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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1077.67081" ID-GBIF-Dataset="66b827c0-65ac-44a6-90e0-b78cd9ef81c2" ID-PMC="PMC8692307" ID-Pensoft-Pub="1313-2970-1077-1" ID-Pensoft-UUID="734088641608531C8E2CC69397B000ED" ID-PubMed="35027858" ID-ZooBank="C28AD65A76F242CFBED7DFB3702CABCE" ModsDocID="1313-2970-1077-1" checkinTime="1639570129388" checkinUser="pensoft" docAuthor="De Mattia, Willy, Reier, Susanne &amp; Haring, Elisabeth" docDate="2021" docId="1DD47A1A356B5385A4D3DEAA8AA05FA4" docLanguage="en" docName="ZooKeys 1077: 1-175" docOrigin="ZooKeys 1077" docPubDate="2021-12-14" docSource="http://dx.doi.org/10.3897/zookeys.1077.67081" docTitle="Siciliaria tiberii subsp. tiberii tiberii (A. Schmidt 1868" docType="treatment" docVersion="4" id="734088641608531C8E2CC69397B000ED" lastPageNumber="1" masterDocId="734088641608531C8E2CC69397B000ED" masterDocTitle="Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)" masterLastPageNumber="175" masterPageNumber="1" pageNumber="1" updateTime="1668151235153" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>De Mattia, Willy</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0056-467X</mods:nameIdentifier>
<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
<mods:nameIdentifier type="email">willy.demattia@icgeb.org</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reier, Susanne</mods:namePart>
<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Haring, Elisabeth</mods:namePart>
<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-12-14</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1077</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>175</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1077.67081</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1077.67081</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1077-1</mods:identifier>
<mods:identifier type="ZooBank">C28AD65A76F242CFBED7DFB3702CABCE</mods:identifier>
<mods:identifier type="Pensoft-UUID">734088641608531C8E2CC69397B000ED</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="191645975" LSID="urn:lsid:plazi:treatment:1DD47A1A356B5385A4D3DEAA8AA05FA4" httpUri="http://treatment.plazi.org/id/1DD47A1A356B5385A4D3DEAA8AA05FA4" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="1DD47A1A-356B-5385-A4D3-DEAA8AA05FA4" authority="tiberii (A. Schmidt, 1868)" authorityName="tiberii (A. Schmidt" authorityYear="1868" baseAuthorityName="A. Schmidt" baseAuthorityYear="1868" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. tiberii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subSpecies="tiberii">Siciliaria tiberii tiberii (A. Schmidt, 1868)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Wagner's (1913, 1925) and Brandt's (1961) Siciliaria genital drawings A Siciliaria septemplicata, Wagner, 1913: plate 572, fig. 14 B Siciliaria grohmanniana grohmanniana, Wagner, 1925: plate 1, fig. 8 C Siciliaria calcarae calcarae, 1925: plate 3, fig. 25 D Siciliaria ferrox, Brandt, 1961: fig. 1 E overall Siciliaria / Sicania and Sicilian Mauritanica distribution F Siciliaria / Sicania and Sicilian Mauritanica distribution of taxa." figureDoi="10.3897/zookeys.1077.67081.figure1" httpUri="https://binary.pensoft.net/fig/622793" pageId="0" pageNumber="1">Figs 1.F, 25.1-25.2, 26.1-26.5, 29.1-29.3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="A. Schmidt" authorityYear="1868" class="Gastropoda" family="Clausiliidae" genus="Clausilia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Clausilia tiberii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tiberii">Clausilia tiberii</taxonomicName>
<bibRefCitation author="Schmidt, A" journalOrPublisher="Ruthenica, Supplement" pageId="0" pageNumber="1" refId="B89" refString="Schmidt, A, 1868. System der europaeischen Clausilien und ihrer naechsten Verwandten. Cassel, 175 pp." title="System der europaeischen Clausilien und ihrer naechsten Verwandten. Cassel, 175 pp." year="1868">Schmidt 1868</bibRefCitation>
: 40.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Paulucci" authorityYear="1878" class="Gastropoda" family="Clausiliidae" genus="Clausilia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Clausilia tiberiana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tiberiana">Clausilia tiberiana</taxonomicName>
<bibRefCitation author="Benoit, L" journalOrPublisher="Bullettino della Societa Malacologica Italiana" pageId="0" pageNumber="1" refId="B6" refString="Benoit, L, 1876. Illustrazione sistematica critica iconografica de'testacei estramarini della Sicilia ulteriore e delle isole circostanti. Nobile, Napoli, [I-XVI,] 248 pp. [Tav. I-IX, XI-XII]" title="Illustrazione sistematica critica iconografica de'testacei estramarini della Sicilia ulteriore e delle isole circostanti. Nobile, Napoli, [I-XVI,] 248 pp. [Tav. I-IX, XI-XII]" year="1876">Benoit 1876</bibRefCitation>
: 152.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName=", s. l." baseAuthorityName="A. Schmidt" baseAuthorityYear="1868" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tiberii">Siciliaria tiberii</taxonomicName>
-
<bibRefCitation author="Boettger, O" journalOrPublisher="Bullettino della Societa Malacologica Italiana" pageId="0" pageNumber="1" refId="B8" refString="Boettger, O, 1877. Clausilienstudien. Palaeontographica (N. F.) Suppl.3: 1-122. [4 pls. Cassel]" title="Clausilienstudien. Palaeontographica (N. F.) Suppl. 3: 1 - 122. [4 pls. Cassel]" year="1877">Boettger 1877</bibRefCitation>
: 33.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName=", s. l." baseAuthorityName="A. Schmidt" baseAuthorityYear="1868" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria (Siciliaria) tiberii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tiberii" subGenus="Siciliaria">Siciliaria (Siciliaria) tiberii</taxonomicName>
-
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" refId="B60" refString="Nordsieck, H, 1979. Zur Anatomie und Systematik der Clausilien, XXI. Das System der Clausilien, II: Die rezenten europaeischen Clausilien. Archiv fuer Molluskenkunde 109(4/6): 249-275." title="Zur Anatomie und Systematik der Clausilien, XXI. Das System der Clausilien, II: Die rezenten europaeischen Clausilien. Archiv fuer Molluskenkunde 109 (4 / 6): 249 - 275." year="1979">Nordsieck 1979</bibRefCitation>
: 259.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="A. Schmidt" baseAuthorityYear="1868" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria tiberii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tiberii">Charpentieria tiberii</taxonomicName>
-
<bibRefCitation DOI="https://doi.org/10.1127/arch.moll/133/2004/185" author="Beckmann, KH" journalOrPublisher="Basteria" pageId="0" pageNumber="1" refId="B4" refString="Beckmann, KH, 2004. Zur Verbreitung der endemischen nordwestsizilianischen Clausiliidae der Untergattung Charpentieria (Siciliaria) mit Beschreibung von zwei neuen Unterarten (Gastropoda: Stylommatophora: Clausiliidae). Archiv fuer Molluskenkunde 133(1/2): 185-191. https://doi.org/10.1127/arch.moll/133/2004/185" title="Zur Verbreitung der endemischen nordwestsizilianischen Clausiliidae der Untergattung Charpentieria (Siciliaria) mit Beschreibung von zwei neuen Unterarten (Gastropoda: Stylommatophora: Clausiliidae). Archiv fuer Molluskenkunde 133 (1 / 2): 185 - 191." url="https://doi.org/10.1127/arch.moll/133/2004/185" year="2004">Beckmann 2004</bibRefCitation>
: 188.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authority="tiberii" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria (Siciliaria) tiberii subsp. tiberii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subGenus="Siciliaria" subSpecies="tiberii">Charpentieria (Siciliaria) tiberii tiberii</taxonomicName>
-
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" refId="B65" refString="Nordsieck, H, 2007. Worldwide Door Snails (Clausiliidae), recent and fossil. Conchbooks Hackenheim, 214 pp." title="Worldwide Door Snails (Clausiliidae), recent and fossil. Conchbooks Hackenheim, 214 pp." year="2007">Nordsieck 2007</bibRefCitation>
: 54.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName=", s. l." baseAuthorityName="A. Schmidt" baseAuthorityYear="1868" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tiberii">Siciliaria tiberii</taxonomicName>
-
<bibRefCitation author="Welter-Schultes, F" journalOrPublisher="Planet Poster Editions, Goettingen" pageId="0" pageNumber="1" refId="B101" refString="Welter-Schultes, F, 2012. European non-marine molluscs, a quide for species identification. Planet Poster Editions, Goettingen" title="European non-marine molluscs, a quide for species identification." year="2012">Welter-Schultes 2012</bibRefCitation>
: 343.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName=", s. l." baseAuthorityName="A. Schmidt" baseAuthorityYear="1868" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tiberii">Siciliaria tiberii</taxonomicName>
-
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" publicationUrl="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" refId="B69" refString="Nordsieck, H, 2013b. Revisory remarks on the species of Siciliaria Vest from N. W. Sicily. http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" title="Revisory remarks on the species of Siciliaria Vest from N. W. Sicily." url="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" year="2013 b">Nordsieck 2013b</bibRefCitation>
: 8.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="A. Schmidt" baseAuthorityYear="1868" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria tiberii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tiberii">Charpentieria tiberii</taxonomicName>
-
<bibRefCitation author="De Mattia, W" journalOrPublisher="Molecular Ecology Resources" pageId="0" pageNumber="1" publicationUrl="https://www.iucnredlist.org/species/171154/1322070" refId="B19" refString="De Mattia, W, 2017f. Charpentieria tiberii. The IUCN Red List of Threatened Species. https://www.iucnredlist.org/species/171154/1322070" title="Charpentieria tiberii. The IUCN Red List of Threatened Species." url="https://www.iucnredlist.org/species/171154/1322070" year="2017 f">De Mattia 2017f</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Specimens examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation accessionNumber="MW758919, MW757116, MW757117, MW758920" collectingDate="2007-12-16" collectorName="Lab ID, W. De Mattia, J. Macor" country="Italy" elevation="30" latitude="38.13863" location="Capo Rama" longLatPrecision="1" longitude="13.053892" municipality="Terrasini" specimenCount="1" stateProvince="Sicily">
<collectingCountry name="Italy">Italy</collectingCountry>
,
<collectingRegion country="Italy" name="Sicilia">Sicily</collectingRegion>
,
<collectingMunicipality>Terrasini</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:1DD47A1A356B5385A4D3DEAA8AA05FA4:36BA2C1693084CB6D652635936FC5BBA" country="Italy" latitude="38.13863" longLatPrecision="1" longitude="13.053892" municipality="Terrasini" name="Capo Rama" stateProvince="Sicily">Capo Rama</location>
,
<elevation metricMagnitude="1" metricUnit="m" metricValue="3.0" unit="m" value="30.0">
<quantity metricMagnitude="1" metricUnit="m" metricValue="3.0" unit="m" value="30.0">30 m</quantity>
asl
</elevation>
,
<geoCoordinate degrees="38" direction="north" minutes="08" orientation="latitude" precision="1" seconds="19.06" value="38.13863">38°08'19.06&quot;N</geoCoordinate>
,
<geoCoordinate degrees="13" direction="east" minutes="03" orientation="longitude" precision="1" seconds="14.01" value="13.053892">13°03'14.01&quot;E</geoCoordinate>
, [
<collectorName>Lab ID</collectorName>
19_1, COI:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MW758919">MW758919</accessionNumber>
, ITS2:
<accessionNumber isEnumeration="true">MW757116, MW757117</accessionNumber>
;
<collectorName>Lab ID</collectorName>
19_2, COI:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MW758920">MW758920</accessionNumber>
],
<collectorName>W. De Mattia</collectorName>
and
<collectorName>J. Macor</collectorName>
leg.,
<collectingDate value="2007-12-16">16.xii.2007</collectingDate>
. 3 dissected spm
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="shell">
<paragraph pageId="0" pageNumber="1">Shell</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
(Figs
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Siciliaria tiberii tiberii (A. Schmidt, 1868), Capo Rama, Terrasini 26.1 shell 26.2 clausiliar plate double side 26.3 shell 26.4 detail of the aperture 26.5 clausiliar plate double side. Siciliaria tiberii alcamoensis Brandt, 1961 comb. nov., Piano Margi, Carini 26.6 shell 26.7 clausiliar plate double side 26.8 detail of the aperture 26.9 shell." figureDoi="10.3897/zookeys.1077.67081.figure26" httpUri="https://binary.pensoft.net/fig/622818" pageId="0" pageNumber="1">26</figureCitation>
.1-26.5, 29.1-29.3).
</emphasis>
Shell decollate; whorls ribbed; dorsal keel prominent; inferior lamella high to very high; anterior upper palatal plica strong and long, separated from the lunella; sometimes the anterior upper palatal plica presents a distal knob, sometimes another lower weak upper palatal plica is present close to the ALPP; anterior lower upper palatal plica strong and long; palatal edge of clausilium plate distally not receding, palatal edge distally more or less strongly bent upwards.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements</paragraph>
<paragraph pageId="0" pageNumber="1">
(n = 40, decollate): shell height 16.3
<normalizedToken originalValue="±">+/-</normalizedToken>
1.2, whorl width 4.0
<normalizedToken originalValue="±">+/-</normalizedToken>
0.1, aperture height 3.1
<normalizedToken originalValue="±">+/-</normalizedToken>
0.2, aperture width 2.3
<normalizedToken originalValue="±">+/-</normalizedToken>
0.2.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="external morphology of the genital organs">
<paragraph pageId="0" pageNumber="1">External morphology of the genital organs</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
(Fig.
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Siciliaria tiberii tiberii (A. Schmidt, 1868), Capo Rama, Terrasini 25.1 whole distal genital organs 25.2 internal distal part of genital organs. Siciliaria tiberii alcamoensis Brandt, 1961 comb. nov., Piano Margi, Carini 25.3 whole distal genital organs 25.4 internal distal part of genital organs 25.5 detail of the pseudopapilla." figureDoi="10.3897/zookeys.1077.67081.figure25" httpUri="https://binary.pensoft.net/fig/622817" pageId="0" pageNumber="1">25</figureCitation>
.1).
</emphasis>
The FO is longer than the V (FO/V range 2.0-2.3). The VD is thin along its whole course except for the proximal part where it is larger in diameter. The FDBC is longer than the BC+SDBC (FDBC/BC+SDBC range 1.5-1.7). The BC+SDBC is club-like to cylindrical and slightly longer than the V (BC+SDBC/V range 1.2-1.5), with no clear distinction between the SDBC and the BC. The apex is big and rounded. The D is slightly longer than the V (D/V range 1.7-1.9) and slightly longer that the BC+SDBC (D/BC+SDBC range 1.1-1.5), thinner than the BC+SDBC and with a small and round apex. The V is big and cylindrical. The A is large and long. The PC is longer than the V (P+E/V range 2.9-3.4). The PR is very short and robust. The ET is clearly visible. The E is longer than the P (E/P range 1.1-1.7) and gradually shrinking and turning into the VD.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="internal morphology of the genital organs">
<paragraph pageId="0" pageNumber="1">Internal morphology of the genital organs</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
(Fig.
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Siciliaria tiberii tiberii (A. Schmidt, 1868), Capo Rama, Terrasini 25.1 whole distal genital organs 25.2 internal distal part of genital organs. Siciliaria tiberii alcamoensis Brandt, 1961 comb. nov., Piano Margi, Carini 25.3 whole distal genital organs 25.4 internal distal part of genital organs 25.5 detail of the pseudopapilla." figureDoi="10.3897/zookeys.1077.67081.figure25" httpUri="https://binary.pensoft.net/fig/622817" pageId="0" pageNumber="1">25</figureCitation>
.2).
</emphasis>
The A shows few fleshy weak irregular folds. The P presents a set of transverse smooth pleats. The pleats are slightly irregular and tend to fade towards the A. The penial wall is smooth. The PP is big, elongated and markedly wrinkled with a pointed apex. It originates from one transverse epiphallar ring. The E shows two main fringed pleats that run as far as the VD. The V shows a weak irregular pattern of smooth pleats.
</paragraph>
<caption doi="10.3897/zookeys.1077.67081.figure24" httpUri="https://binary.pensoft.net/fig/622816" pageId="0" pageNumber="1" start="Figure 24" startId="F24">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 24.</emphasis>
<taxonomicName authority="calcarae" authorityName="calcarae (Philippi" authorityYear="1844" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria calcarae subsp. calcarae" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="calcarae" subSpecies="calcarae">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria calcarae calcarae</emphasis>
</taxonomicName>
(Philippi, 1844), Visicari, Castellammare del Golfo
<emphasis bold="true" pageId="0" pageNumber="1">24.1</emphasis>
palatal plicae
<emphasis bold="true" pageId="0" pageNumber="1">24.2</emphasis>
parietal lamellae.
<taxonomicName baseAuthorityName="Brandt" baseAuthorityYear="1961" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria calcarae subsp. belliemii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="calcarae" subSpecies="belliemii">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria calcarae belliemii</emphasis>
</taxonomicName>
(Brandt, 1961), Monte Belliemi, Partinico
<emphasis bold="true" pageId="0" pageNumber="1">24.3</emphasis>
palatal plicae.
<taxonomicName baseAuthorityName="Brandt" baseAuthorityYear="1961" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria calcarae subsp. belliemii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="calcarae" subSpecies="belliemii">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria calcarae belliemii</emphasis>
</taxonomicName>
(Brandt, 1961), Castello di Calatubo, Alcamo
<emphasis bold="true" pageId="0" pageNumber="1">24.4</emphasis>
palatal plicae.
<taxonomicName authorityName="De Mattia &amp; Reier &amp; Haring" authorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria calcarae subsp. borgettensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="calcarae" subSpecies="borgettensis">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria calcarae borgettensis</emphasis>
</taxonomicName>
ssp. nov., road to Romitello, Borghetto
<emphasis bold="true" pageId="0" pageNumber="1">24.5-24.6</emphasis>
palatal plicae.
<taxonomicName authorityName="De Mattia &amp; Reier &amp; Haring" authorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria calcarae subsp. jatinensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="calcarae" subSpecies="jatinensis">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria calcarae jatinensis</emphasis>
</taxonomicName>
ssp. nov. San Giuseppe Jato
<emphasis bold="true" pageId="0" pageNumber="1">24.7</emphasis>
palatal plicae
<emphasis bold="true" pageId="0" pageNumber="1">24.8</emphasis>
parietal lamellae.
<taxonomicName authorityName="De Mattia &amp; Reier &amp; Haring" authorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria calcarae subsp. parajatinensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="calcarae" subSpecies="parajatinensis">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria calcarae parajatinensis</emphasis>
</taxonomicName>
ssp. nov., west side Monte Kumeta
<emphasis bold="true" pageId="0" pageNumber="1">24.9-24.10</emphasis>
palatal plicae.
<taxonomicName authorityName="Liberto, Reitano, Giglio, Colomba &amp; Sparacio" authorityYear="2016" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria calcarae subsp. orlandoi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="calcarae" subSpecies="orlandoi">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria calcarae orlandoi</emphasis>
</taxonomicName>
Liberto, Reitano, Giglio, Colomba &amp; Sparacio, 2016, Bosco Ficuzza
<emphasis bold="true" pageId="0" pageNumber="1">24.11</emphasis>
palatal plicae
<emphasis bold="true" pageId="0" pageNumber="1">24.12</emphasis>
parietal lamellae.
<taxonomicName authorityName="De Mattia &amp; Reier &amp; Haring" authorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria calcarae subsp. cruenta" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="calcarae" subSpecies="cruenta">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria calcarae cruenta</emphasis>
</taxonomicName>
ssp. nov., Monte Gibilmesi
<emphasis bold="true" pageId="0" pageNumber="1">24.13-24.14</emphasis>
palatal plicae.
<taxonomicName authorityName="Brandt" authorityYear="1961" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria ferrox" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ferrox">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria ferrox</emphasis>
</taxonomicName>
(Brandt, 1961). Trabia
<normalizedToken originalValue="Sant’Onofrio">Sant'Onofrio</normalizedToken>
<emphasis bold="true" pageId="0" pageNumber="1">24.15</emphasis>
palatal plicae
<emphasis bold="true" pageId="0" pageNumber="1">24.16</emphasis>
parietal lamellae.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="ecology">
<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authority="tiberii" authorityName="tiberii (A. Schmidt" authorityYear="1868" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. tiberii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subSpecies="tiberii">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria tiberii tiberii</emphasis>
</taxonomicName>
is found under stones, among scree, rocky debris and on dry stonewalls along the coastline, on a limestone plateau. According to
<bibRefCitation author="De Mattia, W" journalOrPublisher="Molecular Ecology Resources" pageId="0" pageNumber="1" publicationUrl="https://www.iucnredlist.org/species/171154/1322070" refId="B19" refString="De Mattia, W, 2017f. Charpentieria tiberii. The IUCN Red List of Threatened Species. https://www.iucnredlist.org/species/171154/1322070" title="Charpentieria tiberii. The IUCN Red List of Threatened Species." url="https://www.iucnredlist.org/species/171154/1322070" year="2017 f">De Mattia (2017f)</bibRefCitation>
, the taxon is Near Threatened.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation DOI="https://doi.org/10.1127/arch.moll/133/2004/185" author="Beckmann, KH" journalOrPublisher="Basteria" pageId="0" pageNumber="1" refId="B4" refString="Beckmann, KH, 2004. Zur Verbreitung der endemischen nordwestsizilianischen Clausiliidae der Untergattung Charpentieria (Siciliaria) mit Beschreibung von zwei neuen Unterarten (Gastropoda: Stylommatophora: Clausiliidae). Archiv fuer Molluskenkunde 133(1/2): 185-191. https://doi.org/10.1127/arch.moll/133/2004/185" title="Zur Verbreitung der endemischen nordwestsizilianischen Clausiliidae der Untergattung Charpentieria (Siciliaria) mit Beschreibung von zwei neuen Unterarten (Gastropoda: Stylommatophora: Clausiliidae). Archiv fuer Molluskenkunde 133 (1 / 2): 185 - 191." url="https://doi.org/10.1127/arch.moll/133/2004/185" year="2004">Beckmann (2004</bibRefCitation>
: 188), following
<normalizedToken originalValue="Nordsieck’s">Nordsieck's</normalizedToken>
personal communications cites
<taxonomicName authority="tiberii" authorityName="tiberii (A. Schmidt" authorityYear="1868" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. tiberii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subSpecies="tiberii">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria tiberii tiberii</emphasis>
</taxonomicName>
from a southern area surrounding San Cipirello (Madonna dei Furi, Monte Palmeto, Monte della Fiera, San Giuseppe Jato and Monte della Scala). As previously stated, this identification revealed to be wrong.
</paragraph>
<paragraph pageId="0" pageNumber="1">
As a result, the actual distributional range of
<taxonomicName authority="tiberii" authorityName="tiberii (A. Schmidt" authorityYear="1868" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. tiberii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subSpecies="tiberii">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria tiberii tiberii</emphasis>
</taxonomicName>
seems to be limited to the area from San Cataldo (Partinico) in the south to Monte Palmeto and Capo Rama (Terrasini) in the north to the northern slopes of Monte Pecoraro. Further field research is needed in order to define its actual distribution. Recent findings of Diego Viola (Trieste, Italy) from a locality between San Giuseppe Jato and Piana degli Albanesi revealed the presence of a population of
<taxonomicName authorityName="Vest" authorityYear="1867" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria</emphasis>
</taxonomicName>
cfr.
<taxonomicName authorityName=", s. l." baseAuthorityName="A. Schmidt" baseAuthorityYear="1868" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tiberii">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria tiberii</emphasis>
</taxonomicName>
that clearly differ, as a shell, from
<taxonomicName lsidName="S. calcarae subsp. jatinensis" pageId="0" pageNumber="1" rank="subSpecies" species="calcarae" subSpecies="jatinensis">
<emphasis italics="true" pageId="0" pageNumber="1">S. calcarae jatinensis</emphasis>
</taxonomicName>
ssp. nov. and
<taxonomicName lsidName="S. calcarae subsp. parajatinensis" pageId="0" pageNumber="1" rank="subSpecies" species="calcarae" subSpecies="parajatinensis">
<emphasis italics="true" pageId="0" pageNumber="1">S. calcarae parajatinensis</emphasis>
</taxonomicName>
ssp. nov.(personal communication). The poor samples available (few damaged shells) did not allow us a thorough investigation.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
Monte San Calogero in the east (cited by Nordsieck as also included in the
<taxonomicName lsidName="S. tiberii" pageId="0" pageNumber="1" rank="species" species="tiberii">
<emphasis italics="true" pageId="0" pageNumber="1">S. tiberii</emphasis>
</taxonomicName>
distribution), is an isolate limestone massif 10 km SE of Termini Imerese, 50 km east of San Cipirello and 65 km SSE of Capo Rama. The presence of
<taxonomicName authority="tiberii" lsidName="S. tiberii subsp. tiberii" pageId="0" pageNumber="1" rank="subSpecies" species="tiberii" subSpecies="tiberii">
<emphasis italics="true" pageId="0" pageNumber="1">S. tiberii tiberii</emphasis>
</taxonomicName>
in this locality, due to its remarkable distance, should be confirmed by further field research and not only inferred from museum samples. Mislabelling or wrong locality assignments could have played a role in this regard (as for
<taxonomicName authorityName="De Mattia &amp; Reier &amp; Haring" authorityYear="2021" baseAuthorityName="De Mattia &amp; Reier &amp; Haring" baseAuthorityYear="1961" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria septemplicata subsp. alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="septemplicata" subSpecies="alcamoensis">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria septemplicata alcamoensis</emphasis>
</taxonomicName>
, see below). The diagnosis provided by
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" publicationUrl="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" refId="B69" refString="Nordsieck, H, 2013b. Revisory remarks on the species of Siciliaria Vest from N. W. Sicily. http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" title="Revisory remarks on the species of Siciliaria Vest from N. W. Sicily." url="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" year="2013 b">Nordsieck (2013b</bibRefCitation>
: 8) &quot;Shell not or only in part decollate; dorsal keel less strong&quot; imply that it might represent another population of
<taxonomicName lsidName="S. calcarae" pageId="0" pageNumber="1" rank="species" species="calcarae">
<emphasis italics="true" pageId="0" pageNumber="1">S. calcarae</emphasis>
</taxonomicName>
s. l., yet more field and taxonomical research is needed.
<taxonomicName authorityName="Paulucci" authorityYear="1878" class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia tiberiana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tiberiana">
<emphasis italics="true" pageId="0" pageNumber="1">Clausilia tiberiana</emphasis>
</taxonomicName>
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.13317" author="Paulucci, M" journalOrPublisher="Paris, [1, I-IV = 1 - 4]" pageId="0" pageNumber="1" refId="B72" refString="Paulucci, M, 1878. Materiaux pour servir a l'etude de la faune malacologique terrestre et fluviatile de l'Italie et de ses iles. Paris, [1, I-IV = 1-4], DOI: https://doi.org/10.5962/bhl.title.13317" title="Materiaux pour servir a l'etude de la faune malacologique terrestre et fluviatile de l'Italie et de ses iles." url="https://doi.org/10.5962/bhl.title.13317" year="1878">Paulucci 1878</bibRefCitation>
is deemed as junior synonym of
<taxonomicName authority="tiberii" authorityName="tiberii (A. Schmidt" authorityYear="1868" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. tiberii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subSpecies="tiberii">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria tiberii tiberii</emphasis>
</taxonomicName>
by
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" publicationUrl="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" refId="B69" refString="Nordsieck, H, 2013b. Revisory remarks on the species of Siciliaria Vest from N. W. Sicily. http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" title="Revisory remarks on the species of Siciliaria Vest from N. W. Sicily." url="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" year="2013 b">Nordsieck (2013b</bibRefCitation>
: 8). The authorship is probably wrong as
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.13317" author="Paulucci, M" journalOrPublisher="Paris, [1, I-IV = 1 - 4]" pageId="0" pageNumber="1" refId="B72" refString="Paulucci, M, 1878. Materiaux pour servir a l'etude de la faune malacologique terrestre et fluviatile de l'Italie et de ses iles. Paris, [1, I-IV = 1-4], DOI: https://doi.org/10.5962/bhl.title.13317" title="Materiaux pour servir a l'etude de la faune malacologique terrestre et fluviatile de l'Italie et de ses iles." url="https://doi.org/10.5962/bhl.title.13317" year="1878">Paulucci (1878</bibRefCitation>
: 13) cites
<taxonomicName authorityName="Paulucci" authorityYear="1878" class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia tiberiana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tiberiana">
<emphasis italics="true" pageId="0" pageNumber="1">Clausilia tiberiana</emphasis>
</taxonomicName>
Benoit:
<bibRefCitation author="Benoit, L" journalOrPublisher="Bullettino della Societa Malacologica Italiana" pageId="0" pageNumber="1" pagination="129 - 163" refId="B5" refString="Benoit, L, 1875. Catalogo delle conchiglie terrestri e fluviatili della Sicilia e delle isole circostanti. Bullettino della Societa Malacologica Italiana 1 (2): 129 - 163" title="Catalogo delle conchiglie terrestri e fluviatili della Sicilia e delle isole circostanti." volume="1" year="1875">Benoit (1875</bibRefCitation>
: 152) introduced the name
<taxonomicName authorityName="Paulucci" authorityYear="1878" class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia tiberiana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tiberiana">
<emphasis italics="true" pageId="0" pageNumber="1">Clausilia tiberiana</emphasis>
</taxonomicName>
as a &quot;Elegante e bella specie
<normalizedToken originalValue="de’">de'</normalizedToken>
dintorni di S. Giuseppe delle Mortelle contrada non lungi da Palermo&quot; and depicted this taxon in 1876 (Table VI, fig. 7).
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.51778" author="Benoit, L" journalOrPublisher="Bullettino della Societa Malacologica Italiana" pageId="0" pageNumber="1" refId="B7" refString="Benoit, L, 1881. Nuovo catalogo delle conchiglie terrestri e fluviatili della Sicilia. Atti della Reale Accademia Peloritana 3: 43-216. [Sep.: I-VI, 7-176]. https://doi.org/10.5962/bhl.title.51778" title="Nuovo catalogo delle conchiglie terrestri e fluviatili della Sicilia. Atti della Reale Accademia Peloritana 3: 43 - 216. [Sep.: I-VI, 7 - 176]." url="https://doi.org/10.5962/bhl.title.51778" year="1881">Benoit (1881</bibRefCitation>
: 107) stated that again that &quot;Questa bellissima e grande conchiglia, non conosciuta dagli autori siciliani vive nella contrada detta S. Giuseppe delle Mortelle, non lungi da Palermo&quot;. Despite intensive collecting in the surrounding of Palermo by one of the authors (WDM), we were unable to identify the position of this old toponym (also by asking locals).
</paragraph>
</subSubSection>
</treatment>
</document>
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