377 lines
55 KiB
XML
377 lines
55 KiB
XML
<document id="44BD5FF29CDB6C5C5E4BD0FA842154DA" ID-CLB-Dataset="7720" ID-DOI="10.11646/zootaxa.4938.5.6" ID-GBIF-Dataset="71162ab5-a8df-4e4d-931e-73e5cb92dbff" ID-ISSN="1175-5326" ID-Zenodo-Dep="4575108" ID-ZooBank="773ED5ED-D56C-4A40-97D0-AEF05EB39CA3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614755784177" checkinUser="plazi" docAuthor="Dimitrov, Dragomir" docDate="2021" docId="770E87D3FFE9FFF0FF3F0B9CFF461E25" docLanguage="en" docName="zootaxa.4938.5.6.pdf" docOrigin="Zootaxa 4938 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dysdera zonsteini Dimitrov 2021, new species" docType="treatment" docVersion="7" lastPageNumber="593" masterDocId="8B37FFABFFE8FFF5FFA80912FFB61A10" masterDocTitle="Description of Dysdera zonsteini n. sp. (Arachnida: Araneae: Dysderidae) from Turkmenistan" masterLastPageNumber="594" masterPageNumber="588" pageNumber="589" updateTime="1698931949484" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FA2CE03E6296ECB8BFE6683591F76C9E">Description of Dysdera zonsteini n. sp. (Arachnida: Araneae: Dysderidae) from Turkmenistan</mods:title>
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<mods:name id="BBBAD1EC2DDB22753B7F46B423F6E717" type="personal">
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<mods:namePart id="5172FC32AA8E02624195AFE2C36FF361">Dimitrov, Dragomir</mods:namePart>
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<mods:title id="E8E506D8955FE178B1196328930C6B2C">Zootaxa</mods:title>
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<mods:part id="23158384A0960166B87D22D20AD8C8F0">
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<mods:date id="B70AB79D1A7D6A7B8F3829ED42E00837">2021</mods:date>
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<mods:number id="70F4FEB8F5AC691084ADC3E0E16DB997">2021-03-03</mods:number>
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<mods:number id="2A309E7D6A411409633CA68AE3E9E998">4938</mods:number>
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<treatment id="770E87D3FFE9FFF0FF3F0B9CFF461E25" ID-DOI="http://doi.org/10.5281/zenodo.4608016" ID-GBIF-Taxon="180584217" ID-Zenodo-Dep="4608016" LSID="urn:lsid:plazi:treatment:770E87D3FFE9FFF0FF3F0B9CFF461E25" httpUri="http://treatment.plazi.org/id/770E87D3FFE9FFF0FF3F0B9CFF461E25" lastPageId="5" lastPageNumber="593" pageId="1" pageNumber="589">
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<subSubSection id="B7BD654EFFE9FFF4FF3F0B9CFE4218B8" box="[151,500,654,681]" pageId="1" pageNumber="589" type="nomenclature">
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<paragraph id="FF1836C5FFE9FFF4FF3F0B9CFE4218B8" blockId="1.[151,500,654,717]" box="[151,500,654,681]" pageId="1" pageNumber="589">
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<heading id="A45081A9FFE9FFF4FF3F0B9CFE4218B8" bold="true" box="[151,500,654,681]" fontSize="11" level="1" pageId="1" pageNumber="589" reason="1">
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<emphasis id="CDD3EAD7FFE9FFF4FF3F0B9CFE4218B8" bold="true" box="[151,500,654,681]" pageId="1" pageNumber="589">
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<taxonomicName id="38A74D46FFE9FFF4FF3F0B9CFED218B9" ID-CoL="98Z24" authority="Dimitrov, 2021" authorityName="Dimitrov" authorityYear="2021" box="[151,356,654,681]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="589" phylum="Arthropoda" rank="species" species="zonsteini" status="sp. nov.">
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<emphasis id="CDD3EAD7FFE9FFF4FF3F0B9CFED218B9" bold="true" box="[151,356,654,681]" italics="true" pageId="1" pageNumber="589">Dysdera zonsteini</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D6E057ACFFE9FFF4FEC30B9DFE4218B8" box="[363,500,654,681]" pageId="1" pageNumber="589" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B7BD654EFFE9FFF4FF3F0BA1FEB118DD" box="[151,263,691,717]" pageId="1" pageNumber="589" type="description">
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<paragraph id="FF1836C5FFE9FFF4FF3F0BA1FEB118DD" blockId="1.[151,500,654,717]" box="[151,263,691,717]" pageId="1" pageNumber="589">
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<figureCitation id="679C2A40FFE9FFF4FF3F0BA1FEB118DD" box="[151,263,691,717]" captionStart-0="FIGURES 1–6" captionStart-1="FIGURES 7–13" captionStartId-0="2.[151,264,1974,1998]" captionStartId-1="3.[151,264,1626,1650]" captionTargetBox-0="[357,1229,181,1950]" captionTargetBox-1="[151,1436,181,1603]" captionTargetId-0="figure-16@2.[357,1229,181,1950]" captionTargetId-1="figure-34@3.[151,1436,181,1603]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1–6. Habitus of D. zonsteini n. sp., 1–2, 5 holotype male, 3–4, 6 paratype female. 1, 3 body dorsal. 2, 4 body ventral. 5–6 chelicerae." captionText-1="FIGURES 7–13. D. zonsteini n. sp., 7–11 holotype male, left palp. 7 prolateral, 8 dorsal, 9 frontal, 10 retrolateral, 11 apical. 12–13 paratype female, vulva. 12 ventral, 13 dorsal." figureDoi-0="http://doi.org/10.5281/zenodo.4575110" figureDoi-1="http://doi.org/10.5281/zenodo.4575112" httpUri-0="https://zenodo.org/record/4575110/files/figure.png" httpUri-1="https://zenodo.org/record/4575112/files/figure.png" pageId="1" pageNumber="589">Figs 1–13</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="B7BD654EFFE9FFF4FF3F0BE9FE54194D" pageId="1" pageNumber="589" type="materials_examined">
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<paragraph id="FF1836C5FFE9FFF4FF3F0BE9FE54194D" blockId="1.[151,1437,762,1977]" pageId="1" pageNumber="589">
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<emphasis id="CDD3EAD7FFE9FFF4FF3F0BE9FEFE1904" bold="true" box="[151,328,762,789]" pageId="1" pageNumber="589">Type material.</emphasis>
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<materialsCitation id="4FCF3C98FFE9FFF4FEFA0BE9FBB01929" ID-GBIF-Occurrence="3053520302" collectingDate="1985-04-21" collectionCode="MNHN" collectorName="Sergei Zonstein" country="Turkmenistan" elevation="400" latitude="38.466667" location="South-Western Kopet Dagh" longLatPrecision="1175" longitude="56.25" municipality="Kara-Kala Town" pageId="1" pageNumber="589" specimenCode="AR 5862" specimenCount="1" typeStatus="holotype">
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<typeStatus id="201C8867FFE9FFF4FEFA0BE9FE081905" box="[338,446,763,789]" pageId="1" pageNumber="589" type="holotype">Holotype</typeStatus>
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. ³.
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<collectingCountry id="87B07655FFE9FFF4FE580BE9FD381905" box="[496,654,763,789]" name="Turkmenistan" pageId="1" pageNumber="589">Turkmenistan</collectingCountry>
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,
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<location id="FA78601EFFE9FFF4FD340BE9FC6F1905" LSID="urn:lsid:plazi:treatment:770E87D3FFE9FFF0FF3F0B9CFF461E25:FA78601EFFE9FFF4FD340BE9FC6F1905" box="[668,985,763,789]" country="Turkmenistan" latitude="38.466667" longLatPrecision="1175" longitude="56.25" municipality="Kara-Kala Town" name="South-Western Kopet Dagh" pageId="1" pageNumber="589">South-Western Kopet Dagh</location>
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, near
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<collectingMunicipality id="1F7CACBFFFE9FFF4FB890BE9FB571905" box="[1057,1249,763,789]" pageId="1" pageNumber="589">Kara-Kala Town</collectingMunicipality>
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, GPS:
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<geoCoordinate id="9A935002FFE9FFF4FA9C0BE9FA211905" box="[1332,1431,763,789]" degrees="38" direction="north" minutes="28" orientation="latitude" pageId="1" pageNumber="589" precision="925" value="38.466667">38°28’N</geoCoordinate>
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,
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<geoCoordinate id="9A935002FFE9FFF4FF3F0A0CFF411929" box="[151,247,798,825]" degrees="56" direction="east" minutes="15" orientation="longitude" pageId="1" pageNumber="589" precision="925" value="56.25">56°15’E</geoCoordinate>
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, alt.
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<quantity id="385F9B20FFE9FFF4FE840A0CFE1A1929" box="[300,428,798,825]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="1" pageNumber="589" unit="m" value="400.0" valueMax="450.0" valueMin="350.0">
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<elevation id="748AD1F6FFE9FFF4FE840A0CFE1A1929" box="[300,428,798,825]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="1" pageNumber="589" unit="m" value="400.0" valueMax="450.0" valueMin="350.0">350–450 m</elevation>
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</quantity>
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,
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<date id="8B191005FFE9FFF4FE100A0CFD8E1929" box="[440,568,798,825]" pageId="1" pageNumber="589" value="1985-04-21">
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<collectingDate id="9B5DE9EDFFE9FFF4FE100A0CFD8E1929" box="[440,568,798,825]" pageId="1" pageNumber="589" value="1985-04-21">21.04.1985</collectingDate>
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</date>
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,
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<collectorName id="52525313FFE9FFF4FDEB0A0DFD451929" box="[579,755,799,825]" pageId="1" pageNumber="589">Sergei Zonstein</collectorName>
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leg. (
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<collectionCode id="99B6AE00FFE9FFF4FC990A0DFC3D1929" box="[817,907,799,825]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="589">MNHN</collectionCode>
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,
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<specimenCode id="AF019EBEFFE9FFF4FC3C0A0DFC481929" box="[916,1022,798,825]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="589">AR 5862</specimenCode>
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)
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</materialsCitation>
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.
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||
<materialsCitation id="4FCF3C98FFE9FFF4FBB80A0DFE68194D" ID-GBIF-Occurrence="3053520301" collectingDate="1985-04-21" collectionCode="MNHN" collectorName="Sergei Zonstein" country="Turkmenistan" elevation="400" latitude="38.466667" location="South-Western Kopet Dagh" longLatPrecision="1175" longitude="56.25" municipality="Kara-Kala Town" pageId="1" pageNumber="589" specimenCode="AR 5862" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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<typeStatus id="201C8867FFE9FFF4FBB80A0DFBC01929" box="[1040,1142,799,825]" pageId="1" pageNumber="589" type="paratype">Paratype</typeStatus>
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.
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<specimenCount id="E9A1FD4CFFE9FFF4FB280A0DFB151929" box="[1152,1187,799,825]" pageId="1" pageNumber="589" type="female">1♀</specimenCount>
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, with same data as for holotype (
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||
<collectionCode id="99B6AE00FFE9FFF4FEA00A51FED4194D" box="[264,354,835,861]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="589">MNHN</collectionCode>
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,
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<specimenCode id="AF019EBEFFE9FFF4FEC40A51FE60194D" box="[364,470,834,861]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="589">AR 5862</specimenCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="B7BD654EFFE9FFF4FF6F0A75FACF1991" box="[199,1401,871,897]" pageId="1" pageNumber="589" type="etymology">
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<paragraph id="FF1836C5FFE9FFF4FF6F0A75FACF1991" blockId="1.[151,1437,762,1977]" box="[199,1401,871,897]" pageId="1" pageNumber="589">
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<emphasis id="CDD3EAD7FFE9FFF4FF6F0A75FEF81991" bold="true" box="[199,334,871,897]" pageId="1" pageNumber="589">Etymology.</emphasis>
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Named after the well-known arachnologist Sergei Zonstein, who collected the
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<typeStatus id="201C8867FFE9FFF4FB6D0A75FB431991" box="[1221,1269,871,897]" pageId="1" pageNumber="589">type</typeStatus>
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specimens.
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</paragraph>
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</subSubSection>
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<subSubSection id="B7BD654EFFE9FFF4FF6F0A98FDD81EFA" pageId="1" pageNumber="589" type="diagnosis">
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<paragraph id="FF1836C5FFE9FFF4FF6F0A98FDD81EFA" blockId="1.[151,1437,762,1977]" pageId="1" pageNumber="589">
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<emphasis id="CDD3EAD7FFE9FFF4FF6F0A98FEF419B4" bold="true" box="[199,322,906,932]" pageId="1" pageNumber="589">Diagnosis.</emphasis>
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Both male and female of
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<taxonomicName id="38A74D46FFE9FFF4FDCB0A99FC9A19B4" authority="Dimitrov, 2021" authorityName="Dimitrov" authorityYear="2021" box="[611,812,907,933]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="589" phylum="Arthropoda" rank="species" species="zonsteini" status="sp. nov.">
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<emphasis id="CDD3EAD7FFE9FFF4FDCB0A99FC9A19B4" box="[611,812,907,933]" italics="true" pageId="1" pageNumber="589">Dysdera zonsteini</emphasis>
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||
</taxonomicName>
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||
<emphasis id="CDD3EAD7FFE9FFF4FC9B0A99FCC719B5" bold="true" box="[819,881,907,933]" pageId="1" pageNumber="589">
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||
<taxonomicNameLabel id="D6E057ACFFE9FFF4FC9B0A99FCC719B5" box="[819,881,907,933]" pageId="1" pageNumber="589" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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are morphologically close to
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<taxonomicName id="38A74D46FFE9FFF4FB160A99FE9319D9" authority="Dunin, 1992" authorityName="Dunin" authorityYear="1992" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="589" phylum="Arthropoda" rank="species" species="kronebergi">
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<emphasis id="CDD3EAD7FFE9FFF4FB160A99FA2A19B5" box="[1214,1436,907,933]" italics="true" pageId="1" pageNumber="589">Dysdera kronebergi</emphasis>
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Dunin, 1992
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</taxonomicName>
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. The two species can be separated by the following diagnostic characters: (1) the smaller size of
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<taxonomicName id="38A74D46FFE9FFF4FAE30ABDFF6219FC" authority="Dimitrov, 2021" authorityName="Dimitrov" authorityYear="2021" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="589" phylum="Arthropoda" rank="species" species="zonsteini" status="sp. nov.">
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<emphasis id="CDD3EAD7FFE9FFF4FAE30ABDFF6219FC" italics="true" pageId="1" pageNumber="589">D. zonsteini</emphasis>
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</taxonomicName>
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||
<emphasis id="CDD3EAD7FFE9FFF4FF730AC1FEAC19FD" bold="true" box="[219,282,979,1005]" pageId="1" pageNumber="589">
|
||
<taxonomicNameLabel id="D6E057ACFFE9FFF4FF730AC1FEAC19FD" box="[219,282,979,1005]" pageId="1" pageNumber="589" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
[carapace length 1.65 vs.
|
||
<quantity id="385F9B20FFE9FFF4FDE80AC0FD3819FD" box="[576,654,978,1005]" metricMagnitude="-2" metricUnit="m" metricValue="6.223" pageId="1" pageNumber="589" unit="in" value="2.45">2.45 in</quantity>
|
||
<taxonomicName id="38A74D46FFE9FFF4FD3D0AC1FC8619FD" authorityName="Dunin" authorityYear="1992" box="[661,816,979,1005]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="589" phylum="Arthropoda" rank="species" species="kronebergi">
|
||
<emphasis id="CDD3EAD7FFE9FFF4FD3D0AC1FC8619FD" box="[661,816,979,1005]" italics="true" pageId="1" pageNumber="589">D. kronebergi</emphasis>
|
||
</taxonomicName>
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||
]. Male of the new species differs also by (2) the strong process (PE) of the embolic division (
|
||
<figureCitation id="679C2A40FFE9FFF4FDE90AE5FD111E01" box="[577,679,1015,1041]" captionStart="FIGURES 7–13" captionStartId="3.[151,264,1626,1650]" captionTargetBox="[151,1436,181,1603]" captionTargetId="figure-34@3.[151,1436,181,1603]" captionTargetPageId="3" captionText="FIGURES 7–13. D. zonsteini n. sp., 7–11 holotype male, left palp. 7 prolateral, 8 dorsal, 9 frontal, 10 retrolateral, 11 apical. 12–13 paratype female, vulva. 12 ventral, 13 dorsal." figureDoi="http://doi.org/10.5281/zenodo.4575112" httpUri="https://zenodo.org/record/4575112/files/figure.png" pageId="1" pageNumber="589">Figs 7–9</figureCitation>
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||
,
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||
<figureCitation id="679C2A40FFE9FFF4FD1B0AE5FD661E01" box="[691,720,1015,1041]" captionStart="FIGURES 14–17" captionStartId="4.[151,264,1919,1943]" captionTargetBox="[157,1436,181,1889]" captionTargetId="figure-16@4.[151,1436,181,1895]" captionTargetPageId="4" captionText="FIGURES 14–17. D. zonsteini n. sp., 14-15 holotype male, left palp. 14 prolateral, 15 retrolateral. 16-17 paratype female, vulva. 16 ventral, 17 dorsal." figureDoi="http://doi.org/10.5281/zenodo.4575114" httpUri="https://zenodo.org/record/4575114/files/figure.png" pageId="1" pageNumber="589">14</figureCitation>
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) that is not seen in
|
||
<taxonomicName id="38A74D46FFE9FFF4FC050AE5FB821E00" authorityName="Dimitrov" authorityYear="2021" box="[941,1076,1015,1040]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="589" phylum="Arthropoda" rank="species" species="zonsteini">
|
||
<emphasis id="CDD3EAD7FFE9FFF4FC050AE5FB821E00" box="[941,1076,1015,1040]" italics="true" pageId="1" pageNumber="589">D. zonsteini</emphasis>
|
||
</taxonomicName>
|
||
(Dunin 1992, figs. 4–5) and (3) the overall shape of the tegulum and the embolic division of the bulb (
|
||
<figureCitation id="679C2A40FFE9FFF4FC020D09FBAB1E25" box="[938,1053,1051,1077]" captionStart="FIGURES 7–13" captionStartId="3.[151,264,1626,1650]" captionTargetBox="[151,1436,181,1603]" captionTargetId="figure-34@3.[151,1436,181,1603]" captionTargetPageId="3" captionText="FIGURES 7–13. D. zonsteini n. sp., 7–11 holotype male, left palp. 7 prolateral, 8 dorsal, 9 frontal, 10 retrolateral, 11 apical. 12–13 paratype female, vulva. 12 ventral, 13 dorsal." figureDoi="http://doi.org/10.5281/zenodo.4575112" httpUri="https://zenodo.org/record/4575112/files/figure.png" pageId="1" pageNumber="589">Figs 7–10</figureCitation>
|
||
,
|
||
<figureCitation id="679C2A40FFE9FFF4FB810D08FBC51E25" box="[1065,1139,1050,1077]" captionStart="FIGURES 14–17" captionStartId="4.[151,264,1919,1943]" captionTargetBox="[157,1436,181,1889]" captionTargetId="figure-16@4.[151,1436,181,1895]" captionTargetPageId="4" captionText="FIGURES 14–17. D. zonsteini n. sp., 14-15 holotype male, left palp. 14 prolateral, 15 retrolateral. 16-17 paratype female, vulva. 16 ventral, 17 dorsal." figureDoi="http://doi.org/10.5281/zenodo.4575114" httpUri="https://zenodo.org/record/4575114/files/figure.png" pageId="1" pageNumber="589">14–15</figureCitation>
|
||
). Female differs by (4) the thinner and straight transversal bar (
|
||
<figureCitation id="679C2A40FFE9FFF4FD860D2DFD061E49" box="[558,688,1087,1113]" captionStart="FIGURES 7–13" captionStartId="3.[151,264,1626,1650]" captionTargetBox="[151,1436,181,1603]" captionTargetId="figure-34@3.[151,1436,181,1603]" captionTargetPageId="3" captionText="FIGURES 7–13. D. zonsteini n. sp., 7–11 holotype male, left palp. 7 prolateral, 8 dorsal, 9 frontal, 10 retrolateral, 11 apical. 12–13 paratype female, vulva. 12 ventral, 13 dorsal." figureDoi="http://doi.org/10.5281/zenodo.4575112" httpUri="https://zenodo.org/record/4575112/files/figure.png" pageId="1" pageNumber="589">Figs 12–13</figureCitation>
|
||
,
|
||
<figureCitation id="679C2A40FFE9FFF4FD140D2DFCB51E4A" box="[700,771,1087,1114]" captionStart="FIGURES 14–17" captionStartId="4.[151,264,1919,1943]" captionTargetBox="[157,1436,181,1889]" captionTargetId="figure-16@4.[151,1436,181,1895]" captionTargetPageId="4" captionText="FIGURES 14–17. D. zonsteini n. sp., 14-15 holotype male, left palp. 14 prolateral, 15 retrolateral. 16-17 paratype female, vulva. 16 ventral, 17 dorsal." figureDoi="http://doi.org/10.5281/zenodo.4575114" httpUri="https://zenodo.org/record/4575114/files/figure.png" pageId="1" pageNumber="589">16–17</figureCitation>
|
||
) vs. more massive and with curved edges in
|
||
<taxonomicName id="38A74D46FFE9FFF4FAA90D2DFA2A1E49" authorityName="Dunin" authorityYear="1992" box="[1281,1436,1087,1113]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="589" phylum="Arthropoda" rank="species" species="kronebergi">
|
||
<emphasis id="CDD3EAD7FFE9FFF4FAA90D2DFA2A1E49" box="[1281,1436,1087,1113]" italics="true" pageId="1" pageNumber="589">D. kronebergi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="9B364B34FFE9FFF4FF370D71FE821E6D" author="Dunin, P. M." box="[159,308,1123,1149]" pageId="1" pageNumber="589" pagination="35 - 76" refId="ref2707" refString="Dunin, P. M. (1992 a) The spider family Dysderidae of the Caucasian fauna (Arachnida Aranei Haplogynae). Arthropoda Selecta, 1 (3), 35 - 76." type="journal article" year="1992">Dunin 1992a</bibRefCitation>
|
||
, fig 6) and by (5) the more oval shape of the dorsal arc of the anterior diverticulum. The spermathecae of the two species are almost identical, but (6) in
|
||
<taxonomicName id="38A74D46FFE9FFF4FD1A0D95FCFA1EB1" authorityName="Dunin" authorityYear="1992" box="[690,844,1159,1185]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="589" phylum="Arthropoda" rank="species" species="kronebergi">
|
||
<emphasis id="CDD3EAD7FFE9FFF4FD1A0D95FCFA1EB1" box="[690,844,1159,1185]" italics="true" pageId="1" pageNumber="589">D. kronebergi</emphasis>
|
||
</taxonomicName>
|
||
it is wider than the dorsal arc of anterior diverticulum (
|
||
<bibRefCitation id="9B364B34FFE9FFF4FF370DB9FE831ED5" author="Dunin, P. M." box="[159,309,1195,1221]" pageId="1" pageNumber="589" pagination="35 - 76" refId="ref2707" refString="Dunin, P. M. (1992 a) The spider family Dysderidae of the Caucasian fauna (Arachnida Aranei Haplogynae). Arthropoda Selecta, 1 (3), 35 - 76." type="journal article" year="1992">Dunin 1992a</bibRefCitation>
|
||
, fig 6), while in
|
||
<taxonomicName id="38A74D46FFE9FFF4FE470DB9FDC31ED4" authority="Dimitrov, 2021" authorityName="Dimitrov" authorityYear="2021" box="[495,629,1195,1220]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="589" phylum="Arthropoda" rank="species" species="zonsteini" status="sp. nov.">
|
||
<emphasis id="CDD3EAD7FFE9FFF4FE470DB9FDC31ED4" box="[495,629,1195,1220]" italics="true" pageId="1" pageNumber="589">D. zonsteini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CDD3EAD7FFE9FFF4FDD40DB9FD0D1ED5" bold="true" box="[636,699,1195,1221]" pageId="1" pageNumber="589">
|
||
<taxonomicNameLabel id="D6E057ACFFE9FFF4FDD40DB9FD0D1ED5" box="[636,699,1195,1221]" pageId="1" pageNumber="589" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
it is narrower and its lateral wings do not exceed the width of the dorsal arc in its base (
|
||
<figureCitation id="679C2A40FFE9FFF4FE230DDDFDBA1EF9" box="[395,524,1231,1257]" captionStart="FIGURES 7–13" captionStartId="3.[151,264,1626,1650]" captionTargetBox="[151,1436,181,1603]" captionTargetId="figure-34@3.[151,1436,181,1603]" captionTargetPageId="3" captionText="FIGURES 7–13. D. zonsteini n. sp., 7–11 holotype male, left palp. 7 prolateral, 8 dorsal, 9 frontal, 10 retrolateral, 11 apical. 12–13 paratype female, vulva. 12 ventral, 13 dorsal." figureDoi="http://doi.org/10.5281/zenodo.4575112" httpUri="https://zenodo.org/record/4575112/files/figure.png" pageId="1" pageNumber="589">Figs 12–13</figureCitation>
|
||
,
|
||
<figureCitation id="679C2A40FFE9FFF4FDBF0DDDFDD71EFA" box="[535,609,1231,1258]" captionStart="FIGURES 14–17" captionStartId="4.[151,264,1919,1943]" captionTargetBox="[157,1436,181,1889]" captionTargetId="figure-16@4.[151,1436,181,1895]" captionTargetPageId="4" captionText="FIGURES 14–17. D. zonsteini n. sp., 14-15 holotype male, left palp. 14 prolateral, 15 retrolateral. 16-17 paratype female, vulva. 16 ventral, 17 dorsal." figureDoi="http://doi.org/10.5281/zenodo.4575114" httpUri="https://zenodo.org/record/4575114/files/figure.png" pageId="1" pageNumber="589">16–17</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B7BD654EFFE9FFF0FF6F0DE0FE671B0D" lastPageId="5" lastPageNumber="593" pageId="1" pageNumber="589" type="description">
|
||
<paragraph id="FF1836C5FFE9FFF4FF6F0DE0FCBF1CF1" blockId="1.[151,1437,762,1977]" pageId="1" pageNumber="589">
|
||
<emphasis id="CDD3EAD7FFE9FFF4FF6F0DE0FEEF1F1C" bold="true" box="[199,345,1266,1292]" pageId="1" pageNumber="589">Description.</emphasis>
|
||
Male. (
|
||
<figureCitation id="679C2A40FFE9FFF4FE190DE1FD851F1D" box="[433,563,1266,1293]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1974,1998]" captionTargetBox="[357,1229,181,1950]" captionTargetId="figure-16@2.[357,1229,181,1950]" captionTargetPageId="2" captionText="FIGURES 1–6. Habitus of D. zonsteini n. sp., 1–2, 5 holotype male, 3–4, 6 paratype female. 1, 3 body dorsal. 2, 4 body ventral. 5–6 chelicerae." figureDoi="http://doi.org/10.5281/zenodo.4575110" httpUri="https://zenodo.org/record/4575110/files/figure.png" pageId="1" pageNumber="589">Figs 1–2, 5</figureCitation>
|
||
,
|
||
<figureCitation id="679C2A40FFE9FFF4FDE80DE1FDCF1F1D" box="[576,633,1267,1293]" captionStart="FIGURES 7–13" captionStartId="3.[151,264,1626,1650]" captionTargetBox="[151,1436,181,1603]" captionTargetId="figure-34@3.[151,1436,181,1603]" captionTargetPageId="3" captionText="FIGURES 7–13. D. zonsteini n. sp., 7–11 holotype male, left palp. 7 prolateral, 8 dorsal, 9 frontal, 10 retrolateral, 11 apical. 12–13 paratype female, vulva. 12 ventral, 13 dorsal." figureDoi="http://doi.org/10.5281/zenodo.4575112" httpUri="https://zenodo.org/record/4575112/files/figure.png" pageId="1" pageNumber="589">7–11</figureCitation>
|
||
,
|
||
<figureCitation id="679C2A40FFE9FFF4FD2E0DE0FD661F1D" box="[646,720,1266,1293]" captionStart="FIGURES 14–17" captionStartId="4.[151,264,1919,1943]" captionTargetBox="[157,1436,181,1889]" captionTargetId="figure-16@4.[151,1436,181,1895]" captionTargetPageId="4" captionText="FIGURES 14–17. D. zonsteini n. sp., 14-15 holotype male, left palp. 14 prolateral, 15 retrolateral. 16-17 paratype female, vulva. 16 ventral, 17 dorsal." figureDoi="http://doi.org/10.5281/zenodo.4575114" httpUri="https://zenodo.org/record/4575114/files/figure.png" pageId="1" pageNumber="589">14–15</figureCitation>
|
||
). Measurements. Total length 4.05; carapace length 1.65, width 1.18; sternum length 1.05, width 0.75; cheliceral length 0.72, width 0.25; clypeus height 0.05; labium length 0.40; gnathocoxa length 0.58; abdomen length 1.95, width 0.85; Eye diameters: AE—0.17, PE—0.14. Leg measurements. I—5.53 (0.92, 1.33, 0.77, 0.97, 1.04, 0.50); II—4.86 (0.83, 1.26, 0.43, 0.93, 0.94, 0.47); III—3.61 (0.65, 0.97, 0.43, 0.55, 0.65, 0.36); IV—4.66 (0.72, 1.13, 0.58, 0.86, 0.97, 0.40). Leg spination: coxae, trochanters and patellae spineless. Legs I and II have only one distal prolateral femoral spine, legs III and IV with well-armed femur and metatarsus. I—1pl, 0, 0; II—1pl, 0, 0; III—1d, 5pl 5rl, 5pl 2rl; IV—4d, 6pl 6rl, 6pl 5rl. Colouration very pale. Carapace, chelicerae and sternum light orange. Sternum a little lighter than the carapace. Abdomen and legs white. Other somatic characters. Body long and slim. Chelicerae dorsally covered with small tubercles. The distance between the AE equal to their diameter. Posterior eye-row very slightly procurved, almost straight. PE touching to each other. PLE touching to AE. Chelicerae with 3 teeth of approximately the same size, arranged in one row. The proximal one positioned near the base of the cheliceral groove, the medial one is close to it and the distal one is approximately at the middle of the groove (
|
||
<figureCitation id="679C2A40FFE9FFF4FDC00FB1FD1B1CAD" box="[616,685,1698,1725]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1974,1998]" captionTargetBox="[357,1229,181,1950]" captionTargetId="figure-16@2.[357,1229,181,1950]" captionTargetPageId="2" captionText="FIGURES 1–6. Habitus of D. zonsteini n. sp., 1–2, 5 holotype male, 3–4, 6 paratype female. 1, 3 body dorsal. 2, 4 body ventral. 5–6 chelicerae." figureDoi="http://doi.org/10.5281/zenodo.4575110" httpUri="https://zenodo.org/record/4575110/files/figure.png" pageId="1" pageNumber="589">Fig. 5</figureCitation>
|
||
). Labium pyramidal, notched distally. Tarsi with 2 claws and claw tuffs. Anterior border of sternum wider than the labium.
|
||
</paragraph>
|
||
<paragraph id="FF1836C5FFE9FFF4FF6F0FF9FEF71DA9" blockId="1.[151,1437,762,1977]" pageId="1" pageNumber="589">
|
||
Palp. (
|
||
<figureCitation id="679C2A40FFE9FFF4FEBB0FF9FE3F1D15" box="[275,393,1771,1797]" captionStart="FIGURES 7–13" captionStartId="3.[151,264,1626,1650]" captionTargetBox="[151,1436,181,1603]" captionTargetId="figure-34@3.[151,1436,181,1603]" captionTargetPageId="3" captionText="FIGURES 7–13. D. zonsteini n. sp., 7–11 holotype male, left palp. 7 prolateral, 8 dorsal, 9 frontal, 10 retrolateral, 11 apical. 12–13 paratype female, vulva. 12 ventral, 13 dorsal." figureDoi="http://doi.org/10.5281/zenodo.4575112" httpUri="https://zenodo.org/record/4575112/files/figure.png" pageId="1" pageNumber="589">Figs 7–11</figureCitation>
|
||
,
|
||
<figureCitation id="679C2A40FFE9FFF4FE310FF8FE551D15" box="[409,483,1770,1797]" captionStart="FIGURES 14–17" captionStartId="4.[151,264,1919,1943]" captionTargetBox="[157,1436,181,1889]" captionTargetId="figure-16@4.[151,1436,181,1895]" captionTargetPageId="4" captionText="FIGURES 14–17. D. zonsteini n. sp., 14-15 holotype male, left palp. 14 prolateral, 15 retrolateral. 16-17 paratype female, vulva. 16 ventral, 17 dorsal." figureDoi="http://doi.org/10.5281/zenodo.4575114" httpUri="https://zenodo.org/record/4575114/files/figure.png" pageId="1" pageNumber="589">14–15</figureCitation>
|
||
). Cymbium length 0.45; Male copulatory bulb length 0.63. Tegulum oval, almost rounded from retrolateral view. Embolic division wide, bent in the middle, unevenly sclerotized, with massive process starting from its base. Embolus opens at the distal end (
|
||
<figureCitation id="679C2A40FFE9FFF4FC960E21FC2A1D5D" box="[830,924,1843,1869]" captionStart="FIGURES 7–13" captionStartId="3.[151,264,1626,1650]" captionTargetBox="[151,1436,181,1603]" captionTargetId="figure-34@3.[151,1436,181,1603]" captionTargetPageId="3" captionText="FIGURES 7–13. D. zonsteini n. sp., 7–11 holotype male, left palp. 7 prolateral, 8 dorsal, 9 frontal, 10 retrolateral, 11 apical. 12–13 paratype female, vulva. 12 ventral, 13 dorsal." figureDoi="http://doi.org/10.5281/zenodo.4575112" httpUri="https://zenodo.org/record/4575112/files/figure.png" pageId="1" pageNumber="589">Figs. 10</figureCitation>
|
||
,
|
||
<figureCitation id="679C2A40FFE9FFF4FC000E20FC711D5D" box="[936,967,1842,1869]" captionStart="FIGURES 14–17" captionStartId="4.[151,264,1919,1943]" captionTargetBox="[157,1436,181,1889]" captionTargetId="figure-16@4.[151,1436,181,1895]" captionTargetPageId="4" captionText="FIGURES 14–17. D. zonsteini n. sp., 14-15 holotype male, left palp. 14 prolateral, 15 retrolateral. 16-17 paratype female, vulva. 16 ventral, 17 dorsal." figureDoi="http://doi.org/10.5281/zenodo.4575114" httpUri="https://zenodo.org/record/4575114/files/figure.png" pageId="1" pageNumber="589">15</figureCitation>
|
||
). Crest positioned diagonally in the distal part of the embolic division (
|
||
<figureCitation id="679C2A40FFE9FFF4FE430E45FDF41D61" box="[491,578,1878,1905]" captionStart="FIGURES 14–17" captionStartId="4.[151,264,1919,1943]" captionTargetBox="[157,1436,181,1889]" captionTargetId="figure-16@4.[151,1436,181,1895]" captionTargetPageId="4" captionText="FIGURES 14–17. D. zonsteini n. sp., 14-15 holotype male, left palp. 14 prolateral, 15 retrolateral. 16-17 paratype female, vulva. 16 ventral, 17 dorsal." figureDoi="http://doi.org/10.5281/zenodo.4575114" httpUri="https://zenodo.org/record/4575114/files/figure.png" pageId="1" pageNumber="589">Fig. 15</figureCitation>
|
||
). Lateral sheet weakly sclerotized, almost transparent. Posterior apophysis short and bent perpendicularly to the embolic division, detached from the tegulum and surrounded by the distal haematodocha.
|
||
</paragraph>
|
||
<caption id="ABD8664DFFEAFFF7FF3F0EA4FE841DE2" ID-DOI="http://doi.org/10.5281/zenodo.4575110" ID-Zenodo-Dep="4575110" httpUri="https://zenodo.org/record/4575110/files/figure.png" pageId="2" pageNumber="590" startId="2.[151,264,1974,1998]" targetBox="[357,1229,181,1950]" targetPageId="2">
|
||
<paragraph id="FF1836C5FFEAFFF7FF3F0EA4FE841DE2" blockId="2.[151,1437,1974,2034]" pageId="2" pageNumber="590">
|
||
<emphasis id="CDD3EAD7FFEAFFF7FF3F0EA4FE851DDE" bold="true" box="[151,307,1974,1998]" pageId="2" pageNumber="590">FIGURES 1–6</emphasis>
|
||
. Habitus of
|
||
<taxonomicName id="38A74D46FFEAFFF7FE060EA5FD901DDE" authority="Dimitrov, 2021" authorityName="Dimitrov" authorityYear="2021" box="[430,550,1975,1998]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="590" phylum="Arthropoda" rank="species" species="zonsteini" status="sp. nov.">
|
||
<emphasis id="CDD3EAD7FFEAFFF7FE060EA5FD901DDE" box="[430,550,1975,1998]" italics="true" pageId="2" pageNumber="590">D. zonsteini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CDD3EAD7FFEAFFF7FD840EA4FDD21DDE" bold="true" box="[556,612,1974,1998]" pageId="2" pageNumber="590">
|
||
<taxonomicNameLabel id="D6E057ACFFEAFFF7FD840EA4FDD21DDE" box="[556,612,1974,1998]" pageId="2" pageNumber="590" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, 1–2, 5 holotype male, 3–4, 6 paratype female. 1, 3 body dorsal. 2, 4 body ventral. 5–6 chelicerae.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="ABD8664DFFEBFFF6FF3F0F48FD1E1C87" ID-DOI="http://doi.org/10.5281/zenodo.4575112" ID-Zenodo-Dep="4575112" httpUri="https://zenodo.org/record/4575112/files/figure.png" pageId="3" pageNumber="591" startId="3.[151,264,1626,1650]" targetBox="[151,1436,181,1603]" targetPageId="3">
|
||
<paragraph id="FF1836C5FFEBFFF6FF3F0F48FD1E1C87" blockId="3.[151,1437,1626,1687]" pageId="3" pageNumber="591">
|
||
<emphasis id="CDD3EAD7FFEBFFF6FF3F0F48FEF41C63" bold="true" box="[151,322,1626,1651]" pageId="3" pageNumber="591">FIGURES 7–13</emphasis>
|
||
.
|
||
<taxonomicName id="38A74D46FFEBFFF6FEF80F4EFE7C1C62" authority="Dimitrov, 2021" authorityName="Dimitrov" authorityYear="2021" box="[336,458,1627,1651]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="zonsteini" status="sp. nov.">
|
||
<emphasis id="CDD3EAD7FFEBFFF6FEF80F4EFE7C1C62" box="[336,458,1627,1651]" italics="true" pageId="3" pageNumber="591">D. zonsteini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CDD3EAD7FFEBFFF6FE790F49FDBA1C63" bold="true" box="[465,524,1627,1651]" pageId="3" pageNumber="591">
|
||
<taxonomicNameLabel id="D6E057ACFFEBFFF6FE790F49FDBA1C63" box="[465,524,1627,1651]" pageId="3" pageNumber="591" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, 7–11 holotype male, left palp. 7 prolateral, 8 dorsal, 9 frontal, 10 retrolateral, 11 apical. 12–13 paratype female, vulva. 12 ventral, 13 dorsal.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FF1836C5FFEBFFF6FF6F0FD7FEC41DCC" blockId="3.[151,1437,1733,2012]" pageId="3" pageNumber="591">
|
||
Female. (
|
||
<figureCitation id="679C2A40FFEBFFF6FE9B0FD7FE0C1CCF" box="[307,442,1733,1759]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1974,1998]" captionTargetBox="[357,1229,181,1950]" captionTargetId="figure-16@2.[357,1229,181,1950]" captionTargetPageId="2" captionText="FIGURES 1–6. Habitus of D. zonsteini n. sp., 1–2, 5 holotype male, 3–4, 6 paratype female. 1, 3 body dorsal. 2, 4 body ventral. 5–6 chelicerae." figureDoi="http://doi.org/10.5281/zenodo.4575110" httpUri="https://zenodo.org/record/4575110/files/figure.png" pageId="3" pageNumber="591">Figs 3–4, 6</figureCitation>
|
||
,
|
||
<figureCitation id="679C2A40FFEBFFF6FE610FD7FDA71CCF" box="[457,529,1733,1759]" captionStart="FIGURES 7–13" captionStartId="3.[151,264,1626,1650]" captionTargetBox="[151,1436,181,1603]" captionTargetId="figure-34@3.[151,1436,181,1603]" captionTargetPageId="3" captionText="FIGURES 7–13. D. zonsteini n. sp., 7–11 holotype male, left palp. 7 prolateral, 8 dorsal, 9 frontal, 10 retrolateral, 11 apical. 12–13 paratype female, vulva. 12 ventral, 13 dorsal." figureDoi="http://doi.org/10.5281/zenodo.4575112" httpUri="https://zenodo.org/record/4575112/files/figure.png" pageId="3" pageNumber="591">12–13</figureCitation>
|
||
,
|
||
<figureCitation id="679C2A40FFEBFFF6FD880FD7FDDC1CF0" box="[544,618,1733,1760]" captionStart="FIGURES 14–17" captionStartId="4.[151,264,1919,1943]" captionTargetBox="[157,1436,181,1889]" captionTargetId="figure-16@4.[151,1436,181,1895]" captionTargetPageId="4" captionText="FIGURES 14–17. D. zonsteini n. sp., 14-15 holotype male, left palp. 14 prolateral, 15 retrolateral. 16-17 paratype female, vulva. 16 ventral, 17 dorsal." figureDoi="http://doi.org/10.5281/zenodo.4575114" httpUri="https://zenodo.org/record/4575114/files/figure.png" pageId="3" pageNumber="591">16–17</figureCitation>
|
||
). Measurements. Total length (including the chelicerae) 4.68; carapace length 1.80, width 1.26; sternum length 1.12, width 0.80; cheliceral length 0.72, width 0.35; clypeus height 0.05; labium length 0.41; gnathocoxa length 0.65; abdomen length 2.88, width 1.44; Eye diameters: AE—0.17, PE— 0.14. Leg measurements. I—5.41 (0.94, 1.25, 0.80, 1.04, 0.98, 0.40); II—4.92 (0.86, 1.12, 0.79, 0.90, 0.90, 0.35); III—3.34 (0.50, 0.76, 0.47, 0.54, 0.77, 0.30); IV—4.64 (0.76, 1.04, 0.58, 0.92, 1.04, 0.30). Leg spination as in male. Coloration very pale. Carapace, chelicerae and sternum light orange. Sternum a little lighter than the carapace. Abdomen and legs white. Anterior border of sternum as wide as the base of labium. Other somatic characters as in male (
|
||
<figureCitation id="679C2A40FFEBFFF6FF750ED3FED21DCC" box="[221,356,1985,2012]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1974,1998]" captionTargetBox="[357,1229,181,1950]" captionTargetId="figure-16@2.[357,1229,181,1950]" captionTargetPageId="2" captionText="FIGURES 1–6. Habitus of D. zonsteini n. sp., 1–2, 5 holotype male, 3–4, 6 paratype female. 1, 3 body dorsal. 2, 4 body ventral. 5–6 chelicerae." figureDoi="http://doi.org/10.5281/zenodo.4575110" httpUri="https://zenodo.org/record/4575110/files/figure.png" pageId="3" pageNumber="591">Figs. 3–4, 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="ABD8664DFFECFFF1FF3F0E6DFE021DAB" ID-DOI="http://doi.org/10.5281/zenodo.4575114" ID-Zenodo-Dep="4575114" httpUri="https://zenodo.org/record/4575114/files/figure.png" pageId="4" pageNumber="592" startId="4.[151,264,1919,1943]" targetBox="[157,1436,181,1889]" targetPageId="4">
|
||
<paragraph id="FF1836C5FFECFFF1FF3F0E6DFE021DAB" blockId="4.[151,1437,1919,1979]" pageId="4" pageNumber="592">
|
||
<emphasis id="CDD3EAD7FFECFFF1FF3F0E6DFEE61D87" bold="true" box="[151,336,1919,1943]" pageId="4" pageNumber="592">FIGURES 14–17</emphasis>
|
||
.
|
||
<taxonomicName id="38A74D46FFECFFF1FEF70E92FE6D1D87" authority="Dimitrov, 2021" authorityName="Dimitrov" authorityYear="2021" box="[351,475,1920,1943]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="592" phylum="Arthropoda" rank="species" species="zonsteini" status="sp. nov.">
|
||
<emphasis id="CDD3EAD7FFECFFF1FEF70E92FE6D1D87" box="[351,475,1920,1943]" italics="true" pageId="4" pageNumber="592">D. zonsteini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CDD3EAD7FFECFFF1FE4C0E6DFDA91D87" bold="true" box="[484,543,1919,1943]" pageId="4" pageNumber="592">
|
||
<taxonomicNameLabel id="D6E057ACFFECFFF1FE4C0E6DFDA91D87" box="[484,543,1919,1943]" pageId="4" pageNumber="592" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, 14-15 holotype male, left palp. 14 prolateral, 15 retrolateral. 16-17 paratype female, vulva. 16 ventral, 17 dorsal.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FF1836C5FFEDFFF0FF6F0985FE671B0D" blockId="5.[151,1437,151,1077]" pageId="5" pageNumber="593">
|
||
Vulva. (
|
||
<figureCitation id="679C2A40FFEDFFF0FE880985FE171AA1" box="[288,417,151,177]" captionStart="FIGURES 7–13" captionStartId="3.[151,264,1626,1650]" captionTargetBox="[151,1436,181,1603]" captionTargetId="figure-34@3.[151,1436,181,1603]" captionTargetPageId="3" captionText="FIGURES 7–13. D. zonsteini n. sp., 7–11 holotype male, left palp. 7 prolateral, 8 dorsal, 9 frontal, 10 retrolateral, 11 apical. 12–13 paratype female, vulva. 12 ventral, 13 dorsal." figureDoi="http://doi.org/10.5281/zenodo.4575112" httpUri="https://zenodo.org/record/4575112/files/figure.png" pageId="5" pageNumber="593">Figs 12–13</figureCitation>
|
||
,
|
||
<figureCitation id="679C2A40FFEDFFF0FE050985FE411AA2" box="[429,503,151,178]" captionStart="FIGURES 14–17" captionStartId="4.[151,264,1919,1943]" captionTargetBox="[157,1436,181,1889]" captionTargetId="figure-16@4.[151,1436,181,1895]" captionTargetPageId="4" captionText="FIGURES 14–17. D. zonsteini n. sp., 14-15 holotype male, left palp. 14 prolateral, 15 retrolateral. 16-17 paratype female, vulva. 16 ventral, 17 dorsal." figureDoi="http://doi.org/10.5281/zenodo.4575114" httpUri="https://zenodo.org/record/4575114/files/figure.png" pageId="5" pageNumber="593">16–17</figureCitation>
|
||
). Transversal bar width 0.38. Spermatheca small, with mushroom-like anterior part and deeply notched base. Dorsal arc of anterior diverticulum shaped as an irregular trapeze, wider than the spermatheca, weakly sclerotized, with almost transparent edges. Transversal bar comparatively straight. Posterior diverticulum wide, membranous.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B7BD654EFFEDFFF0FF6F0834FBFA1B51" box="[199,1100,294,321]" pageId="5" pageNumber="593" type="distribution">
|
||
<paragraph id="FF1836C5FFEDFFF0FF6F0834FBFA1B51" blockId="5.[151,1437,151,1077]" box="[199,1100,294,321]" pageId="5" pageNumber="593">
|
||
<emphasis id="CDD3EAD7FFEDFFF0FF6F0834FED71B50" bold="true" box="[199,353,294,320]" pageId="5" pageNumber="593">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="201C8867FFEDFFF0FDC80835FD261B51" box="[608,656,295,321]" pageId="5" pageNumber="593">type</typeStatus>
|
||
locality in southwestern
|
||
<collectingCountry id="87B07655FFEDFFF0FC010835FBF11B51" box="[937,1095,295,321]" name="Turkmenistan" pageId="5" pageNumber="593">Turkmenistan</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B7BD654EFFEDFFF0FF6F0858FF461E25" pageId="5" pageNumber="593" type="discussion">
|
||
<paragraph id="FF1836C5FFEDFFF0FF6F0858FB56182D" blockId="5.[151,1437,151,1077]" pageId="5" pageNumber="593">
|
||
<emphasis id="CDD3EAD7FFEDFFF0FF6F0858FEFA1B74" bold="true" box="[199,332,330,356]" pageId="5" pageNumber="593">Discussion.</emphasis>
|
||
The new species’ morphologically closest congener
|
||
<taxonomicName id="38A74D46FFEDFFF0FC300859FB851B75" authorityName="Dunin" authorityYear="1992" box="[920,1075,331,357]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="kronebergi">
|
||
<emphasis id="CDD3EAD7FFEDFFF0FC300859FB851B75" box="[920,1075,331,357]" italics="true" pageId="5" pageNumber="593">D. kronebergi</emphasis>
|
||
</taxonomicName>
|
||
was diagnosed as being close to the
|
||
<taxonomicName id="38A74D46FFEDFFF0FF69087DFED31B98" authorityName="Roewer" authorityYear="1928" box="[193,357,367,393]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="gigas">
|
||
<emphasis id="CDD3EAD7FFEDFFF0FF69087DFED31B98" box="[193,357,367,393]" italics="true" pageId="5" pageNumber="593">Dysdera gigas</emphasis>
|
||
</taxonomicName>
|
||
species-group (
|
||
<bibRefCitation id="9B364B34FFEDFFF0FDBF087DFD511B99" author="Dunin, P. M." box="[535,743,367,393]" pageId="5" pageNumber="593" pagination="35 - 76" refId="ref2707" refString="Dunin, P. M. (1992 a) The spider family Dysderidae of the Caucasian fauna (Arachnida Aranei Haplogynae). Arthropoda Selecta, 1 (3), 35 - 76." type="journal article" year="1992">Dunin 1992a: 137</bibRefCitation>
|
||
), referring to
|
||
<bibRefCitation id="9B364B34FFEDFFF0FC2B087DFAE41B99" author="Deeleman-Reinhold, C. L. & Deeleman, P. R." box="[899,1362,367,393]" pageId="5" pageNumber="593" pagination="141 - 269" refId="ref2593" refString="Deeleman-Reinhold, C. L. & Deeleman, P. R. (1988) Revision des Dysderinae (Araneae, Dysderidae), les especes mediterraneennes occidentales exceptees. Tijdschrift voor Entomologie, 131, 141 - 269." type="journal article" year="1988">Deeleman-Reinhold & Deeleman (1988)</bibRefCitation>
|
||
. However, there is no
|
||
<taxonomicName id="38A74D46FFEDFFF0FEE50881FE1D1BBC" authorityName="Roewer" authorityYear="1928" box="[333,427,403,428]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="gigas">
|
||
<emphasis id="CDD3EAD7FFEDFFF0FEE50881FE1D1BBC" box="[333,427,403,428]" italics="true" pageId="5" pageNumber="593">D. gigas</emphasis>
|
||
</taxonomicName>
|
||
-group defined in the cited paper. Instead,
|
||
<taxonomicName id="38A74D46FFEDFFF0FCD60881FC6B1BBC" authorityName="Roewer" authorityYear="1928" box="[894,989,403,428]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="gigas">
|
||
<emphasis id="CDD3EAD7FFEDFFF0FCD60881FC6B1BBC" box="[894,989,403,428]" italics="true" pageId="5" pageNumber="593">D. gigas</emphasis>
|
||
</taxonomicName>
|
||
along with another three similar species are listed under the
|
||
<taxonomicName id="38A74D46FFEDFFF0FED208A5FD8C1BC0" authorityName="Nosek" authorityYear="1905" box="[378,570,439,465]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="asiatica">
|
||
<emphasis id="CDD3EAD7FFEDFFF0FED208A5FD8C1BC0" box="[378,570,439,465]" italics="true" pageId="5" pageNumber="593">Dysdera asiatica</emphasis>
|
||
</taxonomicName>
|
||
-group (
|
||
<bibRefCitation id="9B364B34FFEDFFF0FD3D08A5FB261BC1" author="Deeleman-Reinhold, C. L. & Deeleman, P. R." box="[661,1168,439,465]" pageId="5" pageNumber="593" pagination="141 - 269" refId="ref2593" refString="Deeleman-Reinhold, C. L. & Deeleman, P. R. (1988) Revision des Dysderinae (Araneae, Dysderidae), les especes mediterraneennes occidentales exceptees. Tijdschrift voor Entomologie, 131, 141 - 269." type="journal article" year="1988">Deeleman-Reinhold & Deeleman 1988: 210</bibRefCitation>
|
||
). The authors explicitly mention that the lateral sheet is missing in these species whereas in
|
||
<taxonomicName id="38A74D46FFEDFFF0FC3E08C9FBAB1BE4" authority="Dimitrov, 2021" authorityName="Dimitrov" authorityYear="2021" box="[918,1053,475,500]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="zonsteini" status="sp. nov.">
|
||
<emphasis id="CDD3EAD7FFEDFFF0FC3E08C9FBAB1BE4" box="[918,1053,475,500]" italics="true" pageId="5" pageNumber="593">D. zonsteini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CDD3EAD7FFEDFFF0FB8C08C9FBD51BE5" bold="true" box="[1060,1123,475,501]" pageId="5" pageNumber="593">
|
||
<taxonomicNameLabel id="D6E057ACFFEDFFF0FB8C08C9FBD51BE5" box="[1060,1123,475,501]" pageId="5" pageNumber="593" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
it exists. All the species related to
|
||
<taxonomicName id="38A74D46FFEDFFF0FF5908EDFEE61808" authorityName="Roewer" authorityYear="1928" box="[241,336,511,536]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="gigas">
|
||
<emphasis id="CDD3EAD7FFEDFFF0FF5908EDFEE61808" box="[241,336,511,536]" italics="true" pageId="5" pageNumber="593">D. gigas</emphasis>
|
||
</taxonomicName>
|
||
are known from
|
||
<collectingCountry id="87B07655FFEDFFF0FDB808EDFDE91809" box="[528,607,511,537]" name="Greece" pageId="5" pageNumber="593">Greece</collectingCountry>
|
||
and the Greek islands and their relationship with
|
||
<taxonomicName id="38A74D46FFEDFFF0FB2408EDFA911809" authorityName="Dunin" authorityYear="1992" box="[1164,1319,511,537]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="kronebergi">
|
||
<emphasis id="CDD3EAD7FFEDFFF0FB2408EDFA911809" box="[1164,1319,511,537]" italics="true" pageId="5" pageNumber="593">D. kronebergi</emphasis>
|
||
</taxonomicName>
|
||
suggested by Dunin is doubtful, since neither the genital morphology nor their distribution ranges support it.
|
||
</paragraph>
|
||
<paragraph id="FF1836C5FFEDFFF0FF6F0B55FCCA18DD" blockId="5.[151,1437,151,1077]" pageId="5" pageNumber="593">
|
||
Some important morphological traits of the new species correspond well with the diagnose of the
|
||
<taxonomicName id="38A74D46FFEDFFF0FABF0B55FA251870" authorityName="Nosek" authorityYear="1905" box="[1303,1427,583,608]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="asiatica">
|
||
<emphasis id="CDD3EAD7FFEDFFF0FABF0B55FA251870" box="[1303,1427,583,608]" italics="true" pageId="5" pageNumber="593">D. asiatica</emphasis>
|
||
</taxonomicName>
|
||
-group (sensu
|
||
<bibRefCitation id="9B364B34FFEDFFF0FE870B79FD5A1895" author="Deeleman-Reinhold, C. L. & Deeleman, P. R." box="[303,748,619,645]" pageId="5" pageNumber="593" pagination="141 - 269" refId="ref2593" refString="Deeleman-Reinhold, C. L. & Deeleman, P. R. (1988) Revision des Dysderinae (Araneae, Dysderidae), les especes mediterraneennes occidentales exceptees. Tijdschrift voor Entomologie, 131, 141 - 269." type="journal article" year="1988">Deeleman-Reinhold & Deeleman 1988</bibRefCitation>
|
||
): 1. Chelicerae are shorter than half of the length of the carapace; 2. The embolic division is longer than the tegulum; 3. The presence of crest and lateral sheet on the embolic division of the male palp; 4. The arched spermatheca of the vulva.
|
||
</paragraph>
|
||
<paragraph id="FF1836C5FFEDFFF0FF6F0BC5FDC119D9" blockId="5.[151,1437,151,1077]" pageId="5" pageNumber="593">
|
||
However, the diagnosis of the group itself (sensu
|
||
<bibRefCitation id="9B364B34FFEDFFF0FD5E0BC5FB1918E1" author="Deeleman-Reinhold, C. L. & Deeleman, P. R." box="[758,1199,727,753]" pageId="5" pageNumber="593" pagination="141 - 269" refId="ref2593" refString="Deeleman-Reinhold, C. L. & Deeleman, P. R. (1988) Revision des Dysderinae (Araneae, Dysderidae), les especes mediterraneennes occidentales exceptees. Tijdschrift voor Entomologie, 131, 141 - 269." type="journal article" year="1988">Deeleman-Reinhold & Deeleman 1988</bibRefCitation>
|
||
) has one serious disadvantage: it is named after a species known only by male sex. Thus, all the other members of the group are linked to
|
||
<taxonomicName id="38A74D46FFEDFFF0FF1B0A0DFEC71928" authorityName="Nosek" authorityYear="1905" box="[179,369,799,825]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="asiatica">
|
||
<emphasis id="CDD3EAD7FFEDFFF0FF1B0A0DFEC71928" box="[179,369,799,825]" italics="true" pageId="5" pageNumber="593">Dysdera asiatica</emphasis>
|
||
</taxonomicName>
|
||
based only on the males. Apart from that, as already stated by
|
||
<bibRefCitation id="9B364B34FFEDFFF0FB8F0A0DFEAA194D" author="Deeleman-Reinhold, C. L. & Deeleman, P. R." pageId="5" pageNumber="593" pagination="141 - 269" refId="ref2593" refString="Deeleman-Reinhold, C. L. & Deeleman, P. R. (1988) Revision des Dysderinae (Araneae, Dysderidae), les especes mediterraneennes occidentales exceptees. Tijdschrift voor Entomologie, 131, 141 - 269." type="journal article" year="1988">Deeleman-Reinhold & Deeleman (1988: 210)</bibRefCitation>
|
||
, the group is very rich in species and in Central Asia is neither homogenous nor can be clearly separated from the
|
||
<taxonomicName id="38A74D46FFEDFFF0FF570A75FE341991" authorityName="Kroneberg" authorityYear="1875" box="[255,386,871,897]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="aculeata">
|
||
<emphasis id="CDD3EAD7FFEDFFF0FF570A75FE341991" box="[255,386,871,897]" italics="true" pageId="5" pageNumber="593">D. aculeata</emphasis>
|
||
</taxonomicName>
|
||
-group. The group warrants a thorough new revision. Here, I provisionally place
|
||
<taxonomicName id="38A74D46FFEDFFF0FABE0A75FA2B1990" authority="Dimitrov, 2021" authorityName="Dimitrov" authorityYear="2021" box="[1302,1437,871,896]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="zonsteini" status="sp. nov.">
|
||
<emphasis id="CDD3EAD7FFEDFFF0FABE0A75FA2B1990" box="[1302,1437,871,896]" italics="true" pageId="5" pageNumber="593">D. zonsteini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CDD3EAD7FFEDFFF0FF3F0A99FF6319B5" bold="true" box="[151,213,907,933]" pageId="5" pageNumber="593">
|
||
<taxonomicNameLabel id="D6E057ACFFEDFFF0FF3F0A99FF6319B5" box="[151,213,907,933]" pageId="5" pageNumber="593" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in
|
||
<taxonomicName id="38A74D46FFEDFFF0FF510A99FEC219B4" authorityName="Nosek" authorityYear="1905" box="[249,372,907,932]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="asiatica">
|
||
<emphasis id="CDD3EAD7FFEDFFF0FF510A99FEC219B4" box="[249,372,907,932]" italics="true" pageId="5" pageNumber="593">D. asiatica</emphasis>
|
||
</taxonomicName>
|
||
species-group. Most of the other species, known from
|
||
<collectingCountry id="87B07655FFEDFFF0FC700A99FBC519B5" box="[984,1139,907,933]" name="Turkmenistan" pageId="5" pageNumber="593">Turkmenistan</collectingCountry>
|
||
belong to the
|
||
<taxonomicName id="38A74D46FFEDFFF0FAB90A99FA2419B5" authorityName="Kroneberg" authorityYear="1875" box="[1297,1426,907,933]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="aculeata">
|
||
<emphasis id="CDD3EAD7FFEDFFF0FAB90A99FA2419B5" box="[1297,1426,907,933]" italics="true" pageId="5" pageNumber="593">D. aculeata</emphasis>
|
||
</taxonomicName>
|
||
-group according to the same classification.
|
||
</paragraph>
|
||
<paragraph id="FF1836C5FFEDFFF0FF6F0AC1FF461E25" blockId="5.[151,1437,151,1077]" pageId="5" pageNumber="593">
|
||
Although the data about the distribution of
|
||
<taxonomicName id="38A74D46FFEDFFF0FD6B0AC1FCFA19FC" authority="Dimitrov, 2021" authorityName="Dimitrov" authorityYear="2021" box="[707,844,979,1004]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="zonsteini" status="sp. nov.">
|
||
<emphasis id="CDD3EAD7FFEDFFF0FD6B0AC1FCFA19FC" box="[707,844,979,1004]" italics="true" pageId="5" pageNumber="593">D. zonsteini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CDD3EAD7FFEDFFF0FCFE0AC1FC2E19FD" bold="true" box="[854,920,979,1005]" pageId="5" pageNumber="593">
|
||
<taxonomicNameLabel id="D6E057ACFFEDFFF0FCFE0AC1FC2E19FD" box="[854,920,979,1005]" pageId="5" pageNumber="593" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are insufficient to identify with certainty its zoogeographical status, its occurrence in the southern part of the country implies that it could be an Irano-Turanian species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |