815 lines
101 KiB
XML
815 lines
101 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.3790666" ID-GBIF-Dataset="6b003da1-30a4-4961-839d-53f2f79ae563" ID-GBIF-Taxon="163748827" ID-ISSN="1313–2970" ID-Zenodo-Dep="3790666" ID-ZooBank="A447D3B6-5387-44B6-AC1B-A05988000E43" approvalRequired="18" approvalRequired_for_taxonomicNames="7" approvalRequired_for_textStreams="11" checkinTime="1587964501893" checkinUser="plazi" docAuthor="Huys, Rony & Lee, Wonchoel" docDate="2009" docId="8F78FF1D160CCB4FC3801FFBFC0ED081" docLanguage="en" docName="ZK_article_2079.pdf" docOrigin="ZooKeys 23 (23)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@763ee217" docStyleName="zookeys.2008.journal_article" docTitle="Marbefia carthyi Huys & Lee 2009, comb. n." docType="treatment" docVersion="4" lastPageId="16" lastPageNumber="17" masterDocId="734187651608CB5FC30C1B58FFE2D275" masterDocTitle="Proposal of Marbefia, gen. n. and Inermiphonte, gen. n., including updated keys to the species of Pseudonychocamptus Lang, 1944 and Paralaophonte Lang, 1948 (Copepoda, Harpacticoida, Laophontidae)" masterLastPageNumber="38" masterPageNumber="1" pageId="4" pageNumber="5" updateTime="1643568717307" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Proposal of Marbefia, gen. n. and Inermiphonte, gen. n., including updated keys to the species of Pseudonychocamptus Lang, 1944 and Paralaophonte Lang, 1948 (Copepoda, Harpacticoida, Laophontidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Huys, Rony</mods:namePart>
|
||
<mods:affiliation>Natural History Museum, London, United Kingdom</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Lee, Wonchoel</mods:namePart>
|
||
<mods:affiliation>Hanyang University, Laboratory of Biodiversity, Seoul, Korea, South</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2009</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2009-10-06</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>23</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>23</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>1</mods:start>
|
||
<mods:end>38</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">10.3897/zookeys.23.168</mods:identifier>
|
||
<mods:identifier type="GBIF-Dataset">6b003da1-30a4-4961-839d-53f2f79ae563</mods:identifier>
|
||
<mods:identifier type="ISSN">1313–2970</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">576545</mods:identifier>
|
||
<mods:identifier type="ZooBank">A447D3B6-5387-44B6-AC1B-A05988000E43</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3790666" ID-GBIF-Taxon="163748827" ID-Zenodo-Dep="3790666" LSID="urn:lsid:plazi:treatment:8F78FF1D160CCB4FC3801FFBFC0ED081" httpUri="http://treatment.plazi.org/id/8F78FF1D160CCB4FC3801FFBFC0ED081" lastPageId="16" lastPageNumber="17" pageId="4" pageNumber="5">
|
||
<subSubSection pageId="4" pageNumber="5" type="nomenclature">
|
||
<paragraph blockId="4.[140,670,1187,1249]" box="[140,670,1187,1214]" pageId="4" pageNumber="5">
|
||
<heading box="[140,670,1187,1214]" fontSize="11" level="2" pageId="4" pageNumber="5" reason="3">
|
||
<emphasis bold="true" box="[140,670,1187,1214]" pageId="4" pageNumber="5">
|
||
<taxonomicName authority="(Hamond, 1968)" authorityName="Huys & Lee" authorityYear="2009" baseAuthorityName="Hamond" baseAuthorityYear="1968" box="[140,552,1187,1214]" class="Maxillopoda" family="Laophontidae" genus="Marbefia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="carthyi" status="comb. n.">
|
||
<emphasis bold="true" box="[140,341,1187,1214]" italics="true" pageId="4" pageNumber="5">Marbefia carthyi</emphasis>
|
||
(
|
||
<bibRefCitation author="Hamond R" box="[356,543,1187,1213]" pageId="4" pageNumber="5" pagination="172 - 176" refId="ref18534" refString="Hamond R (1968) Pseudonychocamptus carthyi n. sp. (Harpacticoida) from Hunstanton, Norfolk. Crustaceana supplement 1: 172 - 176." type="journal article" year="1968">Hamond, 1968</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[565,670,1187,1213]" pageId="4" pageNumber="5" rank="species">comb. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="4.[140,670,1187,1249]" box="[140,236,1223,1249]" pageId="4" pageNumber="5">Figs 1–6</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[140,628,1293,1319]" pageId="4" pageNumber="5" type="reference_group">
|
||
<paragraph blockId="4.[140,628,1293,1319]" box="[140,628,1293,1319]" pageId="4" pageNumber="5">
|
||
<taxonomicName authority="Hamond, 1968" authorityName="Hamond" authorityYear="1968" box="[140,628,1293,1319]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="carthyi">
|
||
<emphasis box="[140,444,1293,1319]" italics="true" pageId="4" pageNumber="5">Pseudonychocamptus carthyi</emphasis>
|
||
<bibRefCitation author="Hamond R" box="[451,628,1293,1319]" pageId="4" pageNumber="5" pagination="172 - 176" refId="ref18534" refString="Hamond R (1968) Pseudonychocamptus carthyi n. sp. (Harpacticoida) from Hunstanton, Norfolk. Crustaceana supplement 1: 172 - 176." type="journal article" year="1968">Hamond, 1968</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="11" lastPageNumber="12" pageId="4" pageNumber="5" type="materials_examined">
|
||
<paragraph blockId="4.[140,1108,1363,1672]" pageId="4" pageNumber="5">
|
||
<emphasis bold="true" box="[140,298,1363,1389]" pageId="4" pageNumber="5">
|
||
<emphasis box="[140,198,1363,1389]" inLineHeading="true" pageId="4" pageNumber="5" reason="1">
|
||
<typeStatus box="[140,198,1363,1389]" pageId="4" pageNumber="5">Type</typeStatus>
|
||
</emphasis>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry box="[305,403,1363,1389]" name="United Kingdom" pageId="4" pageNumber="5">England</collectingCountry>
|
||
,
|
||
<collectingRegion box="[414,508,1363,1389]" country="United Kingdom" name="Norfolk" pageId="4" pageNumber="5">Norfolk</collectingRegion>
|
||
, Hunstanton; in washings of colonies of
|
||
<taxonomicName authorityName="Poche" authorityYear="1914" box="[968,1107,1363,1389]" class="Hydrozoa" family="Campanulariidae" genus="Hartlaubella" kingdom="Animalia" order="Leptothecata" pageId="4" pageNumber="5" phylum="Cnidaria" rank="genus">
|
||
<emphasis box="[968,1107,1363,1389]" italics="true" pageId="4" pageNumber="5">Hartlaubella</emphasis>
|
||
</taxonomicName>
|
||
(as
|
||
<taxonomicName authorityName="Lamouroux" authorityYear="1812" box="[178,288,1399,1425]" class="Hydrozoa" family="Campanulariidae" genus="Laomedea" kingdom="Animalia" order="Leptothecata" pageId="4" pageNumber="5" phylum="Cnidaria" rank="genus">
|
||
<emphasis box="[178,288,1399,1425]" italics="true" pageId="4" pageNumber="5">Laomedea</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
<taxonomicName authority="(Pallas, 1766)" baseAuthorityName="Pallas" baseAuthorityYear="1766" box="[304,570,1399,1426]" class="Hydrozoa" family="Campanulariidae" genus="Laomedea" kingdom="Animalia" order="Leptothecata" pageId="4" pageNumber="5" phylum="Cnidaria" rank="species" species="gelatinosa">
|
||
<emphasis box="[304,409,1399,1425]" italics="true" pageId="4" pageNumber="5">gelatinosa</emphasis>
|
||
(
|
||
<bibRefCitation author="Pallas PS" box="[424,563,1399,1426]" pageId="4" pageNumber="5" refId="ref19902" refString="Pallas PS (1766) Elenchus zoophytorum sistens generum adumbrationes generaliores et specierum cognitarum succinctas descriptiones cum selectis auctorum synomymis. F. Varrentrapp, Hagae, xxviii + 451 pp." type="book" year="1766">Pallas, 1766</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName box="[585,701,1399,1425]" class="Hydrozoa" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Cnidaria" rank="class">Hydrozoa</taxonomicName>
|
||
,
|
||
<taxonomicName box="[713,910,1399,1426]" class="Hydrozoa" family="Campanulariidae" kingdom="Animalia" order="Leptothecata" pageId="4" pageNumber="5" phylum="Cnidaria" rank="family">Campanulariidae</taxonomicName>
|
||
) attached to concrete lumps found in little shallow pools under the pier; for associated fauna see
|
||
<bibRefCitation author="Hamond R" pageId="4" pageNumber="5" pagination="172 - 176" refId="ref18534" refString="Hamond R (1968) Pseudonychocamptus carthyi n. sp. (Harpacticoida) from Hunstanton, Norfolk. Crustaceana supplement 1: 172 - 176." type="journal article" year="1968">Hamond (1968: 176)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="4.[140,1108,1363,1672]" pageId="4" pageNumber="5">
|
||
<emphasis bold="true" box="[185,422,1504,1530]" pageId="4" pageNumber="5">
|
||
<emphasis box="[185,286,1504,1530]" inLineHeading="true" pageId="4" pageNumber="5" reason="1">Material</emphasis>
|
||
examined.
|
||
</emphasis>
|
||
(
|
||
<materialsCitation ID-GBIF-Occurrence="2608819377" collectingDate="1957-04-02" collectionCode="R" pageId="4" pageNumber="5" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
|
||
1)
|
||
<emphasis bold="true" box="[480,601,1504,1530]" pageId="4" pageNumber="5">
|
||
<typeStatus box="[480,596,1504,1530]" pageId="4" pageNumber="5">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCount box="[614,629,1505,1531]" pageId="4" pageNumber="5" type="female">♀</specimenCount>
|
||
dissected on one slide (reg. no. NHM 1967.10.2.2), from type locality, collected on
|
||
<date box="[659,805,1539,1566]" pageId="4" pageNumber="5" value="1957-04-02">
|
||
<collectingDate box="[659,805,1539,1566]" pageId="4" pageNumber="5" value="1957-04-02">2 April 1957</collectingDate>
|
||
</date>
|
||
by
|
||
<collectionCode box="[847,869,1540,1566]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="5">R</collectionCode>
|
||
. Hamond;
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph blockId="4.[140,1108,1363,1672]" pageId="4" pageNumber="5">
|
||
(2) one
|
||
<specimenCount box="[288,303,1575,1601]" pageId="4" pageNumber="5" type="female">♀</specimenCount>
|
||
dissected on 12 slides, one ♁ dissected on 9 slides, one
|
||
<specimenCount box="[1029,1044,1575,1601]" pageId="4" pageNumber="5" type="female">♀</specimenCount>
|
||
and one ♁ used for SEM examination, and two ♁♁ in 70% ethanol;
|
||
<materialsCitation ID-GBIF-Occurrence="2608819375" collectingDate="1992-05-06" collectionCode="MB, R" pageId="4" pageNumber="5" specimenCount="1">
|
||
all specimens collected from Station
|
||
<collectionCode box="[418,469,1645,1671]" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="4" pageNumber="5">MB</collectionCode>
|
||
.17 (53°10.34΄ N, 00°56.34΄ E) on
|
||
<date box="[883,1033,1645,1672]" pageId="4" pageNumber="5" value="1992-05-06">
|
||
<collectingDate box="[883,1033,1645,1672]" pageId="4" pageNumber="5" value="1992-05-06">6 May 1992</collectingDate>
|
||
</date>
|
||
by
|
||
<collectionCode box="[1080,1103,1646,1672]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="5">R</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="5.[140,1108,1618,1673]" pageId="5" pageNumber="6">
|
||
<emphasis bold="true" box="[140,241,1618,1640]" pageId="5" pageNumber="6">Figure |.</emphasis>
|
||
<taxonomicName authority="(Hamond, 1968)" authorityName="Huys & Lee" authorityYear="2009" baseAuthorityName="Hamond" baseAuthorityYear="1968" box="[250,575,1618,1640]" class="Maxillopoda" family="Laophontidae" genus="Marbefia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="carthyi" status="comb. n.">
|
||
<emphasis box="[250,402,1619,1640]" italics="true" pageId="5" pageNumber="6">Marbefia carthyi</emphasis>
|
||
(
|
||
<bibRefCitation author="Hamond R" box="[417,568,1618,1640]" pageId="5" pageNumber="6" pagination="172 - 176" refId="ref18534" refString="Hamond R (1968) Pseudonychocamptus carthyi n. sp. (Harpacticoida) from Hunstanton, Norfolk. Crustaceana supplement 1: 172 - 176." type="journal article" year="1968">Hamond, 1968</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[588,675,1618,1640]" pageId="5" pageNumber="6" rank="species">comb. n.</taxonomicNameLabel>
|
||
:
|
||
<emphasis bold="true" box="[688,707,1618,1640]" pageId="5" pageNumber="6">A</emphasis>
|
||
habitus
|
||
<specimenCount box="[792,804,1619,1640]" pageId="5" pageNumber="6" type="female">♀</specimenCount>
|
||
, dorsal
|
||
<emphasis bold="true" box="[889,906,1618,1640]" pageId="5" pageNumber="6">B</emphasis>
|
||
habitus
|
||
<specimenCount box="[991,1003,1619,1640]" pageId="5" pageNumber="6" type="female">♀</specimenCount>
|
||
, lateral
|
||
<emphasis bold="true" box="[1089,1108,1618,1640]" pageId="5" pageNumber="6">C</emphasis>
|
||
habitus ♁, dorsal.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767614" ID-Zenodo-Dep="3767614" httpUri="https://zenodo.org/record/3767614/files/figure.png" pageId="6" pageNumber="7" startId="6.[140,212,1585,1607]" targetBox="[161,1105,197,1555]" targetPageId="6">
|
||
<paragraph blockId="6.[140,1107,1585,1671]" pageId="6" pageNumber="7">
|
||
<emphasis bold="true" box="[140,239,1585,1607]" pageId="6" pageNumber="7">Figure 2.</emphasis>
|
||
<taxonomicName authority="(Hamond, 1968)" authorityName="Huys & Lee" authorityYear="2009" baseAuthorityName="Hamond" baseAuthorityYear="1968" box="[246,566,1585,1607]" class="Maxillopoda" family="Laophontidae" genus="Marbefia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="carthyi" status="comb. n.">
|
||
<emphasis box="[246,396,1585,1606]" italics="true" pageId="6" pageNumber="7">Marbefia carthyi</emphasis>
|
||
(
|
||
<bibRefCitation author="Hamond R" box="[409,559,1585,1607]" pageId="6" pageNumber="7" pagination="172 - 176" refId="ref18534" refString="Hamond R (1968) Pseudonychocamptus carthyi n. sp. (Harpacticoida) from Hunstanton, Norfolk. Crustaceana supplement 1: 172 - 176." type="journal article" year="1968">Hamond, 1968</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[576,660,1585,1607]" pageId="6" pageNumber="7" rank="species">comb. n.</taxonomicNameLabel>
|
||
(♀):
|
||
<emphasis bold="true" box="[713,732,1585,1607]" pageId="6" pageNumber="7">A</emphasis>
|
||
urosome, ventral [excluding P5-bearing somite]
|
||
<emphasis bold="true" box="[214,231,1617,1639]" pageId="6" pageNumber="7">B</emphasis>
|
||
anal somite and caudal rami, dorsal
|
||
<emphasis bold="true" box="[567,586,1617,1639]" pageId="6" pageNumber="7">C</emphasis>
|
||
rostrum, dorsal
|
||
<emphasis bold="true" box="[738,757,1617,1639]" pageId="6" pageNumber="7">D</emphasis>
|
||
antennule, ventral
|
||
<emphasis bold="true" box="[935,950,1617,1639]" pageId="6" pageNumber="7">E</emphasis>
|
||
labrum, anterior
|
||
<emphasis bold="true" box="[140,154,1649,1671]" pageId="6" pageNumber="7">F</emphasis>
|
||
paragnaths, anterior
|
||
<emphasis bold="true" box="[351,371,1649,1671]" pageId="6" pageNumber="7">G</emphasis>
|
||
genital area with P6 [copulatory pore arrowed].
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767616" ID-Zenodo-Dep="3767616" httpUri="https://zenodo.org/record/3767616/files/figure.png" pageId="7" pageNumber="8" startId="7.[140,212,1618,1640]" targetBox="[161,1095,190,1550]" targetPageId="7">
|
||
<paragraph blockId="7.[140,1108,1618,1672]" pageId="7" pageNumber="8">
|
||
<emphasis bold="true" box="[140,238,1618,1640]" pageId="7" pageNumber="8">Figure 3.</emphasis>
|
||
<taxonomicName authority="(Hamond, 1968)" authorityName="Huys & Lee" authorityYear="2009" baseAuthorityName="Hamond" baseAuthorityYear="1968" box="[244,561,1618,1640]" class="Maxillopoda" family="Laophontidae" genus="Marbefia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="carthyi" status="comb. n.">
|
||
<emphasis box="[244,393,1619,1640]" italics="true" pageId="7" pageNumber="8">Marbefia carthyi</emphasis>
|
||
(
|
||
<bibRefCitation author="Hamond R" box="[405,554,1618,1640]" pageId="7" pageNumber="8" pagination="172 - 176" refId="ref18534" refString="Hamond R (1968) Pseudonychocamptus carthyi n. sp. (Harpacticoida) from Hunstanton, Norfolk. Crustaceana supplement 1: 172 - 176." type="journal article" year="1968">Hamond, 1968</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[571,654,1618,1640]" pageId="7" pageNumber="8" rank="species">comb. n.</taxonomicNameLabel>
|
||
(♀):
|
||
<emphasis bold="true" box="[705,724,1618,1640]" pageId="7" pageNumber="8">A</emphasis>
|
||
antenna
|
||
<emphasis bold="true" box="[810,827,1618,1640]" pageId="7" pageNumber="8">B</emphasis>
|
||
mandibular palp
|
||
<emphasis bold="true" box="[992,1011,1618,1640]" pageId="7" pageNumber="8">C</emphasis>
|
||
maxillule, anterior
|
||
<emphasis bold="true" box="[219,238,1650,1672]" pageId="7" pageNumber="8">D</emphasis>
|
||
maxilla, posterior
|
||
<emphasis bold="true" box="[411,426,1650,1672]" pageId="7" pageNumber="8">E</emphasis>
|
||
maxilliped, posterior.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767618" ID-Zenodo-Dep="3767618" httpUri="https://zenodo.org/record/3767618/files/figure.png" pageId="8" pageNumber="9" startId="8.[140,212,1585,1607]" targetBox="[146,1088,177,1488]" targetPageId="8">
|
||
<paragraph blockId="8.[140,1107,1585,1671]" pageId="8" pageNumber="9">
|
||
<emphasis bold="true" box="[140,237,1585,1607]" pageId="8" pageNumber="9">Figure 4.</emphasis>
|
||
<taxonomicName authority="(Hamond, 1968)" authorityName="Huys & Lee" authorityYear="2009" baseAuthorityName="Hamond" baseAuthorityYear="1968" box="[242,557,1585,1607]" class="Maxillopoda" family="Laophontidae" genus="Marbefia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="carthyi" status="comb. n.">
|
||
<emphasis box="[242,391,1585,1606]" italics="true" pageId="8" pageNumber="9">Marbefia carthyi</emphasis>
|
||
(
|
||
<bibRefCitation author="Hamond R" box="[402,550,1585,1607]" pageId="8" pageNumber="9" pagination="172 - 176" refId="ref18534" refString="Hamond R (1968) Pseudonychocamptus carthyi n. sp. (Harpacticoida) from Hunstanton, Norfolk. Crustaceana supplement 1: 172 - 176." type="journal article" year="1968">Hamond, 1968</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[566,650,1585,1607]" pageId="8" pageNumber="9" rank="species">comb. n.</taxonomicNameLabel>
|
||
:
|
||
<emphasis bold="true" box="[659,678,1585,1607]" pageId="8" pageNumber="9">A</emphasis>
|
||
P1 ♀, anterior
|
||
<emphasis bold="true" box="[821,838,1585,1607]" pageId="8" pageNumber="9">B</emphasis>
|
||
P4 ♀, anterior [inner seta on enp-1 arrowed]
|
||
<emphasis bold="true" box="[287,306,1617,1639]" pageId="8" pageNumber="9">C</emphasis>
|
||
P4 endopod ♁, anterior [spinular rows on enp-1 arrowed]
|
||
<emphasis bold="true" box="[849,868,1617,1639]" pageId="8" pageNumber="9">D</emphasis>
|
||
distal part of P4 endopod ♀, posterior.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767620" ID-Zenodo-Dep="3767620" httpUri="https://zenodo.org/record/3767620/files/figure.png" pageId="9" pageNumber="10" startId="9.[140,212,1585,1607]" targetBox="[152,1104,167,1554]" targetPageId="9">
|
||
<paragraph blockId="9.[140,1107,1585,1671]" pageId="9" pageNumber="10">
|
||
<emphasis bold="true" box="[140,238,1585,1607]" pageId="9" pageNumber="10">Figure 5.</emphasis>
|
||
<taxonomicName authority="(Hamond, 1968)" authorityName="Huys & Lee" authorityYear="2009" baseAuthorityName="Hamond" baseAuthorityYear="1968" box="[243,557,1585,1607]" class="Maxillopoda" family="Laophontidae" genus="Marbefia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="carthyi" status="comb. n.">
|
||
<emphasis box="[243,391,1585,1606]" italics="true" pageId="9" pageNumber="10">Marbefia carthyi</emphasis>
|
||
(
|
||
<bibRefCitation author="Hamond R" box="[403,551,1585,1607]" pageId="9" pageNumber="10" pagination="172 - 176" refId="ref18534" refString="Hamond R (1968) Pseudonychocamptus carthyi n. sp. (Harpacticoida) from Hunstanton, Norfolk. Crustaceana supplement 1: 172 - 176." type="journal article" year="1968">Hamond, 1968</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[567,651,1585,1607]" pageId="9" pageNumber="10" rank="species">comb. n.</taxonomicNameLabel>
|
||
:
|
||
<emphasis bold="true" box="[660,679,1585,1607]" pageId="9" pageNumber="10">A</emphasis>
|
||
P2 ♀, anterior,
|
||
<emphasis bold="true" box="[827,844,1585,1607]" pageId="9" pageNumber="10">B</emphasis>
|
||
P2 endopod ♀, outer lateral
|
||
<emphasis bold="true" box="[140,159,1617,1639]" pageId="9" pageNumber="10">C</emphasis>
|
||
P3 ♀, anterior [inner seta on enp-1 arrowed]
|
||
<emphasis bold="true" box="[593,612,1617,1639]" pageId="9" pageNumber="10">D</emphasis>
|
||
P3 endopod ♁ [spinules on enp-1 and apophysis on enp-2 arrowed].
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767622" ID-Zenodo-Dep="3767622" httpUri="https://zenodo.org/record/3767622/files/figure.png" pageId="10" pageNumber="11" startId="10.[140,212,1585,1607]" targetBox="[184,1099,180,1545]" targetPageId="10">
|
||
<paragraph blockId="10.[140,1107,1585,1672]" pageId="10" pageNumber="11">
|
||
<emphasis bold="true" box="[140,239,1585,1607]" pageId="10" pageNumber="11">Figure 6.</emphasis>
|
||
<taxonomicName authority="(Hamond, 1968)" authorityName="Huys & Lee" authorityYear="2009" baseAuthorityName="Hamond" baseAuthorityYear="1968" box="[245,564,1585,1607]" class="Maxillopoda" family="Laophontidae" genus="Marbefia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="carthyi" status="comb. n.">
|
||
<emphasis box="[245,395,1585,1606]" italics="true" pageId="10" pageNumber="11">Marbefia carthyi</emphasis>
|
||
(
|
||
<bibRefCitation author="Hamond R" box="[408,557,1585,1607]" pageId="10" pageNumber="11" pagination="172 - 176" refId="ref18534" refString="Hamond R (1968) Pseudonychocamptus carthyi n. sp. (Harpacticoida) from Hunstanton, Norfolk. Crustaceana supplement 1: 172 - 176." type="journal article" year="1968">Hamond, 1968</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[574,659,1585,1607]" pageId="10" pageNumber="11" rank="species">comb. n.</taxonomicNameLabel>
|
||
:
|
||
<emphasis bold="true" box="[670,689,1585,1607]" pageId="10" pageNumber="11">A</emphasis>
|
||
antennule ♁, ventral [armature on segments 3, 4 and 8 omitted]
|
||
<emphasis bold="true" box="[331,348,1617,1639]" pageId="10" pageNumber="11">B</emphasis>
|
||
antennulary segments 3 (right) and 4 (left) ♁
|
||
<emphasis bold="true" box="[786,805,1617,1639]" pageId="10" pageNumber="11">C</emphasis>
|
||
antennulary segment 8 ♁
|
||
<emphasis bold="true" box="[1056,1075,1617,1639]" pageId="10" pageNumber="11">D</emphasis>
|
||
P5 and P6 ♁, anterior
|
||
<emphasis bold="true" box="[320,335,1649,1671]" pageId="10" pageNumber="11">E</emphasis>
|
||
P5 ♀, anterior.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="11.[140,1108,167,1672]" pageId="11" pageNumber="12">
|
||
Hamond (reg. nos. NHM 1993.312–321); depth
|
||
<quantity box="[746,859,167,193]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="11" pageNumber="12" unit="m" value="12.5" valueMax="13.0" valueMin="12.0">12–13 m</quantity>
|
||
; fine sand with high silt content;
|
||
</paragraph>
|
||
<paragraph blockId="11.[140,1108,167,1672]" pageId="11" pageNumber="12">
|
||
(3) one
|
||
<specimenCount box="[270,285,238,264]" pageId="11" pageNumber="12" type="female">♀</specimenCount>
|
||
(reg. no. NHM 1998.588) re-identified by F. Fiers in 1998, found among
|
||
<taxonomicName authority="(Boeck, 1873)" baseAuthorityName="Boeck" baseAuthorityYear="1873" box="[140,607,272,298]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="koreni">
|
||
<emphasis box="[140,437,272,298]" italics="true" pageId="11" pageNumber="12">Pseudonychocamptus koreni</emphasis>
|
||
(
|
||
<bibRefCitation author="Boeck A" box="[453,599,272,298]" pageId="11" pageNumber="12" pagination="35 - 60" refId="ref17768" refString="Boeck A (1873) Nye Slaegter og Arter af Saltvands-Copepoder. Forhandlinger i Videnskabsselskabet i Kristiania 1872: 35 - 60." type="journal article" year="1873">Boeck, 1873</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
specimens (reg. no. NHM
|
||
<date box="[922,1054,272,299]" pageId="11" pageNumber="12" value="1967-10-31">1967.10.31</date>
|
||
–70) collected during the 1966 University of London Sub-Aqua Club expedition to the
|
||
<collectingRegion country="United Kingdom" name="Isles of Scilly" pageId="11" pageNumber="12">Isles of Scilly</collectingRegion>
|
||
;
|
||
<collectingCountry box="[251,402,343,369]" name="United Kingdom" pageId="11" pageNumber="12">Great Britain</collectingCountry>
|
||
Rock, off St Mary’s; algal debris at LWN, among bryozoans (
|
||
<taxonomicName box="[149,237,378,404]" class="Gymnolaemata" family="Cellariidae" genus="Cellaria" kingdom="Animalia" order="Cheilostomatida" pageId="11" pageNumber="12" phylum="Bryozoa" rank="genus">
|
||
<emphasis box="[149,237,378,404]" italics="true" pageId="11" pageNumber="12">Cellaria</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Johnston" authorityYear="1838" box="[251,341,378,404]" class="Gymnolaemata" genus="Lepralia" kingdom="Animalia" order="Cheilostomatida" pageId="11" pageNumber="12" phylum="Bryozoa" rank="genus">
|
||
<emphasis box="[251,341,378,404]" italics="true" pageId="11" pageNumber="12">Lepralia</emphasis>
|
||
</taxonomicName>
|
||
) and hydroids (
|
||
<taxonomicName box="[523,631,378,404]" class="Hydrozoa" family="Sertulariidae" genus="Sertularia" kingdom="Animalia" order="Leptothecata" pageId="11" pageNumber="12" phylum="Cnidaria" rank="genus">
|
||
<emphasis box="[523,631,378,404]" italics="true" pageId="11" pageNumber="12">Sertularia</emphasis>
|
||
</taxonomicName>
|
||
),
|
||
<quantity box="[655,718,378,404]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="11" pageNumber="12" unit="m" value="27.0">27 m</quantity>
|
||
;
|
||
<date box="[729,840,378,404]" pageId="11" pageNumber="12" value="1966-07">July 1966</date>
|
||
(
|
||
<emphasis box="[856,882,378,404]" italics="true" pageId="11" pageNumber="12">cf.</emphasis>
|
||
<bibRefCitation author="Wells JBJ" box="[888,1078,378,405]" pageId="11" pageNumber="12" pagination="255 - 268" refId="ref20876" refString="Wells JBJ (1970) The marine flora and fauna of the Isles of Scilly. Crustacea: Copepoda: Harpacticoida. Journal of natural History 4 (2): 255 - 268." type="journal article" year="1970">Wells 1970: 267</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12" type="description">
|
||
<paragraph blockId="11.[140,1108,167,1672]" pageId="11" pageNumber="12">
|
||
<emphasis bold="true" box="[185,331,413,439]" pageId="11" pageNumber="12">
|
||
<emphasis box="[185,325,413,439]" inLineHeading="true" pageId="11" pageNumber="12" reason="1">Description</emphasis>
|
||
.
|
||
</emphasis>
|
||
Female.
|
||
<emphasis box="[435,487,413,439]" italics="true" pageId="11" pageNumber="12">Body</emphasis>
|
||
(Figure 1A, B). Total body length
|
||
<quantity box="[890,1037,413,440]" metricMagnitude="-1" metricUnit="m" metricValue="7.17" metricValueMax="7.38" metricValueMin="6.959999999999999" pageId="11" pageNumber="12" unit="mm" value="717.0" valueMax="738.0" valueMin="696.0">696–738 µm</quantity>
|
||
(n=2; mean
|
||
<quantity box="[207,299,448,475]" metricMagnitude="-1" metricUnit="m" metricValue="7.17" pageId="11" pageNumber="12" unit="mm" value="717.0">717 µm</quantity>
|
||
). Largest width at posterior margin of cephalic shield (
|
||
<quantity box="[895,987,448,475]" metricMagnitude="-1" metricUnit="m" metricValue="1.7399999999999998" pageId="11" pageNumber="12" unit="mm" value="174.0">174 µm</quantity>
|
||
). Urosome gradually tapering posteriorly (Figure 1A). Rostrum triangular (
|
||
<figureCitation box="[839,955,483,510]" captionStart="Figure 2" captionStartId="6.[140,212,1585,1607]" captionTargetBox="[161,1105,197,1555]" captionTargetId="figure@6.[140,1108,165,1566]" captionTargetPageId="6" captionText="Figure 2. Marbefia carthyi (Hamond, 1968), comb. n. (♀): A urosome, ventral [excluding P5-bearing somite] B anal somite and caudal rami, dorsal C rostrum, dorsal D antennule, ventral E labrum, anterior F paragnaths, anterior G genital area with P6 [copulatory pore arrowed]." figureDoi="http://doi.org/10.5281/zenodo.3767614" httpUri="https://zenodo.org/record/3767614/files/figure.png" pageId="11" pageNumber="12">Figure 2C</figureCitation>
|
||
), with pair of sensilla near apex; midventral tube-pore in subapical position; completely defined at base. Cephalothorax with posterior margin fringed with small spinules; pleural areas well developed and rounded with lobate posterolateral angles; entire surface covered with tiny spinules as illustrated in Figure 1A, B; sensilla and few pores present as illustrated in Figure 1A, B. Pedigerous somites covered with minute spinules. All prosomites without defined hyaline frills; posterior margins fringed with small spinules. Body constricted between individual somites. Urosome (Figures 1A, B; 2A) 5-segmented, comprising P5-bearing somite, genital double-somite and 3 free abdominal somites. Abdominal half of genital double-somite and second abdominal somite with lateral lobate extensions. All urosomites with surface ornamentation consisting of small spinules dorsally and laterally; ventral surface largely naked except for few rows of spinules and paired tube-pores; ventral hind margin with large spinules laterally and small spinules medially (
|
||
<figureCitation box="[802,917,906,932]" captionStart="Figure 2" captionStartId="6.[140,212,1585,1607]" captionTargetBox="[161,1105,197,1555]" captionTargetId="figure@6.[140,1108,165,1566]" captionTargetPageId="6" captionText="Figure 2. Marbefia carthyi (Hamond, 1968), comb. n. (♀): A urosome, ventral [excluding P5-bearing somite] B anal somite and caudal rami, dorsal C rostrum, dorsal D antennule, ventral E labrum, anterior F paragnaths, anterior G genital area with P6 [copulatory pore arrowed]." figureDoi="http://doi.org/10.5281/zenodo.3767614" httpUri="https://zenodo.org/record/3767614/files/figure.png" pageId="11" pageNumber="12">Figure 2A</figureCitation>
|
||
). Hyaline frills of urosomites not distinct. Genital double-somite (Figures 1A; 2A) with transverse, surface ridge dorsally and laterally, indicating original segmentation; completely fused ventrally. Genital field (
|
||
<figureCitation box="[297,414,1011,1038]" captionStart="Figure 2" captionStartId="6.[140,212,1585,1607]" captionTargetBox="[161,1105,197,1555]" captionTargetId="figure@6.[140,1108,165,1566]" captionTargetPageId="6" captionText="Figure 2. Marbefia carthyi (Hamond, 1968), comb. n. (♀): A urosome, ventral [excluding P5-bearing somite] B anal somite and caudal rami, dorsal C rostrum, dorsal D antennule, ventral E labrum, anterior F paragnaths, anterior G genital area with P6 [copulatory pore arrowed]." figureDoi="http://doi.org/10.5281/zenodo.3767614" httpUri="https://zenodo.org/record/3767614/files/figure.png" pageId="11" pageNumber="12">Figure 2G</figureCitation>
|
||
) with medium-sized copulatory pore (arrowed in
|
||
<figureCitation box="[983,1099,1011,1038]" captionStart="Figure 2" captionStartId="6.[140,212,1585,1607]" captionTargetBox="[161,1105,197,1555]" captionTargetId="figure@6.[140,1108,165,1566]" captionTargetPageId="6" captionText="Figure 2. Marbefia carthyi (Hamond, 1968), comb. n. (♀): A urosome, ventral [excluding P5-bearing somite] B anal somite and caudal rami, dorsal C rostrum, dorsal D antennule, ventral E labrum, anterior F paragnaths, anterior G genital area with P6 [copulatory pore arrowed]." figureDoi="http://doi.org/10.5281/zenodo.3767614" httpUri="https://zenodo.org/record/3767614/files/figure.png" pageId="11" pageNumber="12">Figure 2G</figureCitation>
|
||
) located in median depression; gonopores fused medially forming single genital slit covered on either side by operculum derived from sixth leg; P6 with small protuberance bearing 2 bare setae; small triangular process located on medial side adjacent to protuberance. Anal somite (
|
||
<figureCitation box="[226,336,1152,1178]" captionStart="Figure 2" captionStartId="6.[140,212,1585,1607]" captionTargetBox="[161,1105,197,1555]" captionTargetId="figure@6.[140,1108,165,1566]" captionTargetPageId="6" captionText="Figure 2. Marbefia carthyi (Hamond, 1968), comb. n. (♀): A urosome, ventral [excluding P5-bearing somite] B anal somite and caudal rami, dorsal C rostrum, dorsal D antennule, ventral E labrum, anterior F paragnaths, anterior G genital area with P6 [copulatory pore arrowed]." figureDoi="http://doi.org/10.5281/zenodo.3767614" httpUri="https://zenodo.org/record/3767614/files/figure.png" pageId="11" pageNumber="12">Figure 2B</figureCitation>
|
||
) with spinular ornamentation and pair of dorsal tube-pores; anal operculum well developed, bearing row of minute spinules and flanked by pair of sensilla; anal opening bordered by well developed frill bearing long setular extensions.
|
||
</paragraph>
|
||
<paragraph blockId="11.[140,1108,167,1672]" pageId="11" pageNumber="12">
|
||
<emphasis box="[185,323,1258,1284]" italics="true" pageId="11" pageNumber="12">Caudal rami</emphasis>
|
||
(
|
||
<figureCitation box="[339,487,1258,1284]" captionStart="Figure 2" captionStartId="6.[140,212,1585,1607]" captionTargetBox="[161,1105,197,1555]" captionTargetId="figure@6.[140,1108,165,1566]" captionTargetPageId="6" captionText="Figure 2. Marbefia carthyi (Hamond, 1968), comb. n. (♀): A urosome, ventral [excluding P5-bearing somite] B anal somite and caudal rami, dorsal C rostrum, dorsal D antennule, ventral E labrum, anterior F paragnaths, anterior G genital area with P6 [copulatory pore arrowed]." figureDoi="http://doi.org/10.5281/zenodo.3767614" httpUri="https://zenodo.org/record/3767614/files/figure.png" pageId="11" pageNumber="12">Figure 2A, B</figureCitation>
|
||
). Short, cylindrical, as long as wide; each ramus with 7 setae: seta I subventral, bare and shortest; setae II and III bare;
|
||
<materialsCitation ID-GBIF-Occurrence="2608819374" collectionCode="IV, V" pageId="11" pageNumber="12" specimenCount="1" typeStatus="holotype">
|
||
setae
|
||
<collectionCode box="[921,950,1294,1320]" pageId="11" pageNumber="12">IV</collectionCode>
|
||
and
|
||
<collectionCode box="[1010,1030,1294,1320]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="12" type="Museum">V</collectionCode>
|
||
bipinnate, fused basally, and with predesigned fracture planes [seta
|
||
<collectionCode box="[822,842,1329,1355]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="12" type="Museum">V</collectionCode>
|
||
presumably longest but incomplete in all specimens examined including
|
||
<typeStatus box="[683,783,1363,1389]" pageId="11" pageNumber="12">holotype</typeStatus>
|
||
,
|
||
<emphasis box="[795,820,1364,1390]" italics="true" pageId="11" pageNumber="12">cf.</emphasis>
|
||
<bibRefCitation author="Hamond R" box="[827,1006,1363,1389]" pageId="11" pageNumber="12" pagination="172 - 176" refId="ref18534" refString="Hamond R (1968) Pseudonychocamptus carthyi n. sp. (Harpacticoida) from Hunstanton, Norfolk. Crustaceana supplement 1: 172 - 176." type="journal article" year="1968">Hamond (1968</bibRefCitation>
|
||
:
|
||
<figureCitation box="[1018,1084,1363,1390]" captionStart="Figure 3" captionStartId="7.[140,212,1618,1640]" captionTargetBox="[161,1095,190,1550]" captionTargetId="figure@7.[140,1108,165,1566]" captionTargetPageId="7" captionText="Figure 3. Marbefia carthyi (Hamond, 1968), comb. n. (♀): A antenna B mandibular palp C maxillule, anterior D maxilla, posterior E maxilliped, posterior." figureDoi="http://doi.org/10.5281/zenodo.3767616" httpUri="https://zenodo.org/record/3767616/files/figure.png" pageId="11" pageNumber="12">Fig. 3</figureCitation>
|
||
)]
|
||
</materialsCitation>
|
||
; seta VI bipinnate; seta VII tri-articulate at base. Each ramus with spinules on outer dorsal surface; additional spinular ornamentation present along inner and outer margins and around ventral and dorsal hind margins; long tube-pore present near ventral hind margin.
|
||
</paragraph>
|
||
<paragraph blockId="11.[140,1108,167,1672]" lastBlockId="12.[140,1108,166,1672]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
|
||
<emphasis box="[185,296,1504,1530]" italics="true" pageId="11" pageNumber="12">Antennule</emphasis>
|
||
(
|
||
<figureCitation box="[316,443,1504,1530]" captionStart="Figure 2" captionStartId="6.[140,212,1585,1607]" captionTargetBox="[161,1105,197,1555]" captionTargetId="figure@6.[140,1108,165,1566]" captionTargetPageId="6" captionText="Figure 2. Marbefia carthyi (Hamond, 1968), comb. n. (♀): A urosome, ventral [excluding P5-bearing somite] B anal somite and caudal rami, dorsal C rostrum, dorsal D antennule, ventral E labrum, anterior F paragnaths, anterior G genital area with P6 [copulatory pore arrowed]." figureDoi="http://doi.org/10.5281/zenodo.3767614" httpUri="https://zenodo.org/record/3767614/files/figure.png" pageId="11" pageNumber="12">Figure 2D</figureCitation>
|
||
). Seven-segmented, slender; with well developed sclerite around base of segment 1; several rows of long spinules present along posterior margins of all segments except for apical segment, and along anterior margins of segments 1–4. Segments 1–2 without spinous processes. Segment 2 largest; segment 4 with aesthetasc fused basally to seta and arising from distinct pedestal. Armature formula: 1-[1], 2-[7 + 1 pinnate], 3-[7], 4-[1 + (1 + ae)], 5-[1], 6-[2], 7-[7 + acrothek]. Apical acrothek consisting of small aesthetasc fused basally to 2 bare setae.
|
||
</paragraph>
|
||
<paragraph blockId="12.[140,1108,166,1672]" pageId="12" pageNumber="13">
|
||
<emphasis box="[185,277,237,263]" italics="true" pageId="12" pageNumber="13">Antenna</emphasis>
|
||
(
|
||
<figureCitation box="[293,411,237,263]" captionStart="Figure 3" captionStartId="7.[140,212,1618,1640]" captionTargetBox="[161,1095,190,1550]" captionTargetId="figure@7.[140,1108,165,1566]" captionTargetPageId="7" captionText="Figure 3. Marbefia carthyi (Hamond, 1968), comb. n. (♀): A antenna B mandibular palp C maxillule, anterior D maxilla, posterior E maxilliped, posterior." figureDoi="http://doi.org/10.5281/zenodo.3767616" httpUri="https://zenodo.org/record/3767616/files/figure.png" pageId="12" pageNumber="13">Figure 3A</figureCitation>
|
||
). Three-segmented, comprising coxa, allobasis and free 1-segmented endopod. Coxa small, with 2 rows of spinules. Allobasis not elongate; without distinct surface suture marking original segmentation but with membranous insert around base of exopod; with one abexopodal bipinnate seta in distal half. Exopod small, twice longer than width, with 4 well developed pinnate setae (2 laterally, 2 apically), distal lateral and inner distal ones with finer spinules; one row of coarse spinules on anterior surface. Endopod shorter than allobasis; lateral armature arising in distal half, consisting of one bare seta flanked by 2 strong pinnate spines; apical armature consisting of 2 strong spines and 3 geniculate setae (one geniculate seta fused basally to short seta). Endopod with 2 rows of long spinules laterally and 2 transverse hyaline frills subapically.
|
||
</paragraph>
|
||
<paragraph blockId="12.[140,1108,166,1672]" pageId="12" pageNumber="13">
|
||
<emphasis box="[185,286,589,615]" italics="true" pageId="12" pageNumber="13">Labrum.</emphasis>
|
||
Well developed; spinular ornamentation on anterior surface as in
|
||
<figureCitation captionStart="Figure 2" captionStartId="6.[140,212,1585,1607]" captionTargetBox="[161,1105,197,1555]" captionTargetId="figure@6.[140,1108,165,1566]" captionTargetPageId="6" captionText="Figure 2. Marbefia carthyi (Hamond, 1968), comb. n. (♀): A urosome, ventral [excluding P5-bearing somite] B anal somite and caudal rami, dorsal C rostrum, dorsal D antennule, ventral E labrum, anterior F paragnaths, anterior G genital area with P6 [copulatory pore arrowed]." figureDoi="http://doi.org/10.5281/zenodo.3767614" httpUri="https://zenodo.org/record/3767614/files/figure.png" pageId="12" pageNumber="13">Figure 2E</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="12.[140,1108,166,1672]" pageId="12" pageNumber="13">
|
||
<emphasis box="[185,291,659,685]" italics="true" pageId="12" pageNumber="13">Mandible</emphasis>
|
||
(
|
||
<figureCitation box="[307,425,659,686]" captionStart="Figure 3" captionStartId="7.[140,212,1618,1640]" captionTargetBox="[161,1095,190,1550]" captionTargetId="figure@7.[140,1108,165,1566]" captionTargetPageId="7" captionText="Figure 3. Marbefia carthyi (Hamond, 1968), comb. n. (♀): A antenna B mandibular palp C maxillule, anterior D maxilla, posterior E maxilliped, posterior." figureDoi="http://doi.org/10.5281/zenodo.3767616" httpUri="https://zenodo.org/record/3767616/files/figure.png" pageId="12" pageNumber="13">Figure 3B</figureCitation>
|
||
). With well developed gnathobase bearing several multicuspidate teeth around distal margin and one pinnate spine at dorsal corner as in original description (
|
||
<bibRefCitation author="Hamond R" box="[285,463,730,756]" pageId="12" pageNumber="13" pagination="172 - 176" refId="ref18534" refString="Hamond R (1968) Pseudonychocamptus carthyi n. sp. (Harpacticoida) from Hunstanton, Norfolk. Crustaceana supplement 1: 172 - 176." type="journal article" year="1968">Hamond 1968</bibRefCitation>
|
||
:
|
||
<figureCitation box="[479,550,730,757]" captionStart="Figure 6" captionStartId="10.[140,212,1585,1607]" captionTargetBox="[184,1099,180,1545]" captionTargetId="figure@10.[140,1108,165,1566]" captionTargetPageId="10" captionText="Figure 6. Marbefia carthyi (Hamond, 1968), comb. n.: A antennule ♁, ventral [armature on segments 3, 4 and 8 omitted] B antennulary segments 3 (right) and 4 (left) ♁ C antennulary segment 8 ♁ D P5 and P6 ♁, anterior E P5 ♀, anterior." figureDoi="http://doi.org/10.5281/zenodo.3767622" httpUri="https://zenodo.org/record/3767622/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
|
||
). Palp (
|
||
<figureCitation box="[644,762,730,756]" captionStart="Figure 3" captionStartId="7.[140,212,1618,1640]" captionTargetBox="[161,1095,190,1550]" captionTargetId="figure@7.[140,1108,165,1566]" captionTargetPageId="7" captionText="Figure 3. Marbefia carthyi (Hamond, 1968), comb. n. (♀): A antenna B mandibular palp C maxillule, anterior D maxilla, posterior E maxilliped, posterior." figureDoi="http://doi.org/10.5281/zenodo.3767616" httpUri="https://zenodo.org/record/3767616/files/figure.png" pageId="12" pageNumber="13">Figure 3B</figureCitation>
|
||
) small; endopod and exopod fused to basis, represented by small peduncles bearing 3 plumose and one pinnate seta(e), respectively. Basal armature represented by one plumose seta.
|
||
</paragraph>
|
||
<paragraph blockId="12.[140,1108,166,1672]" pageId="12" pageNumber="13">
|
||
<emphasis box="[185,304,835,861]" italics="true" pageId="12" pageNumber="13">Paragnaths</emphasis>
|
||
(
|
||
<figureCitation box="[322,442,835,862]" captionStart="Figure 2" captionStartId="6.[140,212,1585,1607]" captionTargetBox="[161,1105,197,1555]" captionTargetId="figure@6.[140,1108,165,1566]" captionTargetPageId="6" captionText="Figure 2. Marbefia carthyi (Hamond, 1968), comb. n. (♀): A urosome, ventral [excluding P5-bearing somite] B anal somite and caudal rami, dorsal C rostrum, dorsal D antennule, ventral E labrum, anterior F paragnaths, anterior G genital area with P6 [copulatory pore arrowed]." figureDoi="http://doi.org/10.5281/zenodo.3767614" httpUri="https://zenodo.org/record/3767614/files/figure.png" pageId="12" pageNumber="13">Figure 2F</figureCitation>
|
||
). Strongly developed paired lobes with medially directed hair-like setules, separated by medial lobe covered by dense pattern of short setules.
|
||
</paragraph>
|
||
<paragraph blockId="12.[140,1108,166,1672]" pageId="12" pageNumber="13">
|
||
<emphasis box="[185,288,906,932]" italics="true" pageId="12" pageNumber="13">Maxillule</emphasis>
|
||
(
|
||
<figureCitation box="[301,416,906,932]" captionStart="Figure 3" captionStartId="7.[140,212,1618,1640]" captionTargetBox="[161,1095,190,1550]" captionTargetId="figure@7.[140,1108,165,1566]" captionTargetPageId="7" captionText="Figure 3. Marbefia carthyi (Hamond, 1968), comb. n. (♀): A antenna B mandibular palp C maxillule, anterior D maxilla, posterior E maxilliped, posterior." figureDoi="http://doi.org/10.5281/zenodo.3767616" httpUri="https://zenodo.org/record/3767616/files/figure.png" pageId="12" pageNumber="13">Figure 3C</figureCitation>
|
||
). Praecoxa with few long spinules around outer margin; arthrite strongly developed, with a tube-seta on anterior surface and 8 spines/setae around distal margin; one transverse row of long spinules on posterior surface, and one row of small spinules along inner margin of arthrite. Coxa with cylindrical endite bearing a naked seta and a curved, pinnate spine; with spinular row on anterior surface. Basis with cylindrical endite bearing 2 naked setae and a pinnate spine; with several spinular rows on anterior surface. Endopod completely incorporated into basis, forming cluster of 3 plumose setae; exopod 1-segmented, with 2 plumose setae and several rows of fine spinules on surface.
|
||
</paragraph>
|
||
<paragraph blockId="12.[140,1108,166,1672]" pageId="12" pageNumber="13">
|
||
<emphasis box="[185,268,1187,1213]" italics="true" pageId="12" pageNumber="13">Maxilla</emphasis>
|
||
(
|
||
<figureCitation box="[282,399,1187,1214]" captionStart="Figure 3" captionStartId="7.[140,212,1618,1640]" captionTargetBox="[161,1095,190,1550]" captionTargetId="figure@7.[140,1108,165,1566]" captionTargetPageId="7" captionText="Figure 3. Marbefia carthyi (Hamond, 1968), comb. n. (♀): A antenna B mandibular palp C maxillule, anterior D maxilla, posterior E maxilliped, posterior." figureDoi="http://doi.org/10.5281/zenodo.3767616" httpUri="https://zenodo.org/record/3767616/files/figure.png" pageId="12" pageNumber="13">Figure 3D</figureCitation>
|
||
). Syncoxa with 3 endites; with a row of long spinules along outer margin, several rows of spinules on posterior surface, and several spinules along inner margin; praecoxal endite small and cylindrical, with one bare seta; both coxal endites with a strong pinnate spine and 2 pinnate setae with subapical tubular extension; proximal coxal endite with one row of spinules anteriorly. Allobasis drawn out into strong, slightly curved, distally pinnate claw; accessory armature consisting of 2 pinnate and 2 naked setae. Endopod represented by one bipinnate and 3 naked setae, surrounded by spinules at base.
|
||
</paragraph>
|
||
<paragraph blockId="12.[140,1108,166,1672]" pageId="12" pageNumber="13">
|
||
<emphasis box="[185,304,1434,1460]" italics="true" pageId="12" pageNumber="13">Maxilliped</emphasis>
|
||
(
|
||
<figureCitation box="[320,437,1434,1460]" captionStart="Figure 3" captionStartId="7.[140,212,1618,1640]" captionTargetBox="[161,1095,190,1550]" captionTargetId="figure@7.[140,1108,165,1566]" captionTargetPageId="7" captionText="Figure 3. Marbefia carthyi (Hamond, 1968), comb. n. (♀): A antenna B mandibular palp C maxillule, anterior D maxilla, posterior E maxilliped, posterior." figureDoi="http://doi.org/10.5281/zenodo.3767616" httpUri="https://zenodo.org/record/3767616/files/figure.png" pageId="12" pageNumber="13">Figure 3E</figureCitation>
|
||
). Syncoxa elongate, with 2 plumose setae and two rows of spinules. Basis with one row of spinules along outer margin, palmar margin completely bare. Endopod drawn out into long claw; with one short accessory seta anteriorly, and one tube-pore posteriorly.
|
||
</paragraph>
|
||
<paragraph blockId="12.[140,1108,166,1672]" pageId="12" pageNumber="13">
|
||
<emphasis box="[185,439,1575,1601]" italics="true" pageId="12" pageNumber="13">Swimming legs. P1–P4</emphasis>
|
||
(
|
||
<figureCitation box="[455,615,1575,1601]" captionStart="Figure 4" captionStartId="8.[140,212,1585,1607]" captionTargetBox="[146,1088,177,1488]" captionTargetId="figure@8.[140,1108,165,1566]" captionTargetPageId="8" captionText="Figure 4. Marbefia carthyi (Hamond, 1968), comb. n.: A P1 ♀, anterior B P4 ♀, anterior [inner seta on enp-1 arrowed] C P4 endopod ♁, anterior [spinular rows on enp-1 arrowed] D distal part of P4 endopod ♀, posterior." figureDoi="http://doi.org/10.5281/zenodo.3767618" httpUri="https://zenodo.org/record/3767618/files/figure.png" pageId="12" pageNumber="13">Figures 4A, B</figureCitation>
|
||
;
|
||
<figureCitation box="[628,641,1574,1601]" captionStart="Figure 5" captionStartId="9.[140,212,1585,1607]" captionTargetBox="[152,1104,167,1554]" captionTargetId="figure@9.[140,1108,165,1566]" captionTargetPageId="9" captionText="Figure 5. Marbefia carthyi (Hamond, 1968), comb. n.: A P2 ♀, anterior, B P2 endopod ♀, outer lateral C P3 ♀, anterior [inner seta on enp-1 arrowed] D P3 endopod ♁ [spinules on enp-1 and apophysis on enp-2 arrowed]." figureDoi="http://doi.org/10.5281/zenodo.3767620" httpUri="https://zenodo.org/record/3767620/files/figure.png" pageId="12" pageNumber="13">5</figureCitation>
|
||
A–C). With wide intercoxal sclerites and well developed praecoxae. Praecoxae, coxae and bases with anterior rows of surface spinules as figured. Exopods 3-segmented, endopods 2-segmented.
|
||
</paragraph>
|
||
<paragraph blockId="13.[140,1108,167,1672]" pageId="13" pageNumber="14">
|
||
<emphasis box="[185,217,167,193]" italics="true" pageId="13" pageNumber="14">P1</emphasis>
|
||
(
|
||
<figureCitation box="[230,348,167,193]" captionStart="Figure 4" captionStartId="8.[140,212,1585,1607]" captionTargetBox="[146,1088,177,1488]" captionTargetId="figure@8.[140,1108,165,1566]" captionTargetPageId="8" captionText="Figure 4. Marbefia carthyi (Hamond, 1968), comb. n.: A P1 ♀, anterior B P4 ♀, anterior [inner seta on enp-1 arrowed] C P4 endopod ♁, anterior [spinular rows on enp-1 arrowed] D distal part of P4 endopod ♀, posterior." figureDoi="http://doi.org/10.5281/zenodo.3767618" httpUri="https://zenodo.org/record/3767618/files/figure.png" pageId="13" pageNumber="14">Figure 4A</figureCitation>
|
||
). Coxa large; with several spinular rows and patches as figured. Basis with bipinnate seta on distal pedestal near insertion of endopod, long setules along inner margin and one bipinnate spine and several spinules along outer margin; anterior tube-pore near articulation with coxa. Exp-1 with one bipinnate seta; exp-2 with one naked outer seta; exp-3 with 2 virtually naked and 2 geniculate setae. Enp-1 1.8 times as long as exopod, with long spinules along proximal half of inner margin; enp-2 with one strong, minutely denticulate claw, and one small naked seta; several spinules along outer margin and around inner distal corner.
|
||
</paragraph>
|
||
<paragraph blockId="13.[140,1108,167,1672]" pageId="13" pageNumber="14">
|
||
<emphasis box="[185,261,448,474]" italics="true" pageId="13" pageNumber="14">P2–P4</emphasis>
|
||
(
|
||
<figureCitation box="[275,398,448,474]" captionStart="Figure 4" captionStartId="8.[140,212,1585,1607]" captionTargetBox="[146,1088,177,1488]" captionTargetId="figure@8.[140,1108,165,1566]" captionTargetPageId="8" captionText="Figure 4. Marbefia carthyi (Hamond, 1968), comb. n.: A P1 ♀, anterior B P4 ♀, anterior [inner seta on enp-1 arrowed] C P4 endopod ♁, anterior [spinular rows on enp-1 arrowed] D distal part of P4 endopod ♀, posterior." figureDoi="http://doi.org/10.5281/zenodo.3767618" httpUri="https://zenodo.org/record/3767618/files/figure.png" pageId="13" pageNumber="14">Figures 4B</figureCitation>
|
||
;
|
||
<figureCitation box="[409,476,448,475]" captionStart="Figure 5" captionStartId="9.[140,212,1585,1607]" captionTargetBox="[152,1104,167,1554]" captionTargetId="figure@9.[140,1108,165,1566]" captionTargetPageId="9" captionText="Figure 5. Marbefia carthyi (Hamond, 1968), comb. n.: A P2 ♀, anterior, B P2 endopod ♀, outer lateral C P3 ♀, anterior [inner seta on enp-1 arrowed] D P3 endopod ♁ [spinules on enp-1 and apophysis on enp-2 arrowed]." figureDoi="http://doi.org/10.5281/zenodo.3767620" httpUri="https://zenodo.org/record/3767620/files/figure.png" pageId="13" pageNumber="14">5A, B</figureCitation>
|
||
). Coxae and bases with spinular rows along outer margin and on anterior surface; basis with tube-pore on anterior surface; outer margin of basis with bipinnate spine (P2) or naked seta (P3–P4); exopodal and endopodal segments with elaborate spinular/setular ornamentation as figured. P4 outer exopodal spines with long spinules. P2–P4 endopods with short enp-1 and long, inflated enp-2; outer margin of enp-2 weakly chitinized, with 2 parallel rows of flimsy setular extensions, possibly surrounding glandular structure (
|
||
<figureCitation box="[517,634,659,686]" captionStart="Figure 5" captionStartId="9.[140,212,1585,1607]" captionTargetBox="[152,1104,167,1554]" captionTargetId="figure@9.[140,1108,165,1566]" captionTargetPageId="9" captionText="Figure 5. Marbefia carthyi (Hamond, 1968), comb. n.: A P2 ♀, anterior, B P2 endopod ♀, outer lateral C P3 ♀, anterior [inner seta on enp-1 arrowed] D P3 endopod ♁ [spinules on enp-1 and apophysis on enp-2 arrowed]." figureDoi="http://doi.org/10.5281/zenodo.3767620" httpUri="https://zenodo.org/record/3767620/files/figure.png" pageId="13" pageNumber="14">Figure 5B</figureCitation>
|
||
). P3–P4 enp-2 with tube-pore near distal margin. P2 enp-2 twice times longer than enp-1; endopod reaching to distal margin of exp-2; exp-3 longest. P3 enp-2 2.5 times longer than enp-1; endopod reaching to distal margin of exp-2; exp-3 longest. P4 enp-2 2.6 times longer than enp-1, posterior distal margin with spinular row (
|
||
<figureCitation box="[453,576,800,826]" captionStart="Figure 4" captionStartId="8.[140,212,1585,1607]" captionTargetBox="[146,1088,177,1488]" captionTargetId="figure@8.[140,1108,165,1566]" captionTargetPageId="8" captionText="Figure 4. Marbefia carthyi (Hamond, 1968), comb. n.: A P1 ♀, anterior B P4 ♀, anterior [inner seta on enp-1 arrowed] C P4 endopod ♁, anterior [spinular rows on enp-1 arrowed] D distal part of P4 endopod ♀, posterior." figureDoi="http://doi.org/10.5281/zenodo.3767618" httpUri="https://zenodo.org/record/3767618/files/figure.png" pageId="13" pageNumber="14">Figure 4D</figureCitation>
|
||
); endopod reaching to about distal margin of exp-2; exp-3 subequal to exp-1. Outer distal seta of P4 enp-2 clearly shorter than others; inner seta of enp-1 very long (arrowed in
|
||
<figureCitation box="[657,770,871,897]" captionStart="Figure 4" captionStartId="8.[140,212,1585,1607]" captionTargetBox="[146,1088,177,1488]" captionTargetId="figure@8.[140,1108,165,1566]" captionTargetPageId="8" captionText="Figure 4. Marbefia carthyi (Hamond, 1968), comb. n.: A P1 ♀, anterior B P4 ♀, anterior [inner seta on enp-1 arrowed] C P4 endopod ♁, anterior [spinular rows on enp-1 arrowed] D distal part of P4 endopod ♀, posterior." figureDoi="http://doi.org/10.5281/zenodo.3767618" httpUri="https://zenodo.org/record/3767618/files/figure.png" pageId="13" pageNumber="14">Figure 4B</figureCitation>
|
||
) and much longer than in P3 enp-1 (arrowed in
|
||
<figureCitation box="[348,464,906,933]" captionStart="Figure 5" captionStartId="9.[140,212,1585,1607]" captionTargetBox="[152,1104,167,1554]" captionTargetId="figure@9.[140,1108,165,1566]" captionTargetPageId="9" captionText="Figure 5. Marbefia carthyi (Hamond, 1968), comb. n.: A P2 ♀, anterior, B P2 endopod ♀, outer lateral C P3 ♀, anterior [inner seta on enp-1 arrowed] D P3 endopod ♁ [spinules on enp-1 and apophysis on enp-2 arrowed]." figureDoi="http://doi.org/10.5281/zenodo.3767620" httpUri="https://zenodo.org/record/3767620/files/figure.png" pageId="13" pageNumber="14">Figure 5C</figureCitation>
|
||
). Spine and setal formulae of swimming legs as for genus.
|
||
</paragraph>
|
||
<paragraph blockId="13.[140,1108,167,1672]" pageId="13" pageNumber="14">
|
||
<emphasis box="[185,373,941,967]" italics="true" pageId="13" pageNumber="14">Fifth pair of legs</emphasis>
|
||
(
|
||
<figureCitation box="[392,513,941,968]" captionStart="Figure 6" captionStartId="10.[140,212,1585,1607]" captionTargetBox="[184,1099,180,1545]" captionTargetId="figure@10.[140,1108,165,1566]" captionTargetPageId="10" captionText="Figure 6. Marbefia carthyi (Hamond, 1968), comb. n.: A antennule ♁, ventral [armature on segments 3, 4 and 8 omitted] B antennulary segments 3 (right) and 4 (left) ♁ C antennulary segment 8 ♁ D P5 and P6 ♁, anterior E P5 ♀, anterior." figureDoi="http://doi.org/10.5281/zenodo.3767622" httpUri="https://zenodo.org/record/3767622/files/figure.png" pageId="13" pageNumber="14">Figure 6E</figureCitation>
|
||
). Baseoendopods not medially fused, with distinct medial extension. Exopod and baseoendopod discrete, each with pattern of spinules as figured; spinular pattern on exopod denser than on baseoendopod. Baseoendopod forming long, outer setophore bearing basal seta and rows of spinules; with 3 tubepores on anterior surface; endopodal lobe just extending to proximal area of exopod, with 2 apical and 3 lateral bipinnate setae, proximal and inner middle ones minutely pinnate. Exopod elongate, about 3 times longer than wide; with one naked terminal seta, one bipinnate inner seta, and one bare and 3 pinnate setae along outer margin; terminal seta arising from short cylindrical process.
|
||
</paragraph>
|
||
<paragraph blockId="13.[140,1108,167,1672]" box="[185,782,1258,1284]" pageId="13" pageNumber="14">
|
||
<emphasis box="[185,272,1259,1284]" italics="true" pageId="13" pageNumber="14">Egg-sac.</emphasis>
|
||
Number of eggs and arrangement unknown.
|
||
</paragraph>
|
||
<paragraph blockId="13.[140,1108,167,1672]" pageId="13" pageNumber="14">
|
||
<emphasis bold="true" box="[185,335,1293,1319]" pageId="13" pageNumber="14">
|
||
<emphasis box="[185,330,1293,1319]" inLineHeading="true" pageId="13" pageNumber="14" reason="1">Description</emphasis>
|
||
.
|
||
</emphasis>
|
||
Male.
|
||
<emphasis box="[414,468,1293,1319]" italics="true" pageId="13" pageNumber="14">Body</emphasis>
|
||
(Figure 1C). Body length
|
||
<quantity box="[769,917,1293,1320]" metricMagnitude="-1" metricUnit="m" metricValue="5.785" metricValueMax="5.87" metricValueMin="5.7" pageId="13" pageNumber="14" unit="mm" value="578.5" valueMax="587.0" valueMin="570.0">570–587 µm</quantity>
|
||
(n=3; mean
|
||
<quantity metricMagnitude="-1" metricUnit="m" metricValue="5.79" pageId="13" pageNumber="14" unit="mm" value="579.0">579 µm</quantity>
|
||
). Largest width at posterior margin of cephalic shield (
|
||
<quantity box="[786,879,1328,1355]" metricMagnitude="-1" metricUnit="m" metricValue="1.44" pageId="13" pageNumber="14" unit="mm" value="144.0">144 µm</quantity>
|
||
). Urosome narrower than prosome. Prosome (Figure 1C) 4-segmented, comprising cephalothorax and 3 free pedigerous somites. Cephalothorax with smooth posterior margin; pleural areas well developed and rounded, with lobate posterolateral angles; whole surface covered with tiny spinules as in
|
||
<specimenCount box="[410,425,1470,1496]" pageId="13" pageNumber="14" type="female">♀</specimenCount>
|
||
. Rostrum as in
|
||
<specimenCount box="[614,629,1470,1496]" pageId="13" pageNumber="14" type="female">♀</specimenCount>
|
||
. Pedigerous somites covered with minute spinules as in
|
||
<specimenCount box="[302,317,1505,1531]" pageId="13" pageNumber="14" type="female">♀</specimenCount>
|
||
. Urosome (Figure 1C) 6-segmented, comprising P5-bearing somite, genital somite and 4 abdominal somites. All urosomites with surface ornamentation consisting of tiny spinules dorsally and laterally; ventral surface smooth except for few rows of small spinules as in
|
||
<specimenCount box="[445,460,1610,1636]" pageId="13" pageNumber="14" type="female">♀</specimenCount>
|
||
; ventral posterior margin with large spinules laterally and minute spinules medially. Hyaline frills of urosomites not distinct.
|
||
</paragraph>
|
||
<paragraph blockId="14.[140,1108,167,1672]" pageId="14" pageNumber="15">
|
||
<emphasis box="[185,296,167,193]" italics="true" pageId="14" pageNumber="15">Antennule</emphasis>
|
||
(
|
||
<figureCitation box="[318,419,167,194]" captionStart="Figure 6" captionStartId="10.[140,212,1585,1607]" captionTargetBox="[184,1099,180,1545]" captionTargetId="figure@10.[140,1108,165,1566]" captionTargetPageId="10" captionText="Figure 6. Marbefia carthyi (Hamond, 1968), comb. n.: A antennule ♁, ventral [armature on segments 3, 4 and 8 omitted] B antennulary segments 3 (right) and 4 (left) ♁ C antennulary segment 8 ♁ D P5 and P6 ♁, anterior E P5 ♀, anterior." figureDoi="http://doi.org/10.5281/zenodo.3767622" httpUri="https://zenodo.org/record/3767622/files/figure.png" pageId="14" pageNumber="15">Figure 6</figureCitation>
|
||
A-C). Eight-segmented; subchirocer with geniculation between segments 5 and 6. Segments 1 and 2 with several rows of long spinules along anterior and posterior margins; without protuberances. Segment 3 (
|
||
<figureCitation box="[922,1038,237,264]" captionStart="Figure 6" captionStartId="10.[140,212,1585,1607]" captionTargetBox="[184,1099,180,1545]" captionTargetId="figure@10.[140,1108,165,1566]" captionTargetPageId="10" captionText="Figure 6. Marbefia carthyi (Hamond, 1968), comb. n.: A antennule ♁, ventral [armature on segments 3, 4 and 8 omitted] B antennulary segments 3 (right) and 4 (left) ♁ C antennulary segment 8 ♁ D P5 and P6 ♁, anterior E P5 ♀, anterior." figureDoi="http://doi.org/10.5281/zenodo.3767622" httpUri="https://zenodo.org/record/3767622/files/figure.png" pageId="14" pageNumber="15">Figure 6B</figureCitation>
|
||
) with few spinules along anterior margin. Segment 4 represented by small U-shaped sclerite around anterior margin (see insert in
|
||
<figureCitation box="[572,689,307,334]" captionStart="Figure 6" captionStartId="10.[140,212,1585,1607]" captionTargetBox="[184,1099,180,1545]" captionTargetId="figure@10.[140,1108,165,1566]" captionTargetPageId="10" captionText="Figure 6. Marbefia carthyi (Hamond, 1968), comb. n.: A antennule ♁, ventral [armature on segments 3, 4 and 8 omitted] B antennulary segments 3 (right) and 4 (left) ♁ C antennulary segment 8 ♁ D P5 and P6 ♁, anterior E P5 ♀, anterior." figureDoi="http://doi.org/10.5281/zenodo.3767622" httpUri="https://zenodo.org/record/3767622/files/figure.png" pageId="14" pageNumber="15">Figure 6B</figureCitation>
|
||
). Segment 5 swollen. Segment 8 triangular (
|
||
<figureCitation box="[240,361,343,370]" captionStart="Figure 6" captionStartId="10.[140,212,1585,1607]" captionTargetBox="[184,1099,180,1545]" captionTargetId="figure@10.[140,1108,165,1566]" captionTargetPageId="10" captionText="Figure 6. Marbefia carthyi (Hamond, 1968), comb. n.: A antennule ♁, ventral [armature on segments 3, 4 and 8 omitted] B antennulary segments 3 (right) and 4 (left) ♁ C antennulary segment 8 ♁ D P5 and P6 ♁, anterior E P5 ♀, anterior." figureDoi="http://doi.org/10.5281/zenodo.3767622" httpUri="https://zenodo.org/record/3767622/files/figure.png" pageId="14" pageNumber="15">Figure 6C</figureCitation>
|
||
). Segmental homologies: 1-(I), 2-(II–VIII), 3-(IX–XII), 4-(XIII), 5-(XIV–XX), 6-(XXI–XXII), 7-(XXIII), 8-(XXIV–XXVIII). Armature formula: 1-[1], 2-[1 pinnate + 8], 3-[8], 4-[2], 5-[9 + 1 pinnate + 1 modified + (1 + ae)], 6-[3 modified spinous elements], 7-[1], 8-[8 + acrothek]. Apical acrothek consisting of minute aesthetasc and 2 naked setae.
|
||
</paragraph>
|
||
<paragraph blockId="14.[140,1108,167,1672]" pageId="14" pageNumber="15">
|
||
<emphasis box="[185,438,519,545]" italics="true" pageId="14" pageNumber="15">Swimming legs. P2–P4</emphasis>
|
||
(
|
||
<figureCitation box="[453,582,519,545]" captionStart="Figure 4" captionStartId="8.[140,212,1585,1607]" captionTargetBox="[146,1088,177,1488]" captionTargetId="figure@8.[140,1108,165,1566]" captionTargetPageId="8" captionText="Figure 4. Marbefia carthyi (Hamond, 1968), comb. n.: A P1 ♀, anterior B P4 ♀, anterior [inner seta on enp-1 arrowed] C P4 endopod ♁, anterior [spinular rows on enp-1 arrowed] D distal part of P4 endopod ♀, posterior." figureDoi="http://doi.org/10.5281/zenodo.3767618" httpUri="https://zenodo.org/record/3767618/files/figure.png" pageId="14" pageNumber="15">Figures 4C</figureCitation>
|
||
;
|
||
<figureCitation box="[595,635,518,545]" captionStart="Figure 5" captionStartId="9.[140,212,1585,1607]" captionTargetBox="[152,1104,167,1554]" captionTargetId="figure@9.[140,1108,165,1566]" captionTargetPageId="9" captionText="Figure 5. Marbefia carthyi (Hamond, 1968), comb. n.: A P2 ♀, anterior, B P2 endopod ♀, outer lateral C P3 ♀, anterior [inner seta on enp-1 arrowed] D P3 endopod ♁ [spinules on enp-1 and apophysis on enp-2 arrowed]." figureDoi="http://doi.org/10.5281/zenodo.3767620" httpUri="https://zenodo.org/record/3767620/files/figure.png" pageId="14" pageNumber="15">5D</figureCitation>
|
||
). Shape and surface ornamentation of intercoxal sclerites and protopods as in
|
||
<specimenCount box="[587,602,554,580]" pageId="14" pageNumber="15" type="female">♀</specimenCount>
|
||
. P2–P3 enp-2 with double row of setular extensions along outer lateral margin as in
|
||
<specimenCount box="[643,658,590,616]" pageId="14" pageNumber="15" type="female">♀</specimenCount>
|
||
(absent in P4;
|
||
<figureCitation box="[849,969,589,615]" captionStart="Figure 4" captionStartId="8.[140,212,1585,1607]" captionTargetBox="[146,1088,177,1488]" captionTargetId="figure@8.[140,1108,165,1566]" captionTargetPageId="8" captionText="Figure 4. Marbefia carthyi (Hamond, 1968), comb. n.: A P1 ♀, anterior B P4 ♀, anterior [inner seta on enp-1 arrowed] C P4 endopod ♁, anterior [spinular rows on enp-1 arrowed] D distal part of P4 endopod ♀, posterior." figureDoi="http://doi.org/10.5281/zenodo.3767618" httpUri="https://zenodo.org/record/3767618/files/figure.png" pageId="14" pageNumber="15">Figure 4C</figureCitation>
|
||
). No sexual dimorphism on P2. Endopodal segments of P3–P4 modified.
|
||
</paragraph>
|
||
<paragraph blockId="14.[140,1108,167,1672]" pageId="14" pageNumber="15">
|
||
<emphasis box="[185,360,660,686]" italics="true" pageId="14" pageNumber="15">
|
||
P3 (
|
||
<figureCitation box="[230,347,660,686]" captionStart="Figure 5" captionStartId="9.[140,212,1585,1607]" captionTargetBox="[152,1104,167,1554]" captionTargetId="figure@9.[140,1108,165,1566]" captionTargetPageId="9" captionText="Figure 5. Marbefia carthyi (Hamond, 1968), comb. n.: A P2 ♀, anterior, B P2 endopod ♀, outer lateral C P3 ♀, anterior [inner seta on enp-1 arrowed] D P3 endopod ♁ [spinules on enp-1 and apophysis on enp-2 arrowed]." figureDoi="http://doi.org/10.5281/zenodo.3767620" httpUri="https://zenodo.org/record/3767620/files/figure.png" pageId="14" pageNumber="15">Figure 5D</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Exopod as in
|
||
<specimenCount box="[517,532,660,686]" pageId="14" pageNumber="15" type="female">♀</specimenCount>
|
||
. Endopod 2-segmented, modified, not reaching to distal margin of exp-2. Enp-1 with several spinules on inner distal margin (arrowed in
|
||
<figureCitation box="[140,259,730,757]" captionStart="Figure 5" captionStartId="9.[140,212,1585,1607]" captionTargetBox="[152,1104,167,1554]" captionTargetId="figure@9.[140,1108,165,1566]" captionTargetPageId="9" captionText="Figure 5. Marbefia carthyi (Hamond, 1968), comb. n.: A P2 ♀, anterior, B P2 endopod ♀, outer lateral C P3 ♀, anterior [inner seta on enp-1 arrowed] D P3 endopod ♁ [spinules on enp-1 and apophysis on enp-2 arrowed]." figureDoi="http://doi.org/10.5281/zenodo.3767620" httpUri="https://zenodo.org/record/3767620/files/figure.png" pageId="14" pageNumber="15">Figure 5D</figureCitation>
|
||
); inner seta absent. Enp-2 produced distally into short, straight, spiniform apophysis (homologous with outer spine of enp-
|
||
<quantity box="[700,746,765,791]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="14" pageNumber="15" unit="in" value="2.0">2 in</quantity>
|
||
<specimenCount box="[756,771,766,792]" pageId="14" pageNumber="15" type="female">♀</specimenCount>
|
||
; arrowed in
|
||
<figureCitation box="[927,1046,765,792]" captionStart="Figure 5" captionStartId="9.[140,212,1585,1607]" captionTargetBox="[152,1104,167,1554]" captionTargetId="figure@9.[140,1108,165,1566]" captionTargetPageId="9" captionText="Figure 5. Marbefia carthyi (Hamond, 1968), comb. n.: A P2 ♀, anterior, B P2 endopod ♀, outer lateral C P3 ♀, anterior [inner seta on enp-1 arrowed] D P3 endopod ♁ [spinules on enp-1 and apophysis on enp-2 arrowed]." figureDoi="http://doi.org/10.5281/zenodo.3767620" httpUri="https://zenodo.org/record/3767620/files/figure.png" pageId="14" pageNumber="15">Figure 5D</figureCitation>
|
||
) surrounded at base by row of coarse spinules.
|
||
</paragraph>
|
||
<paragraph blockId="14.[140,1108,167,1672]" pageId="14" pageNumber="15">
|
||
<emphasis box="[185,358,836,862]" italics="true" pageId="14" pageNumber="15">
|
||
P4 (
|
||
<figureCitation box="[230,344,836,862]" captionStart="Figure 4" captionStartId="8.[140,212,1585,1607]" captionTargetBox="[146,1088,177,1488]" captionTargetId="figure@8.[140,1108,165,1566]" captionTargetPageId="8" captionText="Figure 4. Marbefia carthyi (Hamond, 1968), comb. n.: A P1 ♀, anterior B P4 ♀, anterior [inner seta on enp-1 arrowed] C P4 endopod ♁, anterior [spinular rows on enp-1 arrowed] D distal part of P4 endopod ♀, posterior." figureDoi="http://doi.org/10.5281/zenodo.3767618" httpUri="https://zenodo.org/record/3767618/files/figure.png" pageId="14" pageNumber="15">Figure 4C</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Exopod as in
|
||
<specimenCount box="[515,530,836,862]" pageId="14" pageNumber="15" type="female">♀</specimenCount>
|
||
. Endopod relatively shorter than in
|
||
<specimenCount box="[943,958,836,862]" pageId="14" pageNumber="15" type="female">♀</specimenCount>
|
||
. Enp-1 with several spinules along inner margin (arrowed in
|
||
<figureCitation box="[696,814,871,897]" captionStart="Figure 4" captionStartId="8.[140,212,1585,1607]" captionTargetBox="[146,1088,177,1488]" captionTargetId="figure@8.[140,1108,165,1566]" captionTargetPageId="8" captionText="Figure 4. Marbefia carthyi (Hamond, 1968), comb. n.: A P1 ♀, anterior B P4 ♀, anterior [inner seta on enp-1 arrowed] C P4 endopod ♁, anterior [spinular rows on enp-1 arrowed] D distal part of P4 endopod ♀, posterior." figureDoi="http://doi.org/10.5281/zenodo.3767618" httpUri="https://zenodo.org/record/3767618/files/figure.png" pageId="14" pageNumber="15">Figure 4C</figureCitation>
|
||
); inner seta absent; outer lateral margin without setular extensions found in
|
||
<specimenCount box="[713,728,906,932]" pageId="14" pageNumber="15" type="female">♀</specimenCount>
|
||
. Enp-2 twice longer than enp-1; both apical setae about equally long and longer than in
|
||
<specimenCount box="[768,783,942,968]" pageId="14" pageNumber="15" type="female">♀</specimenCount>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="14.[140,1108,167,1672]" pageId="14" pageNumber="15">
|
||
<emphasis box="[185,364,976,1002]" italics="true" pageId="14" pageNumber="15">Fifth pair of legs</emphasis>
|
||
(
|
||
<figureCitation box="[380,503,976,1003]" captionStart="Figure 6" captionStartId="10.[140,212,1585,1607]" captionTargetBox="[184,1099,180,1545]" captionTargetId="figure@10.[140,1108,165,1566]" captionTargetPageId="10" captionText="Figure 6. Marbefia carthyi (Hamond, 1968), comb. n.: A antennule ♁, ventral [armature on segments 3, 4 and 8 omitted] B antennulary segments 3 (right) and 4 (left) ♁ C antennulary segment 8 ♁ D P5 and P6 ♁, anterior E P5 ♀, anterior." figureDoi="http://doi.org/10.5281/zenodo.3767622" httpUri="https://zenodo.org/record/3767622/files/figure.png" pageId="14" pageNumber="15">Figure 6D</figureCitation>
|
||
). Baseoendopods fused medially, with spinular patch medially. Baseoendopod with large setophore bearing outer basal seta; endopodal lobe vestigial bearing one pinnate and one bare seta; with one tube-pore along medial margin, and one tube-pore near articulation with exopod; with few coarse spinules at base of setophore. Exopod about twice as long as maximum width; with one inner, one apical and 2 outer setae; all setae pinnate except for distal outer one being smooth; anterior surface with several large spinules.
|
||
</paragraph>
|
||
<paragraph blockId="14.[140,1108,167,1672]" pageId="14" pageNumber="15">
|
||
<emphasis box="[185,371,1223,1249]" italics="true" pageId="14" pageNumber="15">Sixth pair of legs</emphasis>
|
||
(
|
||
<figureCitation box="[388,512,1223,1250]" captionStart="Figure 6" captionStartId="10.[140,212,1585,1607]" captionTargetBox="[184,1099,180,1545]" captionTargetId="figure@10.[140,1108,165,1566]" captionTargetPageId="10" captionText="Figure 6. Marbefia carthyi (Hamond, 1968), comb. n.: A antennule ♁, ventral [armature on segments 3, 4 and 8 omitted] B antennulary segments 3 (right) and 4 (left) ♁ C antennulary segment 8 ♁ D P5 and P6 ♁, anterior E P5 ♀, anterior." figureDoi="http://doi.org/10.5281/zenodo.3767622" httpUri="https://zenodo.org/record/3767622/files/figure.png" pageId="14" pageNumber="15">Figure 6D</figureCitation>
|
||
). Asymmetrical; represented on both sides by small plate (fused to ventral wall of supporting somite on one side; articulating at base and covering gonopore on other side; dextral and sinistral configurations observed); outer distal corner produced into cylindrical process bearing several spinules, one bipinnate inner and one naked outer seta.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15" type="discussion">
|
||
<paragraph blockId="14.[140,1108,167,1672]" lastBlockId="15.[140,1108,167,1672]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
|
||
<emphasis bold="true" box="[185,324,1398,1424]" pageId="14" pageNumber="15">
|
||
<emphasis box="[185,318,1398,1424]" inLineHeading="true" pageId="14" pageNumber="15" reason="1">Discussion</emphasis>
|
||
.
|
||
</emphasis>
|
||
<bibRefCitation author="Hamond R" box="[336,542,1399,1426]" pageId="14" pageNumber="15" pagination="172 - 176" refId="ref18534" refString="Hamond R (1968) Pseudonychocamptus carthyi n. sp. (Harpacticoida) from Hunstanton, Norfolk. Crustaceana supplement 1: 172 - 176." type="journal article" year="1968">Hamond’s (1968)</bibRefCitation>
|
||
original description of the female of
|
||
<taxonomicName authorityName="Hamond" authorityYear="1968" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="carthyi">
|
||
<emphasis italics="true" pageId="14" pageNumber="15">Pseudonychocamptus carthyi</emphasis>
|
||
</taxonomicName>
|
||
is detailed by contemporary standards. Our redescription revealed some minor observational errors: (1) the entire body is covered by fine spinules instead “… a few long limp hairs, widely scattered here and there”; (2) the third antennulary segment has 7 setae instead of 5; (3) the distal lateral and inner distal setae of the antennary exopod have fine spinules instead of membranous laminae; (4) the antennary endopod has an additional subdistal spine and the outermost geniculate seta is fused at the base to a minute seta; (5) the basis and coxal endite of the maxillule have an ad- ditional seta; (6) the maxilla has a unisetose praecoxal endite and the endopod 4 setae instead of 2; (7) the P1 displays distinct spinule rows along the outer margin of the praecoxa and coxa, and enp-2 has a minute accessory seta at the base of the claw; (8) the outer spines of P2–P3 exp-2 and -3 are bipinnate instead of smooth; and (9) the caudal ramus has 7 setae (seta I was overlooked).
|
||
</paragraph>
|
||
<paragraph blockId="15.[140,1108,167,1672]" pageId="15" pageNumber="16">
|
||
The discovery of the male of
|
||
<taxonomicName authorityName="Hamond" authorityYear="1968" box="[537,644,343,369]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="carthyi">
|
||
<emphasis box="[537,644,343,369]" italics="true" pageId="15" pageNumber="16">P. carthyi</emphasis>
|
||
</taxonomicName>
|
||
provides conclusive evidence not only for the inclusion of this species in Lee and Huys’ (1999)
|
||
<emphasis box="[829,911,379,404]" italics="true" pageId="15" pageNumber="16">PWPH</emphasis>
|
||
-clade (grouping the genera
|
||
<taxonomicName authorityName="Noodt" authorityYear="1952" box="[269,353,413,439]" class="Maxillopoda" family="Laophontidae" genus="Pilifera" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[269,353,413,439]" italics="true" pageId="15" pageNumber="16">Pilifera</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName box="[368,601,413,439]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[368,601,413,439]" italics="true" pageId="15" pageNumber="16">Pseudonychocamptus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Willen" authorityYear="1996" box="[616,808,413,439]" class="Maxillopoda" family="Laophontidae" genus="Weddellaophonte" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[616,808,413,439]" italics="true" pageId="15" pageNumber="16">Weddellaophonte</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Lee & Huys" authorityYear="1999" box="[866,1098,413,439]" class="Maxillopoda" family="Laophontidae" genus="Heteronychocamptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[866,1098,413,439]" italics="true" pageId="15" pageNumber="16">Heteronychocamptus</emphasis>
|
||
</taxonomicName>
|
||
) but also for its exclusion from the genus
|
||
<taxonomicName box="[639,872,448,474]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[639,872,448,474]" italics="true" pageId="15" pageNumber="16">Pseudonychocamptus</emphasis>
|
||
</taxonomicName>
|
||
. The sexual dimorphism in swimming leg armature (the inner seta of P3–P4 enp-1 being absent in the male) unequivocally links this species to the
|
||
<emphasis box="[771,853,519,544]" italics="true" pageId="15" pageNumber="16">PWPH</emphasis>
|
||
-clade (Table 1). The sexually dimorphic P4 endopod, being 2-segmented in the female but reduced to a minute bisetose segment in the male, is the most significant apomorphy of the genus
|
||
<taxonomicName box="[215,448,624,650]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[215,448,624,650]" italics="true" pageId="15" pageNumber="16">Pseudonychocamptus</emphasis>
|
||
</taxonomicName>
|
||
, however, no such reduction is displayed by the male of
|
||
<taxonomicName authorityName="Hamond" authorityYear="1968" box="[140,245,659,685]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="carthyi">
|
||
<emphasis box="[140,245,659,685]" italics="true" pageId="15" pageNumber="16">P. carthyi</emphasis>
|
||
</taxonomicName>
|
||
. This species also displays the most primitive armature on P2–P5 within the
|
||
<emphasis box="[188,270,695,720]" italics="true" pageId="15" pageNumber="16">PWPH</emphasis>
|
||
-clade (Table 1), lending additional support to a different generic assignment. The only other genus exhibiting two inner setae on P3–P4 exp-3 is
|
||
<taxonomicName authorityName="Willen" authorityYear="1996" box="[140,332,765,791]" class="Maxillopoda" family="Laophontidae" genus="Weddellaophonte" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[140,332,765,791]" italics="true" pageId="15" pageNumber="16">Weddellaophonte</emphasis>
|
||
</taxonomicName>
|
||
, however, in
|
||
<taxonomicName authorityName="Hamond" authorityYear="1968" box="[488,593,765,791]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="carthyi">
|
||
<emphasis box="[488,593,765,791]" italics="true" pageId="15" pageNumber="16">P. carthyi</emphasis>
|
||
</taxonomicName>
|
||
two additional setae are found on P3 enp-2 (formula 321).
|
||
<taxonomicName authorityName="Hamond" authorityYear="1968" box="[322,640,800,826]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="carthyi">
|
||
<emphasis box="[322,640,800,826]" italics="true" pageId="15" pageNumber="16">Pseudonychocamptus carthyi</emphasis>
|
||
</taxonomicName>
|
||
is here fixed as the
|
||
<typeStatus box="[868,919,801,827]" pageId="15" pageNumber="16">type</typeStatus>
|
||
of a new genus,
|
||
<taxonomicName authority="Huys & Lee, 2009" authorityName="Huys & Lee" authorityYear="2009" box="[140,244,835,861]" class="Maxillopoda" family="Laophontidae" genus="Marbefia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis box="[140,244,835,861]" italics="true" pageId="15" pageNumber="16">Marbefia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[264,352,836,862]" pageId="15" pageNumber="16" rank="genus">gen. n.</taxonomicNameLabel>
|
||
, which can be defined by the following autapomorphies: (1) P2–P4 enp-2 (except P4 enp-
|
||
<quantity box="[496,543,871,897]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="15" pageNumber="16" unit="in" value="2.0">2 in</quantity>
|
||
♁) inflated, with weakly chitinized outer margin bearing two parallel rows of flimsy setular extensions, possibly surrounding a glandular structure; a similar but non-homologous structure is found on the P4 endopod of species of
|
||
<taxonomicName authority="Huys & Lee, 2000" authorityName="Huys & Lee" authorityYear="2000" box="[401,761,976,1003]" class="Maxillopoda" family="Laophontidae" genus="Corbulaseta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[401,536,976,1002]" italics="true" pageId="15" pageNumber="16">Corbulaseta</emphasis>
|
||
<bibRefCitation author="Huys R & Lee W" box="[543,761,976,1003]" pageId="15" pageNumber="16" pagination="49 - 107" refId="ref18979" refString="Huys R, Lee W (2000) Basal resolution of laophontid phylogeny and the paraphyly of Esola Edwards. Bulletin of the Natural History Museum, London (Zoology) 66: 49 - 107." type="journal article" year="2000">Huys & Lee, 2000</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Huys R & Lee W" box="[777,1011,976,1003]" pageId="15" pageNumber="16" pagination="49 - 107" refId="ref18979" refString="Huys R, Lee W (2000) Basal resolution of laophontid phylogeny and the paraphyly of Esola Edwards. Bulletin of the Natural History Museum, London (Zoology) 66: 49 - 107." type="journal article" year="2000">Huys and Lee 2000</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Gomez S & Boyko CB" pageId="15" pageNumber="16" pagination="3 - 70" refId="ref18447" refString="Gomez S, Boyko CB (2006) On a small collection of harpacticoids from Easter Island: the family Laophontidae T. Scott (Crustacea: Copepoda: Harpacticoida). Zootaxa 1352: 3 - 70." type="journal article" year="2006">Gómez and Boyko 2006</bibRefCitation>
|
||
); (2) P4 with sexually dimorphic setation on enp-2 (outer distal seta reduced in
|
||
<specimenCount box="[327,342,1047,1073]" pageId="15" pageNumber="16" type="female">♀</specimenCount>
|
||
, very long in ♁); and (3) antennule
|
||
<specimenCount box="[788,803,1047,1073]" pageId="15" pageNumber="16" type="female">♀</specimenCount>
|
||
with spinules along posterior margins of segments 1–6.
|
||
</paragraph>
|
||
<paragraph blockId="15.[140,1108,167,1672]" lastBlockId="16.[140,1108,167,756]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
|
||
<bibRefCitation author="Fiers F" box="[185,388,1117,1143]" pageId="15" pageNumber="16" pagination="211 - 222" refId="ref18246" refString="Fiers F (1992) A redescription of Hemilaophonte janinae Jakubisiak (Copepoda, Harpacticoida), a laophontid living in the gill chambers of the common spider crab. Belgian Journal of Zoology 122: 211 - 222." type="journal article" year="1992">Fiers (1992: 213)</bibRefCitation>
|
||
hinted at a relationship between
|
||
<taxonomicName box="[786,1009,1117,1143]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[786,1009,1117,1143]" italics="true" pageId="15" pageNumber="16">Pseudonychocamptus</emphasis>
|
||
</taxonomicName>
|
||
and the monotypic genus
|
||
<taxonomicName authority="T. Scott, 1906" authorityName="T. Scott" authorityYear="1906" box="[341,627,1152,1179]" class="Maxillopoda" family="Laophontidae" genus="Harrietella" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[341,460,1152,1178]" italics="true" pageId="15" pageNumber="16">Harrietella</emphasis>
|
||
T.
|
||
<bibRefCitation author="Scott T" box="[495,627,1152,1178]" pageId="15" pageNumber="16" pagination="458 - 466" refId="ref20716" refString="Scott T (1906) Notes on British Copepoda: change of names. Annals and Magazine of Natural History 7 (17): 458 - 466." type="journal article" year="1906">Scott, 1906</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus box="[642,695,1152,1178]" pageId="15" pageNumber="16">type</typeStatus>
|
||
:
|
||
<taxonomicName authority="T. Scott, 1894 a" authorityName="T. Scott" authorityYear="1894" box="[705,1099,1152,1179]" class="Maxillopoda" family="Laophontidae" genus="Laophonte" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="simulans">
|
||
<emphasis box="[705,919,1152,1178]" italics="true" pageId="15" pageNumber="16">Laophonte simulans</emphasis>
|
||
T.
|
||
<bibRefCitation author="Scott T" box="[955,1099,1152,1178]" pageId="15" pageNumber="16" pagination="231 - 271" refId="ref20515" refString="Scott T (1894 a) Additions to the fauna of the Firth of Forth. Part VI. Reports of the Fishery Board for Scotland, Edinburgh 12 (3): 231 - 271." type="journal article" year="1894">Scott, 1894a</bibRefCitation>
|
||
</taxonomicName>
|
||
) but provided no supporting evidence for this claim. Presumably his assertion was adopted from
|
||
<bibRefCitation author="Lang K" box="[304,520,1223,1249]" pageId="15" pageNumber="16" refId="ref19237" refString="Lang K (1948) Monographie der Harpacticiden. Hakan Ohlsson, Lund, 1682 pp. (2 volumes)." type="book" year="1948">Lang (1948: 1449)</bibRefCitation>
|
||
who recognized a similar relationship based on the shared more or less dorsoventrally flattened body and the 2-segmented P3 endopod and 1-segmented P4 endopod in the male, in addition to the overall similarity in the maxilliped, P1 and male P3 exopod. Most
|
||
<taxonomicName box="[683,906,1328,1354]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[683,906,1328,1354]" italics="true" pageId="15" pageNumber="16">Pseudonychocamptus</emphasis>
|
||
</taxonomicName>
|
||
species, however, do not have a dorsoventrally depressed body and the general habitus is considerably different from that of
|
||
<taxonomicName box="[391,613,1399,1425]" class="Maxillopoda" family="Laophontidae" genus="Harrietella" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="simulans">
|
||
<emphasis box="[391,613,1399,1425]" italics="true" pageId="15" pageNumber="16">Harrietella simulans</emphasis>
|
||
</taxonomicName>
|
||
(T.
|
||
<bibRefCitation author="Scott T" box="[662,802,1399,1425]" pageId="15" pageNumber="16" pagination="231 - 271" refId="ref20515" refString="Scott T (1894 a) Additions to the fauna of the Firth of Forth. Part VI. Reports of the Fishery Board for Scotland, Edinburgh 12 (3): 231 - 271." type="journal article" year="1894">Scott 1894a</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Sars GO" box="[815,946,1399,1426]" pageId="15" pageNumber="16" pagination="73 - 92" refId="ref20467" refString="Sars GO (1920 b) Copepoda Supplement. Parts VII & VIII. Harpacticoida (continued). An account of the Crustacea of Norway, with short descriptions and figures of all the species 7: 73 - 92 + pls. 49 - 64." type="journal article" year="1920">Sars 1920b</bibRefCitation>
|
||
). In
|
||
<taxonomicName class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">Pseudonychocamptus</emphasis>
|
||
</taxonomicName>
|
||
the P4 endopod is suppressed during development in the male but not in the female, resulting in different segmentation and setation patterns, whereas in
|
||
<taxonomicName authorityName="T. Scott" authorityYear="1906" class="Maxillopoda" family="Laophontidae" genus="Harrietella" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">Harrietella</emphasis>
|
||
</taxonomicName>
|
||
both sexes display a 1-segmented endopod and no sexual dimorphism in setation is discernible (
|
||
<emphasis box="[357,393,1540,1565]" italics="true" pageId="15" pageNumber="16">e.g.</emphasis>
|
||
<bibRefCitation author="Vervoort W" box="[400,567,1539,1566]" pageId="15" pageNumber="16" pagination="297 - 305" refId="ref20772" refString="Vervoort W (1950) Harrietella simulans (T. Scott, 1894), a commensal copepod on Limnoria lignorum (Rathke). Zoologische Mededelingen Leiden 30: 297 - 305." type="journal article" year="1950">Vervoort 1950</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Hicks GRF" box="[580,716,1539,1566]" pageId="15" pageNumber="16" pagination="101 - 117" refId="ref18720" refString="Hicks GRF (1989) Harpacticoid copepods fom biogenic substrata in offshore waters of New Zealand. 2. Partial revisions of Dactylopodella Sars and Amphiascus Sars (varians - group) including new species, and a new record for Harrietella simulans (T. Scott). National Museum of New Zealand Records 3 (10): 101 - 117." type="journal article" year="1989">Hicks 1989</bibRefCitation>
|
||
). The 2-segmented P3 endopod in the male, displaying a straight spinous outgrowth on the distal segment, is indicative of a relationship with
|
||
<taxonomicName box="[395,618,1610,1636]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[395,618,1610,1636]" italics="true" pageId="15" pageNumber="16">Pseudonychocamptus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Noodt" authorityYear="1952" box="[634,713,1610,1636]" class="Maxillopoda" family="Laophontidae" genus="Pilifera" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[634,713,1610,1636]" italics="true" pageId="15" pageNumber="16">Pilifera</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Huys & Lee, 2009" authorityName="Huys & Lee" authorityYear="2009" box="[730,876,1610,1636]" class="Maxillopoda" family="Laophontidae" genus="Inermiphonte" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis box="[730,876,1610,1636]" italics="true" pageId="15" pageNumber="16">Inermiphonte</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[892,971,1610,1636]" pageId="15" pageNumber="16" rank="genus">gen. n.</taxonomicNameLabel>
|
||
(see below) and
|
||
<taxonomicName authority="Huys & Lee, 2009" authorityName="Huys & Lee" authorityYear="2009" box="[192,292,1645,1671]" class="Maxillopoda" family="Laophontidae" genus="Marbefia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis box="[192,292,1645,1671]" italics="true" pageId="15" pageNumber="16">Marbefia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[310,392,1646,1672]" pageId="15" pageNumber="16" rank="genus">gen. n.</taxonomicNameLabel>
|
||
, however,
|
||
<taxonomicName authorityName="T. Scott" authorityYear="1906" box="[517,636,1645,1671]" class="Maxillopoda" family="Laophontidae" genus="Harrietella" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[517,636,1645,1671]" italics="true" pageId="15" pageNumber="16">Harrietella</emphasis>
|
||
</taxonomicName>
|
||
does not display the sexual dimorphism (on P3–P4 enp-1) that defines Lee and Huys’ (1999)
|
||
<emphasis box="[782,862,167,192]" italics="true" pageId="16" pageNumber="17">PWPH</emphasis>
|
||
-clade. Although this absence is conceivably secondary as a result of the loss of the inner seta on P3 enp-
|
||
<quantity box="[1068,1108,202,228]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="16" pageNumber="17" unit="in" value="1.0">1 in</quantity>
|
||
the female, confirmation of its membership of this clade will probably have to await the arrival of molecular sequence data.
|
||
<taxonomicName authorityName="T. Scott" authorityYear="1906" box="[594,713,272,298]" class="Maxillopoda" family="Laophontidae" genus="Harrietella" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[594,713,272,298]" italics="true" pageId="16" pageNumber="17">Harrietella</emphasis>
|
||
</taxonomicName>
|
||
is here tentatively regarded as the sistergroup of
|
||
<taxonomicName authority="Huys & Lee, 2009" authorityName="Huys & Lee" authorityYear="2009" box="[301,447,307,333]" class="Maxillopoda" family="Laophontidae" genus="Inermiphonte" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis box="[301,447,307,333]" italics="true" pageId="16" pageNumber="17">Inermiphonte</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[462,539,308,334]" pageId="16" pageNumber="17" rank="genus">gen. n.</taxonomicNameLabel>
|
||
(see below).
|
||
</paragraph>
|
||
<paragraph blockId="16.[140,1108,167,756]" pageId="16" pageNumber="17">
|
||
Although the mosaic distribution of unique plesiomorphies in single species [e.g.
|
||
<taxonomicName authority="(Sars, 1905)" baseAuthorityName="Sars" baseAuthorityYear="1905" box="[140,580,378,405]" class="Maxillopoda" family="Laophontidae" genus="Heteronychocamptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="exiguus">
|
||
<emphasis box="[140,440,378,404]" italics="true" pageId="16" pageNumber="17">Heteronychocamptus exiguus</emphasis>
|
||
(
|
||
<bibRefCitation author="Sars GO" box="[454,572,378,405]" pageId="16" pageNumber="17" pagination="371 - 414" refId="ref20184" refString="Sars GO (1905) Pacifische Plankton-Crustaceen. (Ergebnisse einer Reise nach dem Pacific. Schauinsland 1896 - 1897). II. Brackwasser-Crustaceen von den Chatham-Inseln. Zoologische Jahrbucher fur Systematik 21 (4): 371 - 414." type="journal article" year="1905">Sars, 1905</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
with inner seta on P2 enp-1 (
|
||
<emphasis box="[912,937,378,404]" italics="true" pageId="16" pageNumber="17">cf.</emphasis>
|
||
<bibRefCitation author="Sars GO" box="[943,1055,378,405]" pageId="16" pageNumber="17" pagination="371 - 414" refId="ref20184" refString="Sars GO (1905) Pacifische Plankton-Crustaceen. (Ergebnisse einer Reise nach dem Pacific. Schauinsland 1896 - 1897). II. Brackwasser-Crustaceen von den Chatham-Inseln. Zoologische Jahrbucher fur Systematik 21 (4): 371 - 414." type="journal article" year="1905">Sars 1905</bibRefCitation>
|
||
: Fig. 125);
|
||
<taxonomicName authority="Hamond, 1969" authorityName="Hamond" authorityYear="1969" box="[207,611,413,439]" class="Maxillopoda" family="Laophontidae" genus="Laophonte" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="danversae">
|
||
<emphasis box="[207,428,413,439]" italics="true" pageId="16" pageNumber="17">Laophonte danversae</emphasis>
|
||
<bibRefCitation author="Hamond R" box="[437,611,413,439]" pageId="16" pageNumber="17" pagination="1 - 14" refId="ref18561" refString="Hamond R (1969) The Laophontidae (Copepoda, Harpacticoida) of the shore at West Runton, Norfolk, England. Crustaceana 16: 1 - 14." type="journal article" year="1969">Hamond, 1969</bibRefCitation>
|
||
</taxonomicName>
|
||
with 5 elements on ♁ P5 exopod (
|
||
<emphasis box="[1010,1035,413,439]" italics="true" pageId="16" pageNumber="17">cf.</emphasis>
|
||
<bibRefCitation author="Hicks GRF" pageId="16" pageNumber="17" pagination="302 - 307" refId="ref18684" refString="Hicks GRF (1982) New records of harpacticoid copepods from the east coast of Britain, and a description of the true male of Laophonte danversae Hamond. Crustaceana 42: 302 - 307." type="journal article" year="1982">Hicks 1982</bibRefCitation>
|
||
:
|
||
<figureCitation box="[211,274,448,475]" captionStart="Figure 6" captionStartId="10.[140,212,1585,1607]" captionTargetBox="[184,1099,180,1545]" captionTargetId="figure@10.[140,1108,165,1566]" captionTargetPageId="10" captionText="Figure 6. Marbefia carthyi (Hamond, 1968), comb. n.: A antennule ♁, ventral [armature on segments 3, 4 and 8 omitted] B antennulary segments 3 (right) and 4 (left) ♁ C antennulary segment 8 ♁ D P5 and P6 ♁, anterior E P5 ♀, anterior." figureDoi="http://doi.org/10.5281/zenodo.3767622" httpUri="https://zenodo.org/record/3767622/files/figure.png" pageId="16" pageNumber="17">Fig. 6</figureCitation>
|
||
);
|
||
<taxonomicName authorityName="Hamond" authorityYear="1968" box="[295,391,448,474]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="carthyi">
|
||
<emphasis box="[295,391,448,474]" italics="true" pageId="16" pageNumber="17">P. carthyi</emphasis>
|
||
</taxonomicName>
|
||
with 3 inner setae on
|
||
<specimenCount box="[633,648,449,475]" pageId="16" pageNumber="17" type="female">♀</specimenCount>
|
||
P3 enp-2 (
|
||
<figureCitation box="[775,889,448,475]" captionStart="Figure 5" captionStartId="9.[140,212,1585,1607]" captionTargetBox="[152,1104,167,1554]" captionTargetId="figure@9.[140,1108,165,1566]" captionTargetPageId="9" captionText="Figure 5. Marbefia carthyi (Hamond, 1968), comb. n.: A P2 ♀, anterior, B P2 endopod ♀, outer lateral C P3 ♀, anterior [inner seta on enp-1 arrowed] D P3 endopod ♁ [spinules on enp-1 and apophysis on enp-2 arrowed]." figureDoi="http://doi.org/10.5281/zenodo.3767620" httpUri="https://zenodo.org/record/3767620/files/figure.png" pageId="16" pageNumber="17">Figure 5C</figureCitation>
|
||
)] hampers the analysis of the basal relationships within the
|
||
<emphasis box="[578,657,484,509]" italics="true" pageId="16" pageNumber="17">PWPH</emphasis>
|
||
-clade, a well defined group of five genera can be distinguished based on the sexual dimorphism of the P3 endopod. In males of
|
||
<taxonomicName authority="Huys & Lee, 2009" authorityName="Huys & Lee" authorityYear="2009" box="[140,238,554,580]" class="Maxillopoda" family="Laophontidae" genus="Marbefia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis box="[140,238,554,580]" italics="true" pageId="16" pageNumber="17">Marbefia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[252,328,554,580]" pageId="16" pageNumber="17" rank="genus">gen. n.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName box="[341,558,554,580]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[341,558,554,580]" italics="true" pageId="16" pageNumber="17">Pseudonychocamptus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Noodt" authorityYear="1952" box="[572,649,554,580]" class="Maxillopoda" family="Laophontidae" genus="Pilifera" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[572,649,554,580]" italics="true" pageId="16" pageNumber="17">Pilifera</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Huys & Lee, 2009" authorityName="Huys & Lee" authorityYear="2009" box="[663,806,554,580]" class="Maxillopoda" family="Laophontidae" genus="Inermiphonte" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis box="[663,806,554,580]" italics="true" pageId="16" pageNumber="17">Inermiphonte</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[820,895,554,580]" pageId="16" pageNumber="17" rank="genus">gen. n.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName authorityName="T. Scott" authorityYear="1906" box="[950,1066,554,580]" class="Maxillopoda" family="Laophontidae" genus="Harrietella" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[950,1066,554,580]" italics="true" pageId="16" pageNumber="17">Harrietella</emphasis>
|
||
</taxonomicName>
|
||
the ramus is 2-segmented (instead of 3-segmented), bearing a straight (rather than a sigmoid) apophysis on the outer margin of the distal endopod segment (Table 1). Within this lineage,
|
||
<taxonomicName authorityName="Huys & Lee" authorityYear="2009" box="[189,287,659,685]" class="Maxillopoda" family="Laophontidae" genus="Marbefia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[189,287,659,685]" italics="true" pageId="16" pageNumber="17">Marbefia</emphasis>
|
||
</taxonomicName>
|
||
is considered to be the basal offshoot, the other four genera sharing a suite of synapomorphic reductions on the swimming legs: (1) P3–P4 exp-3 with 1 inner seta, (2) P2 enp-1 with 1 inner seta, and (3) P3 enp-
|
||
<specimenCount box="[663,698,730,756]" pageId="16" pageNumber="17" type="female">2 ♀</specimenCount>
|
||
with at most 2 inner setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |