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<document id="86BEEF54A896121E3E1F1EEFE75766DD" ID-DOI="10.5281/zenodo.209019" ID-GBIF-Dataset="69418131-0628-4fa0-8601-43960ff6e45f" ID-ISSN="1175-5326" ID-Zenodo-Dep="209019" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460517985539" checkinUser="plazi" docAuthor="Hippa, Heikki &amp; Kurina, Olavi" docDate="2012" docId="D9558784FFE7FFEA1EDFFA355BBFFCAD" docLanguage="en" docName="zt03455p048.pdf" docOrigin="Zootaxa 3455" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Manota dissidens Hippa &amp; Kurina, 2012, sp.n." docType="treatment" docVersion="9" lastPageNumber="16" masterDocId="256CFFFCFFE9FFE51E48FF8D5F71FFF1" masterDocTitle="New species and new records of Afrotropical Manota Williston (Diptera, Mycetophilidae), with a key to the species" masterLastPageNumber="48" masterPageNumber="1" pageNumber="15" updateTime="1698314294007" updateUser="plazi">
<mods:mods id="F8DEE1A9267293E426875F4378A3C1BE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D30F438ADDE2254056F306E0507286BA">
<mods:title id="73C38E50B0660B9AECE82DDD34C23648">New species and new records of Afrotropical Manota Williston (Diptera, Mycetophilidae), with a key to the species</mods:title>
</mods:titleInfo>
<mods:name id="3C8F87C9263955EB0B6443DFC994398D" type="personal">
<mods:role id="A8C56A966201C1F8553E558265132202">
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</mods:role>
<mods:namePart id="F9D151EB412668C1C9B4C157779D3B00">Hippa, Heikki</mods:namePart>
</mods:name>
<mods:name id="4988F6771BFFEF29AB0B602D02DD3487" type="personal">
<mods:role id="E15B91D13BD04AAC197FA97BA1370CFF">
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<mods:namePart id="F8E99024197C3FFDAEFF8D237857CBF6">Kurina, Olavi</mods:namePart>
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<mods:title id="E8F08F40E31F52B3626C7E20B7EFC6C4">Zootaxa</mods:title>
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<mods:part id="9E7FB65C63035A468682FA87E0449AC8">
<mods:date id="0F1C72B02CC85CDF3A5910A8869A79CF">2012</mods:date>
<mods:detail id="2EBACCC0A34192357458F522E415696F" type="volume">
<mods:number id="3E0AFEC3BED992E1A14E1F7392C122DF">3455</mods:number>
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<mods:classification id="0E34D60334AE210945DD9037C954AB9B">journal article</mods:classification>
<mods:identifier id="CD2C317DECFB680B5ECA4878B74A47AB" type="DOI">10.5281/zenodo.209019</mods:identifier>
<mods:identifier id="CC0EF41C48226C281B51A7019E1389E5" type="GBIF-Dataset">69418131-0628-4fa0-8601-43960ff6e45f</mods:identifier>
<mods:identifier id="11928482FFF4772855CB1DC368AEC453" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8BE1E3BC8104E30D91733A2F3621E32E" type="Zenodo-Dep">209019</mods:identifier>
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<treatment id="D9558784FFE7FFEA1EDFFA355BBFFCAD" ID-DOI="http://doi.org/10.5281/zenodo.6177567" ID-GBIF-Taxon="119524784" ID-Zenodo-Dep="6177567" LSID="urn:lsid:plazi:treatment:D9558784FFE7FFEA1EDFFA355BBFFCAD" httpUri="http://treatment.plazi.org/id/D9558784FFE7FFEA1EDFFA355BBFFCAD" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<subSubSection id="19E66519FFE7FFEA1EDFFA355EE1FD56" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="51433692FFE7FFEB1EDFFA355EDAFA23" blockId="14.[151,427,1464,1524]" box="[151,427,1464,1490]" pageId="14" pageNumber="15">
<heading id="0A0B81FEFFE7FFEB1EDFFA355EDAFA23" bold="true" box="[151,427,1464,1490]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="6388EA80FFE7FFEB1EDFFA355EDAFA23" bold="true" box="[151,427,1464,1490]" pageId="14" pageNumber="15">
<taxonomicName id="96FC4D11FFE7FFEB1EDFFA355E18FA23" box="[151,361,1464,1490]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="dissidens" status="sp. nov.">
<emphasis id="6388EA80FFE7FFEB1EDFFA355E18FA23" bold="true" box="[151,361,1464,1490]" italics="true" pageId="14" pageNumber="15">Manota dissidens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="78BB57FBFFE7FFEB1F38FA355EDAFA23" box="[368,427,1464,1490]" pageId="14" pageNumber="15" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="51433692FFE7FFEB1EDFFA565E6BFA05" blockId="14.[151,427,1464,1524]" box="[151,282,1499,1524]" pageId="14" pageNumber="15">
<figureCitation id="C9C72A17FFE7FFEB1EDFFA565FADFA02" box="[151,220,1499,1523]" captionStart="FIGURE 8" captionStartId="14.[151,250,1330,1353]" captionTargetBox="[197,1377,208,1233]" captionTargetId="figure@14.[188,1398,195,1307]" captionTargetPageId="14" captionText="FIGURE 8. Manota dissidens sp. n. (holotype). A. Antennal flagellomere 4, lateral view. B. Hypopygium, ventral view. C. Hypopygium, dorsal view. D. Gonostylus, dorsal view. Scale 0.1 mm." httpUri="https://zenodo.org/record/209027/files/figure.png" pageId="14" pageNumber="15">Figs 8</figureCitation>
A–D
</paragraph>
<paragraph id="51433692FFE7FFEA1EDFF9AE5CF6FD75" blockId="14.[151,1436,1571,2027]" lastBlockId="15.[151,1436,151,860]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
Male.
<emphasis id="6388EA80FFE7FFEB1E96F9AE5E41F9CD" bold="true" box="[222,304,1571,1596]" pageId="14" pageNumber="15">Colour</emphasis>
. Both studied specimens seem to be rather faded. Head pale brown, vertex and dorsal part of occiput dark brown. Antenna brown, scapus and pedicellus may be paler than the more distal parts. Mouthparts pale yellowish. Thorax pale brown. Legs pale yellowish, apical third of femur 3 probably infuscated but the colour is very difficult to observe in both slides. Wing pale brown; haltere pale brown with dark brown knob. Abdomen brown, sternites paler than tergites. All setosity pale, yellowish or brownish, the thicker setae seeming darker than the finer setae and trichia.
<emphasis id="6388EA80FFE7FFEB1F94F95A5D6DF91E" bold="true" box="[476,540,1751,1775]" pageId="14" pageNumber="15">Head</emphasis>
. Antennal flagellomere 4,
<figureCitation id="C9C72A17FFE7FFEB1D16F95A5CD4F91E" box="[862,933,1751,1775]" captionStart="FIGURE 8" captionStartId="14.[151,250,1330,1353]" captionTargetBox="[197,1377,208,1233]" captionTargetId="figure@14.[188,1398,195,1307]" captionTargetPageId="14" captionText="FIGURE 8. Manota dissidens sp. n. (holotype). A. Antennal flagellomere 4, lateral view. B. Hypopygium, ventral view. C. Hypopygium, dorsal view. D. Gonostylus, dorsal view. Scale 0.1 mm." httpUri="https://zenodo.org/record/209027/files/figure.png" pageId="14" pageNumber="15">Fig. 8</figureCitation>
A. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 5 apically expanded curved sensilla; palpomere 4 with parasegment, palpomere 5 ca. 1.4 times longer than palpomere 4. Number of strong postocular setae 10–11.
<emphasis id="6388EA80FFE7FFEB1B74F8925AE4F8C6" bold="true" box="[1340,1429,1823,1847]" pageId="14" pageNumber="15">Thorax</emphasis>
. Anepisternum setose, with 45–55 setae; anterior basalare non-setose; preepisternum 2 setose, with ca. 15 setae; laterotergite non-setose; episternum 3 setose, with ca. 18 setae.
<emphasis id="6388EA80FFE7FFEB1D28F8EA5CE9F88E" bold="true" box="[864,920,1895,1919]" pageId="14" pageNumber="15">Legs</emphasis>
. Mid and hind tibial organs absent.
<emphasis id="6388EA80FFE7FFEB1B64F8EB5A1DF88E" bold="true" box="[1324,1388,1894,1919]" pageId="14" pageNumber="15">Wing</emphasis>
. R1 meeting C well on the basal half of the costal margin; the sclerotized part of M extending to the level of the tip of R1, wing length
<quantity id="96049B77FFE7FFEB1F11F8225E97F836" box="[345,486,1967,1991]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="14" pageNumber="15" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.5–1.7 mm</quantity>
.
<emphasis id="6388EA80FFE7FFEB1FBBF8225DE4F839" bold="true" box="[499,661,1967,1992]" pageId="14" pageNumber="15">Hypopygium,</emphasis>
<figureCitation id="C9C72A17FFE7FFEB1CD7F8225D96F836" box="[671,743,1967,1991]" captionStart="FIGURE 8" captionStartId="14.[151,250,1330,1353]" captionTargetBox="[197,1377,208,1233]" captionTargetId="figure@14.[188,1398,195,1307]" captionTargetPageId="14" captionText="FIGURE 8. Manota dissidens sp. n. (holotype). A. Antennal flagellomere 4, lateral view. B. Hypopygium, ventral view. C. Hypopygium, dorsal view. D. Gonostylus, dorsal view. Scale 0.1 mm." httpUri="https://zenodo.org/record/209027/files/figure.png" pageId="14" pageNumber="15">Figs 8</figureCitation>
B–D: Sternite 9 half the length of gonocoxa, lateral margin sharply delimited, posterior margin transverse, anterior margin deeply incised, the setae similar to the ventral setae of gonocoxa. Ventral mesial margin of gonocoxa simple, slightly angled at the middle. Parastylar lobe subtriangular with 2 setae at anterior end, the setae arising from small basal bodies. Paraapodemal lobe distinct, exposed in ventral view. Dorsal mesial margin of gonocoxa simple, convex, posteriorly with an aggregation of setae deviating from the other gonocoxal setosity; posterior margin simple, forming an obtuse rounded angle with the mesial margin. On the ventral side of the latter there is a finger-like lobe bearing a strong apical seta. Two juxtagonostylar setae present, both rather unmodified blunted megasetae arising from a common basal body which is almost as long as the megasetae. Gonostylus about half the length of gonocoxa, roughly sickle-shaped with pointed apex and a heel-like basomesial angle and a lobe-like crest laterodorsally; there are two long and strong setae at the basomesial angle, one similar at the middle of the mesial side and two shorter ones at the apex; the setae on the ventral surface similar to those on the gonocoxa, on the dorsal side short and numerous mesially, laterally largely lacking except for the crest which has a couple of long setae distally. Aedeagus elongate subtriangular, with lateral shoulders, the apex curved ventrad. Hypoproct posteriorly extending to the basal part of gonostylus, the ventral setae forming a mesial longitudinal stripe of ca. 5 setae on each half. Cerci medially separate, simple, the setae short, the longest ones about twice the apical width of cercus.
</paragraph>
<paragraph id="51433692FFE6FFEA1E8FFD025EE1FD56" blockId="15.[151,1436,151,860]" box="[199,400,655,679]" pageId="15" pageNumber="16">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="19E66519FFE6FFEA1E8FFD3E5AF0FD1E" pageId="15" pageNumber="16" type="discussion">
<paragraph id="51433692FFE6FFEA1E8FFD3E5AF0FD1E" blockId="15.[151,1436,151,860]" pageId="15" pageNumber="16">
<emphasis id="6388EA80FFE6FFEA1E8FFD3E5E34FD3D" bold="true" box="[199,325,691,716]" pageId="15" pageNumber="16">Discussion</emphasis>
. In the key
<taxonomicName id="96FC4D11FFE6FFEA1F8EFD385DF9FD3D" box="[454,648,692,716]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="dissidens">
<emphasis id="6388EA80FFE6FFEA1F8EFD385DF9FD3D" box="[454,648,692,716]" italics="true" pageId="15" pageNumber="16">Manota dissidens</emphasis>
</taxonomicName>
follows
<taxonomicName id="96FC4D11FFE6FFEA1CA0FD385C17FD3D" box="[744,870,692,716]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="redunca">
<emphasis id="6388EA80FFE6FFEA1CA0FD385C17FD3D" box="[744,870,692,716]" italics="true" pageId="15" pageNumber="16">M. redunca</emphasis>
</taxonomicName>
and
<taxonomicName id="96FC4D11FFE6FFEA1DD3FD385B66FD3D" box="[923,1047,693,716]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="toomasi">
<emphasis id="6388EA80FFE6FFEA1DD3FD385B66FD3D" box="[923,1047,693,716]" italics="true" pageId="15" pageNumber="16">M. toomasi</emphasis>
</taxonomicName>
but is distinguished from them and any other species particularly by the 2 + 1 long mesial setae and the dorsolateral lobe on its gonostylus (
<figureCitation id="C9C72A17FFE6FFEA1B5CFD5A5A27FD1E" box="[1300,1366,727,751]" captionStart="FIGURE 8" captionStartId="14.[151,250,1330,1353]" captionTargetBox="[197,1377,208,1233]" captionTargetId="figure@14.[188,1398,195,1307]" captionTargetPageId="14" captionText="FIGURE 8. Manota dissidens sp. n. (holotype). A. Antennal flagellomere 4, lateral view. B. Hypopygium, ventral view. C. Hypopygium, dorsal view. D. Gonostylus, dorsal view. Scale 0.1 mm." httpUri="https://zenodo.org/record/209027/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
D).
</paragraph>
</subSubSection>
<subSubSection id="19E66519FFE6FFEA1E8FFD715BBFFCAD" pageId="15" pageNumber="16" type="etymology">
<paragraph id="51433692FFE6FFEA1E8FFD715A6AFCE5" blockId="15.[151,1436,151,860]" box="[199,1307,764,788]" pageId="15" pageNumber="16">
<emphasis id="6388EA80FFE6FFEA1E8FFD715E3FFCE5" bold="true" box="[199,334,764,788]" pageId="15" pageNumber="16">Etymology.</emphasis>
The name is Latin,
<taxonomicName id="96FC4D11FFE6FFEA1C65FD715DE5FCE5" box="[557,660,764,788]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="dissidens">
<emphasis id="6388EA80FFE6FFEA1C65FD715DE5FCE5" box="[557,660,764,788]" italics="true" pageId="15" pageNumber="16">dissidens</emphasis>
</taxonomicName>
, differing, referring to the unusual gonostylar chaetotaxy.
</paragraph>
<paragraph id="51433692FFE6FFEA1E8FFCAD5BBFFCAD" blockId="15.[151,1436,151,860]" pageId="15" pageNumber="16">
<emphasis id="6388EA80FFE6FFEA1E8FFCAD5E7BFCC9" bold="true" box="[199,266,800,824]" pageId="15" pageNumber="16">
<typeStatus id="8E478830FFE6FFEA1E8FFCAD5E7BFCC9" box="[199,266,800,824]" pageId="15" pageNumber="16">Types</typeStatus>
</emphasis>
.
<emphasis id="6388EA80FFE6FFEA1F53FCAD5EF3FCC9" box="[283,386,800,824]" italics="true" pageId="15" pageNumber="16">
<typeStatus id="8E478830FFE6FFEA1F53FCAD5EF3FCC9" box="[283,386,800,824]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
</emphasis>
. Male,
<collectingCountry id="29EB7602FFE6FFEA1F90FC925D31FCC6" box="[472,576,799,823]" name="Ghana" pageId="15" pageNumber="16">GHANA</collectingCountry>
, Central Region, Kakum nr. Entwi Hrom, Malaise trap
<date id="25421052FFE6FFEA1AFBFC925A39FCC6" box="[1203,1352,799,823]" pageId="15" pageNumber="16" value="1999-11-08" valueMax="1999-11-15" valueMin="1999-11-08">8–15.xi.1999</date>
, leg. T. Andersson (on slide, in MZLU).
<emphasis id="6388EA80FFE6FFEA1C40FCC85D0BFCAD" box="[520,634,837,860]" italics="true" pageId="15" pageNumber="16">
<typeStatus id="8E478830FFE6FFEA1C40FCC85D0BFCAD" box="[520,634,837,860]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<emphasis id="6388EA80FFE6FFEA1CC0FCCE5DA7FCAD" bold="true" box="[648,726,835,860]" pageId="15" pageNumber="16">
<specimenCount id="47FAFD1BFFE6FFEA1CC0FCCE5DA7FCAD" box="[648,726,835,860]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
</emphasis>
, same data as
<typeStatus id="8E478830FFE6FFEA1D31FCC95CAAFCAD" box="[889,987,836,860]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
(on slide, in MZLU).
</paragraph>
</subSubSection>
</treatment>
</document>
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