297 lines
29 KiB
XML
297 lines
29 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1019.61496" ID-GBIF-Dataset="5e7c2faa-0a3e-4533-9104-0cd524762834" ID-PMC="PMC7921071" ID-Pensoft-Pub="1313-2970-1019-141" ID-Pensoft-UUID="616BD8CDECF259DA91B015E9A7737DAC" ID-PubMed="33679172" ID-ZooBank="8CE802531BA74F80820675CA0C003101" ModsDocID="1313-2970-1019-141" checkinTime="1614092291421" checkinUser="pensoft" docAuthor="Kanavalova, Liliana, Grootaert, Patrick, Kubik, Stepan & Bartak, Miroslav" docDate="2021" docId="80E40E7744EF5E21A3D1235F0EEC16C8" docLanguage="en" docName="ZooKeys 1019: 141-162" docOrigin="ZooKeys 1019" docPubDate="2021-02-22" docSource="http://dx.doi.org/10.3897/zookeys.1019.61496" docTitle="Megagrapha starki Bartak & Grootaert 2021, sp. nov." docType="treatment" docUuid="0023393E-CB52-4C09-A30D-4D1AC3D3D336" docUuidSource="ZooBank" docVersion="4" id="616BD8CDECF259DA91B015E9A7737DAC" lastPageNumber="141" masterDocId="616BD8CDECF259DA91B015E9A7737DAC" masterDocTitle="Four new West Palaearctic species and new distributional records of Hybotidae (Diptera)" masterLastPageNumber="162" masterPageNumber="141" pageNumber="141" updateTime="1668149814991" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Four new West Palaearctic species and new distributional records of Hybotidae (Diptera)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kanavalova, Liliana</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4029-8265</mods:nameIdentifier>
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<mods:affiliation>Department of Zoology and Fisheries, Faculty of Agrobiology, Food and Natural Resources, Czech University of Life Sciences Prague, Kamycka 129, 165 00, Praha-Suchdol, Czech Republic</mods:affiliation>
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<mods:nameIdentifier type="email">kanavalova@af.czu.cz</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Grootaert, Patrick</mods:namePart>
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<mods:affiliation>Department of Entomology, Royal Belgian Institute of Natural Sciences, Rue Vautier 29, B - 1000, Brussels, Belgium</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kubik, Stepan</mods:namePart>
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<mods:affiliation>Department of Zoology and Fisheries, Faculty of Agrobiology, Food and Natural Resources, Czech University of Life Sciences Prague, Kamycka 129, 165 00, Praha-Suchdol, Czech Republic</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bartak, Miroslav</mods:namePart>
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<mods:affiliation>Department of Zoology and Fisheries, Faculty of Agrobiology, Food and Natural Resources, Czech University of Life Sciences Prague, Kamycka 129, 165 00, Praha-Suchdol, Czech Republic</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-02-22</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1019</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>141</mods:start>
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<mods:end>162</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1019.61496</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1019.61496</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1019-141</mods:identifier>
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<mods:identifier type="ZooBank">8CE802531BA74F80820675CA0C003101</mods:identifier>
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<mods:identifier type="Pensoft-UUID">616BD8CDECF259DA91B015E9A7737DAC</mods:identifier>
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<treatment ID-GBIF-Taxon="179905020" LSID="urn:lsid:zoobank.org:act:0023393E-CB52-4C09-A30D-4D1AC3D3D336" httpUri="http://treatment.plazi.org/id/80E40E7744EF5E21A3D1235F0EEC16C8" lastPageNumber="141" pageId="0" pageNumber="141">
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<subSubSection pageId="0" pageNumber="141" type="nomenclature">
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<paragraph pageId="0" pageNumber="141">
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<taxonomicName LSID="http://zoobank.org/0023393E-CB52-4C09-A30D-4D1AC3D3D336" authority="Barták & Grootaert" authorityName="Bartak & Grootaert" authorityYear="2021" class="Insecta" family="Hybotidae" genus="Megagrapha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megagrapha starki" order="Diptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="starki" status="sp. nov.">
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Megagrapha starki
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<normalizedToken originalValue="Barták">Bartak</normalizedToken>
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& Grootaert
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</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="141">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megagrapha starki sp. nov. Male habitus, paratype." figureDoi="10.3897/zookeys.1019.61496.figure3" httpUri="https://binary.pensoft.net/fig/512623" pageId="0" pageNumber="141">Figs 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megagrapha starki sp. nov., male antenna, paratype." figureDoi="10.3897/zookeys.1019.61496.figure4" httpUri="https://binary.pensoft.net/fig/512624" pageId="0" pageNumber="141">, 4</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Megagrapha starki sp. nov., male wing, paratype." figureDoi="10.3897/zookeys.1019.61496.figure5" httpUri="https://binary.pensoft.net/fig/512625" pageId="0" pageNumber="141">, 5</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megagrapha starki sp. nov., male terminalia A right epandrial lamella B epandrium with cerci, dorsal C left epandrial lamella D detail of phallus. Abbr. hyp: hypandrium; lc: left cercus; lel: left epandrial lamella; ll: lower lobe of left surstylus; ul: upper lobe of left surstylus; rel: right epandrial lamella. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1019.61496.figure6" httpUri="https://binary.pensoft.net/fig/512626" pageId="0" pageNumber="141">, 6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="141" type="type material">
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<paragraph pageId="0" pageNumber="141">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="141">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="141">Holotype</emphasis>
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♂,
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<emphasis bold="true" pageId="0" pageNumber="141">Slovakia</emphasis>
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:
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<normalizedToken originalValue="Muráňska">Muranska</normalizedToken>
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planina NPR,
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<normalizedToken originalValue="Šarkanica">Sarkanica</normalizedToken>
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, 540 m, MT,
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<geoCoordinate degrees="48" direction="north" minutes="42" orientation="latitude" precision="15" seconds="46" value="48.71278">48°42'46"N</geoCoordinate>
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,
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<geoCoordinate degrees="19" direction="east" minutes="59" orientation="longitude" precision="15" seconds="52" value="19.997778">19°59'52"E</geoCoordinate>
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, M.
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<normalizedToken originalValue="Tkoč">Tkoc</normalizedToken>
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, 26.iv.-15.vi.2017 (CULSP).
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<emphasis bold="true" pageId="0" pageNumber="141">
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<emphasis italics="true" pageId="0" pageNumber="141">Paratypes</emphasis>
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: Slovakia
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</emphasis>
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, 1♂,
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<normalizedToken originalValue="Muráňska">Muranska</normalizedToken>
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planina NP, 580 m, MT,
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<geoCoordinate degrees="48" direction="north" minutes="42" orientation="latitude" precision="15" seconds="46" value="48.71278">48°42'46"N</geoCoordinate>
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,
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<geoCoordinate degrees="19" direction="east" minutes="59" orientation="longitude" precision="15" seconds="56" value="19.99889">19°59'56"E</geoCoordinate>
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, J.
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<normalizedToken originalValue="Roháček">Rohacek</normalizedToken>
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, 14.vi.-11.vii.2017 (dissect., RBINS);
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<emphasis bold="true" pageId="0" pageNumber="141">Poland</emphasis>
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: 1♂, Bialowieza, 'Bialowieza
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<normalizedToken originalValue="Forest’">Forest'</normalizedToken>
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, Promotion Area,
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<normalizedToken originalValue="‘Lesna’">'Lesna'</normalizedToken>
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Reserve, 15 km SE of core zone of NP, 160 m a.s.l., 120 year old Carpinus betulus fogged with pyrethrum,
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<geoCoordinate degrees="52" direction="north" minutes="37" orientation="latitude" precision="15" seconds="41" value="52.628056">52°37'41"N</geoCoordinate>
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,
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<geoCoordinate degrees="23" direction="east" minutes="46" orientation="longitude" precision="15" seconds="12" value="23.77">23°46'12"E</geoCoordinate>
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, A. Floren, 28.vi. 2001 (CAStH);
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<emphasis bold="true" pageId="0" pageNumber="141">Russia</emphasis>
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: 1♂, SW Moscow, Moskovskiy, forest edge, YPT traps,
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<geoCoordinate degrees="55" direction="north" minutes="35" orientation="latitude" precision="15" seconds="16" value="55.587776">55°35'16"N</geoCoordinate>
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,
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<geoCoordinate degrees="37" direction="east" minutes="19" orientation="longitude" precision="15" seconds="58" value="37.332775">37°19'58"E</geoCoordinate>
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, K. Tomkovich, 1-4.viii.2014 (ZMMU).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="141" type="diagnosis">
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<paragraph pageId="0" pageNumber="141">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="141">
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A species of
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<taxonomicName class="Insecta" family="Hybotidae" genus="Megagrapha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megagrapha" order="Diptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="141">Megagrapha</emphasis>
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</taxonomicName>
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Melander with patterned wings and modified male fore and mid basitarsi, very similar to Nearctic
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<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. exquisita" order="Diptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="exquisita">
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<emphasis italics="true" pageId="0" pageNumber="141">M. exquisita</emphasis>
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</taxonomicName>
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(Malloch, 1923). However, the newly described species differs in the colour of the mesoscutum (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megagrapha starki sp. nov. Male habitus, paratype." figureDoi="10.3897/zookeys.1019.61496.figure3" httpUri="https://binary.pensoft.net/fig/512623" pageId="0" pageNumber="141">3</figureCitation>
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), the dark pattern on the wing (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Megagrapha starki sp. nov., male wing, paratype." figureDoi="10.3897/zookeys.1019.61496.figure5" httpUri="https://binary.pensoft.net/fig/512625" pageId="0" pageNumber="141">5</figureCitation>
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), the shape of the postpedicel (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megagrapha starki sp. nov., male antenna, paratype." figureDoi="10.3897/zookeys.1019.61496.figure4" httpUri="https://binary.pensoft.net/fig/512624" pageId="0" pageNumber="141">4</figureCitation>
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) and the male terminalia (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megagrapha starki sp. nov., male terminalia A right epandrial lamella B epandrium with cerci, dorsal C left epandrial lamella D detail of phallus. Abbr. hyp: hypandrium; lc: left cercus; lel: left epandrial lamella; ll: lower lobe of left surstylus; ul: upper lobe of left surstylus; rel: right epandrial lamella. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1019.61496.figure6" httpUri="https://binary.pensoft.net/fig/512626" pageId="0" pageNumber="141">6</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.1019.61496.figure3" httpUri="https://binary.pensoft.net/fig/512623" pageId="0" pageNumber="141" start="Figure 3" startId="F3">
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<paragraph pageId="0" pageNumber="141">
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<emphasis bold="true" pageId="0" pageNumber="141">Figure 3.</emphasis>
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<taxonomicName authorityName="Bartak & Grootaert" authorityYear="2021" class="Insecta" family="Hybotidae" genus="Megagrapha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megagrapha starki" order="Diptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="starki">
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<emphasis italics="true" pageId="0" pageNumber="141">Megagrapha starki</emphasis>
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</taxonomicName>
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sp. nov. Male habitus, paratype.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="141" type="etymology">
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<paragraph pageId="0" pageNumber="141">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="141">
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The species epithet,
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<taxonomicName authorityName="Bartak & Grootaert" authorityYear="2021" class="Insecta" family="Hybotidae" genus="Megagrapha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megagrapha starki" order="Diptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="starki">
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<emphasis italics="true" pageId="0" pageNumber="141">Megagrapha starki</emphasis>
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</taxonomicName>
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, is a Latin genitive patronym given in honour of German dipterist Andreas Stark (Halle (Saale)), who first recognised this species as new, acknowledging his contribution to the knowledge of world
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<taxonomicName kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="141" rank="superFamily" superFamily="Empidoidea">Empidoidea</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="141" type="description">
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<paragraph pageId="0" pageNumber="141">Description.</paragraph>
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<paragraph pageId="0" pageNumber="141">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="141">Male head</emphasis>
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black, rounded in anterior view and ovoid in lateral view. Large eyes with dense and relatively long ommatrichia, all facets subequal in size. Frons very narrow (linear, narrower in lower part than diameter of one ommatidium, widening towards ocellar triangle). Slightly prominent ocellar triangle and adjacent parts of frons and vertex sublustrous, with 2 pairs of short, subequally long and pale ocellars (about 0.10 mm long), with several similar setae posteriorly. Occiput microtrichose (subshiny), with dense yellowish-brown to black setae on dorsal third similar in length to ocellars, ventrally with fewer short, pale setae, posteriorly with several black setae. Face very narrow dorsally and widening ventrally, microtrichose, without setae, gena indistinct. Palpus brown, relatively large (0.22 mm long) and ovoid, slightly narrowing towards tip, covered with relatively long black setae. Labrum yellow, only slightly longer than palpus, labellum narrow with several setae. Antenna yellow (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megagrapha starki sp. nov., male antenna, paratype." figureDoi="10.3897/zookeys.1019.61496.figure4" httpUri="https://binary.pensoft.net/fig/512624" pageId="0" pageNumber="141">4</figureCitation>
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). Scape without setae, 0.08 mm long, pedicel (0.05 mm long) with circlet of short black setae, postpedicel (0.24 mm long) elongated, with rather long narrow apical half. Stylus (0.15 mm long) with several slightly elongated darkened setulae (especially in apical part).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="141">Thorax</emphasis>
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yellow with brown pattern in form of 2 longitudinal stripes on mesoscutum spreading from black antepronotum posteriorly, bent near scutellum towards postalar calli and connected with a broad stripe medially in front of scutellum (dark pattern in dorsal view roughly U-shaped). Postpronotum and adjacent parts of mesoscutum, including notopleuron, brown, slightly brownish parts occur below wing base including mediotergite. Scutellum black and microtrichose except basolateral corners. Mesoscutum mostly lustrous, pleura shiny except dorsoposterior part of anepisternum. Both anterior and posterior spiracles deep black. Chaetotaxy: antepronotum with 4-5 pale setulae on each side, 3 notopleurals, 1 short postalar, 3 pairs of pale scutellars. Mesoscutum densely covered with black setulae anteriorly (also on postpronotum and adjacent areas) and white setulae posteriorly, setae longer in presutural area with 1 not very prominent prescutellar pair. Pleura with several black setae dorsally on anepisternum.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="141">Legs</emphasis>
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including coxae very pale (almost whitish-) yellow, rather densely covered with short, whitish setulae, some longer setae only preapically on ventral side of all femora. Basitarsus of fore leg with narrow ventral process and basitarsus of mid leg narrowed basally and widened apically, with short ventral apical projection. Second mid tarsomere with dense setulae ventrally.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="141">Wing</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Megagrapha starki sp. nov., male wing, paratype." figureDoi="10.3897/zookeys.1019.61496.figure5" httpUri="https://binary.pensoft.net/fig/512625" pageId="0" pageNumber="141">5</figureCitation>
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) with distinct dark pattern, covered on both sides with relatively long microtrichia. Basal costal seta only slightly differentiated. Sc incomplete, closely approximated to R1 and vanishing at about middle. Costa ends at tip of M1 vein. Halter yellow, calypter black with yellow margin and long black fringes.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="141">Abdomen</emphasis>
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subshiny, with brown (and brown setose) tergites and yellow (pale setose) sternites, lateromarginal setae on tergites as long as or longer than segments. Terminalia as in Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megagrapha starki sp. nov., male terminalia A right epandrial lamella B epandrium with cerci, dorsal C left epandrial lamella D detail of phallus. Abbr. hyp: hypandrium; lc: left cercus; lel: left epandrial lamella; ll: lower lobe of left surstylus; ul: upper lobe of left surstylus; rel: right epandrial lamella. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1019.61496.figure6" httpUri="https://binary.pensoft.net/fig/512626" pageId="0" pageNumber="141">6A-D</figureCitation>
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. Cerci narrow, equally long, shorter than right epandrial lamella. Left surstylus composed of lower lobe (ll) and digitiform upper lobe (ul), bearing setae longer than lobe.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="141">Length</emphasis>
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: body 2.4 mm, wing 2.6 mm.
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</paragraph>
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<caption doi="10.3897/zookeys.1019.61496.figure4" httpUri="https://binary.pensoft.net/fig/512624" pageId="0" pageNumber="141" start="Figure 4" startId="F4">
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<paragraph pageId="0" pageNumber="141">
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<emphasis bold="true" pageId="0" pageNumber="141">Figure 4.</emphasis>
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<taxonomicName authorityName="Bartak & Grootaert" authorityYear="2021" class="Insecta" family="Hybotidae" genus="Megagrapha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megagrapha starki" order="Diptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="starki">
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<emphasis italics="true" pageId="0" pageNumber="141">Megagrapha starki</emphasis>
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</taxonomicName>
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sp. nov., male antenna, paratype.
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</paragraph>
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</caption>
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<caption doi="10.3897/zookeys.1019.61496.figure5" httpUri="https://binary.pensoft.net/fig/512625" pageId="0" pageNumber="141" start="Figure 5" startId="F5">
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<paragraph pageId="0" pageNumber="141">
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<emphasis bold="true" pageId="0" pageNumber="141">Figure 5.</emphasis>
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<taxonomicName authorityName="Bartak & Grootaert" authorityYear="2021" class="Insecta" family="Hybotidae" genus="Megagrapha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megagrapha starki" order="Diptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="starki">
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<emphasis italics="true" pageId="0" pageNumber="141">Megagrapha starki</emphasis>
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</taxonomicName>
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sp. nov., male wing, paratype.
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</paragraph>
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</caption>
|
||
<caption doi="10.3897/zookeys.1019.61496.figure6" httpUri="https://binary.pensoft.net/fig/512626" pageId="0" pageNumber="141" start="Figure 6" startId="F6">
|
||
<paragraph pageId="0" pageNumber="141">
|
||
<emphasis bold="true" pageId="0" pageNumber="141">Figure 6.</emphasis>
|
||
<taxonomicName authorityName="Bartak & Grootaert" authorityYear="2021" class="Insecta" family="Hybotidae" genus="Megagrapha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megagrapha starki" order="Diptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="starki">
|
||
<emphasis italics="true" pageId="0" pageNumber="141">Megagrapha starki</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., male terminalia
|
||
<emphasis bold="true" pageId="0" pageNumber="141">A</emphasis>
|
||
right epandrial lamella
|
||
<emphasis bold="true" pageId="0" pageNumber="141">B</emphasis>
|
||
epandrium with cerci, dorsal
|
||
<emphasis bold="true" pageId="0" pageNumber="141">C</emphasis>
|
||
left epandrial lamella
|
||
<emphasis bold="true" pageId="0" pageNumber="141">D</emphasis>
|
||
detail of phallus. Abbr. hyp: hypandrium; lc: left cercus; lel: left epandrial lamella; ll: lower lobe of left surstylus; ul: upper lobe of left surstylus; rel: right epandrial lamella. Scale bar: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="141">
|
||
<emphasis bold="true" pageId="0" pageNumber="141">Female.</emphasis>
|
||
Unknown. However, all allied species are sexually dimorphic with dark-coloured females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="141" type="remarks">
|
||
<paragraph pageId="0" pageNumber="141">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="141">
|
||
<bibRefCitation author="Stark, A" editor="Floren, A" journalOrPublisher="Bioform Entomology, Nuernberg" pageId="0" pageNumber="141" pagination="105 - 117" refId="B37" refString="Stark, A, 2008. Species of the genus Oedalea Meigen, 1820 (Diptera: Hybotidae): An element of the canopy fauna in European forests? In: Floren, A, Schmidl, J, Eds., Canopy Arthropod Research in Europe. Bioform Entomology, Nuernberg: 105 - 117" title="Species of the genus Oedalea Meigen, 1820 (Diptera: Hybotidae): An element of the canopy fauna in European forests?" volumeTitle="Canopy Arthropod Research in Europe." year="2008">Stark (2008)</bibRefCitation>
|
||
already recognized the species as new and announced a separate paper on
|
||
<taxonomicName class="Insecta" family="Hybotidae" genus="Megagrapha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megagrapha" order="Diptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="141">Megagrapha</emphasis>
|
||
</taxonomicName>
|
||
: "Stark A (2009), in press. Description of two species of the genus
|
||
<taxonomicName class="Insecta" family="Hybotidae" genus="Megagrapha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megagrapha" order="Diptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="141">Megagrapha</emphasis>
|
||
</taxonomicName>
|
||
Melander (
|
||
<taxonomicName kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="141" rank="order">Diptera</taxonomicName>
|
||
,
|
||
<taxonomicName kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="141" rank="superFamily" superFamily="Empidoidea">Empidoidea</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" family="Hybotidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="family">Hybotidae</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Collin" authorityYear="1961" lsidName="" pageId="0" pageNumber="141" rank="tribe" tribe="Drapetini">Drapetini</taxonomicName>
|
||
) from the canopy of trees in European primeval forests. Studia dipterologica 16(2)". However, this paper was never published. Andreas Stark kindly sent us illustrations and locality data and his male was included in the type series.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="141">
|
||
The new species described above is very similar to the Nearctic
|
||
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. exquisita" order="Diptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="exquisita">
|
||
<emphasis italics="true" pageId="0" pageNumber="141">M. exquisita</emphasis>
|
||
</taxonomicName>
|
||
as redescribed by
|
||
<bibRefCitation DOI="https://doi.org/10.4039/Ent1151291-10" author="Chillcott, JG" journalOrPublisher="The Canadian Entomologist" pageId="0" pageNumber="141" pagination="1291 - 1328" refId="B9" refString="Chillcott, JG, Teskey, HJ, 1983. A revision of the NewWorld genera allied to Megagrapha Melander (Diptera: Empididae). The Canadian Entomologist 115 (10): 1291 - 1328, DOI: https://doi.org/10.4039/Ent1151291-10" title="A revision of the NewWorld genera allied to Megagrapha Melander (Diptera: Empididae)." url="https://doi.org/10.4039/Ent1151291-10" volume="115" year="1983">Chillcott and Teskey (1983)</bibRefCitation>
|
||
and by
|
||
<bibRefCitation DOI="https://doi.org/10.4039/Ent90608-10" author="Chillcott, JG" journalOrPublisher="The Canadian Entomologist" pageId="0" pageNumber="141" pagination="608 - 611" refId="B8" refString="Chillcott, JG, 1958. Notes on a rare fly, Megagrapha pubescens (Loew) (Diptera: Empididae). The Canadian Entomologist 90 (10): 608 - 611, DOI: https://doi.org/10.4039/Ent90608-10" title="Notes on a rare fly, Megagrapha pubescens (Loew) (Diptera: Empididae)." url="https://doi.org/10.4039/Ent90608-10" volume="90" year="1958">Chillcott (1958</bibRefCitation>
|
||
- male as
|
||
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. pubescens" order="Diptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="pubescens">
|
||
<emphasis italics="true" pageId="0" pageNumber="141">M. pubescens</emphasis>
|
||
</taxonomicName>
|
||
(Loew, 1862)) in the patterned wings and modified basitarsi of the fore and mid legs. However, both species strikingly differ in the colour of the mesoscutum (reddish yellow in
|
||
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. exquisita" order="Diptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="exquisita">
|
||
<emphasis italics="true" pageId="0" pageNumber="141">M. exquisita</emphasis>
|
||
</taxonomicName>
|
||
- ME, with brown stripes in
|
||
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. starki" order="Diptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="starki">
|
||
<emphasis italics="true" pageId="0" pageNumber="141">M. starki</emphasis>
|
||
</taxonomicName>
|
||
- MS), in the dark pattern on the wing (dark around crossveins of basal cells in ME, light in MS), in the shape of the postpedicel (pear-shaped in ME, with long narrow apical half in MS; Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megagrapha starki sp. nov., male antenna, paratype." figureDoi="10.3897/zookeys.1019.61496.figure4" httpUri="https://binary.pensoft.net/fig/512624" pageId="0" pageNumber="141">4</figureCitation>
|
||
) and in the male terminalia (left surstylus longer and more slender in ME than in MS, with bent tip, and apical bristles on upper lobe of digitiform left surstylus shorter than lobe in ME but longer in MS; Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megagrapha starki sp. nov., male terminalia A right epandrial lamella B epandrium with cerci, dorsal C left epandrial lamella D detail of phallus. Abbr. hyp: hypandrium; lc: left cercus; lel: left epandrial lamella; ll: lower lobe of left surstylus; ul: upper lobe of left surstylus; rel: right epandrial lamella. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1019.61496.figure6" httpUri="https://binary.pensoft.net/fig/512626" pageId="0" pageNumber="141">6</figureCitation>
|
||
).
|
||
<taxonomicName authorityName="Papp & Foeldvari" authorityYear="2001" class="Insecta" family="Hybotidae" genus="Megagrapha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megagrapha europaea" order="Diptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="europaea">
|
||
<emphasis italics="true" pageId="0" pageNumber="141">Megagrapha europaea</emphasis>
|
||
</taxonomicName>
|
||
Papp &
|
||
<normalizedToken originalValue="Földvári">Foeldvari</normalizedToken>
|
||
, 2001, the only other known European representative of the genus, has clear wings in the male and mesoscutum yellow without dark stripes. For further differences, see comments under
|
||
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. europaea" order="Diptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="europaea">
|
||
<emphasis italics="true" pageId="0" pageNumber="141">M. europaea</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="141" type="distribution">
|
||
<paragraph pageId="0" pageNumber="141">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="141">Slovakia, Poland, European Russia.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |