261 lines
60 KiB
XML
261 lines
60 KiB
XML
<document id="5AC337309732DBEED38CA982A5A86A0B" ID-CLB-Dataset="55625" ID-DOI="10.11646/zootaxa.5133.3.3" ID-GBIF-Dataset="02d6b967-3efb-4c75-96d3-e9ac8541bb3c" ID-ISSN="1175-5326" ID-Zenodo-Dep="6522677" ID-ZooBank="44C7B333-7D3E-4235-A158-60E70849ED60" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1651756637829" checkinUser="plazi" docAuthor="Haddad, Charles R., Jin, Chi & Platnick, Norman I." docDate="2022" docId="AE2487D1B52FFF4952DF96FECE12FA02" docLanguage="en" docName="zootaxa.5133.3.3.pdf" docOrigin="Zootaxa 5133 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Orthobula Simon 1897" docType="treatment" docVersion="8" lastPageNumber="358" masterDocId="521DFFA9B52CFF4F5248952CCB73FFE9" masterDocTitle="A revision of the spider genus Orthobula Simon, 1897 (Araneae: Trachelidae) in the Afrotropical Region. I. Continental species" masterLastPageNumber="382" masterPageNumber="355" pageNumber="358" updateTime="1699329851161" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="700DD401223A5868BAFD912626E7DF72">
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<mods:title id="226CB2246A6D3BD08804987583C38002">A revision of the spider genus Orthobula Simon, 1897 (Araneae: Trachelidae) in the Afrotropical Region. I. Continental species</mods:title>
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<mods:name id="D07A34DE88610E144C0A3A9DD30E4257" type="personal">
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<mods:roleTerm id="D1DACC84E94910CD188384FD91154B31">Author</mods:roleTerm>
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<mods:namePart id="3386C7C80297F216FF1A27881C72672D">Haddad, Charles R.</mods:namePart>
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<mods:affiliation id="71CB20318E0785E41B807C0C26B40443">Department of Zoology & Entomology, University of the Free State, P. O. Box 339, Bloemfontein 9300, South Africa.</mods:affiliation>
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<mods:name id="239C6321311E54FFE35AA62C53EB3F74" type="personal">
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<mods:roleTerm id="A74013B3727CC70E58FE0FFB7072CCA2">Author</mods:roleTerm>
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<mods:namePart id="9D4549CBF704212DF46049C0678E3363">Jin, Chi</mods:namePart>
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<mods:affiliation id="4DB05E0BAFBBAC689386A8EBADAA234B">School of Landscape and Ecological Engineering, Hebei University of Engineering, Handan, Hebei 056038, P. R. China.</mods:affiliation>
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<mods:namePart id="B46EB87FF89B94923E0CD38B5272D58E">Platnick, Norman I.</mods:namePart>
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<mods:nameIdentifier id="B09A3E39F57F2C0C2724B469677E3587" type="ORCID">0000-0003-4868-7529</mods:nameIdentifier>
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<mods:affiliation id="75932B89DFC2C6F5EB914B14D95BE0EC">Division of Invertebrate Zoology, American Museum of Natural History, Central Park West at 79 th Street, New York, NY 10024, U. S. A. https: // orcid. org / 0000 - 0003 - 4868 - 7529</mods:affiliation>
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<mods:title id="8E63A9C9B15119D60EB9D446E2C4F77D">Zootaxa</mods:title>
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<mods:part id="9721001A3B8ED0E0E3D3939E54D891D8">
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<mods:date id="1BCBE59630994164E3AB6418437223B4">2022</mods:date>
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<mods:number id="38C65D2F631B22A3C067AD654EDF4463">2022-05-05</mods:number>
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<mods:number id="D4DE95B5704AE4394B918DE8B56400A3">5133</mods:number>
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<treatment id="AE2487D1B52FFF4952DF96FECE12FA02" ID-DOI="http://doi.org/10.5281/zenodo.6521598" ID-GBIF-Taxon="195277971" ID-Zenodo-Dep="6521598" LSID="urn:lsid:plazi:treatment:AE2487D1B52FFF4952DF96FECE12FA02" httpUri="http://treatment.plazi.org/id/AE2487D1B52FFF4952DF96FECE12FA02" lastPageId="6" lastPageNumber="358" pageId="3" pageNumber="358">
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<subSubSection id="6E97654CB52FFF4C52DF96FEC97EFC05" box="[151,525,978,1005]" pageId="3" pageNumber="358" type="nomenclature">
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<paragraph id="263236C7B52FFF4C52DF96FEC97EFC05" blockId="3.[151,525,978,1005]" box="[151,525,978,1005]" pageId="3" pageNumber="358">
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<heading id="7D7A81ABB52FFF4C52DF96FEC97EFC05" bold="true" box="[151,525,978,1005]" fontSize="11" level="1" pageId="3" pageNumber="358" reason="1">
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<emphasis id="14F9EAD5B52FFF4C52DF96FEC97EFC05" bold="true" box="[151,525,978,1005]" pageId="3" pageNumber="358">
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Genus
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<taxonomicName id="E18D4D44B52FFF4C52A696FEC97EFC05" ID-CoL="63GT2" ID-ENA="1956877" authority="Simon, 1897" authorityName="Simon" authorityYear="1897" box="[238,525,978,1005]" class="Arachnida" family="Trachelidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="358" phylum="Arthropoda" rank="genus">
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<emphasis id="14F9EAD5B52FFF4C52A696FECA1FFC04" bold="true" box="[238,364,978,1005]" italics="true" pageId="3" pageNumber="358">Orthobula</emphasis>
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Simon, 1897
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="6E97654CB52FFF4C52DF913BC867FBC6" box="[151,788,1047,1071]" pageId="3" pageNumber="358" type="reference_group">
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<paragraph id="263236C7B52FFF4C52DF913BC867FBC6" blockId="3.[151,788,1047,1071]" box="[151,788,1047,1071]" pageId="3" pageNumber="358">
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<treatmentCitationGroup id="069D11E9B52FFF4C52DF913BC867FBC6" box="[151,788,1047,1071]" pageId="3" pageNumber="358">
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<treatmentCitation id="A72C10D6B52FFF4C52DF913BCAB9FBC6" author="Simon, E." box="[151,458,1047,1071]" page="152" pageId="3" pageNumber="358" year="1897">
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<taxonomicName id="E18D4D44B52FFF4C52DF913BCAB9FBC6" ID-CoL="63GT2" authority="Simon, 1897 a: 152" authorityName="Simon" authorityPageNumber="152" authorityYear="1897" box="[151,458,1047,1071]" class="Arachnida" family="Trachelidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="358" phylum="Arthropoda" rank="genus">
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<emphasis id="14F9EAD5B52FFF4C52DF913BCA72FBC6" box="[151,257,1047,1071]" italics="true" pageId="3" pageNumber="358">Orthobula</emphasis>
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<bibRefCitation id="421C4B36B52FFF4C534F913BCAB9FBC6" author="Simon, E." box="[263,458,1047,1071]" pageId="3" pageNumber="358" refId="ref23471" refString="Simon, E. (1897 a) Histoire naturelle des araignees. Tome Second. Premier Fascicule. Librairie Encyclopedique de Roret, Paris, 192 pp." type="book" year="1897">Simon, 1897a: 152</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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;
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<treatmentCitation id="A72C10D6B52FFF4C539D913BC863FBC6" ID-DOI="http://doi.org/10.5281/zenodo.5575641" ID-Zenodo-Dep="5575641" author="Deeleman-Reinhold, C. L." box="[469,784,1047,1071]" httpUri="http://treatment.plazi.org/id/03B3FE72C921FFC664C089AEDEABFD79" page="431" pageId="3" pageNumber="358" year="2001">
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<bibRefCitation id="421C4B36B52FFF4C539D913BC863FBC6" author="Deeleman-Reinhold, C. L." box="[469,784,1047,1071]" pageId="3" pageNumber="358" refId="ref22628" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]). Brill, Leiden, 591 pp. https: // doi. org / 10.1163 / 9789004475588" type="book" year="2001">Deeleman-Reinhold 2001: 431</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="6E97654CB52FFF4C52DF9179C809FB85" box="[151,890,1108,1133]" pageId="3" pageNumber="358" type="type_taxon">
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<paragraph id="263236C7B52FFF4C52DF9179C809FB85" blockId="3.[151,890,1108,1133]" box="[151,890,1108,1133]" pageId="3" pageNumber="358">
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<emphasis id="14F9EAD5B52FFF4C52DF9179CA50FB85" bold="true" box="[151,291,1108,1133]" pageId="3" pageNumber="358">
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<typeStatus id="F9368865B52FFF4C52DF9179CBBFFB84" box="[151,204,1109,1133]" pageId="3" pageNumber="358">Type</typeStatus>
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species.
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||
</emphasis>
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||
<taxonomicName id="E18D4D44B52FFF4C53629178C90DFB84" ID-CoL="6SXDC" authority="Simon, 1897" authorityName="Simon" authorityYear="1897" box="[298,638,1108,1133]" class="Arachnida" family="Trachelidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="358" phylum="Arthropoda" rank="species" species="impressa">
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<emphasis id="14F9EAD5B52FFF4C53629178CA86FB85" box="[298,501,1108,1132]" italics="true" pageId="3" pageNumber="358">Orthobula impressa</emphasis>
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Simon, 1897
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</taxonomicName>
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||
, by original designation.
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="6E97654CB52FFF4C52DF91B6C9EBFA58" pageId="3" pageNumber="358" type="diagnosis">
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<paragraph id="263236C7B52FFF4C52DF91B6C9EBFA58" blockId="3.[151,1437,1178,2033]" pageId="3" pageNumber="358">
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||
<emphasis id="14F9EAD5B52FFF4C52DF91B6CA61FB5D" bold="true" box="[151,274,1178,1204]" pageId="3" pageNumber="358">Diagnosis.</emphasis>
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<taxonomicName id="E18D4D44B52FFF4C535391B7CAE3FB5C" authorityName="Simon" authorityYear="1897" box="[283,400,1179,1205]" class="Arachnida" family="Trachelidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="358" phylum="Arthropoda" rank="genus">
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||
<emphasis id="14F9EAD5B52FFF4C535391B7CAE3FB5C" box="[283,400,1179,1205]" italics="true" pageId="3" pageNumber="358">Orthobula</emphasis>
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</taxonomicName>
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shares with
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<taxonomicName id="E18D4D44B52FFF4C506B91B7C9E2FB5C" authorityName="Haddad, Jin, Platnick & Booysen" authorityYear="2021" box="[547,657,1179,1205]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="358" phylum="Arthropoda" rank="genus">
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<emphasis id="14F9EAD5B52FFF4C506B91B7C9E2FB5C" box="[547,657,1179,1205]" italics="true" pageId="3" pageNumber="358">Capobula</emphasis>
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</taxonomicName>
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the large number of paired ventral spines on the anterior tibiae and metatarsi, and the absence of ventral cusps in both sexes. In somatic morphology, Afrotropical species of
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<taxonomicName id="E18D4D44B52FFF4C57609193CEEEFB30" authorityName="Simon" authorityYear="1897" box="[1320,1437,1215,1241]" class="Arachnida" family="Trachelidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="358" phylum="Arthropoda" rank="genus">
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<emphasis id="14F9EAD5B52FFF4C57609193CEEEFB30" box="[1320,1437,1215,1241]" italics="true" pageId="3" pageNumber="358">Orthobula</emphasis>
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</taxonomicName>
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have pits across the entire carapace surface, including along the midline (
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<figureCitation id="BEB62A42B52FFF4C518691CFCF23FB17" box="[974,1104,1251,1278]" captionStart="FIGURES 13–21" captionStartId="4.[151,264,1248,1272]" captionTargetBox="[151,1436,181,1214]" captionTargetId="figure-26@4.[151,1436,181,1214]" captionTargetPageId="4" captionText="FIGURES 13–21. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 male (13–15) and female (16–21): 13. Carapace, dorsal view; 14. Detail of carapace texture, showing fine tubercles on posterior slope; 15, 16. Eye region, dorsal view; 17. Mouthparts, ventral view; 18. Right chelicera, distal view, arrow indicating promarginal escort seta; 19. Left endite, serrula; 20. Sternum, ventral view; 21. Same, details of pits and tubercles." figureDoi="http://doi.org/10.5281/zenodo.6522683" httpUri="https://zenodo.org/record/6522683/files/figure.png" pageId="3" pageNumber="358">Figs 13–16</figureCitation>
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), while absent along the midline in
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<taxonomicName id="E18D4D44B52FFF4C52AD902BCA20FAC8" authorityName="Haddad, Jin, Platnick & Booysen" authorityYear="2021" box="[229,339,1287,1313]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="358" phylum="Arthropoda" rank="genus">
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<emphasis id="14F9EAD5B52FFF4C52AD902BCA20FAC8" box="[229,339,1287,1313]" italics="true" pageId="3" pageNumber="358">Capobula</emphasis>
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</taxonomicName>
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||
(
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<bibRefCitation id="421C4B36B52FFF4C532A902BC94EFAC8" author="Haddad, C. R. & Jin, C. & Platnick, N. I. & Booysen, R." box="[354,573,1287,1313]" pageId="3" pageNumber="358" pagination="41 - 71" refId="ref22952" refString="Haddad, C. R., Jin, C., Platnick, N. I. & Booysen, R. (2021) Capobula gen. nov., a new Afrotropical dark sac spider genus related to Orthobula Simon, 1897 (Araneae: Trachelidae). Zootaxa, 4942 (1), 41 - 71. https: // doi. org / 10.11646 / zootaxa. 4942.1.2" type="journal article" year="2021">
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||
Haddad
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||
<emphasis id="14F9EAD5B52FFF4C538A902BCA89FAC8" box="[450,506,1287,1313]" italics="true" pageId="3" pageNumber="358">et al.</emphasis>
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2021
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||
</bibRefCitation>
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||
: figs 11, 14) and some Asian species of
|
||
<taxonomicName id="E18D4D44B52FFF4C51B4902BCF02FAC8" authorityName="Simon" authorityYear="1897" box="[1020,1137,1287,1313]" class="Arachnida" family="Trachelidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="358" phylum="Arthropoda" rank="genus">
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||
<emphasis id="14F9EAD5B52FFF4C51B4902BCF02FAC8" box="[1020,1137,1287,1313]" italics="true" pageId="3" pageNumber="358">Orthobula</emphasis>
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</taxonomicName>
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||
. Female
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||
<taxonomicName id="E18D4D44B52FFF4C5690902BCE3EFAC8" authorityName="Simon" authorityYear="1897" box="[1240,1357,1287,1313]" class="Arachnida" family="Trachelidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="358" phylum="Arthropoda" rank="genus">
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||
<emphasis id="14F9EAD5B52FFF4C5690902BCE3EFAC8" box="[1240,1357,1287,1313]" italics="true" pageId="3" pageNumber="358">Orthobula</emphasis>
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||
</taxonomicName>
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||
can be distinguished by the posteriorly positioned spermathecae (as opposed to laterally positioned in
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<taxonomicName id="E18D4D44B52FFF4C569E9007CE37FAAC" authorityName="Haddad, Jin, Platnick & Booysen" authorityYear="2021" box="[1238,1348,1323,1349]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="358" phylum="Arthropoda" rank="genus">
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||
<emphasis id="14F9EAD5B52FFF4C569E9007CE37FAAC" box="[1238,1348,1323,1349]" italics="true" pageId="3" pageNumber="358">Capobula</emphasis>
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</taxonomicName>
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||
), while male
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<taxonomicName id="E18D4D44B52FFF4C529C9063CA3AFA80" authorityName="Simon" authorityYear="1897" box="[212,329,1359,1385]" class="Arachnida" family="Trachelidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="358" phylum="Arthropoda" rank="genus">
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||
<emphasis id="14F9EAD5B52FFF4C529C9063CA3AFA80" box="[212,329,1359,1385]" italics="true" pageId="3" pageNumber="358">Orthobula</emphasis>
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||
</taxonomicName>
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can be recognized by the considerably inflated palpal tegulum (only as wide as cymbium in
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<taxonomicName id="E18D4D44B52FFF4C571E9063CBBAFA64" authorityName="Haddad, Jin, Platnick & Booysen" authorityYear="2021" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="358" phylum="Arthropoda" rank="genus">
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<emphasis id="14F9EAD5B52FFF4C571E9063CBBAFA64" italics="true" pageId="3" pageNumber="358">Capobula</emphasis>
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</taxonomicName>
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), the narrow, finely coiled embolus (longer, uncoiled and retrolaterally directed in
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<taxonomicName id="E18D4D44B52FFF4C562E905FCFA7FA64" authorityName="Haddad, Jin, Platnick & Booysen" authorityYear="2021" box="[1126,1236,1395,1421]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="358" phylum="Arthropoda" rank="genus">
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<emphasis id="14F9EAD5B52FFF4C562E905FCFA7FA64" box="[1126,1236,1395,1421]" italics="true" pageId="3" pageNumber="358">Capobula</emphasis>
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</taxonomicName>
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), and the absence of a patellar apophysis (present in
|
||
<taxonomicName id="E18D4D44B52FFF4C505190BBC9F4FA58" authorityName="Haddad, Jin, Platnick & Booysen" authorityYear="2021" box="[537,647,1431,1457]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="358" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="14F9EAD5B52FFF4C505190BBC9F4FA58" box="[537,647,1431,1457]" italics="true" pageId="3" pageNumber="358">Capobula</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6E97654CB52FFF49528F9096CE12FA02" lastPageId="6" lastPageNumber="361" pageId="3" pageNumber="358" type="description">
|
||
<paragraph id="263236C7B52FFF49528F9096CE12FA02" blockId="3.[151,1437,1178,2033]" lastBlockId="6.[151,1437,1416,1515]" lastPageId="6" lastPageNumber="361" pageId="3" pageNumber="358">
|
||
<emphasis id="14F9EAD5B52FFF4C528F9096CA2AFA3D" bold="true" box="[199,345,1466,1492]" pageId="3" pageNumber="358">Description.</emphasis>
|
||
Small spiders,
|
||
<quantity id="E1759B22B52FFF4C504D9097C9D2FA3C" box="[517,673,1467,1493]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.7" metricValueMin="1.3" pageId="3" pageNumber="358" unit="mm" value="2.0" valueMax="2.7" valueMin="1.3">1.30–2.70mm</quantity>
|
||
in length; carapace orange-brown to wine-red, frequently with mottled black markings (
|
||
<figureCitation id="BEB62A42B52FFF4C53CC90F3CA8BFA10" box="[388,504,1503,1529]" captionStart="FIGURES 1–12" captionStartId="1.[151,264,1877,1901]" captionTargetBox="[151,1436,489,1854]" captionTargetId="figure-164@1.[151,1436,489,1854]" captionTargetPageId="1" captionText="FIGURES 1–12. Dorsal habitus photographs of Orthobula species: 1. O. aethiopica sp. nov., female; 2. Same, male; 3. O. arca sp. nov., female; 4. Same, male; 5. O. calceata Simon, 1897, female; 6. Same, male; 7. O. marusiki sp. nov., female; 8. Same, male; 9. O. milloti Caporiacco, 1949, female; 10. Same, male; 11. O. radiata Simon, 1897, female; 12. Same, male. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6522679" httpUri="https://zenodo.org/record/6522679/files/figure.png" pageId="3" pageNumber="358">Figs 1–12</figureCitation>
|
||
); carapace oval, broadest at coxae II, eye region narrow; fovea absent, only a short shallow depression (
|
||
<figureCitation id="BEB62A42B52FFF4C53C9932FCAA5F9F4" box="[385,470,1539,1565]" captionStart="FIGURES 13–21" captionStartId="4.[151,264,1248,1272]" captionTargetBox="[151,1436,181,1214]" captionTargetId="figure-26@4.[151,1436,181,1214]" captionTargetPageId="4" captionText="FIGURES 13–21. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 male (13–15) and female (16–21): 13. Carapace, dorsal view; 14. Detail of carapace texture, showing fine tubercles on posterior slope; 15, 16. Eye region, dorsal view; 17. Mouthparts, ventral view; 18. Right chelicera, distal view, arrow indicating promarginal escort seta; 19. Left endite, serrula; 20. Sternum, ventral view; 21. Same, details of pits and tubercles." figureDoi="http://doi.org/10.5281/zenodo.6522683" httpUri="https://zenodo.org/record/6522683/files/figure.png" pageId="3" pageNumber="358">Fig. 13</figureCitation>
|
||
); posterior margin slightly concave; carapace surface finely wrinkled, with series of deep pits forming striate patterns over entire surface (
|
||
<figureCitation id="BEB62A42B52FFF4C50B2930BC80DF9AB" box="[762,894,1575,1602]" captionStart="FIGURES 13–21" captionStartId="4.[151,264,1248,1272]" captionTargetBox="[151,1436,181,1214]" captionTargetId="figure-26@4.[151,1436,181,1214]" captionTargetPageId="4" captionText="FIGURES 13–21. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 male (13–15) and female (16–21): 13. Carapace, dorsal view; 14. Detail of carapace texture, showing fine tubercles on posterior slope; 15, 16. Eye region, dorsal view; 17. Mouthparts, ventral view; 18. Right chelicera, distal view, arrow indicating promarginal escort seta; 19. Left endite, serrula; 20. Sternum, ventral view; 21. Same, details of pits and tubercles." figureDoi="http://doi.org/10.5281/zenodo.6522683" httpUri="https://zenodo.org/record/6522683/files/figure.png" pageId="3" pageNumber="358">Figs 13–16</figureCitation>
|
||
), each pit with central pore, often plugged with unknown exudate (
|
||
<figureCitation id="BEB62A42B52FFF4C533A9367CAB4F98C" box="[370,455,1611,1637]" captionStart="FIGURES 13–21" captionStartId="4.[151,264,1248,1272]" captionTargetBox="[151,1436,181,1214]" captionTargetId="figure-26@4.[151,1436,181,1214]" captionTargetPageId="4" captionText="FIGURES 13–21. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 male (13–15) and female (16–21): 13. Carapace, dorsal view; 14. Detail of carapace texture, showing fine tubercles on posterior slope; 15, 16. Eye region, dorsal view; 17. Mouthparts, ventral view; 18. Right chelicera, distal view, arrow indicating promarginal escort seta; 19. Left endite, serrula; 20. Sternum, ventral view; 21. Same, details of pits and tubercles." figureDoi="http://doi.org/10.5281/zenodo.6522683" httpUri="https://zenodo.org/record/6522683/files/figure.png" pageId="3" pageNumber="358">Fig. 14</figureCitation>
|
||
); surface sparsely covered with scattered fine curved setae, particularly in eye region (
|
||
<figureCitation id="BEB62A42B52FFF4C52D79343CA56F963" box="[159,293,1647,1674]" captionStart="FIGURES 13–21" captionStartId="4.[151,264,1248,1272]" captionTargetBox="[151,1436,181,1214]" captionTargetId="figure-26@4.[151,1436,181,1214]" captionTargetPageId="4" captionText="FIGURES 13–21. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 male (13–15) and female (16–21): 13. Carapace, dorsal view; 14. Detail of carapace texture, showing fine tubercles on posterior slope; 15, 16. Eye region, dorsal view; 17. Mouthparts, ventral view; 18. Right chelicera, distal view, arrow indicating promarginal escort seta; 19. Left endite, serrula; 20. Sternum, ventral view; 21. Same, details of pits and tubercles." figureDoi="http://doi.org/10.5281/zenodo.6522683" httpUri="https://zenodo.org/record/6522683/files/figure.png" pageId="3" pageNumber="358">Figs 13–16</figureCitation>
|
||
). All eyes surrounded by black rings, weakly developed around PME (
|
||
<figureCitation id="BEB62A42B52FFF4C56139343CFA1F960" box="[1115,1234,1647,1673]" captionStart="FIGURES 1–12" captionStartId="1.[151,264,1877,1901]" captionTargetBox="[151,1436,489,1854]" captionTargetId="figure-164@1.[151,1436,489,1854]" captionTargetPageId="1" captionText="FIGURES 1–12. Dorsal habitus photographs of Orthobula species: 1. O. aethiopica sp. nov., female; 2. Same, male; 3. O. arca sp. nov., female; 4. Same, male; 5. O. calceata Simon, 1897, female; 6. Same, male; 7. O. marusiki sp. nov., female; 8. Same, male; 9. O. milloti Caporiacco, 1949, female; 10. Same, male; 11. O. radiata Simon, 1897, female; 12. Same, male. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6522679" httpUri="https://zenodo.org/record/6522679/files/figure.png" pageId="3" pageNumber="358">Figs 1–12</figureCitation>
|
||
); AER procurved in anterior view, recurved in dorsal view (
|
||
<figureCitation id="BEB62A42B52FFF4C503693BFC876F947" box="[638,773,1682,1710]" captionStart="FIGURES 13–21" captionStartId="4.[151,264,1248,1272]" captionTargetBox="[151,1436,181,1214]" captionTargetId="figure-26@4.[151,1436,181,1214]" captionTargetPageId="4" captionText="FIGURES 13–21. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 male (13–15) and female (16–21): 13. Carapace, dorsal view; 14. Detail of carapace texture, showing fine tubercles on posterior slope; 15, 16. Eye region, dorsal view; 17. Mouthparts, ventral view; 18. Right chelicera, distal view, arrow indicating promarginal escort seta; 19. Left endite, serrula; 20. Sternum, ventral view; 21. Same, details of pits and tubercles." figureDoi="http://doi.org/10.5281/zenodo.6522683" httpUri="https://zenodo.org/record/6522683/files/figure.png" pageId="3" pageNumber="358">Figs 15, 16</figureCitation>
|
||
); AME very slightly larger than ALE; AME separated by approximately ½ their diameter, nearly touching ALE (
|
||
<figureCitation id="BEB62A42B52FFF4C514A939BC8F6F93B" box="[770,901,1718,1746]" captionStart="FIGURES 13–21" captionStartId="4.[151,264,1248,1272]" captionTargetBox="[151,1436,181,1214]" captionTargetId="figure-26@4.[151,1436,181,1214]" captionTargetPageId="4" captionText="FIGURES 13–21. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 male (13–15) and female (16–21): 13. Carapace, dorsal view; 14. Detail of carapace texture, showing fine tubercles on posterior slope; 15, 16. Eye region, dorsal view; 17. Mouthparts, ventral view; 18. Right chelicera, distal view, arrow indicating promarginal escort seta; 19. Left endite, serrula; 20. Sternum, ventral view; 21. Same, details of pits and tubercles." figureDoi="http://doi.org/10.5281/zenodo.6522683" httpUri="https://zenodo.org/record/6522683/files/figure.png" pageId="3" pageNumber="358">Figs 15, 16</figureCitation>
|
||
); PER recurved in dorsal view, PME oval, PLE round; PME and PLE similar in diameter; median ocular quadrangle narrower anteriorly than posteriorly, length and posterior width approximately equal. Chilum indistinct, a tiny transverse sclerite; cheliceral promargin with three teeth, separated by approximately their basal width, proximal tooth smallest, median tooth slightly larger than distal; retromargin with two subequal teeth, set close together; cheliceral escort seta present (
|
||
<figureCitation id="BEB62A42B52FFF4C5600926BCFB4F888" box="[1096,1223,1863,1889]" captionStart="FIGURES 13–21" captionStartId="4.[151,264,1248,1272]" captionTargetBox="[151,1436,181,1214]" captionTargetId="figure-26@4.[151,1436,181,1214]" captionTargetPageId="4" captionText="FIGURES 13–21. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 male (13–15) and female (16–21): 13. Carapace, dorsal view; 14. Detail of carapace texture, showing fine tubercles on posterior slope; 15, 16. Eye region, dorsal view; 17. Mouthparts, ventral view; 18. Right chelicera, distal view, arrow indicating promarginal escort seta; 19. Left endite, serrula; 20. Sternum, ventral view; 21. Same, details of pits and tubercles." figureDoi="http://doi.org/10.5281/zenodo.6522683" httpUri="https://zenodo.org/record/6522683/files/figure.png" pageId="3" pageNumber="358">Figs 17, 18</figureCitation>
|
||
); fang with distinct serrula; endites convergent, notched laterally (
|
||
<figureCitation id="BEB62A42B52FFF4C50DF9247C99BF86C" box="[663,744,1899,1925]" captionStart="FIGURES 13–21" captionStartId="4.[151,264,1248,1272]" captionTargetBox="[151,1436,181,1214]" captionTargetId="figure-26@4.[151,1436,181,1214]" captionTargetPageId="4" captionText="FIGURES 13–21. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 male (13–15) and female (16–21): 13. Carapace, dorsal view; 14. Detail of carapace texture, showing fine tubercles on posterior slope; 15, 16. Eye region, dorsal view; 17. Mouthparts, ventral view; 18. Right chelicera, distal view, arrow indicating promarginal escort seta; 19. Left endite, serrula; 20. Sternum, ventral view; 21. Same, details of pits and tubercles." figureDoi="http://doi.org/10.5281/zenodo.6522683" httpUri="https://zenodo.org/record/6522683/files/figure.png" pageId="3" pageNumber="358">Fig. 17</figureCitation>
|
||
), with distinct serrula comprising sharp, ventrally curved denticles (
|
||
<figureCitation id="BEB62A42B52FFF4C52A892A3CA41F840" box="[224,306,1935,1961]" captionStart="FIGURES 13–21" captionStartId="4.[151,264,1248,1272]" captionTargetBox="[151,1436,181,1214]" captionTargetId="figure-26@4.[151,1436,181,1214]" captionTargetPageId="4" captionText="FIGURES 13–21. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 male (13–15) and female (16–21): 13. Carapace, dorsal view; 14. Detail of carapace texture, showing fine tubercles on posterior slope; 15, 16. Eye region, dorsal view; 17. Mouthparts, ventral view; 18. Right chelicera, distal view, arrow indicating promarginal escort seta; 19. Left endite, serrula; 20. Sternum, ventral view; 21. Same, details of pits and tubercles." figureDoi="http://doi.org/10.5281/zenodo.6522683" httpUri="https://zenodo.org/record/6522683/files/figure.png" pageId="3" pageNumber="358">Fig. 19</figureCitation>
|
||
); dense maxillar hair tuft on mesal margins (
|
||
<figureCitation id="BEB62A42B52FFF4C516892A3C8D3F840" box="[800,928,1935,1961]" captionStart="FIGURES 13–21" captionStartId="4.[151,264,1248,1272]" captionTargetBox="[151,1436,181,1214]" captionTargetId="figure-26@4.[151,1436,181,1214]" captionTargetPageId="4" captionText="FIGURES 13–21. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 male (13–15) and female (16–21): 13. Carapace, dorsal view; 14. Detail of carapace texture, showing fine tubercles on posterior slope; 15, 16. Eye region, dorsal view; 17. Mouthparts, ventral view; 18. Right chelicera, distal view, arrow indicating promarginal escort seta; 19. Left endite, serrula; 20. Sternum, ventral view; 21. Same, details of pits and tubercles." figureDoi="http://doi.org/10.5281/zenodo.6522683" httpUri="https://zenodo.org/record/6522683/files/figure.png" pageId="3" pageNumber="358">Figs 17–19</figureCitation>
|
||
); labium trapezoidal, slightly wider than long. Pleural bars sclerotised, isolated; sternum shield-shaped, longer than broad, surface smooth with deep pits without pores, sparsely covered in long straight setae (
|
||
<figureCitation id="BEB62A42B52FFF4C50C692FBC87FF818" box="[654,780,2007,2033]" captionStart="FIGURES 13–21" captionStartId="4.[151,264,1248,1272]" captionTargetBox="[151,1436,181,1214]" captionTargetId="figure-26@4.[151,1436,181,1214]" captionTargetPageId="4" captionText="FIGURES 13–21. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 male (13–15) and female (16–21): 13. Carapace, dorsal view; 14. Detail of carapace texture, showing fine tubercles on posterior slope; 15, 16. Eye region, dorsal view; 17. Mouthparts, ventral view; 18. Right chelicera, distal view, arrow indicating promarginal escort seta; 19. Left endite, serrula; 20. Sternum, ventral view; 21. Same, details of pits and tubercles." figureDoi="http://doi.org/10.5281/zenodo.6522683" httpUri="https://zenodo.org/record/6522683/files/figure.png" pageId="3" pageNumber="358">Figs 20, 21</figureCitation>
|
||
); precoxal triangles present, intercoxal sclerites present between all coxal pairs. Leg formula 4123, legs I and IV often very similar in length; sparsely covered in long fine setae; femora I with mesal convex curvature, all femora strongly constricted proximally, with ventral setal bases tuberculate (
|
||
<figureCitation id="BEB62A42B528FF4B536B90F6CA0BFA1C" box="[291,376,1498,1525]" captionStart="FIGURES 22–30" captionStartId="5.[151,264,1488,1512]" captionTargetBox="[151,1436,181,1464]" captionTargetId="figure-19@5.[151,1436,181,1464]" captionTargetPageId="5" captionText="FIGURES 22–30. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 female (22, 24, 26) and male (23, 25, 27–30): 22. Femur I and palp, ventral view; 23. Tibia I, retrolateral view; 24. Same, ventral view; 25. Same, detail of spine; 26. Same, detail of spine bases; 27. Same, detail of chemosensory seta; 28. Same, detail of trichobothrium base; 29. Metatarsus I, retrolateral view; 30. Same, distal end, arrow indicating metatarsal stopper." figureDoi="http://doi.org/10.5281/zenodo.6522685" httpUri="https://zenodo.org/record/6522685/files/figure.png" pageId="4" pageNumber="359">Fig. 22</figureCitation>
|
||
); patellar indentation on retrolateral side narrow, with lyriform organ at proximal end; anterior legs with strong paired ventral spines, chemosensory setae and trichobothria on tibiae, metatarsi and tarsi (
|
||
<figureCitation id="BEB62A42B528FF4B572290D2CB91F9D4" captionStart-0="FIGURES 22–30" captionStart-1="FIGURES 31–39" captionStartId-0="5.[151,264,1488,1512]" captionStartId-1="6.[151,264,1202,1226]" captionTargetBox-0="[151,1436,181,1464]" captionTargetBox-1="[151,1436,151,1184]" captionTargetId-0="figure-19@5.[151,1436,181,1464]" captionTargetId-1="figure-26@6.[151,1436,151,1184]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 22–30. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 female (22, 24, 26) and male (23, 25, 27–30): 22. Femur I and palp, ventral view; 23. Tibia I, retrolateral view; 24. Same, ventral view; 25. Same, detail of spine; 26. Same, detail of spine bases; 27. Same, detail of chemosensory seta; 28. Same, detail of trichobothrium base; 29. Metatarsus I, retrolateral view; 30. Same, distal end, arrow indicating metatarsal stopper." captionText-1="FIGURES 31–39. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 male (31–35, 37) and female (36, 38, 39): 31. Tarsus I, retrolateral view; 32. Same, detail of trichobothria; 33. Same, detail of chemosensory seta; 34–36. Same, detail of tarsal claws and claw tuft in retrolateral (34), retrolateral-ventral (35) and ventral (36) views; 37. Same, detail of tips of tenent setae; 38. Tarsus III, dorsal view, black arrows indicating trichobothria and white arrow indicating tarsal organ; 39. Same, detail of tarsal organ." figureDoi-0="http://doi.org/10.5281/zenodo.6522685" figureDoi-1="http://doi.org/10.5281/zenodo.6522691" httpUri-0="https://zenodo.org/record/6522685/files/figure.png" httpUri-1="https://zenodo.org/record/6522691/files/figure.png" pageId="4" pageNumber="359">Figs 23–33</figureCitation>
|
||
); metatarsi with single long distal trichobothrium (
|
||
<figureCitation id="BEB62A42B528FF4B5152930EC818F9D4" box="[794,875,1570,1597]" captionStart="FIGURES 22–30" captionStartId="5.[151,264,1488,1512]" captionTargetBox="[151,1436,181,1464]" captionTargetId="figure-19@5.[151,1436,181,1464]" captionTargetPageId="5" captionText="FIGURES 22–30. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 female (22, 24, 26) and male (23, 25, 27–30): 22. Femur I and palp, ventral view; 23. Tibia I, retrolateral view; 24. Same, ventral view; 25. Same, detail of spine; 26. Same, detail of spine bases; 27. Same, detail of chemosensory seta; 28. Same, detail of trichobothrium base; 29. Metatarsus I, retrolateral view; 30. Same, distal end, arrow indicating metatarsal stopper." figureDoi="http://doi.org/10.5281/zenodo.6522685" httpUri="https://zenodo.org/record/6522685/files/figure.png" pageId="4" pageNumber="359">Fig. 29</figureCitation>
|
||
) and dorsal stopper (
|
||
<figureCitation id="BEB62A42B528FF4B561F930ECFD8F9D4" box="[1111,1195,1570,1597]" captionStart="FIGURES 22–30" captionStartId="5.[151,264,1488,1512]" captionTargetBox="[151,1436,181,1464]" captionTargetId="figure-19@5.[151,1436,181,1464]" captionTargetPageId="5" captionText="FIGURES 22–30. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 female (22, 24, 26) and male (23, 25, 27–30): 22. Femur I and palp, ventral view; 23. Tibia I, retrolateral view; 24. Same, ventral view; 25. Same, detail of spine; 26. Same, detail of spine bases; 27. Same, detail of chemosensory seta; 28. Same, detail of trichobothrium base; 29. Metatarsus I, retrolateral view; 30. Same, distal end, arrow indicating metatarsal stopper." figureDoi="http://doi.org/10.5281/zenodo.6522685" httpUri="https://zenodo.org/record/6522685/files/figure.png" pageId="4" pageNumber="359">Fig. 30</figureCitation>
|
||
), metatarsi III and IV with ventral preening comb at distal end; tarsi usually with single or pair of short trichobothria at 1/3 their length and long pair at 2/3 their length (
|
||
<figureCitation id="BEB62A42B528FF4B504B9346C9DEF96C" box="[515,685,1642,1669]" captionStart="FIGURES 31–39" captionStartId="6.[151,264,1202,1226]" captionTargetBox="[151,1436,151,1184]" captionTargetId="figure-26@6.[151,1436,151,1184]" captionTargetPageId="6" captionText="FIGURES 31–39. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 male (31–35, 37) and female (36, 38, 39): 31. Tarsus I, retrolateral view; 32. Same, detail of trichobothria; 33. Same, detail of chemosensory seta; 34–36. Same, detail of tarsal claws and claw tuft in retrolateral (34), retrolateral-ventral (35) and ventral (36) views; 37. Same, detail of tips of tenent setae; 38. Tarsus III, dorsal view, black arrows indicating trichobothria and white arrow indicating tarsal organ; 39. Same, detail of tarsal organ." figureDoi="http://doi.org/10.5281/zenodo.6522691" httpUri="https://zenodo.org/record/6522691/files/figure.png" pageId="4" pageNumber="359">Figs 31, 32, 38</figureCitation>
|
||
); trichobothria with sunken distal plate, distal margin of hood overlapping plate, hood with two curved ridges, roughly concentric (
|
||
<figureCitation id="BEB62A42B528FF4B513A93A2C8BBF940" box="[882,968,1678,1705]" captionStart="FIGURES 22–30" captionStartId="5.[151,264,1488,1512]" captionTargetBox="[151,1436,181,1464]" captionTargetId="figure-19@5.[151,1436,181,1464]" captionTargetPageId="5" captionText="FIGURES 22–30. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 female (22, 24, 26) and male (23, 25, 27–30): 22. Femur I and palp, ventral view; 23. Tibia I, retrolateral view; 24. Same, ventral view; 25. Same, detail of spine; 26. Same, detail of spine bases; 27. Same, detail of chemosensory seta; 28. Same, detail of trichobothrium base; 29. Metatarsus I, retrolateral view; 30. Same, distal end, arrow indicating metatarsal stopper." figureDoi="http://doi.org/10.5281/zenodo.6522685" httpUri="https://zenodo.org/record/6522685/files/figure.png" pageId="4" pageNumber="359">Fig. 28</figureCitation>
|
||
); tarsal organ oval, very slightly elevated from integument, surface finely wrinkled, opening oval and distally placed (
|
||
<figureCitation id="BEB62A42B528FF4B51AF939ECF48F924" box="[999,1083,1714,1741]" captionStart="FIGURES 31–39" captionStartId="6.[151,264,1202,1226]" captionTargetBox="[151,1436,151,1184]" captionTargetId="figure-26@6.[151,1436,151,1184]" captionTargetPageId="6" captionText="FIGURES 31–39. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 male (31–35, 37) and female (36, 38, 39): 31. Tarsus I, retrolateral view; 32. Same, detail of trichobothria; 33. Same, detail of chemosensory seta; 34–36. Same, detail of tarsal claws and claw tuft in retrolateral (34), retrolateral-ventral (35) and ventral (36) views; 37. Same, detail of tips of tenent setae; 38. Tarsus III, dorsal view, black arrows indicating trichobothria and white arrow indicating tarsal organ; 39. Same, detail of tarsal organ." figureDoi="http://doi.org/10.5281/zenodo.6522691" httpUri="https://zenodo.org/record/6522691/files/figure.png" pageId="4" pageNumber="359">Fig. 39</figureCitation>
|
||
); paired tarsal claws short, with four intermediate teeth and moderately dense tenant setae forming claw tufts in between (
|
||
<figureCitation id="BEB62A42B528FF4B563793FACE71F918" box="[1151,1282,1750,1777]" captionStart="FIGURES 31–39" captionStartId="6.[151,264,1202,1226]" captionTargetBox="[151,1436,151,1184]" captionTargetId="figure-26@6.[151,1436,151,1184]" captionTargetPageId="6" captionText="FIGURES 31–39. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 male (31–35, 37) and female (36, 38, 39): 31. Tarsus I, retrolateral view; 32. Same, detail of trichobothria; 33. Same, detail of chemosensory seta; 34–36. Same, detail of tarsal claws and claw tuft in retrolateral (34), retrolateral-ventral (35) and ventral (36) views; 37. Same, detail of tips of tenent setae; 38. Tarsus III, dorsal view, black arrows indicating trichobothria and white arrow indicating tarsal organ; 39. Same, detail of tarsal organ." figureDoi="http://doi.org/10.5281/zenodo.6522691" httpUri="https://zenodo.org/record/6522691/files/figure.png" pageId="4" pageNumber="359">Figs 34–37</figureCitation>
|
||
); palpal claw simple, sharply curved distally. Abdomen oval, clearly larger in females than males, with dorsal scutum in males only; dorsum with very sparse fine setae and two pairs of sigilla, prominent in females, barely distinguishable on scutum in males (
|
||
<figureCitation id="BEB62A42B528FF4B5313926ECABDF8B4" box="[347,462,1858,1885]" captionStart="FIGURES 1–12" captionStartId="1.[151,264,1877,1901]" captionTargetBox="[151,1436,489,1854]" captionTargetId="figure-164@1.[151,1436,489,1854]" captionTargetPageId="1" captionText="FIGURES 1–12. Dorsal habitus photographs of Orthobula species: 1. O. aethiopica sp. nov., female; 2. Same, male; 3. O. arca sp. nov., female; 4. Same, male; 5. O. calceata Simon, 1897, female; 6. Same, male; 7. O. marusiki sp. nov., female; 8. Same, male; 9. O. milloti Caporiacco, 1949, female; 10. Same, male; 11. O. radiata Simon, 1897, female; 12. Same, male. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6522679" httpUri="https://zenodo.org/record/6522679/files/figure.png" pageId="4" pageNumber="359">Figs 1–12</figureCitation>
|
||
); abdominal markings variable, predominantly composed of cream markings forming a X on dark grey or black background (
|
||
<figureCitation id="BEB62A42B528FF4B5074924AC9DCF868" box="[572,687,1894,1921]" captionStart="FIGURES 1–12" captionStartId="1.[151,264,1877,1901]" captionTargetBox="[151,1436,489,1854]" captionTargetId="figure-164@1.[151,1436,489,1854]" captionTargetPageId="1" captionText="FIGURES 1–12. Dorsal habitus photographs of Orthobula species: 1. O. aethiopica sp. nov., female; 2. Same, male; 3. O. arca sp. nov., female; 4. Same, male; 5. O. calceata Simon, 1897, female; 6. Same, male; 7. O. marusiki sp. nov., female; 8. Same, male; 9. O. milloti Caporiacco, 1949, female; 10. Same, male; 11. O. radiata Simon, 1897, female; 12. Same, male. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6522679" httpUri="https://zenodo.org/record/6522679/files/figure.png" pageId="4" pageNumber="359">Figs 1–12</figureCitation>
|
||
), sometimes reduced in
|
||
<taxonomicName id="E18D4D44B528FF4B51F7924BCF32F869" authorityName="Simon" authorityYear="1897" box="[959,1089,1894,1921]" class="Arachnida" family="Trachelidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="359" phylum="Arthropoda" rank="species" species="calceata">
|
||
<emphasis id="14F9EAD5B528FF4B51F7924BCF32F869" box="[959,1089,1894,1921]" italics="true" pageId="4" pageNumber="359">O. calceata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E18D4D44B528FF4B563F924BCF88F869" authority="Haddad & Jin & Platnick, 2022" authorityName="Haddad & Jin & Platnick" authorityYear="2022" box="[1143,1275,1894,1921]" class="Arachnida" family="Trachelidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="359" phylum="Arthropoda" rank="species" species="marusiki" status="sp. nov.">
|
||
<emphasis id="14F9EAD5B528FF4B563F924BCF88F869" box="[1143,1275,1894,1921]" italics="true" pageId="4" pageNumber="359">O. marusiki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="14F9EAD5B528FF4B574A924BCE2FF868" bold="true" box="[1282,1372,1895,1921]" pageId="4" pageNumber="359">
|
||
<taxonomicNameLabel id="0FCA57AEB528FF4B574A924BCE2FF868" box="[1282,1372,1895,1921]" pageId="4" pageNumber="359" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="BEB62A42B528FF4B5722924ACB92F84C" captionStart="FIGURES 70–81" captionStartId="15.[151,264,1931,1955]" captionTargetBox="[151,1436,692,1895]" captionTargetId="figure-270@15.[151,1436,692,1905]" captionTargetPageId="15" captionText="FIGURES 70–81. Dorsal habitus photographs of Orthobula calceata Simon, 1897 females (70–72) and males (73–75) and O. marusiki sp. nov. females (76–78) and males (79–81), all from M’bengué, Cote d’Ivoire, indicating variation in extent of abdominal markings. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6522717" httpUri="https://zenodo.org/record/6522717/files/figure.png" pageId="4" pageNumber="359">Figs 70–81</figureCitation>
|
||
); venter without post-epigastric sclerites, with large ventral sclerite in males that is wider than long, absent in females; small, weakly sclerotised inframamillary sclerite present in females, indistinct in males. Spinnerets (details observed in
|
||
<taxonomicName id="E18D4D44B528FF4B536892FFCAE7F805" authorityName="Simon" authorityYear="1897" box="[288,404,2002,2029]" class="Arachnida" family="Trachelidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="359" phylum="Arthropoda" rank="species" species="radiata">
|
||
<emphasis id="14F9EAD5B528FF4B536892FFCAE7F805" box="[288,404,2002,2029]" italics="true" pageId="4" pageNumber="359">O. radiata</emphasis>
|
||
</taxonomicName>
|
||
only): female (
|
||
<figureCitation id="BEB62A42B528FF4B500992FEC99CF804" box="[577,751,2002,2029]" captionStart="FIGURES 40–44" captionStartId="7.[151,264,1026,1050]" captionTargetBox="[151,1436,151,1008]" captionTargetId="figure-19@7.[151,1436,151,1008]" captionTargetPageId="7" captionText="FIGURES 40–44. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 female (40, 42–44) and male (41): 40, 41. Spinnerets, general view; 42. Anterior lateral spinnerets; 43. Posterior median spinnerets; 44. Posterior lateral spinnerets. Abbreviations: Ac—aciniform gland spigot; Cy—cylindrical gland spigot; mAmp—minor ampullate gland spigot; MAmp—major ampullate gland spigot; Pi—piriform gland spigot." figureDoi="http://doi.org/10.5281/zenodo.6522697" httpUri="https://zenodo.org/record/6522697/files/figure.png" pageId="4" pageNumber="359">Figs 40, 42–44</figureCitation>
|
||
): ALS of female with one major ampullate gland spigot and nine piriform gland spigots; PMS of female with one small minor ampullate gland spigot anteriorly and five large cylindrical gland spigots; PLS of female with two large cylindrical gland spigots anteriorly and four aciniform gland spigots posteriorly; male spinnerets (
|
||
<figureCitation id="BEB62A42B529FF4A507393E6C9FDF90D" box="[571,654,1738,1764]" captionStart="FIGURES 40–44" captionStartId="7.[151,264,1026,1050]" captionTargetBox="[151,1436,151,1008]" captionTargetId="figure-19@7.[151,1436,151,1008]" captionTargetPageId="7" captionText="FIGURES 40–44. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 female (40, 42–44) and male (41): 40, 41. Spinnerets, general view; 42. Anterior lateral spinnerets; 43. Posterior median spinnerets; 44. Posterior lateral spinnerets. Abbreviations: Ac—aciniform gland spigot; Cy—cylindrical gland spigot; mAmp—minor ampullate gland spigot; MAmp—major ampullate gland spigot; Pi—piriform gland spigot." figureDoi="http://doi.org/10.5281/zenodo.6522697" httpUri="https://zenodo.org/record/6522697/files/figure.png" pageId="5" pageNumber="360">Fig. 41</figureCitation>
|
||
) similarly short and stout, but spigots retracted and not examined in detail. Female epigyne with small paired CO positioned anterolaterally in epigyne (
|
||
<figureCitation id="BEB62A42B529FF4A560D93C2CFEAF8E0" box="[1093,1177,1774,1801]" captionStart="FIGURES 45–53" captionStartId="8.[151,264,1202,1226]" captionTargetBox="[151,1436,151,1184]" captionTargetId="figure-26@8.[151,1436,151,1184]" captionTargetPageId="8" captionText="FIGURES 45–53. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 female (45) and male (46–53): 45. Epigyne, ventral view; 46. Left palpal femur, retrolateral view; 47. Same, detail of ventrodistal apophysis; 48. Right palpal patella, tibia and tarsus, ventrolateral view; 49. Same, detail of patellar lyriform organ; 50. Same, detail of pores in lyriform organ; 51. Left palpal tibia, dorsolateral view; 52. Left embolus, retrolateral view; 53. Right embolus, retrolateral view, detail." figureDoi="http://doi.org/10.5281/zenodo.6522701" httpUri="https://zenodo.org/record/6522701/files/figure.png" pageId="5" pageNumber="360">Fig. 45</figureCitation>
|
||
), usually in sclerotised anterior ridges, with CD directed posteriorly towards midline (e.g.
|
||
<figureCitation id="BEB62A42B529FF4A51C1923EC8ACF8C4" box="[905,991,1810,1837]" captionStart="FIGURES 54–58" captionStartId="9.[151,264,1124,1148]" captionTargetBox="[159,1428,158,1096]" captionTargetId="figure-316@9.[151,1436,151,1103]" captionTargetPageId="9" captionText="FIGURES 54–58. Orthobula aethiopica sp. nov., genitalic morphology of holotype female (54, 55) and paratype male (56–58) from Addis Ababa: epigyne, ventral (54) and dorsal (55) views; palp, prolateral (56), ventral (57) and retrolateral (58) views. Scale bars: 0.10 mm. Abbreviations: AG—accessory gland; BU—bursa; CD—copulatory duct; CO—copulatory opening; EM—embolus; FD—fertilisation duct; RFA—retrodistal femoral apophysis; RTA—retrolateral tibial apophysis; SBB—sclerotised base of bursa; ST I—primary spermathecae." figureDoi="http://doi.org/10.5281/zenodo.6522707" httpUri="https://zenodo.org/record/6522707/files/figure.png" pageId="5" pageNumber="360">Figs 54</figureCitation>
|
||
,
|
||
<figureCitation id="BEB62A42B529FF4A51A4923ECF79F8C4" box="[1004,1034,1810,1837]" captionStart="FIGURES 59–63" captionStartId="11.[151,264,1053,1077]" captionTargetBox="[157,1432,188,1025]" captionTargetId="figure-19@11.[151,1436,181,1029]" captionTargetPageId="11" captionText="FIGURES 59–63. Orthobula arca sp. nov., genitalic morphology of female paratype from Bloemfontein (59, 60) and male paratype from Erfenis Dam (61–63): epigyne, ventral (59) and dorsal (60) views; palp, prolateral (61), ventral (62) and retrolateral (63) views. Scale bars: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.6522711" httpUri="https://zenodo.org/record/6522711/files/figure.png" pageId="5" pageNumber="360">59</figureCitation>
|
||
,
|
||
<figureCitation id="BEB62A42B529FF4A565E923ECF46F8C4" box="[1046,1077,1810,1837]" captionStart="FIGURES 65–69" captionStartId="14.[151,264,1064,1088]" captionTargetBox="[168,1424,191,1036]" captionTargetId="figure-26@14.[161,1427,181,1040]" captionTargetPageId="14" captionText="FIGURES 65–69. Orthobula calceata Simon, 1897, genitalic morphology of female (65, 66) and male (67–69) from Bagoé: epigyne, ventral (65) and dorsal (66) views; palp, prolateral (67), ventral (68) and retrolateral (69) views. Scale bars: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.6522715" httpUri="https://zenodo.org/record/6522715/files/figure.png" pageId="5" pageNumber="360">65</figureCitation>
|
||
); CD passing round, sclerotised bases of BU centrally; BU large, membranous, oval or teardrop-shaped, directed anterolaterally, each with small an teromesal accessory gland; short ducts connecting base of BU to transversely oval posterior ST I; fertilization ducts originating on posteromedian surface of ST I (e.g.
|
||
<figureCitation id="BEB62A42B529FF4A508F9252C868F870" box="[711,795,1918,1945]" captionStart="FIGURES 54–58" captionStartId="9.[151,264,1124,1148]" captionTargetBox="[159,1428,158,1096]" captionTargetId="figure-316@9.[151,1436,151,1103]" captionTargetPageId="9" captionText="FIGURES 54–58. Orthobula aethiopica sp. nov., genitalic morphology of holotype female (54, 55) and paratype male (56–58) from Addis Ababa: epigyne, ventral (54) and dorsal (55) views; palp, prolateral (56), ventral (57) and retrolateral (58) views. Scale bars: 0.10 mm. Abbreviations: AG—accessory gland; BU—bursa; CD—copulatory duct; CO—copulatory opening; EM—embolus; FD—fertilisation duct; RFA—retrodistal femoral apophysis; RTA—retrolateral tibial apophysis; SBB—sclerotised base of bursa; ST I—primary spermathecae." figureDoi="http://doi.org/10.5281/zenodo.6522707" httpUri="https://zenodo.org/record/6522707/files/figure.png" pageId="5" pageNumber="360">Figs 55</figureCitation>
|
||
,
|
||
<figureCitation id="BEB62A42B529FF4A516E9252C835F870" box="[806,838,1918,1945]" captionStart="FIGURES 59–63" captionStartId="11.[151,264,1053,1077]" captionTargetBox="[157,1432,188,1025]" captionTargetId="figure-19@11.[151,1436,181,1029]" captionTargetPageId="11" captionText="FIGURES 59–63. Orthobula arca sp. nov., genitalic morphology of female paratype from Bloemfontein (59, 60) and male paratype from Erfenis Dam (61–63): epigyne, ventral (59) and dorsal (60) views; palp, prolateral (61), ventral (62) and retrolateral (63) views. Scale bars: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.6522711" httpUri="https://zenodo.org/record/6522711/files/figure.png" pageId="5" pageNumber="360">60</figureCitation>
|
||
). Male palpal femur with small, finger-like RFA, with single (e.g.
|
||
<figureCitation id="BEB62A42B529FF4A5350928ECA1BF854" box="[280,360,1954,1981]" captionStart="FIGURES 59–63" captionStartId="11.[151,264,1053,1077]" captionTargetBox="[157,1432,188,1025]" captionTargetId="figure-19@11.[151,1436,181,1029]" captionTargetPageId="11" captionText="FIGURES 59–63. Orthobula arca sp. nov., genitalic morphology of female paratype from Bloemfontein (59, 60) and male paratype from Erfenis Dam (61–63): epigyne, ventral (59) and dorsal (60) views; palp, prolateral (61), ventral (62) and retrolateral (63) views. Scale bars: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.6522711" httpUri="https://zenodo.org/record/6522711/files/figure.png" pageId="5" pageNumber="360">Fig. 63</figureCitation>
|
||
) or bifid (e.g.
|
||
<figureCitation id="BEB62A42B529FF4A504F928EC9F6F855" box="[519,645,1954,1980]" captionStart="FIGURES 45–53" captionStartId="8.[151,264,1202,1226]" captionTargetBox="[151,1436,151,1184]" captionTargetId="figure-26@8.[151,1436,151,1184]" captionTargetPageId="8" captionText="FIGURES 45–53. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 female (45) and male (46–53): 45. Epigyne, ventral view; 46. Left palpal femur, retrolateral view; 47. Same, detail of ventrodistal apophysis; 48. Right palpal patella, tibia and tarsus, ventrolateral view; 49. Same, detail of patellar lyriform organ; 50. Same, detail of pores in lyriform organ; 51. Left palpal tibia, dorsolateral view; 52. Left embolus, retrolateral view; 53. Right embolus, retrolateral view, detail." figureDoi="http://doi.org/10.5281/zenodo.6522701" httpUri="https://zenodo.org/record/6522701/files/figure.png" pageId="5" pageNumber="360">Figs 46, 47</figureCitation>
|
||
,
|
||
<figureCitation id="BEB62A42B529FF4A50D8928EC9DEF854" box="[656,685,1954,1981]" captionStart="FIGURES 54–58" captionStartId="9.[151,264,1124,1148]" captionTargetBox="[159,1428,158,1096]" captionTargetId="figure-316@9.[151,1436,151,1103]" captionTargetPageId="9" captionText="FIGURES 54–58. Orthobula aethiopica sp. nov., genitalic morphology of holotype female (54, 55) and paratype male (56–58) from Addis Ababa: epigyne, ventral (54) and dorsal (55) views; palp, prolateral (56), ventral (57) and retrolateral (58) views. Scale bars: 0.10 mm. Abbreviations: AG—accessory gland; BU—bursa; CD—copulatory duct; CO—copulatory opening; EM—embolus; FD—fertilisation duct; RFA—retrodistal femoral apophysis; RTA—retrolateral tibial apophysis; SBB—sclerotised base of bursa; ST I—primary spermathecae." figureDoi="http://doi.org/10.5281/zenodo.6522707" httpUri="https://zenodo.org/record/6522707/files/figure.png" pageId="5" pageNumber="360">58</figureCitation>
|
||
) tip; palpal patella without apophysis, but with retrolateral lyriform organ (
|
||
<figureCitation id="BEB62A42B529FF4A52AD92EACA1BF808" box="[229,360,1990,2017]" captionStart="FIGURES 45–53" captionStartId="8.[151,264,1202,1226]" captionTargetBox="[151,1436,151,1184]" captionTargetId="figure-26@8.[151,1436,151,1184]" captionTargetPageId="8" captionText="FIGURES 45–53. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 female (45) and male (46–53): 45. Epigyne, ventral view; 46. Left palpal femur, retrolateral view; 47. Same, detail of ventrodistal apophysis; 48. Right palpal patella, tibia and tarsus, ventrolateral view; 49. Same, detail of patellar lyriform organ; 50. Same, detail of pores in lyriform organ; 51. Left palpal tibia, dorsolateral view; 52. Left embolus, retrolateral view; 53. Right embolus, retrolateral view, detail." figureDoi="http://doi.org/10.5281/zenodo.6522701" httpUri="https://zenodo.org/record/6522701/files/figure.png" pageId="5" pageNumber="360">Figs 49, 50</figureCitation>
|
||
); palpal tibia with small triangular RTA (
|
||
<figureCitation id="BEB62A42B529FF4A517A92EAC8C7F808" box="[818,948,1990,2017]" captionStart="FIGURES 45–53" captionStartId="8.[151,264,1202,1226]" captionTargetBox="[151,1436,151,1184]" captionTargetId="figure-26@8.[151,1436,151,1184]" captionTargetPageId="8" captionText="FIGURES 45–53. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 female (45) and male (46–53): 45. Epigyne, ventral view; 46. Left palpal femur, retrolateral view; 47. Same, detail of ventrodistal apophysis; 48. Right palpal patella, tibia and tarsus, ventrolateral view; 49. Same, detail of patellar lyriform organ; 50. Same, detail of pores in lyriform organ; 51. Left palpal tibia, dorsolateral view; 52. Left embolus, retrolateral view; 53. Right embolus, retrolateral view, detail." figureDoi="http://doi.org/10.5281/zenodo.6522701" httpUri="https://zenodo.org/record/6522701/files/figure.png" pageId="5" pageNumber="360">Figs 48, 51</figureCitation>
|
||
); tegulum subtriangular, greatly expanded proximally on prolateral side, tapering distally, base nearly twice as broad as cymbium, with sperm duct running down retrolateral side of tegulum in ventral view, with single proximal loop (e.g.
|
||
<figureCitation id="BEB62A42B52AFF49565E9080CF1BFA2E" box="[1046,1128,1452,1479]" captionStart="FIGURES 54–58" captionStartId="9.[151,264,1124,1148]" captionTargetBox="[159,1428,158,1096]" captionTargetId="figure-316@9.[151,1436,151,1103]" captionTargetPageId="9" captionText="FIGURES 54–58. Orthobula aethiopica sp. nov., genitalic morphology of holotype female (54, 55) and paratype male (56–58) from Addis Ababa: epigyne, ventral (54) and dorsal (55) views; palp, prolateral (56), ventral (57) and retrolateral (58) views. Scale bars: 0.10 mm. Abbreviations: AG—accessory gland; BU—bursa; CD—copulatory duct; CO—copulatory opening; EM—embolus; FD—fertilisation duct; RFA—retrodistal femoral apophysis; RTA—retrolateral tibial apophysis; SBB—sclerotised base of bursa; ST I—primary spermathecae." figureDoi="http://doi.org/10.5281/zenodo.6522707" httpUri="https://zenodo.org/record/6522707/files/figure.png" pageId="6" pageNumber="361">Fig. 57</figureCitation>
|
||
); embolus with broad transverse ridge at base, spike-like, with fine coils along its length (
|
||
<figureCitation id="BEB62A42B52AFF49511C90FCC8DAFA02" box="[852,937,1488,1515]" captionStart="FIGURES 45–53" captionStartId="8.[151,264,1202,1226]" captionTargetBox="[151,1436,151,1184]" captionTargetId="figure-26@8.[151,1436,151,1184]" captionTargetPageId="8" captionText="FIGURES 45–53. Scanning electron microscope photographs of Orthobula radiata Simon, 1897 female (45) and male (46–53): 45. Epigyne, ventral view; 46. Left palpal femur, retrolateral view; 47. Same, detail of ventrodistal apophysis; 48. Right palpal patella, tibia and tarsus, ventrolateral view; 49. Same, detail of patellar lyriform organ; 50. Same, detail of pores in lyriform organ; 51. Left palpal tibia, dorsolateral view; 52. Left embolus, retrolateral view; 53. Right embolus, retrolateral view, detail." figureDoi="http://doi.org/10.5281/zenodo.6522701" httpUri="https://zenodo.org/record/6522701/files/figure.png" pageId="6" pageNumber="361">Fig. 52</figureCitation>
|
||
); tegulum without any other structures.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |