCF1FD7CF872" box="[548,648,1929,1954]" italics="true" pageId="14" pageNumber="355">verbatim</emphasis>
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||
): 1. Yellow label, handwritten by A. Raffray: “
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||
<emphasis id="B95C7665FF87FF80A18EFCF1FAFDF872" box="[1189,1289,1929,1954]" italics="true" pageId="14" pageNumber="355">Sufamila</emphasis>
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||
[or
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||
<emphasis id="B95C7665FF87FF80A011FCF1FA68F872" box="[1338,1436,1929,1954]" italics="true" pageId="14" pageNumber="355">Jutamila</emphasis>
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||
or
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||
<emphasis id="B95C7665FF87FF80A59CFCD5FED7F816" box="[183,291,1965,1990]" italics="true" pageId="14" pageNumber="355">Sutonnila</emphasis>
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||
or similar, partly illegible] /
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||
<emphasis id="B95C7665FF87FF80A744FCD6FD52F816" box="[623,678,1966,1990]" italics="true" pageId="14" pageNumber="355">Raff.</emphasis>
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” (
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<emphasis id="B95C7665FF87FF80A7E8FCD6FC96F816" box="[707,866,1965,1990]" italics="true" pageId="14" pageNumber="355">nomen nudum</emphasis>
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, unavailable). 2. White label, printed: “El Hajaz / Millinger [sic!]”. 3. White label, printed: “A. Raffray”. 4. White label, printed: “E. Wasmann / vidit 1904”.
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</paragraph>
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||
<caption id="DF57FAFFFF86FF81A5BCFC12FBBAF80C" ID-DOI="http://doi.org/10.5281/zenodo.3770235" ID-Zenodo-Dep="3770235" httpUri="https://zenodo.org/record/3770235/files/figure.png" pageId="15" pageNumber="356" startId="15.[151,250,1898,1920]" targetBox="[403,1206,208,1867]" targetPageId="15">
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||
<paragraph id="8B97AA77FF86FF81A5BCFC12FBBAF80C" blockId="15.[151,1436,1898,2012]" pageId="15" pageNumber="356">
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||
<emphasis id="B95C7665FF86FF81A5BCFC12FEE2F8AF" bold="true" box="[151,278,1898,1920]" pageId="15" pageNumber="356">FIGURE 2.</emphasis>
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<taxonomicName id="4C28D1F4FF86FF81A40AFC13FDBDF850" authority="Abdel-Dayem & Rasool & Elgharbawy & Nagel & Aldhafer, 2018" authorityName="Abdel-Dayem & Rasool & Elgharbawy & Nagel & Aldhafer" authorityYear="2018" box="[289,585,1898,1920]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Hylotorus">
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<emphasis id="B95C7665FF86FF81A40AFC13FDBDF850" box="[289,585,1898,1920]" italics="true" pageId="15" pageNumber="356">Paussus (Hylotorus) abditus</emphasis>
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||
</taxonomicName>
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||
<emphasis id="B95C7665FF86FF81A778FC12FD7BF8AF" bold="true" box="[595,655,1898,1919]" pageId="15" pageNumber="356">
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||
<taxonomicNameLabel id="A26FCB1EFF86FF81A778FC12FD7BF8AF" box="[595,655,1898,1919]" pageId="15" pageNumber="356" rank="species">sp. n.</taxonomicNameLabel>
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||
</emphasis>
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||
holotype specimen from the Hejaz mountains. The illustration shows the specimen with the appendages of the right side at their broadest view, while the left antennae and legs are shown twisted through 90 degrees, i.e. at their narrowest view (cf.
|
||
<bibRefCitation id="EFB9D786FF86FF81A797FCDFFCCCF86D" author="Nagel, P." box="[700,824,1959,1981]" pageId="15" pageNumber="356" pagination="181 - 202" refId="ref22612" refString="Nagel, P. (2006) Ant Nest Beetles of the Carnegie Museum (Coleoptera: Carabidae: Paussinae: Paussini). Annals of Carnegie Museum, 75 (3), 181 - 202. https: // doi. org / 10.2992 / 0097 - 4463 (2006) 75 [181: ANBOTC] 2.0. CO; 2" type="journal article" year="2006">Nagel 2006</bibRefCitation>
|
||
). The inset shows the “collar” of the anterior pronotum at frontal view. Scale bar: 1 mm. Illustration: E. Weber, Basel, Switzerland. Copyright: P. Nagel.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B97AA77FF99FF9EA5ECFBEFFEDEFF04" blockId="16.[151,1437,151,2035]" pageId="16" pageNumber="357">
|
||
<emphasis id="B95C7665FF99FF9EA5ECFBEFFE99FF60" bold="true" box="[199,365,151,176]" pageId="16" pageNumber="357">Added labels:</emphasis>
|
||
1. Red, rectangular, printed: “
|
||
<typeStatus id="549314D5FF99FF9EA7E1FBEFFCB5FF60" box="[714,833,151,176]" pageId="16" pageNumber="357" type="holotype">Holotypus</typeStatus>
|
||
/
|
||
<taxonomicName id="4C28D1F4FF99FF9EA677FBE1FBE0FF60" authorityName="Abdel-Dayem & Rasool & Elgharbawy & Nagel & Aldhafer" authorityYear="2018" box="[860,1044,151,176]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="357" phylum="Arthropoda" rank="species" species="abditus">
|
||
<emphasis id="B95C7665FF99FF9EA677FBE1FBE0FF60" box="[860,1044,151,176]" italics="true" pageId="16" pageNumber="357">Paussus abditus</emphasis>
|
||
</taxonomicName>
|
||
/ P. Nagel, 2018”. 2. White label, printed: “
|
||
<specimenCount id="9D2E61FEFF99FF9EA429FBC4FEEDFF04" box="[258,281,188,212]" pageId="16" pageNumber="357" type="male">♂</specimenCount>
|
||
”.
|
||
</paragraph>
|
||
<paragraph id="8B97AA77FF99FF9EA5ECFBA7FB79FF28" blockId="16.[151,1437,151,2035]" box="[199,1165,223,248]" pageId="16" pageNumber="357">
|
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<materialsCitation id="3B40A02AFF99FF9EA5ECFBA7FB79FF28" ID-GBIF-Occurrence="2608963685" box="[199,1165,223,248]" collectionCode="MNHN" country="France" location="Museum National d'Histoire Naturelle" municipality="Paris" pageId="16" pageNumber="357" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B95C7665FF99FF9EA5ECFBA7FE4AFF28" bold="true" box="[199,446,223,248]" pageId="16" pageNumber="357">
|
||
<typeStatus id="549314D5FF99FF9EA5ECFBA7FEC0FF28" box="[199,308,223,248]" pageId="16" pageNumber="357" type="holotype">Holotype</typeStatus>
|
||
repository:
|
||
</emphasis>
|
||
<location id="8EF7FCACFF99FF9EA4EEFB98FC8DFF28" LSID="urn:lsid:plazi:treatment:03811B61FF87FF9CA5BCFDA0FE29FF27:8EF7FCACFF99FF9EA4EEFB98FC8DFF28" box="[453,889,223,248]" country="France" municipality="Paris" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="357">Muséum National d’Histoire Naturelle</location>
|
||
,
|
||
<collectingMunicipality id="6BF3300DFF99FF9EA6AFFBA7FC4BFF28" box="[900,959,223,248]" pageId="16" pageNumber="357">Paris</collectingMunicipality>
|
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,
|
||
<collectingCountry id="F33FEAE7FF99FF9EA6E1FB98FBE2FF27" box="[970,1046,224,247]" name="France" pageId="16" pageNumber="357">France</collectingCountry>
|
||
(
|
||
<collectionCode id="ED3932B2FF99FF9EA10EFB98FB75FF28" box="[1061,1153,224,248]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="357">MNHN</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B97AA77FF99FF9EA5ECFA7CFC0EFE90" blockId="16.[151,1437,151,2035]" pageId="16" pageNumber="357">
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<emphasis id="B95C7665FF99FF9EA5ECFA7CFEC9FECD" bold="true" box="[199,317,260,285]" pageId="16" pageNumber="357">
|
||
<typeStatus id="549314D5FF99FF9EA5ECFA7CFECCFECD" box="[199,312,260,285]" pageId="16" pageNumber="357" type="paratype">Paratype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Female; dry-mounted, glued on pointed, pinned card; right antennal club absent, tip of abdomen opened and rearranged, gonostyli dissected and glued on card with specimen.
|
||
</paragraph>
|
||
<paragraph id="8B97AA77FF99FF9EA5ECFA34FC36FE7C" blockId="16.[151,1437,151,2035]" pageId="16" pageNumber="357">
|
||
<emphasis id="B95C7665FF99FF9EA5ECFA34FE8BFEB5" bold="true" box="[199,383,332,357]" pageId="16" pageNumber="357">Original labels</emphasis>
|
||
(white, rectangular) (
|
||
<emphasis id="B95C7665FF99FF9EA7A2FA34FD18FEB5" box="[649,748,332,357]" italics="true" pageId="16" pageNumber="357">verbatim</emphasis>
|
||
): 1. printed, “
|
||
<collectingCountry id="F33FEAE7FF99FF9EA6B4FA34FC28FEB4" box="[927,988,332,356]" name="Saudi Arabia" pageId="16" pageNumber="357">KSA</collectingCountry>
|
||
,
|
||
<collectingRegion id="49EC6495FF99FF9EA6C4FA34FBD5FEB5" box="[1007,1057,332,357]" country="Saudi Arabia" name="'Asir" pageId="16" pageNumber="357">Asir</collectingRegion>
|
||
, Abha, Raydah /
|
||
<geoCoordinate id="EE1CCCB0FF99FF9EA02BFA34FA68FEB4" box="[1280,1436,332,357]" degrees="18" direction="north" minutes="12.095" orientation="latitude" pageId="16" pageNumber="357" precision="1" value="18.201584">18°12.095' N</geoCoordinate>
|
||
<geoCoordinate id="EE1CCCB0FF99FF9EA5BCFA17FEDBFE57" box="[151,303,367,392]" degrees="42" direction="east" minutes="24.536" orientation="longitude" pageId="16" pageNumber="357" precision="1" value="42.40893">42°24.536' E</geoCoordinate>
|
||
,
|
||
<quantity id="4CD00792FF99FF9EA415FA17FE6EFE57" box="[318,410,367,392]" metricMagnitude="3" metricUnit="m" metricValue="2.578" pageId="16" pageNumber="357" unit="m" value="2578.0">2578 m</quantity>
|
||
. /
|
||
<date id="FF968CB7FF99FF9EA497FA08FDCAFE58" box="[444,574,368,392]" pageId="16" pageNumber="357" value="2014-06-08">
|
||
08.
|
||
<collectionCode id="ED3932B2FF99FF9EA4F6FA08FE09FE58" box="[477,509,368,392]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="16" pageNumber="357">VI</collectionCode>
|
||
.2014
|
||
</date>
|
||
,
|
||
<collectionCode id="ED3932B2FF99FF9EA765FA08FD8CFE58" box="[590,632,368,392]" pageId="16" pageNumber="357">PT-</collectionCode>
|
||
3 / Al Dhafer, H.; Fadl, H.; Abdel-Dayem, / M.S., El Torkey, A.; El Gharbawy, A.”. 2. printed, “
|
||
<specimenCount id="9D2E61FEFF99FF9EA4FEFAECFE1CFE7C" box="[469,488,404,428]" pageId="16" pageNumber="357" type="female">♀</specimenCount>
|
||
”. 3. handwritten, “sp.1543 /
|
||
<taxonomicName id="4C28D1F4FF99FF9EA607FAEDFC7CFE7C" box="[812,904,405,428]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="357" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95C7665FF99FF9EA607FAEDFC7CFE7C" box="[812,904,405,428]" italics="true" pageId="16" pageNumber="357">Paussus</emphasis>
|
||
</taxonomicName>
|
||
sp.”.
|
||
</paragraph>
|
||
<paragraph id="8B97AA77FF99FF9EA5ECFACFFAC5FE00" blockId="16.[151,1437,151,2035]" box="[199,1329,439,464]" pageId="16" pageNumber="357">
|
||
<emphasis id="B95C7665FF99FF9EA5ECFACFFE94FE00" bold="true" box="[199,352,439,464]" pageId="16" pageNumber="357">Added label:</emphasis>
|
||
White, red frame, rectangular, printed:
|
||
<emphasis id="B95C7665FF99FF9EA637FACFFCDEFE00" bold="true" box="[796,810,439,464]" pageId="16" pageNumber="357">“</emphasis>
|
||
<typeStatus id="549314D5FF99FF9EA601FAC0FC6CFE00" box="[810,920,440,464]" pageId="16" pageNumber="357" type="paratype">Paratypus</typeStatus>
|
||
<emphasis id="B95C7665FF99FF9EA68BFACFFC5CFE00" bold="true" box="[928,936,439,464]" pageId="16" pageNumber="357">/</emphasis>
|
||
<taxonomicName id="4C28D1F4FF99FF9EA685FAC1FB90FE00" authorityName="Abdel-Dayem & Rasool & Elgharbawy & Nagel & Aldhafer" authorityYear="2018" box="[942,1124,439,464]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="357" phylum="Arthropoda" rank="species" species="abditus">
|
||
<emphasis id="B95C7665FF99FF9EA685FAC1FB90FE00" box="[942,1124,439,464]" italics="true" pageId="16" pageNumber="357">Paussus abditus</emphasis>
|
||
</taxonomicName>
|
||
/ P. Nagel, 2018”.
|
||
</paragraph>
|
||
<paragraph id="8B97AA77FF99FF9EA5ECFAA4FE34FDC8" blockId="16.[151,1437,151,2035]" pageId="16" pageNumber="357">
|
||
<materialsCitation id="3B40A02AFF99FF9EA5ECFAA4FE49FDC8" ID-GBIF-Occurrence="2608963635" collectionCode="KSMA" country="Saudi Arabia" location="King Saud University Museum of Arthropods" municipality="King Saud University" pageId="16" pageNumber="357" specimenCount="1" stateProvince="Ar Riyad" typeStatus="paratype">
|
||
<emphasis id="B95C7665FF99FF9EA5ECFAA4FE4BFE25" bold="true" box="[199,447,476,501]" pageId="16" pageNumber="357">
|
||
<typeStatus id="549314D5FF99FF9EA5ECFAA4FEC0FE25" box="[199,308,476,501]" pageId="16" pageNumber="357" type="paratype">Paratype</typeStatus>
|
||
repository:
|
||
</emphasis>
|
||
<location id="8EF7FCACFF99FF9EA4EDFAA4FC3EFE25" LSID="urn:lsid:plazi:treatment:03811B61FF87FF9CA5BCFDA0FE29FF27:8EF7FCACFF99FF9EA4EDFAA4FC3EFE25" box="[454,970,476,501]" country="Saudi Arabia" municipality="King Saud University" name="King Saud University Museum of Arthropods" pageId="16" pageNumber="357" stateProvince="Ar Riyad">King Saud University Museum of Arthropods</location>
|
||
(
|
||
<collectionCode id="ED3932B2FF99FF9EA6F2FAA4FBDBFE24" box="[985,1071,476,500]" pageId="16" pageNumber="357">KSMA</collectionCode>
|
||
),
|
||
<collectingMunicipality id="6BF3300DFF99FF9EA169FAA4FACDFE25" box="[1090,1337,476,501]" pageId="16" pageNumber="357">King Saud University</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49EC6495FF99FF9EA06FFAA4FA6CFE25" box="[1348,1432,476,501]" country="Saudi Arabia" name="Ar Riyad" pageId="16" pageNumber="357">Riyadh</collectingRegion>
|
||
,
|
||
<collectingCountry id="F33FEAE7FF99FF9EA5BCFA87FE49FDC8" box="[151,445,511,536]" name="Saudi Arabia" pageId="16" pageNumber="357">Kingdom of Saudi Arabia</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B97AA77FF99FF9EA5ECF95CFB52FDED" blockId="16.[151,1437,151,2035]" box="[199,1190,548,573]" pageId="16" pageNumber="357">
|
||
<emphasis id="B95C7665FF99FF9EA5ECF95CFE9DFDED" bold="true" box="[199,361,548,573]" pageId="16" pageNumber="357">Type locality:</emphasis>
|
||
Arabian Peninsula, Western
|
||
<collectingCountry id="F33FEAE7FF99FF9EA785F95CFC27FDED" box="[686,979,548,573]" name="Saudi Arabia" pageId="16" pageNumber="357">Kingdom of Saudi Arabia</collectingCountry>
|
||
, Hejaz Mountains.
|
||
</paragraph>
|
||
<paragraph id="8B97AA77FF99FF9EA5ECF93FFE75FD78" blockId="16.[151,1437,151,2035]" pageId="16" pageNumber="357">
|
||
<emphasis id="B95C7665FF99FF9EA5ECF93FFEFCFDB0" bold="true" box="[199,264,583,608]" pageId="16" pageNumber="357">Note:</emphasis>
|
||
The
|
||
<typeStatus id="549314D5FF99FF9EA46AF930FE85FDB0" box="[321,369,584,608]" pageId="16" pageNumber="357">type</typeStatus>
|
||
specimens were collected by Charles Millingen, M.D., Edinburgh at around the 1870ies (labels with collector “Millingen” exist with localities El Hejaz / Hajaz, Jeddah and
|
||
<collectingCountry id="F33FEAE7FF99FF9EA122F915FBAEFD54" box="[1033,1114,621,644]" name="Yemen" pageId="16" pageNumber="357">Yemen</collectingCountry>
|
||
, and the name is sometimes printed “Millinger”).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C332F9FCFF99FF9EA5ECF9CCFAF7FD20" pageId="16" pageNumber="357" type="etymology">
|
||
<paragraph id="8B97AA77FF99FF9EA5ECF9CCFAF7FD20" blockId="16.[151,1437,151,2035]" pageId="16" pageNumber="357">
|
||
<emphasis id="B95C7665FF99FF9EA5ECF9CCFEA5FD1D" bold="true" box="[199,337,692,717]" pageId="16" pageNumber="357">Etymology:</emphasis>
|
||
The specific epithet is the Latin adjective for “hidden” or “undiscovered”. It refers to the fact that this species remained undiscovered for more than 100 years until a second specimen was discovered.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C332F9FCFF99FF9EA5ECF984FE7BFC18" pageId="16" pageNumber="357" type="diagnosis">
|
||
<paragraph id="8B97AA77FF99FF9EA5ECF984FE7BFC18" blockId="16.[151,1437,151,2035]" pageId="16" pageNumber="357">
|
||
<emphasis id="B95C7665FF99FF9EA5ECF984FEB5FCC5" bold="true" box="[199,321,764,789]" pageId="16" pageNumber="357">Diagnosis.</emphasis>
|
||
A small
|
||
<taxonomicName id="4C28D1F4FF99FF9EA48CF985FDF7FCC4" box="[423,515,765,788]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="357" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95C7665FF99FF9EA48CF985FDF7FCC4" box="[423,515,765,788]" italics="true" pageId="16" pageNumber="357">Paussus</emphasis>
|
||
</taxonomicName>
|
||
of the
|
||
<taxonomicName id="4C28D1F4FF99FF9EA779F985FD15FCC5" box="[594,737,764,789]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="357" phylum="Arthropoda" rank="species" species="cucullatus">
|
||
<emphasis id="B95C7665FF99FF9EA779F985FD15FCC5" box="[594,737,764,789]" italics="true" pageId="16" pageNumber="357">P. cucullatus</emphasis>
|
||
</taxonomicName>
|
||
group
|
||
<emphasis id="B95C7665FF99FF9EA61AF985FC56FCC5" box="[817,930,764,789]" italics="true" pageId="16" pageNumber="357">sensu lato</emphasis>
|
||
; characterized by a tumid antennal club with small, projecting anterior basal angle and large, apically broadly rounded posterior basal angle; antennal club with excavation not extending up to apex; head with vertex produced, forming two apically rounded protuberances; two cephalic openings vestigial, hardly discernible; anterior pronotum with blunt and low transverse collar; pronotal trichome well developed; pygidium with lower (posterior) margin with dense fringe of hair; hind tibiae compressed and strongly widened.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C332F9FCFF99FF9FA5ECF8ACFF4BFDED" lastPageId="17" lastPageNumber="358" pageId="16" pageNumber="357" type="description">
|
||
<paragraph id="8B97AA77FF99FF9EA5ECF8ACFD54FB35" blockId="16.[151,1437,151,2035]" pageId="16" pageNumber="357">
|
||
<emphasis id="B95C7665FF99FF9EA5ECF8ACFCFBFC3D" bold="true" box="[199,783,980,1005]" pageId="16" pageNumber="357">
|
||
Description of male
|
||
<typeStatus id="549314D5FF99FF9EA49DF8ACFDE9FC3D" box="[438,541,980,1005]" pageId="16" pageNumber="357" type="holotype">holotype</typeStatus>
|
||
specimen (
|
||
<figureCitation id="1313B6F2FF99FF9EA78AF8ACFCF3FC3D" box="[673,775,980,1005]" captionStart="FIGURE 2" captionStartId="15.[151,250,1898,1920]" captionTargetBox="[403,1206,208,1867]" captionTargetId="figure@15.[364,1228,193,1884]" captionTargetPageId="15" captionText="FIGURE 2. Paussus (Hylotorus) abditus sp. n. holotype specimen from the Hejaz mountains. The illustration shows the specimen with the appendages of the right side at their broadest view, while the left antennae and legs are shown twisted through 90 degrees, i.e. at their narrowest view (cf. Nagel 2006). The inset shows the “collar” of the anterior pronotum at frontal view. Scale bar: 1 mm. Illustration: E. Weber, Basel, Switzerland. Copyright: P. Nagel." figureDoi="http://doi.org/10.5281/zenodo.3770235" httpUri="https://zenodo.org/record/3770235/files/figure.png" pageId="16" pageNumber="357">Figure 2</figureCitation>
|
||
)
|
||
</emphasis>
|
||
. Standardized body length from tip of head to tip of elytra
|
||
<quantity id="4CD00792FF99FF9EA5BCF880FF07FBDF" box="[151,243,1016,1040]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="16" pageNumber="357" unit="mm" value="4.1">4.1 mm</quantity>
|
||
, width across mid-elytra
|
||
<quantity id="4CD00792FF99FF9EA734F88FFD88FBDF" box="[543,636,1015,1040]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="357" unit="mm" value="1.5">1.5 mm</quantity>
|
||
. Body with dorsal and lower side, legs and antennae brown, frontal margins of head and frontal margins of posterior part of pronotum narrowly black. Head, pronotum, elytra with dorsal pubescence of scattered, short, light, lanceolate, narrowly scaliform, apically rounded setae; middle of posterior pronotum glabrous, smooth, glossy; pubescence of appendages equally scattered yet of thinner setae; setae mostly erect to slanting, more appressed at lateral head and antennomere I. Surface shiny except frontal head, neck and antennomere I (scape) with fine, matte microsculpture; punctuation inconspicuous, only scape with coarse, dense punctures; elytra weakly rugose.
|
||
</paragraph>
|
||
<paragraph id="8B97AA77FF99FF9EA5ECFF97FF35F8B3" blockId="16.[151,1437,151,2035]" pageId="16" pageNumber="357">
|
||
Head almost as long as wide, with frontal margin notched in the middle and frontal lobes broadly rounded; eyes slightly vaulted; temples not projecting, at dorsal view forming a regular curve with eye; vertex produced in two rounded protuberances, separated by shallow longitudinal emargination; cephalic openings on top of protuberances vestigial, closed, indicated by small, shallow, oval impressions, difficult to see. Antennomere I devoid of well-marked longitudinal edges; antennal club boat shaped, 1.5 times longer than wide (basal tooth disregarded), tumid; frontal margin of the club with two fenestriform pits near base; anterior basal angle of club small, acute; posterior basal projection large, apically rounded; hind part of the club with large posterior excavation comprising the basal two thirds only; at posterior view dorsal and ventral margins of excavation undulate of four and five tumid tubercles, each with 1 to 3 apical setae; club with distinct tuft of setae (trichome) near ventral base. Labrum with anterior margin weakly emarginate. Maxillary palpomeres similar to Figs 6G,H in
|
||
<bibRefCitation id="EFB9D786FF99FF9EA1DDFD56FEC3F9BB" author="Robertson, J. & Moore, W." pageId="16" pageNumber="357" pagination="134 - 170" refId="ref23316" refString="Robertson, J. A & Moore, W. (2016) Phylogeny of Paussus L. (Carabidae: Paussinae): unravelling morphological convergence associated with myrmecophilous life histories. Systematic Entomology, 42 (1), 134 - 170, figs. S 1 - S 2, tabs. S 1 - S 5 [(January 2017): 28 Sep 2016: version of record online, first published, pp. 170, end of text] https: // doi. org / 10.1111 / syen. 12205" type="journal article" year="2016">Robertson and Moore (2016)</bibRefCitation>
|
||
; palpomere II large, approximately as long as wide, with mesal margin almost straight, mesal apical angle broadly rounded, outer margin rounded with apical angle not produced; maxillary palpomere II three times wider than following palpomere; palpomere III and IV subequal in length, III little wider than terminal palpomere. Lacinia bifalcate (as usual), moderately setose, with 7–8 setae at anterior margin. Labial palpomeres similar to Figs 6C, 7N in
|
||
<bibRefCitation id="EFB9D786FF99FF9EA439FD98FDA4F929" author="Robertson, J. & Moore, W." box="[274,592,1760,1785]" pageId="16" pageNumber="357" pagination="134 - 170" refId="ref23316" refString="Robertson, J. A & Moore, W. (2016) Phylogeny of Paussus L. (Carabidae: Paussinae): unravelling morphological convergence associated with myrmecophilous life histories. Systematic Entomology, 42 (1), 134 - 170, figs. S 1 - S 2, tabs. S 1 - S 5 [(January 2017): 28 Sep 2016: version of record online, first published, pp. 170, end of text] https: // doi. org / 10.1111 / syen. 12205" type="journal article" year="2016">Robertson & Moore (2016)</bibRefCitation>
|
||
; palpomere III long, narrow, 4.5 times as long as wide, apically rounded. Ligula at ventral view with longitudinal carinula on disc; apical margin bisinuate with apical median margin broadly rounded and lateral anterior angles produced anteriorly. Gula: width/length ratio of gula at narrowest point 0.6.
|
||
</paragraph>
|
||
<paragraph id="8B97AA77FF99FF9EA5ECFC17FD91F822" blockId="16.[151,1437,151,2035]" pageId="16" pageNumber="357">Pronotum hardly wider than long, transversely bipartite, with large trichomes at both ends of groove; anterior part little wider than head, low, with transverse dorsal edge of collar-like structure broadly rounded, slightly emarginated in the middle, lateral angles little produced; posterior part narrowed towards base. Elytra with even, scattered pubescence; hindwings present.</paragraph>
|
||
<paragraph id="8B97AA77FF98FF9FA5ECFBEFFE64FEB5" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="358">
|
||
Fused abdominal ventrites 1–3 with scraper of stridulatory organ present (synapomorphic character of
|
||
<taxonomicName id="4C28D1F4FF98FF9FA06BFBE1FA68FF60" box="[1344,1436,153,176]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95C7665FF98FF9FA06BFBE1FA68FF60" box="[1344,1436,153,176]" italics="true" pageId="17" pageNumber="358">Paussus</emphasis>
|
||
</taxonomicName>
|
||
L.
|
||
<emphasis id="B95C7665FF98FF9FA59CFBC4FF23FF05" box="[183,215,188,213]" italics="true" pageId="17" pageNumber="358">s.l.</emphasis>
|
||
). Pygidium with disc flat and lower (anatomically posterior) margin narrowly explanate, dorsally (apically) set with dense fringe of long hair, ventrally with series of short, separate setae. Legs (femora and tibiae) from the front to the back increasingly more compressed and dilated, hind tibiae twice as long as wide; tibial terminal spurs absent; terminal tarsomere of posterior tarsus as long as three preceding ones together; tarsomeres ventrally without setulose pads.
|
||
</paragraph>
|
||
<paragraph id="8B97AA77FF98FF9FA5ECFA17FF4BFDED" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="358">
|
||
Additional description of female
|
||
<typeStatus id="549314D5FF98FF9FA767FA08FD45FE58" box="[588,689,368,392]" pageId="17" pageNumber="358" type="paratype">paratype</typeStatus>
|
||
: The description of the male
|
||
<typeStatus id="549314D5FF98FF9FA124FA17FB85FE58" box="[1039,1137,367,392]" pageId="17" pageNumber="358" type="holotype">holotype</typeStatus>
|
||
specimen fits also to the female
|
||
<typeStatus id="549314D5FF98FF9FA5C6FAEDFEA4FE7D" box="[237,336,405,429]" pageId="17" pageNumber="358" type="paratype">paratype</typeStatus>
|
||
, with a few additions as follows. The pubescence is partially rubbed off; the cephalic pores are virtually absent and at close inspection only recognizable as a minute, longitudinal, slightly darkened structure, possibly a remnant of part of the pores’ margin; eyes little smaller than in male and hardly vaulted; hind tibiae even shorter and wider than in the male; the gonocoxae were dissected, the diverging tips are of normal shape of
|
||
<taxonomicName id="4C28D1F4FF98FF9FA06BF979FA68FDC8" box="[1344,1436,513,536]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="genus" sensu="lato">
|
||
<emphasis id="B95C7665FF98FF9FA06BF979FA68FDC8" box="[1344,1436,513,536]" italics="true" pageId="17" pageNumber="358">Paussus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26FCB1EFF98FF9FA5BCF95CFF4DFDED" box="[151,185,548,573]" pageId="17" pageNumber="358" sensu="lato">s.l.</taxonomicNameLabel>
|
||
;
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C332F9FCFF98FF9FA5ECF93FFEE0FD1C" pageId="17" pageNumber="358" type="biology_ecology">
|
||
<paragraph id="8B97AA77FF98FF9FA5ECF93FFE5FFDB0" blockId="17.[151,1437,151,2013]" box="[199,427,583,608]" pageId="17" pageNumber="358">
|
||
<emphasis id="B95C7665FF98FF9FA5ECF93FFEC2FDB0" bold="true" box="[199,310,583,608]" pageId="17" pageNumber="358">Host ant:</emphasis>
|
||
Unknown
|
||
</paragraph>
|
||
<paragraph id="8B97AA77FF98FF9FA5ECF914FEE0FD1C" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="358">
|
||
<emphasis id="B95C7665FF98FF9FA5ECF914FEDBFD55" bold="true" box="[199,303,620,645]" pageId="17" pageNumber="358">Ecology:</emphasis>
|
||
The female
|
||
<typeStatus id="549314D5FF98FF9FA494F915FDEBFD55" box="[447,543,621,645]" pageId="17" pageNumber="358" type="paratype">paratype</typeStatus>
|
||
specimen was sampled in the African pencil cedar woodland at
|
||
<quantity id="4CD00792FF98FF9FA1D5F914FAA0FD54" box="[1278,1364,620,645]" metricMagnitude="3" metricUnit="m" metricValue="2.578" pageId="17" pageNumber="358" unit="m" value="2578.0">2578 m</quantity>
|
||
in the GRNR (
|
||
<figureCitation id="1313B6F2FF98FF9FA5DCF9F7FEA8FD78" box="[247,348,655,680]" captionStart="FIGURE 3" captionStartId="18.[151,250,1638,1660]" captionTargetBox="[348,1255,336,1597]" captionTargetId="figure@18.[312,1274,303,1616]" captionTargetPageId="18" captionText="FIGURE 3. Photographs of the type locality for paratype of Paussus (Hylotorus) abditus sp. n. in the African pencil cedar woodland at 2578 m, at Garf Raydah Nature Reserve, Asir Province, southwest of the Kingdom of Saudi Arabia." figureDoi="http://doi.org/10.5281/zenodo.3770237" httpUri="https://zenodo.org/record/3770237/files/figure.png" pageId="17" pageNumber="358">Figure 3</figureCitation>
|
||
). Description of habitat see chapter “study area” above. The specimen was recovered from a pitfall trap.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C332F9FCFF98FF9FA5ECF9AFFD57FCE8" pageId="17" pageNumber="358" type="distribution">
|
||
<paragraph id="8B97AA77FF98FF9FA5ECF9AFFEDBFCC4" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="358">
|
||
<emphasis id="B95C7665FF98FF9FA5ECF9AFFE94FD20" bold="true" box="[199,352,727,752]" pageId="17" pageNumber="358">Distribution.</emphasis>
|
||
<collectingCountry id="F33FEAE7FF98FF9FA441F9AFFDF1FD20" box="[362,517,727,752]" name="Saudi Arabia" pageId="17" pageNumber="358">Saudi Arabia</collectingCountry>
|
||
: Hejaz Mountains (exact locality unknown);
|
||
<collectingRegion id="49EC6495FF98FF9FA13BF9AFFBB6FD20" box="[1040,1090,727,752]" country="Saudi Arabia" name="'Asir" pageId="17" pageNumber="358">Asir</collectingRegion>
|
||
, Abha, Raydah,
|
||
<geoCoordinate id="EE1CCCB0FF98FF9FA02EF9AFFA68FD3F" box="[1285,1436,727,752]" degrees="18" direction="north" minutes="12.095" orientation="latitude" pageId="17" pageNumber="358" precision="1" value="18.201584">18°12.095' N</geoCoordinate>
|
||
<geoCoordinate id="EE1CCCB0FF98FF9FA5BCF984FEDFFCC4" box="[151,299,764,789]" degrees="42" direction="east" minutes="24.536" orientation="longitude" pageId="17" pageNumber="358" precision="1" value="42.40893">42°24.536' E</geoCoordinate>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B97AA77FF98FF9FA5ECF867FD57FCE8" blockId="17.[151,1437,151,2013]" box="[199,675,799,824]" pageId="17" pageNumber="358">
|
||
<emphasis id="B95C7665FF98FF9FA5ECF867FEBFFCE8" bold="true" box="[199,331,799,824]" pageId="17" pageNumber="358">Life forms.</emphasis>
|
||
Myrmecophilous (M) species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C332F9FCFF98FF9CA5ECF83CFE29FF27" lastPageId="18" lastPageNumber="359" pageId="17" pageNumber="358" type="discussion">
|
||
<paragraph id="8B97AA77FF98FF9FA5ECF83CFDCAFC3D" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="358">
|
||
<emphasis id="B95C7665FF98FF9FA5ECF83CFECFFC8D" bold="true" box="[199,315,836,861]" pageId="17" pageNumber="358">Remarks.</emphasis>
|
||
The new species is assigned to
|
||
<taxonomicName id="4C28D1F4FF98FF9FA7E2F83DFCD1FC8C" box="[713,805,837,860]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95C7665FF98FF9FA7E2F83DFCD1FC8C" box="[713,805,837,860]" italics="true" pageId="17" pageNumber="358">Paussus</emphasis>
|
||
</taxonomicName>
|
||
subgenus
|
||
<taxonomicName id="4C28D1F4FF98FF9FA68DF83CFB3EFC8C" authority="Dalman, 1823" authorityName="Dalman" authorityYear="1823" box="[934,1226,836,861]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="subGenus" subGenus="Hylotorus">
|
||
<emphasis id="B95C7665FF98FF9FA68DF83CFBE2FC8D" box="[934,1046,836,861]" italics="true" pageId="17" pageNumber="358">Hylotorus</emphasis>
|
||
Dalman, 1823
|
||
</taxonomicName>
|
||
, according to the phylogenetically based classification of
|
||
<bibRefCitation id="EFB9D786FF98FF9FA77CF81FFC69FC50" author="Robertson, J. & Moore, W." box="[599,925,871,896]" pageId="17" pageNumber="358" pagination="134 - 170" refId="ref23316" refString="Robertson, J. A & Moore, W. (2016) Phylogeny of Paussus L. (Carabidae: Paussinae): unravelling morphological convergence associated with myrmecophilous life histories. Systematic Entomology, 42 (1), 134 - 170, figs. S 1 - S 2, tabs. S 1 - S 5 [(January 2017): 28 Sep 2016: version of record online, first published, pp. 170, end of text] https: // doi. org / 10.1111 / syen. 12205" type="journal article" year="2016">Robertson and Moore (2016)</bibRefCitation>
|
||
(cf.
|
||
<bibRefCitation id="EFB9D786FF98FF9FA6FBF81FFB55FC50" author="Nagel, P. & Robertson, J. & Moore, W." box="[976,1185,871,896]" pageId="17" pageNumber="358" pagination="12 - 18" refId="ref22676" refString="Nagel, P., Robertson, J. A & Moore, W. (2017 a) New Nomenclatural and Taxonomic Acts, and Comments. Carabidae: Paussinae: Paussini. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata- Myxophaga-Adephaga. Revised and Updated Edition. Brill, Leiden and Boston, pp. 12 - 18." type="book chapter" year="2017">
|
||
Nagel
|
||
<emphasis id="B95C7665FF98FF9FA131F811FBA6FC50" box="[1050,1106,871,896]" italics="true" pageId="17" pageNumber="358">et al.</emphasis>
|
||
2017a
|
||
</bibRefCitation>
|
||
). The description of
|
||
<taxonomicName id="4C28D1F4FF98FF9FA0ACF811FF1EFC75" authority="Abdel-Dayem & Rasool & Elgharbawy & Nagel & Aldhafer, 2018" authorityName="Abdel-Dayem & Rasool & Elgharbawy & Nagel & Aldhafer" authorityYear="2018" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov.">
|
||
<emphasis id="B95C7665FF98FF9FA0ACF811FF1EFC75" italics="true" pageId="17" pageNumber="358">P. abditus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95C7665FF98FF9FA5DAF8F4FEC6FC75" bold="true" box="[241,306,908,933]" pageId="17" pageNumber="358">
|
||
<taxonomicNameLabel id="A26FCB1EFF98FF9FA5DAF8F4FEC6FC75" box="[241,306,908,933]" pageId="17" pageNumber="358" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
given here above matches generally well the diagnosis and characters used in their key. Deviations are the very long and narrow labial palpomere III and the slightly higher number of setae at the anterior margin of the lacinia in the present new species.
|
||
</paragraph>
|
||
<paragraph id="8B97AA77FF98FF9FA5ECF881FE41FADC" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="358">
|
||
<taxonomicName id="4C28D1F4FF98FF9FA5ECF881FDFEFBC0" authority="Abdel-Dayem & Rasool & Elgharbawy & Nagel & Aldhafer, 2018" authorityName="Abdel-Dayem & Rasool & Elgharbawy & Nagel & Aldhafer" authorityYear="2018" box="[199,522,1015,1040]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Hylotorus">
|
||
<emphasis id="B95C7665FF98FF9FA5ECF881FDFEFBC0" box="[199,522,1015,1040]" italics="true" pageId="17" pageNumber="358">Paussus (Hylotorus) abditus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95C7665FF98FF9FA738F88FFDA1FBC0" bold="true" box="[531,597,1015,1040]" pageId="17" pageNumber="358">
|
||
<taxonomicNameLabel id="A26FCB1EFF98FF9FA738F88FFDA1FBC0" box="[531,597,1015,1040]" pageId="17" pageNumber="358" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is a member of the
|
||
<taxonomicName id="4C28D1F4FF98FF9FA66BF881FC25FBC0" box="[832,977,1015,1040]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="cucullatus">
|
||
<emphasis id="B95C7665FF98FF9FA66BF881FC25FBC0" box="[832,977,1015,1040]" italics="true" pageId="17" pageNumber="358">P. cucullatus</emphasis>
|
||
</taxonomicName>
|
||
group
|
||
<emphasis id="B95C7665FF98FF9FA10EF881FB63FBC0" box="[1061,1175,1015,1040]" italics="true" pageId="17" pageNumber="358">sensu lato</emphasis>
|
||
, an African lineage of
|
||
<taxonomicName id="4C28D1F4FF98FF9FA5BCFF65FE3DFBE5" authorityName="Dalman" authorityYear="1823" box="[151,457,1052,1077]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="subGenus" subGenus="Hylotorus">
|
||
<emphasis id="B95C7665FF98FF9FA5BCFF65FF07FBE4" box="[151,243,1053,1076]" italics="true" pageId="17" pageNumber="358">Paussus</emphasis>
|
||
subgen.
|
||
<emphasis id="B95C7665FF98FF9FA472FF64FE3DFBE5" box="[345,457,1052,1077]" italics="true" pageId="17" pageNumber="358">Hylotorus</emphasis>
|
||
</taxonomicName>
|
||
. The new species is distinguished from the Afrotropical
|
||
<taxonomicName id="4C28D1F4FF98FF9FA17AFF65FB15FBE5" box="[1105,1249,1052,1077]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="cucullatus">
|
||
<emphasis id="B95C7665FF98FF9FA17AFF65FB15FBE5" box="[1105,1249,1052,1077]" italics="true" pageId="17" pageNumber="358">P. cucullatus</emphasis>
|
||
</taxonomicName>
|
||
group s. str. (all species considered in the key of
|
||
<bibRefCitation id="EFB9D786FF98FF9FA4D4FF47FD62FB88" author="Nagel, P." box="[511,662,1087,1112]" pageId="17" pageNumber="358" pagination="181 - 202" refId="ref22612" refString="Nagel, P. (2006) Ant Nest Beetles of the Carnegie Museum (Coleoptera: Carabidae: Paussinae: Paussini). Annals of Carnegie Museum, 75 (3), 181 - 202. https: // doi. org / 10.2992 / 0097 - 4463 (2006) 75 [181: ANBOTC] 2.0. CO; 2" type="journal article" year="2006">Nagel (2006)</bibRefCitation>
|
||
, including the recently described
|
||
<taxonomicName id="4C28D1F4FF98FF9FA127FF39FF25FBAD" authority="Schule & Bednarik, 2015" authorityName="Schule & Bednarik" authorityYear="2015" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="huamboensis">
|
||
<emphasis id="B95C7665FF98FF9FA127FF39FB4DFB88" box="[1036,1209,1087,1112]" italics="true" pageId="17" pageNumber="358">P. huamboensis</emphasis>
|
||
Schüle & Bednařík, 2015
|
||
</taxonomicName>
|
||
) by its tumid antennal club with much smaller excavation, the low anterior part of pronotum and the vestigial cephalic openings.
|
||
<taxonomicName id="4C28D1F4FF98FF9FA451FFF1FDA5FB70" authorityName="Westwood" authorityYear="1833" box="[378,593,1161,1184]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="excavatus">
|
||
<emphasis id="B95C7665FF98FF9FA451FFF1FDA5FB70" box="[378,593,1161,1184]" italics="true" pageId="17" pageNumber="358">Paussus excavatus</emphasis>
|
||
</taxonomicName>
|
||
has an elongated, flattened antennal club and well-developed cephalic openings.
|
||
<taxonomicName id="4C28D1F4FF98FF9FA427FFD5FD93FB15" authority="Kolbe, 1896" authorityName="Kolbe" authorityYear="1896" box="[268,615,1196,1221]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="conradti">
|
||
<emphasis id="B95C7665FF98FF9FA427FFD5FE3BFB15" box="[268,463,1196,1221]" italics="true" pageId="17" pageNumber="358">Paussus conradti</emphasis>
|
||
Kolbe, 1896
|
||
</taxonomicName>
|
||
, also has the cephalic openings present, with the anterior basal angle of the antennal club obliterated and the lower (posterior) pygidial margin set with a series of separated long tufts of hair (
|
||
<bibRefCitation id="EFB9D786FF98FF9FA5FAFF8CFEACFADD" author="Nagel, P." box="[209,344,1268,1293]" pageId="17" pageNumber="358" pagination="181 - 202" refId="ref22612" refString="Nagel, P. (2006) Ant Nest Beetles of the Carnegie Museum (Coleoptera: Carabidae: Paussinae: Paussini). Annals of Carnegie Museum, 75 (3), 181 - 202. https: // doi. org / 10.2992 / 0097 - 4463 (2006) 75 [181: ANBOTC] 2.0. CO; 2" type="journal article" year="2006">Nagel 2006</bibRefCitation>
|
||
;
|
||
<figureCitation id="1313B6F2FF98FF9FA448FF8CFE53FADC" box="[355,423,1268,1293]" captionStart="FIGURE 2" captionStartId="15.[151,250,1898,1920]" captionTargetBox="[403,1206,208,1867]" captionTargetId="figure@15.[364,1228,193,1884]" captionTargetPageId="15" captionText="FIGURE 2. Paussus (Hylotorus) abditus sp. n. holotype specimen from the Hejaz mountains. The illustration shows the specimen with the appendages of the right side at their broadest view, while the left antennae and legs are shown twisted through 90 degrees, i.e. at their narrowest view (cf. Nagel 2006). The inset shows the “collar” of the anterior pronotum at frontal view. Scale bar: 1 mm. Illustration: E. Weber, Basel, Switzerland. Copyright: P. Nagel." figureDoi="http://doi.org/10.5281/zenodo.3770235" httpUri="https://zenodo.org/record/3770235/files/figure.png" pageId="17" pageNumber="358">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B97AA77FF98FF9FA5ECFE61FECAFA4D" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="358">
|
||
<taxonomicName id="4C28D1F4FF98FF9FA5ECFE61FE8BFAE0" authority="Abdel-Dayem & Rasool & Elgharbawy & Nagel & Aldhafer, 2018" authorityName="Abdel-Dayem & Rasool & Elgharbawy & Nagel & Aldhafer" authorityYear="2018" box="[199,383,1303,1328]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov.">
|
||
<emphasis id="B95C7665FF98FF9FA5ECFE61FE8BFAE0" box="[199,383,1303,1328]" italics="true" pageId="17" pageNumber="358">Paussus abditus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95C7665FF98FF9FA4A3FE6FFE3EFAE0" bold="true" box="[392,458,1303,1328]" pageId="17" pageNumber="358">
|
||
<taxonomicNameLabel id="A26FCB1EFF98FF9FA4A3FE6FFE3EFAE0" box="[392,458,1303,1328]" pageId="17" pageNumber="358" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="4C28D1F4FF98FF9FA74DFE61FBE6FAE0" authority="Raffray, 1886" authorityName="Raffray" authorityYear="1886" box="[614,1042,1303,1328]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="cyathiger">
|
||
<emphasis id="B95C7665FF98FF9FA74DFE61FD1AFAE0" box="[614,750,1303,1328]" italics="true" pageId="17" pageNumber="358">P. cyathiger</emphasis>
|
||
<bibRefCitation id="EFB9D786FF98FF9FA7D3FE6FFC60FAE0" author="Raffray, A." box="[760,916,1303,1328]" pageId="17" pageNumber="358" pagination="307 - 359" refId="ref23048" refString="Raffray, A. (1886) Materiaux pour servir a 1 ' etude des coleopteres de la famille des Paussides. Nouvelles Archives du Museum d'Histoire Naturelle, Paris, 2 (8), 307 - 359, pls. 15 - 19." type="journal article" year="1886">Raffray, 1886</bibRefCitation>
|
||
(
|
||
<collectingCountry id="F33FEAE7FF98FF9FA68EFE6FFBFEFAE0" box="[933,1034,1303,1328]" name="Ethiopia" pageId="17" pageNumber="358">Ethiopia</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
, the
|
||
<typeStatus id="549314D5FF98FF9FA160FE6FFB59FAE0" box="[1099,1197,1303,1328]" pageId="17" pageNumber="358" type="holotype">holotype</typeStatus>
|
||
and one
|
||
<typeStatus id="549314D5FF98FF9FA030FE61FA8FFAE1" box="[1307,1403,1305,1329]" pageId="17" pageNumber="358" type="paratype">paratype</typeStatus>
|
||
of which was made available to us due to the courtesy of Ms. Taghavian-Azari (MNHN). This new species differs from
|
||
<taxonomicName id="4C28D1F4FF98FF9FA5FFFE19FEAEFAA8" authorityName="Raffray" authorityYear="1886" box="[212,346,1375,1400]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="cyathiger">
|
||
<emphasis id="B95C7665FF98FF9FA5FFFE19FEAEFAA8" box="[212,346,1375,1400]" italics="true" pageId="17" pageNumber="358">P. cyathiger</emphasis>
|
||
</taxonomicName>
|
||
mainly by the posterior basal angle produced into a broad process rather than a thin, long peg as in
|
||
<emphasis id="B95C7665FF98FF9FA59FFEFDFECAFA4D" box="[180,318,1412,1437]" italics="true" pageId="17" pageNumber="358">
|
||
<taxonomicName id="4C28D1F4FF98FF9FA59FFEFDFECDFA4D" authorityName="Raffray" authorityYear="1886" box="[180,313,1412,1437]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="cyathiger">P. cyathiger</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B97AA77FF98FF9FA5ECFED1FD61F9FD" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="358">
|
||
NB: the specimen illustrated and cited as
|
||
<taxonomicName id="4C28D1F4FF98FF9FA7B7FED1FCD5FA10" authorityName="Raffray" authorityYear="1886" box="[668,801,1447,1472]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="cyathiger">
|
||
<emphasis id="B95C7665FF98FF9FA7B7FED1FCD5FA10" box="[668,801,1447,1472]" italics="true" pageId="17" pageNumber="358">P. cyathiger</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFB9D786FF98FF9FA667FED1FB95FA10" author="Luna de Carvalho, E." box="[844,1121,1447,1472]" pageId="17" pageNumber="358" pagination="377 - 412" refId="ref21994" refString="Luna de Carvalho, E. (1974) Paussideos da Abissina (Coleoptera Carabidae Paussinae). (33 a. Contribuicao para o estudo monografico dos Paussideos). Revue de zoologie et de botanique africaines, 88 (2), 377 - 412." type="journal article" year="1974">Luna de Carvalho (1974</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFB9D786FF98FF9FA146FED0FB52FA10" author="Luna de Carvalho, E." box="[1133,1190,1448,1472]" pageId="17" pageNumber="358" pagination="1 - 1028" refId="ref22042" refString="Luna de Carvalho, E. (1989) Essai monographique des Coleopteres Protopaussines et Paussines. Memorias do Instituto de Investigacao Cientifica Tropical, 2, 70 (1987), 1 - 1028." type="journal article" year="1989">1989</bibRefCitation>
|
||
) is a still undescribed species. The head of the actual
|
||
<taxonomicName id="4C28D1F4FF98FF9FA4D2FEB5FD8BFA35" authorityName="Raffray" authorityYear="1886" box="[505,639,1484,1509]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="cyathiger">
|
||
<emphasis id="B95C7665FF98FF9FA4D2FEB5FD8BFA35" box="[505,639,1484,1509]" italics="true" pageId="17" pageNumber="358">P. cyathiger</emphasis>
|
||
</taxonomicName>
|
||
is devoid of an anterolateral trichome (setose pad), the upper and hind margins of the antennal club are both devoid of tubercles or teeth, and the posterior basal angle of the club is produced into a long, thin, apically blunt peg.
|
||
</paragraph>
|
||
<paragraph id="8B97AA77FF98FF9FA5ECFD41FB3CF948" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="358">
|
||
<taxonomicName id="4C28D1F4FF98FF9FA5ECFD41FE72F980" authority="Abdel-Dayem & Rasool & Elgharbawy & Nagel & Aldhafer, 2018" authorityName="Abdel-Dayem & Rasool & Elgharbawy & Nagel & Aldhafer" authorityYear="2018" box="[199,390,1591,1616]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov.">
|
||
<emphasis id="B95C7665FF98FF9FA5ECFD41FE72F980" box="[199,390,1591,1616]" italics="true" pageId="17" pageNumber="358">Paussus abditus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95C7665FF98FF9FA4BEFD4FFE2AF980" bold="true" box="[405,478,1591,1616]" pageId="17" pageNumber="358">
|
||
<taxonomicNameLabel id="A26FCB1EFF98FF9FA4BEFD4FFE2AF980" box="[405,478,1591,1616]" pageId="17" pageNumber="358" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
seems to be most similar to
|
||
<taxonomicName id="4C28D1F4FF98FF9FA677FD41FA6FF980" authority="Lorenz, 1998" authorityName="Lorenz" authorityYear="1998" box="[860,1435,1592,1616]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="rougemontianus">
|
||
<emphasis id="B95C7665FF98FF9FA677FD41FB89F980" box="[860,1149,1593,1616]" italics="true" pageId="17" pageNumber="358">Paussus rougemontianus</emphasis>
|
||
Lorenz, 1998 (
|
||
<collectingCountry id="F33FEAE7FF98FF9FA068FD40FA61F980" box="[1347,1429,1592,1616]" name="Yemen" pageId="17" pageNumber="358">Yemen</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
(replacement name for
|
||
<taxonomicName id="4C28D1F4FF98FF9FA495FD24FBBFF9A4" authority="Luna de Carvalho, 1989" authorityName="Luna de Carvalho" authorityYear="1989" box="[446,1099,1628,1653]" class="Insecta" family="Carabidae" genus="Cochliopaussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="rougemonti">
|
||
<emphasis id="B95C7665FF98FF9FA495FD24FCF7F9A4" box="[446,771,1628,1653]" italics="true" pageId="17" pageNumber="358">Cochliopaussus rougemonti</emphasis>
|
||
<bibRefCitation id="EFB9D786FF98FF9FA63EFD25FBBFF9A4" author="Luna de Carvalho, E." box="[789,1099,1628,1653]" pageId="17" pageNumber="358" pagination="1 - 1028" refId="ref22042" refString="Luna de Carvalho, E. (1989) Essai monographique des Coleopteres Protopaussines et Paussines. Memorias do Instituto de Investigacao Cientifica Tropical, 2, 70 (1987), 1 - 1028." type="journal article" year="1989">Luna de Carvalho, 1989</bibRefCitation>
|
||
</taxonomicName>
|
||
). The antennal club of
|
||
<taxonomicName id="4C28D1F4FF98FF9FA0ACFD25FEB8F948" authorityName="Lorenz" authorityYear="1998" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="rougemontianus">
|
||
<emphasis id="B95C7665FF98FF9FA0ACFD25FEB8F948" italics="true" pageId="17" pageNumber="358">P. rougemontianus</emphasis>
|
||
</taxonomicName>
|
||
has the posterior excavation shallow and restricted to the basal half of the club.
|
||
</paragraph>
|
||
<paragraph id="8B97AA77FF98FF9FA5ECFDDCFA88F869" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="358">
|
||
The other Arabian member of
|
||
<taxonomicName id="4C28D1F4FF98FF9FA730FDDDFD83F96C" box="[539,631,1701,1724]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95C7665FF98FF9FA730FDDDFD83F96C" box="[539,631,1701,1724]" italics="true" pageId="17" pageNumber="358">Paussus</emphasis>
|
||
</taxonomicName>
|
||
subgenus
|
||
<taxonomicName id="4C28D1F4FF98FF9FA7C6FDDCFCA9F96D" authorityName="Dalman" authorityYear="1823" box="[749,861,1700,1725]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="subGenus" subGenus="Hylotorus">
|
||
<emphasis id="B95C7665FF98FF9FA7C6FDDCFCA9F96D" box="[749,861,1700,1725]" italics="true" pageId="17" pageNumber="358">Hylotorus</emphasis>
|
||
</taxonomicName>
|
||
is
|
||
<taxonomicName id="4C28D1F4FF98FF9FA656FDDDFB47F96D" authority="Raffray, 1886" authorityName="Raffray" authorityYear="1886" box="[893,1203,1700,1725]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="cephalotes">
|
||
<emphasis id="B95C7665FF98FF9FA656FDDDFBE5F96D" box="[893,1041,1700,1725]" italics="true" pageId="17" pageNumber="358">P. cephalotes</emphasis>
|
||
<bibRefCitation id="EFB9D786FF98FF9FA13CFDDCFB47F96D" author="Raffray, A." box="[1047,1203,1700,1725]" pageId="17" pageNumber="358" pagination="307 - 359" refId="ref23048" refString="Raffray, A. (1886) Materiaux pour servir a 1 ' etude des coleopteres de la famille des Paussides. Nouvelles Archives du Museum d'Histoire Naturelle, Paris, 2 (8), 307 - 359, pls. 15 - 19." type="journal article" year="1886">Raffray, 1886</bibRefCitation>
|
||
</taxonomicName>
|
||
. This bizarre species is unmistakable by the shape of the antennal club and the two horn-like projections of the head. All Arabian
|
||
<taxonomicName id="4C28D1F4FF98FF9FA5BCFD94FEF1F8D5" box="[151,261,1772,1797]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="subFamily" subFamily="Paussinae">Paussinae</taxonomicName>
|
||
are members of the species-rich genus
|
||
<taxonomicName id="4C28D1F4FF98FF9FA7FBFD95FC89F8D5" authority="L. Till" authorityName="L. Till" box="[720,893,1772,1797]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95C7665FF98FF9FA7FBFD95FCD8F8D4" box="[720,812,1773,1796]" italics="true" pageId="17" pageNumber="358">Paussus</emphasis>
|
||
L. Till
|
||
</taxonomicName>
|
||
date 8 species were reported from the Arabian Peninsula
|
||
<emphasis id="B95C7665FF98FF9FA427FC69FE8AF8F8" box="[268,382,1807,1832]" italics="true" pageId="17" pageNumber="358">sensu lato</emphasis>
|
||
, as follows (
|
||
<bibRefCitation id="EFB9D786FF98FF9FA73BFC77FD6CF8F8" author="Nagel, P." box="[528,664,1807,1832]" pageId="17" pageNumber="358" pagination="99 - 107" refId="ref22579" refString="Nagel, P. (1982) Insects of Saudi Arabia. Coleoptera: Fam. Carabidae, subfam. Paussinae. Fauna of Saudi Arabia, 4, 99 - 107." type="journal article" year="1982">Nagel 1982</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB9D786FF98FF9FA78EFC68FC36F8F8" author="Luna de Carvalho, E." box="[677,962,1807,1832]" pageId="17" pageNumber="358" pagination="51 - 52" refId="ref22133" refString="Luna de Carvalho, E. (2000 b) Nova contribuicao para o estudo dos Carabideos Paussineos do Yemen (Insecta: Carabidae: Paussinae). Boletin de la Sociedad Entomologica Aragonesa, 27, 51 - 52." type="journal article" year="2000">Luna de Carvalho 2000b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB9D786FF98FF9FA6E4FC77FB53F8F8" author="Nagel, P. & Robertson, J. A. & Moore, W." box="[975,1191,1807,1832]" pageId="17" pageNumber="358" pagination="466 - 470" refId="ref22755" refString="Nagel, P., Robertson, J. A. & Moore, W. (2017 b) Catalogue, family Carabidae Latreille, 1802, subfamily Paussinae Latreille, 1806. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata-Myxophaga-Adephaga. Revised and Updated Edition. Brill, Leiden and Boston, pp. 466 - 470." type="book chapter" year="2017">
|
||
Nagel
|
||
<emphasis id="B95C7665FF98FF9FA131FC69FBA0F8F8" box="[1050,1108,1807,1832]" italics="true" pageId="17" pageNumber="358">et al.</emphasis>
|
||
2017b
|
||
</bibRefCitation>
|
||
):
|
||
<taxonomicName id="4C28D1F4FF98FF9FA190FC69FF27F89D" authority="Raffray, 1886" authorityName="Raffray" authorityYear="1886" class="Insecta" family="Carabidae" genus="Metadromius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="arabicus">
|
||
<emphasis id="B95C7665FF98FF9FA190FC69FACDF8F8" box="[1211,1337,1807,1832]" italics="true" pageId="17" pageNumber="358">P. arabicus</emphasis>
|
||
<bibRefCitation id="EFB9D786FF98FF9FA06AFC77FF27F89D" author="Raffray, A." pageId="17" pageNumber="358" pagination="307 - 359" refId="ref23048" refString="Raffray, A. (1886) Materiaux pour servir a 1 ' etude des coleopteres de la famille des Paussides. Nouvelles Archives du Museum d'Histoire Naturelle, Paris, 2 (8), 307 - 359, pls. 15 - 19." type="journal article" year="1886">Raffray, 1886</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C28D1F4FF98FF9FA5F5FC4DFDBEF89D" authority="Reichensperger, 1957" authorityName="Reichensperger" authorityYear="1957" box="[222,586,1844,1869]" class="Insecta" family="Carabidae" genus="Metadromius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="brittoni">
|
||
<emphasis id="B95C7665FF98FF9FA5F5FC4DFEBAF89D" box="[222,334,1844,1869]" italics="true" pageId="17" pageNumber="358">P. brittoni</emphasis>
|
||
Reichensperger, 1957
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C28D1F4FF98FF9FA77DFC4DFC7AF89D" authority="Raffray, 1886" authorityName="Raffray" authorityYear="1886" box="[598,910,1844,1869]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="cephalotes">
|
||
<emphasis id="B95C7665FF98FF9FA77DFC4DFD1EF89D" box="[598,746,1844,1869]" italics="true" pageId="17" pageNumber="358">P. cephalotes</emphasis>
|
||
<bibRefCitation id="EFB9D786FF98FF9FA7DAFC4CFC7AF89D" author="Raffray, A." box="[753,910,1844,1869]" pageId="17" pageNumber="358" pagination="307 - 359" refId="ref23048" refString="Raffray, A. (1886) Materiaux pour servir a 1 ' etude des coleopteres de la famille des Paussides. Nouvelles Archives du Museum d'Histoire Naturelle, Paris, 2 (8), 307 - 359, pls. 15 - 19." type="journal article" year="1886">Raffray, 1886</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C28D1F4FF98FF9FA6B1FC4DFB5BF89C" authority="Fiori, 1914" authorityName="Fiori" authorityYear="1914" box="[922,1199,1844,1869]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="cirenaicus">
|
||
<emphasis id="B95C7665FF98FF9FA6B1FC4DFBDDF89C" box="[922,1065,1845,1868]" italics="true" pageId="17" pageNumber="358">P. cirenaicus</emphasis>
|
||
Fiori, 1914
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C28D1F4FF98FF9FA197FC4DFF27F8A0" authority="Saulcy, 1874" authorityName="Saulcy" authorityYear="1874" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="piochardi">
|
||
<emphasis id="B95C7665FF98FF9FA197FC4DFAB7F89D" box="[1212,1347,1844,1869]" italics="true" pageId="17" pageNumber="358">P. piochardi</emphasis>
|
||
Saulcy, 1874
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C28D1F4FF98FF9FA5F6FC21FDB8F8A0" authority="Lorenz, 1998" authorityName="Lorenz" authorityYear="1998" box="[221,588,1880,1904]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="rougemontianus">
|
||
<emphasis id="B95C7665FF98FF9FA5F6FC21FE5AF8A0" box="[221,430,1881,1904]" italics="true" pageId="17" pageNumber="358">P. rougemontianus</emphasis>
|
||
Lorenz, 1998
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C28D1F4FF98FF9FA773FC21FC74F8A0" authority="Reiche, 1860" authorityName="Reiche" authorityYear="1860" box="[600,896,1879,1904]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="thomsonii">
|
||
<emphasis id="B95C7665FF98FF9FA773FC21FD16F8A0" box="[600,738,1879,1904]" italics="true" pageId="17" pageNumber="358">P. thomsonii</emphasis>
|
||
Reiche, 1860
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C28D1F4FF98FF9FA6A0FC21FB28F8A0" authority="(Frivaldszky 1835)" baseAuthorityName="Frivaldszky" baseAuthorityYear="1835" box="[907,1244,1879,1904]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="turcicus">
|
||
<emphasis id="B95C7665FF98FF9FA6A0FC21FC08F8A0" box="[907,1020,1881,1904]" italics="true" pageId="17" pageNumber="358">P. turcicus</emphasis>
|
||
(Frivaldszky 1835)
|
||
</taxonomicName>
|
||
(the latter has its southernmost localities in
|
||
<collectingCountry id="F33FEAE7FF98FF9FA497FC04FE0EF845" box="[444,506,1916,1941]" name="Israel" pageId="17" pageNumber="358">Israel</collectingCountry>
|
||
and
|
||
<collectingCountry id="F33FEAE7FF98FF9FA71BFC04FD74F845" box="[560,640,1916,1941]" name="Jordan" pageId="17" pageNumber="358">Jordan</collectingCountry>
|
||
). The specific identity of
|
||
<taxonomicName id="4C28D1F4FF98FF9FA6B1FC05FBD5F845" authorityName="Saulcy" authorityYear="1874" box="[922,1057,1916,1941]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="piochardi">
|
||
<emphasis id="B95C7665FF98FF9FA6B1FC05FBD5F845" box="[922,1057,1916,1941]" italics="true" pageId="17" pageNumber="358">P. piochardi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C28D1F4FF98FF9FA17DFC05FB15F844" authorityName="Fiori" authorityYear="1914" box="[1110,1249,1917,1940]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="cirenaicus">
|
||
<emphasis id="B95C7665FF98FF9FA17DFC05FB15F844" box="[1110,1249,1917,1940]" italics="true" pageId="17" pageNumber="358">P. cirenaicus</emphasis>
|
||
</taxonomicName>
|
||
within the more inclusive
|
||
<taxonomicName id="4C28D1F4FF98FF9FA428FCD9FDA7F869" authority="Raffray, 1886" authorityName="Raffray" authorityYear="1886" box="[259,595,1952,1977]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="crenaticornis">
|
||
<emphasis id="B95C7665FF98FF9FA428FCD9FE46F868" box="[259,434,1953,1976]" italics="true" pageId="17" pageNumber="358">P. crenaticornis</emphasis>
|
||
<bibRefCitation id="EFB9D786FF98FF9FA491FCD8FDA7F869" author="Raffray, A." box="[442,595,1952,1977]" pageId="17" pageNumber="358" pagination="307 - 359" refId="ref23048" refString="Raffray, A. (1886) Materiaux pour servir a 1 ' etude des coleopteres de la famille des Paussides. Nouvelles Archives du Museum d'Histoire Naturelle, Paris, 2 (8), 307 - 359, pls. 15 - 19." type="journal article" year="1886">Raffray, 1886</bibRefCitation>
|
||
</taxonomicName>
|
||
group is debated.
|
||
<taxonomicName id="4C28D1F4FF98FF9FA60AFCD9FC7AF869" authority="Abdel-Dayem & Rasool & Elgharbawy & Nagel & Aldhafer, 2018" authorityName="Abdel-Dayem & Rasool & Elgharbawy & Nagel & Aldhafer" authorityYear="2018" box="[801,910,1952,1977]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov.">
|
||
<emphasis id="B95C7665FF98FF9FA60AFCD9FC7AF869" box="[801,910,1952,1977]" italics="true" pageId="17" pageNumber="358">P. abditus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95C7665FF98FF9FA6BDFCD8FC21F869" bold="true" box="[918,981,1952,1977]" pageId="17" pageNumber="358">
|
||
<taxonomicNameLabel id="A26FCB1EFF98FF9FA6BDFCD8FC21F869" box="[918,981,1952,1977]" pageId="17" pageNumber="358" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
increases the number to nine species.
|
||
</paragraph>
|
||
<paragraph id="8B97AA77FF98FF9CA5ECFCBCFE29FF27" blockId="17.[151,1437,151,2013]" lastBlockId="18.[151,1437,151,248]" lastPageId="18" lastPageNumber="359" pageId="17" pageNumber="358">
|
||
According to the current state of knowledge four species are exclusively known from the Arabian Peninsula
|
||
<emphasis id="B95C7665FF9BFF9CA5BCFBE1FEFBFF60" box="[151,271,151,176]" italics="true" pageId="18" pageNumber="359">sensu lato</emphasis>
|
||
and are possibly endemics (END_AR):
|
||
<taxonomicName id="4C28D1F4FF9BFF9CA7D7FBE1FC85FF60" authority="Abdel-Dayem & Rasool & Elgharbawy & Nagel & Aldhafer, 2018" authorityName="Abdel-Dayem & Rasool & Elgharbawy & Nagel & Aldhafer" authorityYear="2018" box="[764,881,151,176]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="359" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov.">
|
||
<emphasis id="B95C7665FF9BFF9CA7D7FBE1FC85FF60" box="[764,881,151,176]" italics="true" pageId="18" pageNumber="359">P. abditus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95C7665FF9BFF9CA654FBEFFC31FF60" bold="true" box="[895,965,151,176]" pageId="18" pageNumber="359">
|
||
<taxonomicNameLabel id="A26FCB1EFF9BFF9CA654FBEFFC31FF60" box="[895,965,151,176]" pageId="18" pageNumber="359" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(only known from SA),
|
||
<taxonomicName id="4C28D1F4FF9BFF9CA1D7FBE1FA86FF60" authorityName="Reichensperger" authorityYear="1957" box="[1276,1394,151,176]" class="Insecta" family="Carabidae" genus="Metadromius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="359" phylum="Arthropoda" rank="species" species="brittoni">
|
||
<emphasis id="B95C7665FF9BFF9CA1D7FBE1FA86FF60" box="[1276,1394,151,176]" italics="true" pageId="18" pageNumber="359">P. brittoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B95C7665FF9BFF9CA0ACFBE1FE18FF04" italics="true" pageId="18" pageNumber="359">
|
||
<taxonomicName id="4C28D1F4FF9BFF9CA0ACFBE1FEE5FF05" authorityName="Raffray" authorityYear="1886" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="359" phylum="Arthropoda" rank="species" species="cephalotes">P. cephalotes</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C28D1F4FF9BFF9CA437FBC5FE18FF04" authorityName="Lorenz" authorityYear="1998" box="[284,492,189,212]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="359" phylum="Arthropoda" rank="species" species="rougemontianus">P. rougemontianus</taxonomicName>
|
||
</emphasis>
|
||
. Their range comprises
|
||
<collectingCountry id="F33FEAE7FF9BFF9CA7D1FBC5FCBCFF04" box="[762,840,189,212]" name="Yemen" pageId="18" pageNumber="359">Yemen</collectingCountry>
|
||
and the Hejaz and
|
||
<collectingRegion id="49EC6495FF9BFF9CA135FBC4FBBAFF05" box="[1054,1102,188,213]" country="Saudi Arabia" name="'Asir" pageId="18" pageNumber="359">Asir</collectingRegion>
|
||
mountains in
|
||
<collectingCountry id="F33FEAE7FF9BFF9CA1C6FBC4FAD1FF04" box="[1261,1317,188,212]" name="Saudi Arabia" pageId="18" pageNumber="359">KSA</collectingCountry>
|
||
which run parallel to the Red Sea coast.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |